taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03B7D164FFE5FFC8F561FC0AFAFA3218.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004168/files/figure.png	https://doi.org/10.5281/zenodo.19004168	Fig. 1. Quapawjapyx osage Sendra gen. et sp. nov. A. Paratype, ♂ (TAMU-ENTO X1831026). B, D–E. Paratype, ♂ (TAMU-ENTO X1832015). C. Holotype, ♀ (TAMU-ENTO X1831147). A. Habitus. B. Last antennomere; placoid sensilla remarked by spotted line. C. Lacinia with interior laminae and mandible. D. Dorsal portion on third antennomere with pores. E. Metathoracic claws. Abbreviations: see Material and methods.	Fig. 1. Quapawjapyx osage Sendra gen. et sp. nov. A. Paratype, ♂ (TAMU-ENTO X1831026). B, D–E. Paratype, ♂ (TAMU-ENTO X1832015). C. Holotype, ♀ (TAMU-ENTO X1831147). A. Habitus. B. Last antennomere; placoid sensilla remarked by spotted line. C. Lacinia with interior laminae and mandible. D. Dorsal portion on third antennomere with pores. E. Metathoracic claws. Abbreviations: see Material and methods.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE5FFC8F561FC0AFAFA3218.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004174/files/figure.png	https://doi.org/10.5281/zenodo.19004174	Fig. 2. Quapawjapyx osage Sendra gen. et sp. nov. A–C. Paratype, ♂ (TAMU-ENTO X1832015). D. Paratype, ♂ (MUVHN-ZE8000). A. Head, dorsal view. B. Pronotum. C. Mesonotum. D. Prothorax, mesothorax and metathorax sternites. Abbreviations: see Material and methods.	Fig. 2. Quapawjapyx osage Sendra gen. et sp. nov. A–C. Paratype, ♂ (TAMU-ENTO X1832015). D. Paratype, ♂ (MUVHN-ZE8000). A. Head, dorsal view. B. Pronotum. C. Mesonotum. D. Prothorax, mesothorax and metathorax sternites. Abbreviations: see Material and methods.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE5FFC8F561FC0AFAFA3218.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004177/files/figure.png	https://doi.org/10.5281/zenodo.19004177	Fig. 3. Quapawjapyx osage Sendra gen. et sp. nov. A–D, F. Paratype, ♂ (TAMU-ENTO X1832015). E. Paratype, ♂ (MUVHN-ZE8000). A. Metanotum. B. First urotergite. C. Fifth urotergite. D. Eighth urotergite. E. Urite X, dorsal view. F. Cerci, proximal portion. Abbreviations: see Material and methods.	Fig. 3. Quapawjapyx osage Sendra gen. et sp. nov. A–D, F. Paratype, ♂ (TAMU-ENTO X1832015). E. Paratype, ♂ (MUVHN-ZE8000). A. Metanotum. B. First urotergite. C. Fifth urotergite. D. Eighth urotergite. E. Urite X, dorsal view. F. Cerci, proximal portion. Abbreviations: see Material and methods.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE5FFC8F561FC0AFAFA3218.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004184/files/figure.png	https://doi.org/10.5281/zenodo.19004184	Fig. 4. Quapawjapyx osage Sendra gen. et sp. nov., paratype, ♂ (TAMU-ENTO X1832015). A. Posterolateral angle of sixth urotergite. B. Posterolateral angle of seventh urotergite. C. Posterolateral angle of eighth urotergite.	Fig. 4. Quapawjapyx osage Sendra gen. et sp. nov., paratype, ♂ (TAMU-ENTO X1832015). A. Posterolateral angle of sixth urotergite. B. Posterolateral angle of seventh urotergite. C. Posterolateral angle of eighth urotergite.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE5FFC8F561FC0AFAFA3218.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004189/files/figure.png	https://doi.org/10.5281/zenodo.19004189	Fig. 5. Quapawjapyx osage Sendra gen. et sp. nov. A–C. Paratype, ♂ (MUVHN-ZE8000). A. Fifth urosternite. B. Eighth urosternite. C. Urite X, ventral view. Abbreviations: see Material and methods.	Fig. 5. Quapawjapyx osage Sendra gen. et sp. nov. A–C. Paratype, ♂ (MUVHN-ZE8000). A. Fifth urosternite. B. Eighth urosternite. C. Urite X, ventral view. Abbreviations: see Material and methods.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE5FFC8F561FC0AFAFA3218.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004193/files/figure.png	https://doi.org/10.5281/zenodo.19004193	Fig 6. Quapawjapyx osage Sendra gen. et sp. nov. A–C. Holotype, ♀ (TAMU-ENTO X1831147). D–F. Paratype, ♂ (TAMU-ENTO X1832015). A. Latero-posterior border of the first urosternite. B. Detail of lateral organ. C. Central organ of the first urosternite. D. Latero-posterior border of the first urosternite. E. Detail of lateral organ. F. Central organ of the first urosternite. Abbreviations: see Material and methods.	Fig 6. Quapawjapyx osage Sendra gen. et sp. nov. A–C. Holotype, ♀ (TAMU-ENTO X1831147). D–F. Paratype, ♂ (TAMU-ENTO X1832015). A. Latero-posterior border of the first urosternite. B. Detail of lateral organ. C. Central organ of the first urosternite. D. Latero-posterior border of the first urosternite. E. Detail of lateral organ. F. Central organ of the first urosternite. Abbreviations: see Material and methods.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE5FFC8F561FC0AFAFA3218.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004195/files/figure.png	https://doi.org/10.5281/zenodo.19004195	Fig 7. Quapawjapyx osage Sendra gen. et sp. nov., paratype, ♂ (TAMU-ENTO X1832015). A. Genital papillae. B. Central organ of third urosternite.	Fig 7. Quapawjapyx osage Sendra gen. et sp. nov., paratype, ♂ (TAMU-ENTO X1832015). A. Genital papillae. B. Central organ of third urosternite.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE5FFC8F561FC0AFAFA3218.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004201/files/figure.png	https://doi.org/10.5281/zenodo.19004201	Fig. 8. Quapawjapyx osage Sendra gen. et sp. nov., ♀ (ASPC), cerci under electron scanning microscope. A. Right cercus latero-interior view. B. Left cercus, dorsal view. C. Detail distal portion of right cercus, view of postdental margin. D. Left cercus central portion of interior margin.	Fig. 8. Quapawjapyx osage Sendra gen. et sp. nov., ♀ (ASPC), cerci under electron scanning microscope. A. Right cercus latero-interior view. B. Left cercus, dorsal view. C. Detail distal portion of right cercus, view of postdental margin. D. Left cercus central portion of interior margin.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE5FFC8F561FC0AFAFA3218.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004203/files/figure.png	https://doi.org/10.5281/zenodo.19004203	Fig. 9. Quapawjapyx osage Sendra gen. et sp. nov. A–D. Paratype, ♀ (TAMU-ENTO X1832015). A. Left cercus, dorsal view. B. Right cercus, dorsal view. C. Left cercus, ventral view. D. Right cercus, ventral view. Abbreviations: see Material and methods.	Fig. 9. Quapawjapyx osage Sendra gen. et sp. nov. A–D. Paratype, ♀ (TAMU-ENTO X1832015). A. Left cercus, dorsal view. B. Right cercus, dorsal view. C. Left cercus, ventral view. D. Right cercus, ventral view. Abbreviations: see Material and methods.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE5FFC8F561FC0AFAFA3218.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004236/files/figure.png	https://doi.org/10.5281/zenodo.19004236	Fig. 19. Distribution of North American japygids. In yellow: karst areas (source: Chen et al. 2017); in orange: deserts (source: Olson & Dinerstein 2002); in blue: ice cover during the Last Glacial Maximum (source: Ehlers et al. 2011); green dots: known records of soil-adapted of the species of Japygidae Haliday, 1864; red colour symbols cave-adapted species of Japygidae: red triangles = Holjapyx nimiipuu Sendra sp. nov.; red diamonds = Quapawjapyx osage Sendra gen. et sp. nov.; red square = Occasjapyx kofoidi (Silvestri, 1928); red circles = Mixojapyx reddelli Muegge, 1992; red striped line: 64º N parallel, the known northern limit of Campodeidae.	Fig. 19. Distribution of North American japygids. In yellow: karst areas (source: Chen et al. 2017); in orange: deserts (source: Olson & Dinerstein 2002); in blue: ice cover during the Last Glacial Maximum (source: Ehlers et al. 2011); green dots: known records of soil-adapted of the species of Japygidae Haliday, 1864; red colour symbols cave-adapted species of Japygidae: red triangles = Holjapyx nimiipuu Sendra sp. nov.; red diamonds = Quapawjapyx osage Sendra gen. et sp. nov.; red square = Occasjapyx kofoidi (Silvestri, 1928); red circles = Mixojapyx reddelli Muegge, 1992; red striped line: 64º N parallel, the known northern limit of Campodeidae.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE9FFDCF532FD28FB4531A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004205/files/figure.png	https://doi.org/10.5281/zenodo.19004205	Fig. 10. Holjapyx nimiipuu Sendra sp. nov., holotype, ♀ (TAMU-ENTO X1831140). A. Habitus. B. Last antennomere. C. Distal portion of urite X including cerci. D. Left cercus. E. Right cercus; placoid sensilla remarked by dash line.	Fig. 10. Holjapyx nimiipuu Sendra sp. nov., holotype, ♀ (TAMU-ENTO X1831140). A. Habitus. B. Last antennomere. C. Distal portion of urite X including cerci. D. Left cercus. E. Right cercus; placoid sensilla remarked by dash line.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE9FFDCF532FD28FB4531A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004207/files/figure.png	https://doi.org/10.5281/zenodo.19004207	Fig. 11. Holjapyx nimiipuu Sendra sp. nov. A–E, G. Holotype, ♀ (TAMU-ENTO X1831140). F. Paratype, ♂ (TAMU-ENTO X185622). A. Habitus. B. Laminae pectinate. C. Spines (marked with arrows) besides insertion of third lamina pectinae. D. Detail surface urotergite VI. E. Claw of metathoracic leg. F. Genital papilla. G. Genital papilla.	Fig. 11. Holjapyx nimiipuu Sendra sp. nov. A–E, G. Holotype, ♀ (TAMU-ENTO X1831140). F. Paratype, ♂ (TAMU-ENTO X185622). A. Habitus. B. Laminae pectinate. C. Spines (marked with arrows) besides insertion of third lamina pectinae. D. Detail surface urotergite VI. E. Claw of metathoracic leg. F. Genital papilla. G. Genital papilla.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE9FFDCF532FD28FB4531A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004209/files/figure.png	https://doi.org/10.5281/zenodo.19004209	Fig. 12. Holjapyx nimiipuu Sendra sp. nov., holotype, ♀ (TAMU-ENTO X1831140). A. Pronotum. B. Prosternum. Abbreviations: see Material and methods.	Fig. 12. Holjapyx nimiipuu Sendra sp. nov., holotype, ♀ (TAMU-ENTO X1831140). A. Pronotum. B. Prosternum. Abbreviations: see Material and methods.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE9FFDCF532FD28FB4531A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004215/files/figure.png	https://doi.org/10.5281/zenodo.19004215	Fig. 13. Holjapyx nimiipuu Sendra sp. nov., holotype, ♀ (TAMU-ENTO X1831140). A. Mesonotum and metanotum. B. Mesosternum and metasternum. Abbreviations: see Material and methods.	Fig. 13. Holjapyx nimiipuu Sendra sp. nov., holotype, ♀ (TAMU-ENTO X1831140). A. Mesonotum and metanotum. B. Mesosternum and metasternum. Abbreviations: see Material and methods.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE9FFDCF532FD28FB4531A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004219/files/figure.png	https://doi.org/10.5281/zenodo.19004219	Fig. 14. Holjapyx nimiipuu Sendra sp. nov., holotype, ♀ (TAMU-ENTO X1831140). A. First urotergite. B. Second urotergite. C. Third urotergite. D. Sixth urotergite. Abbreviations: see Material and methods; M* = supernumerary macrosetae urotergal; * = setae instead of macrosetae.	Fig. 14. Holjapyx nimiipuu Sendra sp. nov., holotype, ♀ (TAMU-ENTO X1831140). A. First urotergite. B. Second urotergite. C. Third urotergite. D. Sixth urotergite. Abbreviations: see Material and methods; M* = supernumerary macrosetae urotergal; * = setae instead of macrosetae.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE9FFDCF532FD28FB4531A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004225/files/figure.png	https://doi.org/10.5281/zenodo.19004225	Fig. 15. Holjapyx nimiipuu Sendra sp. nov., holotype, ♀ (TAMU-ENTO X1831140). A. Seventh urotergite. B. Eighth urotergite. Abbreviations: see Material and methods; M*= supernumerary macrosetae urotergal..	Fig. 15. Holjapyx nimiipuu Sendra sp. nov., holotype, ♀ (TAMU-ENTO X1831140). A. Seventh urotergite. B. Eighth urotergite. Abbreviations: see Material and methods; M*= supernumerary macrosetae urotergal..	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE9FFDCF532FD28FB4531A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004229/files/figure.png	https://doi.org/10.5281/zenodo.19004229	Fig. 16. Holjapyx nimiipuu Sendra sp. nov., holotype, ♀ (TAMU-ENTO X1831140). A. First urosternite. B. Second urosternite. C. Eighth and ninth urosternites. Abbreviations: see Material and methods.	Fig. 16. Holjapyx nimiipuu Sendra sp. nov., holotype, ♀ (TAMU-ENTO X1831140). A. First urosternite. B. Second urosternite. C. Eighth and ninth urosternites. Abbreviations: see Material and methods.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE9FFDCF532FD28FB4531A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004232/files/figure.png	https://doi.org/10.5281/zenodo.19004232	Fig. 17. Holjapyx nimiipuu Sendra sp. nov. A, C. Paratype, ♀ (MUVHN-ZE8001). B, D. Paratype, ♂ (TAMU-ENTO X1856622). A–B. First urosternite. C. Central organ first urosternite. D. Sac organ of third urosternite. Abbreviations: see Material and methods.	Fig. 17. Holjapyx nimiipuu Sendra sp. nov. A, C. Paratype, ♀ (MUVHN-ZE8001). B, D. Paratype, ♂ (TAMU-ENTO X1856622). A–B. First urosternite. C. Central organ first urosternite. D. Sac organ of third urosternite. Abbreviations: see Material and methods.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE9FFDCF532FD28FB4531A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004234/files/figure.png	https://doi.org/10.5281/zenodo.19004234	Fig. 18. Holjapyx nimiipuu Sendra sp. nov., holotype, ♀ (TAMU-ENTO X1831140). A. Urite X, dorsal view. B. Urite X, ventral view. Abbreviations: see Material and methods.	Fig. 18. Holjapyx nimiipuu Sendra sp. nov., holotype, ♀ (TAMU-ENTO X1831140). A. Urite X, dorsal view. B. Urite X, ventral view. Abbreviations: see Material and methods.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
03B7D164FFE9FFDCF532FD28FB4531A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19004236/files/figure.png	https://doi.org/10.5281/zenodo.19004236	Fig. 19. Distribution of North American japygids. In yellow: karst areas (source: Chen et al. 2017); in orange: deserts (source: Olson & Dinerstein 2002); in blue: ice cover during the Last Glacial Maximum (source: Ehlers et al. 2011); green dots: known records of soil-adapted of the species of Japygidae Haliday, 1864; red colour symbols cave-adapted species of Japygidae: red triangles = Holjapyx nimiipuu Sendra sp. nov.; red diamonds = Quapawjapyx osage Sendra gen. et sp. nov.; red square = Occasjapyx kofoidi (Silvestri, 1928); red circles = Mixojapyx reddelli Muegge, 1992; red striped line: 64º N parallel, the known northern limit of Campodeidae.	Fig. 19. Distribution of North American japygids. In yellow: karst areas (source: Chen et al. 2017); in orange: deserts (source: Olson & Dinerstein 2002); in blue: ice cover during the Last Glacial Maximum (source: Ehlers et al. 2011); green dots: known records of soil-adapted of the species of Japygidae Haliday, 1864; red colour symbols cave-adapted species of Japygidae: red triangles = Holjapyx nimiipuu Sendra sp. nov.; red diamonds = Quapawjapyx osage Sendra gen. et sp. nov.; red square = Occasjapyx kofoidi (Silvestri, 1928); red circles = Mixojapyx reddelli Muegge, 1992; red striped line: 64º N parallel, the known northern limit of Campodeidae.	2026-03-11	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario		Zenodo	biologists	Sendra, Alberto;Jiménez-Valverde, Alberto;Selfa, Jesús;Cupello, Mario			
