identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B8879FFFB3FFB4AED6A5B1DDD7F8D1.text	03B8879FFFB3FFB4AED6A5B1DDD7F8D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Indicopleustes Distant 1908	<div><p>Indicopleustes Distant, 1908</p><p>Indicopleustes Distant, 1908: 25; Funkhouser, 1927: 48; Goding, 1939: 337; Funkhouser, 1951: 233; Metcalf &amp; Wade, 1965: 106; Yuan &amp; Chou, 2002: 199; Wallace &amp; Deitz, 2004: 245.</p><p>Formocentrus Kato, 1928: 223 .</p><p>Type species. Indicopleustes albomaculatus Distant, 1908</p><p>Diagnosis. Medium to large sized. Black or brown with conspicuous covering of setae. Face nearly quadrangular; vertex with dorsal margins slightly arcuate, and ventral margins wave-shaped or slightly oblique. Ocelli ovoid or ballshaped, closer to inner margin of eyes than to each other or equidistant. Frontoclypeal margins parallel or roundly convergent; frontoclypeal lobes distinct. Humeral angles triangular with apices somewhat blunt. Metopidium slightly convex. Suprahumeral horns stout at base, short or long, apex sharp or truncate, forming triangle or rectangle in the dorsal view. Posterior pronotal process strongly elevated above scutellum at base; undulate in profile, with single or double curve; apex sharp and extended beyond forewing anal angle or resting at anal angle. Scutellum entirely exposed, base elevated; posterior margin emarginate, apex flat or slightly convex. Forewing with brown spots or uniform coloration and veins with setae, basal one-fifth to one-half with opaque sclerotization, veins M and Cu fused basally to approximately one-fifth of wing length then strongly divergent, veins M+Cu and R fused basally, with 1 m-cu, 1 r-m, and 1 s crossveins. Hindwing vein R branched into R 1, R 2+3, and R 4+5, vein M branched into M 1+2 and M 3+4, R 4+5 and M 1+2 veins not fused (four apical cells present), 1 r-m and 1 m-cu crossveins present, apical limbus broad, with wrinkles.</p><p>Distribution. China, Indonesia, Sri Lanka (Fig. 1).</p><p>Checklist and distributions of species of Indicopleustes Distant, 1908</p><p>I. albomaculatus Distant, 1908; Sri Lanka (Peradeniya)</p><p>I. apicatus Melichar, 1914; Indonesia (Java)</p><p>I. curvatus (Melichar, 1903); Sri Lanka (Weligama)</p><p>I. fuscomaculatus Kato, 1928; China (Taiwan)</p><p>I. zizhongi Li, Chen &amp; Yang, sp. nov.; China (Guangxi, Hainan)</p><p>I. typicus (Kato, 1928); China (Taiwan)</p><p>I. hainanensis Yuan, 2002; China (Hainan)</p><p>I. yunnanensis Yuan, 2002; China (Guangdong, Guangxi, Yunnan)</p><p>Key to species of Indicopleustes Distant, 1908</p><p>1 Suprahumeral horns with apices truncate............................................................... 2</p><p>– Suprahumeral horns with apices sharp or blunt........................................................... 3</p><p>2 Apex of posterior pronotal process extended to 2A vein of forewing (Kato 1928: fig. 6a–c; fig. 9A–C)............................................................................................... I. typicus (Kato, 1928)</p><p>– Apex of posterior pronotal process extended nearly to M 3+4 vein of forewing (fig. 4)........ I. curvatus (Melichar, 1903)</p><p>3 Suprahumeral horns with apices blunt (Distant 1908: fig. 23; fig. 2A)................. I. albomaculatus Distant, 1908</p><p>– Suprahumeral horns with apices sharp.................................................................. 4</p><p>4 Posterior pronotal process with single undulation in profile................................................. 5</p><p>– Posterior pronotal process with a double undulation in profile............................................... 7</p><p>5 Suprahumeral horns extended horizontally in anterior view (figs 6E, F).......... I. zizhongi Li, Chen &amp; Yang, sp. nov.</p><p>– Suprahumeral horns extended diagonally upward......................................................... 6</p><p>6 Apex of posterior pronotal process upturned, slightly divergent from margins of forewing (Yuan &amp; Chou 2002: fig. 85; fig. 13)......................................................................... I. yunnanensis Yuan, 2002</p><p>– Apex of posterior pronotal process extended straight to anal angle of forewing (Yuan &amp; Chou 2002: fig. 84; fig. 10)................................................................................. I. hainanensis Yuan, 2002</p><p>7 Posterior pronotal process in profile with two undulations of uniform amplitude; suprahumeral horns short and extended horizontally in anterior view (Melichar 1914: fig. 7, 7a; fig. 3A)......................... I. apicatus Melichar, 1914</p><p>– Posterior pronotal process in profile with amplitude of the first undulation considerably more than that of second; suprahumeral horns long and extended diagonally upward (Kato 1928: fig. 5a–b; fig. 5A–B)................................................................................................. I. fuscomaculatus Kato, 1928</p></div>	https://treatment.plazi.org/id/03B8879FFFB3FFB4AED6A5B1DDD7F8D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Li, Fenge;Long, Jiankun;Chang, Zhimin;Chen, Xiangsheng;Yang, Lin	Li, Fenge, Long, Jiankun, Chang, Zhimin, Chen, Xiangsheng, Yang, Lin (2026): Review of the genus Indicopleustes Distant, 1908 (Hemiptera, Membracidae, Centrotinae) with description of a new species from China. Zootaxa 5856 (1): 145-162, DOI: 10.11646/zootaxa.5856.1.22, URL: https://doi.org/10.11646/zootaxa.5856.1.22
03B8879FFFB0FFB5AED6A028D8A2F8C0.text	03B8879FFFB0FFB5AED6A028D8A2F8C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Indicopleustes albomaculatus Distant 1908	<div><p>Indicopleustes albomaculatus Distant, 1908</p><p>(Figure 2)</p><p>Indicopleustes albomaculatus Distant, 1908: 25; Goding, 1939: 337.</p><p>Indicopleustes albomaculata Metcalf &amp; Wade, 1965: 106 .</p><p>Description. Coloration. General color pale brown.</p><p>Head and thorax. Ocelli closer to inner margins of eyes than to each other (Fig. 2B). Frontoclypeal margins roundly convergent; frontoclypeal lobes slender (Fig. 2B). Suprahumeral horns nearly triangular in lateral view (Fig. 2A). Posterior pronotal process with single-wave profile in basal half, remaining half slightly curved, without contacting forewing (Fig. 2A).</p><p>Male genitalia. Lateral plate with fingerlike dorsoapical lobe nearly as long as base; base triangular (Fig. 2D). Style clasp oriented laterally, triangular, with basal thickening, apex blunt (Fig. 2E).</p><p>Female genitalia. Valvulae II (Fig. 2F) knife-shaped, distal half of dorsal margin finely serrated, proximal tooth particularly prominent; slightly broadened distally.</p><p>Distribution. Sri Lanka (Peradeniya).</p><p>Note. The present interpretation of the species is based on photographs published by Wallace and Dietz (2004).</p></div>	https://treatment.plazi.org/id/03B8879FFFB0FFB5AED6A028D8A2F8C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Li, Fenge;Long, Jiankun;Chang, Zhimin;Chen, Xiangsheng;Yang, Lin	Li, Fenge, Long, Jiankun, Chang, Zhimin, Chen, Xiangsheng, Yang, Lin (2026): Review of the genus Indicopleustes Distant, 1908 (Hemiptera, Membracidae, Centrotinae) with description of a new species from China. Zootaxa 5856 (1): 145-162, DOI: 10.11646/zootaxa.5856.1.22, URL: https://doi.org/10.11646/zootaxa.5856.1.22
03B8879FFFB1FFB2AED6A01DD929F832.text	03B8879FFFB1FFB2AED6A01DD929F832.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Indicopleustes apicatus Melichar 1914	<div><p>Indicopleustes apicatus Melichar, 1914</p><p>(Figure 3)</p><p>Indicopleustes apicatus Melichar, 1914: 113 .</p><p>Indicopleustes apicata Metcalf &amp; Wade, 1965: 106 .</p><p>Description. Coloration. General color dark brown, eyes dark yellow, ocelli pale yellow. Thorax dark with white pubescence. Forewing most dark brown, apical third yellowish-brown, white spot present at anal angle; veins dark brown. Legs pale brown, middle or proximal part of tibia yellowish (Fig. 3A–B).</p><p>Head and thorax. Ocelli slightly closer to inner margins of eyes than to each other (Fig. 3A). Frontoclypeal margins long, extended beyond ventral margins of vertex, apex truncate; frontoclypeal lobes relatively wide (Fig. 3A). Suprahumeral horns thick basally, apices flat; in dorsal view base nearly rectangular, apex acute (Fig. 3A, C). Posterior pronotal process with obvious double-wave shape in profile, posterior wave is shallower, apex even with middle of fifth apical cells of forewing (Fig. 3B).</p><p>Distribution. Indonesia (Java).</p><p>Note. The present interpretation of the species is based on photographs of the holotype provided by the Moravské Zemské Muzeum.</p></div>	https://treatment.plazi.org/id/03B8879FFFB1FFB2AED6A01DD929F832	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Li, Fenge;Long, Jiankun;Chang, Zhimin;Chen, Xiangsheng;Yang, Lin	Li, Fenge, Long, Jiankun, Chang, Zhimin, Chen, Xiangsheng, Yang, Lin (2026): Review of the genus Indicopleustes Distant, 1908 (Hemiptera, Membracidae, Centrotinae) with description of a new species from China. Zootaxa 5856 (1): 145-162, DOI: 10.11646/zootaxa.5856.1.22, URL: https://doi.org/10.11646/zootaxa.5856.1.22
03B8879FFFB7FFB0AED6A7DDD929FF3F.text	03B8879FFFB7FFB0AED6A7DDD929FF3F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Indicopleustes curvatus (Melichar 1903)	<div><p>Indicopleustes curvatus (Melichar, 1903)</p><p>(Figure 4)</p><p>Centrotus curvatus Melichar, 1903: 112; Distant, 1908: 26</p><p>Indicopleustes curvata Metcalf &amp; Wade, 1965: 107 .</p><p>Description. Coloration. General color brown with yellow setae (Fig. 4A–C). Head dark brown. Ocelli pale yellow. Eyes yellowish brown. Margins of humeral angles and suprahumeral horns black. Forewing dark brown hyaline. Thorax dark with white pubescence. Legs yellowish brown. Abdomen brown.</p><p>Head and thorax. Ocelli closer to inner margins of eyes than to each other (Fig. 4A). Frontoclypeal margins roundly tapered, distal two-thirds extended beyond ventral margins of vertex; frontoclypeal lobes narrow (Fig. 4A). Suprahumeral horns nearly rectangular in dorsal view; anterior margins slightly shorter than posterior margins (Fig. 4A, C). Posterior pronotal process with obvious double-wave profile, posterior wave slightly shallower, apex slightly divergent from hind margin of the forewing (Fig. 4B).</p><p>Distribution. Sri Lanka (Weligama).</p><p>Note. The present interpretation of the species is based on photographs of the holotype provided by the Moravské Zemské Muzeum.</p></div>	https://treatment.plazi.org/id/03B8879FFFB7FFB0AED6A7DDD929FF3F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Li, Fenge;Long, Jiankun;Chang, Zhimin;Chen, Xiangsheng;Yang, Lin	Li, Fenge, Long, Jiankun, Chang, Zhimin, Chen, Xiangsheng, Yang, Lin (2026): Review of the genus Indicopleustes Distant, 1908 (Hemiptera, Membracidae, Centrotinae) with description of a new species from China. Zootaxa 5856 (1): 145-162, DOI: 10.11646/zootaxa.5856.1.22, URL: https://doi.org/10.11646/zootaxa.5856.1.22
03B8879FFFB4FFB0AED6A600DC6CFC80.text	03B8879FFFB4FFB0AED6A600DC6CFC80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Indicopleustes fuscomaculatus Kato 1928	<div><p>Indicopleustes fuscomaculatus Kato, 1928</p><p>(Figure 5A–B)</p><p>Indicopleustes fuscomaculatus Kato, 1928: 223 .</p><p>Indicopleustes fuscomaculata Metcalf &amp; Wade, 1965: 107 .</p><p>Indicopleustes reticulatus Emmrich, 1973: 267; Mckamey, 1998: 93.</p><p>Description. Coloration. General color brown reddish brown except suprahumeral horns black, and tibia yellowish brown. Eyes yellowish brown. Thorax black with white pubescence. Forewing smoky-yellow.</p><p>Head and thorax. In lateral view suprahumeral horns narrowly triangular; positioned distinctly dorsal to posterior pronotal process. (Fig. 5A). Posterior pronotal process with double-wave profile, amplitude of first wave considerably larger than that of second (Fig. 5B).</p><p>Distribution. China (Taiwan).</p><p>Note. The present interpretation of the species is based on photographs of the specimens provided by The University Museum, The University of Tokyo and illustrations from Kato (1928).</p></div>	https://treatment.plazi.org/id/03B8879FFFB4FFB0AED6A600DC6CFC80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Li, Fenge;Long, Jiankun;Chang, Zhimin;Chen, Xiangsheng;Yang, Lin	Li, Fenge, Long, Jiankun, Chang, Zhimin, Chen, Xiangsheng, Yang, Lin (2026): Review of the genus Indicopleustes Distant, 1908 (Hemiptera, Membracidae, Centrotinae) with description of a new species from China. Zootaxa 5856 (1): 145-162, DOI: 10.11646/zootaxa.5856.1.22, URL: https://doi.org/10.11646/zootaxa.5856.1.22
03B8879FFFB4FFBEAED6A16EDA4FF865.text	03B8879FFFB4FFBEAED6A16EDA4FF865.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Indicopleustes zizhongi Li, Chen & Yang 2026	<div><p>Indicopleustes zizhongi Li, Chen &amp; Yang, sp. nov.</p><p>(Figures 6–8)</p><p>Material examined. Holotype. 1♂, China, Hainan Province, Changjiang Li Autonomous County, Bawangling National Nature Reserve, 19-IV-2017, Liang-Jing Yang leg. Paratypes. 2♀♀, China, Hainan Province, Qiongzhong Li and Miao Autonomous County, Limu Mountain National Forest Park, 4-V-2017, Lan Zhang leg ; 3♀♀ 1♂, China, Hainan Province, Wuzhishan City, Wuzhishan National Nature Reserve, 19-V-2017, Yongshun Ding and Liang-Jing Yang leg ; 1♂, China, Hainan Province, Changjiang Li Autonomous County, Bawangling National Nature Reserve, 5-VI-2019, Yan Jiang leg ; 1♂, China, Hainan Province, Qiongzhong Li and Miao Autonomous County, Limu Mountain National Forest Park, 23-IV-2021, Feng-E Li leg ; 1♀, China, Hainan Province, Baisha Li Autonomous County, Yuanmen Township, Hongkan, 15-V-2021, Feng-E Li leg ; 1♂, Guangxi Zhuang Autonomous Region, Chongzuo City, Longzhou County, Nonggang National Nature Reserve, 12-VII-2018, Yong-Jin Sui leg.</p><p>Description. Body length: male 7.8–7.9 mm (N = 5), female 8.2–8.8 mm (N = 6); forewing length: male 6.8–7.0 mm (N = 5), female 7.1–7.8 mm (N = 6); width between humeral angles: male 3.2–3.6 mm (N = 5), female 3.4–3.6 mm (N = 6); width between suprahumeral horn apices: male 4.6–4.9 mm (N = 5), female 5.3–5.5 mm (N = 6).</p><p>Coloration. General color brown with yellow setae, black in male. Head black (Fig. 6A–F). Ocelli orange. Eyes pale yellow brown with brown border. Suprahumeral horns and posterior pronotal process black, median carina brown. Forewing pale brown subhyaline, veins darker; basal fifth black; anal angle with transparent area flanked by two brown spots, another brown spot near apex (Fig. 6G). Thorax dark with white or yellow pubescence. Trochanter, femur, black brown; tibia and tarsus brown; tarsal claw dark brown. Abdomen black with yellow hairs (Fig. 6B, D).</p><p>Head and thorax. Ocelli equidistant between each other and inner margins of eyes (Fig. 6E–F). Frontoclypeal margins rectangular, apex truncate, distal two-thirds extended beyond ventral margins of vertex; frontoclypeal lobes distinct, margins curved (Fig. 6E–F). Suprahumeral horns extended horizontally to both sides, curved slightly backward in dorsal view; apices sharp (Fig. 6A, C, E–F). Posterior pronotal process with single undulation terminated near anal angle of forewing; apex slightly concave in profile, curved weakly away from margin of forewing in female, flat and touching forewing edge in male (Fig. 6B, D).</p><p>Male genitalia. Pygofer nearly trapezoidal in lateral view, dorsal margin longer than ventral margin (Fig. 7A); sternite IX rectangular shaped (Fig. 7D). Lateral plate with fingerlike dorsoapical lobe extended dorsally, rest ovoid with setae (Fig. 6G). Subgenital plates fused for more than half length ventral view (Fig. 7D). Style clasp oriented laterally, triangular, with basal thickening; connective n-shaped, arms slightly divergent and widened apically (Fig. 7C). Aedeagus in lateral view nearly uncinate, most of surface and margin with retrorse serrations (Fig. 7E); gonopore at apex in posterior view, ovoid, nearly half length of shaft (Fig. 7F).</p><p>Female genitalia. Pygofer in lateral view irregularly shoe-shaped, base acuminate, apex tubular (Fig.8A); in ventral view elliptical (Fig. 8B). Sternite VII in ventral view with posterior margin deeply concave, bearing a triangular membranous area on either side (Fig. 8C). Valvifer I semicircular and thin (Fig. 8D); valvulae I knife-shaped, half of dorsal margin surface, and ventral margin of the apex surface with linear sculpture. Valvifer II shoe-shaped in lateral view (Fig. 8F). Valvulae II (Fig. 8E) knife-shaped, nearly half of dorsal margin serrated, with proximal tooth particularly prominent; slightly broadened distally. Gonoplac (Fig. 8F) with apical half expanded, ventral margin with setae.</p><p>Etymology. The name “ zizhongi ” is in honor of Professor Zi-Zhong Li, a Chinese entomologist who specializes in Hemiptera at Guizhou University.</p><p>Distribution. China (Guangxi, Hainan).</p><p>Remarks. This species is similar to I. hainanensis Yuan, 2002, but differs from the latter as follows: (1) suprahumeral horns extended horizontally (extended dorsolaterally in I. hainanensis); (2) forewing surface without hairlike setae in basal half (surface with hairlike setae in I. hainanensis); (3) forewing pale brown, anal angle with a transparent area small; veins dark (yellowish-brown, the anal angle with triangular transparent band; veins Sc, R, M dark, the rest yellow in I. hainanensis); (4) posterior pronotal process with relatively gentle undulations (undulations slightly larger in I. hainanensis).</p></div>	https://treatment.plazi.org/id/03B8879FFFB4FFBEAED6A16EDA4FF865	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Li, Fenge;Long, Jiankun;Chang, Zhimin;Chen, Xiangsheng;Yang, Lin	Li, Fenge, Long, Jiankun, Chang, Zhimin, Chen, Xiangsheng, Yang, Lin (2026): Review of the genus Indicopleustes Distant, 1908 (Hemiptera, Membracidae, Centrotinae) with description of a new species from China. Zootaxa 5856 (1): 145-162, DOI: 10.11646/zootaxa.5856.1.22, URL: https://doi.org/10.11646/zootaxa.5856.1.22
03B8879FFFBBFFBDAED6A144D965FE32.text	03B8879FFFBBFFBDAED6A144D965FE32.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Indicopleustes typicus (Kato 1928)	<div><p>Indicopleustes typicus (Kato, 1928)</p><p>(Figure 9A–C)</p><p>Formocentrus typicus Kato, 1928: 223 .</p><p>Sipylus typicus Funkhouser, 1943: 271 .</p><p>Indicopleustes typicus Esaki, 1932: 1733 .</p><p>Indicopleustes typica Metcalf &amp; Wade, 1965: 107 .</p><p>Indicopleustes katoi Yuan &amp; Chou, 2002: 199; Dmitriev, 2022.</p><p>Indicopleustes esakii Dmitriev &amp; Deitz, 2017: 95; Dmitriev, 2022.</p><p>Description. Coloration. General color brown to orange-brown. Head dark brown. Eyes dark brown. Forewing pale black hyaline marked with many brownish spots (Fig. 9C).</p><p>Head and thorax. In lateral view, suprahumeral horns short and shape, with truncated apices in the dorsal view (Fig. 9B–C). Posterior pronotal process with single-wave profile of low amplitude; apex touching forewing margin (Fig. 9C).</p><p>Distribution. China (Taiwan).</p><p>Note. The present interpretation of the species is based on photographs of the specimens provided by The University Museum, The University of Tokyo and illustrations from Kato (1928).</p><p>Initially placed by Kato in Formocentrus in 1928, the species was transferred to Sipylus later that year (Kato, 1928; Funkhouser, 1943). Following further study, it was eventually placed in Indicopleustes (Yuan &amp; Chou, 2002; Dmitriev, 2022). During this period, the species was recorded by Esaki (1932), with illustrations included in the publication. Subsequently, Metcalf and Wade (1965) incorrectly attributed Indicopleustes typicus to Esaki, 1932 rather than Kato, 1928, creating an objective homonymy and leading Dmitriev &amp; Deitz (2017) to propose a replacement name, Indicopleustes esakii Dmitriev &amp; Deitz, 2017 . This name is a synonym of Indicopleustes typicus (Kato, 1928) .</p></div>	https://treatment.plazi.org/id/03B8879FFFBBFFBDAED6A144D965FE32	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Li, Fenge;Long, Jiankun;Chang, Zhimin;Chen, Xiangsheng;Yang, Lin	Li, Fenge, Long, Jiankun, Chang, Zhimin, Chen, Xiangsheng, Yang, Lin (2026): Review of the genus Indicopleustes Distant, 1908 (Hemiptera, Membracidae, Centrotinae) with description of a new species from China. Zootaxa 5856 (1): 145-162, DOI: 10.11646/zootaxa.5856.1.22, URL: https://doi.org/10.11646/zootaxa.5856.1.22
03B8879FFFB9FFBBAED6A572DAC0FD0B.text	03B8879FFFB9FFBBAED6A572DAC0FD0B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Indicopleustes hainanensis Yuan & Chou 2002	<div><p>Indicopleustes hainanensis Yuan, 2002</p><p>(Figures 10–12)</p><p>Indicopleustes hainanensis Yuan &amp; Chou, 2002: 196 .</p><p>Material examined. 1♂, China, Hainan Province, Baisha Li Autonomous County, Yinggeling National Nature Reserve, 10-IV-2017, Hong-Xing Li leg ; 1♀, China, Hainan Province, Baisha Li Autonomous County, Yinggeling National Nature Reserve, 16-V-2021, Feng-E Li leg ; 1♀ 1♂, China, Hainan Province, Leduo Li Autonomous County, Jianfengling National Nature Reserve, 16-V-2021, Nian Gong and Feng-E Li leg.</p><p>Description. Body length: male 7.7–7.9 mm (N = 2), female 7.9–8.2 mm (N = 2); forewing length: male 6.7–6.9 mm (N = 2), female 6.9–7.2 mm (N = 2); width between humeral angle apices: male 3.0– 3.2 mm (N = 2), female 3.1–3.3 mm (N = 2); width between suprahumeral horn apices: male 3.2–3.4 mm (N = 2), female 3.2–3.5 mm (N = 2).</p><p>Coloration. General coloration brownish-gray with yellow setae, except suprahumeral horns and lateral carina of posterior pronotal process black. Head black (Fig. 10A–C). Ocelli pale yellow; eyes deep burgundy. Forewing smoky-yellow with transparent wide triangular band at anal angle, narrow triangle dark brown spot posterad of anal angle; veins brown with dark brown on few (r-m, base of R bifurcation, Cu 1, m-cu) (Fig. 10B, D). Thorax black with yellow pubescence; coxae, trochanter, femur mostly black; apices of femur, tibia and tarsus brown, claws dark brown.</p><p>Head and thorax. Ocelli equidistant to each other and inner margins of eyes. Frontoclypeal margins rectangular, extended beyond ventral margins of vertex for most of length; frontoclypeal lobes relatively broad, apices extended to middle of frontoclypeal margins (Fig. 10A). Suprahumeral horns triangular, apices sharp; extended diagonally upward, apices not surpassing humeral angles (Fig. 10A–C). Posterior pronotal process with single undulating wave, apex extended straight to anal angle of forewing (Fig. 10B).</p><p>Male genitalia. Pygofer irregularly rectangular in lateral view, dorsal margin longer than ventral margin (Fig. 11A); sternite IX rectangular, with margins slightly curved (Fig. 11B). Lateral plate with fingerlike dorsoapical lobe extended dorsally, apex blunt (Fig. 11G). Subgenital plates fused in basal half in ventral view (Fig. 11B). Style clasp oriented laterally, triangular, with basal thickening, with more than ten setae; connective n-shaped (Fig. 11C). Aedeagus in lateral view nearly uncinate, most of surface and margin with retrorse serrations; gonopore at apex of shaft in posterior view, ovoid, nearly half length of shaft (Fig. 11E–F).</p><p>Female genitalia. Pygofer in lateral view irregularly shoe-shaped, base acuminate, apex tubular (Fig. 12A); in ventral view ovoid (Fig. 12B). Sternite VII in ventral view wide and flat, with posterior margin concave, bearing triangular membranous area on either side (Fig. 12C). Valvulae I knife-shaped, curved slightly upward at ends, more than half of dorsal margin surface with linear sculpture. Valvifer II shoe-shaped in lateral view (Fig. 12F). Valvulae II (Fig. 12E) knife-shaped, nearly half of dorsal margin serrated, distal two-thirds slightly broadened. Gonoplac with apical half expanded, ventral margin with setae (Fig. 12F).</p><p>Distribution. China (Guangxi, Hainan).</p></div>	https://treatment.plazi.org/id/03B8879FFFB9FFBBAED6A572DAC0FD0B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Li, Fenge;Long, Jiankun;Chang, Zhimin;Chen, Xiangsheng;Yang, Lin	Li, Fenge, Long, Jiankun, Chang, Zhimin, Chen, Xiangsheng, Yang, Lin (2026): Review of the genus Indicopleustes Distant, 1908 (Hemiptera, Membracidae, Centrotinae) with description of a new species from China. Zootaxa 5856 (1): 145-162, DOI: 10.11646/zootaxa.5856.1.22, URL: https://doi.org/10.11646/zootaxa.5856.1.22
03B8879FFFBCFFA6AED6A7DDDB54FE4F.text	03B8879FFFBCFFA6AED6A7DDDB54FE4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Indicopleustes yunnanensis Yuan & Chou 2002	<div><p>Indicopleustes yunnanensis Yuan, 2002</p><p>(Figures 13–15)</p><p>Indicopleustes yunnanensis Yuan &amp; Chou, 2002: 197 .</p><p>Material examined. 1♂, China, Guangdong Province, Huizhou City, Longmen County, Nankunshan National Forest Park , 22-V-2021, Baoxiang Zhan leg ; 1♀, China, Guangxi Zhuang Autonomous Region, Hechi City, Huanjiang Maonan Autonomous County, Mulun Township, 8-VII-2019, Xiao-Ya Wang leg.</p><p>Description. Body length: male 7.3 mm (N = 1), female 8.6 mm (N = 1); forewing length: male 6.2 mm (N = 1), female 7.6 mm (N = 1); width between humeral angles apices: male 3.0 mm (N = 1), female 3.5 mm (N = 1); width between suprahumeral horns apices: male 3.2mm (N = 1), female 3.8 mm (N = 1).</p><p>Coloration. General coloration black with yellow setae (Fig. 13A–C). Ocelli pale yellow hyaline; eyes yellowish-gray with yellowish-brown border. Forewing smoky-yellow with transparent wide triangular band at anal angle, small triangular dark brown spot posterad of anal angle on apical limbus; veins brown with dark brown on a few (r-m, base of R bifurcation, R 1, R 2+3, R 3+4, m-cu). Thorax black with yellow and white pubescence; leg most black, except tibia brown, claw dark brown.</p><p>Head and thorax. Ocelli equidistant between each other and inner margins of eyes (Fig. 13A). Frontoclypeal margins roundly tapered, distal two-thirds extended beyond ventral margins of the vertex; frontoclypeal lobes narrow, apices extended to near middle of frontoclypeal margins (Fig. 13A). Suprahumeral horns triangular, apices sharp; extended diagonally upward, apices not surpassing suprahumeral angles (Fig. 13A–C). Posterior pronotal process with single undulating wave, apex extended posterodorsad and terminating above anal angle of forewing (Fig. 13B).</p><p>Male genitalia. Pygofer irregularly trapezoidal in lateral view, dorsal margin longer than ventral margin, slightly concave (Fig. 14A); sternite IX rectangular shaped, ventral margin slightly concave. Lateral plate with fingerlike dorsoapical lobe extended dorsally, apex triangular (Fig. 14G). Basal half of subgenital plates fused in ventral view (Fig. 14D). Style clasp oriented laterally, triangular, with basal thickening, with more than ten setae; connective n-shaped (Fig. 14C). Aedeagus in lateral view nearly uncinate, most of surface and margin with retrorse serrations; gonopore at the apex of shaft in posterior view, ovoid, one-third length of shaft; margin slightly concave ar middle in posterior view (Fig. 14E–F).</p><p>Female genitalia. Pygofer in lateral view irregularly shoe-shaped, base acuminate, apex tubular (Fig. 15A); ovoid in ventral view (Fig. 15B). Sternite VII in ventral view with posterior margin deeply concave, bearing triangular membranous area on either side (Fig. 15C). Valvifer I semicircular, and thin (Fig. 15D); valvulae I knife-shaped, curved slightly upward at ends, more than half of dorsal margin with linear sculpture. Valvifer II shoe-shaped in lateral view (Fig. 15F). Valvulae II (Fig. 15E) knife-shaped, nearly half of dorsal margin serrated, distal two-thirds slightly broadened. Gonoplac with apical half expanded, ventral margin with setae (Fig. 15F).</p><p>Distribution. China (Guangdong, Guangxi, Yunnan).</p></div>	https://treatment.plazi.org/id/03B8879FFFBCFFA6AED6A7DDDB54FE4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Li, Fenge;Long, Jiankun;Chang, Zhimin;Chen, Xiangsheng;Yang, Lin	Li, Fenge, Long, Jiankun, Chang, Zhimin, Chen, Xiangsheng, Yang, Lin (2026): Review of the genus Indicopleustes Distant, 1908 (Hemiptera, Membracidae, Centrotinae) with description of a new species from China. Zootaxa 5856 (1): 145-162, DOI: 10.11646/zootaxa.5856.1.22, URL: https://doi.org/10.11646/zootaxa.5856.1.22
