identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03970350FFD6FFFF21DCF0AFD3F06AA2.text	03970350FFD6FFFF21DCF0AFD3F06AA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus quadraticeps Huang & Janák & Zhou & Ge 2026	<div><p>1. Diochus quadraticeps sp. nov.</p><p>(Figs 1D; 2-1; 2-2)</p><p>Type locality. Thailand: Chanthaburi: Khao Sabap Nat. Park .</p><p>Type material. Holotype: male, THAILAND: Chanthaburi: Khao Sabap Nat. Park, 150–300 m, 23–24. XI. 85, Burckhardt-Löbl (MHNG) . Paratypes: THAILAND: Chanthaburi: 18 males, 27 females, same data as holotype (MHNG, JJRC); Chiang Mai: 5 females, Doi Suthep, 1050 m, 5. XI. 1985, Burckhardt-Löbl (MHNG) ; NE Bangkok: 1 male, 2 females, Khao Yai Nat. Park, Khao Khieo, 1150 m, 28. XI. 1985, Burckhardt-Löbl (MHNG) . Holotype with additional labels: “ HOLOTYPE ” and “ Diochus quadraticeps sp. nov. Huang, Janák, Zhou &amp; Ge, T. Huang det. 2025”. Paratypes with additional labels: “ PARATYPE ” and “ Diochus quadraticeps sp. nov. Huang, Janák, Zhou &amp; Ge, T. Huang det. 2025”.</p><p>Measurements. BL = 5.19 mm, FL = 2.39 mm, HL = 0.64 mm, HW = 0.50 mm, EyL = 0.17 mm, TL = 0.36 mm, ANL = 1.46 mm, ANT1 = 0.24 mm, ANT2 = 0.14 mm, ANT3 = 0.16 mm, ANT4 = 0.12 mm, ANT 11 = 0.15 mm, PL = 0.86 mm, PW = 0.66 mm, EL = 0.71 mm, EW = 0.91 mm, SL = 0.63 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, medium-sized. Body brown to light brown, abdominal intersegmental membrane lighter. Legs yellowish brown, tarsi distinctly lighter. Antennae yellowish brown, apex of each antennomere lighter. Maxillary and labial palpi yellowish brown.</p><p>Head (Fig 2-1A). Rounded square in shape, 1.28 times as long as wide. Temples distinctly widened posteriad, posterior angles distinct, nearly in right angle. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye medium-sized, distinctly protruding laterad, longitudinal diameter slightly shorter than half the length of temple (eye: temple = 0.46). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.14 mm) distinctly longer than distance from antenna to eye (0.10 mm). Ventral surface with scattered punctures, interspaces between them as large as about 4 puncture diameters. Gular sutures deep, confluent near the base.</p><p>Antennae (Fig 2-1B). Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 distinctly elongate; antennomere 3 distinctly longer than 2; length of antennomere 4 about twice of width, antennomere 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.</p><p>Mouthparts (Fig 2-1A). Labrum oval-shaped, lateral margin rounded, widest at basal 1/2; width of anterior margin about 1/2 the width at the widest point. Mandibles (Fig 2-1D) falciform, left one with three small teeth, whereas right one with two.</p><p>Neck (Fig 2-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.15 mm, shorter than 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, transverse median furrow not distinct.</p><p>Prothorax (Fig 2-1A). Pronotum elongate (PL to PW ratio 1.30), distinctly longer and wider than head. Anterior region near anterior angles slightly deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures comprised of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards.</p><p>Pterothorax (Fig 2-1A). Mesoscutellum triangular, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; paired oblique furrows short but deep; carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.</p><p>Elytra (Fig 2-1A). Elytra transverse (EL to EW ratio 0.87), shorter but wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 5 punctures on disc, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 2-1C). First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi distinctly longer than 2−4 together, that of meso-, meta- slightly shorter than the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal microsculpture, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male (Figs 2-1E–J; 2-2A–E). Head with small round elevation between eyes. Posterior margin of sternite VIII deeply emarginate medially, also shallowly emarginate at both sides; posterior angles not rounded, but sharped (Figs 2-1E, L; 2-2B). Posterior margin of tergite VIII arcuate (Figs 2-1F; 2-2C). Tergite IX (Figs 2-1H; 2-2D) symmetrical, connected mediobasally, narrowed apically. Sternite IX (Figs 2-1G; 2-2E) symmetrical, widest at basal 1/3 to 1/2; basal margin straight, apical margin slightly emarginated; width of apical margin about half the length of basal margin. Tergite X (Figs 2-1H; 2-2D) symmetrical, triangular. Aedeagus (Figs 2-1 I−J; 2-2A) symmetrical, medium-sized, ca. 0.60 mm, normally sclerotized. Parameres symmetrical, thin, reaching to about apical 1/5 of median lobe. Internal structures sclerotized, paired irregular shaped sclerites and paired rectangular sclerites in middle, in brown to black color; an oval sclerite in black color and an elongated sclerite in brown color; an oval sclerite at bottom, in brown color. Sperm pump coiled, thinned toward apex.</p><p>Female (Figs 2-2 F−I). Head without elevation between eyes. Sternite VIII (Fig 2-2F) with posterior margin rounded. Tergite IX bearing numerous setae, elongate and sharply pointed apically. Sternite IX (Fig 2-2H) symmetrical, with deep concave margin in middle. Tergite X broad, basal margin curved. Sternite X (Fig 2-2G) slender and rhomboidal, with numerous setae in lower half. Tube of spermatheca short and sclerotized, with only one bend (Fig 2-2I).</p><p>Distribution. Thailand.</p><p>Diagnosis. This new species is very similar to D. punctipennis (Motschulsky, 1858) in habitus and also has similar structures in both male aedeagus and female spermatheca. The main difference is the shape of male sternite VIII. Posterior angles of male sternite VIII in this new species are sharp, while in D. punctipennis they are round. In this new species, the inner sclerites of male aedeagus are distinctly shorter than those of D. punctipennis . Moreover, the posterior angles of head in this new species are nearly at right angles, while in D. punctipennis they are evenly rounded.</p><p>Etymology. The species epithet is derived from the Latin word quadratus and - ceps, which indicates the square head.</p></div>	https://treatment.plazi.org/id/03970350FFD6FFFF21DCF0AFD3F06AA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang;Ge, Si-Qin	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang, Ge, Si-Qin (2026): Seven new species of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini). Zootaxa 5768 (4): 451-478, DOI: 10.11646/zootaxa.5768.4.1, URL: http://dx.doi.org/10.7717/peerj.20736
03970350FFD1FFF321DCF69BD2416EFE.text	03970350FFD1FFF321DCF69BD2416EFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus sinuatus Huang & Janák & Zhou & Ge 2026	<div><p>2. Diochus sinuatus sp. nov.</p><p>(Figs 1A; 3-1; 3-2)</p><p>Type locality. Thailand: Mts.n Umphang .</p><p>Type material. Holotype: male, THAILAND: Mts.n Umphang, Mae sot Umphang, 1250 m, 10. 2. 93, Schwendinger (MHNG) . Paratypes: THAILAND: 3 females, same data as holotype (MHNG) . Holotype with additional labels: “ HOLOTYPE ” and “ Diochus sinuatus sp. nov. Huang, Janák, Zhou &amp; Ge, T. Huang det. 2025”. Paratype with additional labels: “ PARATYPE ” and “ Diochus sinuatus sp. nov. Huang, Janák, Zhou &amp; Ge, T. Huang det. 2025”.</p><p>Measurements. BL = 6.09 mm, FL = 2.80 mm, HL = 0.78 mm, HW = 0.58 mm, EyL = 0.19 mm, TL = 0.43 mm, ANT1 = 0.20 mm, ANT2 = 0.12 mm, ANT3 = 0.19 mm, ANT4 = 0.15 mm, PL = 0.96 mm, PW = 0.75 mm, EL = 0.93 mm, EW = 1.06 mm, SL = 0.69 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, medium-sized. Body blackish brown, abdominal intersegmental membrane lighter. Legs dark brown, tarsi distinctly lighter. Antennae dark brown, apex of each antennomere lighter. Maxillary and labial palpi brown.</p><p>Head (Fig 3-1A). Rounded rectangular in shape, 1.34 times as long as wide. Temples slightly widened posteriad, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye medium-sized, slightly protruding laterad, longitudinal diameter slightly shorter than half the length of temple (eye: temple = 0.45). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.15 mm) distinctly longer than distance from antenna to eye (0.10 mm). Ventral surface with scattered punctures, interspaces between them as large as about 4 puncture diameters. Gular sutures deep, confluent near the base.</p><p>Antennae (Fig 3-1B). Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 distinctly elongate; antennomere 3 distinctly longer than 2; length of antennomere 4 about twice of width, antennomere 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.</p><p>Mouthparts (Fig 3-1A). Labrum oval-shaped, lateral margin rounded, widest at basal 1/2; width of anterior margin about 1/2 the width at the widest point. Mandibles (Fig 3-1D) falciform, left one with three small teeth, whereas right one with two.</p><p>Neck (Fig 3-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.17 mm, shorter than 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.</p><p>Prothorax (Fig 3-1A). Pronotum elongate (PL to PW ratio 1.28), distinctly longer and wider than head. Anterior region near anterior angles slightly deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures comprised of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards.</p><p>Pterothorax (Fig 3-1A). Mesoscutellum triangular, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; paired oblique furrows short but deep; carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.</p><p>Elytra (Fig 3-1A). Elytra transverse (EL to EW ratio 0.88), shorter but wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 5 punctures on disc, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 3-1C). First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi distinctly longer than 2−4 together, that of meso-, meta- slightly shorter than the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal microsculpture, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male (Figs 3-1E–I; 3-2A–E). Head with small round elevation between eyes. Posterior margin of sternite VIII shallowly emarginate medially, also with deeper emarginations on both sides; posterior angles sharped (Figs 3-1E; 3-2B). Posterior margin of tergite VIII arcuate (Figs 3-1F; 3-2C). Tergite IX (Figs 3-1H; 3-2D) symmetrical, connected mediobasally, narrowed apically. Sternite IX (Figs 3-1G; 3-2E) symmetrical, widest at basal 1/3 to 1/2; basal margin deeply emarginate, apical margin normally emarginate; width of apical margin about half the length of basal margin. Tergite X (Figs 3H; 3-2D) symmetrical, triangular. Aedeagus (Figs 3-1I; 3-2A) symmetrical, medium-sized, ca. 0.52 mm, normally sclerotized. Parameres symmetrical, in normal length, reaching to about apical 1/5 of median lobe. Internal structures sclerotized, paired rectangular sclerites and symmetrical sclerites in middle, in brown color; paired elongated sclerites on both sides; an elongated sclerite at bottom, in light brown color. Sperm pump coiled, thinned toward apex.</p><p>Female (Figs 3-1J; 3-2F–I). Head without elevation between eyes. Sternite VIII (Fig 3-2F) with posterior margin rounded. Tergite IX bearing numerous setae, elongate and sharply pointed apically. Sternite IX (Fig 3-2H) symmetrical, deeply emarginate at middle. Tergite X broad, basal margin curved. Sternite X (Fig 3-2G) slender and rhomboidal, with numerous setae in lower half. Tube of spermatheca short and sclerotized, with four bends (Figs 3-1J; 3-2I).</p><p>Distribution. Thailand.</p><p>Diagnosis. This new species is similar to Myanmar species D. myanmarensis Huang, Janák &amp; Zhou, 2025 and Thailand species D. bogorensis Huang, Janák &amp; Zhou, 2025 in the waved posterior margin of male sternite VIII. Compared to D. bogorensis, the medial emargination of male sternite VIII in this new species is narrower and aedeagal structures are different: the parameres are distinctly longer in this new species; internal sac with four inner sclerites but six in D. bogorensis . Compared to D. myanmarensis, this new species has smaller eyes, the basal margin of male sternite IX is more deeply emarginate, the internal sac of aedeagus is without paired triangular sclerites on both sides, and the female spermatheca has more bends.</p><p>Etymology. The species epithet is derived from the Latin word sinuatus, which indicates the waved shape of male sternite VIII.</p></div>	https://treatment.plazi.org/id/03970350FFD1FFF321DCF69BD2416EFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang;Ge, Si-Qin	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang, Ge, Si-Qin (2026): Seven new species of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini). Zootaxa 5768 (4): 451-478, DOI: 10.11646/zootaxa.5768.4.1, URL: http://dx.doi.org/10.7717/peerj.20736
03970350FFDEFFF121DCF21CD660684F.text	03970350FFDEFFF121DCF21CD660684F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus moluccanus Huang & Janák & Zhou & Ge 2026	<div><p>3. Diochus moluccanus sp. nov.</p><p>(Figs 1B; 4-1; 4-2)</p><p>Type locality. Indonesia: Moluccas, Kai Besar Island, Turkan .</p><p>Type material. Holotype: female, INDONESIA: INDO MALUC, Kai Besar G, Turkan, 300 m, 7. IX. 1991, Leaf Litter, Agosti (MHNG) . Holotype with additional labels: “HOLOTYPE” and “ Diochus moluccanus sp. nov. Huang, Janák, Zhou &amp; Ge, T. Huang det. 2025” .</p><p>Measurements. BL = 4.74 mm, FL = 1.70 mm, HL = 0.47 mm, HW = 0.39 mm, EyL = 0.12 mm, TL = 0.28 mm, ANL = 1.39 mm, ANT1 = 0.19 mm, ANT2 = 0.12, ANT3 = 0.16 mm, ANT4 = 0.11 mm, ANT 11 = 0.13 mm, PL = 0.60 mm, PW = 0.46 mm, EL = 0.57 mm, EW = 0.68 mm, SL = 0.45 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, medium-sized. Body light brown, abdominal intersegmental membrane lighter. Legs light brown, tarsi distinctly lighter. Antennae brown, apex of each antennomere lighter. Maxillary and labial palpi brown.</p><p>Head (Fig 4-1A). Rounded trapezoid in shape, 1.20 times as long as wide. Temples distinctly widened posteriad, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye small-sized, slightly protruding laterad, longitudinal diameter slightly shorter than half the length of temple (eye: temple = 0.42). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.10 mm) distinctly longer than distance from antenna to eye (0.08 mm). Ventral surface with scattered punctures, interspaces between them as large as about 4 puncture diameters. Gular sutures deep, confluent near the base.</p><p>Antennae (Fig 4-1B). Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 distinctly elongate; antennomere 3 distinctly longer than 2; length of antennomere 4 about twice of width, antennomere 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.</p><p>Mouthparts (Fig 4-1A). Labrum oval-shaped, lateral margin rounded, widest at basal 1/2; width of anterior margin about 1/2 the width at the widest point. Mandibles (Fig 4-1D) falciform, left one with three small teeth, whereas right one with two.</p><p>Neck (Fig 4-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.13 mm, about 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, transverse median furrow not distinct.</p><p>Prothorax (Fig 4-1A). Pronotum slightly elongate (PL to PW ratio 1.29), distinctly longer and wider than head. Anterior region near anterior angles slightly deflexed, lateral margins nearly parallel, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures comprised of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards.</p><p>Pterothorax (Fig 4-1A). Mesoscutellum triangular, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; paired oblique furrows short but deep; carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.</p><p>Elytra (Fig 4-1A). Elytra transverse (EL to EW ratio 0.83), shorter but wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 5 punctures on disc, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 4-1C). First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi distinctly longer than 2−4 together, that of meso-, meta- slightly shorter than the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal microsculpture, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male. Unknown.</p><p>Female (Figs 4-1E−G; 4-2A−D). Head without elevation between eyes. Sternite VIII with posterior margin rounded (Figs 4-1E; 4-2A). Tergite IX bearing numerous setae, elongate and sharply pointed apically. Sternite IX symmetrical, with deep concaved margin in middle (Figs 4-1F; 4-2C). Tergite X broad, basal margin curved. Sternite X slender and rhomboidal, with numerous setae in lower half (Figs 4-1F; 4-2B). Tube of spermatheca sclerotized, in normal length, with five bends (Figs 4G; 4-2D).</p><p>Distribution. Indonesia (Moluccas).</p><p>Diagnosis. According to the body size, light coloration and parallel lateral margin of pronotum, this new species is similar to Thailand species D. bogorensis Huang, Janák &amp; Zhou, 2025 . In this new species, temples are distinctly widened toward base and the posterior angles are more distinct, while in D. bogorensis, temples are nearly parallel. Moreover, the first four segments of protarsi in this new species are distinctly smaller and not dilated, which is also different than in D. bogorensis .</p><p>Etymology. The species epithet is derived from the Latin name of type locality, Moluccas (Indonesia).</p></div>	https://treatment.plazi.org/id/03970350FFDEFFF121DCF21CD660684F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang;Ge, Si-Qin	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang, Ge, Si-Qin (2026): Seven new species of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini). Zootaxa 5768 (4): 451-478, DOI: 10.11646/zootaxa.5768.4.1, URL: http://dx.doi.org/10.7717/peerj.20736
03970350FFDCFFF521DCF3B3D2136BD6.text	03970350FFDCFFF521DCF3B3D2136BD6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus ullrichi Huang & Janák & Zhou & Ge 2026	<div><p>4. Diochus ullrichi sp. nov.</p><p>(Figs 1E; 5-1; 5-2; 5-3)</p><p>Type locality. Papua New Guinea: W. Milds., Baiyer River .</p><p>Type material. Holotype: male, PAPUA NEW GUINEA: W. Milds., Baiyer R., 4000’‚ 6.VII.1974 / S. Peck ber. 281 (CNC) . Paratypes: PAPUA NEW GUINEA: 1 male, 3 females: same data as holotype (CNC, JJRC); 1 female: env. Madang, Nagada, VI. 1979, W. G. Ullrich (MHNG) . Holotype with additional label: “ HOLOTYPUS Diochus ullrichi sp. nov. Huang, Janák, Zhou &amp; Ge, T. Huang det. 2025”. Paratypes with additional labels: “ PARATYPE ” and “ Diochus ullrichi sp. nov. Huang, Janák, Zhou &amp; Ge, T. Huang det. 2025” or “ PARATYPUS Diochus ullrichi Huang, Janák, Zhou &amp; Ge, T. Huang det. 2025”.</p><p>Measurements. BL = 4.8 mm, FL = 2.29 mm, HL = 0.60 mm, HW = 0.48 mm, EyL = 0.15 mm, TL = 0.37 mm, ANL = 1.40 mm, ANT1 = 0.20 mm, ANT2 = 0.12 mm, ANT3 = 0.14 mm, ANT4 = 0.12 mm, ANT 14 = 0.13 mm, PL = 0.87 mm, PW = 0.64 mm, EL = 0.81 mm, EW = 0.91 mm, SL = 0.59 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, small-sized. Body dark brown, elytra sometimes slightly lighter, abdominal intersegmental membrane lighter. Legs brown, tarsi distinctly lighter. Antennae dark brown, apex of each antennomere lighter. Maxillary and labial palpi yellowish brown.</p><p>Head (Fig 5-1A). Rounded square in shape, 1.24 times as long as wide. Temples slightly widened posteriad, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye medium sized, distinctly protruding laterad, longitudinal diameter smaller than half of length of temple (eye: temple = 0.40). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.18 mm) distinctly longer than distance from antenna to eye (0.11 mm). Ventral surface with scattered punctures, interspaces between them about 4 puncture diameters. Gular sutures deep, confluent near the base.</p><p>Antennae (Fig 5-1B). Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 slightly elongate; antennomere 3 nearly equal to 2; length of antennomere 4 about two times as long as wide, 4 to 10 gradually shortened, antennomere 4 about as long as 10; antennomere 11 distinctly longer than 10.</p><p>Mouthparts (Fig 5-1A). Labrum nearly hexagonal, lateral margin straight, widest at basal 1/3 to 1/2; width of anterior margin is about 1/3 the width at the widest point. Mandibles (Fig 5-1D) falciform, left one with three small teeth, whereas right one with two.</p><p>Neck (Fig 5-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.14 mm, shorter than 1/3 of head width. Dorsal surface with distinct groove; Ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.</p><p>Prothorax (Fig 5-1A). Pronotum dark brown, elongate (PL to PW ratio 1.34), distinctly longer and wider than head. Anterior region near anterior angles deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures comprised of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing a transverse ridge between basisternum and furcasternum, angularly backwards.</p><p>Pterothorax (Fig 5-1A). Mesoscutellum triangular, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; the paired oblique furrows short but deep; the carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.</p><p>Elytra (Fig 5-1A). Elytra dark brown, slightly transverse (EL to EW ratio 0.89), distinctly longer and wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, two rows of 4–6 punctures on disc, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 5-1C). First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi slightly longer than 2−4 together, that of meso-, meta- slightly shorter than to the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and transverse microstriae; each tergite with dense yellow pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one possessing only one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal reticulum, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male (Figs 5-1E–K; 5-3A–E). Head with elongate elevation between eyes. Segment VIII very densely and finely setose, without long macrosetae. Sternite VIII (Figs 5-1E–F; 5-3B) with trapezoid emargination on posterior margin, macrosetae thin and short. Tergite VIII (Figs 5-1G; 5-3C) with posterior margin rounded. Sternite IX (Figs 5-1H; 5-3E) symmetrical, widest at basal 1/3 to 1/2; basal margin narrow and rounded, apical margin normally emarginate; width of apical margin about twice as wide as width of basal margin. Tergite X (Figs 5-1I; 5-3D) symmetrical, triangular.Aedeagus (Figs 5-1J–K; 5-3A) symmetrical, medium-sized, ca. 0.59 mm, normally sclerotized. Parameres symmetrical, very short, not reaching to apical half of median lobe. Internal structures strongly sclerotized, central rounded triangular sclerite surrounded by two elongate sclerites situated laterally in middle part, small horse-shoe shaped and trapezoidal central sclerites surrounded by two elongate sclerites on at bottom, all in dark brown color; sclerites less visible due to internal sac covered densely with dark more or less elongate fine scales forming also a small dark subrounded form proximally to first sclerites; apical part of median lobe with two elongate projections in light yellowish color, markedly darkened on inner side.</p><p>Female (Figs 5-2A−D; 5-3F−I). Smaller than males (FL = 2.0– 2.1 mm). Head without elevation between eyes. Sternite VIII (Fig 5-3F) with posterior margin rounded. Tergite IX bearing numerous setae, elongate and sharply pointed apically. Sternite IX (Fig 5-3H) symmetrical, with deep concave margin in middle. Tergite X broad, basal margin curved. Sternite X (Fig 5-3G) slender and rhomboidal, with numerous setae in lower half. Tube of spermatheca sclerotized, very long and strongly coiled (Figs 5-2A−D; 5-3I).</p><p>Distribution. Papua New Guinea.</p><p>Diagnosis. Compared to other species from Papua New Guinea, this new species is similar to D. chambrieri Huang, Janák &amp; Zhou, 2024 in appearance. The main differences are the shape of male sternite VIII, sternite X and aedeagi. In the new species, the medial emargination is trapezoid, while in D. chambrieri it is more rounded. The basal margin of sternite X in the new species is distinctly narrower than in D. chambrieri . In this new species, a small dark subrounded form proximally to the first sclerites of aedeagus is present, but absent in D. chambrieri . Although the spermatheca is also similarly shaped in both species, the dilated part of the tube is shorter and thicker in this new species.</p><p>Variability. Variability between males and females (from both type localities) are observed: females are slightly smaller (FL = 2.0– 2.1 mm) with less elongate head (about 1.2 times as long as wide).</p><p>Etymology. The species epithet is dedicated to the collector of a specimen of the type series, Mr. W. G. Ullrich (Switzerland).</p></div>	https://treatment.plazi.org/id/03970350FFDCFFF521DCF3B3D2136BD6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang;Ge, Si-Qin	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang, Ge, Si-Qin (2026): Seven new species of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini). Zootaxa 5768 (4): 451-478, DOI: 10.11646/zootaxa.5768.4.1, URL: http://dx.doi.org/10.7717/peerj.20736
03970350FFD8FFE821DCF124D7C76E0E.text	03970350FFD8FFE821DCF124D7C76E0E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus madangensis Huang & Janák & Zhou & Ge 2026	<div><p>5. Diochus madangensis sp. nov.</p><p>(Figs 1G; 6-1; 6-2)</p><p>Type locality. Papua New Guinea: env. Madang, Nagada .</p><p>Type material. Holotype: female, PAPUA NEW GUINEA: env. Madang, Nagada, VI. 1979, W. G. Ullrich (specimens identified as D. papuanus by ??) (MHNG) . Paratype: PAPUA NEW GUINEA: 1 female, Madang Dist., Finisterre Mts. Damanti 3,550 ft 2.-11.x.1966, M.E. Bacchus BM 1965-120 (NHMUK). Holotype with additional labels: “HOLOTYPE” and “ Diochus madangensis sp. nov. Huang, Janák, Zhou &amp; Ge, T. Huang det. 2025”. Paratype with additional label: “ PARATYPUS Diochus madangensis sp. nov. Huang, Janák, Zhou &amp; Ge, T. Huang det. 2025”.</p><p>Measurements. BL = 3.3 mm, FL = 1.81 mm, HL = 0.51 mm, HW = 0.44 mm, EyL = 0.14 mm, TL = 0.29 mm, ANT1 = 0.14 mm, ANT2 = 0.11 mm, ANT3 = 0.11 mm, ANT4 = 0.08 mm, PL = 0.66 mm, PW = 0.55 mm, EL = 0.64 mm, EW = 0.70 mm, SL = 0.48 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, small-sized. Body dark brown, abdominal intersegmental membrane lighter.Legs brown, tarsi distinctly lighter.Antennae dark brown, apex of each antennomere lighter. Maxillary and labial palpi yellowish brown.</p><p>Head (Fig 6-1A). Rounded square in shape, 1.16 times as long as wide. Temples distinctly widened posteriad, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye medium-sized, distinctly protruding laterad, longitudinal diameter about half of length of temple (eye: temple = 0.48). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.13 mm) distinctly longer than distance from antenna to eye (0.07 mm). Ventral surface with scattered punctures, interspaces between them about 4 puncture diameters. Gular sutures deep, confluent near the base.</p><p>Antennae (Fig 6-1B). Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 slightly elongate, antennomere 3 nearly equal to 2; length of antennomere 4 about 1.5 times as long as wide.</p><p>Mouthparts (Fig 6-1A). Labrum nearly hexagonal, lateral margin straight, widest at basal 1/3 to 1/2; width of anterior margin is about 1/3 the width at the widest point. Mandibles (Fig 6-1D) falciform, left one with two small teeth, basal one long and sharp. Right mandible with two small teeth.</p><p>Neck (Fig 6-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.15 mm, shorter than 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.</p><p>Prothorax (Fig 6-1A). Pronotum dark brown, elongate (PL to PW ratio 1.20), distinctly longer and wider than head. Anterior region near anterior angles deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures comprised of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing a transverse ridge between basisternum and furcasternum, angularly backwards.</p><p>Pterothorax (Fig 6-1A). Mesoscutellum triangular, surface shiny, bearing distinct microstriae, but without any punctures.</p><p>Elytra (Fig 6-1A). Elytra dark brown, slightly transverse (EL to EW ratio 0.91), distinctly longer and wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 5 punctures on disc, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 6-1C). First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi slightly longer than 2−4 together, that of meta- slightly shorter than the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and transverse microstriae; each tergite with dense yellow pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal reticulum, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male. Unknown.</p><p>Female (Figs 6-1E−G; 6-2A−D). Head without elevation between eyes. Sternite VIII (Figs 6-1E; 6-2A) with posterior margin rounded. Tergite IX bearing numerous setae, elongate and sharply pointed apically. Sternite IX (Figs 6-1F; 6-2C) symmetrical, with deep concave margin in middle. Tergite X broad, basal margin curved. Sternite X (Figs 6-1F; 6-2B) slender and rhomboidal, with numerous setae in lower half. Tube of spermatheca sclerotized and long, weakly coiled (Figs 6-1G; 6-2D).</p><p>Distribution. Papua New Guinea.</p><p>Diagnosis. According to the small size, this new species is similar to D. loebli and D. ullrichi sp. nov. from Papua New Guinea. Compared to D. loebli, the tube of spermatheca of this species is distinctly shorter and the dilated part of the tube is also shorter. Compared to D. ullrichi sp. nov., the tube of spermatheca of this species is distinctly shorter, and not so coiled. Moreover, the new species can be distinguished from other species by the long and sharp tooth near the base of the left mandible.</p><p>Etymology. The species epithet derived from the type locality: Madang (Papua New Guinea).</p></div>	https://treatment.plazi.org/id/03970350FFD8FFE821DCF124D7C76E0E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang;Ge, Si-Qin	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang, Ge, Si-Qin (2026): Seven new species of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini). Zootaxa 5768 (4): 451-478, DOI: 10.11646/zootaxa.5768.4.1, URL: http://dx.doi.org/10.7717/peerj.20736
03970350FFC5FFEC21DCF587D76268CF.text	03970350FFC5FFEC21DCF587D76268CF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus insularis Huang & Janák & Zhou & Ge 2026	<div><p>6. Diochus insularis sp. nov.</p><p>(Figs 1F; 7-1; 7-2; 7-3)</p><p>Type locality. China (Taiwan): Taitung Hsien, Hsinkangshan above Chengkang.</p><p>Type material. Holotype: male: CHINA: Taiwan: “TAIWAN, Taitung Hsien, Hsinkangshan above Chengkang 550-600m 22.IV.98, A. Smetana [T187]” (CNC). Paratypes: CHINA: Taiwan: 3 females: same data as holotype (CNC, JJRC); 3 males, 3 females: Taichung Hsien, Wufeng, Wushi (Riv.) 100m, 21.IV.90, A. Smetana [T13] (CNC, JJRC); 1 female: Taitung Hsien, Foothills of Hsingkangshan nr. Chengkung, 350-400m, 19.VII.93, A. Smetana [T155] (CNC); 1 female: Taitung Hsien, Hsinkangshan above Chengkung 800m 27.IV.1995, A. Smetana [T168] (CNC); 1 male, 1 female: Kankau (Koshun), Formosa, H. Sauter VII.1909 (specimens identified as D. antennatus by Bernhauer) (SDEI); 1 male: Formosa, Takezaki near Mt. Ari, 12-12-1937, Coll. Yosio Yano (BMNH); 1 male: Formosa, Mt. Yangming, 22.VII.1966, T. Kobayashi (YHKJ); 1 male, 2 females: Taipei Co., Parc TianMulu, Beitou, Shipai Rd., date stamping litter, 30.XII.2008, leg. Vít (VAHG, JJRC), 1 female: Taipei Co., Yangmingshan, N Beitou, Shamao Rd., 250 m, decaying tree trunk, 23.X.2008, leg. Vít (VAHG). Hong Kong: 1 female: # 2 of Hong Kong, Ping Shan Chai, 14-29.vi.2014., Leg: C. Barthelemy. Malaise Trap (GROG); 1 female: # 7 of Hong Kong, Ping Shan Chai, 12-26.vii.2014., Leg: C. Barthelemy. Malaise Trap ” (JJRC). JAPAN: 1 male: Okinava Pref ., Ishigaki Jima, W-coast, Mt. Jarabu, dry litter, 26.XII.2008, leg. S. Vit (VAHG). MAURITIUS [Ile Maurice]: 1 female: Grand’ Baie, 17. V. 77, I. Sieber (specimens identified as D. punctipennis by ???) (MHNG); 1 male, 2 females: SE Mauritius, Dalais, Riv des Créoles, 10 m, 20°23ʼ44ʼʼS, 57°41ʼ03ʼʼE, 14.1.2006, river bank, sifting, J. Janák lgt. (JJRC); 1 male, 1 female: Albion, 8.V.70, C. M. Courtois (MNHNP). RÉUNION: 1 male: Ravine de St. Gilles, Bassin Cormoran, 13.- 14.1.1992, J. Janák lgt. (JJRC); 1 male: Ravine de St. Gilles, Bassin Cormoran, 22.- 23.1.1992, J. Janák lgt. (JJRC); 1 male: St. Gilles le B., tamisages, Y. Gomy (MNHNP); 1 female, Ravine S. Bassins, Y. Gomy (MNHNP); 1 male: St. Paul, Mafate, Bras Ste. Suzanne, 25-VII-1951, J. Hamon (MNHNP). RODRIGUEZ: 4 males, 1 female: Caverne Patate, Pl. Corail, 11.V.72, Y. Gomy (MNHNP). SEYCHELLES/ MAURITIUS: 1 female: Mahé (Séchelles), Curepipe (Maurice) (specimen with additional labels: “duplicatus Fvl.” [unpublished name] (IRSNB). Holotype with additional label: “ HOLOTYPUS Diochus insularis sp. nov. J. Janák det. 2017”. Paratypes with additional labels: “ PARATYPUS Diochus insularis sp. nov. J. Janák det. 2017” or “ PARATYPE ” and “ Diochus insularis sp. nov. Huang, Janák, Zhou &amp; Ge, T. Huang det. 2025”.</p><p>Measurements. BL = 4.30 mm, FL = 2.24 mm, HL = 0.59 mm, HW = 0.48 mm, EyL = 0.15 mm, TL = 0.34 mm, ANL = 1.42 mm, ANT1 = 0.21 mm, ANT2 = 0.14, ANT3 = 0.16 mm, ANT4 = 0.12 mm, ANT 11 = 0.20 mm, PL = 0.78 mm, PW = 0.64 mm, EL = 0.76 mm, EW = 0.81 mm, SL = 0.56 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, small-sized. Body brown, pronotum and tip of abdomen often reddish brown, abdominal intersegmental membrane lighter. Legs light brown, tarsi distinctly lighter. Antennae brown, apex of each antennomere lighter. Maxillary and labial palpi brown.</p><p>Head (Fig 7-1A). Rounded trapezoid in shape, 1.23 times as long as wide. Temples distinctly widened posteriad, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye medium-sized, distinctly protruding laterad, longitudinal diameter about half of length of temple (eye: temple = 0.44). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.13 mm) distinctly longer than distance from antenna to eye (0.07 mm). Ventral surface with scattered punctures, interspaces between them about 4 puncture diameters. Gular sutures deep, confluent near the base.</p><p>Antennae (Fig 7-1B). Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 distinctly elongate, antennomere 3 slightly longer than 2; length of antennomere 4 about 1.5 times as long as wide, 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.</p><p>Mouthparts (Fig 7-1A). Labrum oval-shaped, lateral margin rounded, widest at basal 1/2; width of anterior margin is about half the width at the widest point. Mandibles (Fig 7-1D) falciform, left one with three small teeth, with basal one long and sharp. Right mandible with two small teeth.</p><p>Neck (Fig 7-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.17 mm, shorter than 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.</p><p>Prothorax (Fig 7-1A). Pronotum dark brown, elongate (PL to PW ratio 1.22), distinctly longer and wider than head. Anterior region near anterior angles deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures comprised of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing a transverse ridge between basisternum and furcasternum, angularly backwards.</p><p>Pterothorax (Fig 7-1A). Mesoscutellum triangular, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; the paired oblique furrows short but deep; the carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.</p><p>Elytra (Fig 7-1A). Elytra dark brown, slightly transverse (EL to EW ratio 0.93), distinctly longer and wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 5 punctures on disc, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 7-1C). First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi distinctly longer than 2−4 together, that of meso-, meta- slightly shorter than to the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and transverse microstriae; each tergite with dense yellow pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal reticulum, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male (Figs 7-1E−K; 7-3A−E). Head with elongate elevation between eyes. Sternite VIII broadly rounded or very slightly almost invisible emarginate medially on posterior margin (Figs 7-1E–F; 7-3B). Tergite VIII (Figs 7-1G; 7-3C) with posterior margin rounded. Sternite IX (Figs 7-1H; 7-3E) symmetrical, widest at basal 1/3; basal margin deeply emarginate, apical margin normally emarginate; width of apical margin about 0.8 as wide as width of basal margin. Tergite X (Figs 7-1I; 7-3D) symmetrical, triangular. Aedeagus (Figs 7-1J–K; 7-3A) symmetrical, medium-sized, ca. 0.53 mm long, weakly sclerotized. Parameres symmetrical, very short, reaching about to apical half of median lobe. Internal structures strongly sclerotized, paired half circle sclerite in middle, two elongate sclerites on both sides connected with elongate triangular central sclerite at bottom, all in brown color.</p><p>Female (Figs 7-2A−C; 7-3 F−I). Head without elevation between eyes. Sternite VIII (Fig 7-3F) with posterior margin rounded. Tergite IX bearing numerous setae, elongate and sharply pointed apically. Sternite IX (Fig 7-3H) symmetrical, with deep concave margin in middle. Tergite X broad, basal margin curved. Sternite X (Fig 7-3G) slender and rhomboidal, with numerous setae in lower half. Tube of spermatheca sclerotized, very thick and short, with only one bend; apical dilated lobe large (Figs 7-2A−C; 7-3I).</p><p>Distribution. Widely distributed species, currently known from China (Taiwan, Hong Kong), Mauritius (Grand Baie), Reunion, Rodriguez and Japan (Okinawa). Records of D. punctipennis (Motschulsky, 1858) in Mascarene Islands (Reunion, Mauritius) recorded by various authors (e.g. Lecoq, 1990; Gomy, 2000; Janák, 2014) refers to D. insularis sp. nov.</p><p>Diagnosis. This new species can be easily distinguished from congeners by the combination of: small size; pale color; the very slightly emarginate posterior margin of the male sternite VIII, male aedeagus with paired, semilunar sclerites in middle, and elongate triangular central sclerite basally; and the thick and short tube of female spermatheca.</p><p>Etymology. The species epithet derived from its distribution on various islands.</p></div>	https://treatment.plazi.org/id/03970350FFC5FFEC21DCF587D76268CF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang;Ge, Si-Qin	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang, Ge, Si-Qin (2026): Seven new species of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini). Zootaxa 5768 (4): 451-478, DOI: 10.11646/zootaxa.5768.4.1, URL: http://dx.doi.org/10.7717/peerj.20736
03970350FFC1FFE021DCF040D2356E22.text	03970350FFC1FFE021DCF040D2356E22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus parvoculatus Huang & Janák & Zhou & Ge 2026	<div><p>7. Diochus parvoculatus sp. nov.</p><p>(Figs 1C; 8-1; 8-2)</p><p>Type locality. Peru: Loreto: Rio Mazan .</p><p>Type material. Holotype: male, PERU: Loreto: Rio Mazan, 10. X. 80, Cl. Vaucher (MHNG) . Paratype: PERU: Loreto: 1 female, same data as holotype (MHNG) . Holotype with additional labels: “ HOLOTYPE ” and “ Diochus parvoculatus sp. nov. Huang, Janák, Zhou &amp; Ge, T. Huang det. 2025”. Paratype with additional labels: “ PARATYPE ” and “ Diochus parvoculatus sp. nov. Huang, Janák, Zhou &amp; Ge, T. Huang det. 2025”.</p><p>Measurements. BL = 5.56 mm, FL = 2.29 mm, HL = 0.68 mm, HW = 0.50 mm, EyL = 0.15 mm, TL = 0.41 mm, ANL = 1.27 mm, ANT1 = 0.21 mm, ANT2 = 0.11 mm, ANT3 = 0.16 mm, ANT4 = 0.10 mm, ANT 11 = 0.15 mm, PL = 0.84 mm, PW = 0.67 mm, EL = 0.80 mm, EW = 0.82 mm, SL = 0.63 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, medium-sized. Body reddish brown to brown, pronotum yellowish brown, abdominal intersegmental membrane lighter. Legs yellowish brown, tarsi distinctly lighter. Antennae brown, apex of each antennomere lighter. Maxillary and labial palpi yellowish brown.</p><p>Head (Fig 8-1A). Rounded trapezoid in shape, 1.34 times as long as wide. Temples distinctly widened posteriad, posterior angles distinct. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye small-sized, distinctly protruding laterad, longitudinal diameter distinctly shorter than half the length of temple (eye: temple = 0.38). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.14 mm) distinctly longer than distance from antenna to eye (0.10 mm). Ventral surface with scattered punctures, interspaces between them as large as about 4 puncture diameters. Gular sutures deep, approaching each other near the base, but not confluent.</p><p>Antennae (Fig 8-1B). Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 distinctly elongate; antennomere 3 distinctly longer than 2; antennomere 4 about 1.5 times as long as wide, antennomere 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.</p><p>Mouthparts (Fig 8-1A). Labrum oval-shaped, lateral margin rounded, widest at basal 1/2; width of anterior margin about 1/2 the width at the widest point. Mandibles falciform, left one with three small teeth, whereas right one with two.</p><p>Neck (Fig 8-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.13 mm, shorter than 1/3 of head width. Dorsal surface without groove; ventral surface with gular sutures forming obvious triangular region, with distinct transverse median furrow.</p><p>Prothorax (Fig 8-1A). Pronotum elongate (PL to PW ratio 1.25), distinctly longer and wider than head. Anterior region near anterior angles slightly deflexed, lateral margins widened posteriad to basal 1/5 (widest at 1/5), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures comprised of 5 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins).</p><p>Pterothorax (Fig 8-1A). Mesoscutellum triangular, surface shiny, bearing distinct microstriae, but without any punctures.</p><p>Elytra (Fig 8-1A). Elytra slightly transverse (EL to EW ratio 0.98), shorter but wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with numerous irregular punctures, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 8-1C). First four segments of protarsi heart-shaped, strongly dilated, and those of meso- and metatarsi slender. Last segment of protarsi distinctly longer than 2−4 together, that of meso-, meta- slightly shorter than the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal microsculpture, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male (Figs 8-1D–H; 8-2A–D). Head without elevation between eyes. Posterior margin of sternite VIII slightly emarginate medially; posterior angles rounded (Figs 8-1D; 8-2B). Posterior margin of tergite VIII arcuate. Tergite IX (Figs 8-1F; 8-2C) symmetrical, connected mediobasally, narrowed apically. Sternite IX (Figs 8-1E; 8-2D) symmetrical, widest at basal 1/3; basal margin straight, apical margin slightly emarginate; width of apical margin slightly shorter than width of basal margin. Tergite X (Figs 8-1F; 8-2C) symmetrical, triangular. Aedeagus (Figs 8-1G−H; 8-2A) not symmetrical, large-sized, ca. 1.03 mm long, normally sclerotized. Parameres symmetrical and long, not reaching apex of median lobe, widened at apex. Internal structures sclerotized, apical coiled duct with only one coil; seminal vesical distinctly elongate, about half length of median lobe.</p><p>Female (Figs 8-1I; 8-2E−I). Head without elevation between eyes. Sternite VIII (Fig 8-2F) with posterior margin rounded. Tergite IX (Fig 8-2E) bearing numerous setae, elongate and sharply pointed apically. Sternite IX (Fig 8-2I) symmetrical, with deep concave margin in middle. Tergite X (Fig 8-2E) broad, basal margin curved. Sternite X (Fig 8-2H) slender and rhomboidal, with numerous setae in lower half. Coiled duct of spermatheca very long and strongly coiled, with bursa copulatrix in central position (Fig 8-1I).</p><p>Species group. According to the key to species groups for Neotropical Diochus by Irmler, 2017, this new species should be included in D. inornatus -group.</p><p>Distribution. Peru.</p><p>Diagnosis. Aedeagus structures of this new species are similar to D. hibbsi Irmler, 2017 (described from Ecuador) in D. inornatus -group. Both have apical coiled duct with only one coil (size of the coil is smaller in the new species), and the tips of parameres in these two species are also of similar shape. The main differences are as follows: body is distinctly larger in this new species; head is nearly rectangular in this new species, with posterior angles distinct, (posterior angles are rounded in D. hibbsi); eyes are smaller in this new species; pronotum is unicolored; antennal segments are wider; seminal vesical of this new species is distinctly elongate (rounded in D. hibbsi).</p><p>Etymology. The species epithet is derived from the Latin word parvus and oculatus, which indicates the small-sized eyes.</p></div>	https://treatment.plazi.org/id/03970350FFC1FFE021DCF040D2356E22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang;Ge, Si-Qin	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang, Ge, Si-Qin (2026): Seven new species of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini). Zootaxa 5768 (4): 451-478, DOI: 10.11646/zootaxa.5768.4.1, URL: http://dx.doi.org/10.7717/peerj.20736
