taxonID	type	description	language	source
03BD879EFFC3BF63FF78F8FCFB7A3482.taxon	type_taxon	Type species: Incasarus garciai Gonzalez, Rasmussen & Engel 2013	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFC3BF63FF78F8FCFB7A3482.taxon	diagnosis	Diagnosis: Incasarus can be distinguished from other genera of Protandrenini — except Liphanthus — by i) a narrow pterostigma, with its greatest width less than twice the width of prestigma (Fig. 8), and ii) the margin within the marginal cell mostly straight (Fig. 8). All other genera in the tribe have a wider pterostigma, with its greatest width more than twice or so the width of the prestigma (at least as in Fig. 9), and the margin within marginal cell at least weakly convex (at least as in Fig. 9). From Liphanthus, Incasarus can be readily separated by the number of submarginal cells: Incasarus has iii) two submarginal cells, while most Liphanthus species have three. Among the nine described Liphanthus species with two submarginal cells, Incasarus can be recognized by any of the following characteristics: iv) upper paraocular and vertexal areas of head, as well as the mesoscutum, with dense to crowded punctures (at least as in Fig. 10); v) anterior portion of the tegula strongly sculptured (Fig. 10); vi) postgradular area of T 2 not strongly depressed — not more so than in other terga (Fig. 12); vii) head and mesosoma with weak metallic reflections, visible under certain lighting (Fig. 10); viii) head and mesosoma bearing minute hairs, visible under certain angles; and ix) S 7 apical lobe fused to apodeme (Figs. 23, 33, 48). In contrast, all Liphanthus with two submarginal cells, the head and mesoscutum have less dense, never crowded punctures (as in Fig. 11); the anterior portion of the tegula is unsculptured or weakly sculptured (Fig. 11); the T 2 postgradular area is strongly depressed (Fig. 13), more than in other terga, and the S 7 apical lobe is differentiated from the apodeme (Fig. 47). The females known in Incasarus also display characteristics i – viii, except that the mesoscutum of some species exhibits somewhat sparser punctation than in males and the microsculpture is weaker, making the surface shiny between punctures (as in Fig. 42). Females of Liphanthus with two submarginal cells can also exhibit distinct punctation but it is not nearly as dense as in any female of Incasarus (compare Figs. 41 and 42).	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDDBF63FF78FAE6FE8C379E.taxon	description	(Figs. 1, 3, 7 – 8, 14 – 15, 49, 53) This species was described in some detail by Gonzalez et al. (2013) and this information is not repeated here, although a new diagnosis and images are provided to facilitate its separation from congeners.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDDBF63FF78FAE6FE8C379E.taxon	diagnosis	Diagnosis: Males can be distinguished from all other species in the genus by any the following characters: presence of a distinct tuft of long setae on the outer mandibular groove (Figs. 14, 49); outer mandibular ridge forming an acute process that surpasses the lower margin of the mandible (Figs. 14, 49); S 1 and S 2 with apicolateral bands of dense and long setae (Figs. 15, 51); and the gonostylus differentiated from the gonocoxa (Figs. 3, 53). Other members of the genus lack both the tuft on outer groove, and the acute process of the outer ridge of the mandible, and the apicolateral bands with dense and long hairs on S 2 and S 3 of I. garciai. The female remains unknown.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDDBF63FF78FAE6FE8C379E.taxon	materials_examined	New record: PERU. 1 ♂; Huánuco, CP [Centro Poblado] Nauyan Rondos, 9 ° 53 ’ 28.27 ” S, 76 ° 17 ’ 44.98 ” W [- 9.891, - 76.296], 3008 m, vi. 2019, I. Medina — MUSM.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDDBF63FF78FAE6FE8C379E.taxon	discussion	Comments: This species was described based upon a sole male specimen from PERU, Ayacucho {- 13.16, - 74.21}, 2750 m, 22. xi. 1971. However, the specimen could not be located at MUSM where it was reportedly deposited (Gonzalez et al. 2013; Mabel Alvarado, pers. comm.; Victor Gonzalez, pers. comm.). The specimen from Huánuco examined in this study largely agrees with the published description in most external morphological features and in the structure of the terminalia, and the differences between this new specimen and the description of the holotype seem minor. Further, as the two localities for the specimens are separated by 426 km, we assume conspecificity.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDCBF62FF78FF45FAAF3565.taxon	description	(Figs. 7, 16 – 17, 44, 64, 71, 73, 75, 89 – 90, 94)	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDCBF62FF78FF45FAAF3565.taxon	discussion	Comments: Direct examination of the female lectotype and male paralectotype of Panurginus aeneiventris Friese, 1908, currently Rhophitulus aeneiventris following Moure (2022), deposited at the ZMHB (Figs. 16 – 17), revealed that it is conspecific with Liphanthus fritzi Mir Sharifi & Packer, 2019. The most noticeable difference between the female lectotype of R. aeneiventris we both studied at ZMHB and the female specimen of L. fritzi is the slightly reddish tints of the metasoma in the former; however, we suspect that this coloration may be due to the method of collection or specimen preservation, and not taxonomically significant. The only other minor difference observed was in the IOD: OOD ratio (0.6 in R. aeneiventris versus 0.5 in L. fritzi), which falls within the range of variation observed for other species of Incasarus and is incorporated into the identification key provided below. No notable morphological differences were detected between the paralectotype male of R. aeneiventris and the male holotype and male paratype of L. fritzi. Therefore, Incasarus aeneiventris (Friese, 1908) is regarded here as a senior synonym of Liphanthus fritzi Mir Sharifi & Packer syn. nov. This species was redescribed in detail in Mir Sharifi et al. (2019), and that information is not repeated here; however, we include type specimen information, additional locality records and new images to aid with identification.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDCBF62FF78FF45FAAF3565.taxon	materials_examined	Lectotype and paratype locality: ARGENTINA. Lectotype ♀; Salta, 2500 m, [iii]. 1905, Steinbach. Paratype ♂; same data except 1200 m. New records: ARGENTINA. 1 ♂, 1 ♀; Catamarca, Quirós {- 28.783, - 65.116, 300 m}, 28. xi. 1941 — MLPA. 3 ♂, 1 ♀; Catamarca, Rodeo {- 28.207, - 65.875}, 1500 m, 8 – 28. i. 1954. R. Golbach — IFML. 1 ♂, 1 ♀; Jujuy, Río Chico {- 22.062, - 65.969, 3000 m}, 18. ii. 1985, J. F — MLPA. One 1 ♂; Jujuy, Valle Grande, Finca Arrayanaí, - 23.466, - 64.944, 1709 m, 16. xi. 2023, L. Álvarez & M. Lucía — MLPA. 1 ♂; Salta, Yacochuya, {- 26.043, - 66.020}, 2400 m, 04. iv. 1969, A. Willink — IFML. 8 ♂, 1 ♀; Salta, Finca El Rey {- 24.676, - 64.606, 1200 m}, 31. xi. 1952, A. Ogloblin — MLPA. 1 ♀; Tucumán, Horco Molle {- 26.793, - 65.315, 600 m}, 01. i. 1966, L. A. Stange — IFML. 1 ♀; Tucumán, camino a [road to] Hualinchay, - 26.313, - 65.613, {2000 m}, 5. xi. 2013, L. Álvarez & M. Lucía — MLPA. 1 ♂, 1 ♀; Tucumán, Colalao, {- 26.248, - 65.489, 1000 m}, Arnau — MLPA.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDCBF62FF78FF45FAAF3565.taxon	description	The following four species were described for the first time and in detail in Mir Sharifi et al. (2019), and that information is not repeated here. When applicable, new records and additional comments are provided.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDFBF61FF78FF45FB8F313E.taxon	description	(Figs. 7, 70, 77, 79)	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDFBF61FF78FF45FB8F313E.taxon	discussion	Comments: Female remains unknown. No additional records are currently known.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDFBF61FF78FE79FC7432F5.taxon	description	(Figs. 7, 68, 78, 80, 86, 88, 91 – 92, 96)	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDFBF61FF78FE79FC7432F5.taxon	materials_examined	New records: ARGENTINA. 3 ♂; Catamarca, El Portezuelo, {- 28.373, - 65.641, 800 m}, 27. xi. 1991 — MLPA. 1 ♀; Catamarca, N. Andalgalá, {1500 m}, 14 – 15. ii. 2003, L. Packer — PCYU.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDFBF61FF78FD31FC4A338A.taxon	description	(Figs. 7, 55, 60, 62, 81, 83)	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDFBF61FF78FD31FC4A338A.taxon	discussion	Comments: Mir sharifi et al. (2019) recognized this as a very distinctive species with numerous unique characteristics, including the sexual dimorphism in colour (brown metasoma of the females versus red of the males), the male modified S 6 and unique genitalia. No additional records are currently known.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDFBF61FF78FBEDFDCC356F.taxon	description	(Figs. 7, 10, 42, 56, 66 – 67, 69, 72, 74, 76, 95)	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDFBF61FF78FBEDFDCC356F.taxon	description	No additional records are currently known.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDFBF61FF78FA88FE7B37ED.taxon	description	(Figs. 7, 85, 87)	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDFBF61FF78FA88FE7B37ED.taxon	materials_examined	New record: ARGENTINA. 1 ♀; Jujuy, 20 Km S Abra Pampa, - 22.891 - 65.619, 3682 m, 11. iv. 2018, L. Packer — MACN.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDFBF61FF78FA88FE7B37ED.taxon	discussion	Comments: This species was described based on a female specimen. The male is unknown. The authors noted the dense punctation on the head and mesoscutum, which suggested it could be the female of I. abotorabi (Mir Sharifi & Packer); however, they argued against this possibility due to the presence of pale markings on the legs, a feature absent in females of other species in the group. Although we did not include this species in our morphological analysis because most characters were scored from males, we hereby reassign it to Incasarus because of the combination of narrow pterostigma, two submarginal cells; dense punctation on head, crowded punctation on mesoscutum; anterior portion of tegula with strong surface sculpture; and bronze reflections on the integument of head and mesoscutum, characteristics seen in the females of this genus but not in those of Liphanthus with two submarginal cells.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDEBF67FF78FF45FBD03556.taxon	description	(Figs. 7, 12, 18 – 25, 40, 48, 50, 52, 54, 57, 59, 61)	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDEBF67FF78FF45FBD03556.taxon	diagnosis	Diagnosis: Males can be readily distinguished from all other Incasarus by the presence of cream markings on the lower paraocular area (Figs. 18, 20 – 21, 59), which is entirely dark in other members (Figs. 14, 26, 60, 63 – 64, 77 – 78). Additionally, the combination of a mandible with a broadened apex, appearing to bear a preapical tooth (Figs. 18, 50), and the presence of comb-like patches of thick setae on S 4 (Figs. 19, 22, 61) is unique within the genus.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDEBF67FF78FF45FBD03556.taxon	description	Description. Holotype male: Dimensions: Approximate body length: ~ 5.2 mm; head width 1.5 mm; forewing length 3.3 mm; ITW 0.9 mm. Coloration: General integument black to dark brown, with subtle dark silver reflections on head and mesosoma, under certain angles. The following parts yellow: mandible (dark brown basally, becoming reddish-brown to orange subapically and dorsally, apex red); apical ~ 1 / 3 of clypeus, slightly more extensive adjacent to epistomal lobe; lower paraocular area (except for a narrow strip adjacent to compound eye); pro- and mesofemora apicodorsally; metafemur apicodorsal spot; pro- and mesotibiae extreme base and apex; metatibia dorsally for basal half extending apically to the anterior half; metatibia apical spot; all basitarsi dorsally (suffused with brown ventrally). Tegula, wing membrane and apical impressed areas of metasomal terga light brown. Sculpture: Labral process unsculptured, shiny; portion below process dull. Clypeus extreme apex weakly sculptured to unsculptured, shiny, becoming areolate toward dark portion of clypeus; pale portion punctures shallow, obscure medially, mostly large and dense laterally (i = 1 – 1.5 d). Clypeus dark portion areolate, somewhat shiny, impunctate except sparsely punctate adjacent to yellow portion. Supraclypeal area areolate to moderately imbricate, shiny; punctures obscure below, deep and dense between antennal sockets (i = 0.5 d). Subantennal sclerite strigate, somewhat dull above, shinier below; punctures dense (i = 0.5 – 1 d). Lower paraocular area moderately imbricate, somewhat dull, punctures small, somewhat sparse (i = 1 – 3 d). Paraocular area mesad of midlength of compound eye areolate, somewhat dull around antennal socket, shinier adjacent to compound eye; punctures deep, somewhat sparse (i = 1 – 2.5 d), denser (i <0.5 d) below level of facial fovea. Upper paraocular and vertexal areas somewhat dull, microsculpture obscure due to dense to crowded punctures; areolate, with punctures becoming sparser (i = 1 d) approaching occipital area. Genal and hypostomal areas strongly imbricate, somewhat shiny; punctures shallow and somewhat obscured by microsculpture, dense (i <0.5 d) near compound eye, otherwise irregularly spaced (i = 1 – 4 d). Mesoscutum dull, microsculpture obscured by deep and dense, almost crowded punctation. Scutellum and metanotum strongly areolate, dull; punctures large (i = 1 – 1.5 d and i = 1 – 2 d, respectively). Propodeum dorsal surface areolate, shiny, with weak short longitudinal striae only medially. Hypoepimeral area and mesopleuron strongly to moderately imbricate, dull to somewhat shiny; punctures shallow, dense (i = 1 d), sparser ventrally (i = 1.5 – 2 d). Metapleuron and sides of propodeum strongly areolate, somewhat shiny; dorsolateral area of propodeum areolate to moderately imbricate, shiny, impunctate. Metasomal terga moderately to strongly imbricate, shiny; T 1 – T 2 punctures small, dense (i = 1 d), except anterior declivitous area of T 1 punctures shallow, sparse (i> 4 d). T 3 punctures minute, moderately dense (i = 1 – 2 d). T 4 – T 6 punctures shallow, irregular in size and spacing (i ≤ 3 d); apical impressed areas of terga imbricate, somewhat shiny, impunctate. Metasomal sterna moderately imbricate, somewhat shiny; punctures shallow, obscure. Pubescence: Generally white, sparse, and short. Head and mesoscutum minute, dense hairs, only visible from certain angles. Labrum with short ≤ 0.5 MOD, branched, dense hairs apically, sparser laterally. Clypeus with long ≤ 2 MOD hairs subapicolaterally, ≤ 3 MOD on extreme apicolateral margin. Lower paraocular area and scape with long ≤ 2 MOD hairs; ≤ 1.5 MOD on vertexal area; ~ 1 MOD on genal and hypostomal areas. Pronotum lateral surface branched hairs ≤ 1 MOD, Mesoscutum posteriorly ≤ 1.2 MOD; scutellum laterally and metanotum ≤ 2 MOD; posterior surface of propodeum branched, dense hairs ≤ 2.5 MOD. Leg hairs ≤ 1.2 MOD, except metatibia with fine apicallyoriented hairs, dorsally increasing in length toward apex 1 – 4 MOD, equally or slightly thicker than elsewhere on metatibia; metabasitarsus hairs dense, brown basoventrally, paler brown elsewhere ≤ 1.3 MOD. Metasomal terga generally with minute, moderately dense hairs except longer ~ 1.5 MOD on T 1 anterior declivitous surface and towards sides; row of fine and sparse hairs ~ 1 – 2 MOD at junction of disc and apical impressed areas of T 2 – T 6, T 7 laterally recurved area hairs ≤ 2.5 MOD. Sterna generally with minute, sparse hairs. S 4 bearing two comb-like patches of thick black setae on apical margin (Figs. 19, 22). S 5 – S 6 with branched medially oriented hairs, ~ 1.2 MOD. S 5 apical margin with a medial fringe of simple, posteriorly oriented hairs ≤ 2 MOD. Structure: Head: ~ 1.2 × as wide as long (81: 65). Mandible ~ 3.4 × longer than basal depth (66: 19). Labrum ~ 2.2 × as wide as long (45: 20), labral process rectangular, marginal ridge strong apically, weaker laterally. Clypeus ~ 3.2 × as wide as long (36: 11), convex in profile with weak medial swelling ~ 1 / 5 length from the base, extreme apex weakly recurved ventrally, in profile. Inner and outer subantennal sutures straight, subparallel, subantennal sclerite parallel-sided; anterior tentorial pit deep, at junction of outer subantennal and epistomal sutures; epistomal suture transverse between inner subantennal sutures. Frontal line weak but distinct, flat above, a raised ridge between antennal sockets. IAD> AOD (10: 7). Inner margins of compound eyes strongly convergent below, UOD> LOD (42: 35). Facial fovea small, ~ 1 / 3 as long as scape, oval, twice as long as wide (4: 2). IOC <OOC (9: 13). Scape more than 3 × as long as greatest width (13: 4), length subequal to pedicel and F 1 combined (14); pedicel length and width subequal (4: 5); F 1 more than 3 × longer than wide (10: 3); F 2 1.5 × longer than wide (6: 4); [remaining flagellomeres lengths and widths subequal (~ 6: 6), except F 11 ~ 2.2 × as long as greatest width (11: 5)]. Mesosoma: Mesoscutum ~ 1.3 × as wide as long (41: 32). Scutellum as long as metanotum and dorsal surface of propodeum combined, 13: 8: 5. Marginal cell almost as long as distance between its apex and wing tip (32: 36). Metatibia ~ 4.5 × longer than greatest depth (50: 11). Metabasitarsus weakly swollen basoventrally,> 7 × longer than deep at mid-length (50: 7). Metatibial spur apex curved, posterior spur longer than anterior. Metasoma: Broadest at midlength of T 2. S 1 – S 3 unmodified. S 4 apical margin concave medially between setal patches. S 5 – S 6 narrowly and transversely concave, distinctly impressed relative to other sterna. S 6 with shallow U-shape apicomedial emargination. Terminalia: S 7: Apodemes enormous, long and broad, anteromedial margin concave, forming an angle of less than 10 ° at point of attachment; apical lobes length and width subequal, fused to apodeme, apex truncate, except acute angulation longer than lobe, oriented obliquely at an angle of ~ 45 ° relative to the longitudinal axis of the lobe, lacking hairs. S 8: Spiculum anterior margin strongly concave, anterior lobe with greatest width and length subequal, lateral margins strongly concave, diverging anteriorly; lateral lobe large, subtriangular; apical lobe with lateral margins converging towards narrow and subtruncate apex, covered in dense, posterolaterally oriented hairs. Gonocoxa anterodorsally convex, outer margins convex, inner margins relatively straight. laterally, strongly concave towards apex, medial margin approximately straight for most of length, briefly concave towards apex; gonostylus fused to gonocoxa, narrow, ~ 4 × longer than medial width, apex rounded with short, mostly medioventrally oriented dense hairs; penis valve narrow, lateral margins mostly straight narrowing to truncate apex, hair short, almost indetectable. Female: unknown.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDEBF67FF78FF45FBD03556.taxon	materials_examined	Material studied: PERU. Holotype ♂, and 4 ♂ paratypes; Arequipa, La Joya. 16 ° 22 ’ 58.52 S, 71 ° 44 ’ 4.08 ” W [- 16.383, - 71.734], 2000 m, 03. iv. 2022, G. Sarabia. All specimens are currently housed at PCYU but will be returned to MUSM upon completion of ongoing studies. See Fig. 7 for the geographic distribution of this species.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDEBF67FF78FF45FBD03556.taxon	etymology	Etymology: The specific epithet honours Arequipa, the Province of Peru where the specimens were collected, and the latin suffix - ensis, which translates to ‘ from Arequipa’. The name is masculine.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDBBF69FF78FF45FCD1366E.taxon	description	(Figs. 7, 26 – 38, 46, 48, 58, 63, 65, 82, 84, 93.)	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDBBF69FF78FF45FCD1366E.taxon	diagnosis	Diagnosis: Males of this species can be differentiated from all others in the genus by the combination of the two following characters: inner margins of compound eyes moderately convergent below, UOD: LOD ≤ 1.2 (Fig. 26), as opposed to ≥ 1.4 (compare Figs. 63 and 64); and metabasitarsus with dense patch of long hairs basoventrally (Fig. 27, 56), as opposed to dense hair patch absent (Fig. 55). Incasarus garciai, I. ancashensis and I. arequipensis also have UOD: LOD ≤ 1.2 but these species lack the metabasitarsal hair patch. Female of I. ivari can be differentiated from all other known females of Incasarus by the combination of IOC: OOC ratio of ~ 0.5 (Fig. 28), and strongly imbricate and dull surface sculpture of the clypeus (Fig. 29). Female I. ivari share the first feature only with I. aeneiventris, but can be differentiated from that species by the dull clypeus of I. ivari (compare Figs. 29, 93), as opposed to the somewhat shiny clypeus of I. aeneiventris (compare Figs. 93 and 94).	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDBBF69FF78FF45FCD1366E.taxon	description	Description. Holotype male: Dimensions: approximate body length ~ 5.16 mm; head width 1.66 mm; [forewing length 3.90 mm]; ITW 0.86 mm. Coloration: General integument black to dark brown, with subtle bronze reflections on the head and mesosoma, when observed from certain angles. The following parts yellow: mandible (except apex red); apical ~ 1 / 3 of clypeus (remainder brown). Ventral surface of flagellum dark yellow-brown. Posterior area of tegula and apical impressed areas of metasomal terga orange-brown. All tibiae brown except small orange-brown apicodorsal mark. Sculpture: Face below antennae moderately imbricate, somewhat shiny, yellow portion of clypeus weakly imbricate, somewhat shiny; remainder of face moderately to strongly imbricate and dull due to dense, almost crowded punctation. Clypeus punctures large, irregularly sparse (i = 0.5 – 2 d), except impunctate medially. Mid- and lower paraocular area moderately to densely punctate close to compound eye (i = 0.5 – 1 d), more sparsely punctate further from eye (i> 1.5 d). Genal and hypostomal areas weakly imbricate, shiny, moderately to densely punctate close to compound eye (i = 0.5 – 1 d), further away punctures large, shallower, and sparser (i> 3 d). Mesosoma strongly imbricate, dull. Mesoscutum punctures dense (i <0.5 d); scutellum punctures larger, sparser medially (i = 1 – 1.5 d); metanotum punctures crowded, edges sharp; sides of thorax coarsely imbricate. Hypoepimeral area and posterior portion of mesopleuron punctures moderately sparse (i = 1.5 – 2 d); anterior portion of mesopleuron punctures moderately dense (i = 1 – 1.5 d); metapleuron impunctate; propodeum dorsal surface weakly longitudinally striate, striae not attaining posterior margin except medially. Metasomal terga weakly imbricate, shiny; T 1 – T 2 punctures dense (i = 0.5 – 1 d); T 3 punctures smaller, moderately sparse (i = 1.5 – 2 d); T 4 – T 5 punctures scattered, small and shallow, almost imperceptible; T 6 – T 7 punctures larger and distinct apicomedially (i = 1 – 2 d). Apical impressed areas of T 1 – T 2 strongly imbricate, imbrication increasingly weak on more posterior terga. Metasomal sterna moderately imbricate, somewhat shiny; punctures shallow, difficult to detect. Pubescence: Generally white, branched, and sparse. Apicolateral margin of clypeus, scape, lower paraocular and hypostomal areas ~ 1.5 MOD, upper portion of clypeus <1 MOD. Vertexal and genal areas ~ 1 MOD. Mesopleuron ~ 1.2 MOD. Lateral areas of scutellum and propodeum hairs 1 – 1.5 MOD; pronotum, posteriorly on mesoscutum, anteriorly on metanotum, and laterally on dorsal surface of propodeum <1 MOD. Metatibia with thick, apicodorsally oriented hairs <4 MOD, metabasitarsus with dense patch of brown hairs basoventrally <1.3 MOD. S 2 – S 3 with erect, sparse, short hairs <0.5 MOD; S 4 – S 5 with sparse fringe of apicomedially oriented hairs ~ 1 MOD; S 6 with moderately dense apicomedially oriented hairs,> 2 MOD. Structure: Head: ~ 1.2 × as wide as long (84: 70). Mandible <3 × longer than basal depth (69: 24). Labrum ~ 2 × as broad as long (40: 20), labral process trapezoidal, narrowed apically, carinate laterally and apically. Clypeus ~ 2.3 × as wide as long (90: 39), convex in profile, strongly convex towards apex, angle between basal and apical portions in profile> 90 °; apicolateral margin convex in frontal view, apicomedial margin straight. Inner and outer subantennal sutures almost straight, subparallel, subantennal sclerite thus parallel-sided; anterior tentorial pit at junction of outer subantennal and epistomal sutures; epistomal suture transverse between inner subantennal sutures. Frontal line weak but distinct, flat above, raised and carinate between antennal sockets. IAD <AOD (20: 24). Inner margins of compound eyes moderately convergent below, UOD> LOD (55: 45). Facial fovea 5.5 × as long as greater width (55: 10), closer to compound eye ventrally. IOC <OOC (20: 35). Scape 2.5 × as long as greatest width (30: 12), longer than pedicel and F 1 combined (23); pedicel length and width equal (10: 10), F 1 longer than wide (13: 10); F 2 shorter than wide (7: 10), remaining flagellomeres shorter than wide (10: 14) except F 11 ~ 1.5 × as long as width (20: 13). \ Mesosoma: Mesoscutum ~ 1.3 × as wide as long (80: 60); length of scutellum: metanotum: dorsal surface of propodeum: 28: 16: 14. [Marginal cell ~ 0.8 × as long as distance between its apex to wing tip (40: 45)]. Metatibia ~ 5.5 × longer than greatest width (55: 10). Metabasitarsus produced basally, ~ 7 × longer than wide at midlength (50: 7). Metatibial spurs straight, posterior somewhat longer than anterior. Metasoma: Broadest at midlength of T 2. S 1 – S 5 unmodified. S 6 apicomedially concave, sides of concavity somewhat less than right angular. Terminalia: S 7. Apodeme long and broad, anterior margins mostly straight forming obtuse angle. Apical lobe fused to apodeme, broader at base than medial length, apex truncate, space between lobes narrowly U-shaped, with moderately dense apicomedially oriented hairs, hairs much shorter than width of lobe. S 8. Spiculum anterior margin concave, lateral margins diverging anteriorly; lateral lobe large, broadly rounded; apical lobe broad, subtriangular, lateral margins mostly straight except abruptly converging towards acute apex, covered in short, sparse, posterolaterally oriented hairs. Gonocoxa anterodorsally weakly convex, sides weakly convex, abruptly narrowed towards base of gonostylus, medial margins approximately straight; gonostylus fused to gonocoxa, narrow, ~ 2 × longer than medial width, apex rounded with short, moderately sparse hairs; penis valve somewhat narrow, lateral margins weakly sinuate, gradually narrowing to pointed apex, with short, sparse, posterolaterally oriented hairs. Allotype Female: Dimensions: Approximate body length ~ 5.16 mm; head width 1.61 mm; forewing length 3.96 mm; ITW 0.9 mm. Coloration: Generally black to dark brown. mandible orange except basal half brown and apex black. Tegula and apical impressed areas of metasomal terga translucent light brown. Pygidial plate red except apex black. Head and metasoma with weak bronze reflections. Sculpture: Face below facial fovea moderately imbricate, somewhat shiny; disc of clypeus punctures large, variable in spacing (i <1.5 d), many punctures coalesced, medial line impunctate; lower mid-paraocular area moderately densely punctate (i = 1.5 – 2 d), except dense (i <1.5 d) adjacent to compound eye very dense (i <1.5 d); remainder of face strongly imbricate to areolate, somewhat dull, punctures crowded. Genal and hypostomal area moderately imbricate, shiny; punctures shallow and moderately dense (i = 1 – 3 d). Thorax moderately imbricate, somewhat shiny. Mesoscutum, scutellum and metanotum punctation dense (i <0.5 d); mesopleuron punctures shallow, somewhat sparse (i = 1 – 3 d). Rest of pleura impunctate. Metasomal terga moderately to strongly imbricate, somewhat dull; T 1 punctures small, dense on disc (i = 1 – 1.5 d); T 2 – T 4 punctures minute, scattered, difficult to detect among imbrication. T 5 punctures large, very deep and increasingly dense toward apical margin (i decreasing from 3 d to 1 d). Apical impressed area moderately imbricate, somewhat dull. Metasomal sterna weakly to moderately imbricate, shiny; punctures large and deep. S 2 punctures on disc irregularly sparse (i = 1.5 – 3), dense subapically (i = 1 – 1.5 d). Lateral and subapical areas of S 1 – S 4 punctures dense, i = 1 – 2 d, medial portion punctures absent. Pubescence: Generally white, short-branched, sparse and short. Apicolateral margin of clypeus ~ 1.2 MOD, scape and hypostomal area ~ 1.8 MOD; vertexal area and mesopleuron ~ 1.5 MOD; pronotum and lateral areas of scutellum, posterior area of mesoscutum, anteriorly on metanotum and laterally on dorsal surface of propodeum ~ 1 MOD; <1.5 MOD on legs, except tibial scopa with curved and thick apicodorsally oriented hairs, increasing in length toward apex, 1 – 3 MOD, thicker and denser on dorsal margin than elsewhere. Metasoma generally with minute hairs <0.5 MOD; T 2 – T 4 increasingly longer and denser hairs at junction of disc and apical impressed area ~ 1.3 MOD. Pre-pygidial and pygidial fimbriae dense, branched, brown ~ 2 MOD. Structure: Head: ~ 1.2 × as wide as long (71: 58). Mandible <3 × as long as basal depth (29: 10). Labrum ~ 1.3 × as broad as long (20: 15), labral process trapezoidal, disc depressed medially, margins weakly defined, not carinate. Clypeus ~ 2 × as wide as long (38: 19); convex in profile, apicolateral margin straight in frontal view, apicomedial margin concave. Outer subantennal sutures almost straight, inner weakly arcuate, subantennal sclerite thus parallel-sided; anterior tentorial pit at junction of outer subantennal and epistomal sutures; epistomal suture transverse between inner subantennal sutures. Frontal line distinct, a groove above, raised and weakly ridged between antennal sockets, below. IAD slightly shorter than AOD (10: 12). Inner margins of compound eyes subparallel, UOD ~ LOD (46: 45). Facial fovea deep, long, 5 × as long as greatest width (15: 3), margins sinuate, closer to inner margin of compound eye below. IOC <OOC (9: 15). Scape ~ 3 × as long as greatest width (14: 5), subequal than pedicel and F 1 combined (11); pedicel length and width equal (4: 4), F 1 longer than wide (6: 4); F 2 length and width subequal (4: 4.5), remaining flagellomere lengths subequal to width (~ 5: ~ 5.5) except F 11 ~ 1.7 × as long as width (10: 6). Mesosoma: Mesoscutum ~ 1.2 × as wide as long (42: 34); length of scutellum: metanotum: dorsal surface of propodeum: 13: 7: 8. Marginal cell ~ 0.8 × as long as distance between its apex to wing tip (36: 42). Metatibia ~ 5.5 × longer than greatest depth (50: 9). Metabasitarsus length excluding apical process ~ 7 × depth at mid-length, (28: 4), apical process length 2 × depth at base. Metatibial spurs straight, posterior longer than anterior. Metasoma: Broadest at mid-length of T 2. Apical portion of pygidial plate subtriangular, swollen medially extreme apex rounded.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDBBF69FF78FF45FCD1366E.taxon	materials_examined	Material studied: BOLIVIA. Holotype ♂, allotype ♀, 7 ♂ paratypes and 1 ♀ paratype; La Paz, Achocalla, W. of Mecapaca, - 16.666, - 68.093, 3804 m, 05. iv. 2024, L. Packer. All specimens are currently housed at PCYU but will be returned to ANCB pending completion of ongoing research.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
03BD879EFFDBBF69FF78FF45FCD1366E.taxon	etymology	Etymology: The species is named in honour of Ivar Ortuño who not only drove the junior author to the type locality but, as an engineer, designed the road taken to get there.	en	Romero, Nora, Packer, Laurence (2026): Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae). Zootaxa 5828 (2): 151-195, DOI: 10.11646/zootaxa.5828.2.1, URL: https://doi.org/10.11646/zootaxa.5828.2.1
