taxonID	type	description	language	source
03B6594EC554FFEF7195586D01DB0082.taxon	description	(Figs. 1 – 3)	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC554FFEF7195586D01DB0082.taxon	diagnosis	Diagnosis. Himalusini is distinguished from other aleocharine tribes by the combination of the following character states: paramere distinctive, apical lobe partially or entirely covered with fine scale-like protuberances; lacinia with several large elongate or triangular teeth near apex; labium with extremely broad ligula; labial palpus with two palpomeres; prementum transverse with elongate, delicate and thin lateral lobes of apodeme (component “ r ” of premental sclerite in Weide et al. (2014 )) that converge at their apices; maxillary palpus with a pseudosegment; tarsal formula 4 - 4 - 4 or 4 - 4 - 5 (modified after Maruyama et al. 2014; most of these features shown in their figures 2 – 5).	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC554FFEF7195586D01DB0082.taxon	discussion	Remarks. Parabrachida decipiens Cameron, 1939 (= Himalusa thailandensis Pace, Klimaszewski & Center, 2010, syn. nov.) is known to feed on the leaves of sewer vine (Klimaszewski et al. 2010), and is the only known leaf-feeding species of Aleocharinae, but the feeding habit of other species of this tribe remains unknown.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC557FFE571955D2506190174.taxon	diagnosis	Diagnosis. The genus Protinodes is distinguished from Parabrachida by the transverse labrum, the ligula (Maruyama et al. 2014: fig. 3) not prominent medially, and the symmetrical median lobe of the aedeagus (Figs. 1 D – E; Pace, 2006: figs. 75 – 76). This genus differs from Sinanarchusa in the narrow and styliform apical lobe of the paramere of the aedeagus.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC557FFE571955D2506190174.taxon	discussion	Remarks. The habitus, mouthparts, and aedeagus of the type species of the genus Himalusa, Himalusa annapurnensis Pace, 2006, suggest a close relationship with Protinodes puncticollis Sharp, 1888. In particular, the morphology of the mouthparts — which are often considered critical fo the classification of Aleocharinae — is so similar between the two species that it does not justify their separation at the generic level. According to their original descriptions (Sharp 1888 and Pace 2006), these two species differ in the tarsal formula (Protinodes puncticollis: 4 - 4 - 4; Himalusa annapurnesis: 4 - 4 - 5), a character traditionally considered important for the generic classification within Aleocharinae. However, the hind tarsi of Protinodes puncticollis were previously miscounted and are in fact 5 - segmented. It is worth noting that Maruyama et al. (2014) also mistakenly recorded the hind tarsi as 4 - segmented, due to the difficulty of observing this character accurately under dry conditions. Therefore, we propose the following new synonymy and a new combination: Protinodes = Himalusa syn. nov., Protinodes annapurnensis (Pace, 2006), comb. nov.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC557FFE371955A11068107D4.taxon	description	(Fig. 1)	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC557FFE371955A11068107D4.taxon	materials_examined	Type material. Lectotype, here designated, male, “ Protinodes / puncticollis / Type D. S. / Tokio. Japan // Type [round label with red border] // Tokio. / 25. IX. - 27. IX. 81. // Japan. / G. Lewis. // Sharp Coll. / 1905 - 313. // Lectotype / Protinodes / puncticollis Sharp, 1888 / des. Hashizume et al. 2025 ” (BMNH) (maxilla, labium, abdominal segments VIII – X and aedeagus were dissected and mounted by MM in 2011). Additional material examined. Putative syntypes (see Remarks below): 2 females, “ Suô / IV // Protinodes / puncticollis Sharp / Det. T. Shiraki ” (TARI). Other specimens: 1 male, 1 female, JAPAN: Honshu: Shizuoka-ken, Susono, Suyama, Ônohara, 17 IX 2009, T. Kato leg. (SPMN).	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC557FFE371955A11068107D4.taxon	diagnosis	Diagnosis. Protinodes puncticollis is distinguished from Pr. annapurnensis by the reddish body color and the relatively stout median lobe of aedeagus.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC557FFE371955A11068107D4.taxon	description	Additional description. Measurements and ratios (n = 2). BL: 2.66 – 2.80; FBL: 1.28 – 1.30; HW: 0.44 – 0.48; PL: 0.50; PW: 0.69 – 0.71; EL: 0.64 – 0.65; EW: 0.88 – 0.89. PW / PL: 1.38 – 1.43; PW / HW: 1.43 – 1.63; EW / EL: 1.38; EL / PL: 1.28 – 1.30; EW / PW: 1.25 – 1.27. Tarsal formula 4 - 4 - 5. Male. Tergite VIII (Fig. 1 B) not modified. Female. Spermatheca (Fig. 1 F) simple; bulbus distalis subspherical.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC557FFE371955A11068107D4.taxon	distribution	Distribution. Japan (Honshu).	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC557FFE371955A11068107D4.taxon	discussion	Remarks. Maruyama et al. (2014) illustrated the “ holotype ” of this species; however, Sharp (1888) originally described it based on three specimens, indicating that this specimen is, in fact, one of the syntypes. Here, this specimen is designated as the lectotype. The other two presumed syntypes of this species are housed at TARI. A literal interpretation of the labels suggests that these specimens are not part of the type series. However, both presumed syntypes are mounted on the characteristically thick paper cards frequently used by Sharp, and the method of preparation and use of glue closely resembles those of other specimens prepared by Sharp. In these two specimens, however, this portion of the card appears to have been cut off. The so-called “ Shiraki specimens ” housed at TARI and other institutions are known to have significant unresolved issues (e. g., Suzuki & Minami 2008, 2015). One such issue — still under investigation — is the possibility that Dr. Tokuichi Shiraki removed a large number of the type specimens of beetles described from Asia (including some syntypes) from the British Museum (Natural History) prior to World War II, subsequently altering their labels, and encrypting them. Therefore, there remains a strong possibility that these specimens are indeed syntypes. For a detailed redescription, see Maruyama et al. (2014). The habitus is shown in Fig. 1 A. The paramere and median lobe of the aedeagus are as in Figs. 1 C – E.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC551FFE371955A53013E02F8.taxon	diagnosis	Diagnosis. The genus Parabrachida is distinguished from Protinodes by the anteriorly prominent labrum (Klimaszewski et al. 2010: fig. 4), the ligula prominent medially (Klimaszewski et al. 2010: fig. 8), and the asymmetrical median lobe of the aedeagus (Figs. 2 E – F, 3 E – F). This genus differs from Sinanarchusa in the ligula prominent medially, the asymmetrical median lobe of the aedeagus, and the narrow and styliform apical lobe of the paramere of the aedeagus.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC551FFE371955A53013E02F8.taxon	discussion	Remarks. Parabrachida has traditionally been placed in the tribe Homalotini to date (Newton 2025). Following the definition of Himalusini by Maruyama et al. (2014), we reassign this genus from Homalotini to Himalusini.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC551FFE371955CB105F4008B.taxon	diagnosis	Diagnosis. Protinodes annapurnensis is distinguished from Pr. puncticollis by the black body color and the more slender median lobe of aedeagus.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC551FFE371955CB105F4008B.taxon	distribution	Distribution. Nepal.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC551FFE17195599D057B0669.taxon	description	(Fig. 2)	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC551FFE17195599D057B0669.taxon	materials_examined	Type material examined. Lectotype, here designated, male, “ Type [round label with red border] // Samsingh 1800 / Kalimpong / Bengal. 2. X. 33 / C. F. C. Beeson. // 805 // Parabrachida / decipiens / TYPE Cam. // M. Cameron. / Bequest. / B. M. 1955 - 147. // NHMUK 015665385 ” (BMNH). Paralectotypes, 3 exs, Burma, Ruby Mines, Doherty leg. (BMNH).	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC551FFE17195599D057B0669.taxon	diagnosis	Diagnosis. Parabrachida decipiens is distinguished from Pa. persimilis by the elongate (distinctly longer than wide) male tergite VIII, and the moderately slender median lobe and long flagellum of the aedeagus, and the Yshaped sclerite located slightly right of the flagellum.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC551FFE17195599D057B0669.taxon	description	Measurements and ratios of lectotype. BL: 3.00; FBL: 1.40; HW: 0.64; PL: 0.57; PW: 0.98; EL: 0.70; EW: 1.10. PW / PL: 1.70; PW / HW: 1.53; EW / EL: 1.57; EL / PL: 1.22; EW / PW: 1.13.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC551FFE17195599D057B0669.taxon	distribution	Distribution. India, Myanmar, Thailand.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC551FFE17195599D057B0669.taxon	discussion	Remarks. For detailed descriptions, see Cameron (1939) and Klimaszewski et al. (2010) (as Himalusa thailandensis).	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC553FFE171955DED05720330.taxon	description	(Fig. 3)	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC553FFE171955DED05720330.taxon	materials_examined	Type material examined. Lectotype, here designated, male, “ Cochin / China // M. Cameron. / Bequest. / B. M. 1955 - 147. // NHMUK 015665389 ” (BMNH). Paralectotypes: 2 exs, Cochin China (BMNH).	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC553FFE171955DED05720330.taxon	diagnosis	Diagnosis. Parabrachida persimilis is distinguished from Pa. decipiens by the not elongate (only as long as wide) male tergite VIII, the more slender median lobe of aedeagus (especially in ventral view), the short flagellum of the median lobe of aedeagus, and the nearly triangular sclerite located slightly right of the flagellum.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC553FFE171955DED05720330.taxon	description	Redescription. Measurements of lectotype. BL: 2.02; FBL: 1.22; HW: 0.55; PL: 0.44; PW: 0.72; EL: 0.56; EW: 0.82. Body (Fig. 3 A) black, robust. Head sparsely punctured; punctures shallow with short setae; microsculpture indistinct, composed of slightly transverse meshes. Antennae short and stout. Thorax. Pronotum transverse (PW / PL: 1.64), wider than head (PW / HW: 1.31), sparsely punctured; punctures shallow with short setae; microsculpture indistinct, composed of slightly transverse meshes. Scutellum with short setae; microsculpture composed of transverse meshes. Elytra transverse (EW / EL: 1.47), longer than pronotum (EL / PL: 1.27), wider than pronotum (EW / PW: 1.14), with distinct large scale-shaped microsculpture; entire elytra also covered with fine, indistinct microsculpture composed of broken almost isodiametric to transverse meshes; setae longer than those of pronotum. Abdomen narrowed posteriad. Abdominal tergites III – VI densely punctured, posterior half of tergite VII without distinct puncture; III – V with distinct scale-shaped microsculpture; also covered with fine, indistinct microsculpture composed of broken almost isodiametric to transverse meshes; setae as long as or shorter than those of pronotum. Male. Tergite VIII (Fig. 3 C) truncate apically. Paramere of aedeagus with long styliform apical lobe with small protrusions; with short setae located near apical end. Median lobe of aedeagus slender, asymmetrical, and curved ventrally; with short flagellum and some indistinct sclerites; sclerite located right of flagellum nearly triangular in shape.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC553FFE171955DED05720330.taxon	distribution	Distribution. Vietnam.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC553FFE171955DED05720330.taxon	discussion	Remarks. In Parabrachida decipiens, body size varies considerably, with pronounced differences in the shape of male tergite VIII between large and small individuals (Klimaszewski et al. 2010). If similar body size variation occurs in Pa. persimilis, the shape of male tergite VIII may not be limited to what is shown here. According to Cameron (1944), this species was described based on four specimens, but we were able to examine three of them.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC55DFFEF71955E9A05F40750.taxon	diagnosis	Diagnosis. Sinanarchusa daxuensis is distinguished from other species of Himalusini by the thick and cylindrical apical lobe of the paramere of the aedeagus and the relatively stout median lobe of the aedeagus.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC55DFFEF71955E9A05F40750.taxon	distribution	Distribution. China.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
03B6594EC55DFFEF71955FF407DF05FD.taxon	diagnosis	Diagnosis. The genus Sinanarchusa is distinguished from Protinodes by the ligula not prominent medially, the thick and cylindrical apical lobe of paramere of aedeagus, and the almost symmetrical median lobe of aedeagus. This genus differs from Parabrachida in the thick and cylindrical apical lobe of paramere of aedeagus. This genus also differs from the other two genera in having a tarsal formula of 4 - 4 - 4.	en	Hashizume, Takuto, Maruyama, Munetoshi, Kishimoto, Toshio, Klimaszewski, Jan (2025): Taxonomic review of the tribe Himalusini (Coleoptera: Staphylinidae: Aleocharinae), with new synonymies. Zootaxa 5701 (1): 54-64, DOI: 10.11646/zootaxa.5701.1.2, URL: https://doi.org/10.11646/zootaxa.5701.1.2
