identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A187D23400CF2DFF4EFF217930FD4F.text	03A187D23400CF2DFF4EFF217930FD4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wyeomyia (Leonidasdeanea) Motta, Sallum & Lourenço-de-Oliveira 2026	<div><p>Wyeomyia subgenus Leonidasdeanea Motta, Sallum &amp; Lourenço-de-Oliveira, subgen. nov.</p><p>Type species. Prosopolepis flui Bonne-Wepster &amp; Bonne, 1920 .</p><p>Wyeomyia in part of Dyar &amp; Knab 1906a: 138–140 ( chalcocephala only; chalcocephala description).</p><p>Heliconiamyia in part of Dyar 1919: 115, 123, pl. 5 (new genus, includes galoa male and chalcocephala, genus defined by male genitalia characters, galoa Dyar &amp; Knab, 1906a selected as type species); Edwards 1932: 86 (synonymy with Dendromyia Theobald, 1903).</p><p>Prosopolepis of Bonne-Wepster &amp; Bonne 1920: 169, 170 ( flui description); Dyar &amp; Ludlow 1921: 677, 678 (in part, flui only); Bonne &amp; Bonne-Wepster 1925: 22, 134–137 (in part, flui only).</p><p>Wyeomyia (Prosopolepis) of Bonne-Wepster &amp; Bonne 1923: 124.</p><p>Wyeomyia (Heliconiamyia) of Dyar 1923: 64, 65 (in part, chalcocephala only; galoa male determined as chalcocephala); Dyar &amp; Shannon 1924a: 85 ( Heliconiamyia restricted to chalcocephala; synonymy of galoa with pseudopecten Dyar &amp; Knab, 1906a); Bonne &amp; Bonne-Wepster 1925: 54, 59–61, 85.</p><p>Prosopolepis (Prosopolepis) in part of Dyar &amp; Shannon 1924b: 481, 482 ( flui only).</p><p>Prosopolepis (Heliconiamyia) of Dyar &amp; Shannon 1924b: 481, 482; Dyar 1925: 118–120, 122.</p><p>Dendromyia (Heliconiamyia) of Dyar 1928: 67, 77 (as Heliconiamyia, p. 69).</p><p>Dendromyia (? Prosopolepis) of Dyar 1928: 89, 90 (in part, only flui; Prosopolepis as tentative subgenus of Dendromyia).</p><p>Wyeomyia (Dendromyia) of Edwards 1932: 83, 86, 87 (in part, chalcocephala and flui only; flui as tentative synonym of confusa (Lutz, 1905b)); Del Ponte &amp; Cerqueira 1938: 225, 226, 228, 229–235 (in part, chalcocephala, flui and kerri only; kerri description); Lane 1939: 139–148 (in part, chalcocephala and kerri only; synonymy of flui with confusa); Lane 1951: 335 (in part, chalcocephala, flui and luciae only; synonymy of luciae with chalcocephala, flui as synonym of confusa); Senevet et al. 1942: 336–347 (in part, chalcocephala and luciae only; luciae description); Bruijning 1959: 100, 109–143 (in part, flui and surinamensis only; flui as synonym of confusa, surinamensis description); Stone et al. 1959: 9, 83–88 (in part, chalcocephala, flui, kerri and luciae only; luciae as synonym of chalcocephala, flui as synonym of confusa); Belkin et al. 1965: 5, 6, 22, 23, 26–28, 47–49, 57–59, 62, 66, 68, 71–73, 76, 77 (in part, chalcocephala, flui, luciae and surinamensis only; luciae as synonym of chalcocephala, flui as synonym of confusa); Belkin 1968: 38–43 (in part, chalcocephala, flui, luciae and surinamensis only; flui as synonym of confusa, synonymy of luciae with chalcocephala contested); Knight &amp; Stone 1977: 4, 327–332 (in part, chalcocephala, flui, kerri, luciae and surinamensis only; luciae as synonym of chalcocephala, flui as synonym of confusa); Peyton et al. 1983: 67, 74, 85, 87, 88 ( kerri only); Guimarães 1997: 118, 119, 128, 129 (in part, chalcocephala, flui, kerri, luciae and surinamensis only; flui as synonym of confusa, luciae as synonym of chalcocephala).</p><p>Dendromyia in part of Del Ponte 1939: 536, 540, 541 ( chalcocephala, flui and kerri only).</p><p>Wyeomyia (Dendromyia) Prosopolepis Series in part of Lane &amp; Cerqueira 1942: 536, 537, 589, 615–619 ( flui and kerri only; flui resurrected from synonymy with confusa); Lane 1953: 867, 995–1001 ( flui and kerri only; flui as synonym of confusa).</p><p>Wyeomyia (Dendromyia) Dendromyia Series in part of Lane &amp; Cerqueira 1942: 536, 537, 589–611 ( chalcocephala only); Lane 1953: 868, 946–988 ( chalcocephala and luciae only; luciae as synonym of chalcocephala).</p><p>Wyeomyia (Wyeomyia) in part of Horsfall 1955: 327, 328 ( chalcocephala only; lapsus inclusion in subgenus Wyeomyia); Linton et al. 2013: 102, suppl. data 2 ( chalcocephala only; lapsus inclusion in subgenus Wyeomyia).</p><p>Wyeomyia (subgenus D) of Heinemann &amp; Belkin 1977: 435, 436, 443, 447, 450, 454; Heinemann &amp; Belkin 1978a: 124–126, 128–133, 135, 136, 149–152, 189, 193, 196; Heinemann &amp; Belkin 1978c: 513, 514, 526; 1979: 99, 100, 102, 109.</p><p>Wyeomyia (? Calladimyia) of Heinemann &amp; Belkin 1978a: 163, 188.</p><p>Wyeomyia (Calladimyia) in part of Heinemann &amp; Belkin 1978b: 398, 399, 403, 405, 409, 420-425, 428, 433, 439, 443, 444, 446, 452, 453, 458 ( surinamensis only); Heinemann &amp; Belkin 1979: 97, 98, 109 ( Wyeomyia sp. 28; possibly surinamensis only).</p><p>Wyeomyia (subgenus uncertain) in part of Motta &amp; Lourenço-de-Oliveira 1995: 384 ( chalcocephala, kerri and surinamensis only; 46 species relegated to subgenus uncertain); Lourenço-de-Oliveira et al. 1999: 200, 201, 207–211 ( flui and kerri only; flui resurrected from synonymy with confusa, synonymy of kerri with flui); Nascimento-Pereira et al. 2019: 359, 360 ( chalcocephala, flui and surinamensis only); Wilkerson et al. 2021b: 613, 1028–1031 ( chalcocephala, flui, kerri, luciae and surinamensis only; kerri as synonym of flui, luciae as synonym of chalcocephala).</p><p>Wyeomyia (subgenus uncertain) Flui Group of Motta et al. 2007: 613, 616, 618, 619, 625, 626.</p><p>The subgenus Leonidasdeanea includes some of the largest species of the genus Wyeomyia . The adults are moderately ornamented with brown scutal scaling with weak greenish-blue reflections, pleura with cream-colored scales, pale scales on the mesopostnotum and an evident pattern of white scaling on the mid- and hindlegs. The combination of characters easily distinguishes species of Leonidasdeanea from most species of the genus Wyeomyia . In addition, females of the subgenus Leonidasdeanea have only one spermathecal capsule. Sexual dimorphism is not only evident in the genitalia, but also in the appearance of the antenna, the flagellum of which is conspicuously more verticillate in males. The male genitalia have tergum IX with about three strong setae and the gonostylus composed of an elongate stem apically divided into four lobes, with lobe C bent backwards into a “J” to “L” shape; sternum IX in a bilobular shape; and the gonocoxite with more than 10 tergal setae. Pupae have a trumpet index that usually ranges from 4.5–5.5; paddles not pointed, essentially truncate; seta 5-CT usually double; setae 2-II–VII inserted mesally; seta 6-VII inserted dorsal. Larvae have maxillae with a rudimentary apical tooth; comb scales arranged in more than one uneven row, not borne on a plate; siphon with seta 2a-S usually with 2–4 aciculate branches and a pecten composed of a single row of slender elongate filaments inserted nearly mid-ventrally.</p><p>Bionomics. All species of Wyeomyia (Leonidasdeanea) are sylvatic. Adults are active during the daytime and females attack humans. The immature stages have been collected mostly from the leaf axils of palm trees, especially in buriti ( Mauritia flexuosa L.f), but also in the leaf axils of plants of the family Araceae, and leaf axils and flower bracts of plants of the order Zingiberales .</p><p>Distribution. Species of the subgenus Leonidasdeanea have been found in locations from Honduras to midwestern Brazil.</p><p>Etymology. The subgeneric name (feminine) is a tribute in recognition of the Brazilian parasitologist, malariologist and specialist of anopheline mosquitoes, Leonidas de Mello Deane. The three-letter abbreviation Leo . is recommended for this subgenus.</p></div>	https://treatment.plazi.org/id/03A187D23400CF2DFF4EFF217930FD4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lourenço-De-Oliveira, Ricardo;Pereira, Glauber Rocha;Neves, Maycon Sabastião Alberto Santos;Santos, Stephani Da Silva;Ribeiro, Paulino Siqueira;Nascimento-Pereira, Agostinho Cardoso;Talaga, Stanislas;Pavan, Márcio Galvão;Sallum, Maria Anice Mureb;Motta, Monique Albuquerque	Lourenço-De-Oliveira, Ricardo, Pereira, Glauber Rocha, Neves, Maycon Sabastião Alberto Santos, Santos, Stephani Da Silva, Ribeiro, Paulino Siqueira, Nascimento-Pereira, Agostinho Cardoso, Talaga, Stanislas, Pavan, Márcio Galvão, Sallum, Maria Anice Mureb, Motta, Monique Albuquerque (2026): Systematics of Leonidasdeanea, a new subgenus of Wyeomyia (Diptera: Culicidae), with recognition of Wy. luciae as a valid species and synonymy of Wy. surinamensis with Wy. flui. Zootaxa 5782 (1): 1-43, DOI: 10.11646/zootaxa.5782.1.1, URL: https://doi.org/10.11646/zootaxa.5782.1.1
03A187D23401CF21FF4EFD617F0DFA9F.text	03A187D23401CF21FF4EFD617F0DFA9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wyeomyia (Leonidasdeanea) flui (Bonne-Wepster & Bonne 1920)	<div><p>Wyeomyia (Leonidasdeanea) flui (Bonne-Wepster &amp; Bonne, 1920)</p><p>flui Bonne-Wepster &amp; Bonne, 1920: 169 ( Prosopolepis; ♀).</p><p>Type ♀: Albina (April 1917) and Dam (January 1919), Suriname, J. Bonne-Wepster &amp; C. Bonne coll. (RMNH).</p><p>kerri Del Ponte &amp; Cerqueira, 1938: 228 (♀, ♂, ♂ G*). Holotype ♀ (nº 2210): Cuiabá, Mato Grosso, Brazil (February–July 1935), Gastão Cesar coll. (CEIOC).</p><p>surinamensis Bruijning, 1959: 135 (♂, ♂ G*). Holotype ♂: Ornamibo, Suriname (11 August 1954), C.F.A. Bruijning coll. (RMNH). New synonymy.</p><p>Prosopolepis flui of Dyar &amp; Ludlow 1921: 677 (♀, taxonomy); Bonne &amp; Bonne-Wepster 1925: 134–136, 551 (♀, bionomics, classification, distribution, etymology, key, species list Suriname, taxonomy); Lane 1939: 142, 214 (synonymy with confusa, classification, distribution, record from Panama erroneous); Lane 1953: 997, 1104 (synonym of confusa, taxonomy); Bruijning 1959: 121 (synonym of confusa, taxonomy); Stone et al. 1959: 85, 340 (synonym of confusa, taxonomy, type info.).</p><p>Wyeomyia (Prosopolepis) flui of Bonne-Wepster &amp; Bonne 1923: 124 (species list Suriname); Dyar &amp; Shannon 1924b: 482 (classification, taxonomy); Dyar 1928: 68–70, 90, 608 (♀, classification, distribution, key, taxonomy).</p><p>Wyeomyia flui of Edwards 1932: 87, 231 (tentative synonymy with confusa, classification, distribution, taxonomy); Knight &amp; Stone 1977: 329, 577 (synonym of Wy. confusa, taxonomy, type info.); Guimarães 1997: 129, 272 (synonym of confusa, classification, type info.); Hendy et al. 2023: 10 (ecology, collection method, protected area); Montalvo-Sabino et al. 2025: 867, 868, 871–873, suppl. (♂ G*, DNA sequence, phylogeny, record Peru); Ayala-Sulca et al. 2025: 134 (nomen dubium in error, species list Peru).</p><p>Dendromyia confusa of Lane 1936a: 181 (bionomy, distribution, Mato Grosso State, Brazil).</p><p>Wyeomyia kerri of Kumm &amp; Novis 1938: 501, 503, 507, 508, 511 (bait catch, bionomics, habitat, collection method, endophily, record Brazil); Horsfall 1955: 330, 719 (habitat, distribution); Aitken 1960: 4 (record Trinidad and Tobago, survey); Aitken 1973: 15 (protected area, Trinidad and Tobago); Mattos &amp; Xavier 1965: 287 (distribution, species list Goiás State, Brazil); Aitken et al. 1969: 210 (arbovirus screening, survey); Marchon-Silva et al. 1996: 476 (type info.); Motta &amp; Lourenço-de-Oliveira 2000: 682 (synonym of flui); Harbach 2018: 49, 68 (synonym of flui, nomenclature); Wilkerson et al. 2021b: 1029, 1285 (synonym of flui, taxonomy, type info.).</p><p>Wyeomyia (Dendromyia) flui of Del Ponte &amp; Cerqueira 1938: 225, 228, 231, 236 (♀, key subgenera of Wyeomyia, systematics, taxonomy); Lane 1951: 335 (synonym of confu sa, taxonomy); Belkin et al. 1965: 62, 66, 94 (synonym of Wy. confusa, bionomics, habitat, type info.); Belkin 1968: 40, 45, 67 (synonym of Wy. confusa, depository, taxonomy, type info.).</p><p>Wyeomyia (Dendromyia) kerri of Chagas et al. 1938: 203, 206, Tables 46, 48 (bait catch, bionomics, daily rhythm); Lane 1939: 144, 215 (classification, distribution); Cerqueira 1943: 21 (distribution, species list Bolivia); Cerqueira 1961: 158 (bait catch, bionomics, habitat, species list Brazilian Amazon, vertical stratification); Deane et al. 1953: 103 (bionomics); Stone et al. 1959: 85, 344 (taxonomy, type info.); Prosen et al. 1964: 78, 113 (distribution, species list Bolivia); Aitken et al. 1968: 256 (bionomics); Xavier &amp; Mattos 1970: 453 (distribution, species list Mato Grosso State, Brazil); Xavier &amp; Mattos 1975: 258 (distribution, species list Pará State, Brazil); Xavier &amp; Mattos 1976: 403 (distribution, species list Amazonas State, Brazil); Xavier et al. 1989a: 292 (distribution, species list Rondônia State, Brazil); Xavier et al. 1989b: 304 (distribution, species list Maranhão State, Brazil); Belkin et al. 1971: 11, 33, 47, 52, 62 (bionomics, habitat, depository, taxonomy, type info.); Knight &amp; Stone 1977: 329, 584 (classification, distribution, taxonomy, type info.); Peyton et al. 1983: 67, 74, 75, 85, 87, 88 (bionomics, habitat, distribution Bolivia, survey); Xavier et al. 1989c: 312 (distribution, species list Acre State, Brazil); Guimarães 1997: 119, 275 (classification, type info.); Hutchings et al. 2005a: 18 (synonym of flui, depository).</p><p>Dendromyia kerri of Del Ponte 1939: 536, 541 (♀, identification method, taxonomy).</p><p>Dendromyia flui of Del Ponte 1939: 540 (♀, identification method, taxonomy).</p><p>Wyeomyia (Dendromyia) confusa of Lane 1939: 142 (in part, only specimens from Mato Grosso State, Brazil, and Suriname); Bruijning 1959: 100, 122, 123 (in part, only specimens from Suriname; ♀, taxonomy); Stone et al. 1959: 85, 337 (in part, only specimens from Bolivia, Suriname and Venezuela; distribution, taxonomy, type info.); Stone 1961: 34 (distribution, record French Guiana); Prosen et al. 1964: 77, 113 (bait catch, distribution, species list Bolivia); Deane et al. 1968: 338, 339 (ecology, simian malaria parasite screening, vertical stratification); Xavier &amp; Mattos 1970: 453 (distribution, species list Mato Grosso State, Brazil); Xavier &amp; Mattos 1975: 258 (distribution, species list Pará State, Brazil); Xavier &amp; Mattos 1976: 403 (distribution, species list Amazonas State, Brazil); Panday 1975a: 143 (bionomics, collection method); Panday 1975b: 299 (bionomics, collection method); Knight &amp; Stone 1977: 329 (in part, only specimens from Bolivia, French Guiana, Suriname and Venezuela; classification, distribution, taxonomy, type info.).</p><p>Wyeomyia (Dendromyia) flui of Lane &amp; Cerqueira 1942: 537, 543, 548, 589, 616, 619, 846 (♀, resurrected from synonymy with confusa, classification, key, revision, systematics).</p><p>Wyeomyia (Dendromyia) kerri of Lane &amp; Cerqueira 1942: 537, 541, 548, 589, 619, 744, 745, 847 (♀, ♂, ♂ G*, bionomics, habitat, classification, distribution, keys, record Bolivia, revision, systematics); Lane 1953: 867, 871, 1000, 1001, 1105 (♀, ♂, ♂ G*, classification, distribution, keys, revision, systematic, type depository).</p><p>Wyeomyia (Dendromyia) surinamensis of Stone 1961: 34, 52 (taxonomy, type info.); Belkin et al. 1965: 62, 66, 94 (bionomics, habitat, type info.); Belkin 1968: 42, 45, 56, 69 (depository, taxonomic status questioned, type info.); Knight &amp; Stone 1977: 331, 605 (classification, distribution, taxonomy, type info.).</p><p>Wyeomyia confusa of Cova Garcia et al. 1966: 72, 408 (in part, only females; key Venezuela).</p><p>Wyeomyia surinamensis of Wilkerson et al. 2021a: 288, 599 (DNA sequence); Montalvo-Sabino et al. 2025: 871–873 (molecular phylogeny).</p><p>Wyeomyia (Calladimyia) surinamensis of Heinemann &amp; Belkin 1978b: 403, 409 (bionomics, habitat, distribution).</p><p>Wyeomyia (Calladimyia) sp. 28 (possibly surinamensis) of Heineman &amp; Belkin 1979: 97, 98, 109 (bionomics, distribution, record, Ecuador).</p><p>Wyeomyia (subgenus uncertain) kerri of Motta &amp; Lourenço-de-Oliveira 1995: 384 (classification, systematics); Lourenço-de-Oliveira et al. 1999: 200, 201, 209, 210 (synonymy with flui, classification, distribution, taxonomy).</p><p>Wyeomyia (subgenus uncertain) surinamensis of Motta &amp; Lourenço-de-Oliveira 1995: 384 (classification, systematics); Guimarães 1997: 132, 283 (classification, distribution, type info.); Hutchings et al. 2013: 5, 14 (collection method, bionomics, survey); Talaga et al. 2015: 776, 779, 780, suppl. data (bionomics, habitat, depository, distribution, species list French Guiana, type info.); Talaga et al. 2016: 1140, 1142 (habitat, ecology, interactions networks, morphology); Talaga et al. 2017: 6, 10, suppl. tables 1 and 2 (16S, Barcode Index Number, COI, molecular phylogeny, systematics); Harbach 2018: 126, 217 (etymology, nomenclature, type loc.); Nascimento-Pereira et al. 2019: 360 (classification, subgenus uncertain); Wilkerson et al. 2021b: 1031, 1304 (classification, distribution, etymology, informal name, taxonomy, type info.); Harbach 2026 (classification, taxonomy).</p><p>Wyeomyia (subgenus uncertain) confusa of Guimarães 1997: 128, 129, 270 (in part, only specimens from Bolivia, French Guiana, Suriname and Venezuela; classification, distribution, type info.).</p><p>Wyeomyia (subgenus uncertain) flui of Lourenço-de-Oliveira et al. 1999: 200, 201, 209–211 (synonymy with confusa, bionomics, nomenclature, systematics); Pecor et al. 2000: 214, 217 (bait catch, bionomics, collection method, record Peru); Motta &amp; Lourenço-de-Oliveira 2000: 682 (L, classification, systematics); Hutchings et al. 2002: 113, 116, 118 (collection method, survey); Hutchings et al. 2005a: 18, 25 (depository, distribution Brazil); Hutchings et al. 2005b: 434 (protected area, survey); Motta et al. 2007: 593–601, 605, 607, 610–613, 616, 618, 619, 625, 626 (Flui Group; L, Lmp, P, ♀, ♀ G, ♂, ♂ G, classification, morphology, phylogeny, systematics); Hutchings et al. 2011: 178, 182, 185 (distribution Amazonas State, Brazil; ecology, protected area, survey); Hutchings et al. 2013: 14 (collection method, ecology, survey); Hutchings et al. 2016: 1339 (collection method, survey); Hutchings et al. 2018: 1202 (collection method, distribution Amazonas State, Brazil, survey); Linton et al. 2013: suppl. data 2 (species list Ecuador); Harbach 2018: 49, 68, 169 (etymology, nomenclature); Nascimento-Pereira et al. 2019: 360 (classification, subgenus uncertain); Brilhante et al. 2020: 100 (distribution, species list Acre State, Brazil); Wilkerson et al. 2021b: 1029, 1279 (classification, distribution, etymology, index, informal name, synonymy, taxonomy, type info.); Harbach 2026 (classification, species list, taxonomy).</p><p>Female. Head (Fig.1A–E):Vertex and occiput covered with large brown decumbent scales with blue-gray reflections; vertex with small median spot with pale reflections; occiput with transverse line of short dark semi-erect scales posteriorly; postgena with white scales; ocular line without white scales; ocular setae brown, including 2 long brown interocular setae. Interocular space narrow above and below, without setae and scales. Frons yellowish, without setae. Clypeus brown, pruinose, rounded, without setae, most of margin with laterally directed narrow pearly scales (Fig. 1A). Maxillary palpus short, slightly longer than clypeus, about 0.2 length of proboscis, brown-scaled with elongate brown setae near apex. Proboscis: Length 1.95–2.2 mm (2 mm), expanded at apex; dorsal surface dark brown-scaled, ventral surface dark brown with line of white scales, more distinct from mid-length to distal 0.75, and may extend less conspicuously basally or distally; about 10 basal labial setae, brown, elongate, progressively shorter laterally; labellum with pale integument. Antenna: As long as proboscis; pedicel brown, progressively paler on latero-ventral surface, pruinose, with minute setae and pearly scales dorsomesad. Flagellum moderately verticillate, whorls with dark setae (Fig. 1A,B). Thorax: Scutum with integument brown, covered with elongate brown scales with greenish-blue reflections (Fig. 1B,C); prescutellar area partially bare (Fig. 1D); around 10 strong brown prescutellar setae; anterior promontory with a few white scales mesally and 10–12 darkish setae of different sizes (Fig. 1A,B); no acrostichal or dorsocentral setae. Supraalar and antealar areas with 27–36 darkish brown setae. Scutellum (Fig. 1D,E) covered with scales concolorous with scutum, limited on lobes; median lobe with 5 long and 2–5(3) short darkish brown setae, lateral lobes with 4,5(5) long and 3,4 short darkish brown setae. Mesopostnotum brown, pruinose, with roughly triangular patch of decumbent spatulate pearly white diaphanous scales anterior to tuft of 5–7(6) brown setae (Fig. 1E). Antepronotum entirely covered with brown scales with faint violaceous reflections, lobes well separated, with row of 11–13(12) darkish brown setae dorsally. Postpronotum covered with white scales; dark scales restricted to dorsal area. Pleura with pale brown integument covered with pearly white scales (Fig. 1C); proepisternum, anterior lateral margin of mesokatepisternum, mesomeron, paratergite, metapleural suture and metapostnotum without scales. Pleural chaetotaxy as follows: 2–4(3) light brown prespiracular setae, long, extending beyond spiracle; postspiracular setae absent; 6–8(7) upper proepisternal setae, yellowish, mostly long; 8,9(9) lower mesokatepisternal setae, 2 setae above upper margin of mesomeron; upper mesokatepisternal setae absent; prealar area with 4–6(6) brown setae; lower mesepimeral setae absent; 10–12(12) pale yellowish upper mesepimeral setae, nearly in a row. Halter (Fig. 1C,D): Integument light brown; scabellum and pedicel dark-scaled dorsally, capitellum dark-scaled. Wing: Length 4 mm (3.85–4.25 mm), scales brown with slight bluish reflections, spatulate with rounded apices on all veins. Dorsal scales: veins R 1 –R 3, R 4+5, M 1, M 2 and distal M 3+4 with rather broad spatulate scales with rounded apices; M with long spatulate scales; Cu and 1A with moderately broad scales. Ventral scales: veins M 3+4, distal CuA and 1A with long spatulate scales. Calypter with 2,3(3) brown setae. Alula with 10,11 simple brown setae. Legs: Coxae and trochanters pale, with pearly white scales and elongate yellowish setae. Femora, tibiae and tarsomeres 1 mainly dark-scaled; femora and tibiae white-scaled ventrally; foretarsomeres 1–5 dark-scaled; midtarsomere 1 dark-scaled; midtarsomeres 2–5 white-scaled; midtarsomere 2 entirely white-scaled on dorsal surface, sometimes with white-scaling consisting of a less conspicuous line on proximal 0.3; ventral surface of midtarsomere 2 dark-scaled; white-scaling usually entirely encircling midtarsomeres 3–5, which may be interrupted with a line of few dark scales, especially on distal third of ventral surface; hindtarsomeres 1 and 2 dark-scaled, hindtarsomeres 3 and 4 white-scaled on proximal 0.9–0.95 of ventral surface; hindtarsomere 5 entirely white-scaled ventrally. Ungues simple. Abdomen: Terga covered with brown spatulate scales with bluish-green reflections. Tergum I with numerous yellowish setae forming a row on anterolateral margin. Sterna covered with whitish scales; color on sides separated in nearly a straight line, sometimes a few brown scales invade apical-lateral margins of terga III and IV. Genitalia (Figs 2, 3): Tergum VIII wider than long, covered with minute spicules and spatulate scales; distal margin with short setae aligned longitudinally. Sternum VIII covered with minute spicules and spatulate scales, invaginated in middle posteriorly, almost forming two lobes. Distal margin with short setae that become longer and sparser from middle to lateral margin and thinner from anterior to posterior area. Tergum IX narrow, covered with minute spicules and with a group of 2 and 3 short setae, in line on each weakly defined lobe. Insula rounded, with anterior margin slightly flattened and covered with small spicules, with a group of 3 or 4 setae on each side of apex. Postgenital lobe elongate and narrow, extending beyond the cerci. Dorsal surface with 14 long scattered setae with conspicuous alveoli; ventral surface covered with minute spicules, numerous short setae distributed from base to apex, with conspicuous alveoli. Cerci covered with minute spicules; apices flat, very expanded in inner portion. Spermatheca a single capsule.</p><p>Male. Similar to female except for the following characteristics: Head (Fig. 1F): Antenna with flagellum distinctly more strongly verticillate, except the last 2 flagellomeres. Legs: Hindtarsomere 4 with ventral surface entirely white-scaled or white-scaled except on distal 0.1. Genitalia (Figs 4, 5): Tergum VIII (Fig. 4G) with numerous scales and minute setae on anterior half; anterior margin with long setae laterally and shorter ones in middle; rounded anterior corners, without lobes or tuft of setae. Sternum VIII (Fig. 4H) covered with moderately long setae, numerous scales and minute setae, uniformly distributed on anterior half. Tergum IX (Fig. 4C) with bare interlobar space, lobes well separated and defined, each bearing 3 strong pointed setae with tips facing outward, setae similar in shape and length. Sternum IX fused to tergum, with a median plate divided into 2 lobes (Fig. 4E). Gonocoxite (Fig. 4A,B) slightly ovate; tergal surface with scales and short setae, mainly on margin; tergomesal setae long and numerous, about 12. Basal mesal lobe roughly triangular with about 25 fine setae. Gonostylus (Fig. 4A,B) shorter than gonocoxite with a narrow and slightly curve stem, apically divided into 4 lobes (Fig. 5); lobe A rotund, short, covered with numerous short and curved setae, bearing a strong short seta with curved apex (Fig. 5B); lobe E adjacent to lobe A, large, margin with 15 strong setae with essentially rounded apices, roughly arranged in 2 rows; lobe M elongate with 10 short setae of different sizes, and a long conspicuous seta-like process arising from tergal surface and extends toward the base of the gonostylus (Fig. 5A,C,D); lobe C narrow, elongate and curved backwards, similar in shape to a fishhook (Fig. 5A,B). Aedeagus (Fig. 4F) rounded, submedian tergal arms not fused, apically digitiform, narrowed and turned inward; median sternal plate membranous, folded laterally and serrate on margin; proctiger narrow; paraproct sclerotized dorsoapically, 2 small apical teeth, 4 minute cercal setae (Fig. 4D).</p><p>Pupa (Fig. 6). Position and character of setae as figured, numbers of branches in Table 1. Cephalothorax: Integument brownish, darker on dorsum; seta 1-CT long, strong, sigmoid; seta 5-CT long, aciculate near apex; seta 10-CT branched, more than 0.6 length of seta 11-CT; seta 12-CT much longer than 11-CT. Trumpet: Integument brown, slightly cylindrical, narrowed at base, index 3.7–5.5 (mean 4.7) (width measured at mid-length). Abdomen: Integument brownish, darker from metanotum to tergum III, darker on transverse basal line on segments III to VI. Seta 1-II–VII near posterior margin of tergum, 1-III–VII lateral to seta 2, 1-III lateral to seta 3, 1-IV–VII short, nearly nearly similar in length to seta 3; seta 2-III–VII inserted mesally, seta 3-II,III similar in length, 3-V–VII close to seta 1; seta 4-I,III usually multiple, similar in length, 4-V,VI far from seta 3-V,VI, 4-VII similar in length to seta 6; seta 5-I–III short, nearly similar in length to seta 2, 5-IV–VI very long, almost reaching apex of next tergum, aciculate; seta 6-I–VI long, similar in length; seta 7-I short, similar to seta 4, 7-II dorso-lateral in position, 7-III–V similar in length to seta 8, 7-VI longer than seta 8; seta 8-III–VII lateral to seta 7; seta 9-I–VI minute, lateral in position, 9-VII,VIII multi-branched; seta 10-I absent, 10-V longer than on other terga; seta 11-II–VII minute, 11-VI lateral to seta 10. Paddle little longer than wide, apex slightly straight, midrib well defined; index 1.3–1.5.</p><p>D = dendritic.</p><p>Larva, fourth-instar (Fig. 7). Position and character of setae as figured. Head: Slightly wider than long, light tan. Collar absent, occipital foramen with slit-like extension on either side. Hypostomal suture complete. Dorsomentum with 10–13(11) lateral teeth on either side of a slightly larger median tooth. Lateral teeth similar in development, except lateralmost teeth sometimes very small. Maxilla: Rounded; maxillary brush long, similar in length to seta 4-Mx. Apical tooth absent. Laciniarastrum 1 (LR1) with row of about 11 short teeth of similar development, except for first 2 longer than others; LR2 distributed on third of maxillary body, forming a dense group of long setae close to LR1. Maxillary palpus not fused to maxillary body, with 3 short digitiform setae, 1 far from the others; seta 1-Mx single, stout, forked at apex, inserted in apical third; seta 2-Mx single, short; seta 4-Mx single, long, inserted on anterior margin; seta 6-Mx single. Mandible: Ventral mandibular tooth 0 (VT 0) prominent and larger than VT 1, VT 2, VT 3; ventral mandibular 4 fine, elongate; mandibular sweeper (MnS) 1 with usually 4 setae, MnS2 with usually 6 setae. Antenna: Slender, slightly longer than seta 7-C; seta 1-A usually single, inserted on apical fourth of antenna. Seta 0-C single, minute; seta 1-C single, elongate, curved, stout; setae 3,5-C single; seta 3-C minute; seta 5-C elongate, as long as seta 4-C; seta 4-C with 1–3(2) branches; seta 6-C as long as seta 7-C, normally single; seta 7-C with 2,3(2) branches; setae 8,9,12-C short, usually double, 8,9-C with 2–4(2) branches; seta 10-C with 1–4(2) branches, almost 3 times length of seta 9-C; seta 11-C aciculate, usually with 10–12 branches; seta 14-C elongate, 4 times length of seta 15-C, usually single; seta 15-C short, with 2–4(3,4) branches, inserted almost in line with 14-C. Thorax: Integument smooth. Setae 4-P, 7,13-T inserted on individual pigmented basal plates; pleural groups 5–7-P, 9–12-P,M,T on common basal plates; seta 0-P small, with 4–7(7) branches; seta 1-P with 4–9(4,5) branches, anterior in position and mesad to setae 2,3-P; seta 2-P long, about twice length of setae 1,3-P, single; seta 4-P with about 12 aciculate branches; setae 5,6-P usually single, similar in length; seta 7-P with usually 12,13 branches of different lengths; setae 8,14-P small, branched, nearly half length of setae 1,3-P; setae 5–7,9,10,12-P, 5,6,9,10,12-M and 7,9,12,13-T aciculate; seta 9-P branched; seta 10-P half length of seta 12-P; seta 11-P,M single, slender, spiniform, 11-T stronger than 11-P,M, rather stout, with stiff spike-like branches on distal third; seta 12-P single, similar in length to 9-P; setae 1,2-M with 1–3(1) branches; setae 3,5,6,9–12-M single; setae 1–4-M almost in line; seta 3-M long, around 3 times length of 1-M; seta 7-M small, normally with 3 branches; seta 8-M elongate, slightly longer than 3-M; setae 9,10,12-M well developed; seta 10-M similar in length to 12-M; setae 13,14-M small, almost similar in size, with 4–8 branches; setae 1–3-T almost in line; setae 2–4-T branched, almost similar in length; seta 5-T single; seta 6-T usually double, about 1.5 length of 5-T; seta 7-T similar in size to seta 9-T; seta 10-T single, 0.4 length of seta 12-T; seta 12-T single, longer than 9-T; seta 13-T multi-branched. Abdomen: Seta 1-I–VII branched, well developed, slightly stellate, aciculate, lateral and posterior in position to seta 2-I–VI; seta 2-I–VII single, small, mesad to setae 1–4; setae 3,4-I,III,IV usually branched, small, almost similar in length; setae 3,5-I–VII with 2,3 branches, 3-V single, longer than seta 1, 3-VII single, strong, about 1.5 length of siphon, aciculate; seta 4-I–VII small, with 1‒3(2) branches, 4-IV–VI mesad to seta 1, 4-VI almost in line with seta 1, about 2.0 length of seta 2; seta 5-I–VI normally double, small, about 0.35 length of seta 1, 5-VII very small, similar in length to setae 8 and 9; setae 6,7-I,II usually with 4,5 branches of different lengths, aciculate, inserted on individual pigmented basal plates, seta 6-I shorter than setae 7-T, 6-II, 6-III–VI long, with 2,3 branches, aciculate, 6-VII small, similar in length to setae 7,10,12, normally with 5 branches; seta 7-III–VII small, branched; seta 8-II usually single, nearly same length as seta 5, 8-III–VII small, usually with 3–5 branches; seta 9-I–VII small, nearly same length as seta 2, 9-I with 1,2(1) branches, 9-II–VI single, 9-VII branched; seta 10-I–VII small, branched, 10-I well mesad and posterior to setae 9,11,13, 10-II–VI lateral and posterior to setae 11,13-II–VI; seta 11-I–VII small, shorter than seta 10; seta 12-II,III small, distinctly lateral to seta 13, 12-V–VII mesad to seta 13, 12-II usually with 3,4 branches, almost similar in length to seta 11, 12-III,V usually single, 12-V about 2.0 length of seta 11, 12-VI,VII usually with 4 branches; seta 13-I–V slightly stellate, aciculate, 13-I–III usually with 3,4 branches, anterior to setae 10,11, about 0.7 length of seta 1, 13-IV,V with 2–5(3) branches, longer than seta 1, 13-V almost in line with setae 11,12, 13-VI,VII small, usually with 3 branches. Segment VIII: Comb composed of a group of about 20 elongate scales arranged in more than 1 uneven row, not borne on a plate; scales elongate, essentially of 2 sizes, fringed at apices. Setae 1–4-VIII branched, small; seta 1-VIII usually with 6 branches, slightly shorter than seta 3, nearly 0.2 length of seta 2,4; seta 3-VIII with about 8 branches, 0.7 length of seta 4, mesad to seta 4; seta 4-VIII usually with 5 branches; seta 5-VIII usually single, aciculate, elongate, ≥ 3.0 length of setae 1–4. Siphon: Moderately long; index 1.5–2.7 (mean 2.4; mode 2.5) (width measured at base); tanned, more pigmented at base. Pecten composed of a row of around 10 slender elongate filaments usually limited to mid-length of siphon; seta 1-S normally inserted at distal limit of basal quarter of siphon, with 4–6(4) branches, aciculate; seta 1a-S usually with 2 single mid-ventral setae inserted at distal quarter of siphon; seta 2a-S normally with 4 elongate aciculate setae mostly distributed on proximal half of siphon. Segment X: Saddle slightly more tanned than siphon. Setae 1–4-X strong, aciculate; seta 1-X single, longer than seta 2–4; setae 2–4-X similar in development and length, 2-X usually with 8 branches, 3-X normally with 3 branches, 4-X normally with 5,6 branches, 0.6 length of seta 1.</p><p>Bionomics. Larvae of Wy. flui have been found most frequently in water accumulated in leaf axils of palms and palm stumps. We have most frequently collected the immature stages in the leaf axil of buriti palms (Lourenço-de-Oliveira et al. 1999), but they have also been found in leaf axils of açai palms ( Euterpe oleracea Mart.) (Kumm &amp; Novis 1938). Peyton et al. (1983) collected larvae in high densities in the matacú palm (genus Attalea Kunth) and suggested a close association of Wy. flui and palm forests. However, larvae were also exceptionally reported to be collected from cut or broken bamboo ( Bambusa vulgaris Schrad.) (Heinemann &amp; Belkin 1978b) and a tree hole (Kumm &amp; Novis 1938). We collected one larva of Wy. flui together with Wy. testei Senevet &amp; Abonnenc, 1939, Wy. (Dodecamyia) aphobema Dyar, 1918 and an undetermined species of Toxorhynchites Teobald, 1901a in a terrestrial bromeliad (pineapple) in the forest edge in the Amazon with high density of buriti palms (CCULI, http://cculi. fiocruz.br). Females of Wy. flui have been frequently captured on human bait in the forest during daytime (Bonne &amp; Bonne-Wepster 1925; Kumm &amp; Novis 1938; Deane et al. 1953; Cerqueira 1961; Panday 1975a,b; Lourenço-de-Oliveira et al. 1999; Pecor et al. 2000), but was once reported biting indoors (Kumm &amp; Novis 1938). The species was also captured in a trap baited with aguti (Kumm &amp; Novis 1938).</p><p>Distribution. Wyeomyia flui (as Wy. confusa, Wy. kerri or Wy. surinamensis) has been found in Bolivia, departments of Beni (as Colonias), La Paz, Santa Cruz (Cerqueira 1943); Brazil, states of Acre (Lane &amp; Cerqueira 1942), Amapá, Amazonas (Cerqueira 1961), Bahia (Hutchings et al. 2005a), Goiás (Lane &amp; Cerqueira 1942), Maranhão (Xavier et al. 1989b), Mato Grosso (Del Ponte &amp; Cerqueira 1938), Minas Gerais (this study), Pará (Kumm &amp; Novis 1938), Rondônia (Cerqueira 1961), Roraima (this study); Ecuador, province of Napo (Heinemann &amp; Belkin 1979); French Guiana (Stone 1961); Guyana (Lourenço-de-Oliveira et al. 1999); Peru, department of Loreto (Pecor et al. 2000), province of Bagua, Amazonas (Montalvo-Sabino et al. 2025); Suriname, districts of Marowijne (Bonne-Wepster &amp; Bonne 1920), Para (Panday 1975a), Paramaribo (Panday 1975b), Wanica (Bruijning 1959); Venezuela (Stone et al. 1959) and the island of Trinidad (Aitken 1960). The records from the Brazilian southeastern states of Rio de Janeiro and São Paulo, as well as the Colombian department of Meta (Forattini et al. 1970), and Panamá (Lane 1939), need confirmation.</p><p>Material examined. 57 specimens (10♀, 6♀ G, 1♀ Pe, 13♀ LePe, 4♀ LePeG, 1LePeG ♀, 8♂, 4♂ LePe, 4♂ LePeG, 1 LePe ♂, 5G ♂). LECTOTYPE ♀ (RMNH 3561) with associated genitalia mounted on microscope slide (no. 97/29): Albina and Dam, Suriname, April 1917 and January 1919, J. Bonne-Wepster &amp; C. Bonne coll. PARALECTOTYPES: 6♀, 1G ♀ (RMNH), same data. BRAZIL: 1♀ LePe (CCULI 1190), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-63.02236&amp;materialsCitation.latitude=-9.908639" title="Search Plazi for locations around (long -63.02236/lat -9.908639)">Zoological Garden (Parque Botânico)</a> (9°54'31.1"S 63°01'20.5"W), Ariquemes, state of Rondônia, July 25, 1987, reared from larvae collected from the leaf axils of a buriti palm tree, R. Lourenço-de-Oliveira coll., R. Lourenço-de-Oliveira, M.G. Castro and M.A. Motta det. ; 1♀, 3♀ LePe, 4♀ LePeG, 4♀ G, 1LePeG ♀, 4♂, 3♂ LePe, 2♂ LePeG, 2♂ G, 1LePe ♂ (CCULI 6208, 6209, 6211–6213, 6215, 6285–6298, 6301, 6307, 6308, 6316, 6317), same data except April 18–26, 1994, R. Lourenço-de-Oliveira and M.A. Motta det.; 1♀ LePe, 1♂ LePe (CCULI 6210, 6214), same data except São Miguel, Porto Velho ; 1♀ G (CCULI 6216) with larval mouthparts (Dm, Mn and Mx) mounted on microscope slide, same data except, Linha 651, Candeias do Jamari, reared from larvae collected from the leaf axils of pineapple bromeliad ; 1♂ LePeG (CCULI 5806), Bairro das Orquídeas, Pacaraima, state of Roraima, August 30, 2018, reared from larvae collected from the leaf axils of buriti palm tree, A.C. Nascimento-Pereira and F. Maciel coll., A.C. Nascimento-Pereira det. ; 2♀ (CCULI 4057, 4058), Cruzeiro do Sul, state of Acre, 2014, collected with human bait, M.A. Motta coll., M.A. Motta det. ; 6♀ LePe, 1♀ Pe, 1G ♀, 1♂ (CCULI 2148–2151, 2159, 2280–2281, 6299, 6318), road to Mosqueiro, Belém, state of Pará, November 1992, M.A. Motta coll., M.A. Motta det. ; 1♀ LePe (RMNH), same data; 3G ♂ (CCULI 6241– 6246, CCULI 4997), Córrego da Onça, Fazenda Uruguaia, Corinto, state of Minas Gerais, December 07, 2018, M.A. Motta, T.F. Silva-do-Nascimento and M.S.A.S. Neves coll., M.A. Motta det. Specimens originally identified as Wy. kerri: 1♀ (CMN 9853), Igarapé do Rio Araçacá, Curralinho, state of Pará, Brazil, December 04, 1935, Toco de Capemba de Bussu, leaf of bussu palm ( Manicaria saccifera) lying on ground, O. Nóvis and H.W. Kumm coll., Del Ponte and Cerqueira 1938 det. ; 1♀ G (CMN 9867), same data except Rio Itaucú, Curralinho, state of Pará, Brazil, near to roça of Antonio Pereira where monkey trap is located, larvae focus in açai palm holding water, December 13, 1935. Specimens originally identified as Wy. surinamensis: 1♂ LePeG, 1♀ LePe, (NMNH 139 -10, 141-10), about 5 km SE of Cayenne, just W of Raban, 5 m, French Guiana, March 15, 1967, reared from immature stages collected in cut or broken bamboo ( Bambusa vulgaris based on the locality) in clearing in forest, partial shade, 67, 68 Mosquitoes of Middle America, R.X. Schick coll.</p><p>Wyeomyia surinamensis . HOLOTYPE ♂ (RMNH) with associated genitalia mounted on microscope slide (RMNH.INS.856454): Ornamibo, Suriname, August 11, 1954, C.F.A. Bruijning coll. PARATYPES: 1♂ (RMNH. INS.856456), same data; 1♂ same data except Nassau Mountains, Marowijne, Suriname, February 22, 1952.</p><p>Wyeomyia kerri . HOLOTYPE ♀ (CEIOC) with genitalia mounted on microscope slide: Cuiabá, Mato Grosso, Brazil, May 15, 1935, G. César coll.; PARATYPES: 1♂ (CMN 9867), Rio Itaucú, Curralinho, state of Pará, Brazil, near to roça of Antonio Pereira where monkey trap is located, larvae focus on açai palm holding water, December 13, 1935, H.W. Kumm coll.; 1♂ (CMN 9101) with genitalia mounted on microscope slide (CMN 1394), same data except, Alto Rio Canaticú, human bait behind house 10, rainy weather, between 06:00 and 06:30 a.m., May 06, 1936, A. Azevedo and H.W. Kumm coll.; 1♀ (CMN 4093) with genitalia mounted on microscope slide, Coronel Ponce, Campo Verde, state of Mato Grosso, Brazil, May 15, 1935, G. César coll .</p></div>	https://treatment.plazi.org/id/03A187D23401CF21FF4EFD617F0DFA9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lourenço-De-Oliveira, Ricardo;Pereira, Glauber Rocha;Neves, Maycon Sabastião Alberto Santos;Santos, Stephani Da Silva;Ribeiro, Paulino Siqueira;Nascimento-Pereira, Agostinho Cardoso;Talaga, Stanislas;Pavan, Márcio Galvão;Sallum, Maria Anice Mureb;Motta, Monique Albuquerque	Lourenço-De-Oliveira, Ricardo, Pereira, Glauber Rocha, Neves, Maycon Sabastião Alberto Santos, Santos, Stephani Da Silva, Ribeiro, Paulino Siqueira, Nascimento-Pereira, Agostinho Cardoso, Talaga, Stanislas, Pavan, Márcio Galvão, Sallum, Maria Anice Mureb, Motta, Monique Albuquerque (2026): Systematics of Leonidasdeanea, a new subgenus of Wyeomyia (Diptera: Culicidae), with recognition of Wy. luciae as a valid species and synonymy of Wy. surinamensis with Wy. flui. Zootaxa 5782 (1): 1-43, DOI: 10.11646/zootaxa.5782.1.1, URL: https://doi.org/10.11646/zootaxa.5782.1.1
03A187D2340DCF36FF4EFA117FFAFA3D.text	03A187D2340DCF36FF4EFA117FFAFA3D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wyeomyia (Leonidasdeanea) luciae Senevet, Chabelard & Abonnenc 1942	<div><p>Wyeomyia (Leonidasdeanea) luciae Senevet, Chabelard &amp; Abonnenc, 1942</p><p>luciae Senevet, Chabelard &amp; Abonnenc, 1942: 343–348 (A, ♂ G*, P*, L*, Lmp*). Holotype ♂ (genitalia mounted on a slide M1552 not M1553, see Belkin 1968). Type locality: Port Inini, Tonnegrande River, French Guiana (18 August 1939), E. Abonnenc coll. (see Talaga et al. 2020), MNHN (not FMP, see Matile 1976 and Degallier &amp; Claustre 1980). Stat. nov., resurrected from synonymy.</p><p>Wyeomyia (Dendromyia) luciae of Lane 1951: 335 (synonymy with chalcocephala, taxonomy); Horsfall 1955: 329, 719 (synonym of chalcocephala, distribution); Stone et al. 1959: 84, 346 (synonym of chalcocephala, taxonomy, type info.); Belkin et al. 1965: 23, 26, 94 (synonym of chalcocephala, habitat, type info.).</p><p>Wyeomyia luciae of Lane 1953: 977, 1107 (synonym of chalcocephala, taxonomy); Belkin 1968: 40, 53, 68 (synonymy with chalcocephala questioned, depository, taxonomy, type info.); Knight &amp; Stone 1977: 328, 587 (synonym of chalcocephala, taxonomy, type info.); Degallier &amp; Claustre 1980: 144 (depository); Guimarães 1997: 128, 269 (synonym of chalcocephala, classification, distribution, type info.); Talaga et al. 2015: 779, 780 (synonym of chalcocephala, nomenclature, species list French Guiana, taxonomy); Talaga et al. 2020: 362, 367, 373 (synonym of chalcocephala, nomenclature, list species described from French Guiana, type info., type loc. restriction); Harbach 2018: 30, 76 (synonym of chalcocephala, nomenclature); Wilkerson et al. 2021b: 1029, 1288 (synonym of chalcocephala, taxonomy, type info.).</p><p>Wyeomyia sp. 4 of Motta et al. 2007: 593–601, 605, 606, 608, 611–613, 616, 618, 619, 625, 626 (L, Lmp, P, ♀, ♀ G, ♂, ♂ G, classification, morphology, phylogeny, systematics).</p><p>Wyeomyia (subgenus uncertain) chalcocephala of Talaga et al. 2015: 776, 780 ( luciae as synonym, nomenclature, species list French Guiana, taxonomy).</p><p>Female. Head:As described for Wy.flui, except clypeus brown, rounded, pruinose, without scales and setae. Maxillary palpus short, slightly longer than clypeus, brown-scaled. Proboscis: Length approximately 1.9 mm (1.85–2.0 mm), expanded at apex; dorsal surface dark brown-scaled, ventral surface dark brown with line of white scales more evident from middle to distal 0.9; 8,9 elongate brown basal labial setae, progressively shorter laterally; labellum with pale integument. Antenna as described for Wy. flui . Thorax: Scutum with brown integument, covered with elongate brown scales with greenish-blue reflections; prescutellar area partially bare; 8–10 strong darkish brown prescutellar setae; anterior promontory with 9–12 darkish setae of different lengths and groups of white scales mesally and laterally, latter usually contiguous to concolor scaling on postpronotum; no acrostichal or dorsocentral setae. Supraalar and antealar areas with total of about 25 strong darkish brown setae. Scutellum covered with scales concolorous with scutal scales, limited to the lobes; median lobe with usually 4 long and 4,5 short darkish brown setae, lateral lobes with 5,6(5) long and 4–6 short darkish brown setae. Mesopostnotum, antepronotum, postpronotum and pleura as described for Wy. flui, except mesopostnotum with tuft of 8–11 yellowish to pale brown setae, and antepronotum with row of 11–14 darkish brown setae dorsally. Pleural chaetotaxy as follows: 2–4(3) yellowish prespiracular setae, long, extending beyond spiracle; postspiracular setae absent; 5–8(7) upper proepisternal setae, yellowish, mostly long; 6–9 lower mesokatepisternal setae, 2 setae above upper margin of mesomeron; upper mesokatepisternal setae absent; prealar area with 4,5 pale brown setae; lower mesepimeral setae absent; about 10–12 pale yellowish upper mesepimeral setae, nearly in a row. Halter: Integument light brown; scabellum and pedicel dark-scaled dorsally, capitellum dark-scaled. Wing: Length 4 mm (3.9–4.0 mm), scales brown with slight bluish reflections, spatulate with rounded apices on all veins. Dorsal and ventral scales as described for Wy. flui . Calypter with 1–3(2) brown setae. Alula with 10,11 simple pale brown setae. Legs: Coxae and trochanters pale, with pearly white scales and elongate yellowish setae. Femora, tibiae and tarsomeres 1 mainly dark-scaled; femora and tibiae white-scaled ventrally; foretarsomeres 1–5 dark-scaled; midtarsomere 1 dark-scaled; midtarsomeres 2–4 white-scaled dorsally; midtarsomere 2 usually more conspicuously white-scaled on distal 0.5–0.8 of dorsal surface, with white scaling consisting of a less evident line on basal 0.3, ventral surface of midtarsomeres 2–4 dark-scaled; midtarsomere 5 entirely dark-scaled, dorsal surface of midtarsomere 5 may be unusually invaded by few white scales on basal 0.2; hindtarsomeres 1 and 2 dark-scaled, hindtarsomere 2 sometimes with a few white scales scattered on ventral surface; hindtarsomeres 3 and 4 white-scaled on basal 0.9 of ventral surface; hindtarsomere 5 entirely white-scaled ventrally. Ungues simple. Abdomen: Terga covered with brown spatulate scales with bluish green reflections. Tergum I with numerous yellowish setae. Sterna covered with whitish scales; color on sides separated in nearly a straight line, sometimes with few whitish scales invading the brown-scaling on apical-lateral margins of terga II and III. Genitalia (Fig. 8). Similar to Wy. flui, except for tergum IX with 1 or 2 longer setae on each lobe, and insula with a group of 7 setae on each side of dorsal surface.</p><p>Male. Similar to female except for the following characteristics: Head: Antenna with flagellum more verticillate than female, except last 2 flagellomeres. Legs: Midtarsomeres 1–4 white-scaled dorsally in a continuous line; hindtarsomeres 1–4 white-scaled on at least basal 0.9 of ventral surface; white scaling on ventral surface of hindtarsomere 1 sometimes less conspicuous. Genitalia (Figs 9, 10). Tergum VIII with numerous setae distributed on half of anterior portion, mixed with scales and minute setae; setae in middle shorter than the others; rounded anterior corners, without lateral lobes or tuft of setae. Sternum VIII (Fig. 9H) with a row of setae on anterior margin, numerous scales distributed on anterior half. Tergum IX (Fig. 9C) with slightly straight interlobar space, lobes bearing 3 strong pointed setae with tips facing outward, similar in shape and length, median seta of each lobe inserted slightly more distant from the others. Sternum IX (Fig. 9E) fused to tergum, expanded in middle as a plate divided into 2 lobes. Gonocoxite (Fig. 9A,B) rounded, tapering to apex; tergal surface with numerous scales and setae; about 35 long tergomesal setae. Basal mesal lobe elongate, with about 13 short setae. Gonostylus (Fig. 9A) slightly shorter than gonocoxite with a curve stem gradually tapering to apex bearing 4 lobes (Fig. 10): lobe A conical, elongate, with a hook-shaped seta at apex (Fig. 10A,B); lobe E, adjacent to lobe A, narrow, bearing 9 strong setae (Fig. 10A,B); lobe M ovate with a small spout-like process on middle of distal margin (Fig. 10B–D), with 20 strong somewhat pointed setae on margin; lobe C resembles a narrow and backward curved arm (Fig. 10A,D). Aedeagus (Fig. 9F) rounded, submedian tergal arms not fused, apically digitiform; narrowed and weakly turned inward; median sternal plate membranous, with slightly pointed apex. Proctiger narrow; paraproct sclerotized dorsoapically, about 3 apical teeth, 4 minute cercal setae (Fig. 9D).</p><p>Pupa (Fig. 11). Positions and character of setae as figured, number of branches in Table 2. Cephalothorax: Integument light brown, darker on dorsum in area of trumpet. Seta 1-CT long, strong, sigmoid; seta 5-CT slightly aciculate, more than twice length of seta 7-CT; seta 10-CT very short, around 0.3 length of seta 11-CT; seta 11-CT slightly longer than seta 12-CT. Trumpet: Integument brown, cylindrical; index 5.1–6.4 (mean 5.7) (width measured at mid-length). Abdomen: Integument brownish, darker from metanotum to tergum IV. Seta 1-III–VII near posterior margin of tergum, lateral to seta 2-III–VII, 1-V,VI long, more than twice length of seta 3-V,VI, 2-III–VII far mesal to group of other dorsal setae; seta 3-V,VI inserted close to seta 1; seta 4-I short, about 0.3 length of seta 7-I, 4-VII almost as long as seta 3; seta 5-III shorter than seta 1, 5-IV–VI long, strong, slightly aciculate; seta 6-II longer than 3 times length of seta 3, 6-III–V similar in length to seta 4; seta 7-I more than half length of seta 6, 7-II dorsolateral in position; seta 8-VII as long as seta 7; seta 10-I minute, 10-V longer than on other terga; seta 11-I not present, 11- II–VII minute. Paddle slightly longer than wide, apex slightly extended on inner margin; index 1.4–1.8.</p><p>D = dendritic.</p><p>Larva, fourth-instar (Figs 12, 13). Body generally with pale reddish coloration. Position and character of setae as figured. Head: Slightly wider than long, light tan. Collar absent, occipital foramen with slit-like extension on either side. Hypostomal suture complete. Dorsomentum with 10–12(11) lateral teeth on either side of a slightly larger median tooth (Figs 12, 13A). Lateral teeth similar in development except for most lateral teeth that sometimes are very small. Maxilla (Fig. 13A, B): Rounded with a small protrusion on inner apical margin; apical tooth absent; maxillary brush long, nearly similar in length to seta 4-Mx; laciniarastrum 1 (LR1) with a row of about 13 teeth, progressively longer apically; LR2 third of maxillary body, forming a dense group of long setae close to LR1. Maxillary palpus not fused to maxillary body, with 3 short digitiform setae, 1 far from the others; seta 1-Mx single, stout, forked at apex, inserted in apical third; seta 2-Mx single, short; seta 4-Mx single, long, inserted on anterior margin; seta 6-Mx single. Mandible (Fig. 13B,C): Ventral mandibular tooth 0 (VT 0) prominent and larger than VT 1, VT 2 and VT 3; mandibular sweeper 2 with one group of 4–7 long setae with pectinate filaments. Antenna: Slender, slightly longer than seta 7-C; seta 1-A usually single, inserted on apical 0.25 of shaft. Seta 1-C single, elongate, nearly as long as setae 5–7-C, slightly curved, stout; seta 3-C single, minute; setae 4–13-C branched, seta 4 multiple, slightly shorter than seta 5; setae 5–7,11-C similar in length, 5-C usually double; seta 8-C slightly shorter than setae 9,12-C; seta 10-C much shorter than setae 8 and 9, almost as long as seta 14-C; setae 12,13-C nearly aligned; seta 14-C elongate, around 3 times length of seta 15-C, usually double; seta 15-C short, branched, inserted almost in line with seta 14. Thorax: Integument smooth. Setae 4-P, 5,6-M and 7,13-T inserted on individual pigmented basal plates; pleural groups 5–7-P and 9–12-P,M,T on common pigmented basal plates; seta 0-P small, branched; seta 1-P branched, nearly 0.2 length of seta 3-P, posterior in position and mesad to setae 2,3; seta 2-P usually double, almost in line with seta 3-P; seta 3-P single, longer than seta 2; seta 4-P multiple, aciculate, much longer than seta 3; setae 5,6-P usually single, similar in length; seta 7-P multi-branched, with branches of different lengths; setae 5‒7,9,12- P; 5,6,8,9,12-M and 7,9,12,13-T aciculate; seta 8-P branched, nearly 0.3 length of seta 3; seta 9-P branched; seta 10-P about 0.5 length of seta 12, usually single, slightly aciculate; seta 11-P,M,T single, spiniform, 11-P,M slender, 11-T stronger than 11-P,M; seta 12-P single, much longer than seta 9; seta 14-P multi-branched, much longer than seta 8, nearly similar in length to seta 1; setae 1–4-M essentially in line, 1,4 branched, almost similar in size; setae 2,3,5,6,10-M single, 3-M long, around 3 times length of seta 1; seta 7-M small, normally double; seta 8-M multi-branched, similar in length to seta 7; seta 9-M branched; 10-M single, about 0.7 length of seta 12; setae 13,14-M branched, 13-M slightly shorter than seta 14, seta 14 similar in development and length to seta 14-P, longer than seta 1; setae 1–3-T almost in line, branched, 1-T similar in development to seta 1-P,M, 2-T usually double, twice length of 3-T; seta 4-T usually double, laterad to setae1–3; setae 5,6-T branched, almost similar in length; seta 7-T multiple, almost similar in development and length to seta 9; seta 9-T multi-branched, slightly shorter than seta 12; setae 10,12-T single, 10-T similar in length to seta 10-P; seta 13-T multiple, nearly 0.2 length of seta 9. Abdomen: Seta 1- I–VII branched, 1-I,V mesad to seta 2, 1-II–IV,VI lateral and posterior in position to seta 2, 1-VII lateral and almost aligned with seta 2; seta 2-I,II branched, longer than seta 4, 2-III–VII usually single, very small; seta 3-I,II,IV,VI usually branched, almost similar in length, 3-III,V usually single, elongate, longer than setae 1,5, 3-VII strong, single, about 1.2 length of siphon, aciculate; seta 4-I,II,V,VI branched, 4-III,IV,VII usually single, 4-I,II,IV–VII lateral to seta 1, 4-III in line with seta 1, 4-VI elongate, about 3.5 length of seta 2; seta 5-I usually double, slightly shorter than seta 1, 5-II–VI branched, elongate, similar in length to seta 1, 5-VII elongate, more than 3.0 length of setae 8 and 9; setae 6,7-I,II branched, with branches of different lengths, aciculate, inserted on pigmented basal plates, 6-I longer than setae 7-T and 6-II, 6-III–VI long, double, aciculate, 6-III,IV more than 1.5 length of seta 6, 6-VII multiple, longer than setae 7,10,12; seta 7-III–VII small, branched; seta 8-II usually single, about same length as seta 5, 8-III–VI small, usually double, 8-VII small, multi-branched; seta 9-I–VII branched, small, similar in development, about same length or slightly shorter than seta 2; seta 10-I–VII small, branched, 10-I almost aligned with seta 13 and posterior in position to setae 9,11,13-I; seta 10-II,VI almost aligned and posterior to seta 11, 10- III–V lateral and posterior to setae 11,13; seta 11-I developed like seta 13, 11-II–VII small, shorter than seta 10; seta 12-II,VI,VII branched, 12-III–V usually single, 12-II–VI longer than seta11, 12-II,III,V small, almost aligned and posterior to seta 13, 12-IV,VII laterad to seta13; seta 13-I usually with 5 branches, anterior to setae 10 and 11, moderately developed, longer than seta 1; seta 13-II–V usually 3,4 branched, similar in length or slightly shorter than seta 1, 13-VI,VII small, branched, 13-III–VI much anterior to seta 12. Segment VIII: Comb composed of a group of about 20 elongate scales arranged in more than one uneven row, not borne on a plate. Setae 1,3-VIII branched; seta 1-VIII similar in length to seta 4, nearly 0.2 length of seta 2; seta 2-VIII usually single, similar in length to seta 5; seta 3-VIII multi-branched, small, nearly 0.2 length of and aligned with seta 4; seta 4-VIII usually single; seta 5-VIII usually double, similar in length to seta 2. Siphon: Elongate; index 2.7–3.2 (mean 3.0; mode 3.1) (width measured at base); tanned, more pigmented at base. Pecten composed of a row of 18–20 slender elongate filaments, in ventral median position, usually extending from 0.25‒0,75 from base of siphon; seta 1-S normally inserted at distal limit of basal one-seventh of siphon, with usually 3 aciculate branches; seta 1a-S usually with 2 midventral single setae on distal 0.25 of siphon; seta 2a-S almost similar in length to seta 1, normally with 4 elongate setae mostly distributed on basal half of siphon, with 1,2 slightly aciculate branches. Segment X: Saddle slightly darker than siphon; setae 1–3-X strong, aciculate; seta 1-X single, longer than setae 2–4; seta 2-X branched, 0.6 length of seta 3; seta 3-X normally double, about 0.8 length of seta 1; seta 4-X branched, much less strongly developed than setae 1–3, nearly 0.25 length of seta 1.</p><p>Bionomics. Larvae of Wy. luciae have been found in the leaf axils of buriti palm trees and balourous or sororoca plants ( Phenakospermum guyanense Endl.) (Senevet, et al. 1942; CCULI, http://cculi.fiocruz.br). We have frequently found Wy. luciae in buriti with Wy. flui and Wy. aphobema . Nothing is known about the bionomics of the adults.</p><p>Distribution. Wyeomyia luciae has been collected in French Guiana (Senevet et al. 1942) and Brazil (states of Maranhão, Mato Grosso, Rondônia, Roraima and Tocantins).</p><p>Material examined. 50 specimens (2L, 8♀, 1♀ Pe, 5♀ LePe, 3♀ G, 2♀ LePeG, 2LePe ♀, 1G ♀, 1♂, 4♂ LePe, 1♂ PeG, 12♂ LePeG, 7♂ G, 1LePeG ♂). HOLOTYPE ♂ (MNHN M1552), adult on a microscope slide under a glass cover, with associated LePe and dissected male genitalia on separate microscope slides, Port Inini, Tonnegrande River, French Guiana, August 18, 1939, reared from larva collected from leaf axils of balourous ( Heliconia sp.) (probably Phenakospermum guyanense) . BRAZIL: 1♀ LePe, 1♂ LePeG (CCULI 1007, 1012), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-63.02236&amp;materialsCitation.latitude=-9.908639" title="Search Plazi for locations around (long -63.02236/lat -9.908639)">Zoological Garden (Parque Botânico)</a> (9°54'31.1"S 63°01'20.5"W), Ariquemes, state of Rondônia, July 16, 1986, reared from larvae collected from leaf axils of buriti palm trees, R. Lourenço-de-Oliveir coll., R. Lourenço-de-Oliveira, M.G. Castro and M.A. Motta det. ; 1♂ G, 1♀ (CCULI1346-1347), same data except M.G. Castro coll.; 2L, 6♀, 2♀ G, 1♀ Pe, 4♀ LePe, 1♀ LePeG, 2LePe ♀, 1G ♀, 4♂ G, 1♂ PeG, 4♂ LePe, 8♂ LePeG, 1LePeG ♂ (CCULI6247–6275, 6277–6282), same data except April, 18–26, 1994, R. Lourenço-de-Oliveira coll., R. Lourenço-de-Oliveira and M.A. Motta det.; 1♀ LePeG (CCULI 6276), same data except Guajará Mirim, Km 7, September, 21 1994, reared from larvae collected from leaf axils of a sororoca plant ( Phenakospermum guyanense), D. Couto-Lima coll. ; 1♀, 1♀ G, 1♂ G (CCULI 6283, 6284, 6302), Chapada dos Guimarães National Park, Chapada dos Guimarães, state of Mato Grosso, May 9, 2018, reared from larvae collected from leaf axils of buriti palm trees, A.C. Nascimento-Pereira coll., A.C. Nascimento-Pereira det. ; 1♂ G (CCULI 5636), same data except M.A. Motta det.; 1♂ LePeG, 1♂ (CCULI 6303, 6304), same data except, Bom Jardim, Pastos Bons, state of Maranhão, January 27, 2021, A.C. Nascimento-Pereira coll., A.C. Nascimento-Pereira det. ; 1♂ LePeG (CCULI 4959), same data, except <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-63.02236&amp;materialsCitation.latitude=-9.908639" title="Search Plazi for locations around (long -63.02236/lat -9.908639)">Chácara do Betinho</a> (vicinal 26), São João da Baliza, state of Roraima, July 25, 2018, A.C. Nascimento-Pereira and F. Maciel coll.</p></div>	https://treatment.plazi.org/id/03A187D2340DCF36FF4EFA117FFAFA3D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lourenço-De-Oliveira, Ricardo;Pereira, Glauber Rocha;Neves, Maycon Sabastião Alberto Santos;Santos, Stephani Da Silva;Ribeiro, Paulino Siqueira;Nascimento-Pereira, Agostinho Cardoso;Talaga, Stanislas;Pavan, Márcio Galvão;Sallum, Maria Anice Mureb;Motta, Monique Albuquerque	Lourenço-De-Oliveira, Ricardo, Pereira, Glauber Rocha, Neves, Maycon Sabastião Alberto Santos, Santos, Stephani Da Silva, Ribeiro, Paulino Siqueira, Nascimento-Pereira, Agostinho Cardoso, Talaga, Stanislas, Pavan, Márcio Galvão, Sallum, Maria Anice Mureb, Motta, Monique Albuquerque (2026): Systematics of Leonidasdeanea, a new subgenus of Wyeomyia (Diptera: Culicidae), with recognition of Wy. luciae as a valid species and synonymy of Wy. surinamensis with Wy. flui. Zootaxa 5782 (1): 1-43, DOI: 10.11646/zootaxa.5782.1.1, URL: https://doi.org/10.11646/zootaxa.5782.1.1
03A187D2341ACF0EFF4EF98A79FEFE1B.text	03A187D2341ACF0EFF4EF98A79FEFE1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wyeomyia (Leonidasdeanea) chalcocephala Dyar & Knab 1906	<div><p>Wyeomyia (Leonidasdeanea) chalcocephala Dyar &amp; Knab, 1906</p><p>chalcocephala Dyar &amp; Knab, 1906a: 140. Holotype ♂ (Cat. no. 10.005), with genitalia mounted on slide no. 352: Cacao, Trece Aguas, Alta Vera Paz, Guatemala, Schwarz &amp; Barber coll. (USNM).</p><p>Wyeomyia chalcocephala of Theobald 1910: 624, 646 (record Guatemala); Howard et al. 1913: iii, vi, pls 5, 92 (♂ G*, L*); Howard et al. 1915: 55–57, 145, 146 (♀, ♂, ♂ G, L, bionomics, classification, distribution, key Central and North America, systematics, type info.); Del Ponte &amp; Cerqueira 1938: 231, 232 (♀, systematics); Arnett 1949: 244, 249 (bionomics, habitat, coinhabiting species, ecology); Arnett 1950: 111 (bionomics, habitat); Galindo et al. 1950: 549 (bionomics, vertical stratification); Stone &amp; Knight 1957: 121, 122 (type info.); Fauran &amp; Pajot 1974: 147 (♂ G, taxonomy); Talaga et al. 2020: 362, 367 (nomenclature, taxonomy).</p><p>Wyeomyia galoa in part of Howard et al. 1913: iii, pl. 4 (male only; ♂ G*); Howard et al. 1915: 56, 128 (male only; ♂ G, bionomics, key Central and North America, systematics).</p><p>Heliconiamyia galoa in part of Dyar 1919: 123, 124, pl. 5, fig. 5 (male only; ♂ G*, Wy. galoa as type of Heliconiamyia, bionomics, classification, distribution, key, systematic).</p><p>Heliconiamyia chalcocephala of Dyar 1919: 123, 124 (♀, ♂ G, bionomics, classification, distribution, key, systematic).</p><p>Wyeomyia (Heliconiamyia) galoa in part of Dyar 1923: 64, 65 (male only; determination of the male galoa as chalcocephala, taxonomy).</p><p>Wyeomyia (Heliconiamyia) chalcocephala of Dyar 1923: 64, 65 (♀, ♂, bionomics); Dyar 1928: 68–70, 77, 78, 488 (fig. 47), 605 (♀, ♂, ♂ G*, L*, bionomics, classification, distribution, key New World, taxonomy); Dyar &amp; Shannon 1924a: 85, 86, 91 (L, bionomics, habitat, cohabiting species, key, record Panama); Bonne &amp; Bonne-Wepster 1925: 59, 85, 549 (A, ♂ G, L, chalcocephala as type of Heliconiamyia, bionomics, classification, distribution, key, review, taxonomy).</p><p>Prosopolepis (Heliconiamyia) chalcocephala of Dyar &amp; Shannon 1924b: 482 (classification, taxonomy); Dyar 1925: 118–120, 122 (bionomics, classification, distribution, key, species list Panama).</p><p>Wyeomyia (Dendromyia) chalcocephala of Edwards 1932: 87, 227 (classification, distribution, taxonomy); Lane 1939: 140, 141, 212 (classification, distribution); Lane 1945: 146, 147, 149 (♂ G, taxonomy); Lane 1951: 335 ( luciae as synonym, taxonomy); Galindo et al. 1951: 131 (bionomics, habitat); Stone et al. 1959: 84, 336 (distribution, taxonomy, type info.); Belkin et al. 1965: 28, 94 (bionomics, habitat, type info.); Forattini 1965: 162, 416 (in part, only from Panama; bionomics, vertical stratification); Belkin 1968: 40, 66 (synonymy, taxonomy); Barreto &amp; Lee 1969: 417 (bait catch, bionomics, record Colombia, survey); Knight &amp; Stone 1977: 328, 569 (in part, only specimens from Colombia, Guatemala and Panama; classification, distribution, taxonomy, type info.); Clark-Gil &amp; Darsie 1983: 156, 182, 199, 217, 231, 270, 282 (bionomics, habitat, co-inhabiting species, distribution, key, species list Guatemala); Darsie 1994: 513 (species list Guatemala).</p><p>Dendromyia chalcocephala of Del Ponte 1939: 540 (♀, identification method, taxonomy).</p><p>Wyeomyia (Dendromyia) chalcocephala of Lane &amp; Cerqueira 1942: 539, 540, 547, 589, 607, 845 (in part, only specimens from Guatemala and Panama, ♀, bionomics, habitat, classification, distribution, key, revision, systematics); Lane 1953: 869, 870, 875, 877, 977–979, 1101 (♀, ♂, ♂ G*, L, classification, distribution, key, taxonomy, type depository); Knight 1978: 65, 97 (distribution).</p><p>Wyeomyia (Wyeomyia) chalcocephala of Horsfall 1955: 327, 719 (in part, only specimens from Guatemala, habitat, distribution); Linton et al. 2013: suppl. data 2 (species list Ecuador).</p><p>Wyeomyia (subgenus D) chalcocephala of Heinemann &amp; Belkin 1977: 435, 436, 443, 447, 450, 454 (bionomics, habitat, coinhabiting species, distribution, species list Honduras and Nicaragua); Heinemann &amp; Belkin 1978a: 124–126, 128– 133, 135, 136, 141, 149–152, 189, 193, 196 (bionomics, habitat, coinhabiting species, distribution, species list Panama); Heinemann &amp; Belkin 1979: 99, 100, 102, 109 (bionomics, habitat, coinhabiting species, distribution, record Ecuador, species list Ecuador).</p><p>Wyeomyia (subgenus D) ? chalcocephala of Heinemann &amp; Belkin 1978c: 513, 514, 526 (bionomics, habitat, distribution, tentative specific determination, species list Colombia).</p><p>Wyeomyia (subgenus uncertain) chalcocephala of Motta &amp; Lourenço-de-Oliveira 1995: 384 (classification, systematics); Guimarães 1997: 128, 269 (in part, only specimens from Colombia, Guatemala, Honduras, Panama, classification, distribution, type info.); Peyton et al. 1999: 550 (depository, distribution); Pecor et al. 2002: 246, 250, 269, 276 (L, bionomics, habitat, distribution, record Belize, species list Belize); Motta et al. 2007: 593–601, 605, 611–613, 616, 618, 619 (L, Lmp, P, ♀, ♀ G, ♂, ♂ G, classification, morphology, phylogeny, systematics); Rozo-Lopez &amp; Mengual 2015: 30 (species list Colombia); Ramón et al. 2018: 196 (species list Ecuador); Harbach 2018: 30, 76 (nomenclature, synonym); Harbach 2026 (classification, taxonomy); Nascimento-Pereira et al. 2019: 360 (classification); Wilkerson et al. 2021b: 1029, 1273 (classification, distribution, etymology, informal name, taxonomy, type info.); Naranjo-Díaz et al. 2022: 9, suppl. 1 (distribution, species list Colombia); Viveros-Santos et al. 2022: 123, 127, 130, 133 (distribution Chiapas State, Mexico, ecology, record Mexico, seasonality).</p><p>Female. Head: Occiput and vertex with brown scales, vertex with spot of whitish reflections; narrow interocular space, ocular line with brown scales; gena covered with pale scales. Clypeus and antenna as described for Wy. luciae . Maxillary palpus brown-scaled, length about 0.25 of proboscis. Proboscis straight, length 2.53 mm, completely brown-scaled, slightly expanded on distal 0.25, labella well developed. Thorax: Scutum elongate, covered by brown scales with slightly blue-greenish reflections, prescutellar area with pronounced depression. Scutellum covered with scales concolorous with scutal scales. Supraalar areas with about 10 long and 5 short setae. Mesopostnotum, antepronotum, postpronotum and pleura as described for Wy. flui, except mesopostnotum with 2 decumbent pearly white diaphanous spatulate scales anterior to tuft of 2 short setae (4 insertions). Pleural chaetotaxy as follows: 5 long, brownish upper proepisternal; 4 well developed, brown prespiracular; 7 long, yellowish lower mesokatepisternal; 12 yellowish upper mesepimeral; 3 brown prealar. Mesopostnotum brown with 2 decumbent pearly white diaphanous spatulate scales anterior to tuft of 2 short setae (4 insertions). Wing: Length 4.0 mm. Dorsal and ventral scales as described for Wy. flui; alula with 8 piliform setae; upper calypter with 2 long, brown setae, Legs: Femora and tibiae with a line of whitish scales ventrally; foretarsomeres dark-scaled; midtarsomeres with white scales from apical half of 3 to 5, midtarsomere 3 with white scales starting as a thin line dorsally, midtarsomeres 4 and 5 entirely white-scaled dorsally. Hindtarsomeres 4 and 5 white-scaled ventrally. Abdomen: Terga brown-scaled, sterna covered by creamy white scales. Lateral demarcation between dark and pale scales, slightly in line.</p><p>Male. Similar to female except for the following characteristics: antenna with flagellum more verticillate than female, except last 2 flagellomeres; mid- and hindtarsomeres 1–5 white-scaled on ventral and dorsal surface, respectively, with white scaling more conspicuous on tarsomeres 3–5. G enitalia (Fig.14): Tergum VIII with numerous setae distributed on anterior half, mixed with scales and minute setae; setae shorter in middle and laterally, rounded anterior corners, without lateral lobes or tuft of setae. Sternum VIII with a row of setae (insertions in Fig. 14I) on anterior margin, numerous scales and minute setae on anterior half, uniformly distributed. Tergum IX (Fig. 14G) with slightly straight interlobar space, lobes well separated each bearing 4 strong pointed setae similar in shape and length with tips slightly bent outwardly. Sternum IX (Fig. 14G) fused to tergum, with a deep median emargination separating 2 elongate, roughly digitiform lobes with minute setae. Gonocoxite rounded, tapering to apex; tergal surface with numerous scales and short setae apically on margin; tergomesal setae long and numerous, about 12. Basal mesal lobe roughly triangular with about 15 slender setae (Fig. 14C). Gonostylus (Figs 14, 15) similar in length to gonocoxite, with a narrow and curve stem and apex divided into four lobes (Fig. 15): lobe A digitiform, narrow and elongate, with a strong seta with apical curved tip (Fig. 15B); lobe E adjacent to lobe A, large, with a row of 11 modified setae on margin, arranged in ascending order of size toward outer margin, first 3 are fine, next 4 stout and short, last 4 are elongate; a row of 5 minute mesal setae (Fig. 15A,B); lobe M (Figs 14B, 15C,D) expanded with a membranous process, mesal margin with a row of 13 setae of different shapes and lengths; lobe C narrow and elongated like a curved arm (Fig. 15B). Aedeagus (Fig. 14D,E) rounded, submedian tergal arms not fused, elongate, apically digitiform; narrowed and weakly turned inward; median sternal plate membranous, folded at edge, like a tight collar at apex. Proctiger narrow; paraproct sclerotized dorsoapically, about 7 small apical teeth; 3 minute cercal setae (Fig. 14F).</p><p>Pupa (Fig. 16). Positions and character of setae as figured, number of branches in Table 3. Cephalothorax: Seta 1-CT long, strong, sigmoid; seta 5-CT slightly aciculate, usually triple, about twice length of seta 7-CT; seta 10-CT slightly more than half length of seta 11-CT; seta 10-I not found; seta 11-I–III minute. Trumpet: Integument brown, cylindrical. Abdomen: Integument brownish. Seta 1-III–VII near to posterior margin of tergum, lateral to seta 2, 1- IV–VII nearly similar in length to seta 3; seta 2-II–VII mesal, 3-VII short; seta 4-VI usually triple, more than half length of seta 6, 4-VII much longer than seta 6; seta 5-II much longer than seta 2, 5-IV–VI slightly aciculate; seta 6-I nearly twice length of seta 7; seta 7-II dorsal in position, 7-VII usually single; seta 8-VII much shorter than seta 7. Paddle much longer than wide, apex essentially truncate, slightly emarginate at apex of well-marked midrib.</p><p>D = dendritic.</p><p>Larva, fourth-instar (Fig. 17). Position and character of setae as figured. Head: Wider than long. Collar absent, occipital foramen slit-like extension on either side. Hypostomal suture complete. Dorsomentum with 9–11(9) lateral teeth on either side of a slightly larger median tooth. Lateral teeth similar in development, except most lateral teeth sometimes very small. Antenna: Slender, slightly longer than seta 7-C; seta 1-A single, inserted on apical 0.25 of shaft. Seta 1-C single, slightly curved stout; seta 3-C single, minute; setae 4,6-C similar in length, multi-branched; seta 5-C much shorter than setae 4,6, branched; seta 7-C longer than setae 4,6, multi-branched; setae 8–10,12-C short, similar in length, branched; seta 11-C almost similar in length seta seta 7, multi-branched; seta 14-C elongate, 4 times longer than seta 15, usually double; seta 15-C short, branched, inserted much posterior to seta 14-C. Thorax: Integument smooth. Setae 4-P and 7,13-T inserted on individual pigmented basal plates; pleural groups 5–7-P and 9– 12-P,M,T on common pigmented basal plates; seta 0-P small, branched; seta 1-P branched, mesad to seta 2,3, longer than and slightly posterior to seta 3 and in line with seta 2; seta 2-P long, about twice length of seta 3, single; seta 4-P with multiple branches; setae 5,6-P usually single, similar in length; seta 7-P with multiple branches of different sizes; setae 8,14-P small, branched; seta 8-P about half length of seta 1; setae 4,5–7,9,10,12-P, 5,6,9,10,12-M and 7,9,10,12,13-T aciculate; seta 9-P with branches of different lengths, munch longer than seta 12; seta 10-P about 0.6 length of seta 12; seta 11-P,M,T single, spiniform; seta 11-T more strongly developed than seta 11-P,M; seta 12-P single; setae 2,3,5,6,12-M single; setae 1–4-M almost in line, 1,4-M branched, 3-M long, about 4.0 length of seta 1; seta 7-M small, normally with 3 branches; seta 8-M elongate, similar in size to seta 9; seta 9-M branched, almost 2.0 length of seta 10-M; seta 10-M usually double, about 0.4 length of seta 12-M; seta 13-M branched, small, about 0.4 length of seta 14; seta 14-M multiple, slightly longer than seta 1; setae 1–3,5-T almost in line, 2-T much longer than setae 3–6, 5-T branched; seta 6-T branched, shorter than seta 5; seta 7-T shorter than seta 9; seta 10-T usually double; seta 12-T single, longer than seta 9; seta 13-T multiple. Abdomen: Seta 1-I–IV,VII branched, posterior to seta 2, 1-I mesad to seta 2, 1-II–IV slightly lateral to or aligned with seta 2, 1-VII aciculate, nearly same length as seta 4; seta 2-I–IV,VII small, 2-I–IV branched, mesad to seta 3,4, 2-VII single; setae 3,4-I–IV,VI branched; seta 3-I similar in length to seta 4, 3-II–IV much longer than seta 4, 3-VII, single, about 1.1 length of siphon, aciculate; seta 4-I–IV branched, lateral to setae 1,2, 4-I nearly in line with and 0.7 length of seta 1, 4-II posterior to seta 1, 4-III,IV anterior to seta1, 4-VII usually single, similar in length to seta 1, in line with seta 3; seta 5-I–IV,VII branched, 5-I small, about 0.5 length of seta 1, 5-II–IV longer than or similar in length to seta 1, 5-VII very small, similar in length to setae 8,9; seta 6-I–VI branched, with branches of different lengths, inserted on pigmented basal plates, aciculate, 6-I longer than seta 7, shorter than 6-II, 6-VII branched, small, longer than setae 7,10,12; seta 7-I,II branched, aciculate, inserted on pigmented basal plates; seta 7-III,VII small, branched; seta 8-II usually single, about same length as seta 5, 8-III,IV,VII small, branched; seta 9-I–IV with 1,2 branches, small, about same length or slightly longer than seta 2, 9-VII branched, small; seta 10-I–IV,VII small, branched, 10-I almost aligned with seta 13, in line with setae 9,11, 10-II–IV lateral and posterior in position to setae 11,13; seta 11-I–IV,VII small, branched, 11-I about 0.2 length of seta 10, 11-II–IV shorter than seta10; seta 12-II–IV,VII branched, small, 12-II,III almost aligned with seta 13, 12-II,IV about 2.0 length of seta 11, 12-III similar in length to seta 11; seta 13-I–IV branched, anterior to setae 10–12-I–IV, 13-I about 1.5 length of seta 1, 13-II branched, slightly longer than seta1, 13-III,IV branched, similar in size to seta 1, 13-VII small, lateral to setae 10,12. Segment VIII: Comb composed of a group of about 20 elongate scales not borne on a plate. Setae 1,3–5-VIII branched; seta 1-VIII multi-branched, much longer than seta 3; seta 3-VIII 0.2 length of seta 4, almost aligned with seta 4; seta 4-VIII similar in length to seta 5, posterior to setae 1–3,5, usually double; seta 5-VIII distinctly anterior to seta 4. Siphon: Moderately long; index 2.5 (width measured at base); tanned, more pigmented at the base. Pecten composed of a row of 6–10 slender elongate filaments, usually limited to mid-length of siphon; seta 1-S normally inserted near distal limit of basal 0.25 of siphon, with 4 branches, slightly aciculate; seta 1a-S usually with 1 midventral double seta at distal 0.25 of siphon; seta 2a-S much shorter than seta 1S, normally with 3 elongate setae mostly distributed on basal half of siphon, with 2,3 slightly aciculate branches. Segment X: Saddle slightly more tanned than siphon; setae 1,3-X single, aciculate, similar in length; setae 2,4-X branched; seta 2-X almost 0.2 length of seta 3; seta 4-X not well developed, about 0.2 length of seta 1.</p><p>Bionomics. Females of Wy. chalcocephala have been reported feeding on humans during daytime, especially near the forest floor (Galindo et al. 1950, 1951; Barreto &amp; Lee 1969). Larvae of Wy. chalcocephala have been found in water in the leaf axils and flower bracts of Heliconia and axils of banana trees (Dyar 1919, 1923, 1928; Dyar &amp; Shannon 1924a; Arnett 1949; Heinemann &amp; Belkin 1977; Clark-Gil &amp; Darsie 1983), and also in the leaf axils of plants of the family Araceae (skunk cabbage, elephant ear, dumb-cane, taro) (Arnett 1949; Heinemann &amp; Belkin 1977; Pecor et al. 2002). In Araceae, Wy. chalcocephala was reported to be found with Johnbelkinia longipes (Fabricius, 1805), Jb. ulopus (Dyar &amp; Knab, 1906a), Trichoprosopon pallidiventer (Lutz, 1905a), Limatus durhamii Theobald, 1901b, Wy. melanocephala Dyar &amp; Knab, 1906a, Wy. ( Hystatomyia) sp., Wy. (Dendromyia) complosa (Dyar, 1928), Wy. (Wyeomyia) melanopus Dyar, 1919, Wy. (Wyo.) celaenocephala Dyar &amp; Knab, 1906a, the Wy. (Wyo.) pertinans (Williston, 1896) group, Toxorhynchites (Lynchiella) superbus (Dyar &amp; Knab, 1906b), while in Heliconia larvae were found with Wy. (Decamyia) pseudopecten Dyar &amp; Knab, 1906a and Wy. ( Dec .) ulocoma (Theobald, 1903), and in banana plants with Wy. celaenocephala .</p><p>Distribution. Wyeomyia chalcocephala has been reported to occur in Belize (Pecor et al. 2002), Colombia (Barreto &amp; Lee 1969)], Costa Rica (Wilkerson et al. 2021b), Ecuador (Heinemann &amp; Belkin 1979), Guatemala (Dyar &amp; Knab 1906a), Honduras (Heinemann &amp; Belkin 1977), Mexico (Viveros-Santos et al. 2022), Nicaragua (Heinemann &amp; Belkin 1977), Panama (Galindo et al. 1951; Dyar &amp; Shannon 1924a; Heinemann &amp; Belkin 1978a). The records from Brazil and French Guiana (Lane 1936b; Lane &amp; Cerqueira 1942; Deane et al. 1953; Stone et al. 1959; Cerqueira 1961; Xavier &amp; Mattos 1970, 1975; Forattini et al. 1970; Knight &amp; Stone 1977; Guimarães 1997; Hutchings et al. 2005a; Reis et al. 2010; Wilkerson et al. 2021b) need confirmation. In fact, we have never encountered this species in our numerous mosquito collections made throughout Brazil for over 40 years, and we have found that none of the specimens from Brazil currently identified as Wy. chalcocephala in the CMN belong to this species.</p><p>Material examined. 9 specimens (1♀ PeG, 2♀ Pe, 1♀ LePe, 1♂ G, 2♂ LePe, 2G ♂). HOLOTYPE ♂ (NMNH 10.002) with associated dissected male genitalia on microscope slide (no. 352): Cacao Trece Aguas, Alta Vera Paz, Guatemala, without data, “bred from flower-bract of Heliconia ” (Dyar 1923: 65), Schwarz and Barber coll. PANAMA: 1♀ PeG, 2♀ Pe, 1♀ LePe (NMNH 196-112, 196-113, 196-116, 196-118), Bocas del Toro, about 3 km S of Punta de Pena, El Guabo (17PLV6981), 60 m, April 14, 1963, Heliconia flower bracts in second growth, partial shade, A. Quinonez coll.; 2♂ LePe (NMNH: PA258-108, PA297-108), Close’s Plantation, Cano Saddle, Canal Zone, R.C. Shannon coll.; 1G ♂ (NMNH: 1866), same date. HONDURAS: 1G ♂ (CNM 7a), Tela, June 29, 1924, without date. Specimens originally identified as Wy. galoa: 1G ♂ (NMNH 357), Cacao Trece Aguas, Alta Vera Paz, Guatemala, without data, Schwarz and Barber coll.</p></div>	https://treatment.plazi.org/id/03A187D2341ACF0EFF4EF98A79FEFE1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lourenço-De-Oliveira, Ricardo;Pereira, Glauber Rocha;Neves, Maycon Sabastião Alberto Santos;Santos, Stephani Da Silva;Ribeiro, Paulino Siqueira;Nascimento-Pereira, Agostinho Cardoso;Talaga, Stanislas;Pavan, Márcio Galvão;Sallum, Maria Anice Mureb;Motta, Monique Albuquerque	Lourenço-De-Oliveira, Ricardo, Pereira, Glauber Rocha, Neves, Maycon Sabastião Alberto Santos, Santos, Stephani Da Silva, Ribeiro, Paulino Siqueira, Nascimento-Pereira, Agostinho Cardoso, Talaga, Stanislas, Pavan, Márcio Galvão, Sallum, Maria Anice Mureb, Motta, Monique Albuquerque (2026): Systematics of Leonidasdeanea, a new subgenus of Wyeomyia (Diptera: Culicidae), with recognition of Wy. luciae as a valid species and synonymy of Wy. surinamensis with Wy. flui. Zootaxa 5782 (1): 1-43, DOI: 10.11646/zootaxa.5782.1.1, URL: https://doi.org/10.11646/zootaxa.5782.1.1
