taxonID	type	description	language	source
03AA879E840B3E2262B0F8A0FE65E6B3.taxon	diagnosis	Diagnosis. Small to medium size wasps (1 – 4 mm) with mesoscutum and metasomal terga strongly arched and with apex of metasoma curved down in lateral view [Cenozoic species (Fig. 1 B)] to moderately arched with apex of metasoma directed backwards in lateral view [Mesozoic species (Fig. 2 A, B)]; head transverse and orthognathous; antennal shelf indistinct (Fig. 3 B) to slightly developed (Fig. 3 A, C); antenna 15 - merous in female, 14 - merous in male; males with modified A 3 and A 4 (sex-segments), or only one of them, or both not modified; third maxillary palpomere cylindrical, not enlarged and not broader than the following palpomere (Fig. 3 B – D); notauli completely developed (Fig. 2 C) or reduced to anterior pits (Fig. 1 A); fore wing without stigma but with slightly thickened marginal vein; costal vein complete, sclerotized and tubular (Fig. 1 D) to totally reduced (Figs 1 C, 2 A, B); metasoma distinctly petiolate with second metasomal segment the longest; connection of second metasomal segment (M 2) with petiole more or less monotypic: base of M 2 shorter than petiole in lateral view (Figs 1 B, 4 D) and as wide as to distinctly wider than petiole in dorsal view, with more or less distinct lateral corners (Figs 2 C, 4 B); terga slightly over crossing sterna at sides, at least S 2 with distinct and complete belytoid line; ovipositor entirely retracted to slightly extracted backwards (Figs 1 B, 4 B and D); males, and most females, with one-segmented exposed cerci on T 8 (Figs 1 B, 2 C and D, 3 B).	en	Chemyreva, Vasilisa G., Rasnitsyn, Alexandr P. (2025): A new ismarid wasp (Hymenoptera, Diaprioidea) from mid-Cretaceous Kachin amber. Zootaxa 5737 (1): 96-104, DOI: 10.11646/zootaxa.5737.1.4, URL: https://doi.org/10.11646/zootaxa.5737.1.4
03AA879E84083E2262B0FDEAFD55E453.taxon	type_taxon	Type species Thyburia christophi sp. nov.; by present designation.	en	Chemyreva, Vasilisa G., Rasnitsyn, Alexandr P. (2025): A new ismarid wasp (Hymenoptera, Diaprioidea) from mid-Cretaceous Kachin amber. Zootaxa 5737 (1): 96-104, DOI: 10.11646/zootaxa.5737.1.4, URL: https://doi.org/10.11646/zootaxa.5737.1.4
03AA879E84083E2262B0FDEAFD55E453.taxon	etymology	Etymology. The name of this genus is an arbitrary euphonious combination of letters, with no special meaning.	en	Chemyreva, Vasilisa G., Rasnitsyn, Alexandr P. (2025): A new ismarid wasp (Hymenoptera, Diaprioidea) from mid-Cretaceous Kachin amber. Zootaxa 5737 (1): 96-104, DOI: 10.11646/zootaxa.5737.1.4, URL: https://doi.org/10.11646/zootaxa.5737.1.4
03AA879E84083E2262B0FDEAFD55E453.taxon	diagnosis	Genus diagnosis. As for the type species below.	en	Chemyreva, Vasilisa G., Rasnitsyn, Alexandr P. (2025): A new ismarid wasp (Hymenoptera, Diaprioidea) from mid-Cretaceous Kachin amber. Zootaxa 5737 (1): 96-104, DOI: 10.11646/zootaxa.5737.1.4, URL: https://doi.org/10.11646/zootaxa.5737.1.4
03AA879E84083E2562B0FCF7FB28E58B.taxon	description	Figs 2 – 4	en	Chemyreva, Vasilisa G., Rasnitsyn, Alexandr P. (2025): A new ismarid wasp (Hymenoptera, Diaprioidea) from mid-Cretaceous Kachin amber. Zootaxa 5737 (1): 96-104, DOI: 10.11646/zootaxa.5737.1.4, URL: https://doi.org/10.11646/zootaxa.5737.1.4
03AA879E84083E2562B0FCF7FB28E58B.taxon	etymology	Etymology. The species is named after Dr Christoph Öhm-Kühnle (Germany), who kindly presented the amber with the specimen to the PIN collection.	en	Chemyreva, Vasilisa G., Rasnitsyn, Alexandr P. (2025): A new ismarid wasp (Hymenoptera, Diaprioidea) from mid-Cretaceous Kachin amber. Zootaxa 5737 (1): 96-104, DOI: 10.11646/zootaxa.5737.1.4, URL: https://doi.org/10.11646/zootaxa.5737.1.4
03AA879E84083E2562B0FCF7FB28E58B.taxon	materials_examined	Material studied. Holotype ♀, PIN 5620 / 125 a, a complete specimen in transparent amber housed in the collection of the A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow (PIN). Syninclusion: a single parasyninclusion – ant-like stone beetles Scydmaeninae (Coleoptera, Staphylinidae). Type locality: Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar. Type horizon: Late Albian to early Cenomanian (mid-Cretaceous).	en	Chemyreva, Vasilisa G., Rasnitsyn, Alexandr P. (2025): A new ismarid wasp (Hymenoptera, Diaprioidea) from mid-Cretaceous Kachin amber. Zootaxa 5737 (1): 96-104, DOI: 10.11646/zootaxa.5737.1.4, URL: https://doi.org/10.11646/zootaxa.5737.1.4
03AA879E84083E2562B0FCF7FB28E58B.taxon	diagnosis	Diagnosis. Small wasps (about 1.2 mm); antennae with (?) annellus between A 2 and A 3; clypeus transverse and strongly convex (Fig. 3 A, C, D), ventrally with distinct semicircular clypeal flange projecting forwards; tentorial pits present, small; malar sulcus absent; occipital carinae narrow and smooth; notauli complete, wide and smooth at bottom; all claws simple, without row of strong setae ventrally; fore tibia with one regular spur and one long setae next to it (Fig. 3 A); anterior scutellar pit large, circular and smooth at the bottom (Fig. 2 C); mesopleural pit indistinct; mesopleuron with setose epicnemial pit, distinct epicnemial and subalar bridges (Fig. 3 A); fore wing without costal vein, marginal vein distinctly shorter than radial cell and parastigma; radial and postmarginal veins nebulous in distal 2 / 3; radial vein straight (Fig. 2 A, B); petiole slightly transverse, carinate and setose dorsally; base of T 2 deeply depressed next to petiole, arched posteriorly and with large lateral corners (Figs 2 C, 4 B, D); base of T 2 and S 2 densely setose (Figs 3 A, 4 D).	en	Chemyreva, Vasilisa G., Rasnitsyn, Alexandr P. (2025): A new ismarid wasp (Hymenoptera, Diaprioidea) from mid-Cretaceous Kachin amber. Zootaxa 5737 (1): 96-104, DOI: 10.11646/zootaxa.5737.1.4, URL: https://doi.org/10.11646/zootaxa.5737.1.4
03AA879E84083E2562B0FCF7FB28E58B.taxon	description	Description. Female (holotype). Body length 1.2 mm; antennae about 0.85 mm; wings 1.0 mm. Head. Head smooth and setose throughout, in dorsal view about 1.4 times as wide as long and about as wide as mesosoma; POL: 18; LOL: 10; OOL: 23; ocelli large and convex, LOL 0.6 times as long as largest diameter of lateral ocellus; eye large and bare; distance between toruli about as long as diameter of one torulus; toruli slightly turned to outer sides of antennal shelf (situated not in one plane on it); malar distance short and without malar sulcus; face with strongly convex supraclypeal area above clypeus; clypeus strongly convex, transverse, covered with few, long upstanding setae; epistomal sulcus distinct; tentorial pit distinct; clypeal flange semicircular, bare; pleurostomal distance slightly narrower than shortest distance between eyes; mandibles elongate, narrowed to apex, bidentate. Antenna. Moniliform, stout; A 1 cylindrical, about 4 times as long as wide, almost smooth and covered with scattered setae; A 2 with (?) annellus; A 3 shorter than A 2 and A 4, A 5 – A 14 similar in shape and gradually increase in size and about as long as wide; A 15 longest antennomere, teardrop-shaped; A 3 – A 15 covered with faint surface sculpture and homogeneous setae. Mesosoma about as wide as high; in dorsal view about 1.7 times as long as wide. Pronotal shoulders not angled and not convex in dorsal view; pronotal collar almost vertical, setose and smooth between pronotal shoulders; epomia absent; lateral side of pronotum depressed medially and with sharp carina from pronotal shoulders to ventral lobe of pronotum (Fig. 3 A); fore mesosomal spiracle located under posterior pronotal margin, pronotum extending to tegula; mesoscutum convex, 1.7 times as wide as long, with notauli deep and wide, complete and smooth at bottom; narrow humeral sulcus developed along whole lateral margin of mesosocutum; tegula small, not covering sclerites at base of wings completely (Fig. 2 C); scutellum convex, entirely setose and sculptured along posterior margin, as long as mesoscutum; anterior scutellar pit large, subcircular but slightly transverse, smooth at bottom; axillar depression densely pubescent, mesoscutellar arm smooth; mesopleuron with setose epicnemial depression, distinct mesopleural pit, epicnemial and subalar bridges; mesopleuron anteriorly with visible anterior mesopleural fovea and subalar impression, and posteriorly with weak trace of mesepimeral sulcus (Fig. 3 A); metanotum long, in dorsal view only slightly shorter than propodeum (Fig. 2 C); metascutellum flat, bounded on sides by two carinae convergent posteriorly; metapleuron and lateral side of propodeum with irregular carinae; propodeum with plicae and blade-like median carina, which are interrupted anteriorly with transverse carina and groove in front of it; posterior margin of propodeum simple, without projections (Figs 2 C, 3 A); nuchal area bare and longitudinally carinate. All legs slender; each trochanter elongate, about 0.5 times as long as femur, each trochantellus very short (Fig. 3 A); hind tibiae slightly incrassate, covered with homogeneous thin setae on its outer side and with thick bristles on its inner side (Fig. 4 B). Metasoma. Petiole cylindrical and short, slightly transverse, covered with longitudinal carinae and entirely pubescent; in lateral view gaster not arched and T 2 slightly convex, significantly wider than petiole in dorsal view; base of T 2 and S 2 densely setose, both with short depressions and sculptured medially, forming smooth lateral corners; T 3 – T 6 short, smooth and scarcely setose; T 7 slightly shorter than combined length of T 5 and T 6; T 8 subtriangular in dorsal view, with one-segmented exposed cerci (Fig. 2 C); all sterna setose, S 3 – S 5 short, S 6 distinctly longer than combined length of S 3 – S 5; ovipositor slightly longer than S 6 with stout and weakly exposed posteriorly sheath (Fig. 4 B, D).	en	Chemyreva, Vasilisa G., Rasnitsyn, Alexandr P. (2025): A new ismarid wasp (Hymenoptera, Diaprioidea) from mid-Cretaceous Kachin amber. Zootaxa 5737 (1): 96-104, DOI: 10.11646/zootaxa.5737.1.4, URL: https://doi.org/10.11646/zootaxa.5737.1.4
03AA879E84083E2562B0FCF7FB28E58B.taxon	discussion	Remarks. This new species shares many characteristics with other members of the family Diapriidae, particularly the subfamily Belytinae, including the most plesiomorphic species. For example, the new species has a connection of the petiole with the second metasomal segment (Figs 2 C, 4 B, D), which is similar to that of the probably basal Belytinae (for example Eoaclista Brazidec) and species of Ambositra Masner (Ambositrinae) (Chemyreva et al. 2024 b; Brazidec 2025). It most likely has an annellus similar to those of some Belytinae and Ambositrinae species, as well as a belytoid line similar to those of all Belytinae and probably all Ambositrinae. Finally, it has one-segmented, exposed cerci like the majority of Ambositrinae. Compared to most Cenozoic Ismarinae, T. christophi sp. nov. has several reductions: C (costal vein) is absent; the pronotal collar is without a transverse carina; and the mesepimeral sulcus is almost smooth. However, these reductions are also known for some Cenozoic Ismarinae and / or Belytinae. The abundant pubescence of T. christophi sp. nov. (particularly on the dorsal side of the petiole and at the base of T 2 and S 2) and the slightly exposed ovipositor make it special within the known Ismarinae.	en	Chemyreva, Vasilisa G., Rasnitsyn, Alexandr P. (2025): A new ismarid wasp (Hymenoptera, Diaprioidea) from mid-Cretaceous Kachin amber. Zootaxa 5737 (1): 96-104, DOI: 10.11646/zootaxa.5737.1.4, URL: https://doi.org/10.11646/zootaxa.5737.1.4
