taxonID	type	description	language	source
039E8A34CA1DFFDAFF3DFF35FD7FF880.taxon	materials_examined	The first individual corresponds to an adult female (CBUCES-D 108; Fig. 2) found on December 8 th of 2015 at 22: 50 h in “ Camino Alto de Venados ”, vereda El Salado, municipality of Frontino, department of Antioquia, Colombia (6.624816, - 76.270877 WGS 84; 1544 m asl; Fig. 1). This individual was found while exhaustively removing leaf litter at the edge of a forest trail in a dense montane forest. The second individual corresponds to an adult female (MHUA-R 15796; Fig. 3) found on August 11 th of 2023 at 15: 00 h in the “ Reserva ProAves Las Tángaras ”, Alto de Guaduas, municipality of Carmen de Atrato, department of Chocó, Colombia (5.82026, - 76.1764 WGS 84; 1817 m asl; Fig. 1). This individual was found actively foraging on the ground on a cliff, less than one meter high, located on the edge of a rural road in a dense montane forest with trees 20 m high. Both locations were characterized by high amounts of humid leaf litter with various centimeters deep, bromeliads and bryophytes; below the leaf litter the earth was black with a high amount of organic material and roots.	en	Lozano-Arias, Juan M., Gómez-Mesa, Laura, Durango, Juan P., Franco, Esteban Garzón- (2026): Hidden in the leaf litter: rediscovery of the Largescale Lizard, Alopoglossus grandisquamatus (Rueda, 1985) (Squamata: Alopoglossidae) with comments on novel morphological variation. Zootaxa 5757 (4): 369-381, DOI: 10.11646/zootaxa.5757.4.5, URL: https://doi.org/10.11646/zootaxa.5757.4.5
039E8A34CA1DFFDAFF3DFF35FD7FF880.taxon	conservation	Conservation status Based on the type locality and the new localities here provided, the distribution of this species reaches an Extent of Occurrence (EOO) of 823.51 km 2 and an Area of Occupancy (AOO) of 12 km 2. Thus, we suggest that this species should be categorized within the IUCN criteria as Endangered: EN B 1 ac (i) and B 2 ac (ii), due to the following criteria: (1) EOO <5,000 km 2 (2) Number of localities ≤ 5 (3) fluctuation in the EOO and (1) AOO <500 km 2 (2) Number of locations ≤ 5 (3) fluctuation in AOO.	en	Lozano-Arias, Juan M., Gómez-Mesa, Laura, Durango, Juan P., Franco, Esteban Garzón- (2026): Hidden in the leaf litter: rediscovery of the Largescale Lizard, Alopoglossus grandisquamatus (Rueda, 1985) (Squamata: Alopoglossidae) with comments on novel morphological variation. Zootaxa 5757 (4): 369-381, DOI: 10.11646/zootaxa.5757.4.5, URL: https://doi.org/10.11646/zootaxa.5757.4.5
039E8A34CA1DFFDAFF3DFF35FD7FF880.taxon	description	Morphology Measurements of new individuals and holotypes of A. grandisquamatus and A. eurylepis are given in table 1. Generic placement was made based on tongue morphology (Boulenger, 1885; Harris, 1994; Hernández-Morales et al., 2020). Specimens were identified as A. grandisquamatus based on the presence of prefrontal scales, number of transverse dorsal scale rows, absence of pregulars, and dorsal coloration with narrow dorsolateral black stripes and a faint dark vertebral stripe (Rueda, 1985; Harris, 1994). Morphological variation between new specimens of A. grandisquamatus Specimen CBUCES-D 108 differs from MHUA-R 15796 (character states in parenthesis) in: prefrontals in contact only with first supraocular (in contact with first superciliar and first supraocular). Postparietals 2 (4); supraoculars 4 / 4 (3 / 3); suboculars 3 / 3 (4 / 3); postoculars 2 / 2 (1 / 2); geneials — mentonian shields — divided in the posterior margin of the last pair by a small triangular scale (in contact medially along its entire extension) — Fig. 4 B, C —. Transversal rows of gulars 5 - 3 - 2 - 2 - 2 (4 - 1 - 1 - 1 - 1) — Fig. 4 B, C —; collar with a single transverse row of three quadrangular enlarged plates (a single transverse row of two) — Fig. 4 B, C —. Longitudinal rows at mid body 11 (10); scales around mid-body 15 (12); subcaudals 71 (62). Subdigital lamellae on fingers I: 8, II: 8, III: 12, IV: 12 (I: 5, II: 10, III: 12, IV: 13); supradigital lamellae on fingers I: 3, II: 5, III: 6, IV: 7 (I: 1, II: 5, III: 6, IV: 7). Subdigital lamellae on toes I: 5, II: 8, III: 13, IV: 19 (I: 4, II: 9, II: 13, IV: 17); supradigital lamellae on toes I: 2, II: 5, III: 7, IV: 11 (I: 2, II: 4, III: 10, IV: 10). Limbs or fingers overlap on two dorsal scales when adpressed to the body (limbs or fingers do not contact or overlap when adpressed to the body). Both specimens lack femoral pores. In preservation, the dorsal coloration is tan brown with a pair of dark dorsolateral stripes that extend from the back of the head to the sacral region — Fig 2 A, C — (solid dark brown; Fig. 3 A, C). Lateral region of head and neck presenting a series of irregular white spots bordered with dark brown — resembling ocelli — extending to the rear margin of the anterior limbs — Fig. 2 E — (uniformly brown with white stippling; Fig. 3 E). Ventrally, the last third of tail cream with brown irregular stains — Fig. 1 B — (black; Fig. 3 B). Sides of tail with a series of paired ocelli on the first third visible from dorsal view — Fig. 1 A — (uniformly brown; Fig. 3 A). In life, overall dorsal coloration is similar as in preservative. However, CBUCES-D 108 presents a yellowcream ventral coloration with region between chest and vent with an intense orange color, which also covers the ventral surface of hind limbs and extends as stippling towards the first third of the tail (Fig. 5). Morphological variation between new specimens and holotype Holotype of A. grandisquamatus (ICN 5937, adult male) differs from newly recorded specimens (character states in parentheses) in: frontal contacting first three supraoculars (contacting first two); frontoparietals contacting supraoculars 3 and 4 (contacting 2 and 3); parietals heptagonal (hexagonal); postparietals 5 (2 – 4); first superciliar quadrangular (pentagonal); inferior palpebrals 2 (3 / 2); suboculars extend from the fourth supralabial (from the third); postoculars 2 (1 – 2); supralabials 7 (6); infralabials 6 (5); femoral pores 7 + 7 (absent); second supraocular bigger than frontoparietal (as big as frontoparietal); no nasal suture (nasal suture only below the nostril); gulars arranged in 3 - 2 - 1 - 1 - 1 (5 - 3 - 2 - 2 - 2 or 4 - 1 - 1 - 1 - 1) — Fig. 4 A – C —; Table 2. General coloration of specimen CBUCES-D 108 is most like the ICN 5937 (holotype), but it lacks the dark middorsal stripe (Fig. 2 A; Fig. 6 A). All but MHUA-R 15796 have ocelli-like markings on laterals of head and neck (Fig. 2 E; Fig. 3 E; Fig. 6 E). ICN 5937 and MHUA-R 15796 have a dark brown coloration on the distal part of the tail (Fig. 3 B; Fig. 6 B), which is absent in CBUCES-D 108 (Fig. 2 B), but this may be due to the first two specimens having a regenerated tail, causing it to get darker.	en	Lozano-Arias, Juan M., Gómez-Mesa, Laura, Durango, Juan P., Franco, Esteban Garzón- (2026): Hidden in the leaf litter: rediscovery of the Largescale Lizard, Alopoglossus grandisquamatus (Rueda, 1985) (Squamata: Alopoglossidae) with comments on novel morphological variation. Zootaxa 5757 (4): 369-381, DOI: 10.11646/zootaxa.5757.4.5, URL: https://doi.org/10.11646/zootaxa.5757.4.5
