taxonID	type	description	language	source
0392AE51FFF7FFDDFF053979FE41FBE2.taxon	materials_examined	Type species Coriaceolamprodes rubidus sp. nov. Description. Uniformly dark coloured. Male with tegmina and wings completely developed, surpassing abdominal apex. Tegmina in proximal part strongly sclerotised, with subobsolete venation (veins partly replaced with rows of punctures); venation in distal part of tegmina distinct. Legs short; femora and tibiae distinctly flattened; tibial spines well developed. Fore tibiae distally thickened (Fig. 3). Structure of hind tarsi (Fig. 4): metatarsus slightly longer than other tarsal segments combined, and with two unequal rows of spines along lower margin [complete exterior row (Fig. 4, e. r.) having 13 – 17 spines, and incomplete interior row (Fig. 4, i. r.) having six to seven spines and displaced on lateral side of metatarsus]; 1 st – 4 th segments with small (as compared with metatarsus length) apical euplantula on first segment, with comparatively large euplantulae on 2 nd – 4 th segments, and with a pair of “ additional spines ” (one inside and one outside) bordering each euplantula on 1 st – 3 rd segments, and without other spines on 2 nd – 4 th segments (Figs 4, a. s.); claws symmetrical, simple; arolium distinct, about half as long as claw. Fore and mid tarsi similar to hind ones, but with following differences: they shorter; fore tarsi lacking spines; mid tarsi having single short row of five spines along lower margin of metatarsus; and “ additional spines ” on their 4 th segment sometimes absent. Abdomen without visible glandular specialisations. Spiracles of VIII abdominal tergite located on apices of attenuate posterolateral angles (Fig. 5). Anal plate transverse (Fig. 5). Paraprocts of blaberid-type (Fig. 6). Hypandrium nearly symmetrical, with caudal margin very widely rounded and lacking median incision; styli long, cylindrical and well sclerotised (Fig. 7). Male genitalia (Figs 8 – 14). Right phallomere (R + N) with caudal part well sclerotised and subrectangular in shape (Figs 8, 9, c. p. R 1 T), and with cranial part of R 1 T and R 2 widely rounded; R 3 elongated, with branches of unequal length; R 4 weakly sclerotised; R 5 in shape of long sclerotised strip, fused with sclerite R 2; bristles absent. Sclerite L 2 D (L 1) not divided into basal and apical parts, rod-like, very weakly widened cranially (Fig. 10), curved caudally, and with membranous lobe at apex; bristles absent (Fig. 11). Sclerite L 3 (L 2 d) short (Figs 12 – 14); basal subsclerite developed, ring-like (Figs 12, 13, b. L 3); “ folded structure ” absent; apex of sclerite in shape of pressbutton, not hook-like. Sclerite L 4 U (L 3 d) distinct, in shape of curved plate (Fig. 12). Differential diagnosis. Based on the complex of somatic (shape of pronotum, fore femora, tarsi, anal plate and hypandrium) and genital (right phallomere) morphological structures (Princis, 1960; McKittrick, 1964), the genus Coriaceolamprodes gen. nov. belongs to the subfamily Epilamprinae. This genus is similar to the genera Calolamprodes Bey-Bienko, 1969, Pseudocalolampra Roth et Princis, 1971 and Paracalolamprodes Anisyutkin, 2015 in the structure of both tarsi (all their tarsi with two unequal rows of spines along lower margin of tarsal segments; Fig. 4) and right phallomere (sclerite R 3 elongated, sclerite R 4 weakly sclerotised and fused with caudal part of R 1 T; Figs 8, 9). The new genus clearly differs from these genera in the following characters: – from Calolamprodes, in nearly symmetric hypandrium, more sclerotised caudal part of sclerite R 1 T, R 5 fused with R 2, absence of bristles on right phallomere, undivided L 2 D, and shape of L 3 [compare Figs 7, 8 – 14 with figures in Anisyutkin, 1999 (figs 18 – 33, 37), 2006 (figs 6, 12, 14, 15 – 41)]; – from Pseudocalolampra, in the structure of paraprocts (in the latter genus, both paraprocts with hook-shaped caudomedial process), nearly symmetric hypandrium, shape of sclerites R 1 T, R 2 and R 5, undivided L 2 D, and shape of L 3 [compare Figs 6, 7, 8 – 14 with figures in Anisyutkin, 2018 (figs 74 – 86)]; – from Paracalolamprodes, in completely developed tegmina and wings of male, nearly symmetric hypandrium, shape of sclerite R 5, undivid- ed L 2 D, and shape of L 3 [compare Figs 7, 8 – 14 with figures in Anisyutkin, 2015 (figs 7 – 15)]. Included species. Type species only. Etymology. The new name is derived from the Latin adjective coriaceus (leathery) and the part of the taxonomic name Calolamprodes. Gender is masculine. Note. The shape of sclerite L 3 (Figs 12 – 14) of the male genitalia of Coriaceolamprodes gen. nov. is unique for Epilamprinae. This sclerite is not hook-like, as usual in cockroaches, but it rath- er has the shape of a pressbutton. The reduction of L 3 is reported for some Panesthiinae (Roth, 1977). At least many representatives of Panesthiinae have a reclusive habit of life in decaying logs or straight-through burrows (Roth, 1977). Some morphological features of C. rubidus sp. nov. may indicate a hidden mode of live: comparatively short legs and cerci, distally thickened fore tibiae, and sclerotised tegmina. Perhaps this explains the features of the structure of sclerite L 3. The peculiarity of this structure is probably reflects a specialised type of copulation. Unfortunately, at the present time nothing is known about the habit of life of this species.	en	Anisyutkin, L. N. (2019): New genus and species of cockroaches, Coriaceolamprodes rubidusgen. et sp. nov. (Dictyoptera: Blaberidae: Epilamprinae), from the island of Borneo (Sarawak). Zoosystematica Rossica (China) 28 (2): 317-322, DOI: 10.31610/zsr/2019.28.2.317, URL: https://doi.org/10.31610/zsr/2019.28.2.317
0392AE51FFF2FFDCFF0A3996FDD8FC89.taxon	description	(Figs 1 – 14) Holotype. Male, Malaysia, Borneo I., State of Sarawak, ~ 30 km from Bintulu City, Similajau National Park, 3 ° 25´26 ´´ N 113 ° 13´59 ´´ E, primary forest not far from sea, almost at sea level, 12 – 16 Nov. 2016, A. Gorochov, M. Berezin, E. Tkatsheva, I. Kamskov, and N. Grigoreva coll. (ZIN). Paratypes. 4 males, same data as for holotype (ZIN). Description. Male (holotype). General colour reddish brown; eyes black; areas around ocellar spots and anteclypeus pale (Fig. 1); antennae yellowish brown; distal halves of tegmina yellowish; metathorax from above and abdominal tergites yellow. Surfaces smooth and lustrous; distal parts of antennae (from 9 – 10 th segment) dull; facial part of head and pronotum with weak punctuation; facial part of head with weak wrinkles above clypeus (Fig. 1). Head wide and rounded (Fig. 1); ocellar spots small, but distinct; distance between eyes about as long as eye; distance between antennal sockets 2.5 times as long as scape (~ 0.7 mm); approximate lengths of 3 rd – 5 th segments of maxillary palps related to each other as 1.4: 1.0: 1.5. Pronotum transverse (Fig. 2); anterior and lateral margins widely rounded; posterior margin distinctly projected caudally. Anterior margin of fore femur armed as in Fig. 3 (type B), with three spines including one apical spine. Structure of legs and hind tarsi as described for this genus (see above). Anal plate (tergite X) with widely round- ed caudal margin and weak median incision (Fig. 5); caudal part of anal plate separated by distinct transverse furrow (Fig. 5: t. f.). Cerci short, flatten, with segments distinctly separated in distal part. Male genitalia (Figs 8 – 14) as described for this genus (see above). Variations. Head with distance between eyes as 0.8 – 1.0 eye length; distance between antennal sockets as 2.1 – 2.6 scape length (~ 0.7 – 0.9 mm); relative lengths of 3 rd – 5 th segments of maxillary palps approximately 1.1 – 1.4: 1.0: 1.4 – 1.6. Mid metatarsi with single short row consisting of three – five spines; hind metatarsi with ten – seventeen spines in exterior row and with five – seven spines in interior row. Hypandrium sometimes with caudal margin more projected than in holotype. Female unknown. Measurements (in mm). Head length 3.2 – 3.5 (3.4); head width 3.4 – 3.6 (3.5); pronotum length 4.8 – 5.5 (5.0); pronotum width 6.5 – 7.2 (6.9); tegmen length 17.5 – 18.0 (17.8); tegmen width 5.6 – 6.0 (6.0). The measurements in parenthesis are those of the holotype. Etymology. This species name is the Latin adjective rubidus – dark red. Comparison. As given for the genus. Remark. Right hind tarsus of the holotype has only four segments. Probably the fourth tarsomere is absent. The reduction of tarsal segments is usual for the subfamily Epilamprinae.	en	Anisyutkin, L. N. (2019): New genus and species of cockroaches, Coriaceolamprodes rubidusgen. et sp. nov. (Dictyoptera: Blaberidae: Epilamprinae), from the island of Borneo (Sarawak). Zoosystematica Rossica (China) 28 (2): 317-322, DOI: 10.31610/zsr/2019.28.2.317, URL: https://doi.org/10.31610/zsr/2019.28.2.317
