taxonID	type	description	language	source
03B687CFF21FFFFFFDADFBDB447BF9A7.taxon	type_taxon	Type species Araneus pallidulus Clerck, 1757.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF21FFFFFFDADFBDB447BF9A7.taxon	discussion	Remarks The genus Clubiona has often functioned as a repository for species that could not be confidently assigned to other, better-defined genera. And due to the high species diversity in Clubiona, several subgenera and species groups have been proposed, resulting in a rather complex taxonomic situation. Clubiona is also one of the spider genera with the largest number of generic-level synonyms; currently, 14 genera or subgenera are considered junior synonyms (Zhang et al. 2021 a, 2021 c, 2021 d, 2025; WSC 2025). This reflects the considerable taxonomic complexity within the genus.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF218FFF8FDEFFC0843A9F981.taxon	vernacular_names	ŀ 样ú巢ss	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF218FFF8FDEFFC0843A9F981.taxon	description	Fig. 1	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF218FFF8FDEFFC0843A9F981.taxon	diagnosis	Diagnosis and description See Jäger & Dankittipakul (2010) for the female; Zhang et al. (2021 a) for the male. Female habitus as in Fig. 1 A – B; epigyne as in Fig. 1 C – D.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF218FFF8FDEFFC0843A9F981.taxon	materials_examined	Material examined CHINA • 1 ♀; Guangxi Zhuang Autonomous Region, Fangchenggang City, Shiwandashan National Nature Reserve, Pinglong Station; 21 ° 50 ′ N, 107 ° 53 ′ E; 527 m a. s. l.; 29 Apr. 2021; J. Liu leg; HNU 1396.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF218FFF8FDEFFC0843A9F981.taxon	distribution	Distribution China (Guangxi Zhuang Autonomous Region, Yunnan), Laos.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF218FFF4FE33F9C04375FEE5.taxon	vernacular_names	ḲNJú 巢ss	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF218FFF4FE33F9C04375FEE5.taxon	description	Fig. 2	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF218FFF4FE33F9C04375FEE5.taxon	diagnosis	Diagnosis and description See Barrion et al. (2012) for the original description of the male, and Zhang et al. (2020) for a redescription of the male and first description of the female. Male habitus as in Fig. 2 A – B; palp as in Fig. 2 C – F.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF218FFF4FE33F9C04375FEE5.taxon	materials_examined	Material examined CHINA • 1 ♂; Guangxi Zhuang Autonomous Region, Fangchenggang City, Shiwandashan National Nature Reserve, Songbai Station; 21 ° 58 ′ N, 108 ° 01 ′ E; 303 m a. s. l.; 1 May 2021; J. Liu leg; HNU 1402.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF218FFF4FE33F9C04375FEE5.taxon	distribution	Distribution China (Guangxi Zhuang Autonomous Region, Hainan).	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF214FFF7FDB2FE3D415EFCA3.taxon	vernacular_names	彦之ú巢ss	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF214FFF7FDB2FE3D415EFCA3.taxon	description	Figs 3 – 4	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF214FFF7FDB2FE3D415EFCA3.taxon	diagnosis	Diagnosis The male C. yanzhii resembles that of C. subapplanata Wang, Chen & Zhang, 2018 in the shape of the palp (compare Fig. 3 C – E with Wang et al. 2018: figs 14 a – b, 15 c – d) but can be distinguished by the absence of the LTA, the membranous VTA, the transparent, membranous conductor, the small EB, and the TA with a straight and pointed distal end (Fig. 3 C – E) (vs presence of the LTA, a distinctly sclerotized VTA, absence of the conductor, a distinctly swollen EB, and a twisted distal end of the TA in C. subapplanata, as in Wang et al. 2018: figs 14 a – b, 15 c – d). For the diagnosis of the female, see Zhang & Yu (2020).	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF214FFF7FDB2FE3D415EFCA3.taxon	materials_examined	Material examined CHINA • 1 ♂; Guangxi Zhuang Autonomous Region, Fangchenggang City, Shiwandashan National Nature Reserve, Pinglong Station; 21 ° 50 ′ N, 107 ° 53 ′ E; 749 m a. s. l.; 28 Apr. 2021; A. He, J. Liu, Y. Liang, R. Liao, Q. Li, H. Zhang, S. Zhu, Z. Zhao and Y. Liu leg; HNU 1372 • 2 ♀♀; same data as for preceding; 21 ° 50 ′ N, 107 ° 53 ′ E; 619 m a. s. l.; 28 Apr. 2021; A. He, J. Liu, Y. Liang, R. Liao, Q. Li, H. Zhang, S. Zhu, Z. Zhao and Y. Liu leg; HNU 1391 to HNU 1392 • 1 ♀; same data as for preceding, Banba Village, Dongkou Station; 21 ° 43 ′ N, 107 ° 32 ′ E; 820 m a. s. l.; 27 Apr. 2021; A. He, J. Liu, Y. Liang, R. Liao, Q. Li, H. Zhang, S. Zhu, Z. Zhao and Y. Liu leg.; HNU 1393.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF214FFF7FDB2FE3D415EFCA3.taxon	description	Description Male (HNU 1372) Total length 5.08; carapace 2.28 long, 1.60 wide; abdomen 2.66 long, 1.51 wide. Carapace elongate-oval, brown, fovea red; cephalic region distinctly narrowed; cervical groove indistinct, radial grooves distinct; anterior part densely covered with fine white hairs (Fig. 3 A). Eyes: AER distinctly recurved, PER wider than AER and nearly straight in dorsal view. Eye sizes and interdistances: AME 0.10, ALE 0.12, PME 0.12, PLE 0.12, AME – AME 0.09, AME – ALE 0.05, PME – PME 0.21, PME – PLE 0.13, MOQL 0.36, MOQA 0.30, MOQP 0.44. Chelicerae brown, darker than carapace, with five promarginal and four retromarginal teeth. Sternum yellow, oval, with long black hairs along the margins, 1.28 long, 0.77 wide. Labium and endites light brown, longer than wide. Legs colored as sternum, without markings (Fig. 3 B). Leg measurements: I 5.58 (1.56, 0.77, 1.47, 1.15, 0.63), II 6.23 (1.82, 0.75, 1.65, 1.36, 0.65), III 5.30 (1.54, 0.75, 1.13, 1.44, 0.44), IV 7.14 (2.03, 0.83, 1.79, 2.14, 0.62). Leg formula 4213. Abdomen elongate-oval, dark brown, with blackish-brown markings; anterior part flat, with thick tuft of setae (Fig. 3 A). Ventral side colored as dorsal side, with blackish-brown markings; book lung area pale yellow. Spinnerets short, pale yellow (Fig. 3 B). PALP (Fig. 3 C – E). Femur and patella unmodified. Tibia short, ca 2 / 5 as long as cymbium, with two apophyses: RTA bluntly triangular, about 120 °, moderately sclerotized; VTA bluntly rounded, partly membranous, longer than wide. Bulb distinctly bulging and prolapsed, sperm duct indistinct in ventral view. EB represented by enlarged tubercle, ca 2 / 7 as long as embolus at 12 o’clock position on tegulum; free part of embolus filiform, ca 5 / 7 of its length, with tip curved ventrally. Conductor nearly as long as embolus, extending alongside embolus, membranous, lamellar. TA beak-shaped, originating from retrolateral side of tegulum, with broad base and sharp apex pointing prolaterally. Female For diagnosis and description, see Zhang & Yu (2020). Habitus as in Fig. 4 A – B, epigyne as in Fig. 4 C – D.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF214FFF7FDB2FE3D415EFCA3.taxon	distribution	Distribution China (Guangxi Zhuang Autonomous Region, Hunan).	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF214FFF7FDB2FE3D415EFCA3.taxon	discussion	Remarks The structure labeled as TA in Fig. 3 D has been identified as the conductor in the similar species C. subapplanata (Wang et al. 2018). However, we consider this structure to be the tegular apophysis (TA) instead, since it is a sclerotized projection arising from the distal margin of the tegulum. In our specimen, a membranous conductor accompanying the embolus is clearly visible, further supporting our interpretation.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF217FFF7FDBEFCE44346FBB7.taxon	vernacular_names	Dkss 属	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF217FFF7FDBEFCE44346FBB7.taxon	type_taxon	Type species Clubiona brachyptera Zhu & Chen, 2012; gender feminine.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF217FFF1FE05FBF2423EFA6A.taxon	vernacular_names	Ḃ 氏Dkss	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF217FFF1FE05FBF2423EFA6A.taxon	description	Fig. 5	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF217FFF1FE05FBF2423EFA6A.taxon	diagnosis	Diagnosis The female of this species can be distinguished from that of all other congeners by its relatively short DPS that is ca ½ as long as bursa, and the visibility of the CD in dorsal view (Fig. 5 E – F) (vs DPS relatively long, more than ⅔ as long, CD covered by large SP in dorsal view in all other species of Femorbiona, such as F. brachyptera, F. pinglong sp. nov. and F. triangula sp. nov.; as in Zhang et al. 2021 b: fig. 2 c – d; Figs 7 E – F, 9 E – F). It can also be distinguished from F. brachyptera, F. phami, F. shenzhen and F. triangula sp. nov. by the CO being centrally located on the epigynal plate (Fig. 5 C – D) (vs CO located posteriorly in the latter four species, as in Zhang et al. 2021 b: figs 2 a – b, 4 a – b, 6 a – b; Fig. 9 C – D).	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF217FFF1FE05FBF2423EFA6A.taxon	materials_examined	Material examined CHINA • 1 ♀; Guangxi Zhuang Autonomous Region, Fangchenggang City, Shiwandashan National Nature Reserve, Banba Village, Dongkou Station; 21 ° 43 ′ N, 107 ° 32 ′ E; 820 m a. s. l.; 27 Apr. 2021; J. Liu leg; HNU 1399.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF217FFF1FE05FBF2423EFA6A.taxon	description	Description Female (HNU 1399) Total length 3.24; carapace 1.34 long, 0.95 wide; abdomen 1.79 long, 1.10 wide. Carapace oval, light orange, fovea red; cephalic region slightly narrowed; cervical and radial grooves indistinct (Fig. 5 A). Eyes: AER distinctly recurved, PER wider than AER, almost straight in dorsal view. Eye sizes and interdistances: AME 0.06, ALE 0.09, PME 0.08, PLE 0.07, AME – AME 0.07, AME – ALE 0.06, PME – PME 0.14, PME – PLE 0.10, MOQL 0.22, MOQA 0.18, MOQP 0.31. Chelicerae light orange, with five promarginal and four retromarginal teeth. Sternum yellow, 0.77 long, 0.55 wide. Labium and endites colored as chelicerae, longer than wide. Legs colored as sternum, without markings. Leg measurements: I 2.27 (0.59, 0.30, 0.57, 0.52, 0.29), II 2.31 (0.55, 0.42, 0.68, 0.40, 0.26), III 2.19 (0.68, 0.35, 0.33, 0.46, 0.37), IV 3.14 (0.67, 0.43, 0.73, 0.69, 0.62). Leg formula 4213. Abdomen yellowish-white, uniformly colored, densely clothed with setae, without patterns, with two pairs of muscle depressions situated centrally on dorsal side (Fig. 5 A). Ventral side colored as dorsal side, without patterns. Spinnerets short, yellow (Fig. 5 B). EPIGYNE (Fig. 5 C – F). Epigynal plate wider than long, with SP and BS distinctly visible through integument in ventral view (Fig. 5 C). CO small and indistinct, situated medially on epigynal plate. CD sclerotized, separated by about twice their own diameter, visible in dorsal view, extending anteriorly to connect with BS. SP tubular and sinuous; DPS slightly convoluted, extending anteriorly; VPS about 2 × as long as wide, nearly parallel and closely positioned, each bearing digitiform glands. FD indistinct. BS reniform, translucent, occupying nearly ⅔ of the epigynal plate, slightly longer than wide, with smooth surface. Male Unknown.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF217FFF1FE05FBF2423EFA6A.taxon	distribution	Distribution China (Guangxi Zhuang Autonomous Region, Taiwan).	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF217FFF1FE05FBF2423EFA6A.taxon	discussion	Remarks Femorbiona kayashimai comb. nov. was originally described as Clubiona kayashimai by Ono (1994), based solely on a female specimen with simple hand-drawn illustrations of the genitalia. Since then, the species has not been reported again for nearly 30 years. Although we were unable to examine the type material of C. kayashimai, we contacted Hirotsugu Ono, who confirmed that our identification of the present specimen is correct.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF211FFEDFDDBF9A9433CFC35.taxon	vernacular_names	平DZDkss	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF211FFEDFDDBF9A9433CFC35.taxon	description	urn: lsid: zoobank. org: act: FE 3 DFBBE- 6 D 52 - 4 AAD- 8452 - 9 E 6 C 213 D 18 F 9 Figs 6 – 7	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF211FFEDFDDBF9A9433CFC35.taxon	diagnosis	Diagnosis The male of the new species can be easily distinguished from that of all other congeners by its membranous and transparent conductor (Fig. 6 D – E) (vs weakly sclerotized in F. triangula sp. nov. and F. brachyptera (Zhu & Chen, 2012), as in Fig. 8 D and Zhang et al. 2021 b: fig. 7 a, respectively; absent in F. phami Yu & Li, 2021 and F. shenzhen Yu & Li, 2021, as in Zhang et al. 2021 b: fig. 7 b – c). It can also be distinguished from other species of Femorbiona by the absence of PPA (Fig. 6 C – D) (vs present, such as in F. brachyptera, F. phami, F. Shenzhen, F. triangula sp. nov.; as in Zhang et al. 2021 b: fig. 7 a – c; Fig. 8 C – D). The female of this species can be distinguished from that of other species of Femorbiona by its DPS, which extends anteriorly and then twists into an S-shaped terminal portion in dorsal view (Fig. 7 E – F) (vs a DPS that extends anteriorly with only a slight downward bend, such as F. phami in Zhang et al. 2021 b: fig. 4 c – d and F. triangula sp. nov. in Fig. 9 E – F).	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF211FFEDFDDBF9A9433CFC35.taxon	etymology	Etymology The specific epithet refers to the type locality; noun in apposition.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF211FFEDFDDBF9A9433CFC35.taxon	materials_examined	Type material Holotype CHINA • ♂; Guangxi Zhuang Autonomous Region, Fangchenggang City, Shiwandashan National Nature Reserve, Pinglong Station; 21 ° 50 ′ N, 107 ° 53 ′ E; 619 m a. s. l.; 28 Apr. 2021; J. Liu leg; HNU 1368. Paratypes CHINA • 1 ♂; same data as for holotype; HNU 1371 • 1 ♀; same data as for holotype; HNU 1398 • 1 ♂; same data as for holotype; 21 ° 50 ′ N, 107 ° 53 ′ E; 339 m a. s. l.; 29 Apr. 2021, J. Liu leg; HNU 1370.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF211FFEDFDDBF9A9433CFC35.taxon	description	Description Male (holotype HNU 1368) Total length 3.93; carapace 1.82 long, 1.30 wide; abdomen 2.10 long, 1.18 wide. Carapace oval, yellow, with anterior part light orange and smooth, while remaining area densely covered with short, fine setae, fovea red; cephalic region slightly narrowed; cervical and radial grooves indistinct (Fig. 6 A). Eyes: AER distinctly recurved; PER wider than AER and nearly straight in dorsal view. Eye sizes and interdistances: AME 0.09, ALE 0.11, PME 0.08, PLE 0.11; AME – AME 0.05, AME – ALE 0.05, PME – PME 0.22, PME – PLE 0.11; MOQL 0.32, MOQA 0.23, MOQP 0.43. Chelicerae light orange, with six promarginal and five retromarginal teeth. Sternum yellow with long black hairs along margins, 1.06 long, 0.73 wide. Labium and endites light orange, longer than wide. Legs same color as sternum, without markings (Fig. 6 A – B). Leg measurements: I 3.88 (1.00, 0.55, 1.10, 0.80, 0.43), II 4.46 (1.34, 0.62, 1.34, 0.85, 0.31), III 3.60 (1.04, 0.46, 0.88, 0.90, 0.32), IV 4.57 (1.57, 0.37, 1.29, 0.93, 0.41). Leg formula: 4213. Abdomen yellow, anterior part flat with a thick tuft of setae (Fig. 6 A). Ventral side slightly paler than dorsal; book lung area pale yellow, with brownish patch located centrally between book lungs. Spinnerets short, yellow (Fig. 6 B). PALP (Fig. 6 C – E). Femur with finger-shaped retrolateral apophysis (FA), ca ½ – ⅓ as long as femur. Patella unmodified. Tibia short, ca 3 / 5 as long as patella, with VTA bearing blunt apex. Bulb distinctly swollen and protruding; tegulum ca 1.5 × as long as wide, SD indistinct in ventral view. Embolus ca 1 / 7 as long as tegulum, arising at 11 o’clock position; EB swollen, abruptly tapering toward short embolic apex. Conductor short, translucent, membranous, and indistinct, ca ⅓ as long as embolus. Female (HNU 1398) Total length 4.75; carapace 1.55 long, 1.04 wide; abdomen 2.69 long, 1.50 wide. Carapace oval, light orange, fovea red; cephalic region slightly narrowed, orange; cervical and radial grooves indistinct (Fig. 7 A). Eyes: AER distinctly recurved, PER wider than AER, almost straight in dorsal view. Eye sizes and interdistances: AME 0.07, ALE 0.08, PME 0.08, PLE 0.09, AME – AME 0.06, AME – ALE 0.06, PME – PME 0.16, PME – PLE 0.11, MOQL 0.23, MOQA 0.19, MOQP 0.33. Chelicerae orange, with four promarginal and five retromarginal teeth. Sternum yellow, 0.91 long, 0.61 wide. Labium and endites light orange, longer than wide. Legs colored as sternum, without markings. Leg measurements: I 3.22 (0.90, 0.54, 0.77, 0.57, 0.44), II 3.76 (1.14, 0.52, 1.03, 0.65, 0.42), III 3.31 (1.00, 0.41, 0.69, 0.75, 0.46), IV 4.08 (1.36, 0.47, 1.14, 0.62, 0.49). Leg formula 4231. Abdomen grey, uniformly colored, densely clothed with setae, lacking distinct patterns; two pairs of muscle depressions present medially on dorsal side (Fig. 7 A). Ventral side colored as dorsal side, without patterns. Spinnerets short, yellow (Fig. 7 B). EPIGYNE (Fig. 7 C – F). Epigynal plate wider than long. CO semicircular, situated medially on epigynal plate. CD sclerotized, closely spaced, concealed by large spermathecae in dorsal view (Fig. 7 E). SP tubular and sinuous; DPS exceptionally long, extending anteriorly and then twisting into S-shaped terminal portion in dorsal view (Fig. 7 E – F); VPS tapering distally (Fig. 7 E – F). FD indistinct. BS reniform, translucent, occupying nearly ⅔ of epigynal plate, longer than wide, with smooth surface.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF211FFEDFDDBF9A9433CFC35.taxon	distribution	Distribution Known only from the type locality in Guangxi Zhuang Autonomous Region, China.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF211FFEDFDDBF9A9433CFC35.taxon	discussion	Remarks We identify this species as a new member of Femorbiona based on the female specimen, which is consistent with the diagnostic characteristics of the genus Femorbiona Yu & Li, 2021 as defined by Zhang et al. (2021 b). However, the male specimen lacks a PPA, which does not fully conform to the current diagnosis of the genus. This indicates that the presence of a PPA may not be a consistent or defining character across all Femorbiona species.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20DFFE9FDD6FC0D44A8FCE3.taxon	vernacular_names	三ĦDkss	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20DFFE9FDD6FC0D44A8FCE3.taxon	description	urn: lsid: zoobank. org: act: DBF 91 FCB-BA 1 C- 4 A 73 - 878 F- 967 EAAACF 70 B Figs 8 – 9	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20DFFE9FDD6FC0D44A8FCE3.taxon	diagnosis	Diagnosis The male of the new species can be readily distinguished from that of its congeners by the conductor, which is slightly sclerotized, lamellar, and triangular in ventral view (Fig. 8 D) (vs elliptical and slightly curved around the embolus in F. brachyptera; absent in F. phami and F. shenzhen; translucent and membranous in F. pinglong, as in Zhang et al. 2021 b: figs 1 d, 3 d, 5 d; Fig. 6 D, respectively). The female of the new species resembles that of F. brachyptera in having the DPS following a double S-shaped course (compare Fig. 9 C – F with Zhang et al. 2021 b: fig. 2 a – d), but differs by the CD being clearly visible through the epigynal plate in ventral view (Fig. 9 C), and by the longer, semicircular connection between DPS and VPS (as indicated by the white dashed line in Fig. 9 F) (vs CD indistinct and the DPS directly connected to the VPS in F. brachyptera, as in Zhang et al. 2021 b: fig. 2 a – d). The female of the new species also resembles that of F. shenzhen in the shape of the CO and the hood, and the CD easily visible through the epigynal plate in ventral view (compare Fig. 9 C with Zhang et al. 2021 b: fig. 6 a), but differs by the DPS strongly convoluted, following a double S-shaped course (Fig. 9 E – F) (vs the DPS not strongly convoluted, following a C-shaped course, as in Zhang et al. 2021 b: fig. 6 c – d).	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20DFFE9FDD6FC0D44A8FCE3.taxon	etymology	Etymology The specific epithet ‘ triangula ’ (Latin adjective, meaning ‘ triangular’) refers to the triangular shape of the conductor.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20DFFE9FDD6FC0D44A8FCE3.taxon	materials_examined	Type material Holotype CHINA • ♂; Guangxi Zhuang Autonomous Region, Fangchenggang City, Shiwandashan National Nature Reserve, Pinglong Station; 21 ° 50 ′ N, 107 ° 53 ′ E; 527 m a. s. l.; 29 Apr. 2021; J. Liu leg; HNU 1373. Paratypes CHINA • 1 ♀; same data as for holotype; 21 ° 51 ′ N, 107 ° 54 ′ E; 339 m a. s. l.; 29 Apr. 2021; A. He, J. Liu, Y. Liang, R. Liao, Q. Li, H. Zhang, S. Zhu, Z. Zhao and Y. Liu leg; HNU 1376 • 2 ♀♀; same data as for holotype, Dalongshan Station; 22 ° 01 ′ N, 108 ° 08 ′ E; 233 m a. s. l.; 2 May 2021; A. He, J. Liu, Y. Liang, R. Liao, Q. Li, H. Zhang, S. Zhu, Z. Zhao and Y. Liu leg; HNU 1374 to HNU 1375 • 3 ♂♂; same data as for holotype; 21 ° 50 ′ N, 107 ° 53 ′ E; 619 m a. s. l.; 28 Apr. 2021; A. He, J. Liu, Y. Liang, R. Liao, Q. Li, H. Zhang, S. Zhu, Z. Zhao and Y. Liu leg; HNU 1377 to HNU 1379 • 4 ♀♀; same data as for holotype; 21 ° 50 ′ N, 107 ° 53 ′ E; 619 m a. s. l.; 28 Apr. 2021; A. He, J. Liu, Y. Liang, R. Liao, Q. Li, H. Zhang, S. Zhu, Z. Zhao and Y. Liu leg; HNU 1380 to HNU 1383 • 1 ♂; same data as for holotype; 21 ° 50 ′ N, 107 ° 53 ′ E; 749 m a. s. l.; 28 Apr. 2021; A. He, J. Liu, Y. Liang, R. Liao, Q. Li, H. Zhang, S. Zhu, Z. Zhao and Y. Liu leg; HNU 1384 • 3 ♀♀; same data as for holotype; 21 ° 50 ′ N, 107 ° 53 ′ E; 749 m a. s. l.; 28 Apr. 2021; A. He, J. Liu, Y. Liang, R. Liao, Q. Li, H. Zhang, S. Zhu, Z. Zhao and Y. Liu leg; HNU 1385 to HNU 1387 • 1 ♂; same data as for holotype; 21 ° 51 ′ N, 107 ° 54 ′ E; 339 m a. s. l.; 29 Apr. 2021; A. He, J. Liu, Y. Liang, R. Liao, Q. Li, H. Zhang, S. Zhu, Z. Zhao and Y. Liu leg; HNU 1388 • 1 ♀; same data as for holotype; 21 ° 51 ′ N, 107 ° 54 ′ E; 339 m a. s. l.; 29 Apr. 2021; A. He, J. Liu, Y. Liang, R. Liao, Q. Li, H. Zhang, S. Zhu, Z. Zhao and Y. Liu leg; HNU 1389 • 1 ♀; same data as for holotype, Wangle Station; 21 ° 54 ′ N, 107 ° 54 ′ E; 309 m a. s. l.; 30 Apr. 2021; A. He, J. Liu, Y. Liang, R. Liao, Q. Li, H. Zhang, S. Zhu, Z. Zhao and Y. Liu leg; HNU 1390.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20DFFE9FDD6FC0D44A8FCE3.taxon	description	Description Male (holotype HNU 1373) Total length 3.42; carapace 1.54 long, 1.09 wide; abdomen 1.76 long, 1.07 wide. Carapace oval, yellow, densely covered with short, fine setae, fovea black; cephalic region slightly narrowed; cervical and radial grooves indistinct (Fig. 8 A). Eyes: AER slightly recurved; PER wider than AER and almost straight in dorsal view. Eye sizes and interdistances: AME 0.07, ALE 0.10, PME 0.08, PLE 0.08; AME – AME 0.06, AME – ALE 0.04, PME – PME 0.19, PME – PLE 0.10; MOQL 0.23, MOQA 0.20, MOQP 0.36. Chelicerae yellow, with five promarginal and five retromarginal teeth. Sternum yellowish white, 0.86 long, 0.73 wide. Labium and endites yellow, same in color as chelicerae, longer than wide. Legs colored as sternum, without markings. Leg measurements: I 2.62 (0.75, 0.37, 0.64, 0.54, 0.32), II 2.89 (0.90, 0.45, 0.73, 0.52, 0.29), III 2.57 (0.81, 0.41, 0.55, 0.60, 0.20), IV 3.67 (1.05, 0.54, 0.89, 0.92, 0.27). Leg formula 4213. Abdomen elongate-oval, pale brown, densely clothed with setae, dorsally marked with numerous brown spots (Fig. 8 A). Ventral side similar in color to dorsal side, laterally marked with numerous brown spots. Spinnerets short, yellowish white (Fig. 8 B). PALP (Fig. 8 C – F). Femur with retrolateral apophysis near distal margin, and apophysis with broad base and slender finger-shaped distal end. Patella ca 2 × as long as and 1.3 × as wide as tibia, bearing round PPA. Tibia short, ca 2 / 7 as long as cymbium, with small, distally blunt, and slightly sclerotized RTA. Bulb inflated and prolapsed; SD faintly visible in ventral view. Embolus needle-like, black, short, situated at 12: 30 o’clock position; apex sharp, directed anterolaterally. EB forming enlarged tubercle. Conductor slightly shorter than EB, weakly sclerotized, lamellar, and triangular in ventral view. Female (paratype HNU 1376) Total length 3.40; carapace 1.53 long, 1.05 wide; abdomen 1.76 long, 1.03 wide. Carapace oval, light orange, densely covered with short, fine setae; fovea red; cephalic region slightly narrowed; cervical and radial grooves indistinct (Fig. 9 A). Eyes: AER distinctly recurved, PER slightly recurved and wider than AER in dorsal view. Eye sizes and interdistances: AME 0.09, ALE 0.08, PME 0.10, PLE 0.07, AME – AME 0.07, AME – ALE 0.05, PME – PME 0.19, PME – PLE 0.10, MOQL 0.23, MOQA 0.21, MOQP 0.34. Chelicerae light orange, with five promarginal and five retromarginal teeth. Sternum yellow, 0.82 long, 0.55 wide. Labium and endites same in color as chelicerae, longer than wide. Legs yellow, same as sternum, without markings. Leg measurements: I 2.56 (0.77, 0.37, 0.59, 0.49, 0.34), II 2.83 (0.73, 0.52, 0.70, 0.54, 0.34), III 2.46 (0.64, 0.31, 0.54, 0.68, 0.29), IV 3.32 (0.72, 0.46, 0.93, 0.85, 0.36). Leg formula 4213. Abdomen brown, uniformly colored, densely covered with setae, lacking distinct dorsal markings (Fig. 9 A). Ventral side slightly lighter in color than dorsal side, without markings. Spinnerets short, pale yellow (Fig. 9 B). EPIGYNE (Fig. 9 C – F). Epigynal plate wider than long, with SP and BS visible through the integument in ventral view (Fig. 9 C). Hood located posteriorly on epigynal plate, ca 2 / 5 as wide as epigyne, slightly procurved, arcuate. CO circular, located laterally at margin of hood, separated by ca 2 × their own diameter. CD thick, strongly sclerotized, obscured by large spermathecae in dorsal view. SP tubular and sinuous; DPS strongly convoluted like intestines, forming double S-shaped course directed anteriorly; VPS nearly parallel and closely positioned, extending forward for ca ⅔ of epigynal plate length. Connection between DPS and VPS long, semicircular (as indicated by the white dashed line in Fig. 9 F). BS oval, translucent, and large, nearly occupying entire epigynal plate, longer than wide, with smooth surface.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20DFFE9FDD6FC0D44A8FCE3.taxon	distribution	Distribution Known only from the type locality in Guangxi Zhuang Autonomous Region, China.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF209FFE9FDDDFB32438DF8F5.taxon	vernacular_names	Ḟ 形 ṳěss	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF209FFE9FDDDFB32438DF8F5.taxon	description	Fig. 10	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF209FFE9FDDDFB32438DF8F5.taxon	diagnosis	Diagnosis and description See Chen & Huang (2006) for the male, Huang & Chen (2012) for the female. Habitus as in Fig. 10 A – D; palp as in Fig. 10 E – G; epigyne as in Fig. 10 H – I.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF209FFE9FDDDFB32438DF8F5.taxon	materials_examined	Material examined CHINA • 1 ♀; Guangxi Zhuang Autonomous Region, Fangchenggang City, Shiwandashan National Nature Reserve, Songbai Station; 21 ° 58 ′ N, 108 ° 01 ′ E; 303 m a. s. l.; 1 May 2021; J. Liu leg; HNU 1400 • 1 ♂; same data as for preceding; HNU 1403.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF209FFE9FDDDFB32438DF8F5.taxon	distribution	Distribution China (Guangxi Zhuang Autonomous Region, Taiwan, Yunnan).	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF209FFE9FD9DFC2442DBFBF7.taxon	vernacular_names	ṳěss 属	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF209FFE9FD9DFC2442DBFBF7.taxon	type_taxon	Type species Matidia virens Thorell, 1878; gender feminine.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20BFFEBFDD6FEBB4134FE6D.taxon	vernacular_names	栅ú巢ss属	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20BFFEBFDD6FEBB4134FE6D.taxon	type_taxon	Type species Clubiona clandestina Menge, 1873; gender feminine; synonym of Porrhoclubiona genevensis (L. Koch, 1866).	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20BFFE5FDEAFDA54367FC8F.taxon	vernacular_names	松柏栅ú巢ss	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20BFFE5FDEAFDA54367FC8F.taxon	description	urn: lsid: zoobank. org: act: DE 36 B 5 D 9 - 0 D 2 F- 45 B 9 - BEE 1 - C 2 C 98 DD 85 DDC Fig. 11	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20BFFE5FDEAFDA54367FC8F.taxon	diagnosis	Diagnosis The female of the new species resembles P. pteronetoides (Deeleman-Reinhold, 2001) in having similarly large CO situated near the epigastric furrow, and translucent, wrinkled CD, and reniform SP (compare Fig. 11 C – F with Zhang et al. 2021 c: fig. 16 a – d). However, it can be easily distinguished from P. pteronetoides by the CD being distinctly longer and extending anteriorly beyond the SP, with their distal ends looping around the base of the spermathecae (Fig. 11 C – F) (vs CD not extending beyond the SP anteriorly and their twisted ends connecting directly to the SP in P. pteronetoides, as in Zhang et al. 2021 c: fig. 16 a – d).	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20BFFE5FDEAFDA54367FC8F.taxon	etymology	Etymology The specific epithet refers to the type locality; noun in apposition.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20BFFE5FDEAFDA54367FC8F.taxon	materials_examined	Type material Holotype CHINA • ♀; Guangxi Zhuang Autonomous Region, Fangchenggang City, Shiwandashan National Nature Reserve, Songbai Station; 21 ° 58 ′ N, 108 ° 01 ′ E; 303 m a. s. l.; 1 May 2021; J. Liu leg; HNU 1401.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20BFFE5FDEAFDA54367FC8F.taxon	description	Description Female (holotype HNU 1401) Total length 4.40; carapace 1.75 long, 1.25 wide; abdomen 2.53 long, 1.28 wide. Carapace oval, yellowish brown with anterior part light orange, densely covered with short, fine setae; fovea black, inconspicuous; cephalic region slightly narrowed; cervical groove distinct, radial groove indistinct (Fig. 11 A). Eyes: AER distinctly recurved; PER slightly procurved and wider than AER in dorsal view. Eye sizes and interdistances: AME 0.08, ALE 0.07, PME 0.10, PLE 0.09; AME – AME 0.06, AME – ALE 0.05, PME – PME 0.20, PME – PLE 0.09; MOQL 0.24, MOQA 0.23, MOQP 0.36. Chelicerae yellow, with three promarginal and two retromarginal teeth. Sternum bright yellow, 0.95 long, 0.55 wide. Labium and endites yellowish white, longer than wide. Legs yellowish white, unmarked. Leg measurements: I 3.21 (0.90, 0.54, 0.80, 0.58, 0.39), II 3.52 (1.02, 0.60, 0.91, 0.62, 0.37), III 2.98 (0.91, 0.37, 0.67, 0.66, 0.37), IV 4.48 (1.17, 0.55, 1.26, 1.06, 0.44). Leg formula: 4213. Abdomen grayish brown, uniformly colored, densely covered with setae, lacking distinct dorsal markings, with two pairs of muscle depressions centrally located on dorsal side (Fig. 11 A). Ventral side slightly paler than dorsal one, unmarked. Spinnerets short, yellow (Fig. 11 B). EPIGYNE (Fig. 11 C – F). Epigynal plate longer than wide, with SP and CD visible through integument in ventral view. CO large, oval, located posteriorly on epigynal plate near epigastric furrow. CD tubular, translucent, with wrinkled and ribbed appearance; ascend anteriorly in front of spermathecae, then curve inward, descend posteriorly in ventral view, and finally loop around base of spermathecae before connecting to both SP and BS. SP reniform. BS situated posteriorly, translucent, and globular. Male Unknown.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20BFFE5FDEAFDA54367FC8F.taxon	distribution	Distribution Known only from the type locality in Guangxi Zhuang Autonomous Region, China.	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
03B687CFF20BFFE5FDEAFDA54367FC8F.taxon	discussion	Remarks The new species also resembles Clubiona wulingensis Yu & Chen, 2017, which should likewise be transferred to the genus Porrhoclubiona (given the lack of available material and that this issue is beyond the scope of the present study, we retain its current taxonomic status). It can be distinguished from C. wulingensis by the dorsal part of the CD being ca ⅓ the diameter of the SP and positioned in the front of them in dorsal view (Fig. 11 C – F) (vs dorsal part ca half the diameter of the SP and positioned behind them in C. wulingensis, as in Yu & Chen 2017: figs 7 – 9).	en	Wu, Yecheng, Wen, Yingli, Yin, Haiqiang, Xu, Xiang (2026): A survey of clubionid spiders (Araneae: Clubionidae) from Shiwandashan National Nature Reserve, Guangxi, China. European Journal of Taxonomy 1034: 133-161, DOI: 10.5852/ejt.2026.1034.3151, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3151/14035
