taxonID	type	description	language	source
03AB5F65FFF5BA2EFF457FB7FA14733F.taxon	vernacular_names	[Japanese name: Futo-munekubo-hanekakushi-zoku]	en	Hashizume, Takuto, Maruyama, Munetoshi (2026): Oncerochara bolitocharoides gen. et sp. nov., of the tribe Homalotini (Coleoptera Staphylinidae: Aleocharinae) from Japan. Zootaxa 5748 (1): 113-119, DOI: 10.11646/zootaxa.5748.1.6, URL: https://doi.org/10.11646/zootaxa.5748.1.6
03AB5F65FFF5BA2EFF457FB7FA14733F.taxon	type_taxon	Type species. Oncerochara bolitocharoides Hashizume & Maruyama, sp. nov.	en	Hashizume, Takuto, Maruyama, Munetoshi (2026): Oncerochara bolitocharoides gen. et sp. nov., of the tribe Homalotini (Coleoptera Staphylinidae: Aleocharinae) from Japan. Zootaxa 5748 (1): 113-119, DOI: 10.11646/zootaxa.5748.1.6, URL: https://doi.org/10.11646/zootaxa.5748.1.6
03AB5F65FFF5BA2EFF457FB7FA14733F.taxon	etymology	Etymology. The generic name Oncerochara is a combination of “ ὀγκηρός ” (onkērós), the ancient Greek word meaning “ swollen ”, referring to the broad maxillary galea, and the termination “ - chara ” (originally from the ancient Greek word “ χάρις ” (charis) meaning “ grace, kindness ”) frequently used in generic names within Aleocharinae. The gender is feminine.	en	Hashizume, Takuto, Maruyama, Munetoshi (2026): Oncerochara bolitocharoides gen. et sp. nov., of the tribe Homalotini (Coleoptera Staphylinidae: Aleocharinae) from Japan. Zootaxa 5748 (1): 113-119, DOI: 10.11646/zootaxa.5748.1.6, URL: https://doi.org/10.11646/zootaxa.5748.1.6
03AB5F65FFF5BA2EFF457FB7FA14733F.taxon	diagnosis	Diagnosis. Oncerochara can be distinguished from other bolitocharine genera by the combination of the following character states: body elongate, more or less parallel-sided; maxillary galea very broad, about two times as broad as maxillary lacinia, apical part covered with numerous rows of minute setae; ligula of labium apically divided; two medial setae of prementum arranged transversely; medial pseudopore field narrow without pseudopores; hypomeron visible in lateral view; mesoventral medial longitudinal carina extending to apex of mesoventral process; apex of mesoventral process rounded; isthmus present; male sternite VI without medial lobate projection on posterior margin. This genus is more or less similar in general appearance, and in the morphology of the labrum, and the paramere and median lobe of the aedeagus, to the nine genera treated in the subtribe Bolitocharina by Ashe (1992). Among these nine genera, it shares with Phymatura Sahlberg, 1876, Silusida Casey, 1906, Stictalia Casey, 1906, Neotobia Ashe, 1992, and Austrasilida Ashe, 1992 the character states of the male sternite VI without a small medial lobe and the male sternite VIII with an asetose posterior margin. However, Stictalia, Phymatura, Silusida, and Neotobia differ in having a slender maxillary galea; a narrow, parallel-sided, apically divided ligula; and medially arranged premental setae, whereas Austrasilida differs in having a slender maxillary galea; a deeply divided ligula; and a male tergite VIII with an asetose posterior margin only at the middle portion. Among the genera occurring in Japan, Bolitochara is superficially similar to this genus, but differs in having the mouthparts similar to those of Phymatura and related genera; the male sternite VI with a small medial lobe; and the male sternite VIII lacking an asetose posterior margin.	en	Hashizume, Takuto, Maruyama, Munetoshi (2026): Oncerochara bolitocharoides gen. et sp. nov., of the tribe Homalotini (Coleoptera Staphylinidae: Aleocharinae) from Japan. Zootaxa 5748 (1): 113-119, DOI: 10.11646/zootaxa.5748.1.6, URL: https://doi.org/10.11646/zootaxa.5748.1.6
03AB5F65FFF5BA2EFF457FB7FA14733F.taxon	description	Description. Body (Fig. 1) somewhat stout; forebody with short setae. Head almost circular; eyes moderately large; infraorbital carina complete; occipital carina complete; setae directed anteriorly. Antennae with 11 antennomeres, incrassate. Mandibular molar region with denticles. Labrum (Fig. 2 A) transverse, with more than a dozen pairs of macrosetae; epipharynx (Fig. 2 A) with developed α-sensilla, β-sensilla located next to each other, γ-sensilla small. Maxillary lacinia (Fig. 2 D) with strong setae at apical half of adoral margin; galea (Fig. 2 D) extending beyond apex of lacinia, twice as broad as lacinia, with numerous minute trichae adorning distal lobe; palpus (Fig. 2 D) with four palpomeres; palpomere 1 small; palpomere 2 widest around apical 1 / 3, approximately three times longer than wide; palpomere 3 widest round apical 1 / 3, approximately three times longer than wide, longer than palpomere 2; palpomere 4 shorter than half of length of 3. Mentum (Fig. 2 B) trapezoidal, covered with pseudopores except lateral area; anterior margin moderately arcuate emarginate. Labial palpus (Fig. 2 C) with three palpomeres; palpomeres 1 and 2 fused laterally; palpomere 1 approximately twice longer than wide; palpomere 2 shorter than 1; palpomere 3 slender, longer than 2; ligula (Fig. 2 C) bifid, with a pair of short setae on dorsal side; prementum (Fig. 2 C) strongly transverse, with a pair of setal pores, two pairs of real pores, and several pseudopores; medial setae long, arranged transversely (one beside the other), bases moderately separated; medial pseudopore field narrow, without pseudopores; paraglossa with long thin setae at apical half. Thorax. Pronotum slightly transverse, nearly hexagonal; posterior half almost parallel-sided; setae of midline of pronotum directed anteriorly in anterior half and posteriorly in posterior half; setae of lateral area directed laterally. Hypomeron visible in lateral view. Mesoventrite with medial longitudinal carina extending to apex of mesoventral process mesocoxal cavities separated by intercoxal processes; mesoventral process twice as long as metaventral process, apex of mesoventral process rounded; metaventral process broad, broadly rounded apically; isthmus narrow. Legs slender; tarsal formula 4 - 4 - 5. Abdomen. Abdomen slightly narrowed posteriad; tergite III – V with deep basal impressions; basal impression of tergite VI shallow. Male. Tergite VII with longitudinal tubercle postero-medially; posterior margin of tergite VIII emarginate, with some small denticles; sternite IV without small medial lobe on posterior margin; sternite VIII margin triangularly produced and asetose. Female. Unknown.	en	Hashizume, Takuto, Maruyama, Munetoshi (2026): Oncerochara bolitocharoides gen. et sp. nov., of the tribe Homalotini (Coleoptera Staphylinidae: Aleocharinae) from Japan. Zootaxa 5748 (1): 113-119, DOI: 10.11646/zootaxa.5748.1.6, URL: https://doi.org/10.11646/zootaxa.5748.1.6
03AB5F65FFF5BA2EFF457FB7FA14733F.taxon	distribution	Distribution. Japan.	en	Hashizume, Takuto, Maruyama, Munetoshi (2026): Oncerochara bolitocharoides gen. et sp. nov., of the tribe Homalotini (Coleoptera Staphylinidae: Aleocharinae) from Japan. Zootaxa 5748 (1): 113-119, DOI: 10.11646/zootaxa.5748.1.6, URL: https://doi.org/10.11646/zootaxa.5748.1.6
03AB5F65FFF5BA2EFF457FB7FA14733F.taxon	discussion	Remarks. Several dozen genera of the tribe Homalotini remain unassigned to any subtribe. On the basis of the original descriptions and our examination of the type specimens, this genus can be distinguished from all of them.	en	Hashizume, Takuto, Maruyama, Munetoshi (2026): Oncerochara bolitocharoides gen. et sp. nov., of the tribe Homalotini (Coleoptera Staphylinidae: Aleocharinae) from Japan. Zootaxa 5748 (1): 113-119, DOI: 10.11646/zootaxa.5748.1.6, URL: https://doi.org/10.11646/zootaxa.5748.1.6
03AB5F65FFF6BA29FF457EE2FB9C7251.taxon	description	(Figs. 1 – 3)	en	Hashizume, Takuto, Maruyama, Munetoshi (2026): Oncerochara bolitocharoides gen. et sp. nov., of the tribe Homalotini (Coleoptera Staphylinidae: Aleocharinae) from Japan. Zootaxa 5748 (1): 113-119, DOI: 10.11646/zootaxa.5748.1.6, URL: https://doi.org/10.11646/zootaxa.5748.1.6
03AB5F65FFF6BA29FF457EE2FB9C7251.taxon	vernacular_names	[Japanese name: Futo-munekubo-hanekakushi]	en	Hashizume, Takuto, Maruyama, Munetoshi (2026): Oncerochara bolitocharoides gen. et sp. nov., of the tribe Homalotini (Coleoptera Staphylinidae: Aleocharinae) from Japan. Zootaxa 5748 (1): 113-119, DOI: 10.11646/zootaxa.5748.1.6, URL: https://doi.org/10.11646/zootaxa.5748.1.6
03AB5F65FFF6BA29FF457EE2FB9C7251.taxon	materials_examined	Type material. Holotype: male, (KUM), JAPAN: Honshu: “ º 山県º山İ ̊ ª 湖付ŕ [Toyama-ken, Toyama-shi, near Arimine-ko] / [JAPAN] Toyama-ken / Toyama-shi, nr. Arimine-ko, / 11 – 12. VI. 2022, / S. Imada & S. Yasukawa ”. Paratype. JAPAN: Honshu: 1 male, Tochigi-ken, Torobe, 24 VI 1984, M. Tao leg. (KUM). Type locality. Japan, Honshu, Toyama-ken, Toyama-shi, near Arimine-ko.	en	Hashizume, Takuto, Maruyama, Munetoshi (2026): Oncerochara bolitocharoides gen. et sp. nov., of the tribe Homalotini (Coleoptera Staphylinidae: Aleocharinae) from Japan. Zootaxa 5748 (1): 113-119, DOI: 10.11646/zootaxa.5748.1.6, URL: https://doi.org/10.11646/zootaxa.5748.1.6
03AB5F65FFF6BA29FF457EE2FB9C7251.taxon	diagnosis	Diagnosis. Same as genus; vide supra.	en	Hashizume, Takuto, Maruyama, Munetoshi (2026): Oncerochara bolitocharoides gen. et sp. nov., of the tribe Homalotini (Coleoptera Staphylinidae: Aleocharinae) from Japan. Zootaxa 5748 (1): 113-119, DOI: 10.11646/zootaxa.5748.1.6, URL: https://doi.org/10.11646/zootaxa.5748.1.6
03AB5F65FFF6BA29FF457EE2FB9C7251.taxon	description	Description. BL ≈ 3.38 – 3.43; FBL, 1.56 – 1.70. Body (Fig. 1) blackish brown; anterior and posterior margins of elytra, abdominal segment III – V, antennae, and basal and apical part of tibiae reddish brown; tarsi yellowish brown. Dorsal surface of forebody covered with short setae. Head almost circular; HW / HL: 1.15 – 1.18; HL, 0.44 – 0.47; HW, 0.52 – 0.54; eyes slightly prominent; surface densely punctured, with distinct microsculpture. Antennae slender; length 0.99; relative length of antennomeres 1 – 11 (n = 1): 24.4: 19.4: 17.7: 11.5: 10.8: 11.6: 10.0: 10.7: 11.8: 12.3: 29.5; antennomeres 1 – 4 and 11 longer than wide; 5 – 6 as long as wide; 7 – 10 wider than long (Ratio of length / width of antennomeres 1 – 11 (n = 1): 2.56, 2.26, 2.04, 1.18, 0.82, 0.79, 0.66, 0.70, 0.73, 0.72, 1.81). Thorax. Pronotum somewhat hexagonal, widest around middle, transverse (PW / PL: 1.23 – 1.27), wider than head (PW / HW: 1.25 – 1.32); PL, 0.53 – 0.57; PW, 0.65 – 0.72; punctures slightly coarser than those of head; microsculpture slightly weaker than head. Elytra transverse (EW / EL: 1.42 – 1.52), longer than pronotum (EL / PL: 1.12 – 1.17), wider than pronotum (EW / PW: 1.31 – 1.40); EL, 0.59 – 0.66; EW, 0.90 – 0.94; posterior margins weakly emarginate laterally; surface coarsely punctured, weakly microsculptured. Abdomen subparallel, slightly narrower than elytra; basal impressions of tergite III – V with large punctures; remainder of tergal surfaces with finer punctures; microsculpture isodiametric to transverse on tergite III – VI, almost isodiametric on tergite VII, more distinct on posterior segments. Male. Tergite VII with longitudinal tubercle postero-medially. Posterior margin of tergite VIII (Fig. 3 A) moderately emarginate, with some denticles. Paramere (Fig. 3 B) of aedeagus slender. Median lobe (Fig. 3 C) of aedeagus with relatively small basal bulb; apical process long, thin, curved, and slightly dilated apically; platelike sclerite of inner sac large, nearly oval; flagellum long. Female. Unknown.	en	Hashizume, Takuto, Maruyama, Munetoshi (2026): Oncerochara bolitocharoides gen. et sp. nov., of the tribe Homalotini (Coleoptera Staphylinidae: Aleocharinae) from Japan. Zootaxa 5748 (1): 113-119, DOI: 10.11646/zootaxa.5748.1.6, URL: https://doi.org/10.11646/zootaxa.5748.1.6
03AB5F65FFF6BA29FF457EE2FB9C7251.taxon	distribution	Distribution. Japan (Honshu).	en	Hashizume, Takuto, Maruyama, Munetoshi (2026): Oncerochara bolitocharoides gen. et sp. nov., of the tribe Homalotini (Coleoptera Staphylinidae: Aleocharinae) from Japan. Zootaxa 5748 (1): 113-119, DOI: 10.11646/zootaxa.5748.1.6, URL: https://doi.org/10.11646/zootaxa.5748.1.6
03AB5F65FFF6BA29FF457EE2FB9C7251.taxon	etymology	Etymology. This specific epithet is derived from the genus “ Bolitochara ” and the suffix “ - oides, ” which indicates that the habitus of new species resembles some species of Bolitochara.	en	Hashizume, Takuto, Maruyama, Munetoshi (2026): Oncerochara bolitocharoides gen. et sp. nov., of the tribe Homalotini (Coleoptera Staphylinidae: Aleocharinae) from Japan. Zootaxa 5748 (1): 113-119, DOI: 10.11646/zootaxa.5748.1.6, URL: https://doi.org/10.11646/zootaxa.5748.1.6
