identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03AE87BC0B138C05FF428229FAB2F917.text	03AE87BC0B138C05FF428229FAB2F917.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Weitschatidae Koteja 2008	<div><p>Family Weitschatidae Koteja, 2008</p><p>Amended diagnosis (macropterous male). Antennae each 10-segmented, flagellar segments cylindrical and irregularly binodose, without capitate setae. Head with distinct lateral compound eyes; midcranial ridge well developed on dorsal surface and fused with postoccipital ridge, forming a reversed T-shaped structure. Scutum without a membranous area; membranous lateral areas of scutellum absent; basisternum hexagonal. Forewings hyaline, broad at base, with a narrow alar fold for holding hamuli; subcostal ridge with distinct club-shaped pterostigma; cubital ridge extending beyond middle of wing; both flexing patches present, but weak; radial sector absent. Tarsus two segmented, proximal segment very short and triangular, tarsal digitules undifferentiated; claw with two spine-like digitules not reaching claw apex. Tail tufts distributed on posterior margin of abdominal tergites VI and VII. Penial sheath short and triangular or with aedeagus forming a beak-like structure.</p></div>	https://treatment.plazi.org/id/03AE87BC0B138C05FF428229FAB2F917	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lin, Shan;Yao, Yunzhi;Ren, Dong	Lin, Shan, Yao, Yunzhi, Ren, Dong (2018): A new scale insect of the extinct family Weitschatidae (Insecta: Hemiptera: Coccomorpha) from mid-Cretaceous Burmese amber. Zootaxa 4407 (3): 427-434, DOI: 10.11646/zootaxa.4407.3.9
03AE87BC0B138C00FF42801AFB17FBA9.text	03AE87BC0B138C00FF42801AFB17FBA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Criniverticillus Lin, Yao & Ren 2018	<div><p>Genus Criniverticillus Lin, Yao &amp; Ren gen. nov.</p><p>http://zoobank.org/urn:lsid:zoobank.org:act:A5025B96-B712-418B-958C-FB3A91475B51</p><p>Type species. Criniverticillus longicumulus Lin, Yao &amp; Ren sp. nov.</p><p>Diagnosis (macropterous male). Body large (ca. 2 mm long, Fig. 1); antenna long (ca. 2.1–2.4 mm, Fig. 2F), most antennal setae forming distinct whorls, antennal apical segment with curved bristles (Fig. 2G); hamulohaltere leaf-shaped; basisternum without a median ridge (Fig. 2B); abdominal tergites well developed and sclerotized, first three tergites each with distinct median ridge (Fig. 2A); abdominal tergites VI and VII each with a transverse row of 7–9 tubular ducts (Fig. 2A) extruding wax filaments; a pair of short tubercular projections present on apex of abdominal segment VIII (Fig. 2 C).</p><p>Etymology. The generic name Criniverticillus comes from combination of the prefix ‘ crini- ’ (from the Latin crinis meaning ‘hair’) with the masculine Latin noun ‘ verticillus ’ (meaning ‘whorl’), referring to most of the setae on the antenna forming whorls around the segments.</p><p>Remarks. Koteja (2008) considered that the following extinct genera were xylococcid-like taxa: Arnoldus Koteja; Grohnus Koteja; Lithuanicoccus Koteja; Serafinus Koteja, and Weitschatus Koteja, and classified them in five families: Arnoldidae, Grohnidae, Lithuanicoccidae, Serafinidae and Weitschatidae respectively. After comparing key structures like antennal capitate setae; pterostigma on forewing; radial sector on forewing; hamulohaltere; basisternal median ridge; small caudal extension on tergite VIII; tarsal digitule; penial sheath; anterior margin of scutellum, and membranous lateral areas of scutellum, Vea &amp; Grimaldi (2015) questioned this classification and considered that further analyses of relationships within the xylococcid-like taxa were needed. According to our observations, based on the body and wing shapes, antennae without specialised setae, subtriangular scutellum and waxy tail tufts of our adult male Criniverticillus, we think these genera are very similar to each other, and are inclined to support the latter authors’ point of view.</p><p>Among the Xylococcid-like genera, records from the Cretaceous only comprise Xiphos Vea &amp; Grimaldi (family placement uncertain) from the Early Cretaceous Lebanese amber and Pseudoweitschatus Vea &amp; Grimaldi from the mid-Cretaceous Burmese amber; all other fossil records are from the Eocene Baltic amber. However, the morphological characters of Xiphos are very different from the other genera, such as: body small (ca. 1.45 mm long total); scutum with an oval membranous area; forewings each without anterior flexing patch; claw digitule absent, and penial sheath extremely elongate. The Xylococcid-like taxa therefore only have one important record ( Pseudoweitschatus) from the Cretaceous. The new species described in this paper adds to the record of the Weitschatidae in the mid-Cretaceous and provides a significant new addition to the understanding the past diversity of scale insects.</p><p>The new genus described herein is assigned to the family Weitschatidae based on the presence of the following features (contrasting conditions in the other fossil taxa/families are given in brackets): antennae without capitate setae (capitate setae present in Arnoldus ( Arnoldidae)); radial sector of forewing absent (present in Xylococcus ( Xylococcidae)); tarsal digitules undifferentiated (clavate in Arnoldus and Xiphos); membranous lateral areas of scutellum absent (present in Grohnus (Grohnidae) and Xiphos) and cubital ridge extending beyond middle of wing (cubital ridge very short, reaching posterior wing margin at 1/4 its length in Serafinus ( Serafinidae)). In particular, the shape of the pterostigma in Criniverticillus is very similar to that of Weitschatus and Pseudoweitschatus (Weitschatidae) (absent in other Xylococcid-like taxa).</p><p>Criniverticillus differs from the other two genera in Weitschatidae by the following combination of characters (contrasting conditions in the other taxa are given in brackets): antenna significantly longer, 2.10–2.40 mm in Criniverticillus (1.30–1.33 mm in Pseudoweitschatus); most setae on antenna forming distinct whorls (without whorls in Weitschatus); antennal apical segment with the short ‘curved bristles’ of Vea &amp; Grimaldi (2015) (absent in Weitschatus); hamulohaltere leaf-shaped (bilobed in Weitschatus); first three abdominal tergites with distinct median ridge (median ridge absent in Weitschatus and Pseudoweitschatus); a single row of tubular ducts present on each of abdominal tergites VI and VII (2–4 rows of pores in Weitschatus), and the presence of a pair of short tubercular projections on the apex of abdominal segment VIII (absent in Pseudoweitschatus).</p></div>	https://treatment.plazi.org/id/03AE87BC0B138C00FF42801AFB17FBA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lin, Shan;Yao, Yunzhi;Ren, Dong	Lin, Shan, Yao, Yunzhi, Ren, Dong (2018): A new scale insect of the extinct family Weitschatidae (Insecta: Hemiptera: Coccomorpha) from mid-Cretaceous Burmese amber. Zootaxa 4407 (3): 427-434, DOI: 10.11646/zootaxa.4407.3.9
03AE87BC0B168C01FF42837AFBA6FA19.text	03AE87BC0B168C01FF42837AFBA6FA19.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Criniverticillus longicumulus Lin & Yao & Ren 2018	<div><p>Criniverticillus longicumulus Lin, Yao &amp; Ren sp. nov.</p><p>(Fig 1 and 2)</p><p>http://zoobank.org/urn:lsid:zoobank.org:act:5858E5C9-5433-4C97-9DB7-0E861B6DB24C</p><p>Type material. Holotype: CNU-COC-MA2018001 ( Key Lab of Insect Evolution and Environmental Changes, College of Life Sciences, Capital Normal University, Beijing, China), macropterous male in a 7 × 7 × 2 mm block of yellow, transparent, polished amber; specimen in good condition, with complete body and almost unbroken antenna, both wings well spread, tubular wax projections visible; accessible views: dorsal and ventral.</p><p>Diagnosis. Setae on flagellar segments of antenna long, longest 3–4 times as long as corresponding segment width (Fig. 2F); penial sheath triangular and short.</p><p>Description. Body large and robust (Fig 1A and 1B), 1960 μm long, with greatest width at mesothorax, 560 μm.</p><p>Head (Fig. 2A) transversely oval, dorsoventrally flattened, 440 μm wide, 255 μm long. Compound eyes hemispherical, bulging laterally, each about 175 μm in diameter and with more than 100 ommatidia. Ocelli located dorsad to compound eyes. Midcranial ridge well-developed on dorsal surface and fused with a bent and strongly sclerotized postoccipital ridge, forming a reversed T-shaped structure. Ventral surface of head (Fig. 2B) with ridges forming an irregularly five-armed star, consisting of midcranial, preocular and preoral ridges; no setae observed.</p><p>Antenna (Fig. 2F) 10 segmented and especially long, longer than body length, total length 2360–2400 μm; flagellar segments III to X all filiform, most segments irregularly binodose. Lengths of segments (in μm): scape 85–90; pedicel 70–75; III 185–190; IV 260–265; V 280–290; VI 255–260; VII 290–300; VIII 275–280; IX 285– 300; X 305. Ratio of maximal width to minimal width of segments: scape 3.08; pedicel 1.33; III 2.0; IV 2.0; V 3.7; VI 2.0; VII 2.4; VIII 2.4; IX 3.3; X 3.3. All flagellar segments covered with long setae (each 170–240 μm long), most of them forming whorls each of 5–15 setae, with a few shorter setae (each 110–150 μm long) present mainly between whorls. Apical segment with short ‘curved bristles’ (Fig. 2G) and 4 very short basiconic sensilla (Fig. 2H). Capitate setae absent.</p><p>Thorax separated from head by a weak neck constriction. Prothorax slightly sclerotized on part of edge and center of dorsal surface, with some small setae on dorsum and a slender medial ridge on venter. Structures of mesothorax well developed and strongly sclerotized. Dorsally (Fig. 2A): prescutum transversely oval (ca. 305 μm wide, 195 μm long); scutum solid without any membranous area (410 μm at its longest and 560 μm at its widest); scutellum subtriangular (ca. 230 μm at its longest, 400 μm at its widest), anterior angle almost a right angle; mesopostnotum short (400 μm wide and 90 μm long) and only slightly overlapped by metanotum (410 μm wide and 190 μm long). Ventrally (Fig. 2B): basisternum hexagonal, 430 μm long, 410 μm at its widest, with a complete margin, without a longitudinal medial ridge. Metathorax represented by a large transverse tergite, but sternite indistinct.</p><p>Forewings (Fig. 2I) each oval, broad at base, with slightly tapering apex, 1950 μm long and 960 μm at its widest; subcostal ridge extending from base of wing to more than 4/5 wing length, forming apically a well-defined club-shaped pterostigma (1 640 μm long total); cubital ridge 1150 μm long, starting about 395 μm from base of wing and fading before reaching pterostigma apex level; angle between subcostal ridge and cubital ridge approximately 45°; anterior flexing patch 1100 μm long, inconspicuous but appearing at same location as cubital ridge and separating progressively, fading before reaching pterostigma apex level; posterior flexing patch present but very weak, starting point invisible. Alar setae and microtrichia absent; sensoria present as a row along subcostal ridge. Alar fold and fine sculpturing present. Hamulohalteres large, leaf shaped (Fig. 2J), each about 190 μm long, 80 μm wide, with 4 hamuli attached on anterodistal margin.</p><p>Legs (Fig. 2 D) slender, subequal in length; fore-coxae situated remote from mid-coxae at a distance of about 440 μm; mid- and hind-coxae situated near to each other, about 185 μm apart; dimensions of right mid-leg: coxa, 170 μm long, 70 μm wide; trochanter + femur 600 μm long, 75 μm wide, with some short and hair-like setae; tibia, 540 μm long, 45 μm wide, with short spine-like setae and some setae that seem to be bifurcate (Fig. 2E), each about 25 μm long; tibial spurs not differentiated; tarsus 2-segmented; proximal segment very short and triangular, 200 μm long, 25 μm wide, with spine-like setae and some setae that seem to be bifurcate (Fig. 2E), each about 20 μm long; tarsal digitules undifferentiated; claws each ca. 60 μm long, 20 μm wide at base, strong and slightly bent, with 2 spine-like digitules (Fig. 2E), each 10–15 μm long, not reaching claw apex; claw without a denticle.</p><p>Abdomen (Fig. 2A and 2B) cylindrical, about 840 μm long (815 μm without penial sheath), maximum width 375 μm. Tergites and sternites well developed and segmentation clear, with a pair of slightly sclerotized tubercular projections on apex of abdominal segment VIII (Fig. 2C). Each tergite with one row of small pores on posterior margin, and anterior three tergites with distinct median ridge. Tergites VI and VII each with 1 row of 9 tubular ducts (Fig. 2A) extruding wax filaments (each filament about 3500 μm long).</p><p>Penial sheath (Fig. 2B) narrow and triangular, 220 μm long, 125 μm wide at base, tapering posteriorly; without setae. Aedeagus obscured. Eversible endophallus visible.</p><p>Etymology. The species epithet longicumulus is a noun in apposition. It comes from combination of the Latin prefix ‘ longi- ’ (from the Latin longus meaning ‘long’) and the masculine Latin noun ‘ cumulus ’ (meaning ‘accumulation’), referring to the long and densely crowded setae on the antenna.</p></div>	https://treatment.plazi.org/id/03AE87BC0B168C01FF42837AFBA6FA19	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lin, Shan;Yao, Yunzhi;Ren, Dong	Lin, Shan, Yao, Yunzhi, Ren, Dong (2018): A new scale insect of the extinct family Weitschatidae (Insecta: Hemiptera: Coccomorpha) from mid-Cretaceous Burmese amber. Zootaxa 4407 (3): 427-434, DOI: 10.11646/zootaxa.4407.3.9
03AE87BC0B178C01FF4282EDFA27F82B.text	03AE87BC0B178C01FF4282EDFA27F82B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Weitschatidae Koteja 2008	<div><p>Key to species of Weitschatidae based on adult macropterous males</p><p>1 Antenna with some setae forming distinct whorls and apical segment with curved bristles. Hamulohalteres leaf-shaped. Abdominal tergites VI and VII with tubular ducts in only 1 row................................................ 2</p><p>- Antenna with setae not forming whorls and apical segment without curved bristles. Hamulohalteres bilobate apically. Abdominal tergites VI and VII with tubular ducts in 2 to 4 rows..................................................... 3</p><p>2 Antenna significantly longer, 2.10–2.40 mm; Apex of abdomen with a pair of short tubercular projections................................................................................ Criniverticillus longicumulus gen. &amp; sp. nov.</p><p>- Antenna shorter, 1.30–1.33 mm; Apex of abdomen without a pair of short tubercular projections................................................................................. Pseudoweitschatus audebertis Vea &amp; Grimaldi, 2015</p><p>3 Flagellar setae long, length of longest seta 2–3 times width of segment and inserted almost perpendicular to longitudinal axis of segment. Basisternum without medial ridge................................... Weitschatus stigmatus Koteja, 2008</p><p>- Flagellar setae short, length of longest as long as width of segment and strongly bent at base, adhering to the segments. Basisternum with longitudinal medial ridge at latter part............................ Weitschatus vysniauskasi Koteja, 2008</p></div>	https://treatment.plazi.org/id/03AE87BC0B178C01FF4282EDFA27F82B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lin, Shan;Yao, Yunzhi;Ren, Dong	Lin, Shan, Yao, Yunzhi, Ren, Dong (2018): A new scale insect of the extinct family Weitschatidae (Insecta: Hemiptera: Coccomorpha) from mid-Cretaceous Burmese amber. Zootaxa 4407 (3): 427-434, DOI: 10.11646/zootaxa.4407.3.9
