taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03A687A66006FFF2FF00FF3BFCD8FF43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326618/files/figure.png	https://doi.org/10.5281/zenodo.17326618	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66006FFF2FF00FF3BFCD8FF43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66006FFF2FF00FF3BFCD8FF43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66007FFF1FF00FE8AFE9BFB7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66007FFF1FF00FE8AFE9BFB7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66007FFF1FF00FE8AFE9BFB7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66004FFF7FF00FA96FE5EFCAE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66004FFF7FF00FA96FE5EFCAE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326622/files/figure.png	https://doi.org/10.5281/zenodo.17326622	FIGURE 5. Cochlis degregorioi nov. sp. A1–A4. Paratype, NHMW 1855/0045/0785b, Grund (Austria). B1–B4. Holotype, NHMW 1855/0045/0785a, Grund (Austria). C1–C4. Paratype, NHMW 1855/0045/0785c, Grund (Austria). D1–D4. Paratype, NHMW 1855/0045/0785d, Grund (Austria).	FIGURE 5. Cochlis degregorioi nov. sp. A1–A4. Paratype, NHMW 1855/0045/0785b, Grund (Austria). B1–B4. Holotype, NHMW 1855/0045/0785a, Grund (Austria). C1–C4. Paratype, NHMW 1855/0045/0785c, Grund (Austria). D1–D4. Paratype, NHMW 1855/0045/0785d, Grund (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66004FFF7FF00FA96FE5EFCAE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66004FFF7FF00FA96FE5EFCAE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66002FFEBFF00FC26FBE5FC16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326624/files/figure.png	https://doi.org/10.5281/zenodo.17326624	FIGURE 6. A1–A4. Cochlis eximia (Eichwald, 1830), SPSU 3/459, lectotype, Zalistsi, Ukraine, Middle Miocene, late Badenian. B1–B4. Cochlis eximia (Eichwald, 1830), SPSU 3/455, paralectotype, Zhukivtsi, Ukraine, Middle Miocene, late Badenian. C1–C4. Cochlis eximia (Eichwald, 1830), PIN 5904/119, Varivtsi, Ukraine, Middle Miocene, late Badenian. D1–D4. Cochlis eximia (Eichwald, 1830), 5904/118, Varivtsi, Ukraine, Middle Miocene, late Badenian.	FIGURE 6. A1–A4. Cochlis eximia (Eichwald, 1830), SPSU 3/459, lectotype, Zalistsi, Ukraine, Middle Miocene, late Badenian. B1–B4. Cochlis eximia (Eichwald, 1830), SPSU 3/455, paralectotype, Zhukivtsi, Ukraine, Middle Miocene, late Badenian. C1–C4. Cochlis eximia (Eichwald, 1830), PIN 5904/119, Varivtsi, Ukraine, Middle Miocene, late Badenian. D1–D4. Cochlis eximia (Eichwald, 1830), 5904/118, Varivtsi, Ukraine, Middle Miocene, late Badenian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66002FFEBFF00FC26FBE5FC16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326626/files/figure.png	https://doi.org/10.5281/zenodo.17326626	FIGURE 7. Cochlis eximia (Eichwald, 1830). A1–A4. Cochlis eximia (Eichwald, 1830), PIN 4450/136, Privol’nyi, Russia, Middle Miocene, late Konkian. B1–B4. Cochlis eximia (Eichwald, 1830), PIN 4450/137, Privol’nyi, Russia, Middle Miocene, late Konkian. C1–C4. Cochlis eximia (Eichwald, 1830), PIN 4450/138, well Molakuduk, Kazakhstan, Middle Miocene, late Konkian. D1–D4. Holotype of Cochlis odovychenorum Pedriali, Sosso & Dell’Angelo, 2019, MPUM 11725, Varivtsi (Ukraine).	FIGURE 7. Cochlis eximia (Eichwald, 1830). A1–A4. Cochlis eximia (Eichwald, 1830), PIN 4450/136, Privol’nyi, Russia, Middle Miocene, late Konkian. B1–B4. Cochlis eximia (Eichwald, 1830), PIN 4450/137, Privol’nyi, Russia, Middle Miocene, late Konkian. C1–C4. Cochlis eximia (Eichwald, 1830), PIN 4450/138, well Molakuduk, Kazakhstan, Middle Miocene, late Konkian. D1–D4. Holotype of Cochlis odovychenorum Pedriali, Sosso & Dell’Angelo, 2019, MPUM 11725, Varivtsi (Ukraine).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66002FFEBFF00FC26FBE5FC16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326628/files/figure.png	https://doi.org/10.5281/zenodo.17326628	FIGURE 8. A1–A2. Cochlis eximia (Eichwald, 1830), NHMW 2024/0231/0002, Weitendorf (Austria). B1–B5. Cochlis eximia (Eichwald, 1830), private collection Franz Jiranek, Weitendorf (Austria). C1–C2.? Cochlis operculum, SPSU 3/456, syntype of Natica eximia Eichwald, 1830, Zhukivtsi, Ukraine, Middle Miocene, late Badenian. D. Cochlis eximia (Eichwald, 1830), 5904/120, Varivtsi, Ukraine, Middle Miocene, late Badenian.	FIGURE 8. A1–A2. Cochlis eximia (Eichwald, 1830), NHMW 2024/0231/0002, Weitendorf (Austria). B1–B5. Cochlis eximia (Eichwald, 1830), private collection Franz Jiranek, Weitendorf (Austria). C1–C2.? Cochlis operculum, SPSU 3/456, syntype of Natica eximia Eichwald, 1830, Zhukivtsi, Ukraine, Middle Miocene, late Badenian. D. Cochlis eximia (Eichwald, 1830), 5904/120, Varivtsi, Ukraine, Middle Miocene, late Badenian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66002FFEBFF00FC26FBE5FC16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66002FFEBFF00FC26FBE5FC16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66002FFEBFF00FC26FBE5FC16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326684/files/figure.png	https://doi.org/10.5281/zenodo.17326684	FIGURE 32. A1–A4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/129, Ripyntsi, Ukraine, Middle Miocene, late Badenian. B1–B4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/130, Yaseniv, Ukraine, Middle Miocene, late Badenian. C1–C4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/131, Ripyntsi, Ukraine, Middle Miocene, late Badenian. D1–D4. Pseudolinices staszici (Friedberg, 1923), SPSU 3/458, syntype of Natica eximia Eichwald, 1830, Zalistsi, Ukraine, Middle Miocene, late Badenian.	FIGURE 32. A1–A4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/129, Ripyntsi, Ukraine, Middle Miocene, late Badenian. B1–B4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/130, Yaseniv, Ukraine, Middle Miocene, late Badenian. C1–C4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/131, Ripyntsi, Ukraine, Middle Miocene, late Badenian. D1–D4. Pseudolinices staszici (Friedberg, 1923), SPSU 3/458, syntype of Natica eximia Eichwald, 1830, Zalistsi, Ukraine, Middle Miocene, late Badenian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66002FFEBFF00FC26FBE5FC16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66002FFEBFF00FC26FBE5FC16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326622/files/figure.png	https://doi.org/10.5281/zenodo.17326622	FIGURE 5. Cochlis degregorioi nov. sp. A1–A4. Paratype, NHMW 1855/0045/0785b, Grund (Austria). B1–B4. Holotype, NHMW 1855/0045/0785a, Grund (Austria). C1–C4. Paratype, NHMW 1855/0045/0785c, Grund (Austria). D1–D4. Paratype, NHMW 1855/0045/0785d, Grund (Austria).	FIGURE 5. Cochlis degregorioi nov. sp. A1–A4. Paratype, NHMW 1855/0045/0785b, Grund (Austria). B1–B4. Holotype, NHMW 1855/0045/0785a, Grund (Austria). C1–C4. Paratype, NHMW 1855/0045/0785c, Grund (Austria). D1–D4. Paratype, NHMW 1855/0045/0785d, Grund (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6601EFFEAFF00F963FD9EFAC2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326630/files/figure.png	https://doi.org/10.5281/zenodo.17326630	FIGURE 9. A–C. Cochlis gebhardti nov. sp. A1–A4. Holotype, GBA 1912/004/16/1, Mörtersdorf (Austria). B1–B4. Paratype, GBA 1912/004/16/2, Loibersdorf (Austria). C1–C4. Paratype, GBA 1912/004/16/3, Loibersdorf (Austria). D1–D4. Cochlis kostejana (Boettger, 1902), Lectotype, SFM 366940, Coşteiu de Sus (Romania).	FIGURE 9. A–C. Cochlis gebhardti nov. sp. A1–A4. Holotype, GBA 1912/004/16/1, Mörtersdorf (Austria). B1–B4. Paratype, GBA 1912/004/16/2, Loibersdorf (Austria). C1–C4. Paratype, GBA 1912/004/16/3, Loibersdorf (Austria). D1–D4. Cochlis kostejana (Boettger, 1902), Lectotype, SFM 366940, Coşteiu de Sus (Romania).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6601EFFEAFF00F963FD9EFAC2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6601EFFEAFF00F963FD9EFAC2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6601FFFE9FF00FA0AFED9F857.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326630/files/figure.png	https://doi.org/10.5281/zenodo.17326630	FIGURE 9. A–C. Cochlis gebhardti nov. sp. A1–A4. Holotype, GBA 1912/004/16/1, Mörtersdorf (Austria). B1–B4. Paratype, GBA 1912/004/16/2, Loibersdorf (Austria). C1–C4. Paratype, GBA 1912/004/16/3, Loibersdorf (Austria). D1–D4. Cochlis kostejana (Boettger, 1902), Lectotype, SFM 366940, Coşteiu de Sus (Romania).	FIGURE 9. A–C. Cochlis gebhardti nov. sp. A1–A4. Holotype, GBA 1912/004/16/1, Mörtersdorf (Austria). B1–B4. Paratype, GBA 1912/004/16/2, Loibersdorf (Austria). C1–C4. Paratype, GBA 1912/004/16/3, Loibersdorf (Austria). D1–D4. Cochlis kostejana (Boettger, 1902), Lectotype, SFM 366940, Coşteiu de Sus (Romania).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6601FFFE9FF00FA0AFED9F857.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6601FFFE9FF00FA0AFED9F857.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6601DFFEDFF00FF3BFE4EFDAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6601DFFEDFF00FF3BFE4EFDAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326632/files/figure.png	https://doi.org/10.5281/zenodo.17326632	FIGURE 10. Cochlis micromillepunctata nov. sp. A1–A4, Paratype, NHMW 2024/0233/0001, Baden-Sooss (Austria). B1–B4. Holotype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). C1–C4. Paratype, NHMW 1846/0037/0444b, Bad Vöslau (Austria). D1–D4. Paratype, NHMW 1846/0037/0444c, Bad Vöslau (Austria).	FIGURE 10. Cochlis micromillepunctata nov. sp. A1–A4, Paratype, NHMW 2024/0233/0001, Baden-Sooss (Austria). B1–B4. Holotype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). C1–C4. Paratype, NHMW 1846/0037/0444b, Bad Vöslau (Austria). D1–D4. Paratype, NHMW 1846/0037/0444c, Bad Vöslau (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6601DFFEDFF00FF3BFE4EFDAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326634/files/figure.png	https://doi.org/10.5281/zenodo.17326634	FIGURE 11. Cochlis micromillepunctata nov. sp.A1–A2. MNHN A31070, Lăpugiu de Sus (Romania). B1–B4. Holotype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). C1–C3. Paratype, NHMWA1267, Baden-Sooss (Austria). D. Paratype, NHMW A1267, Baden-Sooss (Austria). E. Paratype, NHMW 2024/0233/0001, Baden-Sooss (Austria).	FIGURE 11. Cochlis micromillepunctata nov. sp.A1–A2. MNHN A31070, Lăpugiu de Sus (Romania). B1–B4. Holotype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). C1–C3. Paratype, NHMWA1267, Baden-Sooss (Austria). D. Paratype, NHMW A1267, Baden-Sooss (Austria). E. Paratype, NHMW 2024/0233/0001, Baden-Sooss (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6601DFFEDFF00FF3BFE4EFDAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6601DFFEDFF00FF3BFE4EFDAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6601DFFEDFF00FF3BFE4EFDAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326641/files/figure.png	https://doi.org/10.5281/zenodo.17326641	FIGURE 13. Cochlis neoobscura nov. sp. A1–A4. Holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). B1–B4. Paratype, NHMW 1855/0045/0967b, Bad Vöslau (Austria). C1–C4. Paratype, NHMW 2024/0235/0001, Gainfarn (Austria). D1–D4. Paratype, NHMW 2007z0088/0015a, Enzesfeld (Austria).	FIGURE 13. Cochlis neoobscura nov. sp. A1–A4. Holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). B1–B4. Paratype, NHMW 1855/0045/0967b, Bad Vöslau (Austria). C1–C4. Paratype, NHMW 2024/0235/0001, Gainfarn (Austria). D1–D4. Paratype, NHMW 2007z0088/0015a, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6601DFFEDFF00FF3BFE4EFDAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326618/files/figure.png	https://doi.org/10.5281/zenodo.17326618	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66018FFE3FF00FD22FB16FD5A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326637/files/figure.png	https://doi.org/10.5281/zenodo.17326637	FIGURE 12. Cochlis neglecta (Mayer, 1858). A1–A4. GBA 1912/004/15/1, Maria Dreieichen (Austria). B1–B4. GBA 1912/004/15/1, MariaDreieichen (Austria).C1–C4.GBA 1912/004/15/1,Maria Dreieichen(Austria).D1–D4. GBA1912/004/15/1, Maria Dreieichen (Austria).	FIGURE 12. Cochlis neglecta (Mayer, 1858). A1–A4. GBA 1912/004/15/1, Maria Dreieichen (Austria). B1–B4. GBA 1912/004/15/1, MariaDreieichen (Austria).C1–C4.GBA 1912/004/15/1,Maria Dreieichen(Austria).D1–D4. GBA1912/004/15/1, Maria Dreieichen (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66018FFE3FF00FD22FB16FD5A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66018FFE3FF00FD22FB16FD5A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66016FFE1FF00FCB2FE79FCC6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66016FFE1FF00FCB2FE79FCC6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326641/files/figure.png	https://doi.org/10.5281/zenodo.17326641	FIGURE 13. Cochlis neoobscura nov. sp. A1–A4. Holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). B1–B4. Paratype, NHMW 1855/0045/0967b, Bad Vöslau (Austria). C1–C4. Paratype, NHMW 2024/0235/0001, Gainfarn (Austria). D1–D4. Paratype, NHMW 2007z0088/0015a, Enzesfeld (Austria).	FIGURE 13. Cochlis neoobscura nov. sp. A1–A4. Holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). B1–B4. Paratype, NHMW 1855/0045/0967b, Bad Vöslau (Austria). C1–C4. Paratype, NHMW 2024/0235/0001, Gainfarn (Austria). D1–D4. Paratype, NHMW 2007z0088/0015a, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66016FFE1FF00FCB2FE79FCC6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326643/files/figure.png	https://doi.org/10.5281/zenodo.17326643	FIGURE 14. A1–A3. Cochlis neoobscura nov. sp. NHMW 2007z0088/0015b, Enzesfeld (Austria).	FIGURE 14. A1–A3. Cochlis neoobscura nov. sp. NHMW 2007z0088/0015b, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66016FFE1FF00FCB2FE79FCC6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66016FFE1FF00FCB2FE79FCC6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66016FFE1FF00FCB2FE79FCC6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326618/files/figure.png	https://doi.org/10.5281/zenodo.17326618	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66014FFE0FF00FC0EFE5EFE3A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326647/files/figure.png	https://doi.org/10.5281/zenodo.17326647	FIGURE 15. A1–A4. Cochlis paragigantica nov. sp., holotype, NHMW 1855/0045/0785e, Grund (Austria).	FIGURE 15. A1–A4. Cochlis paragigantica nov. sp., holotype, NHMW 1855/0045/0785e, Grund (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66014FFE0FF00FC0EFE5EFE3A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66014FFE0FF00FC0EFE5EFE3A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66015FFE6FF00F90EFCFBF99F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66015FFE6FF00F90EFCFBF99F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326649/files/figure.png	https://doi.org/10.5281/zenodo.17326649	FIGURE 16. Cochlis pseudoprotracta nov. sp. A1–A4. Paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). B1–B4. Paratype, NHMW 2024/0236/0001, Bad Vöslau (Austria). C1–C4. Holotype, NHMW 2024/0232/0002, Bad Vöslau (Austria). D1–D4. Paratype, NHMW 2024/0237/0001, Jaroměřice (Czech Republic).	FIGURE 16. Cochlis pseudoprotracta nov. sp. A1–A4. Paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). B1–B4. Paratype, NHMW 2024/0236/0001, Bad Vöslau (Austria). C1–C4. Holotype, NHMW 2024/0232/0002, Bad Vöslau (Austria). D1–D4. Paratype, NHMW 2024/0237/0001, Jaroměřice (Czech Republic).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66015FFE6FF00F90EFCFBF99F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66015FFE6FF00F90EFCFBF99F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66013FFE4FF00F9F6FC22FC16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326651/files/figure.png	https://doi.org/10.5281/zenodo.17326651	FIGURE 17. Cochlis transgrediens (Schaffer, 1912). A1–A4. NHMW 1934/0001/0123b, Loibersdorf (Austria). B1–B4. NHMW 1934/0001/0123a, Loibersdorf (Austria). C1–C4. Coll. Anton Breitenberger, Mörtersdorf (Austria). D1–D4. Coll. Anton Breitenberger, Mörtersdorf (Austria).	FIGURE 17. Cochlis transgrediens (Schaffer, 1912). A1–A4. NHMW 1934/0001/0123b, Loibersdorf (Austria). B1–B4. NHMW 1934/0001/0123a, Loibersdorf (Austria). C1–C4. Coll. Anton Breitenberger, Mörtersdorf (Austria). D1–D4. Coll. Anton Breitenberger, Mörtersdorf (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66013FFE4FF00F9F6FC22FC16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66013FFE4FF00F9F6FC22FC16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66011FFD8FF00FC7EFCA3FE3B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66011FFD8FF00FC7EFCA3FE3B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326653/files/figure.png	https://doi.org/10.5281/zenodo.17326653	FIGURE 18. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019. A1–A4. Holotype, MPUM 11735, Varivtsi (Ukraine). B1–B4. paratype, MPUM 11736, Varivtsi (Ukraine). C1–C4. NHMW 1855/0045/0226c, Steinebrunn (Austria). D1–D4. NHMW 1855/0045/0226a, Steinebrunn (Austria).	FIGURE 18. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019. A1–A4. Holotype, MPUM 11735, Varivtsi (Ukraine). B1–B4. paratype, MPUM 11736, Varivtsi (Ukraine). C1–C4. NHMW 1855/0045/0226c, Steinebrunn (Austria). D1–D4. NHMW 1855/0045/0226a, Steinebrunn (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66011FFD8FF00FC7EFCA3FE3B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326655/files/figure.png	https://doi.org/10.5281/zenodo.17326655	FIGURE 19. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019. A1–A4. NHMW 2024/0238/0001, Steinebrunn (Austria). B1–B4. NHMW 2024/0238/0002, Steinebrunn (Austria). C1–C4. NHMW 2024/0235/0002, Gainfarn (Austria). D1–D4. NHMW 2024/0235/0003, Gainfarn (Austria).	FIGURE 19. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019. A1–A4. NHMW 2024/0238/0001, Steinebrunn (Austria). B1–B4. NHMW 2024/0238/0002, Steinebrunn (Austria). C1–C4. NHMW 2024/0235/0002, Gainfarn (Austria). D1–D4. NHMW 2024/0235/0003, Gainfarn (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66011FFD8FF00FC7EFCA3FE3B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326657/files/figure.png	https://doi.org/10.5281/zenodo.17326657	FIGURE 20. A1–A4. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, PIN 5904/121, Shushkivtsi, Ukraine, Middle Miocene, late Badenian. B1–B4. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, PIN 5904/122, Gorodok, Ukraine, Middle Miocene, late Badenian. C1–C4. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, PIN 5904/123, ravine Zhabyak, Ukraine, Middle Miocene, late Badenian. D1–D4. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, PIN 5904/124, Varivtsi, Ukraine, Middle Miocene, late Badenian.	FIGURE 20. A1–A4. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, PIN 5904/121, Shushkivtsi, Ukraine, Middle Miocene, late Badenian. B1–B4. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, PIN 5904/122, Gorodok, Ukraine, Middle Miocene, late Badenian. C1–C4. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, PIN 5904/123, ravine Zhabyak, Ukraine, Middle Miocene, late Badenian. D1–D4. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, PIN 5904/124, Varivtsi, Ukraine, Middle Miocene, late Badenian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66011FFD8FF00FC7EFCA3FE3B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66011FFD8FF00FC7EFCA3FE3B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6602DFFD8FF00FE52FAFCF8AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326659/files/figure.png	https://doi.org/10.5281/zenodo.17326659	FIGURE 21. Cochlis unica (Bałuk, 1995). A1–A4. Holotype, Z.PAL.U.W. No. BkK-G465, Korytnica (Poland). B1–B3. Cochlis sp., 5904/125, ravine Kiyandy, Kazakhstan, Middle Miocene, late Konkian.	FIGURE 21. Cochlis unica (Bałuk, 1995). A1–A4. Holotype, Z.PAL.U.W. No. BkK-G465, Korytnica (Poland). B1–B3. Cochlis sp., 5904/125, ravine Kiyandy, Kazakhstan, Middle Miocene, late Konkian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6602DFFD8FF00FE52FAFCF8AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6602DFFD8FF00FE52FAFCF8AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6602AFFDDFF00FC43FC9BFED6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6602AFFDDFF00FC43FC9BFED6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326661/files/figure.png	https://doi.org/10.5281/zenodo.17326661	FIGURE 22. Cochlis zachosi nov. sp. A1–A4. Holotype, NHMW 2024/0239/0001, Grund (Austria). B1–B4. Paratype, NHMW 2024/0239/0002, Grund (Austria). C1–C4. Paratype, NHMW 2024/0239/0003, Grund (Austria). D1–D4. Paratype, NHMW 2024/0239/0004, Grund (Austria).	FIGURE 22. Cochlis zachosi nov. sp. A1–A4. Holotype, NHMW 2024/0239/0001, Grund (Austria). B1–B4. Paratype, NHMW 2024/0239/0002, Grund (Austria). C1–C4. Paratype, NHMW 2024/0239/0003, Grund (Austria). D1–D4. Paratype, NHMW 2024/0239/0004, Grund (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6602AFFDDFF00FC43FC9BFED6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6602AFFDDFF00FC43FC9BFED6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66028FFD3FF00FA43FBCFFBC4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326663/files/figure.png	https://doi.org/10.5281/zenodo.17326663	FIGURE 23. Naticarius lucapedrialii nov. sp. A1–A6. Holotype, NHMW 2024/0240/0001, Lăpugiu de Sus (Romania). B1–B6. Paratype, MNHN A31069, Lăpugiu de Sus (Romania).	FIGURE 23. Naticarius lucapedrialii nov. sp. A1–A6. Holotype, NHMW 2024/0240/0001, Lăpugiu de Sus (Romania). B1–B6. Paratype, MNHN A31069, Lăpugiu de Sus (Romania).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66028FFD3FF00FA43FBCFFBC4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66028FFD3FF00FA43FBCFFBC4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326624/files/figure.png	https://doi.org/10.5281/zenodo.17326624	FIGURE 6. A1–A4. Cochlis eximia (Eichwald, 1830), SPSU 3/459, lectotype, Zalistsi, Ukraine, Middle Miocene, late Badenian. B1–B4. Cochlis eximia (Eichwald, 1830), SPSU 3/455, paralectotype, Zhukivtsi, Ukraine, Middle Miocene, late Badenian. C1–C4. Cochlis eximia (Eichwald, 1830), PIN 5904/119, Varivtsi, Ukraine, Middle Miocene, late Badenian. D1–D4. Cochlis eximia (Eichwald, 1830), 5904/118, Varivtsi, Ukraine, Middle Miocene, late Badenian.	FIGURE 6. A1–A4. Cochlis eximia (Eichwald, 1830), SPSU 3/459, lectotype, Zalistsi, Ukraine, Middle Miocene, late Badenian. B1–B4. Cochlis eximia (Eichwald, 1830), SPSU 3/455, paralectotype, Zhukivtsi, Ukraine, Middle Miocene, late Badenian. C1–C4. Cochlis eximia (Eichwald, 1830), PIN 5904/119, Varivtsi, Ukraine, Middle Miocene, late Badenian. D1–D4. Cochlis eximia (Eichwald, 1830), 5904/118, Varivtsi, Ukraine, Middle Miocene, late Badenian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66026FFD3FF00FB09FDDAF941.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326665/files/figure.png	https://doi.org/10.5281/zenodo.17326665	FIGURE 24. Tectonatica distincta (Eichwald, 1829). A1–A 5. Neotype, SPSU 3/465, Bilozirka, Ukraine, Middle Miocene, late Badenian. B1–B3. MPUM 11740, Varivtsi (Ukraine), holotype of Tectonatica pseudoprietoi Pedriali, Sosso & Dell’Angelo, 2019. C1–C4. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). (Austria). D1–D4. Tectonatica anistratenkorum Pedriali, Sosso & Dell’Angelo, 2019, holotype, MPUM 11738, Varivtsi (Ukraine). E1–E4. Tectonatica anistratenkorum Pedriali, Sosso & Dell’Angelo, 2019, MPUM 11739, Varivtsi (Ukraine).	FIGURE 24. Tectonatica distincta (Eichwald, 1829). A1–A 5. Neotype, SPSU 3/465, Bilozirka, Ukraine, Middle Miocene, late Badenian. B1–B3. MPUM 11740, Varivtsi (Ukraine), holotype of Tectonatica pseudoprietoi Pedriali, Sosso & Dell’Angelo, 2019. C1–C4. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). (Austria). D1–D4. Tectonatica anistratenkorum Pedriali, Sosso & Dell’Angelo, 2019, holotype, MPUM 11738, Varivtsi (Ukraine). E1–E4. Tectonatica anistratenkorum Pedriali, Sosso & Dell’Angelo, 2019, MPUM 11739, Varivtsi (Ukraine).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66026FFD3FF00FB09FDDAF941.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66026FFD3FF00FB09FDDAF941.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66026FFD1FF00F884FC95FB5E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326665/files/figure.png	https://doi.org/10.5281/zenodo.17326665	FIGURE 24. Tectonatica distincta (Eichwald, 1829). A1–A 5. Neotype, SPSU 3/465, Bilozirka, Ukraine, Middle Miocene, late Badenian. B1–B3. MPUM 11740, Varivtsi (Ukraine), holotype of Tectonatica pseudoprietoi Pedriali, Sosso & Dell’Angelo, 2019. C1–C4. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). (Austria). D1–D4. Tectonatica anistratenkorum Pedriali, Sosso & Dell’Angelo, 2019, holotype, MPUM 11738, Varivtsi (Ukraine). E1–E4. Tectonatica anistratenkorum Pedriali, Sosso & Dell’Angelo, 2019, MPUM 11739, Varivtsi (Ukraine).	FIGURE 24. Tectonatica distincta (Eichwald, 1829). A1–A 5. Neotype, SPSU 3/465, Bilozirka, Ukraine, Middle Miocene, late Badenian. B1–B3. MPUM 11740, Varivtsi (Ukraine), holotype of Tectonatica pseudoprietoi Pedriali, Sosso & Dell’Angelo, 2019. C1–C4. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). (Austria). D1–D4. Tectonatica anistratenkorum Pedriali, Sosso & Dell’Angelo, 2019, holotype, MPUM 11738, Varivtsi (Ukraine). E1–E4. Tectonatica anistratenkorum Pedriali, Sosso & Dell’Angelo, 2019, MPUM 11739, Varivtsi (Ukraine).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66026FFD1FF00F884FC95FB5E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326667/files/figure.png	https://doi.org/10.5281/zenodo.17326667	FIGURE 25. Eichwald’s labels for Natica distincta Eichwald, 1829, SPSU 3/465: above: earlier label, below: later label; collection number 3/465 appeared in the 1930ies, when the collection inventory was prepared by N.A.Bauler and E.S. Poretskaya in Leningrad State University (Gataulina & Arkadiev 2010).	FIGURE 25. Eichwald’s labels for Natica distincta Eichwald, 1829, SPSU 3/465: above: earlier label, below: later label; collection number 3/465 appeared in the 1930ies, when the collection inventory was prepared by N.A.Bauler and E.S. Poretskaya in Leningrad State University (Gataulina & Arkadiev 2010).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66026FFD1FF00F884FC95FB5E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66026FFD1FF00F884FC95FB5E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326622/files/figure.png	https://doi.org/10.5281/zenodo.17326622	FIGURE 5. Cochlis degregorioi nov. sp. A1–A4. Paratype, NHMW 1855/0045/0785b, Grund (Austria). B1–B4. Holotype, NHMW 1855/0045/0785a, Grund (Austria). C1–C4. Paratype, NHMW 1855/0045/0785c, Grund (Austria). D1–D4. Paratype, NHMW 1855/0045/0785d, Grund (Austria).	FIGURE 5. Cochlis degregorioi nov. sp. A1–A4. Paratype, NHMW 1855/0045/0785b, Grund (Austria). B1–B4. Holotype, NHMW 1855/0045/0785a, Grund (Austria). C1–C4. Paratype, NHMW 1855/0045/0785c, Grund (Austria). D1–D4. Paratype, NHMW 1855/0045/0785d, Grund (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66026FFD1FF00F884FC95FB5E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326618/files/figure.png	https://doi.org/10.5281/zenodo.17326618	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66024FFD0FF00FAB4FC0DFD66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66024FFD0FF00FAB4FC0DFD66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326665/files/figure.png	https://doi.org/10.5281/zenodo.17326665	FIGURE 24. Tectonatica distincta (Eichwald, 1829). A1–A 5. Neotype, SPSU 3/465, Bilozirka, Ukraine, Middle Miocene, late Badenian. B1–B3. MPUM 11740, Varivtsi (Ukraine), holotype of Tectonatica pseudoprietoi Pedriali, Sosso & Dell’Angelo, 2019. C1–C4. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). (Austria). D1–D4. Tectonatica anistratenkorum Pedriali, Sosso & Dell’Angelo, 2019, holotype, MPUM 11738, Varivtsi (Ukraine). E1–E4. Tectonatica anistratenkorum Pedriali, Sosso & Dell’Angelo, 2019, MPUM 11739, Varivtsi (Ukraine).	FIGURE 24. Tectonatica distincta (Eichwald, 1829). A1–A 5. Neotype, SPSU 3/465, Bilozirka, Ukraine, Middle Miocene, late Badenian. B1–B3. MPUM 11740, Varivtsi (Ukraine), holotype of Tectonatica pseudoprietoi Pedriali, Sosso & Dell’Angelo, 2019. C1–C4. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). (Austria). D1–D4. Tectonatica anistratenkorum Pedriali, Sosso & Dell’Angelo, 2019, holotype, MPUM 11738, Varivtsi (Ukraine). E1–E4. Tectonatica anistratenkorum Pedriali, Sosso & Dell’Angelo, 2019, MPUM 11739, Varivtsi (Ukraine).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66024FFD0FF00FAB4FC0DFD66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66024FFD0FF00FAB4FC0DFD66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66022FFD6FF00FF3BFAD6FD8E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326671/files/figure.png	https://doi.org/10.5281/zenodo.17326671	FIGURE 26. Tectonatica scutaria nov. nom.A1–A4. Lectotype, KM FI 1166, Maria Dreieichen (Austria). B1–B4. Paralectotype: KM FI 1167, Maria Dreieichen.	FIGURE 26. Tectonatica scutaria nov. nom.A1–A4. Lectotype, KM FI 1166, Maria Dreieichen (Austria). B1–B4. Paralectotype: KM FI 1167, Maria Dreieichen.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66022FFD6FF00FF3BFAD6FD8E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66022FFD6FF00FF3BFAD6FD8E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66020FFC9FF00FCEEFF6EFF6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66020FFC9FF00FCEEFF6EFF6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326673/files/figure.png	https://doi.org/10.5281/zenodo.17326673	FIGURE 27. Pseudolinices empina (De Gregorio, 1885). A1–A4. Holotype, NHMW 1855/0002/0029a, Grund (Austria). B1– B4. NHMW 2024/0244/0001, Enzesfeld (Austria). C1–C4. NHMW 2024/0236/0020, Bad Vöslau (Austria). D1–D4. NHMW 2024/0236/0021, Bad Vöslau (Austria).	FIGURE 27. Pseudolinices empina (De Gregorio, 1885). A1–A4. Holotype, NHMW 1855/0002/0029a, Grund (Austria). B1– B4. NHMW 2024/0244/0001, Enzesfeld (Austria). C1–C4. NHMW 2024/0236/0020, Bad Vöslau (Austria). D1–D4. NHMW 2024/0236/0021, Bad Vöslau (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66020FFC9FF00FCEEFF6EFF6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326675/files/figure.png	https://doi.org/10.5281/zenodo.17326675	FIGURE 28. A1–A3. Pseudolinices empina (De Gregorio, 1885), NHMW 2024/0236/0022, Gainfarn (Austria).	FIGURE 28. A1–A3. Pseudolinices empina (De Gregorio, 1885), NHMW 2024/0236/0022, Gainfarn (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66020FFC9FF00FCEEFF6EFF6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326677/files/figure.png	https://doi.org/10.5281/zenodo.17326677	FIGURE 29. Pseudolinices empina (De Gregorio, 1885). A1–A3. NHMW 1858/0038/0188, Vienna/Pötzleinsdorf (Austria). B1–B3. NHMW 1876/0037/0437, Enzesfeld (Austria). C1–C2. NHMW A1269, Bad Vöslau (Austria).	FIGURE 29. Pseudolinices empina (De Gregorio, 1885). A1–A3. NHMW 1858/0038/0188, Vienna/Pötzleinsdorf (Austria). B1–B3. NHMW 1876/0037/0437, Enzesfeld (Austria). C1–C2. NHMW A1269, Bad Vöslau (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66020FFC9FF00FCEEFF6EFF6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66020FFC9FF00FCEEFF6EFF6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66020FFC9FF00FCEEFF6EFF6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326618/files/figure.png	https://doi.org/10.5281/zenodo.17326618	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6603CFFC8FF00FEE6FA8BF861.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326679/files/figure.png	https://doi.org/10.5281/zenodo.17326679	FIGURE 30. Pseudolinices pseudoredemptus (Friedberg, 1923). A1–A4. NHMW 2024/0247/0001, Kleinebersdorf (Austria). B1–B4. NHMW 2024/0247/0003, Kleinebersdorf (Austria). C1–C4. NHMW 2024/0247/0004, Kleinebersdorf (Austria). D1–D4. NHMW 2024/0247/0005, Kleinebersdorf (Austria).	FIGURE 30. Pseudolinices pseudoredemptus (Friedberg, 1923). A1–A4. NHMW 2024/0247/0001, Kleinebersdorf (Austria). B1–B4. NHMW 2024/0247/0003, Kleinebersdorf (Austria). C1–C4. NHMW 2024/0247/0004, Kleinebersdorf (Austria). D1–D4. NHMW 2024/0247/0005, Kleinebersdorf (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6603CFFC8FF00FEE6FA8BF861.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6603CFFC8FF00FEE6FA8BF861.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6603AFFCDFF00FF3BFC02F8D3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326682/files/figure.png	https://doi.org/10.5281/zenodo.17326682	FIGURE 31. A1–A4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/132, Bursuc, Moldova, Middle Miocene, late Badenian. B1–B4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/133, Bursuc, Moldova, Middle Miocene, late Badenian. C1–C4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/127, Bursuc, Moldova, Middle Miocene, late Badenian. D1–D4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/128, Bursuc, Moldova, Middle Miocene, late Badenian.	FIGURE 31. A1–A4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/132, Bursuc, Moldova, Middle Miocene, late Badenian. B1–B4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/133, Bursuc, Moldova, Middle Miocene, late Badenian. C1–C4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/127, Bursuc, Moldova, Middle Miocene, late Badenian. D1–D4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/128, Bursuc, Moldova, Middle Miocene, late Badenian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6603AFFCDFF00FF3BFC02F8D3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326684/files/figure.png	https://doi.org/10.5281/zenodo.17326684	FIGURE 32. A1–A4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/129, Ripyntsi, Ukraine, Middle Miocene, late Badenian. B1–B4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/130, Yaseniv, Ukraine, Middle Miocene, late Badenian. C1–C4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/131, Ripyntsi, Ukraine, Middle Miocene, late Badenian. D1–D4. Pseudolinices staszici (Friedberg, 1923), SPSU 3/458, syntype of Natica eximia Eichwald, 1830, Zalistsi, Ukraine, Middle Miocene, late Badenian.	FIGURE 32. A1–A4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/129, Ripyntsi, Ukraine, Middle Miocene, late Badenian. B1–B4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/130, Yaseniv, Ukraine, Middle Miocene, late Badenian. C1–C4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/131, Ripyntsi, Ukraine, Middle Miocene, late Badenian. D1–D4. Pseudolinices staszici (Friedberg, 1923), SPSU 3/458, syntype of Natica eximia Eichwald, 1830, Zalistsi, Ukraine, Middle Miocene, late Badenian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6603AFFCDFF00FF3BFC02F8D3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326686/files/figure.png	https://doi.org/10.5281/zenodo.17326686	FIGURE 33.A1–A4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/126, Yaseniv, Ukraine, Middle Miocene, late Badenian. B1–B4. Pseudolinices sp., PIN 5904/134, Agara, Georgia, Middle Miocene, late Konkian.	FIGURE 33.A1–A4. Pseudolinices staszici (Friedberg, 1923), PIN 5904/126, Yaseniv, Ukraine, Middle Miocene, late Badenian. B1–B4. Pseudolinices sp., PIN 5904/134, Agara, Georgia, Middle Miocene, late Konkian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6603AFFCDFF00FF3BFC02F8D3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6603AFFCDFF00FF3BFC02F8D3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66039FFC2FF00F9B3FD14FAB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326688/files/figure.png	https://doi.org/10.5281/zenodo.17326688	FIGURE 34. Euspira amitrovi nov. sp. A1–A4. Holotype, PIN 5904/115, Novo-Kuvinsk, Russia, Middle Miocene, Tarkhanian. B1–B4. Paratype, PIN 5904/116, Belaya River, Russia, Middle Miocene, Tarkhanian. C1–C4. Paratype, PIN 5904/117, Otradnaya, Russia, Middle Miocene, Tarkhanian.	FIGURE 34. Euspira amitrovi nov. sp. A1–A4. Holotype, PIN 5904/115, Novo-Kuvinsk, Russia, Middle Miocene, Tarkhanian. B1–B4. Paratype, PIN 5904/116, Belaya River, Russia, Middle Miocene, Tarkhanian. C1–C4. Paratype, PIN 5904/117, Otradnaya, Russia, Middle Miocene, Tarkhanian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66039FFC2FF00F9B3FD14FAB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326690/files/figure.png	https://doi.org/10.5281/zenodo.17326690	FIGURE 35. Euspira amitrovi nov. sp. A1–A2. Holotype, PIN 5904/115, Novo-Kuvinsk, Russia, Middle Miocene, Tarkhanian.	FIGURE 35. Euspira amitrovi nov. sp. A1–A2. Holotype, PIN 5904/115, Novo-Kuvinsk, Russia, Middle Miocene, Tarkhanian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66039FFC2FF00F9B3FD14FAB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66039FFC2FF00F9B3FD14FAB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66039FFC2FF00F9B3FD14FAB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66037FFC0FF00FADCFA98FD66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66037FFC0FF00FADCFA98FD66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326692/files/figure.png	https://doi.org/10.5281/zenodo.17326692	FIGURE 36. Euspira eblera (De Gregorio, 1885). A1–A4. Holotype, NHMW 1855/0045/0968a, Bad Vöslau (Austria). B1–B4. NHMW 1855/0045/0968b, Bad Vöslau (Austria). C1–C4. NHMW 1846/0037/00445, Bad Vöslau (Austria). D1–D4. NHMW 1846/0037/00445a, Bad Vöslau (Austria).	FIGURE 36. Euspira eblera (De Gregorio, 1885). A1–A4. Holotype, NHMW 1855/0045/0968a, Bad Vöslau (Austria). B1–B4. NHMW 1855/0045/0968b, Bad Vöslau (Austria). C1–C4. NHMW 1846/0037/00445, Bad Vöslau (Austria). D1–D4. NHMW 1846/0037/00445a, Bad Vöslau (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66037FFC0FF00FADCFA98FD66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66037FFC0FF00FADCFA98FD66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66035FFC0FF00FCEFFE9BF8E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66035FFC0FF00FCEFFE9BF8E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326694/files/figure.png	https://doi.org/10.5281/zenodo.17326694	FIGURE 37. A1–A4. Euspira incus nov. sp. Holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania).	FIGURE 37. A1–A4. Euspira incus nov. sp. Holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66035FFC0FF00FCEFFE9BF8E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66035FFC0FF00FCEFFE9BF8E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66032FFC5FF00FDD4FCA1FF43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66032FFC5FF00FDD4FCA1FF43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326696/files/figure.png	https://doi.org/10.5281/zenodo.17326696	FIGURE 38. Euspira merlei nov. sp. A1–A4. Paratype, NHMW 1890/0001/0336b, Lăpugiu de Sus (Romania). B1–B4. Holotype, NHMW 1890/0001/0336a, Lăpugiu de Sus (Romania). C1–C4. Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). D1–D4. NHMW 2024/0231/0004, Weitendorf (Austria).	FIGURE 38. Euspira merlei nov. sp. A1–A4. Paratype, NHMW 1890/0001/0336b, Lăpugiu de Sus (Romania). B1–B4. Holotype, NHMW 1890/0001/0336a, Lăpugiu de Sus (Romania). C1–C4. Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). D1–D4. NHMW 2024/0231/0004, Weitendorf (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66032FFC5FF00FDD4FCA1FF43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326698/files/figure.png	https://doi.org/10.5281/zenodo.17326698	FIGURE 39. Euspira merlei nov. sp. A1–A3. Paratype, NHMW 1873/1615/0017a, Lăpugiu de Sus (Romania). B1–B3. Paratype, NHMW 1890/0001/0336b, Lăpugiu de Sus (Romania). C1–C3. Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). D1–D3. Paratype, NHMW 1873/1615/0017b, Lăpugiu de Sus (Romania). E1–E2. coll.Anton Breitenberger, Weitendorf (Austria). F1–F2. coll. Anton Breitenberger, Weitendorf (Austria). G1–G2. coll. Anton Breitenberger, Weitendorf (Austria). H1–H2. coll. Anton Breitenberger, Weitendorf (Austria).	FIGURE 39. Euspira merlei nov. sp. A1–A3. Paratype, NHMW 1873/1615/0017a, Lăpugiu de Sus (Romania). B1–B3. Paratype, NHMW 1890/0001/0336b, Lăpugiu de Sus (Romania). C1–C3. Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). D1–D3. Paratype, NHMW 1873/1615/0017b, Lăpugiu de Sus (Romania). E1–E2. coll.Anton Breitenberger, Weitendorf (Austria). F1–F2. coll. Anton Breitenberger, Weitendorf (Austria). G1–G2. coll. Anton Breitenberger, Weitendorf (Austria). H1–H2. coll. Anton Breitenberger, Weitendorf (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66032FFC5FF00FDD4FCA1FF43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66032FFC5FF00FDD4FCA1FF43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66032FFC5FF00FDD4FCA1FF43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326618/files/figure.png	https://doi.org/10.5281/zenodo.17326618	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66031FFBBFF00FF3BFDC2FB87.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326700/files/figure.png	https://doi.org/10.5281/zenodo.17326700	FIGURE 40. Euspira moravica nov. sp. A1–A4. Paratype, NHMW 1863/0015/1011, Hrušovany nad Jevišovkou (Czech Republic). B1–B4. Paratype, NHMW 1863/0015/1011a, Hrušovany nad Jevišovkou (Czech Republic). C1–C4. Paratype, NHMW 1863/0015/1011b, Hrušovany nad Jevišovkou (Czech Republic). D1–D4. Holotype, NHMW 1861/0045/0153, Hrušovany nad Jevišovkou (Czech Republic).	FIGURE 40. Euspira moravica nov. sp. A1–A4. Paratype, NHMW 1863/0015/1011, Hrušovany nad Jevišovkou (Czech Republic). B1–B4. Paratype, NHMW 1863/0015/1011a, Hrušovany nad Jevišovkou (Czech Republic). C1–C4. Paratype, NHMW 1863/0015/1011b, Hrušovany nad Jevišovkou (Czech Republic). D1–D4. Holotype, NHMW 1861/0045/0153, Hrušovany nad Jevišovkou (Czech Republic).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66031FFBBFF00FF3BFDC2FB87.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66031FFBBFF00FF3BFDC2FB87.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6604EFFBEFF00FBCEFA96FB4B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6604EFFBEFF00FBCEFA96FB4B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326702/files/figure.png	https://doi.org/10.5281/zenodo.17326702	FIGURE 41. Euspira protracta (Eichwald, 1830). A1–A4. PIN 5904/135, Yaseniv, Ukraine, Middle Miocene, late Badenian. B1–B2. ZI RAS 62196, lectotype, Tarnoruda, Ukraine, Middle Miocene, late Badenian. C1–C3. SPSU 3/464, paralectotype, Zhukivtsi, Ukraine, Middle Miocene, late Badenian.	FIGURE 41. Euspira protracta (Eichwald, 1830). A1–A4. PIN 5904/135, Yaseniv, Ukraine, Middle Miocene, late Badenian. B1–B2. ZI RAS 62196, lectotype, Tarnoruda, Ukraine, Middle Miocene, late Badenian. C1–C3. SPSU 3/464, paralectotype, Zhukivtsi, Ukraine, Middle Miocene, late Badenian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6604EFFBEFF00FBCEFA96FB4B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326704/files/figure.png	https://doi.org/10.5281/zenodo.17326704	FIGURE 42. Euspira protracta (Eichwald, 1830). A1–A4. NHMW 2024/0236/0005, Bad Vöslau (Austria). B1–B4. NHMW 2024/0236/0006,BadVöslau(Austria).C1–C4.NHMW 2024/0236/0007,BadVöslau (Austria).D1–D4. NHMW2024/0236/0008, Bad Vöslau (Austria).	FIGURE 42. Euspira protracta (Eichwald, 1830). A1–A4. NHMW 2024/0236/0005, Bad Vöslau (Austria). B1–B4. NHMW 2024/0236/0006,BadVöslau(Austria).C1–C4.NHMW 2024/0236/0007,BadVöslau (Austria).D1–D4. NHMW2024/0236/0008, Bad Vöslau (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6604EFFBEFF00FBCEFA96FB4B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326706/files/figure.png	https://doi.org/10.5281/zenodo.17326706	FIGURE 43. A1–A4, Euspira protracta (Eichwald, 1830), PIN 5904/136, Novo-Kuvinsk, Russia, Middle Miocene, Tarkhanian. B1–B4, Euspira protracta (Eichwald, 1830), PIN 5904/137, Shirvanskaya, Russia, Middle Miocene, Tarkhanian. C1–C4, Euspira protracta (Eichwald, 1830), PIN 5904/138, Belaya River, Russia, Middle Miocene, Tarkhanian. D1–D4, Euspira protracta (Eichwald, 1830), PIN 5904/139, Shirvanskaya, Russia, Middle Miocene, Tarkhanian.	FIGURE 43. A1–A4, Euspira protracta (Eichwald, 1830), PIN 5904/136, Novo-Kuvinsk, Russia, Middle Miocene, Tarkhanian. B1–B4, Euspira protracta (Eichwald, 1830), PIN 5904/137, Shirvanskaya, Russia, Middle Miocene, Tarkhanian. C1–C4, Euspira protracta (Eichwald, 1830), PIN 5904/138, Belaya River, Russia, Middle Miocene, Tarkhanian. D1–D4, Euspira protracta (Eichwald, 1830), PIN 5904/139, Shirvanskaya, Russia, Middle Miocene, Tarkhanian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6604EFFBEFF00FBCEFA96FB4B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6604EFFBEFF00FBCEFA96FB4B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326618/files/figure.png	https://doi.org/10.5281/zenodo.17326618	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6604EFFBEFF00FBCEFA96FB4B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6604EFFBEFF00FBCEFA96FB4B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326708/files/figure.png	https://doi.org/10.5281/zenodo.17326708	FIGURE 44. E1–E2, Euspira protracta (Eichwald, 1830), PIN 5904/140, Shirvanskaya, Russia, Middle Miocene, Tarkhanian. B1–B2. Euspira sp. [? Euspira protracta (Eichwald, 1830)], PIN 5904/141, Privol’nyi, Russia, Middle Miocene, Konkian.	FIGURE 44. E1–E2, Euspira protracta (Eichwald, 1830), PIN 5904/140, Shirvanskaya, Russia, Middle Miocene, Tarkhanian. B1–B2. Euspira sp. [? Euspira protracta (Eichwald, 1830)], PIN 5904/141, Privol’nyi, Russia, Middle Miocene, Konkian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6604BFFBEFF00FA82FC86F87D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326710/files/figure.png	https://doi.org/10.5281/zenodo.17326710	FIGURE 45. A1–A4. Euspira sirenkoi Pedriali, Sosso & Dell’Angelo, 2019. Holotype, MPUM 11744, Varivtsi (Ukraine). B1– B4. Euspira styriaca (Bauer 1900). Holotype, UMJG&P 5560, St. Florian (Austria). C1–C4. Euspira sp. 1, PIN 2220/296, Gup, Abkhazia, Late Miocene, early Maeotian. D1–D4. Euspira sp. 2, PIN 2220/297, Gup, Abkhazia, Late Miocene, early Maeotian.	FIGURE 45. A1–A4. Euspira sirenkoi Pedriali, Sosso & Dell’Angelo, 2019. Holotype, MPUM 11744, Varivtsi (Ukraine). B1– B4. Euspira styriaca (Bauer 1900). Holotype, UMJG&P 5560, St. Florian (Austria). C1–C4. Euspira sp. 1, PIN 2220/296, Gup, Abkhazia, Late Miocene, early Maeotian. D1–D4. Euspira sp. 2, PIN 2220/297, Gup, Abkhazia, Late Miocene, early Maeotian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6604BFFBEFF00FA82FC86F87D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6604BFFBEFF00FA82FC86F87D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66048FFBCFF00F935FF6EFD3E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326710/files/figure.png	https://doi.org/10.5281/zenodo.17326710	FIGURE 45. A1–A4. Euspira sirenkoi Pedriali, Sosso & Dell’Angelo, 2019. Holotype, MPUM 11744, Varivtsi (Ukraine). B1– B4. Euspira styriaca (Bauer 1900). Holotype, UMJG&P 5560, St. Florian (Austria). C1–C4. Euspira sp. 1, PIN 2220/296, Gup, Abkhazia, Late Miocene, early Maeotian. D1–D4. Euspira sp. 2, PIN 2220/297, Gup, Abkhazia, Late Miocene, early Maeotian.	FIGURE 45. A1–A4. Euspira sirenkoi Pedriali, Sosso & Dell’Angelo, 2019. Holotype, MPUM 11744, Varivtsi (Ukraine). B1– B4. Euspira styriaca (Bauer 1900). Holotype, UMJG&P 5560, St. Florian (Austria). C1–C4. Euspira sp. 1, PIN 2220/296, Gup, Abkhazia, Late Miocene, early Maeotian. D1–D4. Euspira sp. 2, PIN 2220/297, Gup, Abkhazia, Late Miocene, early Maeotian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66048FFBCFF00F935FF6EFD3E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66048FFBCFF00F935FF6EFD3E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66049FFB3FF00FD57FE5EFED7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326712/files/figure.png	https://doi.org/10.5281/zenodo.17326712	FIGURE 46. A–C. Euspira vinitor nov. sp. A1–A4. Paratype, NHMW 1851/0002/0076b, Grund (Austria). B1–B4. Holotype, NHMW 1851/0002/0076a, Grund (Austria). C1–C4. Paratype, NHMW 1851/0002/0076c, Grund (Austria). D1–D4. Euspira wagramensis nov. sp. Holotype, MFN 21041, Fels am Wagram (Austria).	FIGURE 46. A–C. Euspira vinitor nov. sp. A1–A4. Paratype, NHMW 1851/0002/0076b, Grund (Austria). B1–B4. Holotype, NHMW 1851/0002/0076a, Grund (Austria). C1–C4. Paratype, NHMW 1851/0002/0076c, Grund (Austria). D1–D4. Euspira wagramensis nov. sp. Holotype, MFN 21041, Fels am Wagram (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66049FFB3FF00FD57FE5EFED7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66049FFB3FF00FD57FE5EFED7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66046FFB1FF00F88AFE9BFC5E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326714/files/figure.png	https://doi.org/10.5281/zenodo.17326714	FIGURE 47. Euspira vohlandae nov. sp. A1–A4. Holotype, NHMW 1848/0003/0035a, Ritzing (Austria). B1–B4. Paratype, NHMW 1848/0003/0035c, Ritzing (Austria). C1–C4. Paratype, NHMW 1848/0003/0035d, Ritzing (Austria). D1–D4. Paratype, NHMW 1848/0003/0035f, Ritzing (Austria).	FIGURE 47. Euspira vohlandae nov. sp. A1–A4. Holotype, NHMW 1848/0003/0035a, Ritzing (Austria). B1–B4. Paratype, NHMW 1848/0003/0035c, Ritzing (Austria). C1–C4. Paratype, NHMW 1848/0003/0035d, Ritzing (Austria). D1–D4. Paratype, NHMW 1848/0003/0035f, Ritzing (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66046FFB1FF00F88AFE9BFC5E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66046FFB1FF00F88AFE9BFC5E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66044FFB0FF00FBB7FDBBFE47.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326712/files/figure.png	https://doi.org/10.5281/zenodo.17326712	FIGURE 46. A–C. Euspira vinitor nov. sp. A1–A4. Paratype, NHMW 1851/0002/0076b, Grund (Austria). B1–B4. Holotype, NHMW 1851/0002/0076a, Grund (Austria). C1–C4. Paratype, NHMW 1851/0002/0076c, Grund (Austria). D1–D4. Euspira wagramensis nov. sp. Holotype, MFN 21041, Fels am Wagram (Austria).	FIGURE 46. A–C. Euspira vinitor nov. sp. A1–A4. Paratype, NHMW 1851/0002/0076b, Grund (Austria). B1–B4. Holotype, NHMW 1851/0002/0076a, Grund (Austria). C1–C4. Paratype, NHMW 1851/0002/0076c, Grund (Austria). D1–D4. Euspira wagramensis nov. sp. Holotype, MFN 21041, Fels am Wagram (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66044FFB0FF00FBB7FDBBFE47.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66044FFB0FF00FBB7FDBBFE47.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66045FFB0FF00FD8EFBF1F83A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326710/files/figure.png	https://doi.org/10.5281/zenodo.17326710	FIGURE 45. A1–A4. Euspira sirenkoi Pedriali, Sosso & Dell’Angelo, 2019. Holotype, MPUM 11744, Varivtsi (Ukraine). B1– B4. Euspira styriaca (Bauer 1900). Holotype, UMJG&P 5560, St. Florian (Austria). C1–C4. Euspira sp. 1, PIN 2220/296, Gup, Abkhazia, Late Miocene, early Maeotian. D1–D4. Euspira sp. 2, PIN 2220/297, Gup, Abkhazia, Late Miocene, early Maeotian.	FIGURE 45. A1–A4. Euspira sirenkoi Pedriali, Sosso & Dell’Angelo, 2019. Holotype, MPUM 11744, Varivtsi (Ukraine). B1– B4. Euspira styriaca (Bauer 1900). Holotype, UMJG&P 5560, St. Florian (Austria). C1–C4. Euspira sp. 1, PIN 2220/296, Gup, Abkhazia, Late Miocene, early Maeotian. D1–D4. Euspira sp. 2, PIN 2220/297, Gup, Abkhazia, Late Miocene, early Maeotian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66045FFB0FF00FD8EFBF1F83A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66045FFB0FF00FD8EFBF1F83A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66042FFB7FF00FF3BFF52FB7E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326716/files/figure.png	https://doi.org/10.5281/zenodo.17326716	FIGURE 48.A1–A6. Figovina yasenivensis nov. sp., PIN 5904/142, holotype,Yaseniv, Ukraine, Middle Miocene, late Badenian. B1–B5. Figovina yasenivensis nov. sp., PIN 5904/143, Yaseniv, Ukraine, Middle Miocene, late Badenian. C1–C4. Figovina sarmatica (Friedberg, 1938), NHMW 2024/0242/0001, Siebenhirten (Austria). D1–D4. Figovina sarmatica (Friedberg, 1938), NHMW 2024/0242/0002, Siebenhirten (Austria). E1–E3. Figovina sarmatica (Friedberg, 1938), NHMW 2024/0242/0004, Siebenhirten (Austria).	FIGURE 48.A1–A6. Figovina yasenivensis nov. sp., PIN 5904/142, holotype,Yaseniv, Ukraine, Middle Miocene, late Badenian. B1–B5. Figovina yasenivensis nov. sp., PIN 5904/143, Yaseniv, Ukraine, Middle Miocene, late Badenian. C1–C4. Figovina sarmatica (Friedberg, 1938), NHMW 2024/0242/0001, Siebenhirten (Austria). D1–D4. Figovina sarmatica (Friedberg, 1938), NHMW 2024/0242/0002, Siebenhirten (Austria). E1–E3. Figovina sarmatica (Friedberg, 1938), NHMW 2024/0242/0004, Siebenhirten (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66042FFB7FF00FF3BFF52FB7E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66042FFB7FF00FF3BFF52FB7E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326622/files/figure.png	https://doi.org/10.5281/zenodo.17326622	FIGURE 5. Cochlis degregorioi nov. sp. A1–A4. Paratype, NHMW 1855/0045/0785b, Grund (Austria). B1–B4. Holotype, NHMW 1855/0045/0785a, Grund (Austria). C1–C4. Paratype, NHMW 1855/0045/0785c, Grund (Austria). D1–D4. Paratype, NHMW 1855/0045/0785d, Grund (Austria).	FIGURE 5. Cochlis degregorioi nov. sp. A1–A4. Paratype, NHMW 1855/0045/0785b, Grund (Austria). B1–B4. Holotype, NHMW 1855/0045/0785a, Grund (Austria). C1–C4. Paratype, NHMW 1855/0045/0785c, Grund (Austria). D1–D4. Paratype, NHMW 1855/0045/0785d, Grund (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66042FFB7FF00FF3BFF52FB7E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66042FFB5FF00FA97FDD4FAA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326716/files/figure.png	https://doi.org/10.5281/zenodo.17326716	FIGURE 48.A1–A6. Figovina yasenivensis nov. sp., PIN 5904/142, holotype,Yaseniv, Ukraine, Middle Miocene, late Badenian. B1–B5. Figovina yasenivensis nov. sp., PIN 5904/143, Yaseniv, Ukraine, Middle Miocene, late Badenian. C1–C4. Figovina sarmatica (Friedberg, 1938), NHMW 2024/0242/0001, Siebenhirten (Austria). D1–D4. Figovina sarmatica (Friedberg, 1938), NHMW 2024/0242/0002, Siebenhirten (Austria). E1–E3. Figovina sarmatica (Friedberg, 1938), NHMW 2024/0242/0004, Siebenhirten (Austria).	FIGURE 48.A1–A6. Figovina yasenivensis nov. sp., PIN 5904/142, holotype,Yaseniv, Ukraine, Middle Miocene, late Badenian. B1–B5. Figovina yasenivensis nov. sp., PIN 5904/143, Yaseniv, Ukraine, Middle Miocene, late Badenian. C1–C4. Figovina sarmatica (Friedberg, 1938), NHMW 2024/0242/0001, Siebenhirten (Austria). D1–D4. Figovina sarmatica (Friedberg, 1938), NHMW 2024/0242/0002, Siebenhirten (Austria). E1–E3. Figovina sarmatica (Friedberg, 1938), NHMW 2024/0242/0004, Siebenhirten (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66042FFB5FF00FA97FDD4FAA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326712/files/figure.png	https://doi.org/10.5281/zenodo.17326712	FIGURE 46. A–C. Euspira vinitor nov. sp. A1–A4. Paratype, NHMW 1851/0002/0076b, Grund (Austria). B1–B4. Holotype, NHMW 1851/0002/0076a, Grund (Austria). C1–C4. Paratype, NHMW 1851/0002/0076c, Grund (Austria). D1–D4. Euspira wagramensis nov. sp. Holotype, MFN 21041, Fels am Wagram (Austria).	FIGURE 46. A–C. Euspira vinitor nov. sp. A1–A4. Paratype, NHMW 1851/0002/0076b, Grund (Austria). B1–B4. Holotype, NHMW 1851/0002/0076a, Grund (Austria). C1–C4. Paratype, NHMW 1851/0002/0076c, Grund (Austria). D1–D4. Euspira wagramensis nov. sp. Holotype, MFN 21041, Fels am Wagram (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66042FFB5FF00FA97FDD4FAA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66042FFB5FF00FA97FDD4FAA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66042FFB5FF00FA97FDD4FAA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326618/files/figure.png	https://doi.org/10.5281/zenodo.17326618	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66040FFABFF00FA2EFA04FB9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66040FFABFF00FA2EFA04FB9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326720/files/figure.png	https://doi.org/10.5281/zenodo.17326720	FIGURE 49. Figovina szobiensis (Strausz, 1959). A1–A4. NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). B1–B4. NHMW 2024/0243/0002, Boršov (= Porstendorf) (Czech Republic). C1–C4. NHMW 2024/0243/0003, Boršov (= Porstendorf) (Czech Republic). D1–D4. NHMW 2024/0243/0004, Boršov (= Porstendorf) (Czech Republic).	FIGURE 49. Figovina szobiensis (Strausz, 1959). A1–A4. NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). B1–B4. NHMW 2024/0243/0002, Boršov (= Porstendorf) (Czech Republic). C1–C4. NHMW 2024/0243/0003, Boršov (= Porstendorf) (Czech Republic). D1–D4. NHMW 2024/0243/0004, Boršov (= Porstendorf) (Czech Republic).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66040FFABFF00FA2EFA04FB9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66040FFABFF00FA2EFA04FB9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605EFFA9FF00FBF1FC8AFDAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326724/files/figure.png	https://doi.org/10.5281/zenodo.17326724	FIGURE 50. Payraudeautia badensis (Glibert, 1952). A1–A4. Holotype, No. 2843, Baden (Austria). B1–B4. NHMW 2024/0236/0009,BadVöslau(Austria).C1–C4.NHMW 2024/0236/0010,BadVöslau (Austria).D1–D4. NHMW2024/0236/0012, Bad Vöslau (Austria).	FIGURE 50. Payraudeautia badensis (Glibert, 1952). A1–A4. Holotype, No. 2843, Baden (Austria). B1–B4. NHMW 2024/0236/0009,BadVöslau(Austria).C1–C4.NHMW 2024/0236/0010,BadVöslau (Austria).D1–D4. NHMW2024/0236/0012, Bad Vöslau (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605EFFA9FF00FBF1FC8AFDAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605EFFA9FF00FBF1FC8AFDAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605CFFA9FF00FD22FCE8F82E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326726/files/figure.png	https://doi.org/10.5281/zenodo.17326726	FIGURE 51. Payraudeautia ermesi Pedriali, Sosso & Dell’Angelo, 2019. A1–A4. Holotype, MPUM 11746, Varivtsi (Ukraine). B1–B4. Paratype, MPUM 11747, Varivtsi (Ukraine).	FIGURE 51. Payraudeautia ermesi Pedriali, Sosso & Dell’Angelo, 2019. A1–A4. Holotype, MPUM 11746, Varivtsi (Ukraine). B1–B4. Paratype, MPUM 11747, Varivtsi (Ukraine).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605CFFA9FF00FD22FCE8F82E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605CFFA9FF00FD22FCE8F82E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605DFFAEFF00FF3BFE0CFE46.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605DFFAEFF00FF3BFE0CFE46.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326728/files/figure.png	https://doi.org/10.5281/zenodo.17326728	FIGURE 52. Payraudeautia profunda nov. sp. A1–A4. Holotype, NHMW 1846/0037/0444b, Bad Vöslau (Austria). B1–B4. Paratype, NHMW 2024/0236/0016, Bad Vöslau (Austria). C1–C4. Paratype, NHMW 2024/0236/0017, Bad Vöslau (Austria). D1–D4. Paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria).	FIGURE 52. Payraudeautia profunda nov. sp. A1–A4. Holotype, NHMW 1846/0037/0444b, Bad Vöslau (Austria). B1–B4. Paratype, NHMW 2024/0236/0016, Bad Vöslau (Austria). C1–C4. Paratype, NHMW 2024/0236/0017, Bad Vöslau (Austria). D1–D4. Paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605DFFAEFF00FF3BFE0CFE46.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605DFFAEFF00FF3BFE0CFE46.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605BFFACFF00FD8EFC82F892.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326730/files/figure.png	https://doi.org/10.5281/zenodo.17326730	FIGURE 53. Payraudeautia volhynia (d’Orbigny, 1852). A1–A4. Holotype of Payraudeautia varivtsiana Pedriali, Sosso & Dell’Angelo, 2019, MPUM 11754, Varivtsi (Ukraine). B1–B4. Paratype of Payraudeautia varivtsiana Pedriali, Sosso & Dell’Angelo, 2019, MPUM 11755, Varivtsi (Ukraine). C1–C4. MPUM 11749, Varivtsi (Ukraine). D1–D5. PIN 5904/144, Gorodok, Ukraine, Middle Miocene, late Badenian. E1–E3. 5904/145, ravine Zhabyak, Ukraine, Middle Miocene, late Badenian. F. PIN 5904/146, Gorodok, Ukraine, Middle Miocene, late Badenian.	FIGURE 53. Payraudeautia volhynia (d’Orbigny, 1852). A1–A4. Holotype of Payraudeautia varivtsiana Pedriali, Sosso & Dell’Angelo, 2019, MPUM 11754, Varivtsi (Ukraine). B1–B4. Paratype of Payraudeautia varivtsiana Pedriali, Sosso & Dell’Angelo, 2019, MPUM 11755, Varivtsi (Ukraine). C1–C4. MPUM 11749, Varivtsi (Ukraine). D1–D5. PIN 5904/144, Gorodok, Ukraine, Middle Miocene, late Badenian. E1–E3. 5904/145, ravine Zhabyak, Ukraine, Middle Miocene, late Badenian. F. PIN 5904/146, Gorodok, Ukraine, Middle Miocene, late Badenian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605BFFACFF00FD8EFC82F892.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605BFFACFF00FD8EFC82F892.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605BFFACFF00FD8EFC82F892.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326618/files/figure.png	https://doi.org/10.5281/zenodo.17326618	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	FIGURE 3. Cochlis baconica (Kókay, 1966). A1–A4. holotype, M277, Herend (Hungary). B1–B4. NHMW 2024/0229/0001, Weitendorf (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6605BFFACFF00FD8EFC82F892.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326622/files/figure.png	https://doi.org/10.5281/zenodo.17326622	FIGURE 5. Cochlis degregorioi nov. sp. A1–A4. Paratype, NHMW 1855/0045/0785b, Grund (Austria). B1–B4. Holotype, NHMW 1855/0045/0785a, Grund (Austria). C1–C4. Paratype, NHMW 1855/0045/0785c, Grund (Austria). D1–D4. Paratype, NHMW 1855/0045/0785d, Grund (Austria).	FIGURE 5. Cochlis degregorioi nov. sp. A1–A4. Paratype, NHMW 1855/0045/0785b, Grund (Austria). B1–B4. Holotype, NHMW 1855/0045/0785a, Grund (Austria). C1–C4. Paratype, NHMW 1855/0045/0785c, Grund (Austria). D1–D4. Paratype, NHMW 1855/0045/0785d, Grund (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66056FFA2FF00FF3BFADBF99E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326732/files/figure.png	https://doi.org/10.5281/zenodo.17326732	FIGURE 54. Robbanacca cerovaensis (Harzhauser in Harzhauser, Mandic & Schlögl, 2011). A. Holotype: NHMW 2010/0080/0001, Cerová-Lieskové (Slovakia). B. Paratype, NHMW 2010/0080/0002, Cerová-Lieskové (Slovakia). C. Paratype, NHMW 2010/0080/0002, Cerová-Lieskové (Slovakia).	FIGURE 54. Robbanacca cerovaensis (Harzhauser in Harzhauser, Mandic & Schlögl, 2011). A. Holotype: NHMW 2010/0080/0001, Cerová-Lieskové (Slovakia). B. Paratype, NHMW 2010/0080/0002, Cerová-Lieskové (Slovakia). C. Paratype, NHMW 2010/0080/0002, Cerová-Lieskové (Slovakia).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66057FFA1FF00F9F7FA9DF97B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326734/files/figure.png	https://doi.org/10.5281/zenodo.17326734	FIGURE 55. Tethynices plicatulaeformis (Kittl, 1887). A1–A4. Paralectotype, NHMW 1888/0001/0025a, Orlová (Czech Republic). B1–B4. Lectotype, NHMW 1888/0001/0025b, Orlová (Czech Republic).	FIGURE 55. Tethynices plicatulaeformis (Kittl, 1887). A1–A4. Paralectotype, NHMW 1888/0001/0025a, Orlová (Czech Republic). B1–B4. Lectotype, NHMW 1888/0001/0025b, Orlová (Czech Republic).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66057FFA1FF00F9F7FA9DF97B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66057FFA1FF00F9F7FA9DF97B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66054FFA7FF00F8B6FBF7FDF6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326736/files/figure.png	https://doi.org/10.5281/zenodo.17326736	FIGURE 56. A–B. Neverita manhartensis (Schaffer, 1912). A1–A4. Lectotype, GBA 1912/004/18/1, Gauderndorf (Austria). B1–B4. Paralectotype, GBA 1912/004/18/2, Loibersdorf (Austria). C1–C4. Neverita olla (de Serres, 1829), SFM 366939, Coşteiu de Sus (Romania).	FIGURE 56. A–B. Neverita manhartensis (Schaffer, 1912). A1–A4. Lectotype, GBA 1912/004/18/1, Gauderndorf (Austria). B1–B4. Paralectotype, GBA 1912/004/18/2, Loibersdorf (Austria). C1–C4. Neverita olla (de Serres, 1829), SFM 366939, Coşteiu de Sus (Romania).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66054FFA7FF00F8B6FBF7FDF6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66054FFA7FF00F8B6FBF7FDF6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66052FFA4FF00FD1FFA1AFC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66052FFA4FF00FD1FFA1AFC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326736/files/figure.png	https://doi.org/10.5281/zenodo.17326736	FIGURE 56. A–B. Neverita manhartensis (Schaffer, 1912). A1–A4. Lectotype, GBA 1912/004/18/1, Gauderndorf (Austria). B1–B4. Paralectotype, GBA 1912/004/18/2, Loibersdorf (Austria). C1–C4. Neverita olla (de Serres, 1829), SFM 366939, Coşteiu de Sus (Romania).	FIGURE 56. A–B. Neverita manhartensis (Schaffer, 1912). A1–A4. Lectotype, GBA 1912/004/18/1, Gauderndorf (Austria). B1–B4. Paralectotype, GBA 1912/004/18/2, Loibersdorf (Austria). C1–C4. Neverita olla (de Serres, 1829), SFM 366939, Coşteiu de Sus (Romania).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66052FFA4FF00FD1FFA1AFC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326738/files/figure.png	https://doi.org/10.5281/zenodo.17326738	FIGURE 57. Neverita olla (de Serres, 1829). A1–A4. NHMW 1855/0002/0055a, Enzesfeld (Austria). B1–B4. NHMW 1853/0016/0014, Bad Vöslau (Austria). C1–C4. NHMW 1846/0037/0435, Enzesfeld (Austria). D1–D4. NHMW 2024/0236/0022, Bad Vöslau (Austria). E1–E2. 5904/147, Shushkivtsi, Ukraine, Middle Miocene, late Badenian. F1–F2. 5904/148, ravine Yaman- Dzhalga, Russia, Middle Miocene, Tarkhanian.	FIGURE 57. Neverita olla (de Serres, 1829). A1–A4. NHMW 1855/0002/0055a, Enzesfeld (Austria). B1–B4. NHMW 1853/0016/0014, Bad Vöslau (Austria). C1–C4. NHMW 1846/0037/0435, Enzesfeld (Austria). D1–D4. NHMW 2024/0236/0022, Bad Vöslau (Austria). E1–E2. 5904/147, Shushkivtsi, Ukraine, Middle Miocene, late Badenian. F1–F2. 5904/148, ravine Yaman- Dzhalga, Russia, Middle Miocene, Tarkhanian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66052FFA4FF00FD1FFA1AFC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66052FFA4FF00FD1FFA1AFC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66051FF9AFF00FBF4FB49FE14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326740/files/figure.png	https://doi.org/10.5281/zenodo.17326740	FIGURE 58. A–B. Eunaticina martingrossi nov. nom. A1–A4. UMJG&P 60782, Wetzelsdorf (Austria). B1–B4. Neotype, UMJG&P 60782, Wetzelsdorf (Austria). C1–C4. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. D1–D4. Sigaretotrema vindobonense nov. sp. Holotype, NHMW 1846/0037/0434, Enzesfeld (Austria).	FIGURE 58. A–B. Eunaticina martingrossi nov. nom. A1–A4. UMJG&P 60782, Wetzelsdorf (Austria). B1–B4. Neotype, UMJG&P 60782, Wetzelsdorf (Austria). C1–C4. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. D1–D4. Sigaretotrema vindobonense nov. sp. Holotype, NHMW 1846/0037/0434, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66051FF9AFF00FBF4FB49FE14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66051FF9AFF00FBF4FB49FE14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606FFF9AFF00FE1DFF51F936.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606FFF9AFF00FE1DFF51F936.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326740/files/figure.png	https://doi.org/10.5281/zenodo.17326740	FIGURE 58. A–B. Eunaticina martingrossi nov. nom. A1–A4. UMJG&P 60782, Wetzelsdorf (Austria). B1–B4. Neotype, UMJG&P 60782, Wetzelsdorf (Austria). C1–C4. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. D1–D4. Sigaretotrema vindobonense nov. sp. Holotype, NHMW 1846/0037/0434, Enzesfeld (Austria).	FIGURE 58. A–B. Eunaticina martingrossi nov. nom. A1–A4. UMJG&P 60782, Wetzelsdorf (Austria). B1–B4. Neotype, UMJG&P 60782, Wetzelsdorf (Austria). C1–C4. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. D1–D4. Sigaretotrema vindobonense nov. sp. Holotype, NHMW 1846/0037/0434, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606FFF9AFF00FE1DFF51F936.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606FFF9AFF00FE1DFF51F936.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606FFF9AFF00FE1DFF51F936.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326742/files/figure.png	https://doi.org/10.5281/zenodo.17326742	FIGURE 59. Sculpture details of Paratethyan Sininae. A. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria).B. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. C. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605, Bad Vöslau (Austria). D. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). E. Sinum magnificum nov. sp., holotype, NHMW 1854/0002/0072, Grund (Austria). F. paratype, NHMW 2024/0239/0003, Grund (Austria).	FIGURE 59. Sculpture details of Paratethyan Sininae. A. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria).B. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. C. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605, Bad Vöslau (Austria). D. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). E. Sinum magnificum nov. sp., holotype, NHMW 1854/0002/0072, Grund (Austria). F. paratype, NHMW 2024/0239/0003, Grund (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606FFF98FF00F95FFF52FF43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606FFF98FF00F95FFF52FF43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326740/files/figure.png	https://doi.org/10.5281/zenodo.17326740	FIGURE 58. A–B. Eunaticina martingrossi nov. nom. A1–A4. UMJG&P 60782, Wetzelsdorf (Austria). B1–B4. Neotype, UMJG&P 60782, Wetzelsdorf (Austria). C1–C4. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. D1–D4. Sigaretotrema vindobonense nov. sp. Holotype, NHMW 1846/0037/0434, Enzesfeld (Austria).	FIGURE 58. A–B. Eunaticina martingrossi nov. nom. A1–A4. UMJG&P 60782, Wetzelsdorf (Austria). B1–B4. Neotype, UMJG&P 60782, Wetzelsdorf (Austria). C1–C4. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. D1–D4. Sigaretotrema vindobonense nov. sp. Holotype, NHMW 1846/0037/0434, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606FFF98FF00F95FFF52FF43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606FFF98FF00F95FFF52FF43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606FFF98FF00F95FFF52FF43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326742/files/figure.png	https://doi.org/10.5281/zenodo.17326742	FIGURE 59. Sculpture details of Paratethyan Sininae. A. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria).B. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. C. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605, Bad Vöslau (Austria). D. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). E. Sinum magnificum nov. sp., holotype, NHMW 1854/0002/0072, Grund (Austria). F. paratype, NHMW 2024/0239/0003, Grund (Austria).	FIGURE 59. Sculpture details of Paratethyan Sininae. A. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria).B. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. C. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605, Bad Vöslau (Austria). D. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). E. Sinum magnificum nov. sp., holotype, NHMW 1854/0002/0072, Grund (Austria). F. paratype, NHMW 2024/0239/0003, Grund (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606AFF9DFF00FDC9FC59FA72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606AFF9DFF00FDC9FC59FA72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326742/files/figure.png	https://doi.org/10.5281/zenodo.17326742	FIGURE 59. Sculpture details of Paratethyan Sininae. A. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria).B. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. C. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605, Bad Vöslau (Austria). D. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). E. Sinum magnificum nov. sp., holotype, NHMW 1854/0002/0072, Grund (Austria). F. paratype, NHMW 2024/0239/0003, Grund (Austria).	FIGURE 59. Sculpture details of Paratethyan Sininae. A. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria).B. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. C. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605, Bad Vöslau (Austria). D. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). E. Sinum magnificum nov. sp., holotype, NHMW 1854/0002/0072, Grund (Austria). F. paratype, NHMW 2024/0239/0003, Grund (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606AFF9DFF00FDC9FC59FA72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326744/files/figure.png	https://doi.org/10.5281/zenodo.17326744	FIGURE 60. A1–A2. Sigaretotrema sp., NHMW 1860/0050/0375, Gauderndorf (Austria). Sinum affine (Eichwald, 1830). B1– B2. Sinum affine (Eichwald, 1830), Borsodbóta (Hungary).	FIGURE 60. A1–A2. Sigaretotrema sp., NHMW 1860/0050/0375, Gauderndorf (Austria). Sinum affine (Eichwald, 1830). B1– B2. Sinum affine (Eichwald, 1830), Borsodbóta (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606AFF9DFF00FDC9FC59FA72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326746/files/figure.png	https://doi.org/10.5281/zenodo.17326746	FIGURE 61. Sinum affine (Eichwald, 1830). A1–A4. NHMW 2010/0004/0605a, Bad Vöslau (Austria). B1–B4. NHMW 2010/0004/0605b, Bad Vöslau (Austria). C1–C4. NHMW 2010/0004/0605c, Bad Vöslau (Austria). D1–D4. NHMW 1870/0037/0036, Ritzing (Austria). Sinum affine (Eichwald, 1830). E1–E4. ZI RAS 62208, lectotype, Zhukivtsi, Ukraine, Middle Miocene, late Badenian. F1–F4. PIN 5904/149, Varivtsi, Ukraine, Middle Miocene, late Badenian.	FIGURE 61. Sinum affine (Eichwald, 1830). A1–A4. NHMW 2010/0004/0605a, Bad Vöslau (Austria). B1–B4. NHMW 2010/0004/0605b, Bad Vöslau (Austria). C1–C4. NHMW 2010/0004/0605c, Bad Vöslau (Austria). D1–D4. NHMW 1870/0037/0036, Ritzing (Austria). Sinum affine (Eichwald, 1830). E1–E4. ZI RAS 62208, lectotype, Zhukivtsi, Ukraine, Middle Miocene, late Badenian. F1–F4. PIN 5904/149, Varivtsi, Ukraine, Middle Miocene, late Badenian.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606AFF9DFF00FDC9FC59FA72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A6606AFF9DFF00FDC9FC59FA72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66068FF93FF00F99BFBE1FD12.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326616/files/figure.png	https://doi.org/10.5281/zenodo.17326616	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 2. Protoconchs and early teleoconch whorls. Scale bar = 0.5 mm. A. Cochlis cuniculata nov. sp., holotype, NHMW 2024/0230/0001, Várpalota (Hungary). B. Cochlis degregorioi nov. sp., paratype, NHMW 1855/0045/0785b, Grund (Austria). C. Cochlis micromillepunctata nov. sp., paratype, NHMW 1846/0037/0444a, Bad Vöslau (Austria). D. Cochlis neoobscura nov. sp., holotype, NHMW 1855/0045/0967a, Bad Vöslau (Austria). E. Cochlis pseudoprotracta nov. sp., paratype, NHMW 2024/0236/0002, Bad Vöslau (Austria). F. Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019, NHMW 1855/0045/0226b, Steinebrunn (Austria). G. Cochlis zachosi nov. sp., holotype: NHMW 2024/0239/0001, Grund (Austria). H. Tectonatica tectula (Sacco, 1890), SFM 366841, Coşteiu de Sus (Romania). I. Pseudolinices empina (De Gregorio, 1885), NHMW A1269, Bad Vöslau (Austria). J. Euspira eblera (De Gregorio, 1885), NHMW 1846/0037/00445a, Bad Vöslau (Austria). K. Euspira merlei nov. sp., Paratype, NHMW 1890/0001/0336c, Lăpugiu de Sus (Romania). L. Euspira protracta (Eichwald, 1830), NHMW 2024/0236/0005a, Bad Vöslau (Austria). M. Figovina szobiensis (Strausz, 1959), NHMW 2024/0243/0001, Boršov (= Porstendorf) (Czech Republic). N. Euspira incus nov. sp., holotype, NHMW 2024/0240/0002, Lăpugiu de Sus (Romania). O. Payraudeautia profunda nov. sp., paratype, NHMW 2024/0236/0018, Bad Vöslau (Austria). P. Neverita olla (de Serres, 1829), NHMW 1846/0037/0435, Enzesfeld (Austria). Q. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. R. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria). S. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). T. Sinum magnificum nov. sp., paratype, NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66068FF93FF00F99BFBE1FD12.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326742/files/figure.png	https://doi.org/10.5281/zenodo.17326742	FIGURE 59. Sculpture details of Paratethyan Sininae. A. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria).B. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. C. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605, Bad Vöslau (Austria). D. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). E. Sinum magnificum nov. sp., holotype, NHMW 1854/0002/0072, Grund (Austria). F. paratype, NHMW 2024/0239/0003, Grund (Austria).	FIGURE 59. Sculpture details of Paratethyan Sininae. A. Sigaretotrema vindobonense nov. sp., holotype, NHMW 1846/0037/0434, Enzesfeld (Austria).B. Sigaretotrema exclathratum (Sacco, 1891), holotype, NHMW 2013/0053/0001, locality unknown. C. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605, Bad Vöslau (Austria). D. Sinum affine (Eichwald, 1830), NHMW 2010/0004/0605b, Bad Vöslau (Austria). E. Sinum magnificum nov. sp., holotype, NHMW 1854/0002/0072, Grund (Austria). F. paratype, NHMW 2024/0239/0003, Grund (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66068FF93FF00F99BFBE1FD12.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326748/files/figure.png	https://doi.org/10.5281/zenodo.17326748	FIGURE 62. Sinum magnificum nov. sp. A1–A4. Paratype, NHMW 2024/0239/0005, Grund (Austria). B1–B4. Holotype, NHMW 1854/0002/0072, Grund (Austria). C1–C4. Paratype, NHMW 2024/0239/0003, Grund (Austria). D1–D4. NHMW 1853/0010/0033, Enzesfeld (Austria).	FIGURE 62. Sinum magnificum nov. sp. A1–A4. Paratype, NHMW 2024/0239/0005, Grund (Austria). B1–B4. Holotype, NHMW 1854/0002/0072, Grund (Austria). C1–C4. Paratype, NHMW 2024/0239/0003, Grund (Austria). D1–D4. NHMW 1853/0010/0033, Enzesfeld (Austria).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66068FF93FF00F99BFBE1FD12.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326614/files/figure.png	https://doi.org/10.5281/zenodo.17326614	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	FIGURE 1. Terminology for shell morphology; MD: maximum diameter, AH: aperture height, LWH: last whorl height, SL: shell length, CA: columellar angle, AA: apical angle.	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
03A687A66068FF93FF00F99BFBE1FD12.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17326620/files/figure.png	https://doi.org/10.5281/zenodo.17326620	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	FIGURE 4. Cochlis cuniculata nov. sp. A1–A4. Paratype, NHMW 2024/0230/0002, Várpalota (Hungary). B1–B4. Holotype, NHMW 2024/0230/0001, Várpalota (Hungary). C1–C4. Paratype, NHMW 2024/0230/0003, Várpalota (Hungary). D1–D4. Paratype, NHMW 2024/0230/0004, Várpalota (Hungary).	2025-10-08	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Guzhov, Aleksandr			
