taxonID	type	description	language	source
03BDEA64330FC621FCEEFD86E8E4FC10.taxon	type_taxon	Type species: Beloptera belemnoidea de Blainville, 1825; by original designation.	en	Dernov, Vitaly (2024): Beloptera Longa Naef, 1922 (Cephalopoda: Coleoidea) From The Late Eocene (Palaeogene) Of Ukraine. Acta Palaeontologica Romaniae 20 (2): 3-10, DOI: 10.35463/j.apr.2024.02.01, URL: https://doi.org/10.35463/j.apr.2024.02.01
03BDEA64330FC621FCEEFD86E8E4FC10.taxon	diagnosis	Diagnosis: Chamber length-to-diameter ratio <0.2, except earliest (postembryonic – juvenile) chambers; postalveolar sheath oriented towards venter, length of club variable, lateral extensions well developed, winglike, oriented towards venter, ventral ridge moderately developed (after Fuchs, 2023: p. 8 – 9).	en	Dernov, Vitaly (2024): Beloptera Longa Naef, 1922 (Cephalopoda: Coleoidea) From The Late Eocene (Palaeogene) Of Ukraine. Acta Palaeontologica Romaniae 20 (2): 3-10, DOI: 10.35463/j.apr.2024.02.01, URL: https://doi.org/10.35463/j.apr.2024.02.01
03BDEA64330FC623FC11FC6AEF2FF795.taxon	description	Fig. 4 a – c Synonymy	en	Dernov, Vitaly (2024): Beloptera Longa Naef, 1922 (Cephalopoda: Coleoidea) From The Late Eocene (Palaeogene) Of Ukraine. Acta Palaeontologica Romaniae 20 (2): 3-10, DOI: 10.35463/j.apr.2024.02.01, URL: https://doi.org/10.35463/j.apr.2024.02.01
03BDEA64330FC623FC11FC6AEF2FF795.taxon	materials_examined	Holotype: Not defined.	en	Dernov, Vitaly (2024): Beloptera Longa Naef, 1922 (Cephalopoda: Coleoidea) From The Late Eocene (Palaeogene) Of Ukraine. Acta Palaeontologica Romaniae 20 (2): 3-10, DOI: 10.35463/j.apr.2024.02.01, URL: https://doi.org/10.35463/j.apr.2024.02.01
03BDEA64330FC623FC11FC6AEF2FF795.taxon	diagnosis	Diagnosis: Beloptera with an elongate guard-like sheath that is slightly curved ventrally and with semicircular in outline lateral expansions. The dorsal surface is roof-like, almost smooth with radial grooves on the rostrum; the ventral surface is concave and almost smooth (radial grooves are on the rostrum); the rostrum is nearly triangular in cross-section.	en	Dernov, Vitaly (2024): Beloptera Longa Naef, 1922 (Cephalopoda: Coleoidea) From The Late Eocene (Palaeogene) Of Ukraine. Acta Palaeontologica Romaniae 20 (2): 3-10, DOI: 10.35463/j.apr.2024.02.01, URL: https://doi.org/10.35463/j.apr.2024.02.01
03BDEA64330FC623FC11FC6AEF2FF795.taxon	materials_examined	Material: One moderately preserved specimen (IGS-DYM 01 / 05) from the Priabonian Mandrykivka Beds of the Rybalsky Quarry, Dnipro.	en	Dernov, Vitaly (2024): Beloptera Longa Naef, 1922 (Cephalopoda: Coleoidea) From The Late Eocene (Palaeogene) Of Ukraine. Acta Palaeontologica Romaniae 20 (2): 3-10, DOI: 10.35463/j.apr.2024.02.01, URL: https://doi.org/10.35463/j.apr.2024.02.01
03BDEA64330FC623FC11FC6AEF2FF795.taxon	description	Description: The specimen IGS-DYM 01 / 05 is a moderately preserved elongated (L 2 / W 2 = 2.44, see Table 1), slightly curved ventrally 26.4 - mm-long guard-like sheath with supposedly semicircular in outline lateral expansions inclined towards the venter and gives a convex form to the dorsal surface and a corresponding concavity to the venter. The lateral expansions are thick at the articulation area of the rostrum and sheath and become gradually thinner to the periphery. The dorsal surface is roof-like with a raised narrow medial zone and steep slopes gradually turning into lateral expansions. The rostrum is nearly triangular in cross-section and gradually widens; its apex is rounded in the ventral aspect and slightly pointed in the lateral aspect; the rostrum is slightly curved ventrally and the guard-like sheath is somewhat arched longitudinally. The ventral surface is almost smooth with very fine radial grooves on the rostrum. The dorsal surface is almost smooth with very fine radial grooves are on the rostrum. dimorphism of B. belemnoidea specimens. However, B. belemnoidea significantly differs from B. curta in the longer and less massive rostrum, as well as an elliptical outline guard-like sheath. It is rather difficult to compare B. longa with B. konincki Briart and Cornet, 1880, as I know this species is described from not well preserved material only in Briart & Cornet (1880); images of this species in this work are not of very high quality. Probably, in contrast to B. longa, B. konincki has a rather short guard-like sheath with a more pointed, relatively short rostrum and radial and concentric ornamentation of the ventral surface. Occurrence: Bartonian of England, Priabonian of Ukraine. Specimen L 1 L 2 W 1 W 2 L 2 / W 2 IGS- 13.6 26.4 6.7 10.8 2.44 DYM 01 / 05	en	Dernov, Vitaly (2024): Beloptera Longa Naef, 1922 (Cephalopoda: Coleoidea) From The Late Eocene (Palaeogene) Of Ukraine. Acta Palaeontologica Romaniae 20 (2): 3-10, DOI: 10.35463/j.apr.2024.02.01, URL: https://doi.org/10.35463/j.apr.2024.02.01
03BDEA64330FC623FC11FC6AEF2FF795.taxon	discussion	Remarks: The described specimen does not differ significantly from Beloptera longa figured by Edwards & Wood (1877: pl. 2, fig. 1) and Naef (1922: text-figs 18 a – g, 19). B. longa differs from the most morphologically similar species B. belemnoidea de Blainville, 1825 in the following morphological features: (1) the very convex smooth dorsal surface; (2) the greatly curved ventrally guard-like sheath; (3) the nearly triangular in cross-section rostrum; (4) the slightly pointed in the lateral aspect rostrum. B. longa differs from B. curta (Cossmann, 1896) in a much longer rostrum (relative to the total length of the guard-like sheath) and a more elongated guard-like sheath. In addition, the sheath of B. curta is rather strongly compressed in the dorsal-ventral direction. Vincent (1901) considered B. curta to be the ” anomaly ” of B. belemnoidea, probably referring to the manifestation of intraspecific, probably sexual,	en	Dernov, Vitaly (2024): Beloptera Longa Naef, 1922 (Cephalopoda: Coleoidea) From The Late Eocene (Palaeogene) Of Ukraine. Acta Palaeontologica Romaniae 20 (2): 3-10, DOI: 10.35463/j.apr.2024.02.01, URL: https://doi.org/10.35463/j.apr.2024.02.01
03BDEA64330DC625FC65FD43ECD9FF7E.taxon	description	Fig. 4 d – k	en	Dernov, Vitaly (2024): Beloptera Longa Naef, 1922 (Cephalopoda: Coleoidea) From The Late Eocene (Palaeogene) Of Ukraine. Acta Palaeontologica Romaniae 20 (2): 3-10, DOI: 10.35463/j.apr.2024.02.01, URL: https://doi.org/10.35463/j.apr.2024.02.01
03BDEA64330DC625FC65FD43ECD9FF7E.taxon	materials_examined	Material: Three moderately preserved specimens (IGS-DYM 01 / 06 to IGS-DYM 01 / 08) from the Priabonian Mandrykivka Beds of the Rybalsky Quarry, Dnipro.	en	Dernov, Vitaly (2024): Beloptera Longa Naef, 1922 (Cephalopoda: Coleoidea) From The Late Eocene (Palaeogene) Of Ukraine. Acta Palaeontologica Romaniae 20 (2): 3-10, DOI: 10.35463/j.apr.2024.02.01, URL: https://doi.org/10.35463/j.apr.2024.02.01
03BDEA64330DC625FC65FD43ECD9FF7E.taxon	description	Description: The specimen IGS-DYM 01 / 06 (Fig. 4 d – f) is a moderately preserved elongated (L 2 / W 2 = 2.51; see Table 2) guard-like sheath, which is slightly curved ventrally; it is 34.6 - mm-long with semicircular in outline lateral expansions inclined towards the venter, which gives a convex form to the dorsal surface and a corresponding concavity to the venter. The lateral expansions are thick at the contact area of the rostrum and sheath and become gradually thinner to the periphery. The dorsal surface is roof-like with a raised narrow medial zone and steep slopes gradually turning into lateral expansions. The rostrum is nearly triangular in cross-section and gradually widens; its apex is rounded; the rostrum is slightly curved ventrally and the guard-like sheath is somewhat arched longitudinally. The ventral surface is covered with very small elongate pits at the base of the lateral expansions and very fine radial grooves on the rostrum. The dorsal surface is almost smooth with very fine radial grooves on the rostrum and front zones of the lateral expansions. Specimen IGS-DYM 01 / 07 (Figs. 4 g-i) is a moderately preserved elongated (L 2 / W 2 = 2.0) guard-like sheath that is slightly curved ventrally; it measures 21.4 mm long with lateral expansions that are inclined towards the ventral surface, which causes a convex shape towards the dorsal surface and a corresponding concavity towards the ventral surface. The lateral expansions are thick in the area of contact between the rostrum and the sheath and gradually thinning towards periphery. The dorsal surface is roof-like with a raised narrow medial zone and steep slopes gradually turning into the lateral expansions. The rostrum is nearly triangular in cross-section and gradually widens; its apex is rounded in the ventral aspect and slightly pointed in the lateral aspect; the rostrum is slightly curved ventrally and the guard-like sheath is somewhat arched longitudinally. The ventral surface is covered with very small elongate pits at the base of the lateral expansions and very fine radial grooves on the rostrum. Most of the dorsal surface is smooth; only shallow marks of blood vessels occur on the preserved lateral expansion and very fine radial grooves are on the rostrum. The specimen IGS-DYM 01 / 08 (Fig. 4 j, k) is represented by a fragment of a rostrum that is nearly triangular in cross-section with a part of the lateral expansions. This poorly preserved fossil is indistinguishable from the specimen IGS-DYM 01 / 06.	en	Dernov, Vitaly (2024): Beloptera Longa Naef, 1922 (Cephalopoda: Coleoidea) From The Late Eocene (Palaeogene) Of Ukraine. Acta Palaeontologica Romaniae 20 (2): 3-10, DOI: 10.35463/j.apr.2024.02.01, URL: https://doi.org/10.35463/j.apr.2024.02.01
03BDEA64330DC625FC65FD43ECD9FF7E.taxon	discussion	Remarks: The specimen IGS-DYM 01 / 06 described above is better preserved than the single specimen of Beloptera cf. longa (IGS NASU- 15 / 02) described in the work of Dernov & Demianov (2023: Fig. 6 A – E). In particular, IGS-DYM 01 / 06 shows that, unlike B. belemnoidea, the lateral expansions of B. aff. longa have semicircular margins, while in B. belemnoidea they are wavy (cf. Fig. 3 a in this work). In contrast to B. cf. longa (IGS NASU- 15 / 02), the specimen IGS-DYM 01 / 06 is less ventrally curved and bears thin radial grooves on the anterior part of the dorsal surface of the lateral expansions. The described specimens differ from the specimens of B. longa, figured by Edwards & Wood (1877: pl. 2, fig. 1), Naef (1922: figs 18 a – g) and in Fig. 4 a – c in this paper by a less massive rostrum. B. aff. longa differs from the most morphologically similar species B. belemnoidea in a very convex smooth dorsal surface, a greatly curved ventrally guard-like sheath and the nearly triangular in cross-section rostrum. Occurrence: Bartonian of England, Priabonian of Ukraine.	en	Dernov, Vitaly (2024): Beloptera Longa Naef, 1922 (Cephalopoda: Coleoidea) From The Late Eocene (Palaeogene) Of Ukraine. Acta Palaeontologica Romaniae 20 (2): 3-10, DOI: 10.35463/j.apr.2024.02.01, URL: https://doi.org/10.35463/j.apr.2024.02.01
