identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0388D1230B709B700D8AFB7FFE4DFA67.text	0388D1230B709B700D8AFB7FFE4DFA67.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neatus turcicus Soldati & Leo & Jansson & Avci & Coşkun & Kayiş 2025	<div><p>Neatus turcicus sp. nov. F. Soldati &amp; P. Leo</p><p>(Figs 5, 11, 14, 16–17, 24, 27–28)</p><p>Body length from clypeus top to elytral apex: 14.4–15.8 mm; short and convex; entirely black, with little shine, femora and tibiae brown to reddish-brown.</p><p>Head. Strongly and densely punctate, punctation less dense on clypeus, the latter slightly indented; genae subparallel in front of eyes, hiding insertion of antennae (Fig. 16); temples short and strongly narrowed behind eyes; antennae stout, gradually enlarged apicad; 1 st antennomere globose, a little longer than wide; 2 nd small, not longer than wide, almost quadratic; 3 rd by far the most elongated, about 1.5 times longer than wide; 4 th –6 th scarcely longer than wide, 7 th and 8 th scarcely transverse, 9 th and 10 th very transverse, 11 th and largest antennomere, sub orbicular, not or scarcely longer than wide; mentum sub cordiform, with strong but spaced punctation.</p><p>Pronotum. Very transverse, with evenly rounded sides, fairly thick lateral margin, prominent and acute anterior angles, anterior margin only slightly indented, almost straight in the middle; anterior margin not rimmed along most of its length, edge visible only at anterior angles; basal margin thin and complete; base of pronotum bisinuate in front of 3 rd –5 th elytral interstriae; punctation of pronotum strong, tight and double-sized, with very large punctures mixed with distinctly smaller ones, present mainly on sides but also numerous on disc (Fig. 11); prohypomera fossulate punctate, punctation tending to merge and form longitudinal wrinkles; anterior and posterior edges of prothorax provided with a dense fringe of long yellow bristles.</p><p>Elytra. Subparallel on their anterior half, then slightly widened posteriorly, humeral angles marked but obtuse, lateral margin visible from above along its entire length, clearly widened posteriorly where it forms a gutter; elytral striae well marked, consisting of series of large, tightly packed punctures at least 5 times larger than those of interstriae, striae complete and clearly visible up to the apex with 3 rd and 6 th, as well as 4 th and 5 th striae connected to each other well before apex; interstriae not very convex, but sufficiently so to reveal the furrowed striae, with very fine, tight punctation, always clearly visible, on a micro reticulate background; basal striole, near scutellum, not very visible and partially erased, consisting of 5–10 large punctures similar to those of striae; epipleura strongly punctate and broad to apex; scutellum quite large, triangular, with strong, dense punctation.</p><p>Tibiae. Male protibiae barely sinuate at inner margin with small granules on distal half (Fig. 14); female protibiae shorter, hardly sinuate at inner margin but smooth; male mesotibiae slightly arcuate and with granules at inner margin; similar in female but smooth at inner margin; male metatibiae with granules on distal inner half, smooth in female.</p><p>Ventrites. First three ventrites densely punctate striolate, 4 th and 5 th ventrites simply punctate; last ventrite simply rounded and entirely rimmed in both sexes.</p><p>Aedeagus. Length of 4.2 to 4.3 mm, parameres measuring slightly less than 1/3 of the total length of the organ (Fig. 24), evenly narrowed to apex which is barely spatulate (Fig. 28) and slightly sinuate at the apex, seen in profile (Fig. 27); Rp of 3.37 to 3.96.</p><p>Sexual dimorphism. Hardly perceptible; all male tibiae granulate in distal half of inner margin, smooth in females; male protibiae barely longer and slightly more sinuate at inner margin.</p><p>The nearest Neatus species morphologically and geographically to N. turcicus sp. nov. are N. subaequalis and N. noctivagus . The new species is distinct from N. subaequalis by its proportionally larger aedeagus, with differently shaped parameres, longer and evenly narrowed towards the apex, which is much more acuminate (Figs 24, 28 vs. Figs 21–22) and less sinuate in profile. The male protibiae are also clearly less sinuate on the inner margin. Moreover, the genae are widened in front of the eyes in N. subaequalis, not subparallel at the sides. Neatus turcicus sp. nov. can be distinguished from N. noctivagus by the subparallel genae in front of eyes, strongly and angularly widened in noctivagus (Fig. 16 vs. Fig. 15), short male protibiae with barely sinuate inner margin (Fig. 14 vs. Fig. 13), the aedeagus smaller in length (L= 4.2–4.3 mm vs. 4.5–6.0 mm) and proportion (Rp=3.37–3.96 vs. 2.75–3.33), with parameres narrowed almost in a straight line and only slightly sinuate in profile (Figs 24, 27–28 vs. Figs 18–19, 26). These three species inhabit different areas of Türkiye but may cohabit locally.</p><p>Type material. HOLOTYPE ♂ (Fig. 5) labelled: “ Turkey, Eğirdir, Gelendost, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.970778&amp;materialsCitation.latitude=38.119583" title="Search Plazi for locations around (long 30.970778/lat 38.119583)">AfŞar village</a>, Hollow Q. ithaburensis, 38°07’10.5’’N; 30°58’14.8’’E, P-trap: 18 2015-4/7, M. Avci / N Jansson ” (white label); “Holotype ♂, Neatus turcicus sp. nov., F. Soldati &amp; P. Leo, des. 2025” (red label), Entomological Museum, Isparta University of Applied Sciences, Faculty of Forestry, Isparta, Türkiye (IEM).</p><p>PARATYPES: 5 ♂♂ and 4 ♀♀; 1 ♂, “ Turkey, Kizilen, 50 km N Erdemli, old oak, 3P, 2005-07-07, N. Jansson / M. Coskun ” (white label) ; 1 ♂ and 1 ♀, “ Turkey, 30 km N Erdemli, ihåliga ekar [hollow oaks], 2001-06-01, N. Jansson ” (white label) ; 1 ♀, “ Turkey, Mersin, 40 km N Gülnar, N 36°30’22,5”; E 33°07’43,3”, Köseçobanli / TaŞdüŞtü,, P-trap 16, Hollow Quercus, 2006-08-26, Nicklas Jansson / Mustafa Avci” (white label) ; 2 ♂♂ and 2 ♀♀, “ Turkey, environs of Konya, 9-12.VII.1999, Werner leg. (white label)”; 1 ♂, “ Turkey, environs of Osmaniye ( Adana), VII.2000, Werner &amp; Lizler leg. (white label)”. All paratypes bear a red label: “ PARATYPE ♂ (or ♀), Neatus turcicus sp. nov., F. Soldati &amp; P. Leo des. 2025”. Paratypes are deposited in the IEM, FS and PL collections.</p><p>Etymology. The name turcicus is an adjective derived from Türkiye, corresponding to Turkish.</p><p>Ecology. Several adult specimens of Neatus turcicus sp. nov. were collected in pitfall traps placed in hollow trunks of old oaks (e.g. Quercus ithaburensis Decne.). This species probably inhabits the decayed wood or the wood mould in hollow oak cavities.</p></div>	https://treatment.plazi.org/id/0388D1230B709B700D8AFB7FFE4DFA67	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Soldati, Fabien;Leo, Piero;Jansson, Nicklas;Avci, Mustafa;Coşkun, Mustafa;Kayiş, Tamer	Soldati, Fabien, Leo, Piero, Jansson, Nicklas, Avci, Mustafa, Coşkun, Mustafa, Kayiş, Tamer (2025): A new species of the genus Neatus J. L. Leconte, 1862 (Coleoptera: Tenebrionidae: Tenebrionini) from Türkiye, with new country records and a key to holarctic species. Zootaxa 5659 (2): 179-190, DOI: 10.11646/zootaxa.5659.2.2, URL: https://doi.org/10.11646/zootaxa.5659.2.2
0388D1230B749B710D8AF9BBFB3EFE0F.text	0388D1230B749B710D8AF9BBFB3EFE0F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neatus noctivagus (Mulsant & Rey 1853)	<div><p>Neatus noctivagus (Mulsant &amp; Rey, 1853)</p><p>(Fig. 1)</p><p>Type locality: “la Sicile ”, described from Sicily (Mulsant &amp; Rey 1853).</p><p>Studied material. Greece.— Etolía-Akarnanía, Lessini, 1 to 3-VII-1993, A. Liberto, 3 ♂♂ (FS, PL). Italy.— Basilicata, Matera, Policoro, 24-IV-1986, F.Angelini, 1 ♀ (FS). Idem, 24-IV-1988, F.Angelini, 3 ♂♂ (FS, PL). Idem, 9-IV-1995, F. Izzillo, 1 ♂, 4 ♀♀ (PL). Idem, 27-VIII-1995, F. Angelini, 2 ♀♀ (FS). Basilicata, Matera, Policoro, Bosco Pantano, IV-1986, A. Liberto, 2 ♂♂ (FS, PL). Calabria, Cosenza, Santo Stefano di Rogliano, 7-IV-1978, M. Zampetti, 1 ♀ (CM). Lazio, Roma, Capocotta, 1959, E. Migliaccio, 1 ♂ (PL). Lazio, Roma, Marcigliana, 31-XII-2005, A. B. Biscaccianti, 1 ♀ (DS) . Lazio, Roma, Pomezia, VIII-1986, E Pacieri, 1 ♀ (PL). Idem, 28-VIII-2012, F. Turchetti, 4 ♂♂ (FS, PL). Lazio, Roma, Villa Borghese, VII-1981, M. Mei, 1 ♂ (PL). Sicily, Messina, Montalbano Elicona, Bosco Malabotta, 1100 m , 3-VI-1993, F. Angelini, 1 ♂ (FS) . Sicily, Messina, Portella di Femmina Morta, 1300 m , 3-V-1972, Korell, 1 ♀ (PL). Sicily, Messina, Mistretta, 26-V-1974, B. Massa, 1 ♂ (PL) . Sicily, Palermo, Madonie, Petralia Sottana, Piano Battaglia, 9-V-1996, F. Angelini, 1 ♀ (PL) . Türkiye.— Balıkesir, Ida Mountain, Ayi stream, 10 km N Edremit, 15-VI-2011, T. Öncül, S. Varlı &amp; N. Jansson, 1 ♂ (NJ). Idem, 20-VII-2011, T. Öncül, S. Varlı &amp; N. Jansson, 2 ♀♀ (IEM). Greece and Türkiye are new country records .</p><p>Geographical distribution: From Latium (Italy) to NW Türkiye; Albany, Croatia and Italy (Iwan et al. 2020); in Italy, in the southern half of the country and in Sicily where it only occurs in few localities of the Madonie and Nebrodi mountains (Aliquó et al. 2007). Occurring in Albany, it is not so surprising to discover this species in Greece (Etolía) and in the extreme NW of Türkiye.</p><p>The aedeagus of this species is quite variable, as shown by Schawaller &amp; Grimm (1995: 72), specimens coming from Albany, Greece and Türkiye having apical tip of parameres larger (Fig. 19 vs Fig. 18).</p></div>	https://treatment.plazi.org/id/0388D1230B749B710D8AF9BBFB3EFE0F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Soldati, Fabien;Leo, Piero;Jansson, Nicklas;Avci, Mustafa;Coşkun, Mustafa;Kayiş, Tamer	Soldati, Fabien, Leo, Piero, Jansson, Nicklas, Avci, Mustafa, Coşkun, Mustafa, Kayiş, Tamer (2025): A new species of the genus Neatus J. L. Leconte, 1862 (Coleoptera: Tenebrionidae: Tenebrionini) from Türkiye, with new country records and a key to holarctic species. Zootaxa 5659 (2): 179-190, DOI: 10.11646/zootaxa.5659.2.2, URL: https://doi.org/10.11646/zootaxa.5659.2.2
0388D1230B759B710D8AFDCFFD5AF9B7.text	0388D1230B759B710D8AFDCFFD5AF9B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neatus picipes (Herbst 1797)	<div><p>Neatus picipes (Herbst, 1797)</p><p>(Fig. 2)</p><p>Type locality: “Deutschland”, described from Germany (Herbst 1797).</p><p>Studied material. Bulgaria.— Isskar valley, N Passarell vill., 710 m, 14-VII to 9-VIII-2005, E. Migliaccio, 1 ♂ (EM). Srandzha Mts., E Kondolovo vill., 300 m, 28-VIII to 11-XI-2005, E. Migliaccio, 1 ♀ (EM). Czech Republic.— Moravia, Břeclav, 31-III-2002, P. Čekovski, 1 ♂, 1 ♀ (PL) . Idem, 15-VI-2005, R. Fornosek, 1 ♂ (FS) . Idem, 30-VI-1997, K. Hlavenka, 1 ♂ (FS) . Greece.— Kilkis, Kavaláris, 01-VII-2001, C. Cocquempot, 1 ♂ (FS) . Thessalia, Larissa, Stomio, 90 m, 2-VIII-2006, D. Sechi, 1 ♂ (PL) . France.— Allier, Châtel-de-Neuvre, 28-III-2012, 5-IV-2012, 31-V-2012, L. Velle, 5 ♂♂, 4 ♀♀ (FS, LV) . Toulon-sur-Allier, 23-IX-2014, 24-VII-2015, L. Velle, 1 ♂, 1 ♀ (LV) . Alpes-de-Haute-Provence, Niozelles, XI-1997, A. Coache, 1 ♀ (AC) . Bouches-du-Rhône, Arles, la Tour-du-Valat, 12-VI-1983, J. F. Voisin (MNHN) . Bouches-du-Rhône, Saint-Paul-lès-Durance, Cadarache forest, 5-VII-2011, T. Barnouin, T. Noblecourt &amp; F. Soldati, 1 ♀ (LNEF-ONF) . Lot-et-Garonne, Houeillès, 19-IV-2016, 8-VIII-2016, C. Van Meer, 1 ♂, 1 ♀ (CVM) . Var, Les Mayons, 15-VIII-1997, H. Brustel, 4 ♂♂, 2 ♀♀ (HB, FS) . Italy.— Emilia Romagna, near Modena, 15-IV-1978, Malferrari, 1 ♂ (LS) . Emilia Romagna, Ravenna, Bagnacavallo, 7-XI-1968, E. Contarini, 1 ♂ (PL) . Emilia Romagna, Ravenna, Pineta S. Vitale, 30-III-1985, L. Saltini, 1 ♀ (PL) . Idem, 29-VI-2002, N. Cuffiani, 1 ♂ (PL) . Emilia Romagna, Reggio Emilia, Villa Gaida, 3-XII-2011, M. Violi, 2 ♂♂, 1 ♀ (PL) . Poland.— Rogalin ad Poznań, 22-VI-1997, P. Jaloszińsky, 2 ♀♀ (PL) . Romania.— Dolj, Craiova, 1-V-1994, 26-VII-2005, F. Soldati, 1 ♂, 1 ♀ (FS) . Idem, VII-1995, F. Soldati, 1 ♂ (FS) . Giurgiu, Comana VlaŞca, without date, A. P. Montandon, 2 ♂♂, 2 ♀♀ (MNHN) . Slovakia.— Bratislava, surroundings of Malacky, 3-VII-1997, K. Hlavenka, 1 ♂ (FS) . Switzerland.— Genève, 13-I-1978, P. Verkaik, 1 ♀ (FS) .</p><p>Geographical distribution: Middle Europe and western Asia, from SW France (Van Meer &amp; Bouyon 2019) to Russia. Europe: Austria, Belarus, Bulgaria, Croatia, Czech Republic, Denmark, France, Great Britain, Germany, Greece, Hungary, Italy, Latvia, Poland, Romania, Russia, Slovakia, Switzerland and Ukraine; Asia: Kazakhstan, NE China, South Korea, western Siberia and Nearctic Region (Iwan et al. 2020). Records from Denmark and Great Britain are erroneous, the species being not reported by Hansen (1996) and Barclay (2012) respectively. The ones of NE China, South Korea and Nearctic Region are due to confusions with Neatus ventralis and N. tenebrioides . In Italy, N. picipes only occurs in the NE of the country (Aliquó et al. 2007) and in Poland, the species is not rare but mainly localized along large rivers (Iwan et al. 2012).</p></div>	https://treatment.plazi.org/id/0388D1230B759B710D8AFDCFFD5AF9B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Soldati, Fabien;Leo, Piero;Jansson, Nicklas;Avci, Mustafa;Coşkun, Mustafa;Kayiş, Tamer	Soldati, Fabien, Leo, Piero, Jansson, Nicklas, Avci, Mustafa, Coşkun, Mustafa, Kayiş, Tamer (2025): A new species of the genus Neatus J. L. Leconte, 1862 (Coleoptera: Tenebrionidae: Tenebrionini) from Türkiye, with new country records and a key to holarctic species. Zootaxa 5659 (2): 179-190, DOI: 10.11646/zootaxa.5659.2.2, URL: https://doi.org/10.11646/zootaxa.5659.2.2
0388D1230B759B7E0D8AF917FB20FE9F.text	0388D1230B759B7E0D8AF917FB20FE9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neatus subaequalis Reitter 1920	<div><p>Neatus subaequalis Reitter, 1920</p><p>(Fig. 3)</p><p>Type locality: “ Albanien (Skutari), Kaukasus, Daghestan, Lenkoran”, described from Albany and Transcaucasia (Reitter 1920).</p><p>Studied material. Armenia.— Aragatsotn, Ashtarak, Antarut, 29-V-1983, M. Kalashian, 1 ♂, 1 ♀ (FS) . Greece.— Evritania, Timfristos, VI-1994, P. Verkaik, 1 ♀ (FS) . Evia, between Steni and Stroponas, 1000 m, 4 to 20- V-1991, A. Liberto, 1 ♀ (PL) . Thessalia, Agia, 2, 5 km N Omolio, 1 to 2-VIII-2006, D. Sechi, 1 ♂, 1 ♀ (DS) . Idem, 14 to 16-VIII-2008, 2 ♂♂ (DS) . Thessalia, Larissa, Ossa Oros, 518–900 m, 23 to 31-VII-2006, D. Sechi, 5 ♂♂, 2 ♀♀ (DS, FS, PL) . Idem, 450 m, 25-VI-2005, S. Peslier, 1 ♂ (FS) . Thessalia, Trikala, Metéora, Vlahava, 19-VI-2009, L. Valladarès, 1 ♂ (FS) . Thessalia, Agia, 2, 5 km N Omolio, 1 to 2-VIII-2006, D. Sechi, 1 ♂, 1 ♀ (DS) . Idem, 14 to 16-VIII-2008, 2 ♂♂ (DS).</p><p>Geographical distribution: From southern Balkans to Transcaucasia and Iran; Azerbaijan, Albany, Armenia, Georgia, Greece, Iran, Macedonia, southern Russia and Türkiye (Iwan et al. 2020).</p><p>The aedeagus of this species is quite variable, as shown by Schawaller &amp; Grimm (1995: 72), specimens coming from Greece having longer aedeagus with parameres apically broader (Fig. 22 vs Fig. 21).</p></div>	https://treatment.plazi.org/id/0388D1230B759B7E0D8AF917FB20FE9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Soldati, Fabien;Leo, Piero;Jansson, Nicklas;Avci, Mustafa;Coşkun, Mustafa;Kayiş, Tamer	Soldati, Fabien, Leo, Piero, Jansson, Nicklas, Avci, Mustafa, Coşkun, Mustafa, Kayiş, Tamer (2025): A new species of the genus Neatus J. L. Leconte, 1862 (Coleoptera: Tenebrionidae: Tenebrionini) from Türkiye, with new country records and a key to holarctic species. Zootaxa 5659 (2): 179-190, DOI: 10.11646/zootaxa.5659.2.2, URL: https://doi.org/10.11646/zootaxa.5659.2.2
0388D1230B7A9B7E0D8AFE3BFDBBFCC3.text	0388D1230B7A9B7E0D8AFE3BFDBBFCC3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neatus tenebrioides (Palisot de Beauvois 1812)	<div><p>Neatus tenebrioides (Palisot de Beauvois, 1812)</p><p>(Fig. 4)</p><p>Type locality: “ Caroline du Sud ”, described from the South Carolina State in the USA (Palisot de Beauvois 1812).</p><p>Studied material. Canada.— Québec, Eardley K. 13, 19-V-1996, V. Maly &amp; S. Laplante, 1 ♂, 3 ♀♀ (FS) . USA.— Illinois, Chicago, Palos Park, 13-V-1950, J. B. Hayes, 6 ♂♂, 3 ♀♀ (FS, PL) . Iowa, Cedar Bluff, 05-VI-1958, J. R. Powers, 1 ♂, coll. P. Ardoin (MNHN) . Michigan, Douglas Lake biological station, 09-VII-1962, 1 ♂, coll. P. Ardoin (MNHN) . Michigan, Macomb County, E of Memphis, 07-V-1963, IV-1966, 21-III-1968, C. Brivio, 1 ♂, 2 ♀♀ coll. P. Ardoin (MNHN) .</p><p>Geographical distribution: Widely distributed in North America, cited from 39 states of the USA and 6 provinces of Canada (Bousquet et al. 2018).</p></div>	https://treatment.plazi.org/id/0388D1230B7A9B7E0D8AFE3BFDBBFCC3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Soldati, Fabien;Leo, Piero;Jansson, Nicklas;Avci, Mustafa;Coşkun, Mustafa;Kayiş, Tamer	Soldati, Fabien, Leo, Piero, Jansson, Nicklas, Avci, Mustafa, Coşkun, Mustafa, Kayiş, Tamer (2025): A new species of the genus Neatus J. L. Leconte, 1862 (Coleoptera: Tenebrionidae: Tenebrionini) from Türkiye, with new country records and a key to holarctic species. Zootaxa 5659 (2): 179-190, DOI: 10.11646/zootaxa.5659.2.2, URL: https://doi.org/10.11646/zootaxa.5659.2.2
0388D1230B7A9B7E0D8AFC07FF40FACB.text	0388D1230B7A9B7E0D8AFC07FF40FACB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neatus ventralis (Marseul 1876)	<div><p>Neatus ventralis (Marseul, 1876)</p><p>(Fig. 6)</p><p>Type locality: “Japon”, described from Japan (Marseul 1876).</p><p>Studied material. Japan.— Honshū, Aomori pref., Hirosaki, 1897, R. P. Faurie, 1 ♂, 1 ♀ (MNHN) . Honshū, Miyagi pref., Amazuka, 1895, 1 ♂, 1 ♀ (MNHN) . Honshū, Tochigi pref., Ichikaimachi, 5-IX-1982, T. Niisato, 3 ♀♀ (PL) . Honshū, Tochigi pref., Ichikai, Ishioroshi, 5-IX-1982, T. Niisato, 1 ♂, 1 ♀ (FS) . Kyushu, Nagasaki pref., Nagasaki, 1898, 1 ♂, 3 ♀♀ (MNHN) .</p><p>Geographical distribution: Far East Asia; China, North Korea, Eastern Siberia, Russian Far East and Japan (Iwan et al. 2020). In spite to be distinct by its body shape, sculpture of the pronotum disc, aciculate elytral punctation and even more its aedeagus, this species is considered erroneously as a synonym of N. picipes by Akita &amp; Masumoto (2017).</p></div>	https://treatment.plazi.org/id/0388D1230B7A9B7E0D8AFC07FF40FACB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Soldati, Fabien;Leo, Piero;Jansson, Nicklas;Avci, Mustafa;Coşkun, Mustafa;Kayiş, Tamer	Soldati, Fabien, Leo, Piero, Jansson, Nicklas, Avci, Mustafa, Coşkun, Mustafa, Kayiş, Tamer (2025): A new species of the genus Neatus J. L. Leconte, 1862 (Coleoptera: Tenebrionidae: Tenebrionini) from Türkiye, with new country records and a key to holarctic species. Zootaxa 5659 (2): 179-190, DOI: 10.11646/zootaxa.5659.2.2, URL: https://doi.org/10.11646/zootaxa.5659.2.2
0388D1230B7A9B7F0D8AFA0FFA35FE62.text	0388D1230B7A9B7F0D8AFA0FFA35FE62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neatus J. L. Leconte 1862	<div><p>Key to Neatus species (males)</p><p>1(2) Smaller (11.5–13.5 mm); disc of pronotum finely punctate, larger punctures only on sides (Fig. 10); cuticle shiny; small aedeagus (2.5–3.1 mm), with parameres very short and truncated at apex (Fig. 23); Rp from 4.03 to 4.60; North American species..................................................................................... tenebrioides</p><p>2(1) Larger (13.5–18 mm); disc of pronotum more strongly and tightly punctate, larger punctures more or less numerous and present, very rarely absent (Figs 7–9, 11–12); Palearctic species</p><p>3(6) Aedeagus proportionally small (Rp from 3.67 to 4.83), either very small and short (L=3.0– 3.2 mm) with acuminate apex (Fig. 25), or longer (L= 3.5–4.5 mm) but with parameres strongly truncated at apex (Figs 21–22)</p><p>4(5) Cuticle shiny, punctation of pronotum disc fine, larger punctures few and far between, clearly scattered (Fig. 12); punctation of elytral interstriae fine and aciculate, barely perceptible; aedeagus very small (Rp from 4.70 to 4.83) and very short (L=3.0– 3.2 mm), with parameres acuminate at apex (Fig. 25); Far East Asia .......................................... ventralis</p><p>5(4) Cuticle not very shiny, punctation of pronotum disc strong and thick, larger punctures numerous and less scattered (Fig. 9); punctation of elytral interstriae quite strong and thick, clearly visible; aedeagus distinctly longer (L= 3.5–4.5 mm) and proportionally larger (Rp from 3.67 to 4.38), with parameres strongly truncated at apex (Figs 21–22); from southern Balkans to the Caucasus and Iran .......................................................................... subaequalis</p><p>6(3) Aedeagus differently shaped, proportionally larger (Rp 2.75 to 3.96), parameres with more or less tapered apical tips (Figs 18–20, 24)</p><p>7(8) Punctation of pronotum disc finer, larger punctures few and far between or absent (Fig. 8); rather slender and subparallel, elytral disc not very convex; aedeagus with sinuate parameres on sides and apex tapering (Fig. 20), 3.8 to 4.4 mm long, Rp 3.30 to 3.81; middle Europe.............................................................................. picipes</p><p>8(7) Punctation of pronotum disc strong, larger punctures more or less invading the disc (Figs 7, 11); more convex, elytral disc even more convex posteriorly; aedeagus with apex less tapered (Figs 18–19, 24, 28); southern Europe, from Sicily and Latium in Italy, to Türkiye</p><p>9(10) Genae subparallel in front of eyes (Fig. 16); male protibiae short, their inner margin only barely sinuate (Fig. 14); aedeagus shorter (L= 4.2–4.3 mm), proportionally smaller (Rp from 3.37 to 3.96), with parameres narrowed almost in a straight line to apex which is tapered (Figs 24, 28), barely sinuate at apex, in profile (Fig. 27); southern Türkiye .......... turcicus sp. nov.</p><p>10(9) Genae more or less expanded in front of eyes (Fig. 15); male protibiae slender, with distinctly sinuate inner margin (Fig. 13); aedeagus distinctly longer (L=4.5–6.0 mm), proportionally larger (Rp from 2.75 to 3.33), with sinuate parameres on sides (Figs 18–19), clearly sinuate at apex, in profile (Fig. 26); from Latium and Sicily in Italy, to northwest Türkiye ....... noctivagus</p></div>	https://treatment.plazi.org/id/0388D1230B7A9B7F0D8AFA0FFA35FE62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Soldati, Fabien;Leo, Piero;Jansson, Nicklas;Avci, Mustafa;Coşkun, Mustafa;Kayiş, Tamer	Soldati, Fabien, Leo, Piero, Jansson, Nicklas, Avci, Mustafa, Coşkun, Mustafa, Kayiş, Tamer (2025): A new species of the genus Neatus J. L. Leconte, 1862 (Coleoptera: Tenebrionidae: Tenebrionini) from Türkiye, with new country records and a key to holarctic species. Zootaxa 5659 (2): 179-190, DOI: 10.11646/zootaxa.5659.2.2, URL: https://doi.org/10.11646/zootaxa.5659.2.2
