taxonID	type	description	language	source
03AB87E2FFB5FFE9512E0EC78482B157.taxon	type_taxon	Type species Protoma knockeri Baird, 1870, by monotypy.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB5FFEE51460D108749B1E0.taxon	description	Fig. 2 A – D	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB5FFEE51460D108749B1E0.taxon	materials_examined	Material examined IRAN – Sistan and Baluchestan • 10 shells; Dar Pahn Unit;, GDZU- 0001 to GDZU- 0010.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB5FFEE51460D108749B1E0.taxon	discussion	Remarks Cox (1936) introduced Turritella harrisoni for specimens from the Pliocene ‘ Mekran beds’ near Bandar-e Jask on the western end of the Makran margin in southern Iran. He described the whorls as “ convex ”, and among the five primary spirals, he noted “ the three more posterior ones [are] weaker than the other two ”. This description matches only some of the specimens he illustrated, including the one he selected as holotype (Cox 1936: pl. 8 fig. 26). Two specimens (Cox 1936: pl. 8 figs 27, 29), however, show a nearly flat whorl profile and spirals of almost equal strength. These specimens are now assigned to Protoma kayalensis Dey, 1961 (see below). A specimen illustrated as Turritella angulata from the Talar Sandstone, between Garr Koh and Chatti at the Makran margin in southwestern Pakistan (Crame 1984) also belongs to P. harrisoni. The actual Turritella angulata is an Early Miocene species from Kutch in India, and is much more angular due to a prominent keel near the base of the whorls (see Sowerby 1840: pl. 26 fig. 7, or Kulkarni et al. 2010: fig. 2 g).	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB2FFEE512D0AB7849CB54B.taxon	type_taxon	Type species Neverita josephinia Risso, 1826, by monotypy.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB2FFEE51070C638706B676.taxon	description	Fig. 2 E – G	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB2FFEE51070C638706B676.taxon	materials_examined	Material examined IRAN – Sistan and Baluchestan • 10 shells; Dar Pahn Unit; GDZU- 0011 to GDZU- 0020.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB2FFEE51070C638706B676.taxon	discussion	Remarks Dey’s (1961) description and illustration match the specimens reported here from the Dar Pahn Unit, with the exception that these specimens attain a height of 88 mm, whereas the incomplete specimens from India measure only 32 mm. The species was subsequently reclassified to Protoma by Kulkarni et al. (2010).	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB2FFEF510409068318B2CD.taxon	description	Fig. 3 A – D	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB2FFEF510409068318B2CD.taxon	materials_examined	Material examined IRAN – Sistan and Baluchestan • 2 shells; Dar Pahn Unit; GDZU- 0065, GDZU- 0066.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB2FFEF510409068318B2CD.taxon	discussion	Remarks This extant species is found throughout the Indo-West Pacific Ocean, including the Persian Gulf (Niamaimandi et al. 2017) and has Neogene fossil occurrences ranging from India though Indonesia to Sakhalin Island (Majima 1989; Amano et al. 1996; Lyngdoh et al. 1999; Kase et al. 2008; Harzhauser et al. 2018). The species is often referred to as Glossaulax didymia, but Huelsken et al. (2012) questioned the distinction of Glossaulax Pilsbry, 1929, as a subgenus within Neverita based on molecular data.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB3FFEF512E0D7C853FB19C.taxon	type_taxon	Type species Nerita vitellus Linnaeus, 1758, by subsequent designation.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB3FFEF51060DCE873FB76C.taxon	description	Fig. 3 E – I	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB3FFEF51060DCE873FB76C.taxon	materials_examined	Material examined IRAN – Sistan and Baluchestan • 3 shells; Dar Pahn Unit; GDZU- 0067 to GDZU- 0069.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB3FFEF51060DCE873FB76C.taxon	discussion	Remarks The specimens from the Dar Pahn Unit are very similar to N. vitellus, but appear somewhat taller. The species [including its synonym Natica rufa (Born, 1778)] has Neogene fossil records in the central Indo-West Pacific region, as well as in India and Japan (Mukerjee 1939; Dey 1961; Kanno et al. 1982; Majima 1989).	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB0FFEC515F0E82875DB4BF.taxon	description	urn: lsid: zoobank. org: act: 1203 B 45 A- 1800 - 4 BA 8 - BA 8 F- 668 A 684 A 2 A 53 Fig. 4 A – J	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB0FFEC515F0E82875DB4BF.taxon	diagnosis	Diagnosis The shell is biconical, with six whorls and a strongly constricted base. A central row of spines is present, with 9 spines on the penultimate whorl and 12 spines on the last whorl. The aperture is lenticular to subcircular, with the inner lip featuring beaded spirals and little to no callus. The outer lip bears four strong spines of alternating strength: the sutural spine is long, the central spine is elongate-triangular and shorter, the spine at the basal margin is long and strong, and the spine at the base is short. The areas between the rows of spines have 3 to 6 beaded spiral cords, also alternating in strength.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB0FFEC515F0E82875DB4BF.taxon	etymology	Etymology For the town of Chabahar, located in the Makran region of southeastern Iran, where the specimens were collected.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB0FFEC515F0E82875DB4BF.taxon	materials_examined	Type material Holotype IRAN – Sistan and Baluchestan • Dar Pahn Unit; GDZU- 0082 Paratypes IRAN – Sistan and Baluchestan • 6 shells; Dar Pahn Unit; GDZU- 0083 to GDZU- 0088. Other material studied IRAN – Sistan and Baluchestan • 82 shells; Dar Pahn Unit; GDZU- 0089 to GDZU- 0170.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB0FFEC515F0E82875DB4BF.taxon	discussion	Remarks Bufonaria chabaharensis sp. nov. differs from many other species of Bufonaria by its strongly constricted base. Species with a similarly constricted base include B. crumena (Lamarck, 1816) from Neogene strata of Indonesia, and B. margaritula (Deshayes, 1832) from mostly Pleistocene strata in Indonesia, both of which have a lower spire and weaker spines (see Beu 2005: figs 1 – 5) than B. chabaharensis. Similarly long spines are known from Bufonaria echinata (Link, 1807), which today is common throughout the northern Indian Ocean, including the Red Sea and is rare in the northwestern Pacific Ocean up to southern Japan (Okutani 2000; Beu 2005). Other similar extant species include Bufonaria subgranosa (G. B. Sowerby II, 1836), also known from Neogene – Pleistocene strata in the central Indo-West Pacific (Tesch 1915), and Bufonaria rana (Linnaeus, 1758), both of which, however, have a less strongly constricted basal margin as Bufonaria chabaharensis (see Raven 2021).	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB0FFEC51660E3285DBB2C7.taxon	type_taxon	Type species Gyrineum echinatum Link, 1807, by subsequent designation.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB1FFED513E0E328523B2C7.taxon	type_taxon	Type species Tiara isabella Swainson, 1831, by subsequent designation.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB1FFE251450E8287B6B63E.taxon	description	urn: lsid: zoobank. org: act: D 783 C 058 - D 8 BF- 4492 - 86 A 4 - 66 FF 5 C 645 AA 3 Fig. 4 K – M	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB1FFE251450E8287B6B63E.taxon	diagnosis	Diagnosis A tall, slender, mitriform shell with at least six whorls, each featuring a finely incised suture. The whorls are slightly convex, sculptured by 6 to 7 broad, flat-topped spiral bands, separated by narrow grooves. In the lower third of the final whorl, the spirals become more rounded, and the interspaces widen. The aperture is narrow and elongated, occupying slightly more than half of the shell’s total height. The outer lip is thin, while the inner lip is lightly callused, with four columellar folds. The uppermost fold is the most prominent, with subsequent folds becoming progressively weaker and more oblique toward the base. A fasciole is present at the base of the siphonal column.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB1FFE251450E8287B6B63E.taxon	etymology	Etymology For the Makran margin, where the specimens were collected.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB1FFE251450E8287B6B63E.taxon	materials_examined	Type material Holotype IRAN – Sistan and Baluchestan • Dar Pahn Unit; GDZU- 0172. Paratype IRAN – Sistan and Baluchestan • 1 shell; Dar Pahn Unit; GDZU- 0173.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB1FFE251450E8287B6B63E.taxon	discussion	Remarks The image of holotype GDZU- 0172 figured by Hadi et al. (2024) was photographed at an oblique angle and appears broader than it actually is. Similar specimens were reported by Dey (1961) as Mitra (Tiara) aegra from the Miocene of western India. However, the extant Nebularia aegra (Reeve, 1845) (cf. Fedosov et al. 2018) has a much lower aperture and higher spire; hence, the Miocene Indian specimens do not belong to that extant species. These Indian specimens are here considered conspecific with Cancilla makranensis sp. nov. The most similar fossil species regarding shell shape is Mitra birmanica Vredenburg, 1923 (Vredenburg 1923: pl. 16 fig. 5), but it is sculptured with more numerous and finer spirals. Mitra tittabweensis Vredenburg, 1923 (Vredenburg 1923: pl. 16 fig. 8) is less slender and has a broader aperture. Another common Miocene mitrid from the northwestern Indian Ocean region is Scabricola sowerbyi (d’Orbigny, 1852) [as Mitra (Chrysame) sowerbyi (Vredenburg 1925; Mukerjee 1939; Dey 1961) and Nebularia sowerbyi (Harzhauser 2014: 122, pl. 8 figs 5 – 6)]. This species is currently assigned to Scabricola Swainson, 1840 (cf. Fedosov et al. 2018) and differs from Cancilla makranensis sp. nov. mainly by having a lower spire. Also similar is Mitra granatinaeformis Martin, 1884, from the Miocene of Java, but has only 4 – 5 spiral incisions per whorl, a taller last whorl, less constricted basal margin and longer siphonal rostrum. The specimen illustrated as Mitra granatinaeformis by Mukerjee (1939: 65, pl. 3 fig. 25) from eastern India has four spiral incisions like the specimens from Java, but a somewhat more constricted basal margin.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBEFFE2512A0AC3851DB550.taxon	type_taxon	Type species Arca antiquata Linnaeus, 1758, by original designation.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBEFFE3511509138256B5D0.taxon	description	Fig. 5	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBEFFE3511509138256B5D0.taxon	materials_examined	Material examined IRAN – Sistan and Baluchestan • 23 shells; Dar Pahn Unit; GDZU- 0314 to GDZU- 0337.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBEFFE3511509138256B5D0.taxon	discussion	Remarks The specimens reported from “ the beach off the Ormara Headland, facing the Mekran or Baluchistan coast, 130 miles west of Karachi ” (Newton 1905: 293) are smaller than those reported here (45 mm vs 71 mm in length), but his description of shell outline, inflation, and the number and shape of the ribs leaves no doubt that these specimens belong to the same species. Similar, but less inflated and smaller is Anadara garoensis Mukerjee, 1939, from the Miocene Garo Hills in the eastern Indian province of Meghalaya (Mukerjee 1939: 26, pl. 22 figs 2 – 3); it reaches only about a third of the size of A. blanfordi (25.8 mm vs up to 71 mm in length). Also similar is Arca yawensis Noetling, 1901, from the Miocene of Myanmar (Noetling 1901: 144, pl. 6 figs 4 – 6), which is nearly identical in outline and also very inflated. Noetling reported 29 radial ribs for this species, which falls within the range of the 28 – 31 present on A. blanfordi. However, the species can be distinguished by the difference in the size of the adductor muscle scars, described as “ the posterior one much larger than the anterior one ” for A. yawensis, whereas in A. blanfordi they are of subequal size (Fig. 5 A – D). The Miocene Anadara gourae Dey, 1961, from Kerala, India, is similar to A. blanfordi especially regarding its outline, but it has more numerous and narrower radial ribs (Dey 1961: pl. 1 figs 4 – 8). The most similar extant species is Anadara rufescens (Reeve, 1844), an uncommon species that is, however, widely distributed in the Indo-West Pacific Ocean (Mukerjee 1939; Dey 1961).	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBFFFE052B708558593B137.taxon	type_taxon	Type species Pecten vasselii Fuchs, 1878, by original designation.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBFFFE052B708558593B137.taxon	discussion	Remarks Bifurcating radial ribs as seen in Fascipecten do not occur among species of Pecten. Therefore, we elevate the subgenus Pecten (Fascipecten) to the level of genus. The type species F. vasselii occurs in Mio – Pliocene strata from South Africa via the Gulf of Suez to southern Pakistan (Fuchs 1878; Cox 1927; King 1954; Freneix et al. 1971; Crame 1984). The genus also includes F. shanganiensis Eames & Cox, 1958 originally described from the Late Miocene of East Africa (Eames & Cox 1956), and F. praevasseli Douglas 1928, from the Early Miocene of Iran (Douglas 1928). Thus, Fascipecten is currently restricted to the Neogene of the western Indian Ocean (Freneix et al. 1971).	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBCFFE152F30D3587B5B2A4.taxon	description	Fig. 6 A – M	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBCFFE152F30D3587B5B2A4.taxon	materials_examined	Material examined IRAN – Sistan and Baluchestan • 17 shells; Dar Pahn Unit; GDZU- 0243 to GDZU- 0259.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBCFFE152F30D3587B5B2A4.taxon	discussion	Remarks Identical specimens were described as Pecten fasciculatus from the Late Miocene of East Africa (Eames & Cox 1956). That name, however, was invalid, as no fewer than three species of the same name had been described earlier (Hinds 1844 – 1845; Sandberger 1863; Millet 1866) and it was thus replaced by Pecten shanganiensis.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBDFFE651120A45879AB256.taxon	description	Fig. 7 A – B	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBDFFE651120A45879AB256.taxon	materials_examined	Material examined IRAN – Sistan and Baluchestan • 1 shell; Dar Pahn Unit; GDZU- 0338.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBDFFE651120A45879AB256.taxon	discussion	Remarks This specimen agrees well with previous illustrations and description of D. peralta (Vredenburg 1928; Borkar & Kulkarni 2019), although with a height of 45 mm it is somewhat larger than the maximum height of 35 mm reported by Borkar & Kulkarni (2019). A very similar species is Dosinia boettgeri Martin, 1879 from the Late Miocene of Java (see Leloux & Wesselingh 2009: pl. 17 figs 6 – 8), from which D. peralta differs mainly by its more acute anterodorsal margin. Another similar species is Dosinia exasperata from Pleistocene strata in the Philippines (Shuto 1971), which has more irregular commarginal sculpture and the sculpture is scaly on the posterodorsal margin, unseen in D. peralta. Extant specimens of Dosinia exasperata are more roundish and have more steeply sloping antero- and posterodorsal margins.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBDFFE151310BF7849CB604.taxon	type_taxon	Type species Venus concentrica Born, 1778, by monotypy.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBDFFE151430DF28324B734.taxon	description	Fig. 6 N	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBDFFE151430DF28324B734.taxon	materials_examined	Material examined IRAN – Sistan and Baluchestan • 1 shell; Dar Pahn Unit; GDZU- 0260.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBDFFE151430DF28324B734.taxon	discussion	Remarks The single right valve from the Dar Pahn Unit has 18 – 19 narrow, regularly spaced and evenly sized radial ribs, virtually identical to those described for the left valves of Pecten kilindoniensis, from the Late Miocene of Mafia Island, Tanzania. It resembles that species also in size, outline, and auricle shape.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBDFFE151250EA4852BB177.taxon	type_taxon	Type species Ostrea maxima Linnaeus, 1758, by subsequent designation.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBAFFE651330E1585D8B2E0.taxon	type_taxon	Type species Mactra papyracea Gmelin, 1791, by subsequent designation.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBAFFE651610D6087CFB7B2.taxon	description	Fig. 7 C – E	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBAFFE651610D6087CFB7B2.taxon	materials_examined	Material examined IRAN – Sistan and Baluchestan • 6 shells; Dar Pahn Unit; GDZU- 0264 to GDZU- 0269.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBAFFE651610D6087CFB7B2.taxon	discussion	Remarks This is an extant species with a broad Indo-West Pacific distribution; fossil occurrences range back at least into the Early Miocene and are known across the Indo-West Pacific region (Sowerby 1840; Cossmann 1923; Vredenburg 1928; Beets 1950; Shuto 1969; Mahdi 2007; Ralte 2012), including Japan (Ogasawara 1976).	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBAFFE751690A0F85B4B2DD.taxon	description	Fig. 7 F – I	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBAFFE751690A0F85B4B2DD.taxon	materials_examined	Material examined IRAN – Sistan and Baluchestan • 4 shells; Dar Pahn Unit; GDZU- 0277 to GDZU- 0280.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBAFFE751690A0F85B4B2DD.taxon	discussion	Remarks Tapes protolirata Noetling, 1901 from the Miocene of Myanmar was described as follows: “ ornamentation consists of strong concentric ribs, very regular, flatly rounded, and cover the entire shell uniformly ”. Although the outer shell surface of the available specimens is either obscured by sediment or partially corroded, one specimen (Fig. 7 G) shows indications of concentric sculpture. The drawings provided by Noetling (1901: 212, pl. 13 figs 12 – 13) show specimens with a somewhat straighter dorsal margin than the Iranian specimens, hence they are here only tentatively assigned to Paratapes protolirata. Pullastra? virgata (Sowerby 1840: pl. 25 fig. 9) from the Miocene of Kutch, India, is less rectangular than typical Venerupis Lamarck, 1818 (= Pullastra Sowerby, 1826) and somewhat resembles Paratapes. However, that species has never been revised and was only briefly mentioned in the context of an Eocene species of Tapes Megerle von Mühlfeld, 1811 from northern Italy (Mayer-Eymar 1863). Popenoe & Kleinpell (1978) illustrated a similar specimen as Paphia euglypta (Philippi, 1847) from the Pliocene Vigo Formation in the Philippines. The three accepted extant species of Paratapes, differ from each other only by minute differences, but they seem to have a more gently rounded dorsal margin than Paratapes protolirata (i. e., Thach 2024).	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBAFFE651080BB185CCB643.taxon	type_taxon	Type species Venus textile (Gmelin, 1791), by typification of replaced name.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBBFFE751630AE082D9B496.taxon	description	Fig. 7 K – L	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBBFFE751630AE082D9B496.taxon	materials_examined	Material examined IRAN – Sistan and Baluchestan • 2 shells; Dar Pahn Unit; GDZU- 0339, GDZU- 0340.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBBFFE751630AE082D9B496.taxon	discussion	Remarks This is a quite characteristic extant species, originally reported from the Red Sea (Gmelin 1791; Lamy 1918), but widely distributed in the Indo-West Pacific (Glover et al. 2016). The potentially oldest fossil record (at least of the genus) is from the Early Oligocene of India (Kachhara et al. 2012). Further fossil records of A. inflata are from the Neogene of the Philippines (Shuto 1971; Kanno et al. 1982), the Pliocene of Vanuatu (Abrard 1947), and the Pleistocene of Somalia (Abrard 1941).	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBBFFE751720B918512B661.taxon	type_taxon	Type species Tellina inflata (Gmelin, 1791), by original designation.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFBBFFE751130D4C8278B1AF.taxon	type_taxon	Type species Solen strigilatus Linnaeus, 1758, by subsequent designation (Anton 1838).	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB8FFE451010E3B826DB2D1.taxon	type_taxon	Type species Corbula sulcata Lamarck, 1801, by subsequent designation (Schmidt 1818).	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB8FFFA514E0E918316B2E5.taxon	description	urn: lsid: zoobank. org: act: F 22549 C 7 - 616 F- 4739 - B 4 D 4 - F 2141711 A 3 EB Fig. 7 M – S	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB8FFFA514E0E918316B2E5.taxon	diagnosis	Diagnosis The shell is elongate-oval, moderately inflated, with low, uncurved umbones. The posterior margin is acute and truncated, featuring a blunt ridge. The ventral margin exhibits a distinct angulation or curvature just posterior to the midline. The sculpture consists of fine or rough growth increments, less distinct on the left valve. There is no lunule, and the escutcheon is indistinct.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB8FFFA514E0E918316B2E5.taxon	etymology	Etymology For the Dar Pahn Unit.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB8FFFA514E0E918316B2E5.taxon	materials_examined	Type material Holotype IRAN – Sistan and Baluchestan • Dar Pahn Unit; GDZU- 0289. Paratypes IRAN – Sistan and Baluchestan • 3 shells; Dar Pahn Unit; GDZU- 0290, GDZU- 0291, GDZU- 0293. Other material examined IRAN – Sistan and Baluchestan • 20 shells; Dar Pahn Unit; GDZU- 0294 to GDZU- 0313.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
03AB87E2FFB8FFFA514E0E918316B2E5.taxon	discussion	Remarks Corbula darpahnensis sp. nov. differs from the Late Miocene “ Corbula harpa d’Arch. ” from Myanmar (Noetling 1901) by being elongate rather than triangular; the original C. harpa is from the Early Eocene Ranikot Group in Pakistan (d’Archiac & Haime 1853) and is also triangular in outline. Two similarly elongate species are the Miocene Corbula tunicosulcata Vredenburg, 1928, and the Pliocene Corbula mekranica Vredenburg, 1928, both of which have more regular concentric ornament and a more acute posterior margin (Vredenburg 1928). Also more triangular is the accordingly named Corbula trigonalis Sowerby, 1840 from the Miocene of India (Sowerby 1840: pl. 25 fig. 4), which also has a much sharper posterior ridge (see Halder & Bano 2015: fig. 5). Sowerby also reported C. rugosa Lamarck, 1806 from the Miocene of India (Sowerby 1840: pl. 25 fig. 5), but that specimen was later included in Corbula tunicosulcata (Vredenburg 1928). Similar regarding shell outline and the low umbo is Corbula socialis Martin, 1879, from the Miocene of Java, but that species typically has distinct, blunt commarginal ribs, unlike the specimens from the Dar Pahn Unit. Two further species with a similar outline but distinct commarginal ribs are C. scaphoides Hinds, 1843 and C. laterugata Cossmann, 1923 from the Pliocene of southeastern India (Cossmann 1923). Also similar is Corbula njalindungensis Martin, 1922, from the Middle Miocene of Java. It is rather variable and includes relatively smooth forms and some that are of only moderate inflation (see Leloux & Wesselingh 2009: pls 28 – 30), but typically has a much more distinct and elevated umbo.	en	Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng (2026): New species and records of Late Miocene mollusks from southeastern Iran. European Journal of Taxonomy 1074: 1-22, DOI: 10.5852/ejt.2026.1074.3300, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3300/14534
