identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B187DCFFC7FFC4FF398604FD41FEAD.text	03B187DCFFC7FFC4FF398604FD41FEAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites cagrii Mauss & Yildirim 2022	<div><p>Celonites cagrii Mauss &amp; Yildirim, 2022</p><p>Celonites cagrii Mauss &amp; Yildirim in Mauss et al. 2022a: 115, figs 2, 4f, 5f, 7c, 8c, h, i, 9b, e, g, i, 10f, 11f, 12f, 13f, 14f, ♀ ♂ (type locality: “[Turkey] TR-Erzurum <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=42.34235&amp;materialsCitation.latitude=40.648262" title="Search Plazi for locations around (long 42.34235/lat 40.648262)">Şenkaya-Akşar</a> [40.648262°N, 42.342351°E] … 1275 m … [on Heliotropium ellipticum Ledeb.]”), holotype, ♀, OLML (examined).</p><p>Material examined. PARATYPES: ARMENIA: Erevan [Yerevan], Monti desertici, Aighepat [Aygepat], 40 km SE, 23.VII.1963, 2 ♂ (dbM 5789, 5790), leg. A. Giordani Soika [MSNVE] .</p><p>Distribution. Armenia (Fig. 11); Turkey.</p><p>Trophic relationships. In Turkey, females and males of this species were collected at flowers of Heliotropium ellipticum Ledeb. ( Boraginaceae) (Mauss et al. 2022a). Probably narrowly oligolectic (sensu Müller &amp; Kuhlmann 2008) on the genus Heliotropium Tourn. ex L.</p></div>	https://treatment.plazi.org/id/03B187DCFFC7FFC4FF398604FD41FEAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.	Fateryga, Alexander V., Mauss, Volker, Proshchalykin, Maxim Yu., Maharramov, Mahir M. (2025): Taxonomic review of the pollen wasps (Hymenoptera: Vespidae: Masarinae) from the Caucasus, with the description of two new species, new nest and flower-visiting records, and a key to the species. Zootaxa 5716 (4): 514-540, DOI: 10.11646/zootaxa.5716.4.3, URL: https://doi.org/10.11646/zootaxa.5716.4.3
03B187DCFFC4FFC0FF398080FE4AFE41.text	03B187DCFFC4FFC0FF398080FE4AFE41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites carpenteri Fateryga & Mauss 2025	<div><p>Celonites carpenteri Fateryga &amp; Mauss, sp. nov.</p><p>(Fig. 1)</p><p>Diagnosis. The species is closely related to Celonites hamanni Gusenleitner, 1973 and C. hellenicus Gusenleitner, 1997 . The female of C. carpenteri can be distinguished from females of C. hamanni and C. hellenicus by the posterior margin of terga II–V, which is deflected upwards and not crenulate (Fig. 1B, D) (versus not deflected upwards but strongly to moderately crenulate in C. hamanni, see ZOBODAT 2025a, b, and not deflected upwards but weakly crenulate in C. hellenicus, see ZOBODAT 2025c). The female of C. carpenteri can also be distinguished from that of C. hamanni and C. hellenicus by the outline of the emargination between the lateral lamella and the postero-lateral process of the propodeum being at least 0.5× as broad as deep or even broader (Fig. 1G) (versus a distinctly narrower emargination in C. hamanni, see Gusenleitner 1973: 65, fig. 6c [note that the captions to the figures in Gusenleitner’s paper are misplaced], and C. hellenicus). The female of C. carpenteri differs from that of C. hellenicus (ZOBODAT 2025c) in the punctation on the frons, especially at the ocular sinus, which is less deep and distinct in C. carpenteri (Fig. 1E), and in the punctation on the mesosoma and the metasomal terga, which is coarser in C. carpenteri (Fig. 1A, D); in both characters C. carpenteri is similar to C. hamanni (ZOBODAT 2025a, b).</p><p>Description. Female. Body length (from head to posterior margin of tergum II) 6 mm (total body length 8–9 mm); forewing length 5.5 mm. Head in frontal view about 1.3× as wide as long (from top of median ocellus to ventral margin of clypeus). Proboscis much shorter than body. Clypeus about 1.25× as wide as long; its ventral emargination shallow, about 0.15× as deep as wide, taking about 0.45 of clypeal width, ventro-lateral teeth rounded. Distance between lateral ocellus and occiput about as long as distance between lateral ocelli and longer than distance between lateral ocellus and compound eye. Antenna with articles A8–A12 forming ventrally flattened club about 2.4× as long as wide in dorsal view. Width of gena in front of ocular sinus about equal to diameter of lateral ocellus. Preoccipital carina distinct along whole outer margin of compound eye. Pronotum with anterior side nearly vertical, roundly angled to dorsal surface. Pronotal humerus broadly rounded; in lateral view, pronotal carina distinctly separating antero-ventral area of pronotum from its dorsal area at antero-lateral sides. Scutellum separated from mesoscutum by transverse suture, convex, bluntly angled posteriorly. Axilla of scutellum with distinct lateral projection lying on tegula. Tegula with weak depression in front of lateral projection of axilla; outer side of tegula shallowly emarginate. Epipleural and episternal sulci indistinct. Ventral mesepisternum unevenly rounded ventrally. Epicnemial carina distinct and sharp. Mesepimeron produced posteriorly as small tubercle. Dorsal carinae of propodeum forming sharp lamellar tooth on medial end, adjacent to metanotum. Outline of emargination between lateral lamella and postero-lateral process of propodeum usually 0.5× (0.9 in one specimen) as broad as deep, with its outer side directed at angle of about 45° from posterior margin of lateral lamella and about two times shorter inner side parallel to outer one. Terga I–V medially with straight, not crenulate margins, laterally with tooth; margins of terga II–V deflected upwards. Tergum VI bluntly angulate posteriorly, laterally with blunt tooth. Sternum VI postero-medially with strong acute process. Basitarsus of foreleg about 3.3× as long as broad in dorsal view.</p><p>Clypeus shallowly and irregularly punctate with punctures of various sizes; interstices much narrower than largest punctures, shining, with indistinct microsculpture. Punctation on frons between antennal sockets similar to that on clypeus; punctures becoming larger towards ocular sinus and ocelli. Punctation on vertex behind ocelli denser than that on upper part of frons, interstices less shining. Gena nearly without distinct macropunctures, with reticulate microsculpture, rather dull. Pronotum, mesoscutum, and scutellum densely reticulately-punctate, punctures larger than those anywhere on head (largest punctures about 0.5× as wide as diameter of median ocellus), interstices much narrower than puncture diameter, shining, with indistinct microsculpture. Tegula with few dense macropunctures in anterior fourth, then with scattered micropunctures only, and with sparse deep macropunctures in posterior half; interstices shining. Dorsal and ventral mesepisterna and mesepimeron sculptured similarly to pronotum, mesoscutum, and scutellum. Posterior surface of metanotum with few shallow and indistinct macropunctures and scattered micropunctures; interstices shining. Metapleuron and lateral surface of propodeum dull, longitudinally wrinkled, without punctures. Posterior surface of propodeum sculptured similarly to pronotum, mesoscutum, and scutellum but interstices between punctures becoming larger (about 0.5 of puncture diameter) at lateral side of lateral lamella. Punctation on tergum I less coarse than that on pronotum, mesoscutum, and scutellum; interstices much narrower than puncture diameter on disc and about 0.5 of puncture diameter on depression. Terga II–V with punctures on disc similar to those on pronotum, mesoscutum, and scutellum; punctures becoming sparser and sometimes smaller on depression where some interstices reaching puncture diameter. Tergum VI with punctures similar to those on discs of terga II–V but diminishing in size towards apex. Sternum I with dull reticulate microsculpture only. Sterna II–V with scattered punctures of various sizes, interstices exceeding diameter of largest punctures, rather dull, with reticulate microsculpture. Sternum VI laterally with irregular coarse macropunctures, interstices less than puncture diameter, more shining than those on previous sterna, with indistinct microsculpture; medial line of sternum VI with longitudinal shining impunctate zone.</p><p>Mandible, labrum, and clypeus covered with thin pale brownish setae, reaching in length diameter of median ocellus. Frons and pronotum with two times shorter and sparser setae. Vertex between ocelli with few setae reaching in length diameter of lateral ocellus. Compound eye with very short dust-like setae. Mesoscutum, scutellum, and terga with short unnoticeable erect setae. Sterna mostly with short appressed setae, especially at posterior margins. Coxae with sparse setae as long as those on clypeus; femora with setae as long as those on frons. Tibia and tarsus of foreleg with stiff setae about as long as diameter of median ocellus, those setae on mid and hind legs much shorter. Pollen-collecting brush of stiff setae on inner surface of basitarsus of foreleg not modified. Spur of foretibia evenly curved and equipped with pointed spike at apical end, directed at about 90° from spur body.</p><p>Base color black. Following parts pale yellow: median spot on pronotum, spot on antero-lateral side of pronotum (absent in two specimens), short and narrow band along posterior side of pronotum near tegula, tegula (except antero-central part), lateral projection of axilla of scutellum, postero-median spot on scutellum (nearly reduced in one specimen), lateral side of lateral lamella of propodeum, posterior band on tergum I extended anteriorly at center and lateral sides, posterior bands on terga II–V extended anteriorly at center and replaced by brown at lateral sides (but not at teeth), small basal parts of tibiae. Following parts reddish-brown or ferruginous: distal half of mandible, ventral side of antennal club, antero-central part of tegula, posterior end of tergum VI, tibiae (except small basal parts), tarsi. Yellow color on body usually also outlined by reddish-brown. Wings strongly fuscous; veins and pterostigma dark brown.</p><p>Male. Unknown.</p><p>Material examined. HOLOTYPE: ♀, labeled “AZ, Ordubad, Bilav / 1050 m, 39°02'43"N / 45°49'07"E, 20.VI.2024 / Astrodaucus orientalis / Proshchalykin, Maharramov // Holotypus ♀ / Celonites carpenteri / Fateryga &amp; Mauss [red label] // Zoological Institute / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.81861&amp;materialsCitation.latitude=39.045277" title="Search Plazi for locations around (long 45.81861/lat 39.045277)">St. Petersburg</a> / INS_HYM_0000377 [pale yellow label]” [ZISP] . PARATYPES: AZERBAIJAN. Nakhchivan Autonomous Republic: Tirkesh [Shakhbuz, Türkeş], 26.VII.1937, 1 ♀, leg. M. Ryabov [ZISP] ; Ordubad, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.81861&amp;materialsCitation.latitude=39.045277" title="Search Plazi for locations around (long 45.81861/lat 39.045277)">Bilav</a>, 39°02'43"N, 45°49'07"E, 1050 m, on Astrodaucus orientalis, 20.VI.2024, 2 ♀ (dbM 6335), leg. M. Proshchalykin, M. Maharramov [CAFK, CVMM] .</p><p>Etymology. The new species is named after our colleague James M. Carpenter (American Museum of Natural History, New York, USA), the world’s leading expert on vespid wasps, in recognition of his great contribution to the taxonomy of the subfamily Masarinae .</p><p>Distribution. Azerbaijan (Nakhchivan AR) (Fig. 11).</p><p>Habitat and trophic relationships. This species was observed on dry mountain slopes with sparse herbaceous vegetation and scattered shrubs (Fig. 2A). Three females were collected foraging on inflorescences of Astrodaucus orientalis (L.) Drude ( Apiaceae). There are only a few species of pollen wasps that were previously recorded at flowers of the family Apiaceae: Gayella araucana Willink (Pérez 1989), Ceramius bischoffi Richards (Richards 1963), three species of the genus Jugurtia de Saussure (Bequaert 1940), and Celonites afer Lepeletier (Bequaert 1940) (summarized by Gess 1996). Thus, the only former record of a Celonites species at Apiaceae is that of C. afer, a species normally visiting flowers of the genus Echium Tourn. ex L. ( Boraginaceae) (Bequaert 1940; Mauss &amp; Müller 2014; Mauss et al. 2014). Bequaert (1940) reported C. afer visiting flowers of “ Bupleurum maritimum Linnaeus ”; however, such a taxon does not exist and this record probably refers to Crithmum maritimum L. The trophic relationship of C. carpenteri with Apiaceae, and particularly Astrodaucus orientalis, requires further study. The closely related Celonites hellenicus was recorded at flowers of Pallenis spinosa (L.) Cass. ( Asteraceae) (Fateryga et al. 2023).</p></div>	https://treatment.plazi.org/id/03B187DCFFC4FFC0FF398080FE4AFE41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.	Fateryga, Alexander V., Mauss, Volker, Proshchalykin, Maxim Yu., Maharramov, Mahir M. (2025): Taxonomic review of the pollen wasps (Hymenoptera: Vespidae: Masarinae) from the Caucasus, with the description of two new species, new nest and flower-visiting records, and a key to the species. Zootaxa 5716 (4): 514-540, DOI: 10.11646/zootaxa.5716.4.3, URL: https://doi.org/10.11646/zootaxa.5716.4.3
03B187DCFFC0FFCEFF3980ECFF32FD39.text	03B187DCFFC0FFCEFF3980ECFF32FD39.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites ivanovi Mauss & Fateryga 2022	<div><p>Celonites ivanovi Mauss &amp; Fateryga, 2022</p><p>(Figs 4, 10B, K)</p><p>Celonites ivanovi Mauss &amp; Fateryga in Mauss et al. 2022a: 130, figs 2, 4e, 5b, d, e, 6b, c, 7g, 8b, f, g, 9a, d, f, h, 10e, 11e, 12e, 13e, 14e, 18a–e, g, h, ♀ ♂ (type locality: “[Russia] Dagestan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.97028&amp;materialsCitation.latitude=42.601944" title="Search Plazi for locations around (long 46.97028/lat 42.601944)">Maydanskoye</a> 42°36'16"N 46°58'10"E [corrected to 42°36'07"N, 46°58'13"E in 2021] … on Heliotropium styligerum ”), holotype, ♀, OLML (examined).</p><p>Material examined. HOLOTYPE: ♀, labeled “ Dagestan, Maydanskoye / 42°36'16"N 46°58'10"E / on Heliotropium styligerum / 11.VI.2019 leg. Fateryga // ♀ Celonites sp. aff. cyprius / de Saussure, 1854 / det. Fateryga, 2019 // HOLOTYPUS / Celonites ivanovi MAUSS / &amp; FATERYGA, 2022 ♀ / designated by V. Mauss 2021 / db Mauss Nr.: 5492 [red label]” [OLML]. PARATYPES: RUSSIA. Dagestan: Untsukulskiy District, vicinity of Maydanskoye, on Heliotropium styligerum, 23.VI.2018, 2 ♀ (dbM 5287), leg. A. Fateryga [CAFK, CVMM]; Maydanskoye, 42°36'16"N, 46°58'10"E [actually 42°36'07"N, 46°58'13"E], 11.VI.2019, 8 ♂ (dbM 5497, 5498, 5496, 5499), leg. A. Fateryga [1 ♂, AMNH; 3 ♂, CAFK; 2 ♂, CVMM; 1 ♂, OLML; 1 ♂, ZISP]; ibid., on Heliotropium styligerum, 11.VI.2019, 10 ♀ (dbM 5494, 5490, 5491, 5493), 1 ♂ (dbM 5495), leg. A. Fateryga [1 ♀, AMNH; 5 ♀, CAFK; 3 ♀, 1 ♂, CVMM; 1 ♀, ZISP]; Maydanskoye, 42°36'07"N, 46°58'13"E, on Heliotropium styligerum, 15.VI.2021, 4 ♀ (dbM 5856, 5857, 5858, 5995), leg. A. Fateryga [CVMM]; ibid., 16.VI.2021, 1 ♂ (dbM 5996), leg. A. Fateryga [CVMM]; ibid., on Heliotropium styligerum, 16.VI.2021, 2 ♀ (dbM 5859, 5860), leg. A. Fateryga [CVMM]; Vicinity of Turtsi, 42°11'34"N, 47°09'33"E, on Heliotropium styligerum, 22.VI.2021, 1 ♀ (dbM 5997), leg. A. Fateryga [CVMM]. ADDITIONAL MATERIAL: RUSSIA. Dagestan: Maydanskoye, 42°36'07"N, 46°58'13"E, on Heliotropium styligerum, 28.V.2022, 1 ♀, leg. A. Fateryga [CAFK]; Vicinity of Maydanskoye, 42°35'39"N, 46°58'17"E, 19.VI.2023, 1 ♂, leg. A. Fateryga [CAFK]; Khotoch, 42°24'52"N, 46°57'10"E, 17.VI.2023, 1 ♂, leg. M. Proshchalykin [CAFK].</p><p>Distribution. Russia (Dagestan) (Fig. 11).</p><p>Biology. Four old nests of this species were found in the vicinity of Maydanskoye in Dagestan (42°35'39"N,</p><p>46°58'17"E) on 19.VI.2023. The nesting site was a steep rocky slope facing south; at the bottom of the slope, there was a shale scree covered with sparse herbs predominated by Heliotropium styligerum Trautv. ( Boraginaceae) (Fig. 3A). Three females (Fig. 4A) and one male of C. ivanovi were observed at this locality during 2 h. All four nests were attached to subvertical surfaces of medium-sized stones (Fig. 3B). The nests were made of fine clayey soil with a small proportion of sand grains and consisted of one to three cells. In the case of more than one cell, they were arranged in a longitudinal row in which each subsequent cell abutted with its basal end onto the apical end of the preceding cell. An additional nest covering was not present. Nest no. 1 consisted of one cell oriented vertically, with the entrance opening downwards (Fig. 3C); nest no. 2 consisted of one cell oriented diagonally, with the entrance opening rather upwards (Fig. 3E); nest no. 3 consisted of three cells oriented vertically, with the entrance opening downwards (Fig. 3F); nest no. 4 consisted of two cells oriented diagonally, with the entrance opening rather downwards (Fig. 3G). The lengths of the nests were 9.5, 7.5, 22, and 17 mm, respectively. The outer diameter of the cells was 3.5– 4 mm, with the narrowest part at the plug and the broadest one in 1–2 mm from the basal end. The thickness of the cell walls was about 0.2 mm. Each cell was sealed with a flat plug, also about 0.2 mm thick. The inner length of the cells was 6.0–6.5 mm. Outside around the cell plug, the cell walls continued and covered either the rounded bottom (basal end) of the next cell or the closing plug of the last cell, which had the same structure as the bottom of the cells. The bottom of the next cell or the closing plug were partially fused with the plug of the previous cell at the center and detached from it laterally, resulting in a small hollow space (Fig. 3D). In the case of the last (or a single) cell of the nest, the cell walls continued beyond the cell plug for 1–4 mm, forming a small rim, which had one or two deep notches and was slightly bent outwards (Fig. 3C).</p><p>Nest no. 1 contained an empty cocoon of Celonites, the first cell of nest no. 3 was broken and empty, while five other cells (nest no. 2, the second and the third cells in nest no. 3, and both cells in nest no. 4) contained empty cocoons of a cuckoo wasp, presumably of the genus Spintharina Semenov ( Hymenoptera: Chrysididae). The cocoon of Celonites was whitish and soft, covering the whole inner volume of the cell and hardly detachable from the cell walls, with fecal pellets in the basal third (Fig. 3D). Cocoons of Spintharina were only 3.5– 4 mm long; they covered somewhat more than a basal half of the cell walls and then terminated on the head end by a transverse cap, which was often separated from the remaining part of the cocoon by the emerged imago. The cocoons were also whitish but denser than that of Celonites, parchment-like and semitransparent (Fig. 3G), easy detachable from the cell walls. Separated caps of the cocoons were sometimes visible through the holes that had been made by the emerging cuckoo wasps in lateral walls of the cells (Fig. 3F). After emergence of the imagines, nest no. 1 and the first cell in nest no. 4 were reused by spiders that placed their egg cocoons inside of them.</p><p>Nests of five Palaearctic species of the genus Celonites were described previously: C. abbreviatus (Villers), C. mayeti Richards, C. fischeri Spinola, C. tauricus Kostylev, and C. jousseaumeii du Buysson (Lichtenstein 1869, 1875; Bellmann 1984; Mauss &amp; Müller 2014; Mauss et al. 2016, 2022b). In all cases, the nests consist of usually several rather cylindrical mud cells with thin walls that are attached to either stones or twigs. In most species, at least some cells are attached longitudinally to each other. Linearly arranged brood cells were observed in a few nests of C. abbreviatus (Lichtenstein 1869) and C. fischeri (Mauss &amp; Müller 2014), but in these nests at least a single cell was also attached longitudinally to the others in addition. An exclusively linear arrangement of the brood cells was previously only described in C. jousseaumei (Mauss et al. 2022b) . Taking into account the phylogenetic relationships of C. ivanovi and C. jousseaumei (Mauss et al. 2022 a, 2024; Fateryga et al. 2023), we can assume that this type of linear cell arrangement belongs to the ground pattern of the clade containing C. jousseaumei and the members of the C. cyprius -group (sensu Mauss et al. 2022a). Nests of C. ivanovi are very similar to those of C. jousseaumei, except that the outer cell surface shows a less distinct “fish scale” pattern than in the latter species (Mauss et al. 2022b). However, the slight difference in surface structure could be the result of an aging and weathering process, since the investigated cells of C. jousseaumei were freshly build, while the examined empty cells of C. ivanovi were at least one year old.</p><p>Cuckoo wasps of the genus Spintharina are known to be associated with the pollen wasp genus Celonites . Spintharina versicolor (Spinola) was recorded as a brood parasite of C. abbreviatus (Blüthgen 1961; Erlandsson 1972), while S. arnoldi (Brauns) and S. bispinosa (Mocsáry) were reared from brood cells of two different Afrotropical species of Celonites (Brauns 1913; Gess 1996). A cocoon with a larva of Spintharina innesi (du Buysson) was recently recorded in a brood cell of C. jousseaumei (Mauss et al. 2022b) . There are at least three species of Spintharina in the Caucasus, which are expected to be distributed in its Russian part (Rosa et al. 2019). It is worthy to mention that in the nests of C. ivanovi the cocoons of Spintharina sp. were not located inside of a hostcocoon, which is similar to the position of a cocoon of S. innesi in a nest of C. jousseaumei (Mauss et al. 2022b) . Pauli et al. (2018) found a close phylogenetic relationship between Spintharina and the Nearctic genus Chrysurissa Bohart, which also appears to parasitize predominately pollen wasps (Hicks 1927, 1929; Hungerford 1937; Parker 1967). This phylogenetic placement indicates that both genera are descendants of a last common ancestor which may already have exploited pollen wasps as hosts, suggesting an old and exclusive association with pollen wasps in this clade (Pauli et al. 2018; Mauss et al. 2022b). However, the cocoons of Chrysurissa densa (Cresson) were found inside of the cocoons of their host, Pseudomasaris vespoides (Cresson) (Hicks 1927) . This may indicate a difference between larval behaviors of the two cuckoo-wasp genera: the larva of Spintharina probably consumes a full-grown host larva before it starts cocooning, while the larva of Chrysurissa consumes the host as a prepupa or even pupa that has already finished spinning its cocoon. Cocoons of cuckoo wasps inside host cocoons are common in some other groups of the family Chrysididae, particularly in species associated with bees (Martynova 2020), and Ch. densa is also reported to parasitize bees besides the genus Pseudomasaris Ashmead (Hicks 1927, 1929).</p><p>Imagines of C. ivanovi exclusively visit flowers of Heliotropium styligerum (Mauss et al. 2022 a, 2024) (Fig. 4B). Females of this species demonstrate a special case of narrow oligolecty (sensu Müller &amp; Kuhlmann 2008): pollen collection on only one plant species in the absence of coflowering congenerics. Morphological and behavioral adaptations of C. ivanovi for pollen uptake from flowers of H. styligerum were described in detail by Mauss et al. (2024).</p></div>	https://treatment.plazi.org/id/03B187DCFFC0FFCEFF3980ECFF32FD39	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.	Fateryga, Alexander V., Mauss, Volker, Proshchalykin, Maxim Yu., Maharramov, Mahir M. (2025): Taxonomic review of the pollen wasps (Hymenoptera: Vespidae: Masarinae) from the Caucasus, with the description of two new species, new nest and flower-visiting records, and a key to the species. Zootaxa 5716 (4): 514-540, DOI: 10.11646/zootaxa.5716.4.3, URL: https://doi.org/10.11646/zootaxa.5716.4.3
03B187DCFFCEFFCEFF39820DFEC3FA62.text	03B187DCFFCEFFCEFF39820DFEC3FA62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites laetus Panfilov 1968	<div><p>Celonites laetus Panfilov, 1968</p><p>(Figs 9H, 10D, H)</p><p>Celonites laetus Panfilov, 1968: 40, fig. 3, ♀ (type locality: “ Нахичевань, окрестности Джульфы, пустынные предгорьЯ” [Azerbaijan, Nakhchivan, vicinity of Julfa, desert foothills]), holotype, ♀, ZMMU (examined).</p><p>Celonites phlomis Gusenleitner, 1973: 63, figs 3c, 4c, 5, ♀ ♂ (type locality: “Türkei, Urfa ” [Turkey]), holotype, ♀, OLML</p><p>(examined), synonymized by Fateryga 2025b: 210.</p><p>Material examined. HOLOTYPE: ♀, labeled “ Нахичевань [Nakhchivan] 3.VI.62 / ОКР. ДжУЛьфы / Д. ПанфиЛОв [vicinity of Julfa, D. Panfilov] // МОСКва / ЗООЛОГичеСКий / МУЗей МГУ [Moscow, Zoological Museum MSU] // Holotypus ♀ 1968 / Celonites / laetus Panf. [red label]” [ZMMU] . ADDITIONAL MATERIAL: AZERBAIJAN. Nakhchivan Autonomous Republic: Ordubad, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.81861&amp;materialsCitation.latitude=39.045277" title="Search Plazi for locations around (long 45.81861/lat 39.045277)">Bilav</a>, 39°02'43"N, 45°49'07"E, 1050 m, on Stachys inflata, 22.V.2025, 1 ♀, leg. M. Proshchalykin, M. Maharramov [CAFK] .</p><p>Distribution. Azerbaijan (Nakhchivan AR) (Fig. 11); Turkey.</p><p>Trophic relationships. One female was collected in the Nakhchivan Autonomous Republic of Azerbaijan at Stachys inflata Benth. ( Lamiaceae). In Turkey, females and males of this species were collected at flowers of Phlomis sp. ( Lamiaceae) (Gusenleitner 1973). Probably broadly oligolectic (sensu Müller &amp; Kuhlmann 2008) on Lamiaceae .</p></div>	https://treatment.plazi.org/id/03B187DCFFCEFFCEFF39820DFEC3FA62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.	Fateryga, Alexander V., Mauss, Volker, Proshchalykin, Maxim Yu., Maharramov, Mahir M. (2025): Taxonomic review of the pollen wasps (Hymenoptera: Vespidae: Masarinae) from the Caucasus, with the description of two new species, new nest and flower-visiting records, and a key to the species. Zootaxa 5716 (4): 514-540, DOI: 10.11646/zootaxa.5716.4.3, URL: https://doi.org/10.11646/zootaxa.5716.4.3
03B187DCFFCEFFCEFF3984CDFB5BF8BB.text	03B187DCFFCEFFCEFF3984CDFB5BF8BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites osseus Morawitz 1888	<div><p>Celonites osseus Morawitz, 1888</p><p>(Figs 9F, G, 10E, G)</p><p>Celonites osseus Morawitz, 1888: 268, ♀ (type locality: “in territorio transcaspico (Tschikischljar)” [Turkmenistan]), lectotype (Mauss et al. 2022a: 113), ♀, ZISP (examined).</p><p>Material examined. ARMENIA: Erivan [Yerevan], Nork [Nor Nork], 23.VI.1930, 1 ♀, leg. A. Shelkovnikov [ZISP]; Provinz Erivanj [Yerevan], Zanga [Hrazdan] River, 19.VII.1935, 1 ♀ (dbM 5855), leg. G. Kostylev [ZMMU] ; Ararat Province, Vedi, 9.VIII.1959, 2 ♀, leg. V. Rikhter [CAFK] . AZERBAIJAN. Nakhchivan Autonomous Republic: Ordubad, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.866665&amp;materialsCitation.latitude=39.066666" title="Search Plazi for locations around (long 45.866665/lat 39.066666)">Behrud</a>, 39°04'N, 45°52'E, 1345 m, 7.VIII.2020, 1 ♀, leg. M. Maharramov [CAFK] .</p><p>Distribution. Armenia, * Azerbaijan (Nakhchivan AR) (Fig. 11); Iran, Turkmenistan.</p></div>	https://treatment.plazi.org/id/03B187DCFFCEFFCEFF3984CDFB5BF8BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.	Fateryga, Alexander V., Mauss, Volker, Proshchalykin, Maxim Yu., Maharramov, Mahir M. (2025): Taxonomic review of the pollen wasps (Hymenoptera: Vespidae: Masarinae) from the Caucasus, with the description of two new species, new nest and flower-visiting records, and a key to the species. Zootaxa 5716 (4): 514-540, DOI: 10.11646/zootaxa.5716.4.3, URL: https://doi.org/10.11646/zootaxa.5716.4.3
03B187DCFFCFFFCFFF3981A1FE4AF913.text	03B187DCFFCFFFCFFF3981A1FE4AF913.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites tauricus Kostylev 1935	<div><p>Celonites tauricus Kostylev, 1935</p><p>(Figs 9I, 10A, F, I)</p><p>Celonites abbreviatus: Kostylev 1928: 399, ♀; 1929: 76, ♀.</p><p>Celonites abbreviatus tauricus Kostylev, 1935: 108, [♀] (type locality: “Крым” [Russia, Crimea]), neotype (Mauss et al. 2016: 36), ♀, OLML (examined).</p><p>Celonites tauricus: Mauss et al. 2016: 36.</p><p>Celonites spinosus Gusenleitner, 1966: 359, fig. 2, ♀ ♂ (type locality: “Kusadasi” [Turkey]), holotype, ♀, OLML (examined), synonymized by Mauss et al. 2016: 36.</p><p>Celonites abbreviatus invitus Gusenleitner, 1973: 58, fig. 2b, ♀ ♂ (type locality: “ Türkei, Gürün” [Turkey]), holotype, ♀, OLML (examined), synonymized by Mauss et al. 2016: 36.</p><p>Material examined. RUSSIA. Dagestan: Vicinity of Talgi, 42.8767°N, 47.4451°E, 25.VI.2018, 1 ♂, leg. Yu. Astafurova, K. Fadeev, V. Loktionov, M. Mokrousov, M. Proshchalykin [CAFK]; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.825&amp;materialsCitation.latitude=43.023888" title="Search Plazi for locations around (long 46.825/lat 43.023888)">Vicinity of Talgi</a>, 42°52'36"N, 47°26'42"E, on Teucrium canum, 12.VI.2019, 3 ♀ (dbM 5511), 2 ♂ (dbM 5510), leg. A. Fateryga [2 ♀, 1 ♂, CAFK; 1 ♀, 1 ♂, CVMM]; ibid., on Teucrium canum, 13.VI.2021, 1 ♀, leg. A. Fateryga [CAFK]; ibid., on Teucrium canum, 18.VI.2021, 1 ♀ (dbM 5998), leg. A. Fateryga [CVMM]; Khotoch, 42°24'52"N, 46°57'10"E, 17.VI.2023, 1 ♂, leg. A. Fateryga [CAFK]; Dubki, 43°01'26"N, 46°49'30"E, 21.VI.2023, 1 ♀, leg. A. Fateryga [CAFK] . GEORGIA: Kasbek, 1 ♀ [ZISP] . ARMENIA: Caucasus, Araxesthal, 2 ♀, leg. H. Leder, E. Reitter [ZISP]; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.81861&amp;materialsCitation.latitude=39.045277" title="Search Plazi for locations around (long 45.81861/lat 39.045277)">Parakar</a>, 2.V.1925, 1 ♀ [ZMMU]. AZERBAIJAN. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.81861&amp;materialsCitation.latitude=39.045277" title="Search Plazi for locations around (long 45.81861/lat 39.045277)">Main</a> territory: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.81861&amp;materialsCitation.latitude=39.045277" title="Search Plazi for locations around (long 45.81861/lat 39.045277)">Zuvand</a>, 8.VI.1985, 1 ♂, leg. V. Tobias [FSCV]. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.81861&amp;materialsCitation.latitude=39.045277" title="Search Plazi for locations around (long 45.81861/lat 39.045277)">Nakhchivan Autonomous Republic</a>: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.81861&amp;materialsCitation.latitude=39.045277" title="Search Plazi for locations around (long 45.81861/lat 39.045277)">Ordubad</a> (Arax), 1892, 2 ♀, leg. E. Reitter [ZISP]; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.81861&amp;materialsCitation.latitude=39.045277" title="Search Plazi for locations around (long 45.81861/lat 39.045277)">Julfa</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.81861&amp;materialsCitation.latitude=39.045277" title="Search Plazi for locations around (long 45.81861/lat 39.045277)">Gazanchi</a>, 39°13'N, 45°41'E, 1300 m, 15.VI.2019, 1 ♀, leg. M. Proshchalykin, Kh. Aliyev, M. Maharramov [FSCV]; Babek, Payiz, 39°26'06"N, 45°22'56"E, 1230 m, 25.VI.2024, 3 ♀, leg. M. Proshchalykin, M. Maharramov [FSCV]; ibid., on Nepeta trautvetteri, 28.VI.2024, 2 ♀, leg. M. Proshchalykin, M. Maharramov [CAFK]; Julfa, Daridagh-2, 39°03'58"N, 45°37'35"E, 1100 m, 24.V.2025, 1 ♂, leg. M. Proshchalykin, M. Maharramov [CAFK]; ibid., 27.V.2025, 1 ♀, 2 ♂, leg. M. Proshchalykin, M. Maharramov [1 ♀, 1 ♂, CVMM; 1 ♂, FSCV]; Ordubad, Bilav, 39°02'43"N, 45°49'07"E, 1050 m, on Nepeta trautvetteri, 27.V.2025, 2 ♀, 1 ♂, leg. M. Proshchalykin, M. Maharramov [CAFK]; ibid., on Nepeta trautvetteri, 29.V.2025, 8 ♀, 5 ♂, leg. M. Proshchalykin, M. Maharramov [2 ♀, 2 ♂, CVMM; 3 ♀, 1 ♂, FSCV; 3 ♀, 2 ♂, ZISP] .</p><p>Additional literature records. ARMENIA: Erevan [Yerevan], Monti desertici, Aighepat [Aygepat], 40 km SE (Gusenleitner 1973).</p><p>Distribution. Russia (Crimea, Dagestan), Georgia, Armenia, Azerbaijan (main territory, Nakhchivan AR) (Fig. 11); Greece (Chios, Kos, Rhodes, Samos), Turkey, Cyprus, Syria, Iran.</p><p>Biology. Females of this species construct nests on the underside of small stones. Nests consist of one to three cylindrical mud cells measuring 9 mm in length and 4 mm in outer diameter; cell walls are 0.25 mm thick. The brood cells are abutted longitudinally to each other. Complete nests are covered with an additional layer of mud (Mauss et al. 2016). Celonites tauricus is broadly oligolectic (sensu Müller &amp; Kuhlmann 2008) on Lamiaceae . Morphological and behavioral adaptations of C. tauricus for pollen uptake from nototribic flowers of Lamiaceae were described in detail by Mauss et al. (2016) and Fateryga et al. (2022). Flower-visiting records (all Lamiaceae): Satureja thymbra L. and Thymbra capitata (L.) Cav. (≡ Thymus capitatus (L.) Hoffmanns. &amp; Link) in Kos (Mauss et al. 2016); Teucrium canum Fisch. &amp; C.A. Mey. in Dagestan (Fateryga et al. 2022); Salvia nemorosa subsp. tesquicola (Klokov &amp; Pobed.) Soó, Teucrium capitatum L., T. chamaedrys L., Thymus tauricus Klokov &amp; Des.-Shost., Ziziphora capitata L., and Z. taurica M. Bieb. in Crimea (Mauss et al. 2016; Fateryga et al. 2022); Nepeta trautvetteri Boiss. &amp; Buhse in the Nakhchivan Autonomous Republic of Azerbaijan (new record); Ziziphora clinopodioides Lam. in Turkey (Mauss et al. 2016); Nepeta italica subsp. troodi (Holmboe) A.L. Budantzev (≡ Nepeta troodi Holmboe) in Cyprus (Fateryga et al. 2022).</p></div>	https://treatment.plazi.org/id/03B187DCFFCFFFCFFF3981A1FE4AF913	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.	Fateryga, Alexander V., Mauss, Volker, Proshchalykin, Maxim Yu., Maharramov, Mahir M. (2025): Taxonomic review of the pollen wasps (Hymenoptera: Vespidae: Masarinae) from the Caucasus, with the description of two new species, new nest and flower-visiting records, and a key to the species. Zootaxa 5716 (4): 514-540, DOI: 10.11646/zootaxa.5716.4.3, URL: https://doi.org/10.11646/zootaxa.5716.4.3
03B187DCFFCFFFCAFF39861AFD51FD4D.text	03B187DCFFCFFFCAFF39861AFD51FD4D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites tudesculptus Kostylev 1935	<div><p>Celonites tudesculptus Kostylev, 1935</p><p>(Fig. 5, 9J, 10C, J)</p><p>Celonites tudesculptus Kostylev, 1935: 109, [♀] (type locality: “АрмениЯ” [Armenia]), lectotype (Fateryga 2025b: 219), ♀, ZISP (examined).</p><p>Description of male (hitherto unknown). Body length (from head to posterior margin of tergum II) 4–5 mm (total body length 5–6 mm); forewing length 4–5 mm. Head in frontal view about 1.35× as wide as long (from top of median ocellus to ventral margin of clypeus). Proboscis much longer than body. Clypeus about 1.3× as wide as long; its ventral emargination very shallow and narrowly lamellate, taking about 0.5 of clypeal width, ventro-lateral teeth rounded. Distance between lateral ocellus and occiput about 1.1× as long as distance between lateral ocelli and about 1.9× as long as distance between lateral ocellus and compound eye. Antenna with articles A8–A12 forming ventrally flattened club about 1.7× as long as wide in dorsal view; ventral side of antennal club with three oval-shaped tyloids on A9, A10, and A11; tyloid of A10 taking whole length of article; tyloid of A11 smaller than that of A10 and larger than that of A9. Width of gena in front of ocular sinus about equal to diameter of lateral ocellus. Preoccipital carina distinct along whole outer margin of compound eye. Pronotum with anterior side nearly vertical, roundly angled to dorsal surface. Pronotal humerus angulately rounded; in lateral view, pronotal carina distinctly separating antero-ventral area of pronotum from its dorsal area at antero-lateral sides. Scutellum separated from mesoscutum by transverse suture, moderately convex, bluntly angled posteriorly. Axilla of scutellum with distinct lateral projection lying on tegula. Tegula with very weak depression in front of lateral projection of axilla; outer side of tegula rather rounded. Epipleural and episternal sulci obsolete. Ventral mesepisternum unevenly rounded ventrally. Epicnemial carina distinct and sharp. Mesepimeron produced posteriorly as small tubercle. Dorsal carinae of propodeum weakly recognizable, forming just indistinct tubercles on medial end, adjacent to metanotum. Outline of emargination between lateral lamella and postero-lateral process of propodeum small, comma-like, its medially directed apex just slightly enlarged. Terga I–VI medially with straight, not crenulate margins, laterally with tooth. Tergum VII trilobed; median lobe roundly truncate, lateral lobes tooth-like. Sternum VIII posteriorly broadly emarginate, with short acute median process. Genitalia as in Fig. 5D; harpide bifurcate apically; volsella directed medio-posteriorly; aedoeagus moderately narrowing towards apex.</p><p>Clypeus shining, with shallow sparse irregular punctures, interstices mostly exceeding puncture diameter. Punctation on frons and vertex coarse, interstices less than puncture diameter, moderately shining. Punctation on gena finer and sparser than that on frons and vertex. Pronotum, mesoscutum, and scutellum densely punctate with punctures slightly larger than those on frons and vertex, interstices narrower than puncture diameter, moderately shining. Tegula with moderately dense punctures and dull reticulately microsculptured interstices mostly not reaching puncture diameter.Dorsal and ventral mesepisterna and mesepimeron sculptured similarly to pronotum, mesoscutum, and scutellum. Posterior surface of metanotum dull, reticulately microsculptured, mostly without macropunctures. Metapleuron and lateral surface of propodeum dull, shagreened, without distinct punctures. Posterior surface of propodeum sculptured similarly to pronotum, mesoscutum, and scutellum but interstices between punctures becoming larger (about 0.5 of puncture diameter) at lateral side of lateral lamella. Punctation on metasomal terga about as coarse as that on pronotum, mesoscutum, and scutellum; punctures becoming sparser and sometimes smaller on depression where some interstices reaching puncture diameter. Sternum I with reticulate microsculpture only. Sterna II–VI with irregularly scattered punctures of various sizes, interstices exceeding diameter of largest punctures, rather dull, with distinct reticulate microsculpture. Sternum VII on distal half with band of scattered coarse punctures as well as small punctures.</p><p>Mandible and labrum covered with thin pale setae, reaching in length diameter of median ocellus. Clypeus, frons, vertex, and pronotum mostly with setae as long as 2/3 of diameter of median ocellus. Compound eye with very unnoticeable short dust-like setae. Mesoscutum, scutellum, and terga with short unnoticeable erect setae. Sterna mostly with short appressed setae, especially at posterior margins. Legs from coxae to femora with setae mostly shorter than those on clypeus, frons, vertex, and pronotum. Tibia and tarsus of foreleg with stiff setae about as long as diameter of median ocellus, those setae on mid and hind legs much shorter. Pollen-collecting brush of stiff setae on inner surface of basitarsus of foreleg not modified. Spur of foretibia curved, dorsally with large transparent lamella, apically blunt.</p><p>Base color black. Clypeus mostly whitish except two vertical black lines along lateral margins; often whitish coloration reduced to broad median vertical band (ten specimens of 18) or rarely to medio-ventral spot (two specimens) or medio-dorsal and medio-ventral spots (two specimens). Labrum often with whitish spot (nine specimens). Following parts whitish, often outlined by ferruginous: median spot on pronotum fused with narrow band along its posterior margin, spot on antero-lateral side of pronotum, lateral projection of axilla of scutellum, large spot on dorsal mesepisternum, lateral side of lateral lamella of propodeum, posterior bands on terga I–VI. Distal half of mandible, tyloids of antenna, tegula, metanotum, tergum VII, and legs from distal ends of femora ferruginous. Wings strongly fuscous; veins and pterostigma dark brown.</p><p>Material examined. LECTOTYPE (designated by Fateryga (2025b: 219)): ♀, labeled “[golden disc] // Caucasus. / Araxesthal. / Leder.Reitter. // Celonites ♀ / rudesculptus Typ / G. Kostylev det. 1932 // Lectotype ♀ / Celonites tudesculptus / Kostylev, 1935 / des. Fateryga, 2024 [red label] // Zoological Institute / St. Petersburg / INS_HYM_ 0000329 [pale yellow label]” [ZISP]. PARALECTOTYPES: ARMENIA: Caucasus, Araxesthal, 1 ♀, leg. H. Leder, E. Reitter [ZISP]; Caucasus, Armenisches Gebiet, 1 ♀, leg. H. Leder, E. Reitter [ZISP]. ADDITIONAL MATERIAL: AZERBAIJAN. Nakhchivan Autonomous Republic: Shakhbuz, Kolani, 39°26'51"N, 45°39'13"E, 1330 m, on Nepeta trautvetteri, 21.VI.2024, 6 ♀ (dbM 6336, 6337), leg. M. Proshchalykin, M. Maharramov [2 ♀, CAFK; 2 ♀, CVMM; 2 ♀, FSCV]; ibid., on Nepeta trautvetteri, 28.V.2025, 3 ♀, leg. M. Proshchalykin, M. Maharramov [FSCV]; Ordubad, Bilav, 39°02'43"N, 45°49'07"E, 1050 m, on Nepeta trautvetteri, 22.V.2025, 5 ♀, leg. M. Proshchalykin, M. Maharramov [4 ♀, CAFK; 1 ♀, FSCV]; ibid., on Nepeta trautvetteri, 27.V.2025, 19 ♀, leg. M. Proshchalykin, M. Maharramov [1 ♀, CAFK; 5 ♀, CVMM; 8 ♀, FSCV; 5 ♀, ZISP]; ibid., on Nepeta trautvetteri, 29.V.2025, 18 ♀, leg. M. Proshchalykin, M. Maharramov [FSCV]; Julfa, Daridagh-2, 39°03'58"N, 45°37'35"E, 1100 m, 24.V.2025, 7 ♀, 1 ♂, leg. M. Proshchalykin, M. Maharramov [1 ♂, CAFK; 7 ♀, FSCV]; ibid., 27.V.2025, 5 ♀, 5 ♂, leg. M. Proshchalykin, M. Maharramov [4 ♂, CAFK; 5 ♀, FSCV; 1 ♂, CVMM]; ibid., 29.V.2025, 7 ♀, 7 ♂, leg. M. Proshchalykin, M. Maharramov [2 ♂, CVMM; 7 ♀, 5 ♂, FSCV]; ibid., 30.V.2025, 2 ♀, 5 ♂, leg. M. Proshchalykin, M. Maharramov [2 ♀, FSCV; 5 ♂, ZISP].</p><p>Distribution. Armenia, * Azerbaijan (Nakhchivan AR) (Fig. 11).</p><p>Trophic relationships. Females were collected in the Nakhchivan Autonomous Republic of Azerbaijan at Nepeta trautvetteri ( Lamiaceae). Plants of this family are a common pollen source for the genus Celonites in the Palaearctic region (summarized by Gess 1996 and supplemented by Mauss 2006; Mauss et al. 2016; Fateryga et al. 2022, 2023; Fateryga 2025a). However, it was not documented whether C. tudesculptus collected pollen or nectar (or both) at N. trautvetteri, but the females have characteristically erected setae with ventro-medially curved distal ends on the clypeus that may facilitate pollen uptake from nototribic flowers, like Lamiaceae (cf. Müller 1996; Fateryga et al. 2022, 2023; Fateryga 2025a).</p></div>	https://treatment.plazi.org/id/03B187DCFFCFFFCAFF39861AFD51FD4D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.	Fateryga, Alexander V., Mauss, Volker, Proshchalykin, Maxim Yu., Maharramov, Mahir M. (2025): Taxonomic review of the pollen wasps (Hymenoptera: Vespidae: Masarinae) from the Caucasus, with the description of two new species, new nest and flower-visiting records, and a key to the species. Zootaxa 5716 (4): 514-540, DOI: 10.11646/zootaxa.5716.4.3, URL: https://doi.org/10.11646/zootaxa.5716.4.3
03B187DCFFCAFFCAFF398275FEB4F81B.text	03B187DCFFCAFFCAFF398275FEB4F81B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceramius caucasicus Andre 1884	<div><p>Ceramius caucasicus André, 1884</p><p>(Fig. 9A, D)</p><p>Ceramius caucasicus André, 1884: 820, ♂ (type locality: “ Caucase ” [presumably Armenia]), type depository unknown, probably Muséum National d’Histoire Naturelle, Paris, France (Carpenter 2001).</p><p>Material examined. ARMENIA: Etschmiadsin [Vagharshapat], 1 ♂ [ZISP]; Arsakiand [Arzakan], 22.VI.1904, 1 ♀, leg. N. Briansky [ZISP]; bei Erivan [Yerevan], 6300', 2.VII.1912, 2 ♂ [ZISP]; prope Mis-chana [Hankavan], 7000', 1924, 1 ♂ [ZISP]; Vallis Araxis, Mt. Saraj-bulag [8 km NE Armash], 5000', 8.VI.1926, 1 ♂, leg. A. Shelkovnikov [ZMMU]; Kotaik [Kotayk], prope Eilar [Abovyan], 1 ♂, leg. A. Shelkovnikov [ZMMU]; Kotaik [Kotayk], Mt. Gadis [Hatis], 6000', 1.VII.1926, 9 ♂, leg. A. Shelkovnikov [ZMMU]; Kotaik [Kotayk], Jelidja [4 km E Zovashen], 7100', 24.VII.1926, 4 ♀, 7 ♂, leg. A. Shelkovnikov [ZMMU]; Gokcha [Lake Sevan], northern coast, 19.VII.1927, 1 ♂, leg. Arnoldi [ZISP]; Karabakhlar [Vedi], 3.VI.1959, 2 ♂, leg. G. Viktorov [ZMMU]; Dzhrvezh [Jrvezh], 6.VI.1959, 1 ♂, leg. G. Viktorov [ZMMU]; Ashtarak District, Byurakan, 16.VII.1959, 1 ♂, leg. G. Viktorov [ZMMU] . AZERBAIJAN. Main territory: Adshikent [Hajikend], 2 ♀, 3 ♂ [ZISP]. Nakhchivan <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.9&amp;materialsCitation.latitude=39.1" title="Search Plazi for locations around (long 45.9/lat 39.1)">Autonomous Republic</a>: Ordubad (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.9&amp;materialsCitation.latitude=39.1" title="Search Plazi for locations around (long 45.9/lat 39.1)">Arax</a>), 1892, 1 ♀, 1 ♂, leg. E. Reitter [ZISP]; Ordubad, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.9&amp;materialsCitation.latitude=39.1" title="Search Plazi for locations around (long 45.9/lat 39.1)">Aghdara</a>, 39°06'N, 45°54'E, 2000 m, 17.VI.2019, 1 ♂, leg. M. Proshchalykin, Kh. Aliyev, M. Maharramov [CAFK] .</p><p>Additional literature records. ARMENIA: Erivan [Yerevan] and Kalp [Koghb] (Richards 1962); Kotaik [Kotayk], Keilan [probably Kuyli, 3 km NE Zovashen] (Kostylev 1929) .</p><p>Distribution. Armenia, Azerbaijan (main territory, Nakhchivan AR) (Fig. 11); Turkey, Iran.</p><p>Biology. Nesting of this species was studied in Turkey by Mauss et al. (2005). The only discovered nest was excavated by the female wasp in clayey but quite crumbly soil; the burrow was surmounted by a turret constructed by the female using mud extracted from within the burrow. The nest had a vertical to sub-vertical main shaft of approximately 13 cm length. At its base the main shaft turned outwards and continued into a short sub-horizontal secondary shaft terminated by a cell. The cell was sub-horizontal; there was no constructed mud cell within the excavated cell. Water was used to soften the soil; females visited water collection sites where they stood on wet ground, on or in dense vegetation, or on soil covered by a thin film of water. The provision consisted of pollen from plants of the families Campanulaceae and Lamiaceae . Pollen of the same two families was also present in the alimentary tract of females and males. Furthermore, the alimentary tract of males contained also pollen of Resedaceae, Fabaceae, and Caprifoliaceae (= Dipsacaceae) in lower proportions (Mauss et al. 2005). Therefore, C. caucasicus is a polylectic species. Flower-visiting records in Turkey: Ziziphora clinopodioides ( Lamiaceae), Papaver orientale L. ( Papaveraceae), and Acantholimon venustum Boiss. ( Plumbaginaceae) (Fahringer 1922; Mauss et al. 2005).</p></div>	https://treatment.plazi.org/id/03B187DCFFCAFFCAFF398275FEB4F81B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.	Fateryga, Alexander V., Mauss, Volker, Proshchalykin, Maxim Yu., Maharramov, Mahir M. (2025): Taxonomic review of the pollen wasps (Hymenoptera: Vespidae: Masarinae) from the Caucasus, with the description of two new species, new nest and flower-visiting records, and a key to the species. Zootaxa 5716 (4): 514-540, DOI: 10.11646/zootaxa.5716.4.3, URL: https://doi.org/10.11646/zootaxa.5716.4.3
03B187DCFFCBFFCBFF3981E9FEE2FB72.text	03B187DCFFCBFFCBFF3981E9FEE2FB72.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Jugurtia eurycara Kostylev 1935	<div><p>Jugurtia eurycara Kostylev, 1935</p><p>(Figs 9B, E, 10L)</p><p>Jugurtia eurycara Kostylev, 1935: 97, figs 6, 7, ♀ ♂ (type locality: “Caucasus, Armen. Gab.” [Armenia]), lectotype (Fateryga 2025b: 221), ♀, ZISP (examined).</p><p>Material examined. LECTOTYPE (designated by Fateryga (2025b: 221)): ♀, labeled “[golden disc] // Caucasus. / Armen. Geb. / Leder.Reitter. // Jugurtia ♀ / eurycara Typ / G. Kostylev det. 1932 // Lectotype ♀ / Jugurtia eurycara / Kostylev, 1935 / des. Fateryga, 2024 [red label] // Zoological Institute / St. Petersburg / INS_HYM_0000315 [pale yellow label]” [ZISP]. PARALECTOTYPES: ARMENIA: Caucasus, Armenisches Gebiet, 1 ♂, leg. H. Leder, E. Reitter [ZISP]; Provinz Erivanj [Yerevan], 3.VII.1932, 5 ♀, 1 ♂, leg. G. Kostylev [ZMMU]; ibid., 16.VII.1932, 1 ♀, leg. G. Kostylev [ZISP]; ibid., 18.VII.1932, 1 ♀, leg. G. Kostylev [ZMMU]. ADDITIONAL MATERIAL: AZERBAIJAN. Nakhchivan Autonomous Republic: Babek, 3 km NE Sirab, 39°18'N, 45°32'E, 1250 m, 12.VI.2019, 1 ♀, leg. M. Proshchalykin, Kh. Aliyev, M. Maharramov [CAFK]; Babek, Payiz, 39°26'06"N, 45°22'56"E, 1230 m, 25.VI.2024, 1 ♀, leg. M. Proshchalykin, M. Maharramov [CAFK]; ibid., 28.VI.2024, 1 ♀, leg. M. Proshchalykin, M. Maharramov [FSCV]; Julfa, Gazanchi-2, 39°15'04"N, 45°42'45"E, 1440 m, 24.VI.2024, 1 ♀ (dbM 6342), 1 ♂ (dbM 6343), leg. M. Proshchalykin, M. Maharramov [CVMM]; Ordubad, Bilav, 39°02'43"N, 45°49'07"E, 1050 m, 23.VI.2024, 1 ♂, leg. M. Proshchalykin, M. Maharramov [CAFK]; Shakhbuz, Badamli, 39°28'05"N, 45°33'00"E, 1290 m, 17.VI.2024, 2 ♂, leg. M. Proshchalykin, M. Maharramov [FSCV]; ibid., on Helichrysum rubicundum, 21.VI.2024, 3 ♂, leg. M. Proshchalykin, M. Maharramov [CAFK].</p><p>Distribution. Armenia, Azerbaijan (Nakhchivan AR) (Fig. 11); Turkey, Iran.</p><p>Trophic relationships. Males were collected in the Nakhchivan Autonomous Republic of Azerbaijan at flowers of Helichrysum arenarium subsp. rubicundum (K. Koch) P.H. Davis &amp; Kupicha (≡ H. rubicundum (K. Koch) Bornm., Asteraceae). Species of the family Asteraceae are common forage plants for several Afrotropical species of the genus Jugurtia (Gess 1996; Gess &amp; Gess 1989, 2010; Gess et al. 1997) and were also recorded to be visited by one Palaearctic species, namely J. oraniensis (Lepeletier) in Algeria (Bequaert 1940). However, trophic relationships of Palaearctic species of the genus Jugurtia with forage plants are poorly investigated and require a detailed study.</p></div>	https://treatment.plazi.org/id/03B187DCFFCBFFCBFF3981E9FEE2FB72	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.	Fateryga, Alexander V., Mauss, Volker, Proshchalykin, Maxim Yu., Maharramov, Mahir M. (2025): Taxonomic review of the pollen wasps (Hymenoptera: Vespidae: Masarinae) from the Caucasus, with the description of two new species, new nest and flower-visiting records, and a key to the species. Zootaxa 5716 (4): 514-540, DOI: 10.11646/zootaxa.5716.4.3, URL: https://doi.org/10.11646/zootaxa.5716.4.3
03B187DCFFCBFFCBFF39843AFC82F9F4.text	03B187DCFFCBFFCBFF39843AFC82F9F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Jugurtia irana Kostylev 1935	<div><p>Jugurtia irana Kostylev, 1935</p><p>(Figs 9C, 10M)</p><p>Jugurtia irana Kostylev, 1935: 99, ♀ (type locality: “Kuusha-Larumba, Bampur, S.-O. Persien ” [Iran]), lectotype (Fateryga 2025b: 222), ♀, ZISP (examined).</p><p>Material examined. AZERBAIJAN. Nakhchivan Autonomous Republic: Julfa, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.75&amp;materialsCitation.latitude=39.016666" title="Search Plazi for locations around (long 45.75/lat 39.016666)">Dize</a>, 39°01'N, 45°45'E, 880 m, 20.VI.2019, 3 ♀ (dbM 6341), leg. M. Proshchalykin, Kh. Aliyev, M. Maharramov [CAFK, CVMM, FSCV] .</p><p>Distribution. Azerbaijan (Nakhchivan AR) (Fig. 11); Iran.</p></div>	https://treatment.plazi.org/id/03B187DCFFCBFFCBFF39843AFC82F9F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.	Fateryga, Alexander V., Mauss, Volker, Proshchalykin, Maxim Yu., Maharramov, Mahir M. (2025): Taxonomic review of the pollen wasps (Hymenoptera: Vespidae: Masarinae) from the Caucasus, with the description of two new species, new nest and flower-visiting records, and a key to the species. Zootaxa 5716 (4): 514-540, DOI: 10.11646/zootaxa.5716.4.3, URL: https://doi.org/10.11646/zootaxa.5716.4.3
03B187DCFFCBFFD7FF3987E0FD2BFA9D.text	03B187DCFFCBFFD7FF3987E0FD2BFA9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov 2025	<div><p>Quartinia aliyevi Fateryga, Mauss, Proshchalykin &amp; Maharramov, sp. nov.</p><p>(Figs 6, 7)</p><p>Quartinia sp.: Fateryga et al. 2021: 39, ♀.</p><p>Diagnosis. The species is closely related to Quartinia araxana Giordani Soika, 1960, Q. cretica Gusenleitner, 1994, Q. libanica Richards, 1964, and Q. soikai Richards, 1964 . Both the female and the male of Q. aliyevi can be distinguished from those of Q. araxana by their larger size (total body length about 4 mm in Q. aliyevi versus 3 mm in Q. araxana), a denser macropunctation on the frons and shining interstices with a fine microsculpture (Figs 6D, 7D) (versus sparse macropunctures and dull interstices with a strong reticulate microsculpture in Q. araxana, Fig. 8B), denser and irregular macropunctures of various sizes and shining interstices with indistinct microsculpture on the mesoscutum (Figs 6A, 7A) (versus sparse regular macropunctures and dull interstices with a strong reticulate microsculpture in Q. araxana, Fig. 8A), and a longer tegula with a strongly produced posterior end (Figs 6A, 7A) (versus a shorter one with less produced posterior end in Q. araxana, Fig. 8A). The female of Q. aliyevi has a less deeply emarginate ventral margin of the clypeus (Fig. 6D) than Q. araxana (Fig. 8B), while the male of Q. aliyevi has a longer clypeus (Fig. 7D) than that of Q. araxana (Fig. 8C). The male of Q. aliyevi has the foretibia most similar to that in Q. libanica (according to fig. 3 in Richards 1964), with a more developed bulge (Fig. 7F, G) than that in Q. araxana (Fig. 8D, E). The male genitalia of Q. aliyevi (Fig. 7E) have a broader basal sclerite and less slender parameres (each consisting of fused stipes + harpide) than those of Q. araxana (Fig. 8F).</p><p>The female of Q. aliyevi is most similar to that of Q. libanica (ZOBODAT 2025e), but in Q. aliyevi the frons is covered with moderately spaced distinct small flat macropunctures, in between a dense, finely reticulate microsculpture, whereas these macropunctures are indistinct and nearly absent in Q. libanica . The distribution of the macropunctures on the mesoscutum is quite similar in Q. aliyevi and Q. libanica being dense to moderately spaced along the antero-lateral margins and becoming sparse towards the center, but the macropunctures are larger and deeper in Q. aliyevi and the insterstices are more shining especially on the center, since the fine reticulation of the cuticula is weaker to nearly absent in Q. aliyevi . In Q. aliyevi, the interstices between the dense macropunctation on the dorso-lateral parts of the pronotum and the mesepisternum are raised to little rounded ridges resulting in a distinctly reticulate to horizontally striated appearance, whereas the interstices are only weakly raised in Q. libanica .</p><p>The female of Q. aliyevi is also rather similar to that of Q. cretica but as in Q. libanica, the macropunctures on the frons of Q. cretica females are also less distinct, the macropunctures on the mesoscutum are much finer, and the interstices on the dorso-lateral parts of the pronotum and the mesepisternum are only weakly raised. Moreover, the interstices on the mesoscutum of Q. cretica are only matt shining, owing to a more pronounced, finely reticulate microsculpture (ZOBODAT 2025d). In females of Q. soikai (ZOBODAT 2025f), the sculpture of the frons and the mesoscutum is somewhat similar to that of Q. araxana, with dull interstices with a strong reticulate microsculpture; therefore it can be also easily distinguished from Q. aliyevi . This is especially true, since in Q. soikai the macropunctation is even denser than in Q. araxana and the macropunctures on the mesoscutum are of various sizes.</p><p>Males of Q. cretica and especially Q. soikai (Mauss et al., in preparation) have a broader foretibia in view on the outside, with a more developed proximal bulge and a distally adjacent nearly semicircular deep emargination on its posterior side than in Q. aliyevi; the bulge continues into the distal concavity by a distinct, nearly right angle, while the transition between the bulge and the distal concavity is obliquely rounded forming a blunt angle in Q. aliyevi (Fig. 7F, G). Quartinia orientalis Gusenleitner, 1973, another species of this group, can be distinguished from Q. aliyevi (as well as from Q. araxana, Q. cretica, Q. libanica, and Q. soikai) by a significantly more extended yellow color pattern of the body (ZOBODAT 2025g, h).</p><p>Description. Female. Body length (from head to posterior margin of tergum II) 3 mm (total body length about 4 mm); forewing length 2.5 mm. Head in frontal view about 1.1× as wide as long (from top of median ocellus to ventral margin of clypeus). Clypeus about 1.35× as wide as long; its ventral emargination shallow, about 0.2× as deep as wide, taking about 0.45 of clypeal width, ventro-lateral teeth rounded. Distance between lateral ocellus and occiput about 1.1× as long as distance between lateral ocelli and about 1.2× as long as distance between lateral ocellus and compound eye. Antenna with articles A8–A12 forming club about 2.25× as long as wide in dorsal view. Width of gena in front of ocular sinus about equal to width of compound eye at ocular sinus. Occipital carina complete, forming indistinct obtuse angle at lower part of gena. Pronotum with anterior side nearly vertical, roundly angled to dorsal surface. Pronotal humerus broadly rounded; in lateral view, pronotal carina distinctly separating antero-ventral area of pronotum from its dorsal area at antero-lateral sides. Scutellum separated from mesoscutum by transverse suture, slightly convex, broadly rounded posteriorly. Tegula with posterior end strongly produced medio-posteriorly and narrowly rounded apically; outer side of tegula broadly rounded. Epipleural and episternal sulci weak. Ventral mesepisternum evenly rounded ventrally. Epicnemial carina indistinct. Dorsal carina of propodeum complete, roundly produced dorso-laterally; lateral carina of propodeum complete. Terga I–V and sterna II–V with translucent posterior lamella.</p><p>Clypeus dull, densely irregularly punctate, interstices much smaller than puncture diameter, nearly indistinct. Frons with regular rounded macropunctures more than five times smaller in diameter than median ocellus, interstices about as puncture diameter, shining, with fine reticulate microsculpture. Sculpture becoming denser and duller on vertex behind ocelli where interstices much narrower than puncture diameter. Gena shining, with smoothened indistinct maropunctures and wrinkles more or less parallel to lower part of occipital carina. Pronotum densely irregularly punctate, interstices smaller than macropuncture diameter, shining, with fine reticulate microsculpture. Mesoscutum irregularly punctate with macropunctures of various sizes, largest ones about 0.3× as wide as diameter of median ocellus; interstices usually about as puncture diameter on anterior half and exceeding puncture diameter on posterior half, shining, with indistinct microsculpture. Scutellum sculptured similarly to posterior half of mesoscutum but interstices even larger, reaching several puncture diameters. Tegula shining, without distinct punctures. Dorsal and ventral mesepisterna and mesepimeron sculptured similarly to pronotum. Posterior surface of metanotum rather shining, with smoothened micropunctures. Metapleuron and lateral surface of propodeum rather dull, with shagreened microsculpture, without macropunctures. Dorso-lateral and posterior surfaces of propodeum rather dull, with shallow smoothened macropunctures and microsculptured interstices. Metasomal terga with dull reticulate microsculpture and shallow smoothened macropunctures especially distinct on tergum I and hardly recognizable on other ones; interstices between macropunctures about as puncture diameter. Metasomal sterna I–V sculptured similarly to corresponding terga but macropunctures more distinct, well recognizable on all sterna. Sternum VI densely punctate, interstices between macropunctures less than puncture diameter, with reticulate microsculpture similar to that on previous terga.</p><p>Mandible and labrum covered with thin erect pale setae, exceeding in length diameter of median ocellus.Clypeus, frons, and vertex with short erect pale brownish setae less in length than diameter of median ocellus. Compound eye bare. Pronotum and mesoscutum with very short and sparse pale setae, shorter than those on clypeus, frons, and vertex. Scutellum, mesopleuron, and propodeum with short unnoticeable setae. Metasoma and legs from coxae to femora mostly with short appressed setae. Tibia and tarsus of foreleg with erect stiff setae about as long as diameter of median ocellus, those setae on mid and hind legs much shorter.</p><p>Base color black. Following parts pale yellow: ventral surface of antenna, anterior band on pronotum interrupted medially (sometimes largely reduced), tegula (except central part), postero-median spot on scutellum (often absent or largely reduced), dorsal surface of metanotum (except medial part), posterior bands on terga I–V; legs from distal end of femora (gradually replaced by brownish-yellow on tarsi). Yellow color on body usually outlined by reddish-brown. Mandible (except basal black part), labrum, central part of tegula, posterior surface of metanotum, and metasomal sterna more or less brown. Tergum VI brownish-yellow to ferruginous. Wings transparent; veins and pterostigma brown.</p><p>Male. Structure and size similar to female except following: head in frontal view about 1.2× as wide as long; clypeus about 1.2× as wide as long; its ventral emargination taking about 0.55 of clypeal width; distance between lateral ocellus and occiput about 0.85× as distance between lateral ocelli and about 0.95× as distance between lateral ocellus and compound eye; width of gena in front of ocular sinus less than width of compound eye at ocular sinus. Tergum VII posteriorly with two rounded lobes and deep emargination between them. Sternum VII+VIII shallowly concave, posteriorly tridentate, central tooth broader than lateral ones. Genitalia as in Fig. 7E. Foretibia with lateral bulge in proximal half and concavity in distal one; transition between bulge and this concavity rounded.</p><p>Sculpture and setation as in female except that clypeus finely and much densely punctate, interstices between macropunctures less than puncture diameter. Tergum VII dull, with strong reticulate microsculpture. Sternum VI sculptured as previous sterna; sternum VII+VIII sculptured as sternum VI in female. Coloration resembles that in female but mandible, labrum, and clypeus pale yellow. Tarsi mostly pale yellow as tibiae, not gradually replaced by brownish-yellow.</p><p>Material examined. HOLOTYPE: ♀, labeled “Azerbaijan, Nakhichevan AR / Ordubad, Nurgut / 39°13'N 45°53'E 1900 m / 29.VII 2018 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.883335&amp;materialsCitation.latitude=39.216667" title="Search Plazi for locations around (long 45.883335/lat 39.216667)">Proshchalykin</a>, / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.883335&amp;materialsCitation.latitude=39.216667" title="Search Plazi for locations around (long 45.883335/lat 39.216667)">Aliyev</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.883335&amp;materialsCitation.latitude=39.216667" title="Search Plazi for locations around (long 45.883335/lat 39.216667)">Maharramov</a> // Quartinia spec. ♀ / det. V. Mauss 17.02.2019 / db Mauss Nr.: 5294 // Holotypus ♀ / Quartinia aliyevi / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.883335&amp;materialsCitation.latitude=39.216667" title="Search Plazi for locations around (long 45.883335/lat 39.216667)">Fateryga</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.883335&amp;materialsCitation.latitude=39.216667" title="Search Plazi for locations around (long 45.883335/lat 39.216667)">Mauss</a>, / Proshchalykin &amp; Maharramov [red label] // Zoological Institute / St. Petersburg / INS_HYM_0000378 [pale yellow label]” [ZISP] . PARATYPES: ARMENIA: Caucasus, Araxesthal, 1 ♀, leg. H. Leder, E. Reitter [ZISP] . AZERBAIJAN. Nakhchivan Autonomous Republic: Shakhbuz, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.616665&amp;materialsCitation.latitude=39.35" title="Search Plazi for locations around (long 45.616665/lat 39.35)">Kulus</a>, 39°21'N, 45°37'E, 1395 m, 20.VI.2021, 2 ♀ (dbM 6347, 6348), 1 ♂ (dbM 6346), leg. M. Maharramov [CVMM] ; ibid., 28.V.2025, 1 ♀, leg. M. Proshchalykin, M. Maharramov [CVMM]; Julfa, Daridagh- 2, 39°03'58''N, 45°37'35"E, 1100 m, 22.VI.2024, 1 ♂, leg. M. Proshchalykin, M. Maharramov [CAFK]; ibid., 26.VI.2024, 1 ♀, leg. M. Proshchalykin, M. Maharramov [CAFK]; ibid., 22.V.2025, 2 ♀, leg. M. Proshchalykin, M. Maharramov [CAFK]; ibid., 24.V.2025, 1 ♀, leg. M. Proshchalykin, M. Maharramov [CVMM]; ibid., 29.V.2025, 2 ♀, leg. M. Proshchalykin, M. Maharramov [FSCV, ZISP]; ibid., 30.V.2025, 1 ♀, leg. M. Proshchalykin, M. Maharramov [FSCV]; Babek, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.38222&amp;materialsCitation.latitude=39.435" title="Search Plazi for locations around (long 45.38222/lat 39.435)">Payiz</a>, 39°26'06"N, 45°22'56"E, 1230 m, 25.VI.2024, 1 ♂, leg. M. Proshchalykin, M. Maharramov [ZISP] ; ibid., 28.VI.2024, 1 ♀, leg. M. Proshchalykin, M. Maharramov [CAFK]; Ordubad, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.62722&amp;materialsCitation.latitude=39.364166" title="Search Plazi for locations around (long 45.62722/lat 39.364166)">Bilav</a>, 39°02'43"N, 45°49'07"E, 1050 m, 26.VI.2024, 1 ♀, leg. M. Proshchalykin, M. Maharramov [FSCV]; Shakhbuz, Kulus-2, 39°21'51"N, 45°37'38"E, 1400 m, 28.V.2025, 1 ♀, 1 ♂, leg. M. Proshchalykin, M. Maharramov [1 ♂, FSCV; 1 ♀, ZISP] .</p><p>Etymology. The new species is named in memory of our colleague Khalid A. Aliyev (1949–2019) from the Institute of Zoology of the National Academy of Sciences of Azerbaijan (Baku, Azerbaijan), one of the collectors of the holotype.</p><p>Distribution. Armenia, Azerbaijan (Nakhchivan AR) (Fig. 11).</p><p>Habitat. Specimens of this species were collected with yellow pan traps on dry mountain slopes with sparse herbaceous vegetation and scattered shrubs (Fig. 2B).</p></div>	https://treatment.plazi.org/id/03B187DCFFCBFFD7FF3987E0FD2BFA9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.	Fateryga, Alexander V., Mauss, Volker, Proshchalykin, Maxim Yu., Maharramov, Mahir M. (2025): Taxonomic review of the pollen wasps (Hymenoptera: Vespidae: Masarinae) from the Caucasus, with the description of two new species, new nest and flower-visiting records, and a key to the species. Zootaxa 5716 (4): 514-540, DOI: 10.11646/zootaxa.5716.4.3, URL: https://doi.org/10.11646/zootaxa.5716.4.3
03B187DCFFD7FFD4FF398491FB16FDD1.text	03B187DCFFD7FFD4FF398491FB16FDD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quartinia araxana Giordani Soika 1960	<div><p>Quartinia araxana Giordani Soika, 1960</p><p>(Fig. 8)</p><p>Quartinia araxana Giordani Soika, 1960: 133, ♀, ♂ (type locality “Caucaso: Araxesthal” [Armenia]), lectotype (Giordani Soika 1973: 51), ♀, NHMW (examined).</p><p>Material examined. LECTOTYPE (designated by Giordani Soika (1973: 51)): ♀, labeled “Araxesthal / Reitter 1890 // dilecta Gr / det. F. Kohl. // HOLOTYPUS / Quartinia / araxana / sp. nov. [red label]” [NHMW] . PARALECTOTYPES: ARMENIA: Caucasus, Araxesthal, 2 ♀, 2 ♂, leg. E. Reitter [1 ♀, 1 ♂, MSNVE; 1 ♀, 1 ♂, NHMW] . ADDITIONAL MATERIAL: ARMENIA: Caucasus, Armenisches Gebiet, 2 ♀,leg. H.Leder, E.Reitter [ZISP, ZMMU] . AZERBAIJAN. Nakhchivan Autonomous Republic: Shakhbuz, Kulus-2, 39°21'51"N, 45°37'38"E, 1400 m, 23.V.2025, 2 ♀, leg. M. Proshchalykin, M. Maharramov [CAFK]; ibid., 28.V.2025, 52 ♀, 9 ♂, leg. M. Proshchalykin, M. Maharramov [2 ♀, 3 ♂, CAFK; 40 ♀, 2 ♂, FSCV; 5 ♀, 2 ♂, CVMM; 5 ♀, 2 ♂, ZISP]; Julfa, Daridagh-2, 39°03'58"N, 45°37'35"E, 1100 m, 27.V.2025, 1 ♀, leg. M. Proshchalykin, M. Maharramov [CAFK]; ibid., 29.V.2025, 2 ♀, leg. M. Proshchalykin, M. Maharramov [FSCV]; ibid., 30.V.2025, 1 ♀, leg. M. Proshchalykin, M. Maharramov [FSCV] .</p><p>Distribution. Armenia, * Azerbaijan (Nakhchivan AR) (Fig. 11); Turkey, Iran.</p><p>Remarks. Giordani Soika (1960) reported a series of three females and two males deposited in NHMW. Later (Giordani Soika 1973) he designated a female as the lectotype (as “lecto-olotipo”) and a male as the allotype (as “lecto-allotipo”) and indicated that both were stored in his collection in Venice. At present, there are two females and a male in NHMW and a female and a male in MSNVE. However, both specimens in MSNVE bear red labels “ PARATYPUS ”. At the same time, there is a female labeled as the “ HOLOTYPUS ” and a male labeled as the “ ALLOTYPUS ” in NHMW. Giordani Soika (1960) described another species of vespid wasps in the same paper, Xanthodynerus caucasicus Giordani Soika, 1960, based on the only available specimen (holotype), and also indicated that it was deposited in NHMW. In his later paper (Giordani Soika 1973), the holotype of X. caucasicus was mentioned as stored in Venice, similarly to the lectotype of Q. araxana; however, it is actually present in NHMW (Fateryga et al. 2025). Therefore, we believe that Giordani Soika (1973) mentioned his collection in Venice as the depository of both the lectotype of Q. araxana and the holotype of X. caucasicus by mistake, and the lectotype of Q. araxana designated by him is actually the female in NHMW with the label “ HOLOTYPUS ”.</p></div>	https://treatment.plazi.org/id/03B187DCFFD7FFD4FF398491FB16FDD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.	Fateryga, Alexander V., Mauss, Volker, Proshchalykin, Maxim Yu., Maharramov, Mahir M. (2025): Taxonomic review of the pollen wasps (Hymenoptera: Vespidae: Masarinae) from the Caucasus, with the description of two new species, new nest and flower-visiting records, and a key to the species. Zootaxa 5716 (4): 514-540, DOI: 10.11646/zootaxa.5716.4.3, URL: https://doi.org/10.11646/zootaxa.5716.4.3
