taxonID	type	description	language	source
039F87CAC8437049FF72E59FFD76CD20.taxon	materials_examined	Type species: Cephalochetus indicus Kraatz, 1859, fixed by subsequent designation by Blackwelder 1939: 117.	en	Li, Xiao-Yan, Ma, Hong-Ye, Wang, Lin-Fei, Pan, Zhao-Hui, Lü, Liang (2026): A new species of Cephalochetus Kraatz, 1859 (Coleoptera: Staphylinidae: Paederinae) from Mêdog, China. Zootaxa 5786 (1): 112-118, DOI: 10.11646/zootaxa.5786.1.13, URL: https://doi.org/10.11646/zootaxa.5786.1.13
039F87CAC8437049FF72E59FFD76CD20.taxon	distribution	Distribution. Southeast Asia, Australia, Africa.	en	Li, Xiao-Yan, Ma, Hong-Ye, Wang, Lin-Fei, Pan, Zhao-Hui, Lü, Liang (2026): A new species of Cephalochetus Kraatz, 1859 (Coleoptera: Staphylinidae: Paederinae) from Mêdog, China. Zootaxa 5786 (1): 112-118, DOI: 10.11646/zootaxa.5786.1.13, URL: https://doi.org/10.11646/zootaxa.5786.1.13
039F87CAC843704CFF72E465FC64CCE1.taxon	description	(Figs 1, 2)	en	Li, Xiao-Yan, Ma, Hong-Ye, Wang, Lin-Fei, Pan, Zhao-Hui, Lü, Liang (2026): A new species of Cephalochetus Kraatz, 1859 (Coleoptera: Staphylinidae: Paederinae) from Mêdog, China. Zootaxa 5786 (1): 112-118, DOI: 10.11646/zootaxa.5786.1.13, URL: https://doi.org/10.11646/zootaxa.5786.1.13
039F87CAC843704CFF72E465FC64CCE1.taxon	materials_examined	Type material. HOLOTYPE: CHINA: ♂, Baibung Town [Ä 崩乡], Mêdog County [墨º县], Nyingchi City [ū ĭ Ṅ], 29.2468772 ° N, 95.18382553 ° E, 1059 m, 23 - VII- 2019, sieved from litter in a bamboo grove, leg. Liang LÜ (HBNU). Measurement. BL: 5.73 mm; FL: 3.397 mm; HL: 0.991 mm; AL: 2.150 mm; PL: 1.187 mm; EL: 1.122 mm; HW: 0.926 mm; PW: 1.051 mm; EW: 1.098 mm; ABW: 1.184 mm.	en	Li, Xiao-Yan, Ma, Hong-Ye, Wang, Lin-Fei, Pan, Zhao-Hui, Lü, Liang (2026): A new species of Cephalochetus Kraatz, 1859 (Coleoptera: Staphylinidae: Paederinae) from Mêdog, China. Zootaxa 5786 (1): 112-118, DOI: 10.11646/zootaxa.5786.1.13, URL: https://doi.org/10.11646/zootaxa.5786.1.13
039F87CAC843704CFF72E465FC64CCE1.taxon	description	Description. Habitus as in Fig. 1 A. Head and elytra black or blackish brown; pronotum, abdomen and antennomeres 1 – 6 dark brown; antennomeres 7 – 11 and legs slightly brown. Head (Fig. 1 A, C) dull, longer than wide (HL / HW = 1.07), broadest across eyes. Eyes small, round and prominent, shorter than 1 / 3 of head length (HL / EYL = 3.64). Gena slightly longer than eye and temple together (gena: eye: temple = 1: 0.53: 0.39). Setae restricted to lateral margins. Vertex rough, with sparse small tubercles and slightly elongate-oval depression in middle, longitudinal carina present within fovea. Median area between antennae with triangular glabrous region. Posterior head between eyes with slightly convex and narrow arcuate region bearing punctures. Gena parallel-sided in dorsal view and temples short with both sides protruding laterally. Posterior area of head slightly convex and smooth with coarse punctures. Ventral surface rugose with irregular fine striations. Gular sutures broad in anterior 1 / 3, narrowed posteriorly and parallel and narrow in posterior 2 / 3. Antennae (Fig. 1 A) elongate and slender with antennomere 1 equal to length of 2 – 8, antennomeres 2 – 5 distinctly elongate, 6 – 8 slightly longer than wide, 9 – 10 subcircular, and 11 elongate drop-shaped. Maxillary palpi (Fig. 1 B) long, with palpomere 1 short, 2 – 3 subequal, and 4 small and tuberculate. Pronotum (Fig. 1 C) slightly longer than wide (PL / PW = 1.13), rough and glabrous; broadest approximately at apical 1 / 3, gradually narrowing anteriorly forming semicircular shape, and gradually tapering posteriorly into trapezoidal shape. Margins and broader median area depressed and corrugated, remaining lateral areas shoe-shaped, and slightly convex with punctures small, circular, and dense. Posteromedian area near base with smooth longitudinal elevation approximately 1 / 3 as long as pronotum, and deep median sulcus approximately 1 / 3 as wide as pronotum and deep circular fovea on both sides. Elytra dull, slightly longer than wide (average EL / EW = 1.02); humeri well-developed, slightly expanding posteriorly, with regular lateral margins. Dorsal surface covered with dense, small and circular punctures. Basal portion of the anterior margin glabrous and impunctate; posterolateral angles rugose. Hind wings short, each approximately 1 / 3 as large as elytron, flightless. Abdomen broad and wider than forebody, widest at segment V. Lateral tergites broad; pleurites and tergites fused. Segments III – VII gradually narrowed distally. All tergites with dense setiferous punctation; posterior margins with black setae (Fig. 1 A). Tergite VII (Fig. 2 B) broad and short, apex truncate; tergite VIII (Fig. 2 D) slightly longer than wide, with apex triangular; tergite IX (Fig. 2 E) slender and separate, with apex elongate and acute. Abdominal sternites IV – VI with median area slightly depressed with black setae. Sternite IV (Fig. 1 D) with posterior margin slightly emarginated. Sternite V with circular “ umbilicus ” with cluster of black setae (Fig. 1 D, E). Sternite VII (Fig. 2 A) broad and short, with apex truncate, median area with semicircular emargination, densely setiferous on both sides. Sternite VIII (Fig. 2 C) with deep triangular emargination in middle. Sternite IX (Fig. 2 F) slender, weakly sclerotized at base, gradually tapering distally, with apex truncate. Aedeagus (Fig. 2 G – I) weakly sclerotized, slender and symmetrical, length-to-width ratio = 3.74. Dorsal plate small with median lobe partly unprotected. Ventral plate relatively broad, gradually tapering distally, with apex hooked. Internal sac formless, without obvious sclerites, densely covered with small spines. Female. Unknown.	en	Li, Xiao-Yan, Ma, Hong-Ye, Wang, Lin-Fei, Pan, Zhao-Hui, Lü, Liang (2026): A new species of Cephalochetus Kraatz, 1859 (Coleoptera: Staphylinidae: Paederinae) from Mêdog, China. Zootaxa 5786 (1): 112-118, DOI: 10.11646/zootaxa.5786.1.13, URL: https://doi.org/10.11646/zootaxa.5786.1.13
039F87CAC843704CFF72E465FC64CCE1.taxon	distribution	Distribution. Currently only known from the type locality. The type specimen was sieved from the leaf litter in a bamboo grove in Baibung Town, Mêdog County, Xizang Autonomous Region, Southwest of China, at altitude of 1059 m.	en	Li, Xiao-Yan, Ma, Hong-Ye, Wang, Lin-Fei, Pan, Zhao-Hui, Lü, Liang (2026): A new species of Cephalochetus Kraatz, 1859 (Coleoptera: Staphylinidae: Paederinae) from Mêdog, China. Zootaxa 5786 (1): 112-118, DOI: 10.11646/zootaxa.5786.1.13, URL: https://doi.org/10.11646/zootaxa.5786.1.13
039F87CAC843704CFF72E465FC64CCE1.taxon	etymology	Etymology. The specific epithet honors General Guohua Zhang [张国华], whose sacrifice and contribution, along with that of his men in 1962, made our expedition, as well as any kind of threat-free activities, to the region possible.	en	Li, Xiao-Yan, Ma, Hong-Ye, Wang, Lin-Fei, Pan, Zhao-Hui, Lü, Liang (2026): A new species of Cephalochetus Kraatz, 1859 (Coleoptera: Staphylinidae: Paederinae) from Mêdog, China. Zootaxa 5786 (1): 112-118, DOI: 10.11646/zootaxa.5786.1.13, URL: https://doi.org/10.11646/zootaxa.5786.1.13
039F87CAC843704CFF72E465FC64CCE1.taxon	diagnosis	Diagnosis and notes. Cephalochetus zhangguohuai sp. nov. was found in close geographic proximity to several congeners, including C. kraatzi (Fauvel, 1895), C. aspericollis (Fauvel, 1895), C. brunneus (Motschulsky, 1858), C. rufus, C. apterus Bernhauer, 1928, C. indicus Kraatz, 1859, C. elegans Kraatz, 1859, and C. myrmecocephalus (Lea, 1904). Examination of the original descriptions shows that the posterior margin of sternite VII (often referred to as sternite VI in most works, but as sternite V in Lea 1904) exhibits an emargination of different shapes in C. apterus, C. elegans, and C. myrmecocephalus. In contrast, this feature is absent in C. aspericollis, C. brunneus, C. rufus, and C. indicus (Bernhauer 1928; Fauvel 1895; Kraatz 1859; Lea 1904; Motschulsky 1858). Importantly, none of these species display the distinctive umbilicus-like modification on sternite V that characterizes C. zhangguohuai sp. nov., despite the fact that their abdominal sternites were carefully illustrated in the original descriptions. Furthermore, C. kraatzi was described only from female specimens, but its reported body size (7 mm) is considerably larger than C. zhangguohuai sp. nov. (Fauvel 1895). The new species can be recognized by the combination of following characteristics: a weakly sclerotized aedeagus with notable internal structures, the highly reduced hind wings which limit its ability for long-distance dispersal, and the presence of an “ umbilicus ” on sternite V (Fig. 1 E). We tentatively hypothesize that this structure may be male-specific and could function either as a glandular opening or as a tuft of sensory setae, though its exact role requires further confirmation once female individuals are available.	en	Li, Xiao-Yan, Ma, Hong-Ye, Wang, Lin-Fei, Pan, Zhao-Hui, Lü, Liang (2026): A new species of Cephalochetus Kraatz, 1859 (Coleoptera: Staphylinidae: Paederinae) from Mêdog, China. Zootaxa 5786 (1): 112-118, DOI: 10.11646/zootaxa.5786.1.13, URL: https://doi.org/10.11646/zootaxa.5786.1.13
