taxonID	type	description	language	source
03B487D9FFB81D78FF243DE53E37FD17.taxon	discussion	Discussion. As noted by D. Polhemus (2017), the genus Engytatus is pantropical in distribution, and has undergone a minor insular radiation within the Hawaiian Islands, where 11 endemic species are now known occur, including the new species described below (see Appendix 1). The genus also contains a widespread agricultural pest species, E. modestus (Distant), that has been accidentally introduced to Hawaii. For a more comprehensive discussion of the genus and its taxonomic history in Hawaii, readers are referred to D. Polhemus (2017).	en	Polhemus, Dan A., Nyberg, Ben (2026): Engytatus pali (Heteroptera, Miridae): a new species collected by drone from vertical cliffs on Kauai, Hawaiian Islands. Zootaxa 5853 (3): 415-424, DOI: 10.11646/zootaxa.5853.3.8, URL: https://doi.org/10.11646/zootaxa.5853.3.8
03B487D9FFB81D7DFF243EA038E3FD17.taxon	description	(Figs. 5 – 10)	en	Polhemus, Dan A., Nyberg, Ben (2026): Engytatus pali (Heteroptera, Miridae): a new species collected by drone from vertical cliffs on Kauai, Hawaiian Islands. Zootaxa 5853 (3): 415-424, DOI: 10.11646/zootaxa.5853.3.8, URL: https://doi.org/10.11646/zootaxa.5853.3.8
03B487D9FFB81D7DFF243EA038E3FD17.taxon	materials_examined	Type Material Examined. Holotype, male, HAWAIIAN ISLANDS, Kauai, Waimea Canyon, on steep vertical cliff below Waimea Canyon Lookout, 22.07504 ° N, 159.65431 ° W, 29 April 2025, collected from Schiedea waiahuluensis by B. Nyberg using aerial drone (BPBM). Paratypes: HAWAIIAN ISLANDS, Kauai: 3 males, same data as holotype but 3 April 2024 (BPBM).	en	Polhemus, Dan A., Nyberg, Ben (2026): Engytatus pali (Heteroptera, Miridae): a new species collected by drone from vertical cliffs on Kauai, Hawaiian Islands. Zootaxa 5853 (3): 415-424, DOI: 10.11646/zootaxa.5853.3.8, URL: https://doi.org/10.11646/zootaxa.5853.3.8
03B487D9FFB81D7DFF243EA038E3FD17.taxon	description	Description. Male with general form slender, elongate, parallel-sided; overall length 3.10 mm, length from tip of tylus to cuneal fracture 2.26, maximum width (across base of cuneus) 1.00. Coloration: General color pale yellowish green, eyes dark red, antennal segment I broadly fasciate with dark brown centrally (Fig. 10). Structural characters: Head length 0.51, width across eyes 0.44, pale yellowish green; eyes relatively small, dark reddish-black, length 0.17, width 0.10; vertex width 0.20, bearing scattered moderately long, semi-erect, pale setae. Antennae long, slender, segment I stouter than segments II – IV, narrowly pale yellow at basal and distal ends, central section broadly marked with dark brown, with 3 moderately long, dark, spine-like setae; segments II and III linear, more slender and elongate than segment I; segment IV weakly fusiform; segments II – IV uniformly pale brown, bearing semi-recumbent, pale setae with lengths not exceeding the diameter of the segment on which they arise; lengths of segments I – IV = 0.24, 0.76, 0.75, 0.35. Rostrum length 1.11, reaching slightly beyond apices of hind coxae, pale golden yellow, extreme tip dark. Pronotum length 0.36, width 0.73, uniformly pale yellowish green; anterior lobe demarcated by a shallow transverse sulcus, bearing scattered pale, erect, curving setae; posterior lobe with short, recumbent pale setae, with a few longer pale setae along posterior margin. Scutellum length 0.34, width 0.44, pale yellowish-green, bearing scattered short, recumbent pale setae. Hemelytra translucent, uniformly pale yellowish green; entire hemelytral surface set with scattered simple, recumbent pale setae, these setae somewhat denser on cuneus and along embolar margin; wing membrane including veins hyaline and translucent. Legs slender, elongate, predominantly pale yellowish-green, with tarsi golden brown; all leg segments clothed with very short, pale, recumbent setae; fore coxa with longitudinal row of 3 long, slender, pale, spine-like setae on anterior face; all femora slightly thickened, lacking spines, fore and middle femora with fringes of fine, erect setae on ventral faces; outer margins of middle and hind tibiae bearing scattered sets of paired black spines, lengths of these spines about 1.5 times the diameter of corresponding tibiae on which they occur, such spines absent on fore tibia; tarsi with segment I shortest, segment II longest, segment III shorter than II. Lengths of leg segments as follows: femur, tibia, tarsal 1, tarsal 2, tarsal 3 of fore leg, 0.75: 0.80: 0.06: 0.15: 0.12; of middle leg, 0.74; 0.93: 0.06: 0.17: 0.09; of hind leg, 1.07: 1.45: 0.06: 0.31: 0.11. Ventral surface predominantly pale green, clothed with short, recumbent pale setae, these setae becoming longer and more numerous adjacent to genital cavity. Male genitalia with right paramere slender and finger-like (Fig. 5); left paramere stout basally, with expanded, leaf-like distal arm (Figs. 6 – 8), basal lobe bearing an acuminate tuft of long, dark setae; proctiger bilobate, posterior lobe thicker with rounded apex, lateral lobe strongly recurved distally, apex slightly bulbous (Figs. 5, 6, 7, 9). Female: Unknown.	en	Polhemus, Dan A., Nyberg, Ben (2026): Engytatus pali (Heteroptera, Miridae): a new species collected by drone from vertical cliffs on Kauai, Hawaiian Islands. Zootaxa 5853 (3): 415-424, DOI: 10.11646/zootaxa.5853.3.8, URL: https://doi.org/10.11646/zootaxa.5853.3.8
03B487D9FFB81D7DFF243EA038E3FD17.taxon	biology_ecology	Host plant. Schiedea waiahuluensis W. L. Wagner, Weller, B. Nyberg, & A. K. Sakai (Caryophyllaceae), 2024. Ecological Notes. This species has been collected exclusively on S. waiahuluensis (Figs. 3, 4). Given the host plants’ habitat specificity on vertical dry rock cliffs, surveying and inventorying these areas is nearly impossible using traditional ground-based approaches. Engytatus pali has been observed or collected in March, April, and May, with no presence detected on October collections of S. waihuluensis. While S. waiahuluensis does flower yearround, its peak flowering season in the spring overlaps with the collections E. pali. Dense glandular pubescence covers S. waiahuluensis throughout the stems, leaves, and inflorescence, conforming to the preference of other Engytatus species for sticky plants.	en	Polhemus, Dan A., Nyberg, Ben (2026): Engytatus pali (Heteroptera, Miridae): a new species collected by drone from vertical cliffs on Kauai, Hawaiian Islands. Zootaxa 5853 (3): 415-424, DOI: 10.11646/zootaxa.5853.3.8, URL: https://doi.org/10.11646/zootaxa.5853.3.8
03B487D9FFB81D7DFF243EA038E3FD17.taxon	etymology	Etymology. The name “ pali ” refers to the word for “ cliff ” in the Hawaiian language, and is used as a noun in apposition describing the habitat of this species.	en	Polhemus, Dan A., Nyberg, Ben (2026): Engytatus pali (Heteroptera, Miridae): a new species collected by drone from vertical cliffs on Kauai, Hawaiian Islands. Zootaxa 5853 (3): 415-424, DOI: 10.11646/zootaxa.5853.3.8, URL: https://doi.org/10.11646/zootaxa.5853.3.8
