taxonID	type	description	language	source
03969E197F76322642AA31ADFE7EFAA6.taxon	description	Description (emended) Medium- to large-sized miliolid composed of a spherical proloculus followed by an undivided deuteroloculus with different types of coiling: entirely glomospiral, initially glomospiral and then planispiral (or aligned) involute, glomospiral becoming planispiral evolute or semi-involute, glomospiral then irregular planispiral involute / semi-evolute (?), initially quinqueloculinoid then planispiral involute or evolute. The following chamber is semicircular in axial section and sometimes with flosculinisations. Pseudosepta present in few genera. Thick porcelanaceous wall with buttresses and sometimes with perforations similar to comb teeth-shaped structure. Aperture terminal, simple.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F76322740243660FDC5FE76.taxon	description	Original description Neodiscidae medium - to large-sized, undivided or with weak pseudosepta, involute or with a short evolute terminal stage of coiling, and devoid of flosculinisation.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F76322740243660FDC5FE76.taxon	discussion	Remarks In the original description, the genus Praeneodiscus is tentatively included in the family Neodiscidae by Vachard et al. (2015). After the acceptance of the original taxonomic position of Praeneodiscus under the family Neodiscidae by Vachard (2018), it was later transferred to the subfamily Cornuspirinae Schultze, 1854 nom. translat. Rhumbler 1904 within the family? Cornuspiridae Schultze, 1854 by Krainer et al. (2019). Recently, the genus Praeneodiscus was placed tentatively in the subfamily Hemigordiinae Reitlinger in Vdovenko et al., 1993 (nomen translat. Pronina 1994 pro family) of the family Hemigordiidae Reitlinger in Vdovenko et al. (1993) by Vachard & Krainer (2022). In this study, even the taxonomic status of the genus Praeneodiscus is still controversial, it is concluded that it should be included in the subfamily Neodiscinae due to its test shape, coiling type and devoid of flosculinisation.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F773227433E3084FDE7FA9C.taxon	type_taxon	Type species Agathammina pusilla (Geinitz, 1848) (type by subsequent designation).	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F773227433E3084FDE7FA9C.taxon	description	Description (emended) Test ovate, globular proloculus followed by enrolled undivided tubular second chamber of few gradually enlarging whorls, initially quinqueloculinoid then planispiral involute sometimes with evolute last whorls without septation; wall calcareous, porcelaneous with perforations similar to comb teeth-shaped structure. Aperture terminal, simple.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F77322743313153FC02FC03.taxon	description	Description (emended) Medium-sized miliolid with spherical proloculus followed by a tubular chamber, initially quinqueloculinoid then planispiral involute, sometimes with evolute last whorls. The tubular chamber sometimes shows pseudosepta. Porcelanaceous wall with perforations similar to comb teeth-shaped structure. Aperture terminal, simple.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F7732244041360BFD4FFC64.taxon	description	urn: lsid: zoobank. org: act: 9 BC 6 E 72 C-D 473 - 47 DD- 9681 - 51754 BC 42 BAE	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F7732244041360BFD4FFC64.taxon	type_taxon	Type genus Globidiscus Okuyucu, 2021.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F7732244041360BFD4FFC64.taxon	description	Description Large miliolid tests composed of a spherical proloculus followed by an undivided tubular chamber, diversely coiled: initially glomospiral and finally planispiral involute, initially glomospiral then irregular planispiral involute / semi-evolute (?). Sometimes one or two sided ogival shape of the last whorls. No septa or pseudosepta. Lumen with common flosculinisation. The thick wall with perforations similar to comb teeth-shaped structure. Neosparitization rare to common. Umbilicus slightly depressed or absent. Aperture terminal, simple.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F7732244041360BFD4FFC64.taxon	discussion	Comparison Globidiscinae differ from Neodiscinae by the test shape of the last whorls which has one- or two-sided ogival-shaped last whorls, and is of a coiling type initially glomospiral and finally planispiral involute, initially glomospiral then irregular planispiral involute / semi-evolute (?) and with common flosculinisation; from Baisalininae and Agathammininae by a larger test and devoid of septa / pseudosepta; and from Multidiscinae by a test which has one- or two-sided ogival-shaped last whorls, a larger test and common flosculinisation. Remarks In the original description of the genus Kamurana it was tentatively attributed to the family Hemigordiopsidae Nikitina, 1969 instead of Milioliporidae Brönnimann & Zaninetti, 1971 by Altiner & Zaninetti (1977) due to the probable absence of septation and a homeomorph of the irregular forms of the genus Hemigordius Schubert, 1908. Besides, as indicated by the authors, the presence of perforations in the wall at the adult stage of the genus Kamurana makes its attribution to the Hemigordiopsidae problematic. In this study, after a detailed investigation of the wall structure of the genus Kamurana in the original description compared with recovered well-preserved material, it is clearly shown that the wall of Kamurana does not feature a perforation structure in the adult stage in the sense of authors. This structure is clearly comb teeth-shaped which developed on the porcelaneous wall related to the neosparitization. For this reason, the taxonomic position of the genus Kamurana should be located under the subfamily Globidiscinae subfam. nov.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F743224438030A2FD19F908.taxon	type_taxon	Type species Globidiscus fragilis Okuyucu, 2021 (type by original designation).	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F743224438030A2FD19F908.taxon	description	Description (emended) Test free, very large, bilocular, involute, inflated, lenticular in axial section and subglobular in transverse section with tapering last whorls, sometimes with sharp triangular protrusions. Coiling initially glomospiral and planispiral in the terminal part. A well-developed flosculinisation fills the base of the lumen and generally exceeds the two-thirds height of the chamber. Thick porcelaneous wall with perforations (comb teeth-shaped) related to neosparitization and without buttresses. Aperture simple, terminal.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F743224438030A2FD19F908.taxon	distribution	Range and distribution Discovered in the Middle Permian (late Capitanian) of the Karakaya Complex (= Elmadağ Olistrostrome, this study), northwest Anatolia, Türkiye.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F74322543CA359FFD9DFAB2.taxon	type_taxon	Type species Kamurana bronnimanni Altiner & Zaninetti, 1977 (type by original designation).	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F74322543CA359FFD9DFAB2.taxon	description	Description (emended) Test free, large and spherical to ovoid, formed of a globular proloculus and a undivided tube. Coiling initially glomospiral then irregular planispiral involute / semi-evolute (?). Thick porcelaneous wall often neosparitized with perforations (comb teeth-shaped). Aperture simple, terminal.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F74322543CA359FFD9DFAB2.taxon	discussion	Remarks Compared to the type species of Kamurana (Kamurana bronnimanni), the position of K. chatalovi Trifonova, 1984 from the Lower Triassic strata in the Sveti Iliya Ridge, southeastern Bulgaria is more controversial. Additionally, Trifonova (1984) reassigned Kamurana, previously considered in the family Hemigordiopsidae, to the family Milioliporidae due to its coarse perforation on the wall in the adult stage. Besides, she also noted that her new species differed from K. bronnimanni by its phylogenetically reduced first volutions, whereas K. chatalovi is characterized by the first 2 – 3 irregular coils compared to about 12 glomospirid coils in K. bronnimanni. As indicated above by Trifonova (1984), the typical coiling type of Kamurana (glomospirid-type with more initial volutions) is not observed in the specimens of K. chatalovi. It is also clear that the perforation of the wall in K. bronnimanni (sensu Altiner & Zaninetti 1977) is not a typical structural element (e. g., septa), whereas it is a structure formed by the neosparatization of the thick porcelanous wall. For this reason, K. chatalovi of Trifonava (1984) is excluded from the genus Kamurana in this study. In the original study, Altiner & Zaninetti 1977 stated that the taxonomic position of Kamurana was provisionally based on the difference in the descriptions of the linked families. For the reasons detailed above, the genus Kamurana is included in the Globidiscinae subfamily, which includes many genera with similar characteristics.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F74322543CA359FFD9DFAB2.taxon	distribution	Range and distribution Kamurana bronnimanni was described from the Upper Permian (Upper Dzhulfian) of the Eastern Taurus, Türkiye. Discovered in the Middle Permian (late Capitanian) (this study) of the Elmadağ Olistostrome, Ankara City, Central Türkiye.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F75322A402D3615FA8DFC8E.taxon	description	urn: lsid: zoobank. org: act: 53 AA 5 ECF- 1580 - 4176 - BF 8 A- 9 C 4 BE 232 DCD 5	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F75322A402D3615FA8DFC8E.taxon	type_taxon	Type species Rectogordiopsis kamuranaeformis gen. et sp. nov.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F75322A402D3615FA8DFC8E.taxon	etymology	Etymology From the Latin ‘ rectus ’ (= ‘ straight’) because of the many planispiral coils and the Latin ‘ gordiopsis ’ because of the similarity to the genus Hemigordiopsis Reichel, 1945.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F75322A402D3615FA8DFC8E.taxon	description	Description Test free, very large and globular. Proloculus spherical followed by a tubular undivided chamber. First whorls numerous tightly streptospiral and following whorls planispiral involute. Flosculinisation well developed. Thick porcelaneous wall often neosparitized with perforations (comb teeth-shaped). Aperture simple terminal.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F75322A402D3615FA8DFC8E.taxon	discussion	Comparison Rectogordiopsis gen. nov. differs from the other genera of the Neodiscidae by the type of coiling (initially glomospiral and finally planispiral) involute test, the well-developed flosculinisation, the long streptospiral initial stage and devoid of buttresses. It differs from the morpholocically closest genus Kamurana by a larger and more globular test with planispiral involute coiling without any oscillation in the last whorls. It is also similar to Globidiscus but differs by the last whorls not tapering. Hemigordiopsis can be easily differentiated from the new genus Rectogordiopsis by the varying test shape (globular to discoidal), the very dark or even black porcelaneous wall, the absence of a comb teeth-shaped structure on the wall and a very low and broad lumen.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F75322A402D3615FA8DFC8E.taxon	distribution	Range and distribution Middle Permian (late Capitanian) of the Elmadağ Olistostrome, Ankara City, Central Türkiye.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F7A3229439B3018FA8DF98E.taxon	description	urn: lsid: zoobank. org: act: E 13 B 6453 - 85 E 7 - 4 A 7 B-AAF 0 - A 323 C 865 B 17 F Fig. 3	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F7A3229439B3018FA8DF98E.taxon	diagnosis	Diagnosis Large globular Rectogordiopsis gen. nov. with rounded periphery.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F7A3229439B3018FA8DF98E.taxon	etymology	Etymology Referring to its morphology relatively similar to Kamurana.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F7A3229439B3018FA8DF98E.taxon	materials_examined	Material examined Fifty specimens were observed. 1 holotype and 11 paratypes are illustrated. Holotype (Fig. 3 A – B) TÜRKİYE – Ankara • SE of Kutludüğün village, Kutludüğün Yayla, sample KDY – 1; 39 ° 52 ′ 11.0 ″ N, 33 ° 05 ′ 51.4 ″ E; 20 Jun. 2023; Okuyucu and Akbaş leg.; KTÜN. CO – KDY 1, KDY 1.2 – 36. Paratypes (Fig. 3 C – M) TÜRKİYE – Ankara • 6 specs; same data as for holotype; KTÜN. CO – KDY 1, KDY 1.2 – 3, KDY 1.2 – 35, KDY 1.3 – 29, KDY 1.5 – 48, KDY 1.7 – 15, KDY 1.7 – 27 • 5 specs; same data as for holotype, sample KDY – 2; 39 ° 52 ′ 07.4 ″ N, 33 ° 06 ′ 23.4 ″ E; KTÜN. CO – KDY 2, KDY 2.1 – 14, KDY 2.2 – 24, KDY 2.4 – 10, KDY 2.4 – 11, KDY 2.4 – 24.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F7A3229439B3018FA8DF98E.taxon	description	Description Test large to very large, involute, globular with rounded periphery. Coiling initially glomospiral and finally planispiral. Spherical proloculus, 0.087 – 0.100 mm in diameter, followed by second chamber slowly enlarging in height through whorls. Initial part consists of at least 15 – 17 whorls, thin-walled and tightly coiled. In the terminal part (5 – 8 whorls) the height of the chamber increases gradually. Adult specimens measure: D = 2.08 – 3.25, w = 1.88 – 3.00 mm, the ratio w / D = 0.74 – 1.04, n. w. = 20 – 25. Wall calcareous, porcelaneous with perforations (comb teeth-shaped). The lumen of the tube is relatively high with frequent flosculinisation in adult stage. Aperture simple, terminal.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F7A3229439B3018FA8DF98E.taxon	discussion	Comparison Rectogordiopsis kamuranaeformis gen. et sp. nov. differs from K a murana bronnimanni by the planispiral involute coiling without any oscillation. Microfossil association The associated foraminiferal fauna is listed in Table 1 and illustrated in Figs 4 – 6.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F7A3229439B3018FA8DF98E.taxon	distribution	Range and distribution Middle Permian (late Capitanian) of the Elmadağ Olistostrome, Ankara City, Central Türkiye.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F79323240633518FA8DFA93.taxon	description	urn: lsid: zoobank. org: act: F 4124 BCF- 5 B 42 - 495 B-B 5 C 8 - FE 3 DB 60 FAEDF Fig. 7 Indeterminate Miliolina – Kobayashi & Altiner 2011: pl. 1 fig 30.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F79323240633518FA8DFA93.taxon	diagnosis	Diagnosis Very large oval Rectogordiopsis gen. nov. with flattened to slightly rounded periphery.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F79323240633518FA8DFA93.taxon	etymology	Etymology From the overall oval shape of the test.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F79323240633518FA8DFA93.taxon	materials_examined	Material examined Twelve specimens were observed. 1 holotype and 7 paratypes are illustrated. Holotype (Fig. 7 A) TÜRKİYE – Ankara • SE of Kutludüğün village, Kutludüğün Yayla, sample KDY – 2; 39 ° 52 ′ 07.4 ″ N, 33 ° 06 ′ 23.4 ″ E; 20 Jun. 2023; Okuyucu and Akbaş leg.; KTÜN. CO – KDY 2, KDY 2.1 – 6. Paratypes (Fig. 7 B – I) TÜRKİYE – Ankara • 2 specs; same data as for holotype; KTÜN. CO – KDY 2, KDY 2.1 – 52, KDY 2.2 – 1 • 5 specs; same data as for holotype, sample KDY – 1; 39 ° 52 ′ 11.0 ″ N, 33 ° 05 ′ 51.4 ″ E; KTÜN. CO – KDY 1, KDY 1.1 – 1, KDY 1.2 – 19, KDY 1.3 – 19, KDY 1.5 – 24, KDY 1.6 – 28.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F79323240633518FA8DFA93.taxon	description	Description Test large, involute, oval with slightly flattened to broadly rounded periphery. Coiling at the beginning tightly glomospiral and finally planispiral. Spherical proloculus, 0.06 – 0.08 mm in diameter, followed by second chamber slowly enlarging in height through successive whorls. The initial part consists of at least 25 – 30 whorls with thin-walled. In the terminal part (4 – 6 whorls) the height of the chamber increases gradually. Adult specimens measure: D = 1.95 – 3.43 mm, w = 1.70 – 2.68 mm, the ratio w / D = 0.73 – 0.96, n. w. = 29 – 36. Wall thick, calcareous porcelaneous with perforations (comb teeth-shaped). The lumen of the tube is relatively narrow with some flosculinisation in adult stage. Aperture simple, terminal.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F79323240633518FA8DFA93.taxon	discussion	Comparison This new species differs from the Rectogordiopsis kamuranaeformis gen. et sp. nov. mainly by narrower chamber height, flattened periphery and more tightly coiled inner glomospiroid whorls. Microfossil association The associated foraminiferal fauna is listed in Table 1 and illustrated in Figs 4 – 6.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F79323240633518FA8DFA93.taxon	distribution	Range and distribution Middle Permian (late Capitanian) of the Elmadağ Olistostrome, Ankara City, Central Türkiye.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F623233400F3634FA8DFDFD.taxon	description	urn: lsid: zoobank. org: act: 6 D 884 B 23 - E 7 D 2 - 425 E-A 527 - 7 C 53 DAEFC 6 E 1	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F623233400F3634FA8DFDFD.taxon	type_taxon	Type species Davanella ankaraensis gen. et sp. nov.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F623233400F3634FA8DFDFD.taxon	etymology	Etymology After the Dava spring in Kutludüğün Yayla, southeast of Kutludüğün village, Ankara City, Central Türkiye.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F623233400F3634FA8DFDFD.taxon	description	Description Test free, involute, very large and globular to ovoid. Proloculus spherical followed by a tubular undivided chamber. First whorls numerous tightly streptospiral and adult whorls planispiral with depressed umbilicus on both sides. Flosculinisation well developed. Thick porcelaneous wall often neosparitized with perforations (comb teeth-shaped). Aperture simple, terminal.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F623233400F3634FA8DFDFD.taxon	discussion	Comparison Davanella gen. nov. differs from the similar genus Rectogordiopsis gen. nov. by the depressed umbilicus on both sides of the test.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F623233400F3634FA8DFDFD.taxon	distribution	Range and distribution Middle Permian (late Capitanian) of the Elmadağ Olistostrome, Ankara City, Central Türkiye.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F6332314046312DFA8DFEF8.taxon	description	urn: lsid: zoobank. org: act: 32686 C 24 - 35 E 1 - 49 BD- 9162 - 7204 E 5 E 84947 Fig. 8 A – G	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F6332314046312DFA8DFEF8.taxon	diagnosis	Diagnosis Very large globular to ovoid Davanella gen. nov. with rounded periphery.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F6332314046312DFA8DFEF8.taxon	etymology	Etymology Latin, from Ankara, the capital of Türkiye.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F6332314046312DFA8DFEF8.taxon	materials_examined	Material examined Eight specimens were observed. 1 holotype and 5 paratypes are illustrated. Holotype (Fig. 8 E) TÜRKİYE – Ankara • SE of Kutludüğün village, Kutludüğün Yayla, sample KDY – 1; 39 ° 52 ′ 11.0 ″ N, 33 ° 05 ′ 51.4 ″ E; 20 Jun. 2023; Okuyucu and Akbaş leg.; KTÜN. CO – KDY 1, KDY 1.1 – 16. Paratypes (Fig. 8 A – D, G) TÜRKİYE – Ankara • 5 specs; same data as for holotype; KTÜN. CO – KDY 1, KDY 1.2 – 2, KDY 1.3 – 53, KDY 1.8 – 4, KDY 1.8 – 19, KDY 1.8 – 25.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F6332314046312DFA8DFEF8.taxon	description	Description Test large to very large, involute, globular to ovoid with rounded periphery. A central depression or umbilicus well-developed in axial section. Spherical proloculus, measuring 0.07 – 0.09 mm in outside diameter, followed by second tubular chamber. Initial part consists of approximately 12 – 15 loosely coiled glomospiroid whorls. In all subsequent whorls the height of the chamber increases rapidly. Adult specimens measure: D = 2.13 – 3.43 mm, w = 1.90 – 2.38 mm, the ratio w / D = 0.67 – 0.89, n. w. = 15 – 19. Wall thick, calcareous porcelaneous with perforations (comb teeth-shaped). A weak flosculinisation developed in adult stage. Aperture simple, terminal.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F6332314046312DFA8DFEF8.taxon	discussion	Comparison Davanella ankaraensis gen. et sp. nov. differs from the species of the Rectogordiopsis gen. nov. by the presence of umbilicus. Microfossil association The associated foraminiferal fauna is listed in Table 1 and illustrated in Figs 4 – 6.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F6332314046312DFA8DFEF8.taxon	distribution	Range and distribution Middle Permian (late Capitanian) of the Elmadağ Olistostrome, Ankara City, Central Türkiye.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F6132314049322EFA8DF946.taxon	description	urn: lsid: zoobank. org: act: C 385 F 006 - 1 FB 6 - 4017 - B 23 C-F 70139 AC 58 FA Fig. 8 H – L	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F6132314049322EFA8DF946.taxon	diagnosis	Diagnosis Very large globular to ovoid Davanella gen. nov. with pointed periphery.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F6132314049322EFA8DF946.taxon	etymology	Etymology Latin, ‘ acuminatus ’ (= ‘ sharp, pointed’).	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F6132314049322EFA8DF946.taxon	materials_examined	Material examined Five specimens were observed. 1 holotype and 3 paratypes are illustrated. Holotype (Fig. 8 I – J) TÜRKİYE – Ankara • SE of Kutludüğün village, Kutludüğün Yayla, sample KDY – 2; 39 ° 52 ′ 07.4 ″ N, 33 ° 06 ′ 23.4 ″ E; 20 Jun. 2023; Okuyucu and Akbaş leg.; KTÜN. CO – KDY 2, KDY 2.1 – 17. Paratypes (Fig. 8 H, K – L) TÜRKİYE – Ankara • 3 specs; same data as for holotype; KTÜN. CO – KDY 2, KDY 2.1 – 48, KDY 2.1 – 49, KDY 2.3 – 16.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F6132314049322EFA8DF946.taxon	description	Description Test large to very large, involute, globular to ovoid with pointed periphery. Umbilicus slightly-developed on lateral sides. Spherical proloculus, 0.06 – 0.086 mm in diameter, followed by second tubular chamber. The quite tightly coiled glomospiroid initial part consists of approximately 25 whorls. The height of the chamber in planisipiral coiled part increases gradually. Adult specimens measure: D = 2.65 – 3.40 mm, w = 2.03 – 2.63 mm, the ratio w / D = 0.76 – 0.77, n. w. = approximately 32. Wall relatively thick, calcareous, porcelaneous with perforations (comb teeth-shaped). A flosculinisation fills progressively the base of the chamber almost in all whorls. Aperture simple, terminal.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F6132314049322EFA8DF946.taxon	discussion	Comparison Davanella acuminata gen. et sp. nov. differs from the Davanella ankaraensis gen. et sp. nov. by the pointed periphery, quite tightly coiled inner whorls and gradually developed test. Microfossil association The associated foraminiferal fauna is listed in Table 1 and illustrated in Figs 4 – 6.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F6132314049322EFA8DF946.taxon	distribution	Range and distribution Middle Permian (late Capitanian) of the Elmadağ Olistostrome, Ankara City, Central Türkiye.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F613236400F3440FA8DFC4D.taxon	description	urn: lsid: zoobank. org: act: BC 97972 C- 632 D- 4124 - 9 D 50 - 76 CC 24 B 5 AE 04	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F613236400F3440FA8DFC4D.taxon	type_taxon	Type species Kaganella tekini gen. et sp. nov.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F613236400F3440FA8DFC4D.taxon	etymology	Etymology This genus is named after Professor Dr U. Kagan Tekin (Hacettepe University, Ankara, Türkiye), in honour of his contributions to the knowledge of Radiolarian biostratigraphy.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F613236400F3440FA8DFC4D.taxon	description	Description Test involute moderate to large and globular. Proloculus spherical followed by a tubular undivided chamber. First whorls numerous, tightly glomosproid and following whorls planispiral. Flosculinisation poorly developed. Relatively thick porcelaneous wall neosparitized with rare perforations (comb teeth-shaped). Aperture simple terminal.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F613236400F3440FA8DFC4D.taxon	discussion	Comparison Kaganella gen. nov. differs from the other genera of the subfamily Globidiscinae subfam. nov. by the weakly-developed flosculinisation, rare perforations (comb teeth-shaped) in neosparitized wall and smaller dimensions in similar number of whorls.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F613236400F3440FA8DFC4D.taxon	distribution	Range and distribution Middle Permian (late Capitanian) of the Elmadağ Olistostrome, Ankara City, Central Türkiye.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F663237402F375DFA8DFCEF.taxon	description	urn: lsid: zoobank. org: act: ABA 3 BE 39 - 4 E 76 - 47 C 5 - 9 AA 9 - A 6 D 380 B 32 B 51 Fig. 8 M – R	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F663237402F375DFA8DFCEF.taxon	diagnosis	Diagnosis Moderate to large rounded Kaganella gen. nov. with slightly pointed periphery.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F663237402F375DFA8DFCEF.taxon	etymology	Etymology This species is named after Professor Dr U. Kagan Tekin (Hacettepe University, Ankara, Türkiye), in honour of his contributions to the knowledge of Radiolarian biostratigraphy.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F663237402F375DFA8DFCEF.taxon	materials_examined	Material examined Five specimens were observed. 1 holotype and 4 paratypes are illustrated. Holotype (Fig. 8 M – N) TÜRKİYE – Ankara • SE of Kutludüğün village, Kutludüğün Yayla, sample KDY – 2; 39 ° 52 ′ 07.4 ″ N, 33 ° 06 ′ 23.4 ″ E; 20 Jun. 2023; Okuyucu and Akbaş leg.; KTÜN. CO – KDY 2, KDY 2.2 – 2. Paratypes (Fig. 8 O – R) TÜRKİYE – Ankara • 1 spec.; same data as for holotype; KTÜN. CO – KDY 2, KDY 2.2 – 35 • 3 specs; same data as for holotype, sample KDY – 1; 39 ° 52 ′ 11.0 ″ N, 33 ° 05 ′ 51.4 ″ E; KTÜN. CO – KDY 1, KDY 1.2 – 27, KDY 1.2 – 40, KDY 1.3 – 35.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F663237402F375DFA8DFCEF.taxon	description	Description Test moderate to large, involute, globular with slightly pointed periphery. Coiling at the beginning tightly glomospiral and finally planispiral. Spherical proloculus, 0.06 – 0.10 mm in diameter, followed by second chamber slowly enlarging in height through successive whorls. Initial part consists of approximately 23 – 25 whorls with relatively thin-walled. In the terminal part (planispiral coiled 3 – 4 whorls) the height of the whorls increases gradually. Adult specimens measure: D = 1.23 – 2.38 mm, w = 0.9 – 1.25 mm, the ratio w / D = 0.64 – 0.87, n. w. = 26 – 29. Wall calcareous, porcelaneous with some neosparitizations mainly in last whorls. The lumen of the tube is relatively narrow with rare flosculinisations. Aperture simple, terminal.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F663237402F375DFA8DFCEF.taxon	discussion	Comparison Kaganella tekini gen. et sp. nov. differs from Rectogordiopsis kamuranaeformis gen. et sp. nov. by the smaller size of the test and rare perforations (comb teeth-shaped) in neosparitized wall which observed mainly in last whorls. Microfossil association The associated foraminiferal fauna is listed in Table 1 and illustrated in Figs 4 – 6.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
03969E197F663237402F375DFA8DFCEF.taxon	distribution	Range and distribution Middle Permian (late Capitanian) of the Elmadağ Olistostrome, Ankara City, Central Türkiye.	en	Okuyucu, Cengiz, Akbaş, Melikan (2026): Systematic revision and phylogenetic assessment of the family Neodiscidae (Foraminifera) and new Capitanian foraminiferal genera from the Ankara region, central Türkiye. European Journal of Taxonomy 1043: 33-61, DOI: 10.5852/ejt.2026.1043.3207, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3207/14233
