identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03BA0C3EBE3C533B47ADFDD5FA79FC6B.text	03BA0C3EBE3C533B47ADFDD5FA79FC6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eusphalerum contortispinum Zanetti 1998	<div><p>Eusphalerum contortispinum Zanetti, 1998</p><p>(Fig. 1)</p><p>Eusphalerum contortispinum Zanetti, 1998: 136 .</p><p>Material. TAIWAN: NANTOU: 4 ♂♂, 5 ♀♀: ‘TAIWAN: NANTOU county Ren`ai township, Yuanfeng . 24°07'04'' 121°14'13''E, 2760 m, 14.IV.2025, beating. M. Geiser &amp; C.F. Lee leg. BMNH{E} 2025-22.’ (BMNH).</p><p>Remarks. Eusphalerum contortispinum (Fig. 1) was originally described based on the holotype (male) from “ Taiwan Chiai Hsien Yushan N.P. Ta-Ta Ghia”. Females for these species were unknown. I dissected three females and did not find sclerotized accessory sclerites. Specimens from Nantou were collected in subalpine scrubland with Rhododendron and small coniferous trees by beating a little tree. It is here recorded from Nantou for the first time.</p></div>	https://treatment.plazi.org/id/03BA0C3EBE3C533B47ADFDD5FA79FC6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2026): Three new species and new records of the genus Eusphalerum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Eusphalerini) from Taiwan. Zootaxa 5768 (3): 419-426, DOI: 10.11646/zootaxa.5768.3.8, URL: https://doi.org/10.11646/zootaxa.5768.3.8
03BA0C3EBE3C533B47ADFC06FB88FA9A.text	03BA0C3EBE3C533B47ADFC06FB88FA9A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eusphalerum dubitatum Zanetti 1998	<div><p>Eusphalerum dubitatum Zanetti, 1998</p><p>(Fig. 2)</p><p>Eusphalerum dubitatum Zanetti, 1998: 124 .</p><p>Material. TAIWAN: PINGTUNG: 1 ♂, 2 ♀♀: ‘TAIWAN: PINGTUNG county Chunjih (Chunri) township, Tahanshan, Jinshuiying trail 24°24'27'' 120°45'24''E, 1450 m, 11.IV.2025, M. Geiser, M. Fikáček, B.H. Ho, G.J. Phang leg. BMNH {E} 2025-22.’ (BMNH).</p><p>Remarks. Eusphalerum dubitatum (Fig. 2) was originally described based on a male and a female from “ Taiwan Nantou Hsien Nenkaoshan Tenchi Hut”. Specimens from Pingtung were collected by beating flowering Rhododendron and Maesa perlaria . It is here recorded from Pingtung for the first time.</p></div>	https://treatment.plazi.org/id/03BA0C3EBE3C533B47ADFC06FB88FA9A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2026): Three new species and new records of the genus Eusphalerum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Eusphalerini) from Taiwan. Zootaxa 5768 (3): 419-426, DOI: 10.11646/zootaxa.5768.3.8, URL: https://doi.org/10.11646/zootaxa.5768.3.8
03BA0C3EBE3C533947ADFA48FF28F9A5.text	03BA0C3EBE3C533947ADFA48FF28F9A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eusphalerum geiseri Shavrin 2026	<div><p>Eusphalerum geiseri sp. nov.</p><p>(Figs 3, 7–8, 13)</p><p>Type material. TAIWAN: NANTOU: Holotype ♂ (dissected; Fig. 3): ‘ TAIWAN: NANTOU county | Hsinyi township, Tatachia, | Lulin sacred tree, 16.IV.2025 | 23°28'55'' 120°51'17''E, | 2350-2400m. M. Geiser, C.F. | Lee, Y. Hsiao &amp; Y.T. Chung leg. | BMNH {E} 2025-22.’ &lt;printed&gt;, ‘ Eusphalerum | geiseri sp.n. | Shavrin A.V. des. 2026’ &lt;red, printed&gt; (TARI).</p><p>Paratypes: 4 ♂♂ (two specimens dissected), 7 ♀♀ (three specimens dissected): same data as the holotype, with additional red printed label: ‘PARATYPE | Eusphalerum | geiseri sp.n. | Shavrin A.V. des. 2026’ &lt;printed&gt; (4 ♂♂, 6 ♀♀: BMNH; 1 ♀: TARI).</p><p>Description. Measurements (n=12): HW: 0.45–0.55; HL: 0.27–0.37; OL: 0.12–0.16; TL: 0.05–0.06; AL (holotype): 0.92; PL: 0.42–0.47; PW: 0.56–0.72; ESL: 0.90–1.06; EW: 0.79–1.04; MTbL (holotype): 0.50; MTrL (holotype): 0.30; AW: 0.88–1.10; AedL: 0.45; BL: 2.02–4.20 (holotype: 2.90).</p><p>Habitus as in Fig. 3. Body yellow-brown to reddish-brown, with slightly paler elytra; abdomen yellowish to reddish-brown; mouthparts, antennae and legs yellow to yellow-brown (mouthparts and antennomeres 1–3 sometimes slightly paler). Head with dense, moderately regular and large punctation, finer and sparser on frontal portion; pronotum with dense and regular punctation, similar to that in middle part of head, sometimes finer and sparser in middle and anteromedial portion, and indistinct in posteromedial third in some specimens; scutellum without punctures; punctation of elytra dense, distinctly larger and deeper than that on pronotum, slightly finer in middle along suture; abdomen without punctures. Head with dense and coarse microsculpture, transverse on clypeus, isodiametric in middle and somewhat longitudinal on infraorbital portions; pronotum with dense and coarse isodiametric microreticulation; scutellum with dense isodiametric meshes; elytra without microreticulation; abdominal tergites with dense isodiametric sculpture.</p><p>Head 1.4–1.6 times as broad as long, with slightly elevated middle part and wide deep anteromedian depressions, reaching level of anterior margins of eyes; anteocellar foveae short and sometimes indistinct, about as long as diameter of ocellus; postocular carina obtuse, with distance between posterior margin of eyes and carina as long as diameters of three-four nearest ommatidia; temples convex, distinctly more than twice as long as eyes. Ocelli large, located at level of posterior margins of eyes; distance between ocelli about as long as distance between ocellus and posterior margin of eyes or slightly longer. Antennomeres 6–10 progressively broadened apicad; basal antennomere about twice as long as broad, antennomere 2 slightly shorter and narrower than basal antennomere, 3 about as long as and distinctly narrower than 2, 4 about twice shorter than 3, 5 slightly broader than 4, 6 distinctly broader than 5, 7 slightly broader than 6, 8 slightly longer and broader than 7, 9 slightly longer and distinctly broader than 8, 10 slightly longer and broader than 9, apical antennomere about twice as long as 10, from preapical third strongly narrowed toward acute apex.</p><p>Pronotum 1.3–1.5 times as broad as long, 1.2–1.3 times as broad as head, from widest middle distinctly more narrowed posteriad than anteriad; laterobasal margins distinctly and sometimes widely (in some specimens) concave in front of rounded posterior angles; anterior angles widely rounded; laterobasal portions widely impressed.</p><p>Elytra slightly longer than broad, slightly more than twice as long as pronotum, from about middle slightly broadened posteriad, with widely rounded posterior margins.</p><p>Protibia straight, slightly broadened and curved apically.</p><p>Abdomen slightly broader than elytra, with two large and transverse and sometimes indistinct tomentose spots in middle of abdominal tergite IV; apical margin of abdominal tergite VII with narrow palisade fringe.</p><p>Male. Abdomen darker, reddish-brown. Posterior margins of abdominal tergite VIII and sternite VIII widely concave. Aedeagus with wide basal part, slightly and gradually narrowed toward apex, median lobe with apex moderately narrow, rounded apically (two paratypes with narrower apical part of aedeagus); parameres long, significantly longer than apex of median lobe, with slightly broadened apical lobes, each with several very long apical and several short preapical setae, each inner preapical margins with several short setae; internal sac narrow and moderately long, with paired narrow, transverse and sclerotized structures in basal portion (Fig. 7). Lateral aspect of the aedeagus as in Fig. 8.</p><p>Female. Abdomen paler, yellowish-brown. Posterior margin of abdominal tergite VIII truncate or somewhat rounded. Posterior margin of abdominal sternite VIII rounded. Accessory sclerite as in Fig. 13. Spermatheca not recognized.</p><p>Comparative notes. Based on the presence of dense punctation on the head and the pronotum, and the general shape of the aedeagus, Eu. geiseri sp. nov. is similar to Taiwanese Eu. simpliciphallum Zanetti, 1998, described from Kaohsiung (Zanetti 1998). The new species can be distinguished from it by the paler head and abdomen, the shorter and broader median lobe, significantly longer parameres with longer apical setae, and different structure of the internal sac and the female accessory sclerite.</p><p>Distribution. Eusphalerum geiseri sp. nov. is known only from the type locality in Nantou (Hsinyi township, Tatachia), central Taiwan.</p><p>Etymology. Patronymic, the species is named to honour coleopterologist Michael Geiser (London), one of collectors of the type material.</p><p>Bionomics. Specimens were collected along the road with Formosan Cypress ( Chamaecyparis formosensis) by beating vegetation ( Maesa perlaria, etc.). They were collected together with Eu. leei sp. nov. and Eu. sacrum sp. nov.</p></div>	https://treatment.plazi.org/id/03BA0C3EBE3C533947ADFA48FF28F9A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2026): Three new species and new records of the genus Eusphalerum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Eusphalerini) from Taiwan. Zootaxa 5768 (3): 419-426, DOI: 10.11646/zootaxa.5768.3.8, URL: https://doi.org/10.11646/zootaxa.5768.3.8
03BA0C3EBE3E533F47ADF955FCAAFA11.text	03BA0C3EBE3E533F47ADF955FCAAFA11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eusphalerum leei Shavrin 2026	<div><p>Eusphalerum leei sp. nov.</p><p>(Figs 4, 9–10)</p><p>Type material. TAIWAN: NANTOU: Holotype ♂ (dissected; left and right antennomeres 9–11 missing): ‘TAIWAN: NANTOU county | Hsinyi township, Tatachia, | Lulin sacred tree, 16.IV.2025 | 23°28'55'' 120°51'17''E, | 2350-2400m. M. Geiser, C.F. | Lee, Y. Hsiao &amp; Y.T. Chung leg. | BMNH {E} 2025-22.’ &lt;printed&gt;, ‘HOLOTYPE | Eusphalerum | leei sp.n. | Shavrin A. V. des. 2026’ &lt;red, printed&gt; (TARI).</p><p>Paratypes: 1 ♂, 3 ♀♀ (one specimen dissected): same data as the holotype, with additional red printed label: ‘PARATYPE | Eusphalerum | leei sp.n. | Shavrin A.V. des. 2026’ (1 ♂ 2 ♀♀: BMNH; 1 ♀ TARI).</p><p>Description. Measurements (n=5): HW: 0.45–0.47; HL: 0.28–0.30; OL: 0.12–0.14; TL: 0.04–0.05; AL (paratype): 0.68; PL: 0.42; PW: 0.57–0.62; ESL: 0.90–0.96; EW: 0.70–0.84; MTbL (holotype): 0.41; MTrL (holotype): 0.23; AW: 0.72–0.92; AedL: 0.42; BL: 2.45 (holotype)–2.60.</p><p>Habitus as in Fig. 4. Head, pronotum and abdomen reddish-brown (lateral and basal portions of pronotum yellow-brown); elytra yellowish-brown. Middle part of head with sparse and fine and infraorbital portions with indistinct very sparse and fine punctation; pronotum with relatively sparse punctation, finer and sparser in middle; punctation of elytra dense, distinctly larger and deeper than that on pronotum, finer in middle; abdominal tergites with indistinct fine and sparse punctation. Head with dense and coarse microsculpture, transverse on clypeus, isodiametric in middle, and diagonal on infraorbital portions; punctation of pronotum similar to that on head, coarser in medioapical and finer in mediobasal portions in paratype; abdominal tergites with dense transverse and isodiametric microsculpture.</p><p>Head 1.5–1.6 times as broad as long; anteocellar foveae indistinct (paratype) or deep and wide, each distinctly longer than diameter of ocellus, reaching level of posterior third of eyes; anteromedian depressions long, reaching anteocellar foveae; postocular carina indistinct, obtuse. Distance between ocelli slightly longer than distance between ocellus and posterior margin of eyes. Antennomeres 5–6 about as long as broad, slightly broader than 4, 7 slightly longer and broader than 6, 8–9 slightly longer and broader than 7, 10 slightly longer than 9.</p><p>Pronotum 1.3–1.4 times as broad as long, from widest middle relatively strongly narrowed posteriad; anterior margin somewhat rounded, slightly concave in middle (paratype).</p><p>Elytra 1.1–1.2 times as long as broad.</p><p>Abdomen distinctly broader than elytra, with two transverse tomentose spots in middle of abdominal tergite IV and two indistinct, small and round spots in middle of tergite V (invisible in paratype).</p><p>Male. Posterior margins of abdominal tergite VIII slightly and sternite VIII widely and deeply concave.Aedeagus with wide and moderately short basal part and elongate relatively narrow median lobe, from apical third gradually narrowed toward rounded apex; parameres exceeding median lobe, each with slightly broadened apical lobe, with two very long apical and several short apical and preapical setae; internal sac moderately short and narrow (Fig. 9). Lateral aspect of the aedeagus as in Fig. 10.</p><p>Female. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII rounded. Sclerotized accessory sclerite and spermatheca not recognized.</p><p>Comparative notes. Based on similar features of microsculpture and punctation of the head and the pronotum, and the general shape of the aedeagus, Eu. leei sp. nov. is similar to Taiwanese Eu. dubitatum (see above), from which can be distinguished by the darker pronotum, smaller body, slightly broader apical part of the median lobe, longer parameres with longer apical setae, different morphology of the internal sac, and different structure of the female accessory sclerite.</p><p>Distribution. Eusphalerum leei sp. nov. is known only from the type locality in Nantou (Hsinyi township, Tatachia), central Taiwan.</p><p>Etymology. Patronymic, the species is named to honour Chi-Feng Lee (Taichung), one of collectors of the type material.</p><p>Bionomics. See bionomical notes for Eu. geiseri sp. nov.</p></div>	https://treatment.plazi.org/id/03BA0C3EBE3E533F47ADF955FCAAFA11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2026): Three new species and new records of the genus Eusphalerum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Eusphalerini) from Taiwan. Zootaxa 5768 (3): 419-426, DOI: 10.11646/zootaxa.5768.3.8, URL: https://doi.org/10.11646/zootaxa.5768.3.8
03BA0C3EBE38533F47ADF9C1FCAFF87B.text	03BA0C3EBE38533F47ADF9C1FCAFF87B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eusphalerum luteoides Zanetti 1998	<div><p>Eusphalerum luteoides Zanetti, 1998</p><p>(Fig. 5)</p><p>Eusphalerum luteoides Zanetti, 1998: 140 .</p><p>Material. TAIWAN: NANTOU: 7 ♂♂, 6 ♀♀: ‘TAIWAN: NANTOU county Ren`ai township, Yuanfeng . 24°07'04'' 121°14'13''E, 2760 m, 14.IV.2025, beating. M. Geiser &amp; C.F. Lee leg. BMNH{E} 2025-22.’ (BMNH) .</p><p>Remarks. Eusphalerum luteoides (Fig. 5) was originally described based on 14 specimens from several localities in Pingtung and Kaohsiung, Taiwan. Specimens from Nantou were collected by beating shrubs near Yuanfeng observatory. It is here recorded from Nantou for the first time.</p></div>	https://treatment.plazi.org/id/03BA0C3EBE38533F47ADF9C1FCAFF87B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2026): Three new species and new records of the genus Eusphalerum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Eusphalerini) from Taiwan. Zootaxa 5768 (3): 419-426, DOI: 10.11646/zootaxa.5768.3.8, URL: https://doi.org/10.11646/zootaxa.5768.3.8
03BA0C3EBE3B533D47ADFF60FCAAFCCD.text	03BA0C3EBE3B533D47ADFF60FCAAFCCD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eusphalerum sacrum Shavrin 2026	<div><p>Eusphalerum sacrum sp. nov.</p><p>(Figs 6, 11–12, 14)</p><p>Type material. TAIWAN: NANTOU: Holotype ♂ (dissected): ‘TAIWAN: NANTOU county | Hsinyi township, Tatachia, | Lulin sacred tree, 16.IV.2025 | 23°28'55'' 120°51'17''E, | 2350-2400m. M. Geiser, C.F. | Lee, Y. Hsiao &amp; Y.T. Chung leg. | BMNH {E} 2025-22.’ &lt;printed&gt;, ‘PARATYPE | Eusphalerum | sacrum sp.n. | Shavrin A.V. des. 2026’ &lt;printed&gt; (TARI).</p><p>Paratypes: 6 ♂♂ (one specimen without right antennomeres 8–11), 4 ♀♀ (dissected): same data as the holotype, with additional red printed label: ‘PARATYPE | Eusphalerum | sacrum sp.n. | Shavrin A.V. des. 2026 &lt;printed&gt; (6 ♂♂, 3 ♀♀: BMNH; 1 ♀: TARI).</p><p>Description. Measurements (n=11): HW: 0.50–0.62; HL: 0.29–0.32; OL: 0.12–0.15; TL: 0.03–0.05; AL (holotype): 0.77; PL: 0.36–0.45; PW: 0.59–0.67; ESL: 0.81–1.01; EW: 0.79–0.97; MTbL (holotype): 0.38; MTrL (holotype): 0.22; AW: 0.67–0.96; AedL: 0.45; BL: 2.16–2.65 (holotype: 2.40).</p><p>Habitus as in Fig. 6. Forebody yellowish-brown (neck in holotype and one paratype darker, reddish-brown; scutellum in holotype and two paratypes distinctly darker); abdomen yellowish-brown to reddish-brown (paratergites and apical tergites slightly paler); antennomeres 7–11 usually darkened. Head with sparse and fine punctation, denser in middle, holotype and two paratypes with indistinct punctures; punctation of pronotum moderately sparse, distinctly larger and deeper than that in middle part of head, finer and sparser in middle and posteromedial portions (two paratypes without punctures in posteromedial part); elytra with dense, large and deep punctation, usually finer and sparser in middle. Head and pronotum with very dense, regular, isodiametric microsculpture; abdominal tergites with dense transverse microreticulation.</p><p>Head 1.7–1.9 times as broad as long, with indistinct shallow, narrow and short anteocellar foveae; anteromedian depressions almost reaching anteocellar foveae; postocular carina indistinct, obtuse. Antennomeres 5–6 indistinctly longer and broader than 4, 6 slightly longer and broader than 5, 7 slightly broader than 6, 8 slightly broader than 7, 9 slightly longer and broader than 8, 10 indistinctly longer than 9.</p><p>Pronotum 1.4–1.6 times as broad as long, slightly broader than head, from widest middle moderately strongly narrowed posteriad; anterior margin widely rounded or somewhat straight.</p><p>Abdomen slightly narrower than elytra, with two indistinct transverse tomentose spots in middle of abdominal tergite IV and two indistinct, small and round spots in middle of tergite V (invisible in two paratypes) .</p><p>Male. Abdomen reddish-brown. Posterior margins of abdominal tergite VIII slightly and sternite VIII widely and deeply concave. Aedeagus with wide basal portion, gradually narrowed toward short and narrow median lobe; parameres significantly longer than apex of median lobe, each with broadly widened apical lobes with two long and several short apical and preapical setae, with additional several short setae on widened preapical part of paramere; internal sac wide and short (Fig. 11). Lateral aspect of the aedeagus as in Fig. 12.</p><p>Female. Abdomen yellowish-brown. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII rounded. Accessory sclerite as in Fig. 14. Spermatheca not recognized.</p><p>Comparative notes. Based on the coloration and the length of the body, the coloration of darkened apical antennomeres, and the general shape of the aedeagus, Eu. sacrum sp. nov. is somewhat similar to Taiwanese Eu. pseudoreitteri Zanetti, 1998, also described from Nantou (Zanetti 1998), from which can be reliably distinguished by the shape of shorter median lobe, the longer parameres with narrower apical lobes with different features of the apical setation, and completely different morphology of the internal sac and the female accessory sclerite.</p><p>Distribution. Eusphalerum sacrum sp. nov. is known only from the type locality in Nantou (Hsinyi township, Tatachia), central Taiwan.</p><p>Etymology. The specific epithet is the Latin adjective sacer, feminine sacra, neuter sacrum (holy, sacred). It alludes to the type locality in Tatachia with Lulin sacred tree.</p><p>Bionomics. See bionomical notes for Eu. geiseri sp. nov.</p></div>	https://treatment.plazi.org/id/03BA0C3EBE3B533D47ADFF60FCAAFCCD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2026): Three new species and new records of the genus Eusphalerum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Eusphalerini) from Taiwan. Zootaxa 5768 (3): 419-426, DOI: 10.11646/zootaxa.5768.3.8, URL: https://doi.org/10.11646/zootaxa.5768.3.8
