identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B887D4AC47167FFC92F909FD76FDC7.text	03B887D4AC47167FFC92F909FD76FDC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemiphyllodactylus xiengkhouangensis Luu & Sitthivong & Hoang & Ha & Ha & Xayasith & Ziegler & Grismer 2025	<div><p>Hemiphyllodactylus xiengkhouangensis sp. nov.</p><p>Xiengkhouang Slender Gecko (Figs. 5, 6).</p><p>Holotype. Adult male VNUF R.2025.02 (Field no. KNGAN.XK.02) collected on 19 May 2023 at 19:45 hrs by Saly Sitthivong at the Phouvieng Village, Khuon District, Xiengkhouang Province, Laos (19°14'39"N 103°35'47"E; 1180 m in elevation).</p><p>Paratypes. Adult female VNUF R.2025.03 (Field no. KNGAN.XK.03 and adult male NUOL R.2025.01 (Field no. KNGAN.XK.01) bear the same collection data as the holotype.</p><p>Diagnosis. Hemiphyllodactylus xiengkhouangensis sp. nov. can be separated from all other species of Hemiphyllodactylus by possessing the unique combination of having a maximum SVL of 41.7 mm; chin scales 8–10; postmentals enlarged; circumnasal scales three or four; intersupranasals (=postrostrals) two or three; supralabials 9–12; infralabials 8–11; dorsal scales 16–20; ventral scales 9–11; subdigital lamellae on the first finger four or five; femoral and precloacal pores 20 or 21, in a continuous row in males and 12 pitted scales in the female; no plate - like subcaudal scales; dorsal surfaces of head, body, and limbs light-brown, heavily mottled with darker markings; top of head overlain with a faint, irregular dark pattern; dark stripe extends from the external nares through the eye to the region just posterior of forelimb insertion on the body; ventral surfaces of the trunk and limbs uniformly pale-yellow with darker lateral areas; ventral surface of original tail orange with ventrolaterally dark pigmentation. Three selective color pattern characters (DorBodPatrn, VenBodPatrn, and VenTalPatrn as mentioned above) are scored across closely related Hemiphyllodactylus species in Tables 3 and 4.</p><p>Description of holotype. Adult male, SVL 41.7 mm; triangularly shaped head in dorsal profile, depressed, distinct from neck; lores flat; rostrum moderate in length (SnEye/SVL 0.11); prefrontal region slightly concave; canthal ridge gently rounded, indistinct; snout moderate, dorsally rounded; eye large; ear opening elliptical, small; eye to ear distance greater than diameter of eye; rostral rectangular, wider than high, shortly divided dorsally by straight rostral groove, bordered by enlarge supra - nasals posteriorly; two intersupranasals (=postnasals); external nares bordered anteriorly by rostral, dorsally by supranasal, posteriorly by two postnasals, ventrally by first supralabial; rectangular supralabials 11/12 (R, L); rectangular infralabials 11 (R, L); scales on rostrum, lores, top of head, and occiput small, granular with rostrum scales largest and slightly raised; superciliaries dorsally flat, mostly square, subimbricate, largest anteriorly; mental triangular, bordered laterally by first infralabials, posteriorly by two enlarged postmentals; each postmental in contact with first infralabial; chin scales nine; gular scales small, subimbricate, grading posteriorly into weakly larger, subimbricate throat.</p><p>Body elongate (Trunk/SVL 0.50), dorsoventrally compressed; dorsal scales small, granular, 20 at midbody contained within one eye diameter; ventral scales flat, subimbricate much larger than dorsal scales, 11 at midbody ventral within the length of eye diameter; precloacal scales slightly larger than abdominal scales; continuous femoral and precloacal pore-bearing scales 20; enlarged tubercle on anterior margin of hemipenial swelling single; forelimbs short and robust, dorsally and ventrally covered with flat, subimbricate scales; palmar scales flat, subimbricate; all digits except digit I well - developed; digit I vestigial, clawless bearing five subdigital lamellae; distal subdigital lamellae of digits II–V undivided, angular and U - shaped, lamellae proximal to these transversely expanded; distal lamellar formula of digits II–V 4 - 4 - 4 - 4 (left); claws on digits II–V well-developed, unsheathed; distal portions of digits strongly curved, terminal joint free, arising from central portion of lamellar pad; hindlimbs short, more robust than forelimbs, covered with flat, juxtaposed scales dorsally and larger, flat subimbricate scales ventrally; plantar scales low, flat, subimbricate; all digits except digit I well - developed; digit I vestigial, clawless bearing five subdigital lamellae; distal subdigital lamellae of digits II–V undivided, angular and U - shaped, lamellae proximal to these transversely expanded; distal lamellar formula of digits II–V 4 - 5 - 5 - 5 (left); claws on digits II–V welldeveloped, unsheathed; distal portions of digits strongly curved, terminal joint free, arising from central portion of lamellar pad; tail partially regenerated, 36 mm in length; caudal scales non-whorled; dorsal caudal scales of original tail larger than dorsal body scales, flat, subcycloid, subimbricate; subcaudals weakly larger than dorsal caudals, not plate - like. Morphological characters are presented in Table 9.</p><p>Coloration in life. Dorsal surfaces of head, body, and limbs light-brown, heavily mottled with darker markings; top of head overlain with a faint, irregular dark pattern; dark stripe extends from the external nares through the eye to the region just posterior of forelimb insertion on the body; eye large with a vertical pupil and coppery brown iris speckled with fine black reticulations; paravertebral region with poorly defined dark blotches continuing from nape to tail base transforming into a distinct, black, post - sacral band; cream post - sacral marking immediately posterior to black post - sacral band; bearing light-colored, anteriorly projecting arms; dorsolateral region of trunk with diffuse dark speckling and scattered light spots on the lateral side; upper body with numerous brown spots irregularly distributed; limbs bearing irregularly shaped, dark markings, especially on the hindlimbs; original portion of tail with alternating unclearly dark and light-brown bands; regenerated portion tail grey with scattered dark flecks; gular region pale-yellow to cream; ventral surface of the trunk uniformly pale-yellow with darker lateral areas; ventral surface of limbs; ventral surface of original tail distinctly orange with laterally dark pigmentation; subcaudal of regenerated tail portion grey with dark markings.</p><p>Variation (Figs 5, 6). The dorsal pattern on the head, body, limbs, and tail of the female paratype (VNUF R.2025.03) is faded or nearly absent, whereas the male paratype (NUOL R.2025.01) is darker brown with a more faded pattern compared to the holotype. The ventral surface of the tail of the paratype is uniformly orange, lacking dark lateral spots. Variation in scale counts and morphomeric data are presented in Table 9.</p><p>......continued on the next page</p><p>Distribution. Hemiphyllodactylus xiengkhouangensis sp. nov. is only known from the type locality at Phouvieng Village, Khuon District, Xiengkhouang Province, Laos (Fig. 1).</p><p>Natural History. Hemiphyllodactylus xiengkhouangensis sp. nov. is an upland species, and all individuals were collected from Phouvieng Village at night from 19:00–20:30 hrs, on garden vegetation and houses, at heights of 0.5–1.5 meters above the ground (Fig. 7). The female paratype (VNUF R.2025.03) has two eggs.</p><p>Etymology. The specific name is derived from the type locality of Xiengkhouang Province, Laos.</p><p>Comparisons. Hemiphyllodactylus xiengkhouangensis sp. nov. is the sister species to H. banaensis with uncorrected pairwise sequence divergence of 11.4% (Table 2). H. xiengkhouangensis sp. nov. differs significantly from H. banaensis in mean values of SVL (1.59 vs. 1.69, p = 0.028), Trunk (1.3 vs. 1.36, p &lt;0.001) and HeadW (0.84 vs. 0.87, p = 0.004), and the presence of a cream-colored venter (vs. dark brown in H. banaensis), and the presence of orange coloration in the subcaudal region of the original tail (vs. dark brown H. banaensis). Morphologically, H. xiengkhouangensis sp. nov. (n = 3) is very similar to H. serpispecus (n = 1) in terms of morphometric data. However, the new species can be distinguished from H. serpispecus by the significantly lower mean number of dorsal scales (DS: 18.33 vs. 26; p &lt;0.001), the presence of a dark grey dorsum with irregular zig-zag markings (vs. a chevron pattern in H. serpispecus), and an orange venter of the original tail (vs. cream-colored in H. serpispecus). Raw morphometric, meristic, and categorical data used in the statistical analyses are presented in Tables 3 and 4. Statistically significant differences between Hemiphyllodactylus xiengkhouangensis sp. nov. and other closely related species are provided in Tables 5 and 7.</p></div>	https://treatment.plazi.org/id/03B887D4AC47167FFC92F909FD76FDC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Luu, Vinh Quang;Sitthivong, Saly;Hoang, Tuoi Thi;Ha, Hong Bich;Ha, Huan Van;Xayasith, Santi;Ziegler, Thomas;Grismer, L. Lee	Luu, Vinh Quang, Sitthivong, Saly, Hoang, Tuoi Thi, Ha, Hong Bich, Ha, Huan Van, Xayasith, Santi, Ziegler, Thomas, Grismer, L. Lee (2025): Two more new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Laos and Vietnam identified using an integrative taxonomic approach. Zootaxa 5686 (2): 211-238, DOI: 10.11646/zootaxa.5686.2.3, URL: https://doi.org/10.11646/zootaxa.5686.2.3
03B887D4AC5D1673FC92FD5EFE22FEE7.text	03B887D4AC5D1673FC92FD5EFE22FEE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemiphyllodactylus yenchauensis Luu & Sitthivong & Hoang & Ha & Ha & Xayasith & Ziegler & Grismer 2025	<div><p>Hemiphyllodactylus yenchauensis sp. nov.</p><p>Yenchau Slender Gecko (Fig. 8).</p><p>Holotype. Adult male VNUF R.2025.05 (Field no. Sonla05) collected on 04 October 2023 at 21:10 hrs by Vinh Q. Luu, Le P. Duc, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.401&amp;materialsCitation.latitude=20.861444" title="Search Plazi for locations around (long 104.401/lat 20.861444)">Vilay Phimpasone at the entrance of karst cave of Pa Kha 1 Village, Chieng Tuong Commune</a>, Yen Chau District, Son La Province, Vietnam (20°51'41.2"N 104°24'03.6"E; 1228 m in elevation).</p><p>Paratypes. Subadult male VNUF R.2025.06 (Field no. Sonla06) bears the same collection data as the holotype .</p><p>Diagnosis. Hemiphyllodactylus yenchauensis sp. nov. can be distinguished from all other species of Hemiphyllodactylus by possessing the unique combination of having a maximum SVL of 39.1 mm; chin scales 7 – 10; postmentals enlarged; circumnasal scales two or three; intersupranasals (=postrostrals) two; supralabials 10 or 11; infralabials 10 or 11; dorsal scales 21 and ventral scales 11 or 12; subdigital lamellae on the first finger four or five; subdigital lamellae on the first toe five or six; no femoral or precloacal pores in males; no plate - like subcaudal scales; dorsal surfaces of head, body, and limbs light brown with irregular dark blotches and marbling throughout; dark and light pre and postorbital stripes in parallel extends from posterior nares to forelimb insertion; dorsolateral light - colored spots on trunk; paravertebral region with a series of dark, irregular blotches from nape to base of tail; forming a chevron pattern along the midbody; ventral surface of trunk pale yellowish to cream with faint darker margins; ventral surface of original tail bright orange with ventrolaterally dark pigmentation. Three selective color pattern characters (DorBodPatrn, VenBodPatrn, and VenTalPatrn as montioned above) are scored across closely related Hemiphyllodactylus species in Tables 3 and 4.</p><p>Description of holotype. Adult male SVL 39.1 mm; triangularly shaped head in dorsal view, depressed, distinguish from neck; lores flat; rostrum moderate in length (SnEye/SVL 0.09); prefrontal region slightly concave; canthus rostralis smoothly rounded, barely discernable; snout moderate, rounded when viewed from above; eye large; ear opening small and elliptical, small; eye to ear distance greater than diameter of eye; rostral wider than high, bordered posteriorly by large supranasals; two intersupranasals; external nares bordered anteriorly by rostral, dorsally by supranasal, posteriorly by two postnasals, ventrally by first supralabial; rectangular supralabials tapering to below posterior margin of eye 10/11 (L, R); rectangular infralabials tapering to below posterior margin of eye 10 (L, R); scales of rostral, loreal, cranial, and occipital scales small, elevated, rostral scales largest; dorsal superciliaries flat, mostly square, subimbricate, largest anteriorly; mental triangular, bordered laterally by first infralabials and posteriorly by two enlarged postmentals; each postmental in contact with first infralabial and bordered laterally by a smaller sublabial; chin scales seven; gular scales small, subimbricate, grading posteriorly into slightly larger, subimbricate throat and even larger pectoral scales which grade into slightly larger, subimbricate ventrals.</p><p>Body somewhat elongate (Trunk/SVL 0.53), vertically flattened body shape; dorsal scales small, granular, dorsal scales at midbody contained within one eye diameter 21; ventral scales flat, subimbricate much larger than dorsal scales, ventral scales contained within one eye diameter 11; precloacal scales slightly larger than abdominal scales; no pore-bearing femoral or precloacal scales; enlarged tubercle on lateral margin of tail base single; forelimbs short, robust in stature, covered with flat, subimbricate scales dorsally and ventrally; palmar scales slightly raised, subimbricate; all digits except digit I well - developed; digit I vestigial, clawless bearing five subdigital lamellae; distal subdigital lamellae of digits II–V undivided, angular and U - shaped, lamellae proximal to these transversely expanded; distal lamellar formula of digits II–V (left) 4-5-5-4; claws on digits II–V well developed, unsheathed; distal portions of digits strongly curved, terminal joint free, arising from central portion of lamellar pad; hind limbs short, more robust than forelimbs, covered with slightly raised, juxtaposed scales dorsally and by larger, flat subimbricate scales anteriorly and ventrally; plantar scales slightly raised, subimbricate; all digits except digit I well - developed; digit I vestigial, clawless bearing five subdigital lamellae; distal subdigital lamellae of digits II–V undivided, angular and U - shaped, lamellae proximal to these transversely expanded; distal lamellar formula of digits II–V (left) 4-5-5-4; bearing five subdigital lamellae claws on digits II–V well - developed, unsheathed; distal portions of digits strongly curved, terminal joint free, arising from central portion of lamellar pad; tail partially regenerated, 30.2 mm in length; caudal scales in ring-like whorls; dorsal caudal scales larger than dorsal body scales, flat, nearly circular, slightly overlapping; ventrolateral caudals slightly enlarged, weak anterior flare, fringelike appearance; subcaudals flat, moderately larger than dorsal caudals, non-plate-like. Morphological characters are presented in Table 9.</p><p>Coloration in life (Fig. 8). Dorsal surfaces of head, body, and limbs light brown with irregular, dark blotching and marbling throughout; top of head with faint, dark markings forming a Y-shaped pattern; parallel dark and white stripes from posterior nares, through eye, to forelimb insertion; eye large with vertical pupil and coppery-brown iris interspersed with black reticulations; paravertebral region with a series of dark, irregular blotches from nape to base of tail; forming a chevron pattern along the midbody, followed by a distinct black post-sacral band; cream post-sacral marking present immediately behind the black band; bearing faint, light-colored, anteriorly projecting arms; dorsolateral region of trunk with diffuse dark flecking and scattered light spots laterally; upper body with numerous light brown spots irregularly arranged; limbs mottled with irregular dark markings; original portion of tail with alternating, poorly defined dark and light brown bands; regenerated portion of tail grey-brown with dark markings and flecks extending to the tip; gular region pale pinkish-cream; ventral surface of trunk pale yellowish to cream with faint darker margins; ventral surface of limbs similar in tone to trunk; ventral surface of original tail bright orange with lateral dark pigmentation; subcaudal area of regenerated tail light grey with scattered dark markings.</p><p>Variation (Fig. 8). The color patterns of the paratype generally match that of the holotype, though the dark dorsal markings are less bold and narrower than those of the holotype. Variation in morphometric and meristic data are presented in Table 9.</p><p>Distribution. Hemiphyllodactylus yenchauensis sp. nov. is known only from Pa Kha 1 Village, Chieng Tuong Commune, Yen Chau District, Son La Province, Vietnam (Fig. 1).</p><p>Natural History. Hemiphyllodactylus yenchauensis sp. nov. is an upland species and the type series was collected at night between 20:00 and 20:30 hrs on trees approximately 0.5 m above the ground at the entrance of a karstic cave (Fig. 9). The surrounding habitat consisted of disturbed forest, with the lower area converted to agricultural land (Fig. 10).</p><p>Etymology. The specific epithet is a toponym derived from the type locality of Yen Chau District, Son La Province, Vietnam.</p><p>Comparisons. The molecular analyses indicate that Hemiphyllodactylus yenchauensis sp. nov. is the sister species to H. bonkowskii + H. vanhoensis (Fig. 2) with mean uncorrected pairwise sequence divergences of 4.7% and 4.8%, respectively (Table 2). However, H. yenchauensis sp. nov. differs significantly from H. bonkowskii in mean values of Trunk (1.26 vs. 1.34, p &lt;0.001), HeadW (0.8 vs. 0.93, p &lt;0.001), SnEye (0.53 vs. 0.67, p &lt;0.001), NarEye (0.43 vs. 0.52, p &lt;0.001), EyeD (0.35 vs. 0.45, p &lt;0.006), SnW (0.16 vs. 0.31, p &lt;0.001), VS (11.5 vs. 14.33, p = 0.022), DS (21 vs. 25.17, p = 0.009), the presence of a chevron pattern on the trunk (vs. a yellowish-grey dorsal trunk pattern in H. bonkowskii), and an orange venter of the original tail (vs. cream-colored in H. bonkowskii); differs from H. vanhoensis in mean values of Trunk (1.26 vs. 1.34, p &lt;0.001), HeadL (1.00 vs. 1.04, p = 0.005), HeadW (0.8 vs. 0.86, p &lt;0.001), NarEye (0.43 vs. 0.50, p = 0.018), CN (2.5 vs. 5.17, p &lt;0.001), and the presence of a chevron pattern on the trunk (vs. semi-reticulate dorsal markings in H. vanhoensis), and an orange venter of the original tail (vs. dull yellow with black and white spots in H. vanhoensis). Raw morphometric, meristic, and categorical data used in the statistical analyses are provided in Tables 3 and 4. Statistically significant and discrete categorical differences between H. yenchauensis sp. nov. and all other closely related species are presented in Tables 6 and 7.</p></div>	https://treatment.plazi.org/id/03B887D4AC5D1673FC92FD5EFE22FEE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Luu, Vinh Quang;Sitthivong, Saly;Hoang, Tuoi Thi;Ha, Hong Bich;Ha, Huan Van;Xayasith, Santi;Ziegler, Thomas;Grismer, L. Lee	Luu, Vinh Quang, Sitthivong, Saly, Hoang, Tuoi Thi, Ha, Hong Bich, Ha, Huan Van, Xayasith, Santi, Ziegler, Thomas, Grismer, L. Lee (2025): Two more new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Laos and Vietnam identified using an integrative taxonomic approach. Zootaxa 5686 (2): 211-238, DOI: 10.11646/zootaxa.5686.2.3, URL: https://doi.org/10.11646/zootaxa.5686.2.3
