taxonID	type	description	language	source
03B287920C61FF92FD74AFD9C282FA7E.taxon	materials_examined	Examined material: Holotype: FINLAND: Male, Province of Enontekis Lappmark: Kuttanen, 68 ° 26 ’ N / 22 ° 43 ’ E, 4. VII. 2012, leg / det. Å. Lindelöw. The holotype will be deposited in Swedish Museum of Natural History (Stockholm, Sweden) SMNH. Paratypes: SWEDEN: 1 ex. Torne Lappmark, Årosjokk, 18. VI. 1969, Lars Huggert leg., SMNH; 5 exx., Torne Lappmark, Årosjokk. 67 ° 87 ' N / 19 ° 37 ', 15. VI. 2008, leg. et det. Å. Lindelöw, CÅL; 26 exx., Torne Lappmark, Årosjokk. 67 ° 87 ' N / 19 ° 37 ' 5. VI. 2009, leg. et det. Å. Lindelöw, CÅL; 1 ex. Torne Lappmark, Årosjokk, 67 ° 52 ’ N, 19 ° 22 ’ E, stems of Salix myrsinifolia, 15. VI. 2008, Å. Lindelöw leg. / det., CÅL; 3 exx. Torne Lappmark, Akkar. 67 ° 86 ' N / 19 ° 43 ', 5. VI. 2009, leg. et det. Å. Lindelöw, ZIN; NORWAY: 8 exx., Region of Troms and Finmark, Karasjok: Jergul, 69 ° 25 ’ N / 24 ° 58 ’ E (EIS 166), 3. VII. 2012, stems of Salix myrsinifolia, leg. / det. Å. Lindelöw, CÅL; 1 ex., Region of Troms and Finmark, Karasjok, Jergul, RT 90 7732 / 1858. 3. VII. 2012. Svartvide. Leg et det. Å. Lindelöw, CTK; FINLAND: 8 exx., Province of Enontekis Lappmark: Kuttanen, 68 ° 26 ’ N / 22 ° 43 ’ E, 4. VII. 2012, stems of Salix glauca (Ripvide), 1 - 2 cm thick, leg. et det. Å. Lindelöw, CÅL; 1 ex. Province of Enontekis Lappmark: Kuttanen, 68 ° 26 ’ N / 22 ° 43 ’ E, 4. VII. 2012, stems of Salix glauca (Ripvide), 1 - 2 cm thick, leg. / det. Å. Lindelöw, CTK.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C61FF92FD74AFD9C282FA7E.taxon	description	Description. Body: 1.95 – 2.15 mm long. Head: frons convex, with densely and rather shallow, round, partially fusing punctures. Epistoma impressed, surface reticulated and shining; impression is divided into two parts by small median tubercle; epistomal impression have large fusing punctures. Vertex reticulated, not aciculated. Frontal surface with sparse inclined hair-like yellow setae; setae more abundant at epistomal margin. Eyes oval, with few facets missing behind antennal insertion. Antennae brown, only pedicel yellow. Pronotum: 1.1 times wider than long, with rounded posterolateral angles, broadly rounded; 4 – 8 denticles at anterior margin, arranged in concentric, transverse rows, highest pronotum point behind middle. Pronotum densely punctured near base and at posterolateral angles, brightly shining, faintly reticulated between punctures and evidently reticulated at flat bottoms of individual points; punctured area is covered by abundant and rather short hairs with their apices oriented forward and reaching the base of adjacent hairs. Anterior tuberculated part of pronotum with hairs directed backwards, with longer hairs at anterior margin of pronotum and at epipleura, surface between crenulations also reticulated but shining. Scutellum: flat and shining, strongly punctured, covered with short hairs which are oriented backwards. Elytra: more than 1.5 times as long as wide, 1.8 times longer than pronotum, with developed humeral angles; sides subparallel at basal 2 / 3 and rather narrowly rounded posteriorly. Surface brightly shining, glossy, with very faint reticulation and with strongly developed striae of round punctures, densely set, with less than 1 / 2 of puncture diameter between individual points, interstriae slightly wider than striae, with 3 irregular rows of minute punctures, each puncture bearing short hair-like scale directed backwards; central row of interstrial scales somehow longer, but still less than distance between individual bristles in row. Most of elytra has long, erect and thin hairs. Declivity with inclined hair-like scales, rather gradual, occupying only posterior 1 / 3 of elytral length, with suture not elevated. Individual inclined scales at elytral declivity are strongly serrate at sides and are pointed at apex, recumbent scales at declivity are also strongly serrate. Legs: femur and tibia brown and tarsus are yellowish. A small 8 th abdominal tergum is present in males, but not females. This is a common feature of all former Cryphalini, including Trypophloeini.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C61FF92FD74AFD9C282FA7E.taxon	diagnosis	Diagnosis. Specimens of T. borealis sp. n. from Northern Fennoscandia differs morphologically only subtly from T. nitidus and T. dejevi. T. borealis sp. n. differs from T. nitidus by slightly larger size (1.67 – 2.25 mm vs. 1.7 – 1.9 mm in T. nitidus). The average length of T. borealis sp. n. was 2,03 mm. In long series of “ nitidus / striatulus ” (at BEM) body length was found to be 1.6 – 2.1 mm). The structure of the frons, which is usually medially impressed in T. nitidus, can be very variable.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C61FF92FD74AFD9C282FA7E.taxon	etymology	Etymology. The name borealis is a derivation of boreal, which means “ belonging to the north ”.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C61FF92FD74AFD9C282FA7E.taxon	biology_ecology	Notes on the biology and development of Trypophloeus borealis sp. n. Biology and development in T. borealis are very similar to T. nitidulus in North America (Furniss 2004, 2013), except different host tree choice. Newly emerged adults perform maturation feeding in cave-like galleries in the bark of living trees / bushes of Salix. After hibernation, some adults may leave the host tree and disperse. Suitable trees are colonised for reproduction. In some cases, this may occur in the same gallery in which hibernation took place. Colonisation seems to be related to parts of the tree, which have reduced vitality or damaged or dying tissue. Galleries have been found in the transition zones between dead and alive tissue. Such zones often occur after cracks and other damages caused by snow during the winter. T. nitidus mating is taking place at the bark surface. The female enters the bark first, joined by the male later. The Trypophloeus species are monogamous and only one female and one male are found in a gallery. The eggs are laid in a cluster (Furniss 2004) in the gallery. Thus, the female does not make a separate niche for each egg. This behaviour is characteristic in Cryphalinii (Wood 1982). The development seems to be irregular. In mid-June parent beetles and larvae were found together in the galleries and in early July, adults, pupae and larvae have been observed (Lindelöw and Kvamme 2013). We assume that the development time can be more than one year due to the short summers and harsh climate in the North. Since galleries can be found in small parts of trees it is likely that the species can develop for several generations in the same trunk when the area of dead tissue increases year by year. The first Scandinavian record was probably made by sweeping in June in the area where the species was rediscovered in Salix myrsinifolia (Lindelöw 2009). In Norway the host tree was also S. myrsinifolia. In Finland the species was found in 1 – 2 cm thick trunks of Salix glauca (Lindelöw and Kvamme 2013). S. myrsinifolia (= S. nigricans) is widely distributed in North Europe, eastwards to the Ob Valley. S. glauca is circumpolar and widely distributed, both in Palaearctic as well as Nearctic (Hultén & Fries 1986, Jonsell 2000). Orange tendrils of fungus protruded from the bark surface have been observed several times (Figure 16) (Lindelöw and Kvamme 2013). They resemble the Cytospora sp. that Furniss (2004) observed on the bark surface of Salix alaxensis (Andersson), which was colonised by T. nitidus. Any possible relation between the fungus and the beetles is unknown. Spores found on the surface of the pronotum of T. striatulus did not belong to Cytospora sp. (Furniss 2004).	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C65FF90FD76AC06C034F9DE.taxon	materials_examined	Holotype: ♀, // Alta 30.6. Ut. // 7599 e [in blue ink] Hopk W. Va [in black ink] // Det. Type [in blue ink] No. Hopk. 3 - 1 - 02 // Leg. Mounted [on blue paper] // Trypophloeus punctipennis Hopk. [in black ink] // punctipennis H. 3 [in black pencil] Glyptoderes punctatus n. sp. [in blue ink] Hopk. [on reverse side] // Glyptoderes punctipennis Hopk. [in black ink] Hopk. [printed] // // red rectangle Typ No. 7399, USNM. Trypophloeus nitidus Swaine, 1912 res. n. Additional material. CANADA: 5 exx. 1930 Karl Schedl 737, NHMW; 4 exx. 1930 Karl Schedl 759, NHMW; Ontario: 4 exx. Biscotasing, Alnus incana, 10. V. 1930, K. E. Schedl leg / det, NHMW; Yukon Territory: 5 exx. Elliot Lake, Salix, 25. IX. 1987, M. M. Furniss leg., BEM; USA, Alaska: 6 exx. Cantwell, Salix, 29. VII. 1967, M. M. Furniss leg. / det., BEM; 13 exx. 21 miles E of Cantwell, Salix, 30. VIII. 1967, M. M. Furniss leg. / S. L. Wood det., BEM; 12 exx. Kongakut Riv., Brooks Range, Salix, 26. VIII. 1974, M. M. Furniss leg. / S. L. Wood det., BEM; 3 exx. Shublik Springs. Cunning River, Host Salix sp., 8. X. 1976, M. M. Furniss leg., TAMU; 10 exx. Mount. Mc. Kinley N. P., Igloo camp, Salix, 29. V. 1977, M. M. Furniss leg. / det., BEM; 4 exx. Fairbanks, Salix, 30. VIII. 1977, L. Bordelon leg. / M. M. Furniss det., BEM; USA, Utah: 2 exx. S 36 T 12 NR 2 E, Logan Сanyon., el. 7,000 Ft. Salix scouleriana, 17. VII. 1946, S. L. Wood leg., SEM; 1 ex. Little Bery, Logan Canyon., el. 8,000 Ft., Salix scouleriana 17. V. 1947, S. L. Wood leg., SEM; 1 ex. Logan Canyon, Salix scouleriana, 28. V. 1949, S. L. Wood, SEM; USA, Idaho: 10 exx. Coeur d’Alene Alnus sp., 4. XII. 1916 Evenden JC Colr., BEM; 2 ex. Poison Creek, 20 mi SW Grandview, Salix 31. VII. 1967, M. M. Furniss leg., D. M. Anderson det., BEM; USA, Oregon: 15 exx. Hot Springs C. G. Hart Mtn. Refuge, Lake Co. Salix scouleriana 14. VIII. 1990 M. M. Furniss, J. B. Johnson leg, M. M. Furniss det., BEM Trypophloeus nitidus Swaine, 1912: 349 = T. striatulus auct., nec Mannerheim, 1853: 235 syn. n. = T. punctipennis Hopkins, 1915: 37 Examined material. Type material: Holotype: Trypophloeus striatulus (Mannerheim, 1853): circle of golden paper // “ Cryphalus striatulus Mnnhm (= Mannerheim), Kenai J. ” farther unreadable in ink handwriting, probably “ Russam ”, ZIN. Comment. The type of the species has been considered lost, at least not found in Mannerheim collection, Finland (Wood and Bright 1992). However, in 1853 both Alaska and Finland were still parts of Russian Empire and thus deposition of the holotype in main country collection, i. e. in the former capital of the Empire (St. Petersburg) was logical. Cryphalus striatulus is the original valid name proposed by Mannerheim for a new species. It was described and really belongs to genus Cryphalus and not to genus Trypophloeus as many authors thought. It is not a new synonym, but a senior name and Cryphalus ruficollis is a new junior synonym of Cryphalus striatulus.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C65FF90FD76AC06C034F9DE.taxon	materials_examined	Examined material. Type material: Holotype: Trypophloeus striatulus (Mannerheim, 1853): circle of golden paper // “ Cryphalus striatulus Mnnhm (= Mannerheim), Kenai J. ” farther unreadable in ink handwriting, probably “ Russam ”, ZIN.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C65FF90FD76AC06C034F9DE.taxon	discussion	Comment. The type of the species has been considered lost, at least not found in Mannerheim collection, Finland (Wood and Bright 1992). However, in 1853 both Alaska and Finland were still parts of Russian Empire and thus deposition of the holotype in main country collection, i. e. in the former capital of the Empire (St. Petersburg) was logical. Cryphalus striatulus is the original valid name proposed by Mannerheim for a new species. It was described and really belongs to genus Cryphalus and not to genus Trypophloeus as many authors thought. It is not a new synonym, but a senior name and Cryphalus ruficollis is a new junior synonym of Cryphalus striatulus.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C65FF90FD76AC06C034F9DE.taxon	materials_examined	Lectotype: ♀, 172 [in red ink on small rectangle] // Weymouth NS [in red ink] LECTOTYPE [in black ink, printed] CNC No 9276 // Trypophloeus nitidus Sw. [handwritten in black ink], female, CNCI; Paratype (Paralectotype) 1 ex., Cornell U., No. 292.2 // Trypophloeus nitidus SW. // on Alnus G. E. Sanders // Weymouth 1912 – n. 8, NHMW; 1 ex., 2476 // [silver rectangular piece of paper] // Cryphalus nitidus Swaine Type series S. L. Wood (19) 56 // Trypophloeus nitidus Sokanovský det., ZMMU.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C65FF90FD76AC06C034F9DE.taxon	distribution	General distribution. North America, including Canada (Newfoundland, Nova Scotia, Quebec, Yukon), Alaska and mountains of western USA (Colorado, Idaho, Minnesota, Oregon, Utah) (Wood and Bright, 1992).	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C65FF90FD76AC06C034F9DE.taxon	discussion	Comment. We were not able to find the reference indicating the introduction of the species to Hungary (cf. Knížek, 2011).	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C65FF90FD76AC06C034F9DE.taxon	biology_ecology	Host trees. Alnus crispa, A. rugosa, Salix scouleriana, S. alaxensis.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C65FF90FD76AC06C034F9DE.taxon	description	Re-descripion. Body: length 1.6 – 2.2 mm, 2.7 (2.4 – 3.0) times as long as wide, black or dark brown, brightly shining. Head: frons convex, with Y-shaped impression from upper level of eyes to antennal insertions; surface finely reticulated from upper level of eyes to vertex, shallowly punctured below; pubescence consisting of inconspicuous sparse hairs. Pronotum: widest at base, distinctly transverse, 0.65 – 0.75 as long as wide; summit slightly behind middle; asperities in front of summit, rather large and abundant, forming concentric rows in anterior half; anterior margin with four to eight contiguous teeth. Surface of lateral and posterior areas shining, punctures rather close, coarse and deep; covered with short hair-like setae, slightly longer ispiratete area. Elytra: glossy, 1.6 – 2.0 times as long as wide; punctures on striae and interstriae deeply impressed in anterior two thirds, becoming smaller and shallow near declivity; interstriae with small granules on declivity; surface covered with recumbent hair-like strial and interstrial setae, becoming more scale-like at the declivity, and with rows of erect elongated scales on interstriae throughout elytral length.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C65FF90FD76AC06C034F9DE.taxon	diagnosis	Diagnosis. Distinguished from other species due to reticulated surface of the upper portion of frons and vertex; from T. alni and T. dejevi by more weak punctures in basal half of pronotum and declivity, by elytral vestiture, and by brightly shining body. T. nitidus differs from the North American species T. populi, T. salicis, and T. thatcheri by strong pucturation of elytra organized in rows. These rows of punctures on elytra extending at least two-thirds from the elytral base to declivity. The scale-like pubescence is only present at the declivity.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C64FF8EFD50ADB9C0FDFE3E.taxon	materials_examined	Examined material: Trypophloeus alni (Lindemann, 1875). Type material: Lindemann’s syntypes of Cryphalus alni are not preserved in ZMMU where they were expected to be kept. However, some of Lindemann’s specimens were located in NHMW. Lectotype is designated here from the NHMW. Lectotype: ♀ Trypophloeus alni (Lindemann, 1875). Mosqua // Lindemann // Collect Türk. // Cryphalus alni Lind. Typ [in handwriting] // TYPUS [on red paper, printed] // Lectotypus Cryphalus alni Lindemann, 1875 designated by Mandelshtam (date marked on label as 2006, but in fact lectotype is designated only in the current paper) // Trypophloeus alni (Lindemann, 1875) det. Mandelshtam, 2006 NHMW; Paralectotypes, 7 ex. with same collecting label: Lindemann // Mosqua // NHMW.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C64FF8EFD50ADB9C0FDFE3E.taxon	materials_examined	Additional material. ESTONIA: 2 exx. Estland, Leliland, 10. VIII. 1937, J. Hristian, 113, NHMW. RUSSIA, Leningrad Province: 2 exx. Lesnoy, Petrograd, 25. IV. [1] 915, Spessivtsev leg., ZIN; 2 exx. Beloostrov, Sestra River Bank, riverflooded forest, forest with Alnus and Prunus padus, under bark of fallen Alnus incana thick branches, 23. IV. 1990, M. Yu. Mandelshtam leg., ZIN; 11 exx. Orzhitsy, Lomonosov District, park, under bark of fallen stem of Alnus incana, 20. VIII. 2001, M. Yu. Mandelshtam leg., ZIN; 1 ex. RUSSIA, Pskov Province, Sebezh District, Sebezh National Park, Osyno Forestry, under bark of Alnus incana cut stem, VII. 2002, Mandelshtam leg., ZIN; RUSSIA, Arkhangelsk Province: 12 exx. 1851 alder, Vel’sk, Vologda Government, 23. VIII. 1929, G. K. Pjatnitskiy leg., ZIN; 33 exx. under number 1855, 3. IX. 1929; 57 exx. under number 1854, ZIN; RUSSIA, Yaroslavl Province, 3 exx. Poshekhonsk forestry, Yaroslavl Government, standing dead willow, 20 years, 17. VII. 1928, G. Pjatnitskiy leg., ZIN; RUSSIA, Moscow Province: 1 ex. Bol’shevo Station, in the bark of stem of alder, 15. IV. 1925, S. Shorokhov leg., ZIN; 1 ex. ZMMU; 1 ex. Bol’shevo Station, in the bark of growing drying alder, 06. VI. 1925, S. Shorokhov leg., ZIN; 2 exx. Bol’shevo Station, in the bark of growing drying alder, 14. VI. 1925, S. Shorokhov leg., ZMMU; 2 exx. Bol’shevo Station, alder, 14. VI. 1926, ZIN; 2 exx. Zvenigorod Biological Station of Moscow State University, window trap, 23. V. 1981, M. Samkov leg. (V. Gratchev coll.), ZMMU; 1 ex. Zvenigorod Biological Station of Moscow State University, 30. VI. 1981, M. Samkov leg. (V. Gratchev coll.), ZMMU; 1 ex. Zagorsk District, Malinniki village, waterfall “ Gremyachiy ”, aspen stem, 22. V. 1982, A. Petrov leg., ZMMU; 1 ex. Zvenigorod, 22. III. 1989, M. Yu. Mandelshtam leg., ZIN; 3 exx. Morozky, window trap on the elm No. 2, 25. IV – 18. V. 1994, Nikitsky leg., ZMMU; 2 exx. Mozhaiskyi District, near village Osiritsi, window trap 5, 22. IV. – 28. V. 2009 Nikitsky leg., ZMMU; RUSSIA, Bryansk Province: 4 exx. Bryansk, Orlov Gouvernement, 22. VII. [1] 926, V. Stark leg., ZIN; RUSSIA, Permsk Province: 1 ex. Middle Ural, environment. of Gubakha, Molotov Province (nowadays Permsk Province), alder, water-flooded forest of Kos’va River, square. 265, 28. XI. 1952, Zinovjev leg., ZIN; 2 exx. environment. of Molotov (nowadays – Perm), Mysy village, 07. V. 1953, G. Zinovjev leg., ZIN; 9 exx. environment. of Molotov (nowadays – Perm), Mysy village, 13. V. 1953, G. Zinovjev leg., ZIN; 3 exx. Mysy village, Krasnok [utskiy] District, alder, 07. VI. 1953, G. Zinovjev leg., ZIN; RUSSIA, Kemerovo Province: 1 ex. Kemerovo, on willow, “ intermediate form between T. alni and T. rybinski ” [B. Sokanovskij note], ZMMU; 3 exx. Aspen Plateau, willow, M. Lurie leg.; “ Tr. alni v. rybinskyi Rtt. B. Sokanovskij det. 57 ”, ZIN; PATRIA IGNOTA: 2 exx. 1930 Karl Schedl 135, NHMW; 8 exx. 1930 Karl Schedl 741, NHMW.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C64FF8EFD50ADB9C0FDFE3E.taxon	distribution	General distribution: Austria, Belarus, Estonia, Finland, Germany, Norway, Poland, Switzerland, European part of Russia (Northern and Central Territories), West Siberia (Knížek 2011, Kvamme & Lindelöw 2014). Comments. Reports of the species from Italy is erroneous. It is based on incorrect assignment of Campo-Grosso to Italy, but in fact this locality is in Switzerland. T. alni is listed from China (Hua 2002, Knížek 2011), including Xinjiang and Sichuan. The general distribution indicates that this most probably is erroneous and might refer either to T. dejevi or to a still undescribed species (cf. Knížek 2011). No specimens from Ukraine were found in available collections.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C64FF8EFD50ADB9C0FDFE3E.taxon	biology_ecology	Host trees: Alnus glutinosa, A. incana. Much more common in A. incana.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C64FF8EFD50ADB9C0FDFE3E.taxon	discussion	In China T. alni is mentioned to infest Alnus and Populus davidiana (Hua 2002). However, since we evaluate the occurrence of T. alni in China to be doubtful, we do not delete this information until new documentation is available. Re-description. Body: small, slim and strongly elongated, 1.2 – 1.6 mm, 3.0 – 3.25 times as long as wide, dark brown to black. Frons: weakly convex, shallowly reticulated from upper level of eyes to vertex; vertex entirely reticulated. Frontal pubescence consists of inconspicuous sparse hairs, vertex glabrous. Pronotum: triangular of equal length and width at base, widest posterior of tuberculated area, summit at middle. Asperities in front of summit abundant, rather small, conical, their height is equal to width at base, apically sharpened, basically not fused, not arranged in regular rows. Number of tubercles at pronotal, anterior margin variable. Pronotal vestiture consists of short, fine hairs with hair-like scales intermixed, the latter more abundant nearby base. Setae directed backward at tuberculated area and forward at the posterior area. Elytra: as long as wide, 2.2 (2.08 – 2.33), just a little wider than pronotum (1.0 – 1.1), sides parallel on basal two thirds, narrowly rounded at apex. Striae distinctly impressed throughout their length, with elongated punctures; interstriae slightly raised. Strial punctures with short hair-like setae, interstriae with ground vestiture of recumbent hair-like setae replaced with scale-like setae at declivity and each with a single row of long erect hair-like scales.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C64FF8EFD50ADB9C0FDFE3E.taxon	diagnosis	Diagnosis. Rather similar to Trypophloeus dejevi and T. nitidus and sharing with these two species the reticulated (not aciculated) surface of the upper portion of frons and vertex. Can be distinguished from both species by more elongated elytra and by acute, not merging asperities at tuberculated area. T. rybinskii, one more species without spines and granules at male elytral declivity, has aciculated surface of the upper portion of frons, indistinct elytral striae and much stouter body. The smallest of the species in the species group (Figure 5).	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C7AFF8DFF8BACD9C09DFB3E.taxon	materials_examined	Type material: Lectotype: “ Sajan Mts., Margasan River, 22. VIII. 31, S. Deev leg. ” // “ Mixed forest, willow, stem, 56 ” // “ Trypophloeus dejevi Stark, types, d [et]. Stark, 1933 ” // “ Lectotypus Trypophloeus dejevi Stark, det. Michalski 1965 ”, ZIN; paralectotypes: 7 exx. with the same geographic label, one bearing an additional label “ Trypophloeus deevi Stark, nov. types ”; 2 exx. with labels “ Sajan Mts., Barun-Ingasun River (in Michalsky 1969, name of river is given erroneously as Barun-Intarun), 14. IX. 31., S. Deev leg. ”, “ Mixed forest, alder, stem 79 ” // “ Paralectotypus ”, ZIN.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C7AFF8DFF8BACD9C09DFB3E.taxon	discussion	Comment. Michalski designating the lectotype and used the latinized version of the term on the label, and his label is cited here in full. In addition to the types labeled by J. Michalski there are more specimens in V. N. Stark collection preserved at ZIN. Eight of these specimens have the same label as the lectotype, and 7 specimens carry the labels: “ Sajan Mts., Margasan River 24. VIII. 31, S. Deev leg. ”, “ Mixed forest, willow, stem 62 ”. Trypophloeus niger Stark, 1936. Type material: Lectotype: “ Vladivostok Reg., Maikhe forestry, 2. VIII. 1930 ”, “ 159 ”, “ species? from willow ”, “ Trypophloeus niger Stark, types d. Stark, 1933 ” “ Lectotypus Trypophloeus niger Stark. det. J. Michalski, 1965 ”, ZIN; paralectotype: “ Vladivostok Reg., Maikhe forestry, 2. VIII. 1930 ”, “ 159 ”, “ species? V. Stark det. 1931 ”.;? “ 159, bank of Maikhe River on willow Salix 2. VIII. 1930, Lyubarskyi ”, ZIN. Comment. The original description was based on only two syntypes. No more specimens of this species are known so far.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C7AFF8DFF8BACD9C09DFB3E.taxon	materials_examined	Additional material: RUSSIA, Irkutsk Province: 1 ex. Sajan Mountains, Margasan River, mixed forest, willow, stem 56, 22. VIII. 1931, S. Deev leg., IBSS; 1 ex, Sajan Mountains, Barun-Ingasun River, mixed forest, alder, stem 79, 14. IX [19] 31, IBSS; 1 ex. Baical (sic!), Sajan, 14. IX. 1931 Salix, Dejev leg., NHMW; 1 ex. Baikal, Salix, 16. IX. 1931, Cotype, Dejev leg., NHMW; 7 exx. Nizhneangarsk, alder, without date and collector name, ZMMU; RUSSIA, Chita Province: 4 exx., Kodar Mts., near Sul’han River, 1600 - 1900 m., 10 - 17. VII. 1997, A. Petrov leg., ZIN; RUSSIA, Kamchatka Province: 5 exx. 28. VII. 1958, Kurentsov et Kononov leg., IBSS; 2 exx. 13. VIII. 1958, Kurentsov leg., IBSS; 2 exx. 14. VIII. 1958, Kurentsov leg., IBSS; 3 exx. 17. VIII. 1958, Kurentsov et Kononov leg., IBSS; 4 exx. 27. VIII. 1959, D. Kononov leg., IBSS; 8 exx. 6. VII. 1960, Ivliev leg., IBSS; 7 exx. 8. VII. 1960, Kupyanskaya leg., IBSS; 12 exx. 9. VII. 1960 Lisichenko leg., IBSS; 6 exx. 11. VII. 1960, Ivliev leg., IBSS; 18 exx. 15. VII. 1960, Kupyanskaya leg., IBSS; 2 exx. 17. VII. 1960, Kupyanskaya leg., IBSS; 7 exx. 18. VII. 1960, Kupyanskaya leg., IBSS; 2 exx. without collector name, IBSS; 9 exx. 26. VII. 1960, Lisichenko leg., IBSS; 1 ex. 8. VIII. 1960, Lisichenko leg., IBSS; 14 exx. 8. VIII. 1960, Kupyanskaya leg., IBSS; 5 exx. 10. IX. 1960, Lisichenko leg., IBSS; additional 69 exx. without precise locality, collected in VII-IX 1958 – 1960 by different collectors, IBSS; RUSSIA, Sakhalin Province: 5 exx. Sakhalin, 06. VIII. [19] 51, G. Krivolutskaya leg., ZMMU; 53 exx. Uglegorsk Distrrict, environs of Kotan settlement, sea shore, from alder, 6. VIII. 1951, Krivolutskaya leg. (erroneously labeled as syntypes of Eocryphalus zachvatkini Krivolutzkaja, collected at same place and date), ISEA; 13 exx. Sakhalin, alder, without date, G. Krivolutskaya leg., ZMMU; RUSSIA, Amur Province, 1 ex. Zeiysky Nature Reserve, kordon 52 - nd km. 25. VII. 1978, leg. Zimenko (N. B. Nikitsky coll.), ZMMU; RUSSIA, Primorsk Territory: 3 exx. Lesozavodsk town, from aspen, Naberezhnaya street, 24. VIII. [19] 66, A. N. Kypyanskaya leg., IBSS; 5 exx. Vladivostok, on Populus tremula, 16. IX. 2000, M. Mandelshtam leg., ZIN.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C7AFF8DFF8BACD9C09DFB3E.taxon	distribution	General distribution: Russia: Eastern Siberia (Sajan Mts., Cisbaikalia, Transbaikalia), Sakhalin, Kamchatka, Amur Province, Primorsk Territory, Mongolia (Ulan-Bator, 8. VII. 1956, on willow, leg. V. P. Gretchkin, cf. Sokanovsky 1960), (Knížek 2011).	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C7AFF8DFF8BACD9C09DFB3E.taxon	biology_ecology	Host trees: Salix sachalinensis, Salix spp., Alnus sp., and Populus tremula.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C7AFF8DFF8BACD9C09DFB3E.taxon	description	Re-description. Body: length – 1.85 – 2.15 mm, 2.7 (2.6 – 2.9) as long as wide. Body black, faintly shining. Head: antennae brown, frons slightly impressed from the middle to epistoma; surface finely punctured, more densely at lower half; reticulated from upper level of eyes to vertex; vertex entirely reticulated. Pronotum: widest near base, summit slightly behind middle, distinctly elevated; asperities coarse and forming concentric rows in anterior half; anterior margin with six teeth; surface rough, less densely punctured at posterior half, covered with mixture of hair-like setae and scarce elongated scales. Elytra: 1.85 – 2.0 times as long as wide, strial punctures distinctly impressed throughout elytral length; interstriae with shallow punctures; covered with recumbent hair-like setae replaced with scale-like setae on declivity; 2 nd and 3 rd interstriae on declivity slightly raised. Legs: legs brown colored.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
03B287920C7AFF8DFF8BACD9C09DFB3E.taxon	diagnosis	Diagnosis. T. dejevi can be distinguished from closely related T alni and T. nitidus by stouter body, coarse asperities in anterior half of pronotum, distinct, deep punctures on elytra and pronotum, and by elytral vestiture.	en	Kvamme, Torstein, Mandelshtam, Michail, Salnitska, Мaria, Ojeda, Dario I., Lindelöw, Åke, I., D. (2021): A new cryptic Trypophloeus Fairmaire, 1864 species in Northern Fennoscandia (Coleoptera, Curculionidae) revealed by DNA analyses. Norwegian Journal of Entomology 68: 44-66, DOI: 10.5281/zenodo.15883364
