taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872545/files/figure.png	https://doi.org/10.5281/zenodo.21872545	Fig. 1. External morphological differences between the Phanaeus chalcomelas species group and the P. achilles species group. A–B. Females head in dorsal view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. achilles Boheman, 1858 (CEMT). C. P. arnaudorum sp. nov. (CEMT), elytra in dorsal view. D–E. Anteromedial portion of pronotum and transverse carina of the head of females. D. P. cambeforti (CEMT). E. P. achilles (CEMT). F–G. Pronotal disc of major males in dorsal view. F. P. chalcomelas (Perty, 1830) (CEMT). G. P. achilles (CEMT). H. P. lecourti Arnaud, 2000 (CEMT), elytra in dorsal view.	Fig. 1. External morphological differences between the Phanaeus chalcomelas species group and the P. achilles species group. A–B. Females head in dorsal view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. achilles Boheman, 1858 (CEMT). C. P. arnaudorum sp. nov. (CEMT), elytra in dorsal view. D–E. Anteromedial portion of pronotum and transverse carina of the head of females. D. P. cambeforti (CEMT). E. P. achilles (CEMT). F–G. Pronotal disc of major males in dorsal view. F. P. chalcomelas (Perty, 1830) (CEMT). G. P. achilles (CEMT). H. P. lecourti Arnaud, 2000 (CEMT), elytra in dorsal view.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872594/files/figure.png	https://doi.org/10.5281/zenodo.21872594	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872566/files/figure.png	https://doi.org/10.5281/zenodo.21872566	Fig. 8. Phanaeus divisor sp. nov. A, C–D. Holotype, ♂ (CEMT). A. Dorsal view. B. Paratype, ♀, dorsal view (CEMT). C. Ventral view. D. Head and anterior pronotal portion in lateral view. Arrows indicate the anteromedial pronotal tubercles; asterisk arrow indicates the carina adjacent to anteromedial pronotal margin.	Fig. 8. Phanaeus divisor sp. nov. A, C–D. Holotype, ♂ (CEMT). A. Dorsal view. B. Paratype, ♀, dorsal view (CEMT). C. Ventral view. D. Head and anterior pronotal portion in lateral view. Arrows indicate the anteromedial pronotal tubercles; asterisk arrow indicates the carina adjacent to anteromedial pronotal margin.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872588/files/figure.png	https://doi.org/10.5281/zenodo.21872588	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872590/files/figure.png	https://doi.org/10.5281/zenodo.21872590	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872597/files/figure.png	https://doi.org/10.5281/zenodo.21872597	Fig. 20. Geographic distribution of the Phanaeus cambeforti Arnaud, 1982; P. minos Erichson, 1847 stat. rev.; P. lecourti Arnaud, 2000; P. meleagris Blanchard, 1845; P. divisor sp. nov and P. mariocupelloi sp. nov.	Fig. 20. Geographic distribution of the Phanaeus cambeforti Arnaud, 1982; P. minos Erichson, 1847 stat. rev.; P. lecourti Arnaud, 2000; P. meleagris Blanchard, 1845; P. divisor sp. nov and P. mariocupelloi sp. nov.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872599/files/figure.png	https://doi.org/10.5281/zenodo.21872599	Fig. 21. Geographic distribution of Phanaeus chalcomelas (Perty, 1830). Black circles represent the coordinates from analyzed material. Gray circles sensu Chamorro et al. (2019).	Fig. 21. Geographic distribution of Phanaeus chalcomelas (Perty, 1830). Black circles represent the coordinates from analyzed material. Gray circles sensu Chamorro et al. (2019).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872551/files/figure.png	https://doi.org/10.5281/zenodo.21872551	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872576/files/figure.png	https://doi.org/10.5281/zenodo.21872576	Fig. 11. Phanaeus meleagris Blanchard, 1845 from Santa Cruz, Bolivia (CEMT). A. Major male, dorsal view. B. Female, dorsal view. C. Major male, ventral view. D. Major male, dorsal color.	Fig. 11. Phanaeus meleagris Blanchard, 1845 from Santa Cruz, Bolivia (CEMT). A. Major male, dorsal view. B. Female, dorsal view. C. Major male, ventral view. D. Major male, dorsal color.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872553/files/figure.png	https://doi.org/10.5281/zenodo.21872553	Fig. 3. Carina adjacent to anteromedial pronotal margin of females (frontal view). A. Phanaeus minos Erichson, 1847 stat. rev. (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). A–B. Arrows indicate middle of carina. C. P. cambeforti Arnaud, 1982 (CEMT). D. P. divisor sp. nov., paratype (CEMT).	Fig. 3. Carina adjacent to anteromedial pronotal margin of females (frontal view). A. Phanaeus minos Erichson, 1847 stat. rev. (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). A–B. Arrows indicate middle of carina. C. P. cambeforti Arnaud, 1982 (CEMT). D. P. divisor sp. nov., paratype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872578/files/figure.png	https://doi.org/10.5281/zenodo.21872578	Fig. 12. Phanaeus minos Erichson, 1847 stat. rev. A. Major male from Junín (Peru), dorsal view (CEMT). B. Female from Tachira (Venezuela), dorsal view (CEMT). C. Major male from Junín, ventral view (CEMT). D. Major male from Junín, dorsal color (CEMT).	Fig. 12. Phanaeus minos Erichson, 1847 stat. rev. A. Major male from Junín (Peru), dorsal view (CEMT). B. Female from Tachira (Venezuela), dorsal view (CEMT). C. Major male from Junín, ventral view (CEMT). D. Major male from Junín, dorsal color (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872574/files/figure.png	https://doi.org/10.5281/zenodo.21872574	Fig. 10. Phanaeus mariocupelloi sp. nov. A. Holotype, ♂, dorsal view (CEMT). B. Paratype, ♀, dorsal view (CEMT). C. Holotype, ♂, ventral view. D. Paratype, ♂, head and pronotum in dorsal view. Arrows indicate the anteromedial pronotal tubercles.	Fig. 10. Phanaeus mariocupelloi sp. nov. A. Holotype, ♂, dorsal view (CEMT). B. Paratype, ♀, dorsal view (CEMT). C. Holotype, ♂, ventral view. D. Paratype, ♂, head and pronotum in dorsal view. Arrows indicate the anteromedial pronotal tubercles.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872568/files/figure.png	https://doi.org/10.5281/zenodo.21872568	Fig. 9. Phanaeus lecourti Arnaud, 2000. A–C. Blue form. A. Major male from Junín (Peru), dorsal view (CEMT). B. Female from Cochabamba (Bolivia), dorsal view (CEMT). C. Major male (Junín), ventral view (CEMT). D. Green form ♀, from Leoncio Prado (Peru), dorsal view (CEMT).	Fig. 9. Phanaeus lecourti Arnaud, 2000. A–C. Blue form. A. Major male from Junín (Peru), dorsal view (CEMT). B. Female from Cochabamba (Bolivia), dorsal view (CEMT). C. Major male (Junín), ventral view (CEMT). D. Green form ♀, from Leoncio Prado (Peru), dorsal view (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872580/files/figure.png	https://doi.org/10.5281/zenodo.21872580	Fig. 13. Aedeagus of the Phanaeus chalcomelas species group. I = dorsal view; II = lateral view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). C. P. divisor sp. nov., paratype (CEMT). D. P. minos Erichson, 1847 stat. rev. (CEMT). E. P, lecourti Arnaud, 2000 (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT).	Fig. 13. Aedeagus of the Phanaeus chalcomelas species group. I = dorsal view; II = lateral view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). C. P. divisor sp. nov., paratype (CEMT). D. P. minos Erichson, 1847 stat. rev. (CEMT). E. P, lecourti Arnaud, 2000 (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872555/files/figure.png	https://doi.org/10.5281/zenodo.21872555	Fig. 4. Posteromedial pronotal fossae of males. A. Phanaeus cambeforti Arnaud, 1982 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT).	Fig. 4. Posteromedial pronotal fossae of males. A. Phanaeus cambeforti Arnaud, 1982 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059638FFADB93CF9C2191962FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872559/files/figure.png	https://doi.org/10.5281/zenodo.21872559	Fig. 5. Phanaeus cambeforti Arnaud, 1982. A. Male from Amapá, Brazil (CEMT), dorsal view. B. Female from Villavicencio, Colombia (CEMT), dorsal view. C. Male from Amapá, ventral view (CEMT). D. Head and pronotum of major male from Padmi, Ecuador (CEMT), dorsal view.	Fig. 5. Phanaeus cambeforti Arnaud, 1982. A. Male from Amapá, Brazil (CEMT), dorsal view. B. Female from Villavicencio, Colombia (CEMT), dorsal view. C. Male from Amapá, ventral view (CEMT). D. Head and pronotum of major male from Padmi, Ecuador (CEMT), dorsal view.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059634FFA9B943F88F1DB16397.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872545/files/figure.png	https://doi.org/10.5281/zenodo.21872545	Fig. 1. External morphological differences between the Phanaeus chalcomelas species group and the P. achilles species group. A–B. Females head in dorsal view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. achilles Boheman, 1858 (CEMT). C. P. arnaudorum sp. nov. (CEMT), elytra in dorsal view. D–E. Anteromedial portion of pronotum and transverse carina of the head of females. D. P. cambeforti (CEMT). E. P. achilles (CEMT). F–G. Pronotal disc of major males in dorsal view. F. P. chalcomelas (Perty, 1830) (CEMT). G. P. achilles (CEMT). H. P. lecourti Arnaud, 2000 (CEMT), elytra in dorsal view.	Fig. 1. External morphological differences between the Phanaeus chalcomelas species group and the P. achilles species group. A–B. Females head in dorsal view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. achilles Boheman, 1858 (CEMT). C. P. arnaudorum sp. nov. (CEMT), elytra in dorsal view. D–E. Anteromedial portion of pronotum and transverse carina of the head of females. D. P. cambeforti (CEMT). E. P. achilles (CEMT). F–G. Pronotal disc of major males in dorsal view. F. P. chalcomelas (Perty, 1830) (CEMT). G. P. achilles (CEMT). H. P. lecourti Arnaud, 2000 (CEMT), elytra in dorsal view.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059634FFA9B943F88F1DB16397.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872551/files/figure.png	https://doi.org/10.5281/zenodo.21872551	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059634FFA9B943F88F1DB16397.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872553/files/figure.png	https://doi.org/10.5281/zenodo.21872553	Fig. 3. Carina adjacent to anteromedial pronotal margin of females (frontal view). A. Phanaeus minos Erichson, 1847 stat. rev. (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). A–B. Arrows indicate middle of carina. C. P. cambeforti Arnaud, 1982 (CEMT). D. P. divisor sp. nov., paratype (CEMT).	Fig. 3. Carina adjacent to anteromedial pronotal margin of females (frontal view). A. Phanaeus minos Erichson, 1847 stat. rev. (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). A–B. Arrows indicate middle of carina. C. P. cambeforti Arnaud, 1982 (CEMT). D. P. divisor sp. nov., paratype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059634FFA9B943F88F1DB16397.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872555/files/figure.png	https://doi.org/10.5281/zenodo.21872555	Fig. 4. Posteromedial pronotal fossae of males. A. Phanaeus cambeforti Arnaud, 1982 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT).	Fig. 4. Posteromedial pronotal fossae of males. A. Phanaeus cambeforti Arnaud, 1982 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059634FFA9B943F88F1DB16397.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872559/files/figure.png	https://doi.org/10.5281/zenodo.21872559	Fig. 5. Phanaeus cambeforti Arnaud, 1982. A. Male from Amapá, Brazil (CEMT), dorsal view. B. Female from Villavicencio, Colombia (CEMT), dorsal view. C. Male from Amapá, ventral view (CEMT). D. Head and pronotum of major male from Padmi, Ecuador (CEMT), dorsal view.	Fig. 5. Phanaeus cambeforti Arnaud, 1982. A. Male from Amapá, Brazil (CEMT), dorsal view. B. Female from Villavicencio, Colombia (CEMT), dorsal view. C. Male from Amapá, ventral view (CEMT). D. Head and pronotum of major male from Padmi, Ecuador (CEMT), dorsal view.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059634FFA9B943F88F1DB16397.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872580/files/figure.png	https://doi.org/10.5281/zenodo.21872580	Fig. 13. Aedeagus of the Phanaeus chalcomelas species group. I = dorsal view; II = lateral view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). C. P. divisor sp. nov., paratype (CEMT). D. P. minos Erichson, 1847 stat. rev. (CEMT). E. P, lecourti Arnaud, 2000 (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT).	Fig. 13. Aedeagus of the Phanaeus chalcomelas species group. I = dorsal view; II = lateral view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). C. P. divisor sp. nov., paratype (CEMT). D. P. minos Erichson, 1847 stat. rev. (CEMT). E. P, lecourti Arnaud, 2000 (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059634FFA9B943F88F1DB16397.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872582/files/figure.png	https://doi.org/10.5281/zenodo.21872582	Fig. 14. Endophallus and endophallites of the Phanaeus chalcomelas species group. Respectively P. cambeforti Arnaud, 1982 (CEMT), P. divisor sp. nov., paratype (CEMT) and P. lecourti Arnaud, 2000 (CEMT). A–C. Endophallus in dorsal view. D–F. Medial endophallite (ME), dorsal (I) and lateral views (II). G–I. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. J–L. Superior right peripheral endophallite (SRP), dorsal (I) and ventral views (II).	Fig. 14. Endophallus and endophallites of the Phanaeus chalcomelas species group. Respectively P. cambeforti Arnaud, 1982 (CEMT), P. divisor sp. nov., paratype (CEMT) and P. lecourti Arnaud, 2000 (CEMT). A–C. Endophallus in dorsal view. D–F. Medial endophallite (ME), dorsal (I) and lateral views (II). G–I. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. J–L. Superior right peripheral endophallite (SRP), dorsal (I) and ventral views (II).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059634FFA9B943F88F1DB16397.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872588/files/figure.png	https://doi.org/10.5281/zenodo.21872588	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059634FFA9B943F88F1DB16397.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872590/files/figure.png	https://doi.org/10.5281/zenodo.21872590	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059634FFA9B943F88F1DB16397.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872592/files/figure.png	https://doi.org/10.5281/zenodo.21872592	Fig. 18. A–H. Posterior view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Peru (CEMT). F. P. mariocupelloi sp. nov., holotype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT). I. Phanaeus mariocupelloi sp. nov., paratype, ♀, posterior view (CEMT).	Fig. 18. A–H. Posterior view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Peru (CEMT). F. P. mariocupelloi sp. nov., holotype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT). I. Phanaeus mariocupelloi sp. nov., paratype, ♀, posterior view (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059634FFA9B943F88F1DB16397.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872594/files/figure.png	https://doi.org/10.5281/zenodo.21872594	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059634FFA9B943F88F1DB16397.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872597/files/figure.png	https://doi.org/10.5281/zenodo.21872597	Fig. 20. Geographic distribution of the Phanaeus cambeforti Arnaud, 1982; P. minos Erichson, 1847 stat. rev.; P. lecourti Arnaud, 2000; P. meleagris Blanchard, 1845; P. divisor sp. nov and P. mariocupelloi sp. nov.	Fig. 20. Geographic distribution of the Phanaeus cambeforti Arnaud, 1982; P. minos Erichson, 1847 stat. rev.; P. lecourti Arnaud, 2000; P. meleagris Blanchard, 1845; P. divisor sp. nov and P. mariocupelloi sp. nov.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059632FFB2B938FE861F616783.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872545/files/figure.png	https://doi.org/10.5281/zenodo.21872545	Fig. 1. External morphological differences between the Phanaeus chalcomelas species group and the P. achilles species group. A–B. Females head in dorsal view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. achilles Boheman, 1858 (CEMT). C. P. arnaudorum sp. nov. (CEMT), elytra in dorsal view. D–E. Anteromedial portion of pronotum and transverse carina of the head of females. D. P. cambeforti (CEMT). E. P. achilles (CEMT). F–G. Pronotal disc of major males in dorsal view. F. P. chalcomelas (Perty, 1830) (CEMT). G. P. achilles (CEMT). H. P. lecourti Arnaud, 2000 (CEMT), elytra in dorsal view.	Fig. 1. External morphological differences between the Phanaeus chalcomelas species group and the P. achilles species group. A–B. Females head in dorsal view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. achilles Boheman, 1858 (CEMT). C. P. arnaudorum sp. nov. (CEMT), elytra in dorsal view. D–E. Anteromedial portion of pronotum and transverse carina of the head of females. D. P. cambeforti (CEMT). E. P. achilles (CEMT). F–G. Pronotal disc of major males in dorsal view. F. P. chalcomelas (Perty, 1830) (CEMT). G. P. achilles (CEMT). H. P. lecourti Arnaud, 2000 (CEMT), elytra in dorsal view.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059632FFB2B938FE861F616783.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872551/files/figure.png	https://doi.org/10.5281/zenodo.21872551	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059632FFB2B938FE861F616783.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872562/files/figure.png	https://doi.org/10.5281/zenodo.21872562	Fig. 6. Phanaeus chalcomelas (Perty, 1830), dorsal view. A–B. From Amazonas State, Brazil (CEMT). C–D. From Santa Cruz de la Sierra, Bolivia (CEMT). A, C = males; B, D = females.	Fig. 6. Phanaeus chalcomelas (Perty, 1830), dorsal view. A–B. From Amazonas State, Brazil (CEMT). C–D. From Santa Cruz de la Sierra, Bolivia (CEMT). A, C = males; B, D = females.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059632FFB2B938FE861F616783.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872564/files/figure.png	https://doi.org/10.5281/zenodo.21872564	Fig. 7. Male genitalia of Phanaeus chalcomelas (Perty, 1830) (CEMT). A–B. Aedeagus. A. Dorsal view. B. Lateral view. C–E. Endophalli. C. Lateral view. D. Dorsal view. E. Ventral view. F–G. Medial endophallite (ME). F. Dorsal view. G. Lateral view. H. Additional endophallite (AE), ventral view. I. Superior right peripheral endophallite (SRP), ventral view. Abbreviations: A = axial endophallite; AE = additional endophallite; FLP = fronto-lateral peripheral endophallite; ME = medial endophallite; SA = subaxial endophallite; SRP = superior right peripheral endophallite.	Fig. 7. Male genitalia of Phanaeus chalcomelas (Perty, 1830) (CEMT). A–B. Aedeagus. A. Dorsal view. B. Lateral view. C–E. Endophalli. C. Lateral view. D. Dorsal view. E. Ventral view. F–G. Medial endophallite (ME). F. Dorsal view. G. Lateral view. H. Additional endophallite (AE), ventral view. I. Superior right peripheral endophallite (SRP), ventral view. Abbreviations: A = axial endophallite; AE = additional endophallite; FLP = fronto-lateral peripheral endophallite; ME = medial endophallite; SA = subaxial endophallite; SRP = superior right peripheral endophallite.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059632FFB2B938FE861F616783.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872588/files/figure.png	https://doi.org/10.5281/zenodo.21872588	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059632FFB2B938FE861F616783.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872590/files/figure.png	https://doi.org/10.5281/zenodo.21872590	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059632FFB2B938FE861F616783.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872592/files/figure.png	https://doi.org/10.5281/zenodo.21872592	Fig. 18. A–H. Posterior view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Peru (CEMT). F. P. mariocupelloi sp. nov., holotype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT). I. Phanaeus mariocupelloi sp. nov., paratype, ♀, posterior view (CEMT).	Fig. 18. A–H. Posterior view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Peru (CEMT). F. P. mariocupelloi sp. nov., holotype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT). I. Phanaeus mariocupelloi sp. nov., paratype, ♀, posterior view (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059632FFB2B938FE861F616783.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872594/files/figure.png	https://doi.org/10.5281/zenodo.21872594	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059632FFB2B938FE861F616783.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872627/files/figure.png	https://doi.org/10.5281/zenodo.21872627	Fig. 28. A. Phanaeus chalcomelas (Perty, 1830), holotype, ♀ (SZM). B. Labels. Photographs by Dr Michael Balke.	Fig. 28. A. Phanaeus chalcomelas (Perty, 1830), holotype, ♀ (SZM). B. Labels. Photographs by Dr Michael Balke.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059632FFB2B938FE861F616783.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872599/files/figure.png	https://doi.org/10.5281/zenodo.21872599	Fig. 21. Geographic distribution of Phanaeus chalcomelas (Perty, 1830). Black circles represent the coordinates from analyzed material. Gray circles sensu Chamorro et al. (2019).	Fig. 21. Geographic distribution of Phanaeus chalcomelas (Perty, 1830). Black circles represent the coordinates from analyzed material. Gray circles sensu Chamorro et al. (2019).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805962BFFBCB987FC441EAE6668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872551/files/figure.png	https://doi.org/10.5281/zenodo.21872551	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805962BFFBCB987FC441EAE6668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872553/files/figure.png	https://doi.org/10.5281/zenodo.21872553	Fig. 3. Carina adjacent to anteromedial pronotal margin of females (frontal view). A. Phanaeus minos Erichson, 1847 stat. rev. (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). A–B. Arrows indicate middle of carina. C. P. cambeforti Arnaud, 1982 (CEMT). D. P. divisor sp. nov., paratype (CEMT).	Fig. 3. Carina adjacent to anteromedial pronotal margin of females (frontal view). A. Phanaeus minos Erichson, 1847 stat. rev. (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). A–B. Arrows indicate middle of carina. C. P. cambeforti Arnaud, 1982 (CEMT). D. P. divisor sp. nov., paratype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805962BFFBCB987FC441EAE6668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872555/files/figure.png	https://doi.org/10.5281/zenodo.21872555	Fig. 4. Posteromedial pronotal fossae of males. A. Phanaeus cambeforti Arnaud, 1982 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT).	Fig. 4. Posteromedial pronotal fossae of males. A. Phanaeus cambeforti Arnaud, 1982 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805962BFFBCB987FC441EAE6668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872566/files/figure.png	https://doi.org/10.5281/zenodo.21872566	Fig. 8. Phanaeus divisor sp. nov. A, C–D. Holotype, ♂ (CEMT). A. Dorsal view. B. Paratype, ♀, dorsal view (CEMT). C. Ventral view. D. Head and anterior pronotal portion in lateral view. Arrows indicate the anteromedial pronotal tubercles; asterisk arrow indicates the carina adjacent to anteromedial pronotal margin.	Fig. 8. Phanaeus divisor sp. nov. A, C–D. Holotype, ♂ (CEMT). A. Dorsal view. B. Paratype, ♀, dorsal view (CEMT). C. Ventral view. D. Head and anterior pronotal portion in lateral view. Arrows indicate the anteromedial pronotal tubercles; asterisk arrow indicates the carina adjacent to anteromedial pronotal margin.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805962BFFBCB987FC441EAE6668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872580/files/figure.png	https://doi.org/10.5281/zenodo.21872580	Fig. 13. Aedeagus of the Phanaeus chalcomelas species group. I = dorsal view; II = lateral view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). C. P. divisor sp. nov., paratype (CEMT). D. P. minos Erichson, 1847 stat. rev. (CEMT). E. P, lecourti Arnaud, 2000 (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT).	Fig. 13. Aedeagus of the Phanaeus chalcomelas species group. I = dorsal view; II = lateral view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). C. P. divisor sp. nov., paratype (CEMT). D. P. minos Erichson, 1847 stat. rev. (CEMT). E. P, lecourti Arnaud, 2000 (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805962BFFBCB987FC441EAE6668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872582/files/figure.png	https://doi.org/10.5281/zenodo.21872582	Fig. 14. Endophallus and endophallites of the Phanaeus chalcomelas species group. Respectively P. cambeforti Arnaud, 1982 (CEMT), P. divisor sp. nov., paratype (CEMT) and P. lecourti Arnaud, 2000 (CEMT). A–C. Endophallus in dorsal view. D–F. Medial endophallite (ME), dorsal (I) and lateral views (II). G–I. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. J–L. Superior right peripheral endophallite (SRP), dorsal (I) and ventral views (II).	Fig. 14. Endophallus and endophallites of the Phanaeus chalcomelas species group. Respectively P. cambeforti Arnaud, 1982 (CEMT), P. divisor sp. nov., paratype (CEMT) and P. lecourti Arnaud, 2000 (CEMT). A–C. Endophallus in dorsal view. D–F. Medial endophallite (ME), dorsal (I) and lateral views (II). G–I. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. J–L. Superior right peripheral endophallite (SRP), dorsal (I) and ventral views (II).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805962BFFBCB987FC441EAE6668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872588/files/figure.png	https://doi.org/10.5281/zenodo.21872588	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805962BFFBCB987FC441EAE6668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872590/files/figure.png	https://doi.org/10.5281/zenodo.21872590	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805962BFFBCB987FC441EAE6668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872592/files/figure.png	https://doi.org/10.5281/zenodo.21872592	Fig. 18. A–H. Posterior view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Peru (CEMT). F. P. mariocupelloi sp. nov., holotype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT). I. Phanaeus mariocupelloi sp. nov., paratype, ♀, posterior view (CEMT).	Fig. 18. A–H. Posterior view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Peru (CEMT). F. P. mariocupelloi sp. nov., holotype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT). I. Phanaeus mariocupelloi sp. nov., paratype, ♀, posterior view (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805962BFFBCB987FC441EAE6668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872594/files/figure.png	https://doi.org/10.5281/zenodo.21872594	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805962BFFBCB987FC441EAE6668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872597/files/figure.png	https://doi.org/10.5281/zenodo.21872597	Fig. 20. Geographic distribution of the Phanaeus cambeforti Arnaud, 1982; P. minos Erichson, 1847 stat. rev.; P. lecourti Arnaud, 2000; P. meleagris Blanchard, 1845; P. divisor sp. nov and P. mariocupelloi sp. nov.	Fig. 20. Geographic distribution of the Phanaeus cambeforti Arnaud, 1982; P. minos Erichson, 1847 stat. rev.; P. lecourti Arnaud, 2000; P. meleagris Blanchard, 1845; P. divisor sp. nov and P. mariocupelloi sp. nov.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805962BFFBCB987FC441EAE6668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872559/files/figure.png	https://doi.org/10.5281/zenodo.21872559	Fig. 5. Phanaeus cambeforti Arnaud, 1982. A. Male from Amapá, Brazil (CEMT), dorsal view. B. Female from Villavicencio, Colombia (CEMT), dorsal view. C. Male from Amapá, ventral view (CEMT). D. Head and pronotum of major male from Padmi, Ecuador (CEMT), dorsal view.	Fig. 5. Phanaeus cambeforti Arnaud, 1982. A. Male from Amapá, Brazil (CEMT), dorsal view. B. Female from Villavicencio, Colombia (CEMT), dorsal view. C. Male from Amapá, ventral view (CEMT). D. Head and pronotum of major male from Padmi, Ecuador (CEMT), dorsal view.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059625FFBEB95FFDE0188F61BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872545/files/figure.png	https://doi.org/10.5281/zenodo.21872545	Fig. 1. External morphological differences between the Phanaeus chalcomelas species group and the P. achilles species group. A–B. Females head in dorsal view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. achilles Boheman, 1858 (CEMT). C. P. arnaudorum sp. nov. (CEMT), elytra in dorsal view. D–E. Anteromedial portion of pronotum and transverse carina of the head of females. D. P. cambeforti (CEMT). E. P. achilles (CEMT). F–G. Pronotal disc of major males in dorsal view. F. P. chalcomelas (Perty, 1830) (CEMT). G. P. achilles (CEMT). H. P. lecourti Arnaud, 2000 (CEMT), elytra in dorsal view.	Fig. 1. External morphological differences between the Phanaeus chalcomelas species group and the P. achilles species group. A–B. Females head in dorsal view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. achilles Boheman, 1858 (CEMT). C. P. arnaudorum sp. nov. (CEMT), elytra in dorsal view. D–E. Anteromedial portion of pronotum and transverse carina of the head of females. D. P. cambeforti (CEMT). E. P. achilles (CEMT). F–G. Pronotal disc of major males in dorsal view. F. P. chalcomelas (Perty, 1830) (CEMT). G. P. achilles (CEMT). H. P. lecourti Arnaud, 2000 (CEMT), elytra in dorsal view.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059625FFBEB95FFDE0188F61BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872551/files/figure.png	https://doi.org/10.5281/zenodo.21872551	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059625FFBEB95FFDE0188F61BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872568/files/figure.png	https://doi.org/10.5281/zenodo.21872568	Fig. 9. Phanaeus lecourti Arnaud, 2000. A–C. Blue form. A. Major male from Junín (Peru), dorsal view (CEMT). B. Female from Cochabamba (Bolivia), dorsal view (CEMT). C. Major male (Junín), ventral view (CEMT). D. Green form ♀, from Leoncio Prado (Peru), dorsal view (CEMT).	Fig. 9. Phanaeus lecourti Arnaud, 2000. A–C. Blue form. A. Major male from Junín (Peru), dorsal view (CEMT). B. Female from Cochabamba (Bolivia), dorsal view (CEMT). C. Major male (Junín), ventral view (CEMT). D. Green form ♀, from Leoncio Prado (Peru), dorsal view (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059625FFBEB95FFDE0188F61BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872580/files/figure.png	https://doi.org/10.5281/zenodo.21872580	Fig. 13. Aedeagus of the Phanaeus chalcomelas species group. I = dorsal view; II = lateral view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). C. P. divisor sp. nov., paratype (CEMT). D. P. minos Erichson, 1847 stat. rev. (CEMT). E. P, lecourti Arnaud, 2000 (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT).	Fig. 13. Aedeagus of the Phanaeus chalcomelas species group. I = dorsal view; II = lateral view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). C. P. divisor sp. nov., paratype (CEMT). D. P. minos Erichson, 1847 stat. rev. (CEMT). E. P, lecourti Arnaud, 2000 (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059625FFBEB95FFDE0188F61BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872582/files/figure.png	https://doi.org/10.5281/zenodo.21872582	Fig. 14. Endophallus and endophallites of the Phanaeus chalcomelas species group. Respectively P. cambeforti Arnaud, 1982 (CEMT), P. divisor sp. nov., paratype (CEMT) and P. lecourti Arnaud, 2000 (CEMT). A–C. Endophallus in dorsal view. D–F. Medial endophallite (ME), dorsal (I) and lateral views (II). G–I. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. J–L. Superior right peripheral endophallite (SRP), dorsal (I) and ventral views (II).	Fig. 14. Endophallus and endophallites of the Phanaeus chalcomelas species group. Respectively P. cambeforti Arnaud, 1982 (CEMT), P. divisor sp. nov., paratype (CEMT) and P. lecourti Arnaud, 2000 (CEMT). A–C. Endophallus in dorsal view. D–F. Medial endophallite (ME), dorsal (I) and lateral views (II). G–I. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. J–L. Superior right peripheral endophallite (SRP), dorsal (I) and ventral views (II).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059625FFBEB95FFDE0188F61BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872588/files/figure.png	https://doi.org/10.5281/zenodo.21872588	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059625FFBEB95FFDE0188F61BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872590/files/figure.png	https://doi.org/10.5281/zenodo.21872590	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059625FFBEB95FFDE0188F61BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872592/files/figure.png	https://doi.org/10.5281/zenodo.21872592	Fig. 18. A–H. Posterior view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Peru (CEMT). F. P. mariocupelloi sp. nov., holotype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT). I. Phanaeus mariocupelloi sp. nov., paratype, ♀, posterior view (CEMT).	Fig. 18. A–H. Posterior view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Peru (CEMT). F. P. mariocupelloi sp. nov., holotype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT). I. Phanaeus mariocupelloi sp. nov., paratype, ♀, posterior view (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059625FFBEB95FFDE0188F61BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872594/files/figure.png	https://doi.org/10.5281/zenodo.21872594	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059625FFBEB95FFDE0188F61BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872597/files/figure.png	https://doi.org/10.5281/zenodo.21872597	Fig. 20. Geographic distribution of the Phanaeus cambeforti Arnaud, 1982; P. minos Erichson, 1847 stat. rev.; P. lecourti Arnaud, 2000; P. meleagris Blanchard, 1845; P. divisor sp. nov and P. mariocupelloi sp. nov.	Fig. 20. Geographic distribution of the Phanaeus cambeforti Arnaud, 1982; P. minos Erichson, 1847 stat. rev.; P. lecourti Arnaud, 2000; P. meleagris Blanchard, 1845; P. divisor sp. nov and P. mariocupelloi sp. nov.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059627FFBBB952FA481F4C6061.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872551/files/figure.png	https://doi.org/10.5281/zenodo.21872551	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059627FFBBB952FA481F4C6061.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872555/files/figure.png	https://doi.org/10.5281/zenodo.21872555	Fig. 4. Posteromedial pronotal fossae of males. A. Phanaeus cambeforti Arnaud, 1982 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT).	Fig. 4. Posteromedial pronotal fossae of males. A. Phanaeus cambeforti Arnaud, 1982 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059627FFBBB952FA481F4C6061.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872574/files/figure.png	https://doi.org/10.5281/zenodo.21872574	Fig. 10. Phanaeus mariocupelloi sp. nov. A. Holotype, ♂, dorsal view (CEMT). B. Paratype, ♀, dorsal view (CEMT). C. Holotype, ♂, ventral view. D. Paratype, ♂, head and pronotum in dorsal view. Arrows indicate the anteromedial pronotal tubercles.	Fig. 10. Phanaeus mariocupelloi sp. nov. A. Holotype, ♂, dorsal view (CEMT). B. Paratype, ♀, dorsal view (CEMT). C. Holotype, ♂, ventral view. D. Paratype, ♂, head and pronotum in dorsal view. Arrows indicate the anteromedial pronotal tubercles.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059627FFBBB952FA481F4C6061.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872580/files/figure.png	https://doi.org/10.5281/zenodo.21872580	Fig. 13. Aedeagus of the Phanaeus chalcomelas species group. I = dorsal view; II = lateral view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). C. P. divisor sp. nov., paratype (CEMT). D. P. minos Erichson, 1847 stat. rev. (CEMT). E. P, lecourti Arnaud, 2000 (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT).	Fig. 13. Aedeagus of the Phanaeus chalcomelas species group. I = dorsal view; II = lateral view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). C. P. divisor sp. nov., paratype (CEMT). D. P. minos Erichson, 1847 stat. rev. (CEMT). E. P, lecourti Arnaud, 2000 (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059627FFBBB952FA481F4C6061.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872584/files/figure.png	https://doi.org/10.5281/zenodo.21872584	Fig. 15. Endophallus and endophallites of the Phanaeus chalcomelas species group. Respectively P. meleagris Blanchard, 1845 (CEMT), P. minos Erichson, 1847 stat. rev. (CEMT) and P. mariocupelloi sp. nov., paratype (CEMT). A–C. Endophallus, dorsal view. D–F. Medial endophallite (ME), dorsal (I) and lateral views (II). G–I. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. J–L. Superior right peripheral endophallite (SRP), dorsal (I) and ventral views (II).	Fig. 15. Endophallus and endophallites of the Phanaeus chalcomelas species group. Respectively P. meleagris Blanchard, 1845 (CEMT), P. minos Erichson, 1847 stat. rev. (CEMT) and P. mariocupelloi sp. nov., paratype (CEMT). A–C. Endophallus, dorsal view. D–F. Medial endophallite (ME), dorsal (I) and lateral views (II). G–I. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. J–L. Superior right peripheral endophallite (SRP), dorsal (I) and ventral views (II).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059627FFBBB952FA481F4C6061.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872588/files/figure.png	https://doi.org/10.5281/zenodo.21872588	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059627FFBBB952FA481F4C6061.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872590/files/figure.png	https://doi.org/10.5281/zenodo.21872590	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059627FFBBB952FA481F4C6061.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872592/files/figure.png	https://doi.org/10.5281/zenodo.21872592	Fig. 18. A–H. Posterior view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Peru (CEMT). F. P. mariocupelloi sp. nov., holotype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT). I. Phanaeus mariocupelloi sp. nov., paratype, ♀, posterior view (CEMT).	Fig. 18. A–H. Posterior view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Peru (CEMT). F. P. mariocupelloi sp. nov., holotype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT). I. Phanaeus mariocupelloi sp. nov., paratype, ♀, posterior view (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059627FFBBB952FA481F4C6061.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872594/files/figure.png	https://doi.org/10.5281/zenodo.21872594	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059627FFBBB952FA481F4C6061.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872597/files/figure.png	https://doi.org/10.5281/zenodo.21872597	Fig. 20. Geographic distribution of the Phanaeus cambeforti Arnaud, 1982; P. minos Erichson, 1847 stat. rev.; P. lecourti Arnaud, 2000; P. meleagris Blanchard, 1845; P. divisor sp. nov and P. mariocupelloi sp. nov.	Fig. 20. Geographic distribution of the Phanaeus cambeforti Arnaud, 1982; P. minos Erichson, 1847 stat. rev.; P. lecourti Arnaud, 2000; P. meleagris Blanchard, 1845; P. divisor sp. nov and P. mariocupelloi sp. nov.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059622FF84B93DFBE61DC563A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872551/files/figure.png	https://doi.org/10.5281/zenodo.21872551	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059622FF84B93DFBE61DC563A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872553/files/figure.png	https://doi.org/10.5281/zenodo.21872553	Fig. 3. Carina adjacent to anteromedial pronotal margin of females (frontal view). A. Phanaeus minos Erichson, 1847 stat. rev. (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). A–B. Arrows indicate middle of carina. C. P. cambeforti Arnaud, 1982 (CEMT). D. P. divisor sp. nov., paratype (CEMT).	Fig. 3. Carina adjacent to anteromedial pronotal margin of females (frontal view). A. Phanaeus minos Erichson, 1847 stat. rev. (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). A–B. Arrows indicate middle of carina. C. P. cambeforti Arnaud, 1982 (CEMT). D. P. divisor sp. nov., paratype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059622FF84B93DFBE61DC563A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872576/files/figure.png	https://doi.org/10.5281/zenodo.21872576	Fig. 11. Phanaeus meleagris Blanchard, 1845 from Santa Cruz, Bolivia (CEMT). A. Major male, dorsal view. B. Female, dorsal view. C. Major male, ventral view. D. Major male, dorsal color.	Fig. 11. Phanaeus meleagris Blanchard, 1845 from Santa Cruz, Bolivia (CEMT). A. Major male, dorsal view. B. Female, dorsal view. C. Major male, ventral view. D. Major male, dorsal color.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059622FF84B93DFBE61DC563A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872580/files/figure.png	https://doi.org/10.5281/zenodo.21872580	Fig. 13. Aedeagus of the Phanaeus chalcomelas species group. I = dorsal view; II = lateral view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). C. P. divisor sp. nov., paratype (CEMT). D. P. minos Erichson, 1847 stat. rev. (CEMT). E. P, lecourti Arnaud, 2000 (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT).	Fig. 13. Aedeagus of the Phanaeus chalcomelas species group. I = dorsal view; II = lateral view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). C. P. divisor sp. nov., paratype (CEMT). D. P. minos Erichson, 1847 stat. rev. (CEMT). E. P, lecourti Arnaud, 2000 (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059622FF84B93DFBE61DC563A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872584/files/figure.png	https://doi.org/10.5281/zenodo.21872584	Fig. 15. Endophallus and endophallites of the Phanaeus chalcomelas species group. Respectively P. meleagris Blanchard, 1845 (CEMT), P. minos Erichson, 1847 stat. rev. (CEMT) and P. mariocupelloi sp. nov., paratype (CEMT). A–C. Endophallus, dorsal view. D–F. Medial endophallite (ME), dorsal (I) and lateral views (II). G–I. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. J–L. Superior right peripheral endophallite (SRP), dorsal (I) and ventral views (II).	Fig. 15. Endophallus and endophallites of the Phanaeus chalcomelas species group. Respectively P. meleagris Blanchard, 1845 (CEMT), P. minos Erichson, 1847 stat. rev. (CEMT) and P. mariocupelloi sp. nov., paratype (CEMT). A–C. Endophallus, dorsal view. D–F. Medial endophallite (ME), dorsal (I) and lateral views (II). G–I. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. J–L. Superior right peripheral endophallite (SRP), dorsal (I) and ventral views (II).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059622FF84B93DFBE61DC563A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872588/files/figure.png	https://doi.org/10.5281/zenodo.21872588	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059622FF84B93DFBE61DC563A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872590/files/figure.png	https://doi.org/10.5281/zenodo.21872590	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059622FF84B93DFBE61DC563A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872592/files/figure.png	https://doi.org/10.5281/zenodo.21872592	Fig. 18. A–H. Posterior view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Peru (CEMT). F. P. mariocupelloi sp. nov., holotype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT). I. Phanaeus mariocupelloi sp. nov., paratype, ♀, posterior view (CEMT).	Fig. 18. A–H. Posterior view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Peru (CEMT). F. P. mariocupelloi sp. nov., holotype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT). I. Phanaeus mariocupelloi sp. nov., paratype, ♀, posterior view (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059622FF84B93DFBE61DC563A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872594/files/figure.png	https://doi.org/10.5281/zenodo.21872594	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059622FF84B93DFBE61DC563A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872597/files/figure.png	https://doi.org/10.5281/zenodo.21872597	Fig. 20. Geographic distribution of the Phanaeus cambeforti Arnaud, 1982; P. minos Erichson, 1847 stat. rev.; P. lecourti Arnaud, 2000; P. meleagris Blanchard, 1845; P. divisor sp. nov and P. mariocupelloi sp. nov.	Fig. 20. Geographic distribution of the Phanaeus cambeforti Arnaud, 1982; P. minos Erichson, 1847 stat. rev.; P. lecourti Arnaud, 2000; P. meleagris Blanchard, 1845; P. divisor sp. nov and P. mariocupelloi sp. nov.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059622FF84B93DFBE61DC563A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872578/files/figure.png	https://doi.org/10.5281/zenodo.21872578	Fig. 12. Phanaeus minos Erichson, 1847 stat. rev. A. Major male from Junín (Peru), dorsal view (CEMT). B. Female from Tachira (Venezuela), dorsal view (CEMT). C. Major male from Junín, ventral view (CEMT). D. Major male from Junín, dorsal color (CEMT).	Fig. 12. Phanaeus minos Erichson, 1847 stat. rev. A. Major male from Junín (Peru), dorsal view (CEMT). B. Female from Tachira (Venezuela), dorsal view (CEMT). C. Major male from Junín, ventral view (CEMT). D. Major male from Junín, dorsal color (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805961EFF8FB923FE86181865F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872551/files/figure.png	https://doi.org/10.5281/zenodo.21872551	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	Fig. 2. Anteromedial prominence of metaventrite (Phanaeus chalcomelas species group). A. P. chalcomelas (Perty, 1830), from Santa Cruz de la Sierra, Bolivia (CEMT). B. P. lecourti Arnaud, 2000 (CEMT). C. P. mariocupelloi sp. nov., holotype (CEMT). D. P. divisor sp. nov., holotype (CEMT). E. P. divisor sp. nov., holotype, lateral view (CEMT). F. P. meleagris Blanchard, 1845 (CEMT). G. P. cambeforti Arnaud, 1982 (CEMT). H. P. cambeforti, lateral view (CEMT). I. P. minos Erichson, 1847 stat. rev., from Peru (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805961EFF8FB923FE86181865F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872553/files/figure.png	https://doi.org/10.5281/zenodo.21872553	Fig. 3. Carina adjacent to anteromedial pronotal margin of females (frontal view). A. Phanaeus minos Erichson, 1847 stat. rev. (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). A–B. Arrows indicate middle of carina. C. P. cambeforti Arnaud, 1982 (CEMT). D. P. divisor sp. nov., paratype (CEMT).	Fig. 3. Carina adjacent to anteromedial pronotal margin of females (frontal view). A. Phanaeus minos Erichson, 1847 stat. rev. (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). A–B. Arrows indicate middle of carina. C. P. cambeforti Arnaud, 1982 (CEMT). D. P. divisor sp. nov., paratype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805961EFF8FB923FE86181865F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872578/files/figure.png	https://doi.org/10.5281/zenodo.21872578	Fig. 12. Phanaeus minos Erichson, 1847 stat. rev. A. Major male from Junín (Peru), dorsal view (CEMT). B. Female from Tachira (Venezuela), dorsal view (CEMT). C. Major male from Junín, ventral view (CEMT). D. Major male from Junín, dorsal color (CEMT).	Fig. 12. Phanaeus minos Erichson, 1847 stat. rev. A. Major male from Junín (Peru), dorsal view (CEMT). B. Female from Tachira (Venezuela), dorsal view (CEMT). C. Major male from Junín, ventral view (CEMT). D. Major male from Junín, dorsal color (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805961EFF8FB923FE86181865F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872580/files/figure.png	https://doi.org/10.5281/zenodo.21872580	Fig. 13. Aedeagus of the Phanaeus chalcomelas species group. I = dorsal view; II = lateral view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). C. P. divisor sp. nov., paratype (CEMT). D. P. minos Erichson, 1847 stat. rev. (CEMT). E. P, lecourti Arnaud, 2000 (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT).	Fig. 13. Aedeagus of the Phanaeus chalcomelas species group. I = dorsal view; II = lateral view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. meleagris Blanchard, 1845 (CEMT). C. P. divisor sp. nov., paratype (CEMT). D. P. minos Erichson, 1847 stat. rev. (CEMT). E. P, lecourti Arnaud, 2000 (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805961EFF8FB923FE86181865F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872584/files/figure.png	https://doi.org/10.5281/zenodo.21872584	Fig. 15. Endophallus and endophallites of the Phanaeus chalcomelas species group. Respectively P. meleagris Blanchard, 1845 (CEMT), P. minos Erichson, 1847 stat. rev. (CEMT) and P. mariocupelloi sp. nov., paratype (CEMT). A–C. Endophallus, dorsal view. D–F. Medial endophallite (ME), dorsal (I) and lateral views (II). G–I. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. J–L. Superior right peripheral endophallite (SRP), dorsal (I) and ventral views (II).	Fig. 15. Endophallus and endophallites of the Phanaeus chalcomelas species group. Respectively P. meleagris Blanchard, 1845 (CEMT), P. minos Erichson, 1847 stat. rev. (CEMT) and P. mariocupelloi sp. nov., paratype (CEMT). A–C. Endophallus, dorsal view. D–F. Medial endophallite (ME), dorsal (I) and lateral views (II). G–I. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. J–L. Superior right peripheral endophallite (SRP), dorsal (I) and ventral views (II).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805961EFF8FB923FE86181865F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872588/files/figure.png	https://doi.org/10.5281/zenodo.21872588	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 16. Frontal view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. Phanaeus mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805961EFF8FB923FE86181865F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872590/files/figure.png	https://doi.org/10.5281/zenodo.21872590	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 17. Lateral view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Ecuador (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805961EFF8FB923FE86181865F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872592/files/figure.png	https://doi.org/10.5281/zenodo.21872592	Fig. 18. A–H. Posterior view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Peru (CEMT). F. P. mariocupelloi sp. nov., holotype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT). I. Phanaeus mariocupelloi sp. nov., paratype, ♀, posterior view (CEMT).	Fig. 18. A–H. Posterior view of males of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., holotype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Peru (CEMT). F. P. mariocupelloi sp. nov., holotype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT). I. Phanaeus mariocupelloi sp. nov., paratype, ♀, posterior view (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805961EFF8FB923FE86181865F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872594/files/figure.png	https://doi.org/10.5281/zenodo.21872594	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	Fig. 19. Frontal view of females of the Phanaeus chalcomelas species group. A. P. lecourti Arnaud, 2000 (CEMT). B. P. divisor sp. nov., paratype (CEMT). C. P. cambeforti Arnaud, 1982 (CEMT). D. P. meleagris Blanchard, 1845 (CEMT). E. P. minos Erichson, 1847 stat. rev., from Venezuela (CEMT). F. P. mariocupelloi sp. nov., paratype (CEMT). G. P. chalcomelas (Perty, 1830), from Amazonas, Brazil (CEMT). H. P. chalcomelas, from Santa Cruz de la Sierra, Bolivia (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805961EFF8FB923FE86181865F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872597/files/figure.png	https://doi.org/10.5281/zenodo.21872597	Fig. 20. Geographic distribution of the Phanaeus cambeforti Arnaud, 1982; P. minos Erichson, 1847 stat. rev.; P. lecourti Arnaud, 2000; P. meleagris Blanchard, 1845; P. divisor sp. nov and P. mariocupelloi sp. nov.	Fig. 20. Geographic distribution of the Phanaeus cambeforti Arnaud, 1982; P. minos Erichson, 1847 stat. rev.; P. lecourti Arnaud, 2000; P. meleagris Blanchard, 1845; P. divisor sp. nov and P. mariocupelloi sp. nov.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805961EFF8FB923FE86181865F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872576/files/figure.png	https://doi.org/10.5281/zenodo.21872576	Fig. 11. Phanaeus meleagris Blanchard, 1845 from Santa Cruz, Bolivia (CEMT). A. Major male, dorsal view. B. Female, dorsal view. C. Major male, ventral view. D. Major male, dorsal color.	Fig. 11. Phanaeus meleagris Blanchard, 1845 from Santa Cruz, Bolivia (CEMT). A. Major male, dorsal view. B. Female, dorsal view. C. Major male, ventral view. D. Major male, dorsal color.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059617FF88B950FE8619196668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872545/files/figure.png	https://doi.org/10.5281/zenodo.21872545	Fig. 1. External morphological differences between the Phanaeus chalcomelas species group and the P. achilles species group. A–B. Females head in dorsal view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. achilles Boheman, 1858 (CEMT). C. P. arnaudorum sp. nov. (CEMT), elytra in dorsal view. D–E. Anteromedial portion of pronotum and transverse carina of the head of females. D. P. cambeforti (CEMT). E. P. achilles (CEMT). F–G. Pronotal disc of major males in dorsal view. F. P. chalcomelas (Perty, 1830) (CEMT). G. P. achilles (CEMT). H. P. lecourti Arnaud, 2000 (CEMT), elytra in dorsal view.	Fig. 1. External morphological differences between the Phanaeus chalcomelas species group and the P. achilles species group. A–B. Females head in dorsal view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. achilles Boheman, 1858 (CEMT). C. P. arnaudorum sp. nov. (CEMT), elytra in dorsal view. D–E. Anteromedial portion of pronotum and transverse carina of the head of females. D. P. cambeforti (CEMT). E. P. achilles (CEMT). F–G. Pronotal disc of major males in dorsal view. F. P. chalcomelas (Perty, 1830) (CEMT). G. P. achilles (CEMT). H. P. lecourti Arnaud, 2000 (CEMT), elytra in dorsal view.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059617FF88B950FE8619196668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872607/files/figure.png	https://doi.org/10.5281/zenodo.21872607	Fig. 22. Phanaeus achilles species group in dorsal view. A–B. P. achilles Boheman, 1858 (CEMT). A. Major male. B. Female. C–D. P. arnaudorum sp. nov. C. Holotype, major male (CEMT). D. Paratype, female (CEMT).	Fig. 22. Phanaeus achilles species group in dorsal view. A–B. P. achilles Boheman, 1858 (CEMT). A. Major male. B. Female. C–D. P. arnaudorum sp. nov. C. Holotype, major male (CEMT). D. Paratype, female (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059617FF88B950FE8619196668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872611/files/figure.png	https://doi.org/10.5281/zenodo.21872611	Fig. 23. Phanaeus achilles species group. A. P. achilles Boheman, 1858 ♂ (CEMT), frontal view. B. P. arnaudorum sp. nov., holotype, ♂ (CEMT), frontal view. C. P. achilles Boheman, 1858 ♂ (CEMT), posterior view. D. P. arnaudorum sp. nov., holotype, ♂ (CEMT), posterior view. E. P. achilles Boheman, 1858 ♀ (CEMT), frontal view. F. P. arnaudorum sp. nov., paratype, ♀ (CEMT), frontal view.	Fig. 23. Phanaeus achilles species group. A. P. achilles Boheman, 1858 ♂ (CEMT), frontal view. B. P. arnaudorum sp. nov., holotype, ♂ (CEMT), frontal view. C. P. achilles Boheman, 1858 ♂ (CEMT), posterior view. D. P. arnaudorum sp. nov., holotype, ♂ (CEMT), posterior view. E. P. achilles Boheman, 1858 ♀ (CEMT), frontal view. F. P. arnaudorum sp. nov., paratype, ♀ (CEMT), frontal view.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059617FF88B950FE8619196668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872617/files/figure.png	https://doi.org/10.5281/zenodo.21872617	Fig. 25. Aedeagus of the Phanaeus achilles species group. A. P. achilles Boheman, 1858 (CEMT). B. P. arnaudorum sp. nov., paratype (CEMT), dorsal (I) and lateral (II) views.	Fig. 25. Aedeagus of the Phanaeus achilles species group. A. P. achilles Boheman, 1858 (CEMT). B. P. arnaudorum sp. nov., paratype (CEMT), dorsal (I) and lateral (II) views.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059617FF88B950FE8619196668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872623/files/figure.png	https://doi.org/10.5281/zenodo.21872623	Fig. 27. Geographic distribution of Phanaeus achilles Boheman, 1858 and P. arnaudorum sp. nov. The star is sensu Figueroa et al. (2014).	Fig. 27. Geographic distribution of Phanaeus achilles Boheman, 1858 and P. arnaudorum sp. nov. The star is sensu Figueroa et al. (2014).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059617FF88B950FE8619196668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872615/files/figure.png	https://doi.org/10.5281/zenodo.21872615	Fig. 24. Major males of Phanaeus achilles species group in fronto-lateral view. A, C. P. achilles Boheman, 1858 (CEMT). B, D. P. arnaudorum sp. nov., holotype (CEMT). Arrows indicate the anteromedial pronotal tubercles, larger in P. achilles; asterisk arrows indicate the anterolateral pronotal fossae, larger in P. arnaudorum.	Fig. 24. Major males of Phanaeus achilles species group in fronto-lateral view. A, C. P. achilles Boheman, 1858 (CEMT). B, D. P. arnaudorum sp. nov., holotype (CEMT). Arrows indicate the anteromedial pronotal tubercles, larger in P. achilles; asterisk arrows indicate the anterolateral pronotal fossae, larger in P. arnaudorum.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059617FF88B950FE8619196668.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872619/files/figure.png	https://doi.org/10.5281/zenodo.21872619	Fig. 26. Endophallus and endophallites of the Phanaeus achilles species group. Respectively P. achilles Boheman, 1858 (CEMT) and P. arnaudorum sp. nov., paratype (CEMT). A–B. Endophallus in dorsal view. C–D. Medial endophallite (ME), dorsal (I) and ventral (II) views. E–F. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. G–H. Superior right peripheral endophallite (SRP), dorsal (I) and ventral (II) views.	Fig. 26. Endophallus and endophallites of the Phanaeus achilles species group. Respectively P. achilles Boheman, 1858 (CEMT) and P. arnaudorum sp. nov., paratype (CEMT). A–B. Endophallus in dorsal view. C–D. Medial endophallite (ME), dorsal (I) and ventral (II) views. E–F. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. G–H. Superior right peripheral endophallite (SRP), dorsal (I) and ventral (II) views.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059611FF94B94CFDE018FA669E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872545/files/figure.png	https://doi.org/10.5281/zenodo.21872545	Fig. 1. External morphological differences between the Phanaeus chalcomelas species group and the P. achilles species group. A–B. Females head in dorsal view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. achilles Boheman, 1858 (CEMT). C. P. arnaudorum sp. nov. (CEMT), elytra in dorsal view. D–E. Anteromedial portion of pronotum and transverse carina of the head of females. D. P. cambeforti (CEMT). E. P. achilles (CEMT). F–G. Pronotal disc of major males in dorsal view. F. P. chalcomelas (Perty, 1830) (CEMT). G. P. achilles (CEMT). H. P. lecourti Arnaud, 2000 (CEMT), elytra in dorsal view.	Fig. 1. External morphological differences between the Phanaeus chalcomelas species group and the P. achilles species group. A–B. Females head in dorsal view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. achilles Boheman, 1858 (CEMT). C. P. arnaudorum sp. nov. (CEMT), elytra in dorsal view. D–E. Anteromedial portion of pronotum and transverse carina of the head of females. D. P. cambeforti (CEMT). E. P. achilles (CEMT). F–G. Pronotal disc of major males in dorsal view. F. P. chalcomelas (Perty, 1830) (CEMT). G. P. achilles (CEMT). H. P. lecourti Arnaud, 2000 (CEMT), elytra in dorsal view.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059611FF94B94CFDE018FA669E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872607/files/figure.png	https://doi.org/10.5281/zenodo.21872607	Fig. 22. Phanaeus achilles species group in dorsal view. A–B. P. achilles Boheman, 1858 (CEMT). A. Major male. B. Female. C–D. P. arnaudorum sp. nov. C. Holotype, major male (CEMT). D. Paratype, female (CEMT).	Fig. 22. Phanaeus achilles species group in dorsal view. A–B. P. achilles Boheman, 1858 (CEMT). A. Major male. B. Female. C–D. P. arnaudorum sp. nov. C. Holotype, major male (CEMT). D. Paratype, female (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059611FF94B94CFDE018FA669E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872611/files/figure.png	https://doi.org/10.5281/zenodo.21872611	Fig. 23. Phanaeus achilles species group. A. P. achilles Boheman, 1858 ♂ (CEMT), frontal view. B. P. arnaudorum sp. nov., holotype, ♂ (CEMT), frontal view. C. P. achilles Boheman, 1858 ♂ (CEMT), posterior view. D. P. arnaudorum sp. nov., holotype, ♂ (CEMT), posterior view. E. P. achilles Boheman, 1858 ♀ (CEMT), frontal view. F. P. arnaudorum sp. nov., paratype, ♀ (CEMT), frontal view.	Fig. 23. Phanaeus achilles species group. A. P. achilles Boheman, 1858 ♂ (CEMT), frontal view. B. P. arnaudorum sp. nov., holotype, ♂ (CEMT), frontal view. C. P. achilles Boheman, 1858 ♂ (CEMT), posterior view. D. P. arnaudorum sp. nov., holotype, ♂ (CEMT), posterior view. E. P. achilles Boheman, 1858 ♀ (CEMT), frontal view. F. P. arnaudorum sp. nov., paratype, ♀ (CEMT), frontal view.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059611FF94B94CFDE018FA669E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872615/files/figure.png	https://doi.org/10.5281/zenodo.21872615	Fig. 24. Major males of Phanaeus achilles species group in fronto-lateral view. A, C. P. achilles Boheman, 1858 (CEMT). B, D. P. arnaudorum sp. nov., holotype (CEMT). Arrows indicate the anteromedial pronotal tubercles, larger in P. achilles; asterisk arrows indicate the anterolateral pronotal fossae, larger in P. arnaudorum.	Fig. 24. Major males of Phanaeus achilles species group in fronto-lateral view. A, C. P. achilles Boheman, 1858 (CEMT). B, D. P. arnaudorum sp. nov., holotype (CEMT). Arrows indicate the anteromedial pronotal tubercles, larger in P. achilles; asterisk arrows indicate the anterolateral pronotal fossae, larger in P. arnaudorum.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059611FF94B94CFDE018FA669E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872617/files/figure.png	https://doi.org/10.5281/zenodo.21872617	Fig. 25. Aedeagus of the Phanaeus achilles species group. A. P. achilles Boheman, 1858 (CEMT). B. P. arnaudorum sp. nov., paratype (CEMT), dorsal (I) and lateral (II) views.	Fig. 25. Aedeagus of the Phanaeus achilles species group. A. P. achilles Boheman, 1858 (CEMT). B. P. arnaudorum sp. nov., paratype (CEMT), dorsal (I) and lateral (II) views.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059611FF94B94CFDE018FA669E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872619/files/figure.png	https://doi.org/10.5281/zenodo.21872619	Fig. 26. Endophallus and endophallites of the Phanaeus achilles species group. Respectively P. achilles Boheman, 1858 (CEMT) and P. arnaudorum sp. nov., paratype (CEMT). A–B. Endophallus in dorsal view. C–D. Medial endophallite (ME), dorsal (I) and ventral (II) views. E–F. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. G–H. Superior right peripheral endophallite (SRP), dorsal (I) and ventral (II) views.	Fig. 26. Endophallus and endophallites of the Phanaeus achilles species group. Respectively P. achilles Boheman, 1858 (CEMT) and P. arnaudorum sp. nov., paratype (CEMT). A–B. Endophallus in dorsal view. C–D. Medial endophallite (ME), dorsal (I) and ventral (II) views. E–F. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. G–H. Superior right peripheral endophallite (SRP), dorsal (I) and ventral (II) views.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059611FF94B94CFDE018FA669E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872629/files/figure.png	https://doi.org/10.5281/zenodo.21872629	Fig. 29. A–B. Lectotype of Phanaeus achilles Boheman, 1858 (here designated) (NRMS). A. Typical dorsal color. B. Labels. C–D. Green specimen wrongly cited as holotype of Phanaeus achilles by Edmonds (1994) (NRMS) and its labels.	Fig. 29. A–B. Lectotype of Phanaeus achilles Boheman, 1858 (here designated) (NRMS). A. Typical dorsal color. B. Labels. C–D. Green specimen wrongly cited as holotype of Phanaeus achilles by Edmonds (1994) (NRMS) and its labels.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F8059611FF94B94CFDE018FA669E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872623/files/figure.png	https://doi.org/10.5281/zenodo.21872623	Fig. 27. Geographic distribution of Phanaeus achilles Boheman, 1858 and P. arnaudorum sp. nov. The star is sensu Figueroa et al. (2014).	Fig. 27. Geographic distribution of Phanaeus achilles Boheman, 1858 and P. arnaudorum sp. nov. The star is sensu Figueroa et al. (2014).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805960EFF90B95BFE861F8A6277.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872545/files/figure.png	https://doi.org/10.5281/zenodo.21872545	Fig. 1. External morphological differences between the Phanaeus chalcomelas species group and the P. achilles species group. A–B. Females head in dorsal view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. achilles Boheman, 1858 (CEMT). C. P. arnaudorum sp. nov. (CEMT), elytra in dorsal view. D–E. Anteromedial portion of pronotum and transverse carina of the head of females. D. P. cambeforti (CEMT). E. P. achilles (CEMT). F–G. Pronotal disc of major males in dorsal view. F. P. chalcomelas (Perty, 1830) (CEMT). G. P. achilles (CEMT). H. P. lecourti Arnaud, 2000 (CEMT), elytra in dorsal view.	Fig. 1. External morphological differences between the Phanaeus chalcomelas species group and the P. achilles species group. A–B. Females head in dorsal view. A. P. cambeforti Arnaud, 1982 (CEMT). B. P. achilles Boheman, 1858 (CEMT). C. P. arnaudorum sp. nov. (CEMT), elytra in dorsal view. D–E. Anteromedial portion of pronotum and transverse carina of the head of females. D. P. cambeforti (CEMT). E. P. achilles (CEMT). F–G. Pronotal disc of major males in dorsal view. F. P. chalcomelas (Perty, 1830) (CEMT). G. P. achilles (CEMT). H. P. lecourti Arnaud, 2000 (CEMT), elytra in dorsal view.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805960EFF90B95BFE861F8A6277.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872607/files/figure.png	https://doi.org/10.5281/zenodo.21872607	Fig. 22. Phanaeus achilles species group in dorsal view. A–B. P. achilles Boheman, 1858 (CEMT). A. Major male. B. Female. C–D. P. arnaudorum sp. nov. C. Holotype, major male (CEMT). D. Paratype, female (CEMT).	Fig. 22. Phanaeus achilles species group in dorsal view. A–B. P. achilles Boheman, 1858 (CEMT). A. Major male. B. Female. C–D. P. arnaudorum sp. nov. C. Holotype, major male (CEMT). D. Paratype, female (CEMT).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805960EFF90B95BFE861F8A6277.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872611/files/figure.png	https://doi.org/10.5281/zenodo.21872611	Fig. 23. Phanaeus achilles species group. A. P. achilles Boheman, 1858 ♂ (CEMT), frontal view. B. P. arnaudorum sp. nov., holotype, ♂ (CEMT), frontal view. C. P. achilles Boheman, 1858 ♂ (CEMT), posterior view. D. P. arnaudorum sp. nov., holotype, ♂ (CEMT), posterior view. E. P. achilles Boheman, 1858 ♀ (CEMT), frontal view. F. P. arnaudorum sp. nov., paratype, ♀ (CEMT), frontal view.	Fig. 23. Phanaeus achilles species group. A. P. achilles Boheman, 1858 ♂ (CEMT), frontal view. B. P. arnaudorum sp. nov., holotype, ♂ (CEMT), frontal view. C. P. achilles Boheman, 1858 ♂ (CEMT), posterior view. D. P. arnaudorum sp. nov., holotype, ♂ (CEMT), posterior view. E. P. achilles Boheman, 1858 ♀ (CEMT), frontal view. F. P. arnaudorum sp. nov., paratype, ♀ (CEMT), frontal view.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805960EFF90B95BFE861F8A6277.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872615/files/figure.png	https://doi.org/10.5281/zenodo.21872615	Fig. 24. Major males of Phanaeus achilles species group in fronto-lateral view. A, C. P. achilles Boheman, 1858 (CEMT). B, D. P. arnaudorum sp. nov., holotype (CEMT). Arrows indicate the anteromedial pronotal tubercles, larger in P. achilles; asterisk arrows indicate the anterolateral pronotal fossae, larger in P. arnaudorum.	Fig. 24. Major males of Phanaeus achilles species group in fronto-lateral view. A, C. P. achilles Boheman, 1858 (CEMT). B, D. P. arnaudorum sp. nov., holotype (CEMT). Arrows indicate the anteromedial pronotal tubercles, larger in P. achilles; asterisk arrows indicate the anterolateral pronotal fossae, larger in P. arnaudorum.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805960EFF90B95BFE861F8A6277.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872617/files/figure.png	https://doi.org/10.5281/zenodo.21872617	Fig. 25. Aedeagus of the Phanaeus achilles species group. A. P. achilles Boheman, 1858 (CEMT). B. P. arnaudorum sp. nov., paratype (CEMT), dorsal (I) and lateral (II) views.	Fig. 25. Aedeagus of the Phanaeus achilles species group. A. P. achilles Boheman, 1858 (CEMT). B. P. arnaudorum sp. nov., paratype (CEMT), dorsal (I) and lateral (II) views.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805960EFF90B95BFE861F8A6277.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872619/files/figure.png	https://doi.org/10.5281/zenodo.21872619	Fig. 26. Endophallus and endophallites of the Phanaeus achilles species group. Respectively P. achilles Boheman, 1858 (CEMT) and P. arnaudorum sp. nov., paratype (CEMT). A–B. Endophallus in dorsal view. C–D. Medial endophallite (ME), dorsal (I) and ventral (II) views. E–F. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. G–H. Superior right peripheral endophallite (SRP), dorsal (I) and ventral (II) views.	Fig. 26. Endophallus and endophallites of the Phanaeus achilles species group. Respectively P. achilles Boheman, 1858 (CEMT) and P. arnaudorum sp. nov., paratype (CEMT). A–B. Endophallus in dorsal view. C–D. Medial endophallite (ME), dorsal (I) and ventral (II) views. E–F. Additional endophallite (AE), dorsal (I), lateral (II) and ventral (III) views. G–H. Superior right peripheral endophallite (SRP), dorsal (I) and ventral (II) views.	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
03B6F805960EFF90B95BFE861F8A6277.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21872623/files/figure.png	https://doi.org/10.5281/zenodo.21872623	Fig. 27. Geographic distribution of Phanaeus achilles Boheman, 1858 and P. arnaudorum sp. nov. The star is sensu Figueroa et al. (2014).	Fig. 27. Geographic distribution of Phanaeus achilles Boheman, 1858 and P. arnaudorum sp. nov. The star is sensu Figueroa et al. (2014).	2026-07-23	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.		Zenodo	biologists	Bordin, Bruna Ribeiro;Arias-Buriticá, Jorge;Maldaner, María Eduarda;Vaz-De-Mello, Fernando Z.			
