identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A587CFFFB0FF84FC59F873FD50F93A.text	03A587CFFFB0FF84FC59F873FD50F93A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotinus lahomius (Kheil 1884) Šumpich & Weidenhoffer 2025	<div><p>Allotinus lahomius (Kheil, 1884)</p><p>comb. nov. &amp; stat. restit.</p><p>(Fig. 1)</p><p>Miletus lahomius Kheil, 1884: 27–28, figs 28–29 (original description). Logania regina lahomius: FRUHSTORÞER (1914a): 23 (new combination). Logania lahomius: FRUHSTORÞER (1916a): 806 (new combination).</p><p>Logania marmorata lahomius: CORBET (1940a):112; ELlOT (1986):4, 66 (redescription, distribution, key to the subspecies of L. marmorata); BRlDGES (1988): VII.249, IX.65 (catalogue).</p><p>Type material examined. SYNTYPES (5 JJ 5 ♀♀, NMPC): 1 J, ‘Typo [hw] // Nias [p] // Orig. [p, pink label] // Typus [p, red label] // Coll. N. Kheil [p] // [small yellow label without text] // Photo / Gen. prep. / J. Šumpich / 22/157 [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype J / Miletus lahomius / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’ (Fig. 1A).</p><p>1 J, ‘Typo [hw] // Nias [p] // Orig. [p, pink label] // Typus [p, red label] // Coll. N. Kheil [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype J / Miletus lahomius / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’ (without abdomen).</p><p>3 JJ, ‘Nias [p] // Coll. N. Kheil [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype J / Miletus lahomius / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’ (one male without abdomen).</p><p>5 ♀♀, ‘Nias [p] // Coll.N. Kheil [p] // ex coll. N.M.Kheil / Coll.National / Museum Prague [p] // Syntype ♀ / Miletus lahomius / Kheil, 1884 / labelled by J. Šumpich / &amp; Z.Weidenhoffer,2025 [p, red label]’. One female with the label ‘Photo / Gen. prep. / J. Šumpich / 22/158 [p]’ (Fig. 1B).</p><p>Current status. Allotinus lahomius (Kheil, 1884) comb. nov., valid species (see Remark).</p><p>Distribution. Indonesia: Nias (KHElL 1884),? Sumatra (ELlOT 1986).</p><p>Remarks. Classification in the genus Allotinus C. &amp; R. Felder, [1865] follows SEKl et al. (1991).</p><p>The description of Logania marmorata Moore, 1884 was published on 30 June 1884 (cf. title pages of Journal of the Asiatic Society of Bengal, Vol. LIII). Kheil’s work was published in 1884 without further specification of the publication date, only the author’s preface is dated at the end of February 1884. However, in the same year SCHlLDE (1884) wrote a review of KHElL’ S (1884) paper, which is dated June 1884. It is obvious that J. Schilde had to read Kheil’s work before F. Moore’s paper was published. For this reason, Miletus lahomius Kheil, 1884 has precedence over Logania marmorata Moore, 1884, which becomes a junior subjective synonym of the former, though valid in the subspecies rank. Due to this synonymy the change of the nominotypical subspecies is required (ICZN 1999: 47.2), affecting all its existing subspecies according to ELlOT (1986) (see the list bellow):</p><p>Allotinus lahomius lahomius (Kheil, 1884)</p><p>Allotinus lahomius damis (Fruhstorfer, 1914), stat. nov.</p><p>Allotinus lahomius diehli (Eliot, 1986), stat. nov.</p><p>Allotinus lahomius faustina (Fruhstorfer, 1914), stat. nov. Allotinus lahomius hilaeira (Fruhstorfer, 1914), stat. nov. Allotinus lahomius javanica (Fruhstorfer, 1914), stat. nov. Allotinus lahomius marmorata (Moore, 1884), stat. nov.</p><p>Allotinus lahomius munichya (Fruhstorfer, 1914), stat. nov. Allotinus lahomius palawana (Fruhstorfer, 1914), stat. nov. Allotinus lahomius samosata (Fruhstorfer, 1914), stat. nov.</p></div>	https://treatment.plazi.org/id/03A587CFFFB0FF84FC59F873FD50F93A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Šumpich, Jan;Weidenhoffer, Zdeněk	Šumpich, Jan, Weidenhoffer, Zdeněk (2025): A review of the types of Lycaenidae species (Lepidoptera) described by Napoleon Manuel Kheil deposited in the collection of the National Museum of the Czech Republic. Acta Entomologica Musei Nationalis Pragae 65 (1): 177-195, DOI: 10.37520/aemnp.2025.014, URL: https://doi.org/10.37520/aemnp.2025.014
03A587CFFFB3FF83FF0BF9B7FD28F8CD.text	03A587CFFFB3FF83FF0BF9B7FD28F8CD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotinus unicolor subsp. aphocha (Kheil 1884)	<div><p>Allotinus unicolor aphocha (Kheil, 1884)</p><p>(Figs 2–3)</p><p>Allotinus aphocha Kheil, 1884: 28, fig. 30 (original description).</p><p>Allotinus aphocha: FRUHSTORÞER (1916a): 810 (checklist).</p><p>Paragerydus aphocha: DRUCE (1895): 563 (new combination).</p><p>Allotinus (Paragerydus) unicolor aphocha: CORBET (1939): 70 (new combination), ELlOT (1986): 48 (taxonomic review).</p><p>Type material examined. SYNTYPES (5 JJ 6 ♀♀; NMPC):5 JJ, ‘Nias [p] // Coll. N. Kheil [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype J / Allotinus aphocha / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’. Tree specimens were photographed or dissected and have label containing:‘Photo / Gen. prep. / J. Šumpich [p]’as prefix for number of a photo or slide made by J. Šumpich and following numbers: 22155, 22180 and 22181 (Figs 2A–B).</p><p>1 ♀, ‘Typo [hw] // Nias [p] // Orig. [p, pink label] // Typus [p, red label] // Coll. N. Kheil [p] // [small yellow label without text] // Photo / Gen. prep. / J. Šumpich / 22/157 [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype ♀ / Allotinus aphocha / Kheil, 1884 / labelled by J. Šumpich / &amp; Z.Weidenhoffer, 2025 [p, red label]’ (Fig.2C).</p><p>5 ♀♀, ‘Nias [p] // Coll. N.Kheil [p] // ex coll.N. M. Kheil / Coll. National / Museum Prague [p] // Syntype ♀ / Allotinus aphocha / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’. Three specimens were photographed or dissected and have label containing ‘Photo / Gen. prep. / J. Šumpich [p]’as prefix for number of a photo or slide made by J. Šumpich and following numbers: 22156, 22182 (Fig. 2D) and 22183.</p><p>Additional material examined. INDONESIA: [NIAS]: without faunistic details, 2 ♀♀, coll. N. M. Kheil (NMPC) (Fig. 2E); without faunistic details, 2 ♀♀, coll. J. Šrogl (NMPC).</p><p>Comparative material examined. Allotinus unicolor rekkia Riley &amp; Godfrey, 1921: THAILAND: NAKHON RATCHASIMA PROVINCE: Wang Nam Khio, 14°28′N, 101°55′E, 550 m, 3.xii.2012, 2 JJ 3 ♀♀, Z. Weidenhoffer leg. (ZWPC).</p><p>Current status. Allotinus unicolor aphocha (Kheil, 1884), subspecies of Allotinus unicolor C. &amp; R. Felder, [1865].</p><p>Distribution. Indonesia: Nias (KHElL 1884),?Bangka Island,?Batu Island,?Mentawai Islands,? Sumatra Island (ELlOT 1986).</p><p>Remarks. ELlOT (1986) pointed out in his review paper that it is very difficult to decide which specimens constitute a valid subspecies because of the great range of phenetic variation. He accepted subspecific status based on the study of large series of butterflies stored in BMNH, but highlighted the difficulty or impossibility of identifying a butterfly to a subspecies if the specimen does not bear a locality label. ELlOT (1986) recognised a total of eight subspecies of A. unicolor, writing for the subspecies aphocha: ‘Differs from subsp. unicolor only in that on the underside of the hindwing the postdiscal spot in space 6 is usually placed closer to the spot in space 7. Many specimens are inseparable from unicolor, so the subspecies is of doubtful validity’. However, the text in ELlOT’ s (1986) work indicated that he did not have the opportunity to study the type material of A. unicolor aphocha . For this reason, the distribution of this subspecies as defined by ELlOT (1986) requires future revision.</p><p>Allotinus unicolor unicolor was described from Singapore (FELDER &amp; FELDER [1865]), and according to INAYOSHl (1996–2024), it is distributed on the Malay Peninsula from Singapore in the south to Ranong province in southern Thailand; according to ELlOT (1986) also to south Myanmar. North from Ranong across Myanmar to Assam, the subspecies A. unicolor continentalis Fruhstorfer, 1913 occurs and in northeastern Thailand, Cambodia and southern Vietnam, there is the subspecies A. unicolor rekkia Riley &amp; Godfrey, 1921 (cf. ELlOT 1986, INAYOSHl 1996–2024).</p><p>Allotinus unicolor aphocha was described from Nias and is currently treated as the oldest subspecies of A. unicolor except the nominotypical one. We have for comparison only collection material from central Thailand (Nakhon Ratchasima Province), which we provisionally assign to the subspecies A. unicolor rekkia . However, more specimens of all three subspecies mentioned above are depicted by INA- YOSHl (1996–2024). Obviously, distinguishing these three subspecies is not easy and in some cases it is impossible. It seems also impossible to point out reliable distinguishing characters in the pattern on the underside of the wings. However, the A. unicolor aphocha differs quite markedly from these three subspecies, and therefore we continue to consider this taxon a valid subspecies of A. unicolor . Well visible differences are especially on the upper side of the forewings in females which have reddish tint and instead of a more pronounced discoidal spot, the centre of the wing is only slightly lighter. Other characters, such as the more markedly serrated hindwing margin or the size and position of the small spots on the underside of the wings, are variable and are not reliable. The distinctly elongated distal part of the great lobe of uncus of the male genitalia may also be characteristic for this subspecies (Figs 3A, B).</p><p>KHElL (1884) published one pair of A. unicolor from Nias. There are two other females without locality label in Kheil’s collection in NMPC, one of which is identified as ‘ unicolor Felder’ (Fig. 2E). These two females are not included in the syntype series of A. unicolor aphocha, but most probably belong to this subspecies.</p><p>Tribe Spalgini Toxopeus, 1929</p></div>	https://treatment.plazi.org/id/03A587CFFFB3FF83FF0BF9B7FD28F8CD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Šumpich, Jan;Weidenhoffer, Zdeněk	Šumpich, Jan, Weidenhoffer, Zdeněk (2025): A review of the types of Lycaenidae species (Lepidoptera) described by Napoleon Manuel Kheil deposited in the collection of the National Museum of the Czech Republic. Acta Entomologica Musei Nationalis Pragae 65 (1): 177-195, DOI: 10.37520/aemnp.2025.014, URL: https://doi.org/10.37520/aemnp.2025.014
03A587CFFFB4FF83FEECF80FFAE3F8EC.text	03A587CFFFB4FF83FEECF80FFAE3F8EC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spalgis epeus subsp. fangola (Kheil 1884)	<div><p>Spalgis epeus fangola (Kheil, 1884)</p><p>(Fig. 4)</p><p>Lucia fangola Kheil, 1884: 28, fig. 31 (original description).</p><p>Spalgis epius [sic] fangola: FRUHSTORÞER (1917a):75 (new combination). Spalgis epeus fangola: BRlDGES (1988): IX.118 (catalogue).</p><p>Type material examined. HOLOTYPE: J (NMPC), ‘ Typo [hw] // Nias [p] // Orig. [p, pink label] // Typus [p, red label] // Coll. N. Kheil [p] //</p><p>Photo / Gen. prep. / J. Šumpich / 22/169 [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Holotype J / Lucia fangola / Kheil, 1884 / labelled by J. Šumpich / &amp; Z.Weidenhoffer, 2025 [p, red label]’ (Fig.4A).</p><p>Current status. Spalgis epeus fangola (Kheil, 1884), subspecies of S. epeus (Westwood, [1851]) .</p><p>Distribution. Indonesia: Nias (KHElL 1884), Sumatra (FRUHSTORÞER 1917a).</p><p>Remarks. FRUHSTORÞER (1917a) placed fangola as a subspecies of S. epeus based on its extremely small size and the very fine net pattern on the underside of the wings.</p><p>Subfamily Polyommatinae Swainson, 1827</p><p>Tribe Polyommatini Swainson, 1827</p></div>	https://treatment.plazi.org/id/03A587CFFFB4FF83FEECF80FFAE3F8EC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Šumpich, Jan;Weidenhoffer, Zdeněk	Šumpich, Jan, Weidenhoffer, Zdeněk (2025): A review of the types of Lycaenidae species (Lepidoptera) described by Napoleon Manuel Kheil deposited in the collection of the National Museum of the Czech Republic. Acta Entomologica Musei Nationalis Pragae 65 (1): 177-195, DOI: 10.37520/aemnp.2025.014, URL: https://doi.org/10.37520/aemnp.2025.014
03A587CFFFB4FF80FC29F86FFE88F84B.text	03A587CFFFB4FF80FC29F86FFE88F84B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Jamides talinga (Kheil 1884)	<div><p>Jamides talinga (Kheil, 1884)</p><p>(Fig. 5)</p><p>Plebeius talinga Kheil, 1884: 29, figs 32–33 (original description). Lampides talinga: DE NlCḖVlLLE (1895): 39 (new combination). Lampides kondulana talinga: FRUHSTORÞER (1916a): 10 (new status). Jamides talinga talinga: HlROWATARl (1986): 126 (new combination).</p><p>Type material examined. SYNTYPES (1 J 2 ♀♀, NMPC): 1 J, ‘Nias [p] // Orig. [p, pink label] // Typus [p, red label] // Coll. N. Kheil [p]</p><p>// [small yellow label without text] // Photo / Gen. prep. / J. Šumpich / 22/163 [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype J / Plebeius talinga / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’ (Fig. 5A).</p><p>1 ♀, ‘Typo [hw] // Nias [p] // Orig. [p, pink label] // Typus [p, red label] // Coll. N. Kheil [p] // Photo / Gen. prep. / J. Šumpich / 22/164 [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype ♀ / Plebeius talinga / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’ (Fig. 5B).</p><p>1 ♀, ‘Nias [p] // Coll. N. Kheil [p] // [small yellow label without text] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype ♀ / Plebeius talinga / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’.</p><p>Current status. Jamides talinga (Kheil, 1884), valid species.</p><p>Distribution. Nias (KHElL 1884), Sumatra (DE NlCḖVlLLE (1895), Malaysia (RlLEY &amp; CORBET 1938), Thailand (Trang and southward), Langkawi, Borneo (INAYOSHl 1996–2024).</p></div>	https://treatment.plazi.org/id/03A587CFFFB4FF80FC29F86FFE88F84B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Šumpich, Jan;Weidenhoffer, Zdeněk	Šumpich, Jan, Weidenhoffer, Zdeněk (2025): A review of the types of Lycaenidae species (Lepidoptera) described by Napoleon Manuel Kheil deposited in the collection of the National Museum of the Czech Republic. Acta Entomologica Musei Nationalis Pragae 65 (1): 177-195, DOI: 10.37520/aemnp.2025.014, URL: https://doi.org/10.37520/aemnp.2025.014
03A587CFFFB7FF80FC70FADFFC27F7C5.text	03A587CFFFB7FF80FC70FADFFC27F7C5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Jamides bochus subsp. siraha (Kheil 1884)	<div><p>Jamides bochus siraha (Kheil, 1884)</p><p>(Fig. 6)</p><p>Plebeius siraha Kheil, 1884: 30, fig. 35 (original description).</p><p>Lampides bochus siraha: FRUHSTORÞER (1916a): 36 (new combination). Jamides bochus siraha: RlLEY &amp; CORBET (1938): 150 (taxonomic revision); D’ ABRERA (1986): 643 (taxonomic review).</p><p>Type material examined. SYNTYPES (2 JJ 1 ♀, NMPC): 1 J, ‘Nias [p] // Orig. [p, pink label] // Typus [p, red label] // Coll. N. Kheil [p] // [small yellow label without text] // Photo / Gen. prep. / J. Šumpich / 22/160 [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype J / Plebeius siraha / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’.</p><p>1 J, ‘Nias [p] // Coll. N. Kheil [p] // Photo / Gen. prep. / J. Šumpich / 22/161 [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype J / Plebeius siraha / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’ (Fig. 6A).</p><p>1 ♀, ‘Nias [p] // Coll. N. Kheil [p] /// Photo / Gen. prep. / J. Šumpich / 22/162 [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype ♀ / Plebeius siraha / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’ (Fig. 6B).</p><p>Current status. Jamides bochus siraha (Kheil, 1884), subspecies of Jamides bochus (Stoll, [1782]) .</p><p>Distribution. Nias (KHElL 1884), Sumatra (FRUHSTORÞER 1916a).</p></div>	https://treatment.plazi.org/id/03A587CFFFB7FF80FC70FADFFC27F7C5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Šumpich, Jan;Weidenhoffer, Zdeněk	Šumpich, Jan, Weidenhoffer, Zdeněk (2025): A review of the types of Lycaenidae species (Lepidoptera) described by Napoleon Manuel Kheil deposited in the collection of the National Museum of the Czech Republic. Acta Entomologica Musei Nationalis Pragae 65 (1): 177-195, DOI: 10.37520/aemnp.2025.014, URL: https://doi.org/10.37520/aemnp.2025.014
03A587CFFFB8FF8FFEE0F95AFA34F7E1.text	03A587CFFFB8FF8FFEE0F95AFA34F7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Prosotas nora subsp. kupu (Kheil 1884)	<div><p>Prosotas nora kupu (Kheil, 1884)</p><p>(Fig. 7)</p><p>Plebeius kupu Kheil, 1884: 29, fig. 34 (original description). Nacaduba nora kupu: FRUHSTORÞER (1916b): 117 (new status). Prosotas nora kupu: TlTE (1963): 92 (new combination).</p><p>Type material examined. HOLOTYPE: ♀ (NMPC), ‘Typo [p, pink label] // Nias [p] // Typus [p, red label] // Coll. N. Kheil [p] // Photo / Gen. prep. / J. Šumpich / 22/169 [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype ♀ / Plebeius kupu / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’ (Fig. 7A).</p><p>Current status. Prosotas nora kupu (Kheil, 1884), subspecies of Prosotas nora (Felder, 1860) .</p><p>Distribution. Prosotas nora is widely distributed in Southeast Asia, from Pakistan to Papua New Guinea, also in Australia (BRABY 2004, LEPlÞORUM 2024). The known distribution of the subspecies P.n. kupu to date is restricted to the island of Nias.</p><p>Remarks. TlTE (1963) in his work on this species noted: ‘Type? A specimen from the Fruhstorfer collection in the B.M. (N.H.) is labelled in Fruhstorfer’s writing as the type’, but the description of the species was based on a single specimen (KHElL 1884) and the holotype is deposited in the NMPC.</p><p>The male genitalia of several subspecies of P. nora were depicted by FRUHSTORÞER (1916b) and CORBET &amp; PENDLE- BURY (1992) but not the female genitalia. The genitalia of the subspecies P. n. kupu have not yet been depicted. The holotype of P. nora kupu is a female. Comparisons with female genitalia of related taxa could not be made because comparative material was not available.According to FRUH- STORÞER (1916b), the subspecies P. n. kupu differs from the related subspecies P. n. ardates (Moore, [1875]) only in its lighter colour. For the moment, we accept the taxonomic status of the subspecies but we cannot exclude that P.n. kupu may be synonymous with some other subspecies of P. nora .</p></div>	https://treatment.plazi.org/id/03A587CFFFB8FF8FFEE0F95AFA34F7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Šumpich, Jan;Weidenhoffer, Zdeněk	Šumpich, Jan, Weidenhoffer, Zdeněk (2025): A review of the types of Lycaenidae species (Lepidoptera) described by Napoleon Manuel Kheil deposited in the collection of the National Museum of the Czech Republic. Acta Entomologica Musei Nationalis Pragae 65 (1): 177-195, DOI: 10.37520/aemnp.2025.014, URL: https://doi.org/10.37520/aemnp.2025.014
03A587CFFFBBFF8CFF05FADEFAEDFD4C.text	03A587CFFFBBFF8CFF05FADEFAEDFD4C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acytolepis puspa subsp. puspinus (Kheil 1884)	<div><p>Acytolepis puspa puspinus (Kheil, 1884)</p><p>(Fig. 9)</p><p>Plebeius puspinus Kheil, 1884: 30, figs 37–39 (original description).</p><p>Cyaniris puspinus: FRUHSTORÞER (1910): 287 (new combination).</p><p>Lycaenopsis puspa puspinus: FRUHSTORÞER (1917b): 26 (new combination).</p><p>Acytolepis puspa puspinus: ELlOT &amp; KAWAZOḖ (1983): 179 (new combination).</p><p>Acytolepis puspa puspinus: SAVELLA (2024) (taxonomic review).</p><p>Type material examined. SYNTYPES (8 JJ 8 ♀♀, NMPC): 1 J, ‘Nias [p] // Coll. N. Kheil [p] // Typus [p, red label] // Orig. [p, pink label] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype J / Plebeius puspinus / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label] // Photo / Gen. prep. / J. Šumpich / 22/165 [p]’ (Fig. 9A).</p><p>1 J, ‘Nias [p] // Coll. N. Kheil [p] // Typus [p, red label] // Orig. [p, pink label] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype J / Plebeius puspinus / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’.</p><p>6 JJ, ‘Nias [p] // Coll. N. Kheil [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype J / Plebeius puspinus / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’.</p><p>1 ♀, ‘Nias [p] // Coll. N. Kheil [p] // Typus [p, red label] // Orig. [p, pink label] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype ♀ / Plebeius puspinus / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label] // Photo / Gen. prep. / J. Šumpich / 22/169 [p]’ (Fig. 9B).</p><p>7 ♀♀, ‘Nias [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype ♀ / Plebeius puspinus / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’. One female has very small green label without text.</p><p>Current status. Acytolepis puspa puspinus (Kheil, 1884), subspecies of Acytolepis puspa (Horsfield, [1828]) .</p><p>Distribution. Acytolepis puspa is widely distributed in Southeast Asia (LEPlÞORUM 2024); the occurrence of A. p. puspinus has been reported only from the island of Nias so far (FRUHSTORÞER 1917b).</p><p>Subfamily Theclinae Swainson, 1831</p><p>Tribe Hypolycaenini Swinhoe, 1910</p></div>	https://treatment.plazi.org/id/03A587CFFFBBFF8CFF05FADEFAEDFD4C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Šumpich, Jan;Weidenhoffer, Zdeněk	Šumpich, Jan, Weidenhoffer, Zdeněk (2025): A review of the types of Lycaenidae species (Lepidoptera) described by Napoleon Manuel Kheil deposited in the collection of the National Museum of the Czech Republic. Acta Entomologica Musei Nationalis Pragae 65 (1): 177-195, DOI: 10.37520/aemnp.2025.014, URL: https://doi.org/10.37520/aemnp.2025.014
03A587CFFFBBFF8CFC37FD8EFB34FAF1.text	03A587CFFFBBFF8CFC37FD8EFB34FAF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chliaria tora (Kheil 1884)	<div><p>Chliaria tora (Kheil, 1884)</p><p>(Fig. 10)</p><p>Hypolycaena tora Kheil, 1884: 31, fig. 40 (original description).</p><p>Chliaria amabilis tora: SElTZ (1927): 981 (new combination).</p><p>Chliaria tora: CORBET &amp; PENDLEBURY (1956): 354 (new combination); BECCALONl et al. (2003) (checklist); SAVELLA (2024) (taxonomic review).</p><p>Hypolycaena tora tora: D’ ABRERA (1986): 620 (taxonomic review).</p><p>Type material examined. SYNTYPES (1 J 2 ♀♀, NMPC): 1 J, ‘Nias [p] // Coll. N. Kheil [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype J / Hypolycaena tora / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label] // Photo / Gen. prep. / J. Šumpich / 22/149 [p]’ (Fig. 10A).</p><p>1 ♀, ‘Nias [p] // Coll. N. Kheil [p] // Orig. [p, pink label] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype ♀ / Hypolycaena tora / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label] // Photo / Gen. prep. / J. Šumpich / 22/154 [p]’ (Fig. 10B).</p><p>1 ♀, ‘Nias [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype ♀ / Hypolycaena tora / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’.</p></div>	https://treatment.plazi.org/id/03A587CFFFBBFF8CFC37FD8EFB34FAF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Šumpich, Jan;Weidenhoffer, Zdeněk	Šumpich, Jan, Weidenhoffer, Zdeněk (2025): A review of the types of Lycaenidae species (Lepidoptera) described by Napoleon Manuel Kheil deposited in the collection of the National Museum of the Czech Republic. Acta Entomologica Musei Nationalis Pragae 65 (1): 177-195, DOI: 10.37520/aemnp.2025.014, URL: https://doi.org/10.37520/aemnp.2025.014
03A587CFFFBBFF8CFF2AFFE6FE6AFAA7.text	03A587CFFFBBFF8CFF2AFFE6FE6AFAA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cupido lacturnus subsp. polysperchinus (Kheil 1884)	<div><p>Cupido lacturnus polysperchinus (Kheil, 1884) (Fig. 8)</p><p>Plebeius polysperchinus Kheil, 1884: 29, fig. 36 (original description). Everes polysperchinus: SElTZ (1912–1927): 924 (new combination; Fruhstorfer given wrongly instead of Kheil as the author of the name polysperchinus).</p><p>Elkalyce lacturnus polysperchinus: BECCALONl et al. (2003) (new combination).</p><p>Cupido lacturnus: SAVELLA (2024) (new synonymum,new combination).</p><p>Type material examined. SYNTYPES (8 JJ 3 ♀♀, NMPC):1 J, ‘Nias [p] // Coll. N. Kheil [p] // Typus [p, red label] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype J / Plebeius polysperchinus / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label] // Photo / Gen. prep. / J. Šumpich / 22/170 [p]’ (Fig. 8A).</p><p>7 JJ, ‘Nias [p] // Coll.N. Kheil [p] // ex coll. N.M.Kheil / Coll.National / Museum Prague [p] // Syntype J / Plebeius polysperchinus / Kheil,1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’. One specimen was dissected and has label containing ‘Photo / Gen. prep. / J. Šumpich / 24/082 [p]’.</p><p>1 ♀, ‘Nias [p] // Coll. N. Kheil [p] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype ♀ / Plebeius polysperchinus / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’.</p><p>1 ♀, ‘Nias [p] // Coll. N. Kheil [p] // [small white label with illegible character] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype ♀ / Plebeius polysperchinus / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’ (Fig. 8B).</p><p>1 ♀, ‘Nias [p] // Coll. N. Kheil [p] // [small yellow label without text] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Syntype ♀ / Plebeius polysperchinus / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label]’.</p><p>Current status. Cupido lacturnus polysperchinus (Kheil, 1884), subspecies of Cupido lacturnus (Godart, [1824]) .</p><p>Distribution. Cupido lacturnus is widely distributed in the Oriental Region, Australia and also in southern Japan. Subspecies C. l. polysperchinus is hitherto known only from Nias.</p><p>Remarks. We accept subspecies status for the taxon C. l. polysperchinus according to BECCALONl et al. (2003), but in generic combination in Cupido Schrank, 1801 (SAVELLA 2024, Z. Fric, pers. comm.).</p></div>	https://treatment.plazi.org/id/03A587CFFFBBFF8CFF2AFFE6FE6AFAA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Šumpich, Jan;Weidenhoffer, Zdeněk	Šumpich, Jan, Weidenhoffer, Zdeněk (2025): A review of the types of Lycaenidae species (Lepidoptera) described by Napoleon Manuel Kheil deposited in the collection of the National Museum of the Czech Republic. Acta Entomologica Musei Nationalis Pragae 65 (1): 177-195, DOI: 10.37520/aemnp.2025.014, URL: https://doi.org/10.37520/aemnp.2025.014
03A587CFFFBFFF89FC7BF80EFBF6F998.text	03A587CFFFBFFF89FC7BF80EFBF6F998.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tajuria bagus (Kheil 1884)	<div><p>Tajuria bagus (Kheil, 1884) stat. restit.</p><p>(Fig. 11)</p><p>Jolaus bagus Kheil, 1884: 31, fig. 41 (original description).</p><p>Tajuria cippus bagus: FRUHSTORÞER (1912): 211 (new combination).</p><p>Tajuria cippus bagas [sic!]: SElTZ (1912–1927): 972 (taxonomic review); D’ABRERA (1986): 612 (taxonomic review); SAVELLA (2024) (taxonomic review).</p><p>Iolaus bagus: BECCALONl et al. (2003) (checklist).</p><p>Type material examined. HOLOTYPE: ♀ (NMPC), ‘Nias [p] // Coll. N. Kheil [p] // Typus [p, red label] // Orig. [p, pink label] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Holotype ♀ / Jolaus bagus / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label] // Photo / Gen. prep. / J. Šumpich / 22/167 [p]’ (Figs 11A–C).</p><p>Current status. Tajuria bagus (Kheil, 1884), stat. restit., valid species.</p><p>Distribution. Indonesia: Nias (KHElL 1884).</p><p>Remarks. Tajuria bagus belongs to the least known taxa described by N. M. Kheil. The holotype has not yet been studied by anyone, which resulted in both misidentifications of some Indonesian material in other collections (e.g., WlKlSPEClES 2019), but also its still unclear taxonomic status. FRUHSTORÞER (1912) considered bagus a subspecies of Tajuria cippus (Fabricius, 1798) but only based on the pattern of the underside of the wings presented in Kheil’s publication. Unfortunately, in print of Kheil’s publication, the yellow paint on hindwings merged with the black, creating an artifact that greatly complicated the correct identification of this species (KHElL 1884: fig. 41; Fig. 11C). The pattern on the underside of the wings of T. bagus is more similar to Pratapa deva (Moore, [1858]) than to T. cippus, but the appearance of the upper side is different from both named species and most resembles T. isaeus amardus Fruhstorfer, 1912, also described from Nias (there are 14 specimens of both sexes in Kheil’s collection, cf. KHElL 1884). Due to its considerable differences from all nearby species, we consider T. bagus to be a good species.</p><p>In Kheil’s work (KHElL 1884), only the underside of the wings is depicted (Fig. 11C), where the repair of the right hindwing is visible at higher magnification, while the other hindwing appears to be in its original, undamaged condition. For more details compare the photos of the holotype in Fig. 11A with Fig. 11B, where the arrows point to the repaired parts of the wings.</p></div>	https://treatment.plazi.org/id/03A587CFFFBFFF89FC7BF80EFBF6F998	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Šumpich, Jan;Weidenhoffer, Zdeněk	Šumpich, Jan, Weidenhoffer, Zdeněk (2025): A review of the types of Lycaenidae species (Lepidoptera) described by Napoleon Manuel Kheil deposited in the collection of the National Museum of the Czech Republic. Acta Entomologica Musei Nationalis Pragae 65 (1): 177-195, DOI: 10.37520/aemnp.2025.014, URL: https://doi.org/10.37520/aemnp.2025.014
03A587CFFFBEFF96FC91F9DBFA20FB69.text	03A587CFFFBEFF96FC91F9DBFA20FB69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deudorix calderon Kheil 1884	<div><p>Deudorix calderon Kheil, 1884, stat. restit.</p><p>(Fig. 12)</p><p>Deudorix calderon Kheil, 1884: 33, fig. 25 (original description).</p><p>Deudorix calderon: FRUHSTORÞER (1912): 264 (taxonomic review); BRlDGES (1988): VIII.87 (catalogue).</p><p>Deudorix staudingeri calderon: SElTZ (1912–1927): 998 (new combination).</p><p>Type material examined. HOLOTYPE: J (NMPC), ‘Nias [p] // Coll. N. Kheil [p] // Orig. [p, pink label] // ex coll. N. M. Kheil / Coll. National / Museum Prague [p] // Holotype J / Deudorix calderon / Kheil, 1884 / labelled by J. Šumpich / &amp; Z. Weidenhoffer, 2025 [p, red label] // Photo / Gen. prep. / J. Šumpich / 22/167 [p]’ (Fig. 12A).</p><p>Current status. Deudorix calderon Kheil, 1884, stat. restit., valid species.</p><p>Distribution. Indonesia (Nias) (KHElL 1884), Philippines (Palawan) (SCHRÖDER &amp; TREADAWAY 2013), Myanmar, Thailand, Laos, China (Yunnan, Hainan) (INAYOSHl 1996–2024).</p><p>Remarks. KHElL (1884) did not depict the underside of the holotype wings, which resulted in misplacement of the species as a subspecies of D. staudingeri Druce, 1895 by SElTZ (1912–1927). Paradoxically, D. staudingeri is 11 years younger than D. calderon . SAVELLA (2024) accepts this combination with a question mark; however, this mistake was already recognized by BRlDGES (1988) and perhaps that is why he listed D. calderon as a good species. SEKl et al. (1991) pointed out that D. calderon is likely conspecific with D. hypargyria (Elwes, 1893), but similarly as in the previous case, Kheil’s description is older (!). Unfortunately, we do not have comparative material of D. hypargyria, but adults of that taxon were depicted by ELWES (1893) and CORBET &amp; PENDLEBURY (1992); the upper and underside of this species, and live butterflies from nature as well are depicted also by INAYOSHl (1996–2024). We see no difference in external appearance between D. hypargyria and the holotype of D. calderon, and especially in photos of live males from Thailand, the orange end of the abdomen, which is typical of D. calderon, is also very visible. We therefore have a good reason to follow the opinion of SEKl et al. (1991) and consider D. hypargyria to be a subspecies of D. calderon .</p><p>Deudorix strephanus Druce, 1896 was described as a separate species from Mt. Kinabalu, Borneo. In the original paper, the number of specimens of the type series is not specified and the description appears to have been based on a single male; therefore, TAKANAKl (1989) designated a lectotype, instead of holotype. SAVELLA (2024) considers strephanus to be a good species, but SEKl et al. (1991) classify it as a subspecies of D. hypargyria and point out a possible conspecificity with D. calderon . We agree with the opinion of SEKl et al. (1991) and transfer D. hypargyria strephanus as the subspecies of D. calderon .</p><p>Deudorix hypargyria annawarneckae Schröder &amp; Treadaway, 2013, described from the Palawan island in the Philippines, has so far been the only subspecies of D. hypargyria (SCHRÖDER &amp; TREADAWAY 2013, SAVELLA 2024). As a result of the change in taxonomic status of D. hypargyria, D. h. annawarneckae should be transferred to the subspecies level of D. calderon .</p><p>Updated list of subspecies of Deudorix calderon:</p><p>Deudorix calderon calderon Kheil, 1884, stat. restit.</p><p>Deudorix calderon annawarneckae (Schröder &amp; Treadaway, 2013), stat. nov.</p><p>Deudorix calderon hypargyria (Elwes, 1893), stat. nov.</p><p>Deudorix calderon strephanus (Druce, 1896), stat. nov.</p></div>	https://treatment.plazi.org/id/03A587CFFFBEFF96FC91F9DBFA20FB69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Šumpich, Jan;Weidenhoffer, Zdeněk	Šumpich, Jan, Weidenhoffer, Zdeněk (2025): A review of the types of Lycaenidae species (Lepidoptera) described by Napoleon Manuel Kheil deposited in the collection of the National Museum of the Czech Republic. Acta Entomologica Musei Nationalis Pragae 65 (1): 177-195, DOI: 10.37520/aemnp.2025.014, URL: https://doi.org/10.37520/aemnp.2025.014
