taxonID	type	description	language	source
038D87BEFFAEFFF8FDF5FD21FCA5FCB1.taxon	description	= Phalangochaeta huebneri – Chalumeau & Howden 1984: 87 = Odontolytes huebneri – Krell et al. 2011: 175	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB6FFE1FD84FCF2FA82F863.taxon	diagnosis	Diagnosis Length 3.0 – 5.0 mm. Body sub-cylindrical, testaceous to dark testaceous. Head, disc of pronotum and elytra with a dark spot variably developed; clypeus sinuate or emarginate; gena rounded with distinct pale and long setae. Apical spines of meso- and metatibiae set close, not allowing the first tarsomere to articulate between them. Sexual dimorphism evidenced by the transverse head and apical spine of protarsus in males, these can be lanceolate or laterally flat and downwards bent hook like or truncate. The abdomen of males of some species with a median fovea.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB6FFE1FD84FCF2FA82F863.taxon	discussion	Remarks	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB7FFE0FD97FE95FA83FBA5.taxon	description	Fig. 33 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB7FFE0FD97FE95FA83FBA5.taxon	diagnosis	Diagnosis Length 3.2 mm. Head emarginate medially; clypeus finely punctate, punctation subrugose anteriorly. Pronotum with dense and uneven punctation. Elytral interstriae eight and ten distinctively elevated posteriorly and with fine and scarce punctures. Sexual dimorphism Unknown, it may follow the pattern observed with the other species in the genus, where the male, normally, has a clypeus more transverse and discretely sinuate. Collecting information No information available in the literature.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB7FFE0FD97FE95FA83FBA5.taxon	discussion	Remarks No specimens were available for the present study, thus the diagnosis provided and positioning of the species in the dichotomous key was made based solely on the available literature (Dellacasa et al. 2001 b).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB7FFE6FE3AFBA0FC2FFD68.taxon	description	Figs 1 E – G, 33 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB7FFE6FE3AFBA0FC2FFD68.taxon	diagnosis	Diagnosis Length 3.0 – 4.0 mm. Head with clypeus slightly emarginate and with fine and scarce punctures; gena obtusely rounded; frons with large punctures. Pronotum with punctures of two sizes; punctures symmetrically distributed on disc, but closer on sides; posterior angles of pronotum obtusely rounded. Elytra shagreened; interstriae discretely convex with finely punctate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB7FFE6FE3AFBA0FC2FFD68.taxon	materials_examined	Type material Paratypes BRAZIL • 1 ♀; “ [white paper, printed] BR-MS-Selvíria / UNESP Farm / Black light trap / Brachiaria decumbens pasture / C. A. H. Flechtmann. col / [handwritten to the side] 28 / XI / 1998. // [white paper, handwritten] Alex 08. // [red paper, printed] Aidophus / flechtmanni n. sp. / ALLOTYPE / DELLACASA M & ALII DES. 2001 ”; CEMT • 1 ♀; “ [white paper, printed] BR-MS- Selvíria / UNESP Farm / Black light trap / Brachiaria decumbens pasture / C. A. H. Flechtmann. col / [handwritten to the side] 14 / III / 1996. // [red paper, printed] Aidophus / flechtmanni n. sp. / PARATYPUS / DELLACASA M & ALII DES. 2001 ”; CEMT • 1 ♀; “ [white paper, printed] BR-MS-Selvíria / UNESP Farm / Black light trap / Brachiaria decumbens pasture / C. A. H. Flechtmann. col / [handwritten to the side] 14 / III / 1996. // [white paper, printed RS 24. // [red paper, printed] Aidophus / flechtmanni n. sp. / PARATYPUS / DELLACASA M & ALII DES. 2001 ”; CEMT • 2 ♀♀; “ [white paper, printed] BR-MS-Selvíria / UNESP Farm / Black light trap / Brachiaria decumbens pasture / C. A. H. Flechtmann. col / [handwritten to the side] 28 / XI / 1998. // [red paper, printed] Aidophus / flechtmanni n. sp. / PARATYPUS / DELLACASA M & ALII DES. 2001 ”; CEMT. Other material examined BRAZIL – Mato Grosso do Sul • 4 ♂♂, 2 ♀♀; Selvíria, UNESP Farm; 28 Nov. 1993; C. A. H. Flechtmann leg.; Guzera bovine dropping; Brachiaria decumbens pasture; CEMT • 2 ♂♂; same data as for preceding 31 Oct. 1993; Bubalus bubalis dropping; CEMT • 1 ♂; Inocência, Fazenda Moinho; 19 ° 13 ′ 12 ″ S, 52 ° 9 ′ 34 ″ W; elev. 502 m; 21 Oct. 2018; H. L. Rainho leg; light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB7FFE6FE3AFBA0FC2FFD68.taxon	description	Sexual dimorphism Males with protibial apical spine robust, abruptly downward bent, hook like. Abdomen of males with evident median fovea. Collecting information Individuals can be collected with light traps and from bovine feces in pasture of Urochloa decumbens (Stapf) R. D. Webster (Dellacasa et al. 2001 b; Mesquita et al. 2017).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB1FFE6FDE5FC9FFD8EF9F5.taxon	description	Figs 1 B, 33 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB1FFE6FDE5FC9FFD8EF9F5.taxon	diagnosis	Diagnosis Length 3.8 – 4.2 mm. Head with fine and coarse punctures; punctures symmetrically distributed; coarse punctures mixed with fine ones; coarse punctures more abundant on frons; clypeus with small emargination and weakly gibbous; finely punctate; gena obtusely angled. Pronotum with punctures of two sizes; larger punctures closer near lateral margins; posterior angles rounded. Elytral interstriae microreticulate; finely punctate; elytral interstriae eight and ten more elevated than others. Apical spine of metatibia somewhat shorter than the first tarsomere; first tarsomere slightly longer than the following three tarsomere combine.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB1FFE6FDE5FC9FFD8EF9F5.taxon	materials_examined	Material examined BRAZIL – Piauí • 2 ♂♂, 2 ♀♀; Floriano; 21 Oct. 2010; L. Carvalho leg; Cerrado s. str.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB1FFE6FDE5FC9FFD8EF9F5.taxon	description	Sexual dimorphism Males with head broader and more transverse that of female. Metatibia of male with first tarsomere about the same length of the superior apical spur; first tarsomere slightly longer than the following three tarsomeres. The female first tarsomere is shorter than the superior apical spur and shorter than the following three tarsomeres combined.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB1FFE5FDC5F9F1FD1BFBA7.taxon	description	Figs 1 A, 33 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB1FFE5FDC5F9F1FD1BFBA7.taxon	diagnosis	Diagnosis Length 3.0 – 4.0 mm. Head punctate; visibly downward bent, at least anteriorly; clypeus emarginate, margin rounded, convex and subrugosely granulate; frons with larger and more evident punctures. Pronotum with punctures of two sizes; punctures scarce near lateral margins. Elytra shagreened; interstriae flat with scarce and fine punctation.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB1FFE5FDC5F9F1FD1BFBA7.taxon	materials_examined	Material examined ARGENTINA – Corrientes • 168 specs; Barrio Víctor Colas; 27 ° 28 ′ 50 ″ S, 58 ° 47 ′ 18 ″ W; elev. 64 m; 21 Oct. 2012; M. I. Polesel et al. leg.; CEMT. – Mendoza • 10 specs; Lavalle, Reserva Biológica Telteca; 9 Dec. 2002; Arriágada and Diéguez leg.; light; CEMT • 187 specs; Desaguadero; 5 Feb. 2017; G. Arriágada leg; light; CEMT. – San Luis • 61 spec.; Belgrano, Fdo. El Molle; 33 ° 02 ′ 23 ″ S, 66 ° 30 ′ 48 ″ W; elev. 622 m; 7 Feb. 2017; G. Arriágada leg.; CEMT • 1 spec.; same data as for preceding; 18 Feb. – 8 Mar. 2012; human feces; CEMT • 2 specs; same data as for preceding; 14 – 18 Feb. 2018; J. Jensen leg.; light; CEMT. – Santiago del Estero • 1 spec.; Río Jan [Santiago del Estero, Río Salado]; Wagner leg.; CEMT. BRAZIL – Mato Grosso do Sul • 1 spec.; Selvíria, UNESP Farm; 22 Oct. 1991; C. A. H. Flechtmann leg.; Brachiaria decumbens pasture; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB1FFE5FDC5F9F1FD1BFBA7.taxon	description	Sexual dimorphism Males apical spur of protibia stout and flattened, obtusely truncate apically.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB1FFE5FDC5F9F1FD1BFBA7.taxon	discussion	Collecting information Specimens were collected in pastures of Urochloa decumbens, inside bovine faecal masses and with light and flight intercept traps (Mesquita et al. 2017).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB2FFE5FDDAFBA6FCECF83D.taxon	description	Figs 1 C – D, 33 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB2FFE5FDDAFBA6FCECF83D.taxon	diagnosis	Diagnosis Length 3.0 – 3.5 mm. Head with clypeus sinuate, punctate with fine and spaced punctures; genal suture evident and somewhat raised; frons with punctures larger than those of clypeus. Pronotum with punctures of different sizes; larger punctures more concentrated laterally; posterior angles obtuse. Elytra microreticulate; interstriae flat with small and spaced punctures. First tarsomere of metatarsus a bit longer than larger apical spine and as long as the next three tarsomeres combined.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB2FFE5FDDAFBA6FCECF83D.taxon	materials_examined	Material examined BRAZIL – Mato Grosso • 3 ♂♂, 9 ♀♀; Cocalinho, Fazenda Água Preta; 14 ° 08 ′ 22 ″ S, 51 ° 28 ′ 53.33 ″ W; elev. 250 m; 9 Nov. 2019; M. E. Maldaner and F. Z. Vaz-de-Mello leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB2FFE5FDDAFBA6FCECF83D.taxon	description	Sexual dimorphism Males with head transverse, abdomen with large median fovea. Males with frons finely punctate, females with frons coarsely punctate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB2FFE5FDDAFBA6FCECF83D.taxon	discussion	Collecting data Specimens were collected in pastures of sandy soil.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB3FFE4FD9FFE94FA83F86C.taxon	diagnosis	Diagnosis Length 2.5 – 5.0 mm. Body oval, glabrous, testaceous, red or brown. Head with clypeus not denticulate, emarginate medially; frontoclypeal suture distinct, not tuberculate. Pronotum punctate with punctures of two sizes; posteriorly lacking marginal line. Elytra without humeral teeth. Dorsal surface of protibia punctate. Meso- and metatibiae with distinct transverse carina in its lateral surface; metatibiae, apically fringed by setae of different sizes.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB3FFE4FD9FFE94FA83F86C.taxon	discussion	Remarks Blackburneus is currently comprised of 50 species distributed across the Neotropical, Neartic and Afrotropical (?) regions (Dellacasa et al. 2001 a, 2011; Schoolmeesters 2023). In the New World Blackburneus comprises 18 species (Dellacasa et al. 2011). For keys to all species see Dellacasa et al. (2011).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBCFFE9FE45FE95FD70F988.taxon	description	Figs 3 F, 34 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBCFFE9FE45FE95FD70F988.taxon	diagnosis	Diagnosis Length 3.0 – 3.5 mm. Body light to dark red. Head with punctures of two sizes; fine punctures everywhere distributed; coarser punctures present on frons, sometimes present on clypeus, near frontoclypeal suture; clypeus not sinuate, weakly gibbous; frontoclypeal suture evident; eyes wide, width equivalent to 1 / 3 or 1 / 2 that of frons. Pronotum with posterior margin and angles with line of coarse punctures. Elytra with interstriae of disc flat.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBCFFE9FE45FE95FD70F988.taxon	materials_examined	Type material examined Holotype BRAZIL • 1 ♂; “ First label [white paper, typewritten] Brazil: Amazonia / 70 km N. of Manaus / Fazenda Porto Alegre / MCS / DBFF project / [handwritten] 15 - [typewritten] VI- 86 / Bert Klein // Second label [white paper, typewritten] BAIT: HUMAN DUNG / HAB. MATURE FOREST / RESERVE: [handwritten] 2309 / NO.: / TIME: [handwritten] AM. // [red paper, printed] Blackburneus / amazonicus / Dellacasa M. & Alii, 2010 / HOLOTYPE. // [white paper, printed] // QR to the left; FSCA 00050480 ”; FSCA. Other material examined BRAZIL – Acre • 1 spec.; Assis Brasil; 10 ° 35 ′ 44 ″ S, 69 ° 37 ′ 12 ″ W; 21 Nov. 2016; Ortiz and Borges leg.; CDC light trap; CEMT. – Amapá • 1 spec.; Laranjal do Jari; 0 ° 36 ′ 29 ″ S, 52 ° 13 ′ 48 ″ W; 23 Jun. 2017; S. Milheira leg.; pitfall feces; CEMT. – Amazonas • 1 spec.; 60 km N of Manaus, Fazenda Esteio, ZF- 3, Km 23; 9 Apr. 1985; B. C. Klein leg.; Malaise trap; Reserva 1208; CEMT • 2 specs; same data as for preceding; 16 May 1986; Reserva 1112; INPA • 1 spec.; same data as for preceding; CEMT • 1 spec.; same data as for preceding; 21 May 1986; INPA • 1 spec.; same data as for preceding; 10 Jun. 1986; Reserva 1208; INPA • 1 spec.; same data as for preceding; 11 Jun. 1986; INPA • 1 spec.; Manaus PDBFF, Dimana, 10 ha; 2 ° 20 ′ S, 60 ° 06 ′ W; 16 Mar. 2005; M. R. Radtke leg.; dung-pitfall; INPA • 1 spec.; same data as for preceding; PDBFF, Km 41; 2 ° 26 ′ S, 59 ° 45 ′ W; 8 Apr. 2005; INPA • 1 spec.; Reserva Ducke; 2 ° 55 ′ S; 59 ° 58 ′ W; 1 Mar. 2005; M. R. Radtke leg.; dung-pitfall; CEMT • 2 specs; same data as for preceding; INPA • 1 spec.; Manaus ZF- 03, Km 41; 1996 – 1997; A. Ellen leg.; in feces of Alouatta seniculus; CEMT • 1 spec.; Manaus, P Q. Laranjeiras; 6 Jun. 1981; J. Arias leg.; INPA • 1 spec.; same data as for preceding; 3 Jun. 1981; INPA • 2 specs; Manaus; 29 Nov. 1977; B. C. Ratcliffe leg.; INPA • 1 spec.; same data as for preceding; 6 Dec. 1977; CEMT • 2 specs; same data as preceding; 20 Dec. 1977; CEMT • 9 specs; same data as preceding; INPA • 1 spec.; same data as for preceding; 27 Jun. 1978; CEMT • 2 specs; same data as for preceding; INPA • 1 spec.; same data as for preceding; 25 May 1978; INPA • 1 spec.; Manaus, Reserva Ducke, 26 km NW of Manaus; 25 Oct. 1977; CEMT • 1 spec.; same data as for preceding; INPA • 1 spec.; same data as for preceding; 11 Apr. 1978; CEMT • 2 specs; same data as for preceding; INPA • 1 spec.; same data as for preceding; 30 May 1978; CEMT • 1 spec.; same data as for preceding; 30 May 1978; INPA • 1 spec.; same data as for preceding; 13 Jun. 1978; CEMT • 10 specs; same data as for preceding; INPA • 1 spec.; same data as for preceding; 1 Aug. 1978; INPA • 1 spec.; same data as for preceding; Reserva Ducke, BR 010, Km 26; 4 Oct. 1977; CEMT • 1 spec.; same data as for preceding; 3 Jan. 1978; INPA • 3 specs; same data as for preceding; 10 Jan. 1978; INPA • 1 spec.; same data as for preceding; 17 Jan. 1978; INPA • 1 spec.; same data as for preceding; 31 Jan. 1978; CEMT • 1 spec.; same data as for preceding; INPA • 1 spec.; same data as for preceding; 7 Feb. 1978; INPA • 1 spec.; same data as for preceding; 14 Mar. 1978; INPA • 1 spec.; same data as for preceding; 28 Mar. 1978; CEMT • 1 spec.; same data as for preceding; 15 Aug. 1978; INPA • 1 spec.; Manaus, Reserva Ducke, AM. 010, Km 28; 13 Sep. 1977; INPA • 2 specs; same data as for preceding; Reserva Ducke; Jun. 2011; J. Bonanomi leg.; CEMT • 4 specs; same data as for preceding; 3 ° 00 ′ 20 ″ S, 59 ° 56 ′ 25 ″ W; 16 May 2012; CEMT • 1 spec.; Mamicoré, BR 319, Km 450, Resex Lago Capanã Grande; 5 ° 54 ′ 33 ″ S, 62 ° 23 ′ 56 ″ W; elev. 850 m; 30 Oct. – 1 Nov. 2019; A. R. Azevedo leg.; human feces; CEMT. – Mato Grosso • 1 spec.; Alta Floresta, P. E. Cristalino; 9 ° 32 ′ 45 ″ S, 55 ° 54 ′ 56 ″ W; elev. 248 m; 15 – 17 Nov. 2019; R. A. Azevedo leg.; human feces; CEMT • 3 specs; same data as for preceding; 9 ° 40 ′ 59 ″ S, 55 ° 54 ′ 57 ″ W; elev. 260 m; 15 – 17 Nov. 2019; CEMT • 11 specs; same data as for preceding; 9 ° 35 ′ 50 ″ S, 55 ° 53 ′ 40 ″ W; elev. 287 m; 15 – 17 Nov. 2019; CEMT • 2 specs; Brasnorte, Fazenda Hermínia; 11 ° 57 ′ 02 ″ S, 58 ° 14 ′ 49 ″ W; elev. 311 m; 29 – 30 Nov. 2017; A. F. Machado leg.; pitfall, floresta; CEMT • 2 specs; same data as for preceding; elev. 309 m; 29 Nov. – 1 Dec. 2017; borda, human feces; CEMT • 1 spec.; same data as for preceding; 11 ° 56 ′ 59 ″ S, 58 ° 14 ′ 47 ″ W; elev. 304 m; 29 Nov. – 1 Dec. 2017; CEMT • 1 spec.; same data as for preceding; 11 ° 57 ′ 02 ″ S, 58 ° 14 ′ 49 ″ W; elev. 311 m; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 9 ° 50 ′ 19 ″ S, 58 ° 15 ′ 3 ″ W; 13 Dec. 2009; Vaz-de-Mello leg.; matinha borda [matinha border]; pitfall; CEMT • 2 specs; same data as for preceding; 9 ° 51 ′ 40 ″ S, 58 ° 13 ′′ 12 ″ W; 2 – 4 Nov. 2021; J. Mariano et al. leg.; human feces; prainha; CEMT • 1 spec.; same data as for preceding; 9 ° 50 ′ 47 ″ S, 58 ° 14 ′ 09 ″ W; Talhão 46 b, human feces; CEMT • 1 spec.; same data as for preceding; 9 ° 50 ′ 44 ″ S, 58 ° 14 ′ 3 ″ W; CEMT • 2 specs; same data as for preceding; 9 ° 51 ′ 31 ″ S, 58 ° 14 ′ 23 ″ W; Talhão 37; human feces; CEMT • 1 spec.; same data as for preceding; 9 ° 51 ′ 28 ″ S, 58 ° 14 ′ 23 ″ W; CEMT • 2 specs; same data as for preceding; 9 ° 51 ′ 26 ″ S, 58 ° 14 ′ 24 ″ W; CEMT • 1 spec.; same data as for preceding; 9 ° 50 ′ 53 ″ S, 58 ° 15 ′ 19 ″ W; 4 – 6 Nov. 2021; morrinho; human feces; CEMT • 1 spec.; same data as for preceding; 9 ° 51 ′ 01 ″ S, 58 ° 14 ′ 11 ″ W; 5 – 7 Nov. 2021; Talhão 48 b; human feces; CEMT • 1 spec.; same data as for preceding; 9 ° 49 ′ 29 ″ S, 58 ° 16 ′ 14 ″ W; Talhão 70 A; human feces; CEMT • 1 spec.; same data as for preceding; 9 ° 49 ′ 26 ″ S, 58 ° 16 ′ 15 ″ W; CEMT • 4 specs; same data as for preceding; 9 ° 51 ′ 47 ″ S, 58 ° 15 ′ 50 ″ W; Talhão 14 b; human feces; CEMT • 4 specs; same data as for preceding 9 ° 51 ′ 48 ″ S, 58 ° 15 ′ 50 ″ W; CEMT • 1 spec.; same data as for preceding; 9 ° 50 ′ 57 ″ S, 58 ° 14 ′ 03 ″ W; Talhão 47; human feces; CEMT • 3 specs; same data as for preceding; 9 ° 52 ′ 08 ″ S, 58 ° 15 ′ 38 ″ W; 18 – 20 Oct. 2021; Talhão 19; human feces; CEMT • 1 spec.; same data as for preceding; 9 ° 52 ′ 09 ″ S, 58 ° 15 ′ 39 ″ W; CEMT • 4 specs; same data as for preceding; 9 ° 50 ′ 49 ″ S, 58 ° 14 ′ 4 ″ W; 19 – 21 Oct. 2021; Talhão 46 b; human feces; CEMT • 1 spec.; same data as for preceding; 9 ° 51 ′ 01 ″ S, 58 ° 14 ′ 53 ″ W; Talhão 37; human feces; CEMT • 3 specs; same data as for preceding; 9 ° 49 ′ 15 ″ S, 58 ° 17 ′ 18 ″ W; 27 – 29 Oct. 2021; Trilha Castanheira, human feces; CEMT • 10 specs; same data as for preceding; 9 ° 50 ′ 23 ″ S, 58 ° 15 ′ 13 ″ W; 28 – 30 Oct. 2021; matinha, human feces; CEMT • 6 specs; same data as for preceding; 9 ° 50 ′ 25 ″ S, 58 ° 15 ′ 11 ″ W; CEMT • 5 specs; same data as for preceding; 9 ° 50 ′ 27 ″ S, 58 ° 15 ′ 8 ″ W; CEMT • 5 specs; same data as for preceding; 9 ° 50 ′ 26 ″ S, 58 ° 15 ′ 10 ″ W; CEMT • 1 spec.; same data as for preceding; 9 ° 49 ′ 14 ″ S, 58 ° 15 ′ 10 ″ W; ppbio; human feces; CEMT • 2 specs; same data as for preceding; 9 ° 49 ′ 12 ″ S, 58 ° 15 ′ 30 ″ W; CEMT • 2 specs; same data as for preceding; 9 ° 49 ′ 17 ″ S, 58 ° 15 ′ 30 ″ W; CEMT • 1 spec.; Diamantino; 16 Mar. 2018; L. G. Nunes and E. Carvalho leg.; CEMT • 1 spec.; same data as for preceding; 13 ° 48 ′ 7 ″ S, 56 ° 41 ′ 32 ″ W; L. G. Nunes and E. Carvalho leg.; human feces; CEMT • 1 spec.; same data as for preceding; 13 ° 51 ′ 2 ″ S, 56 ° 41 ′ 8 ″ W; 17 Mar. 2018; CEMT • 2 specs; São José do Rio Claro; 15 ° 51 ′ 32 ″ S, 46 ° 41 ′ 04 ″ W; 16 Mar. 2018; L. G. Nunes and E. Carvalho leg.; human feces; CEMT • 2 specs; same data as for preceding; 15 ° 53 ′ 50 ″ S, 56 ° 40 ′ 42 ″ W; 29 Jun. 2018; CEMT • 3 specs; same data as for preceding; 30 Jun. 2018; CEMT. – Pará • 1 spec.; Almeirim, Jari; 4 Feb. 2011; F. M. França leg.; feces; CEMT • 19 specs; same data as for preceding; 5 Feb. 2011; CEMT • 9 specs; same data as for preceding; 9 Feb. 2011; CEMT • 72 specs; same data as for preceding; 16 Feb. 2011; CEMT • 91 specs; same data as for preceding; 19 Feb. 2011; CEMT • 77 specs; same data as for preceding; 23 Feb. 2011; CEMT • 63 specs; same data as for preceding; 26 Feb. 2011; CEMT • 39 specs; same data as for preceding; 2 Mar. 2011; CEMT • 23 specs; same data as for preceding; 5 Mar. 2011; CEMT • 2 specs; same data as for preceding; 9 Mar. 2011; CEMT • 22 specs; same data as for preceding; 12 Mar. 2011; CEMT • 36 specs; same data as for preceding; 16 Mar. 2011; CEMT • 22 specs; same data as for preceding; 19 Mar. 2011; CEMT • 2 specs; same data as for preceding; 23 Feb. 2016; CEMT • 1 spec.; Belém, Comunidade Boa Vista; 1 ° 30 ′ 57 ″ S, 48 ° 26 ′ 18 ″ W; 12 Feb. 2014; Silva et al. leg.; human feces; CEMT • 2 specs; same data as for preceding; 1 ° 31 ′ 15 ″ S, 48 ° 26 ′ 17 ″ W; CEMT • 2 specs; same data as for preceding; 1 ° 31 ′ 14 ″ S, 48 ° 26 ′ 12 ″ W; CEMT • 1 spec.; same data as for preceding; 1 ° 31 ′ 15 ″ S, 48 ° 26 ′ 18 ″ W; CEMT • 1 spec.; Belterra; 3 ° 27 ′ 52 ″ S, 54 ° 51 ′ 26 ″ W; 15 Mar. 2019; F. França leg.; logging forest fire; human and pig dung; CEMT • 3 specs; same data as for preceding; FLONA-Tapajós; 3 ° 25 ′ 46 ″ S, 54 ° 50 ′ 33 ″ W; logging forest fire; CEMT • 1 spec.; same data as for preceding; 3 ° 20 ′ 9 ″ S, 54 ° 57 ′ 21 ″ W; 20 Mar. 2019; primary forest; CEMT • 9 specs; same data as for preceding; 3 ° 01 ′ 6 ″ S, 55 ° 00 ′ 17 ″ W; 21 Mar. 2019; CEMT • 1 spec.; same data as for preceding; 3 ° 01 ′ 43 ″ S, 55 ° 00 ′ 40 ″ W; CEMT • 4 specs; same data as for preceding; 3 ° 19 ′ 12 ″ S, 54 ° 57 ′ 37 ″ W; 22 Mar. 2019; CEMT • 2 specs; same data as for preceding; 3 ° 20 ′ 9 ″ S, 54 ° 57 ′ 21 ″ W; CEMT • 2 specs; same data as for preceding; 3 ° 16 ′ 58 ″ S, 54 ° 51 ′ 15 ″ W; 25 Mar. 2019; secondary forest; CEMT • 1 spec.; same data as for preceding; 3 ° 1 ′ 6 ″ S, 55 ° 20 ′ 17 ″ W; primary forest; CEMT • 6 specs; Bragança; 1 ° 10 ′ 46 ″ S, 46 ° 40 ′ 54 ″ W; 5 Mar. 2019; F. M. França leg.; primary forest; human and pig dung; CEMT • 1 ♂, 2 specs; same data as for preceding; 1 ° 10 ′ 39 ″ S, 46 ° 39 ′ 54 ″ W; 5 Mar. 2019; F. M. França leg.; secondary forest; human and pig dung; CEMT • 2 specs; Marabá; 5 ° 47 ′ 04 ″ S, 48 ° 54 ′ 55 ″ W; 21 Feb. 2019; F. M. França leg.; logging forest fire; human and pig dung; CEMT • 1 spec.; Mojuí dos Campos; 2 ° 41 ′ 36 ″ S, 54 ° 29 ′ 43 ″ W; 14 Mar. 2019; F. M. França leg.; logging forest fire; human and pig dung; CEMT • 2 specs; Santarém; 3 ° 07 ′ 49 ″ S, 54 ° 51 ′ 24 ″ W; 25 Mar. 2019, F. França and V. Neto leg.; logg. for. fire; human and pig dung; CEMT • 4 specs; same data as for preceding; FLONA-Tapajós; 3 ° 19 ′ 12 ″ S, 55 ° 57 ′ 37 ″ W; 22 Mar. 2019; F. França leg.; primary forest; CEMT • 2 specs; Usina Hidrelétrica de Curuá-una; 2 ° 48 ′ 45 ″ S, 54 ° 17 ′ 34 ″ W; 22 Jun. 2018; Ganança and V. Neto leg.; human feces; CEMT • 1 spec.; same data as for preceding; 2 ° 47 ′ 47 ″ S, 54 ° 17 ′ 42 ″ W; CEMT • 4 specs; same data as for preceding; 23 Jun. 2018; CEMT. – Rondônia • 2 specs; Parque Estadual Guajará-Mirim; 10 ° 18 ′ S, 64 ° 32 ′ W; 23 Nov. 2019; M. A. P. A. Silveira leg.; human feces; CEMT • 1 spec.; same data as for preceding; 25 Nov. 2016; CEMT. – Roraima • 2 specs; Cantá; Nov. 1996; Vaz-de-Mello and Ribeiro leg.; CEMT. FRENCH GUIANA • 6 specs; Cayenne, Monte Proche du Munt; 4 ° 18 ′ 58 ″ N, 53 ° 17 ′ 10 ″ W; 2 Nov. 2013; SAEG leg.; Baruol (BAR), FIT (V); CEMT • 1 spec.; Saint Laurent de Maroni, Belvedère Saül; 3 ° 37 ′ 22 ″ S, 53 ° 12 ′ 57 ″ W; elev. 326 m; 14 Mar. 2011; SAEG leg.; FIT; CEMT • 1 spec.; same data as for preceding; 30 Mar. 2011; FIT-V 5; CEMT • 1 spec.; same data as for preceding; 20 May 2011; FIT (PIV); CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBCFFE9FE45FE95FD70F988.taxon	description	Sexual dimorphism Males with metaventrite plate concave, longitudinal groove evident; females with metaventrite plate somewhat flat, longitudinal groove weak.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBCFFE9FE45FE95FD70F988.taxon	discussion	Collecting information Specimens were collected on human or pig feces from regions of mature tropical forests, secondary and logging forests (Dellacasa et al. 2011).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBEFFE8FDD1F9FFFE39FCD1.taxon	description	Figs 2 B, 34 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBEFFE8FDD1F9FFFE39FCD1.taxon	diagnosis	Diagnosis Length 4.0 – 4.5 mm. Body wide, reddish testaceous. Head with sparse punctures; clypeus with sparse and almost imperceptibly small punctures; emarginate medially; distinctively gibbous; frontoclypeal suture evident, medially depressed; frons with coarse punctures. Pronotum with punctures of two sizes; large punctures scattered, closer near lateral margins. Elytra shagreened; interstriae flat, with two rows of small punctures; interstria near elytral suture darker than the others; fifth interstria, from elytral suture, wider than the rest near the anterior margin of elytra.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBEFFE8FDD1F9FFFE39FCD1.taxon	materials_examined	Material examined BRAZIL – Mato Grosso do Sul • 1 spec.; Selvíria, UNESP farm; 20 Dec. 1991; S. R. Rodrigues; Guzera bovine dropping; Brachiaria decumbens pasture; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBEFFE8FDD1F9FFFE39FCD1.taxon	description	Sexual dimorphism Male with frontoclypeal margin elevated medially; protibial apical spine sub-cylindrical, apically rounded.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBEFFE8FDD1F9FFFE39FCD1.taxon	discussion	Collecting information Individuals were collected at altitudes ranging from 250 to 1100 metres. Specimens collected within feces of Guzera cattle droppings in pastures of Urochloa decumbens and with flight intercept traps (Dellacasa et al. 2011).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBFFFE8FDCFFC14FA83F839.taxon	description	Figs 2 C, 33 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBFFFE8FDCFFC14FA83F839.taxon	diagnosis	Diagnosis Length 2.5 – 3.0 mm. Body brown. Head entirely punctate with moderate in size punctures; punctures on frons no bigger than those of clypeus; clypeus sinuate, weakly gibbous; frontoclypeal suture finely impressed. Pronotum with moderate and large punctures; moderate punctures evenly spread across pronotal surface; large punctures scarce on pronotal disc and near posterior margin, closely grouped near lateral margins. Elytral interstriae finely punctate and microreticulate, microreticulation stronger closer to posterior margins; interstriae somewhat convex. First metatarsomere longer than large apical spur.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBFFFE8FDCFFC14FA83F839.taxon	materials_examined	Material examined BRAZIL – Bahia • 4 specs; Encruzilhada; Nov. 1972; M. Alvarenga leg.; CEMT. – Distrito Federal • 2 specs; Planaltina, Embrapa Cerrados; 15 ° 36 ′ 20 ″ S, 47 ° 42 ′ 26 ″ W; 30 Nov. 2005; C. Oliveira leg.; fragmento cerrado [fragment of cerrado]; light; CEMT. – Mato Grosso • 1 spec.; Chapada dos Guimarães; 15 ° 24 ′ 31 ″ S, 55 ° 45 ′ 53 ″ W; 2 Dec. 2015; A. Frolov and L. Akhmetova leg.; light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBFFFE8FDCFFC14FA83F839.taxon	description	Sexual dimorphism Slight. Males with apical spur of protibia stout, inwardly curved, more strongly microreticulate. Females with apical spur slender and feebly convex.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBFFFE8FDCFFC14FA83F839.taxon	discussion	Collecting information Specimens have been collected at altitudes ranging from 800 to 1380 metres above sea level. Black light and flight intercept traps are methods that can be used for collecting this species (Dellacasa et al. 2011).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB9FFEEFDF0FE95FCE4F825.taxon	description	Figs 3 B – C, 34 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB9FFEEFDF0FE95FCE4F825.taxon	description	Blackburneus furcatus – Paulian 1942: 75.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB9FFEEFDF0FE95FCE4F825.taxon	diagnosis	Diagnosis Length 2.5 – 3.0 mm. Body reddish testaceous. Head, up to the frontoclypeal suture, with fine punctation; coarse punctures restricted to frons; clypeus medially emarginate; with evident, anteriorly furcate carina resulting in two oblique carina; frontoclypeal suture medially depressed, elevated laterally. Pronotum with punctures of two sizes; small punctures distributed throughout pronotal surface; large punctures concentrated near lateral margins. Elytral interstriae shagreened, flat; finely punctate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB9FFEEFDF0FE95FCE4F825.taxon	materials_examined	Material examined ARGENTINA – Córdoba • 5 specs; 10 km NW of Alpa Corral; 8 – 9 Feb. 2009; F. C. Ocampo leg.; CEMT. BRAZIL – Distrito Federal • 2 specs; Gama-Cabeça-de-Veado; 15 ° 56 ′ 49 ″ S, 47 ° 56 ′ 58 ″ W; 15 Dec. 2018; W. L. Cunha leg.; cerrado s. str.; pitfall with feces; CEMT • 1 spec.; same data as for preceding; 29 Nov. 2018; CEMT • 1 spec.; Brasília; elev. 1100 m; Jan. 2001; N. Degallier leg.; light; CEMT. – Mato Grosso • 1 spec.; Chapada dos Guimarães, Fazenda Jardim; 9 Jun. 2012; V. C. Jorge leg.; CEMT • 1 spec.; same data as for preceding; 29 Sep. 2012; CEMT • 1 spec.; same data as for preceding; 13 Oct. 2012; CEMT • 1 spec.; Chapada dos Guimarães, Santuário dos Elefantes; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 5 ″ W; 22 Jan. 2020; H. Sehn et al. leg.; cow feces; CEMT • 1 spec.; Porto Estrela, Estação Ecológica [Ecological Station] Serra das Araras, Trilha Boca do José; Nov. 2017; T. F. Conceição leg.; Cerrado; pitfall; CEMT. – Mato Grosso do Sul • 1 ♀; Selvíria; UNESP farm; 20 ° 22 ′ 41 ″ S, 51 ° 25 ′ 10 ″ W; 12 Jan. 2005; F. W. Mesquita leg.; Brachiaria decumbens pasture; black light; CEMT • 1 ♂; same data as for preceding; 27 Mar. 2005; CEMT • 1 ♂; same data as for preceding; 3 Dec. 2005; CEMT • 1 ♂; same data as for preceding; 20 ° 22 ′ 53 ″ S, 51 ° 24 ′ 39 ″ W; 9 Jan. 2007; CEMT • 1 ♀; same data as for preceding; 20 ° 22 ′ 41 ″ S, 51 ° 26 ′ 10 ″ W; F. Oikawa leg.; CEMT • 1 ♂; Santa Ofélia farm; 1 May 2007; C. A. H. Flechtmann leg.; Brachiaria brizantha pasture; bovine droppings; CEMT. – Minas Gerais • 1 spec.; Itumirim, Serra do Janela; 21.340 ° S, 44.801 ° W; 19 Oct. 2016; L. Vieira and F. Z. Vaz-de-Mello leg.; light; CEMT • 1 spec.; same data as for preceding; 21 ° 20 ′ 24 ″ S, 44 ° 48 ′ 06 ″ W; CEMT. – Pará • 1 spec.; Jaguariaiva, Parque Estadual do Cerrado; 3 Feb. 2011; Grossi and Parizotto leg.; CEMT • 3 specs; Marabá; 5 ° 42 ′ 4 ″ S, 49 ° 13 ′ 45 ″ W; 22 Feb. 2019; F. França leg.; secondary forest; human and pig dung; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB9FFEEFDF0FE95FCE4F825.taxon	description	Sexual dimorphism Subtle. Females with punctures of pronotal disc denser compared with males.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFB9FFEEFDF0FE95FCE4F825.taxon	discussion	Collecting information Blackburneu s furcatus can be found at altitudes up to 1000 m. Specimens can be collected with light traps and flight intercept traps. Some were collected in Urochloa brizantha (A. Rich.) R. D. Webster and Urochloa decumbens pastures with the methods previously mentioned and with traps baited with bovine dung (Dellacasa et al. 2011; Mesquita et al. 2017).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBBFFECFDE7FE95FE34F9DF.taxon	description	Figs 2 A, 33 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBBFFECFDE7FE95FE34F9DF.taxon	diagnosis	Diagnosis Length 3.5 – 4.5 mm. Body elongate, sub-parallel-sided, brown with anterior and lateral margins of head and pronotum lighter. Clypeus sinuate, medially weakly convex; frontoclypeal suture evident. Pronotum punctate with punctures of two sizes; coarse punctures near lateral margins; small punctures present everywhere. Elytra shagreened; interstriae with two rows of small punctures.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBBFFECFDE7FE95FE34F9DF.taxon	materials_examined	Material examined BRAZIL – Distrito Federal • 27 specs; Planaltina, Embrapa Cerrados, fragment of cerrado; 15 ° 36 ′ 20 ″ S, 47 ° 42 ′ 26 ″ W; 23 Sep. 2005; C. Oliveira leg.; light; CEMT. – Mato Grosso • 1 spec.; Chapada dos Guimarães, Santuário dos Elefantes; 2 Nov. 2019; Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; 15 ° 18 ′ 50 ″ S, 55 ° 24 ′ 29 ″ W; 21 Jan. 2020; H. Sehn et al. leg.; elephant feces; CEMT • 2 specs; same data as for preceding; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 14 ″ W; 22 Jan. 2020; CEMT • 1 spec.; same data as for preceding; horse feces; CEMT • 8 spec.; same data as for preceding; 23 Jan. 2020; elephant feces; CEMT • 3 specs; same data as for preceding; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 05 ″ W; 23 Jan. 2020; horse feces; CEMT • 4 specs; same data as for preceding; 15 ° 18 ′ 50 ″ S, 55 ° 24 ′ 12 ″ W; 24 Jan. 2020; elephant feces; CEMT • 9 spec.; same data as for preceding; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 14 ″ W; 25 Jan. 2020, H. Sehn et al. leg.; CEMT • 15 specs; same data as for preceding; H. Sehn et al. leg.; cow feces; CEMT • 5 specs; same data as for preceding; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 07 ″ W; CEMT • 1 spec.; same data as for preceding; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 14 ″ W; 25 Jan. 2020; horse feces; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 9 ° 50 ′ 19 ″ S, 58 ° 15 ′ 3 ″ W; 25 Oct. 2014; A. Asenjo leg.; light; CEMT • 1 spec.; Juína, Fazenda Quiles; 29 Nov. 2012; N. Degallier leg.; carcaça; CEMT • 1 spec.; Santo Antônido do Leverger, Águas quentes; 23 Nov. 2016; A. Frolov and M. Cupello leg.; UV light; CEMT. – Mato Grosso do Sul • 1 spec.; Campo Grande; 1990 – 1992; I. Bianchin leg.; CEMT. – Minas Gerais • 1 spec.; Campanha, Rod. Vital Brazil; 21 ° 47 ′ 49 ″ S, 45 ° 25 ′ 40 ″ W; 5 Jun. 2016; F. França leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBBFFECFDE7FE95FE34F9DF.taxon	description	Sexual dimorphism Males with medial longitudinal groove of metaventrite anteriorly foveate, females without fovea.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBBFFECFDE7FE95FE34F9DF.taxon	discussion	Collecting information Blackburneus indio was found at altitudes up to 980 metres. Specimens can be collected with UV light traps and are attracted to feces of elephant, cattle and Tapirus Brisson, 1762 (Dellacasa et al. 2011; Mesquita et al. 2017).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBBFFD1FDD6F90EFCF7FA45.taxon	description	Figs 3 D – E, 35 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBBFFD1FDD6F90EFCF7FA45.taxon	diagnosis	Diagnosis Length 3.0 – 4.5 mm. Body, light to dark red. Head with punctures of two sizes; fine punctures mixed with coarse punctures, the latter more numerous on frons; clypeus slightly emarginate; strongly gibbous medially; frontoclypeal suture evident, medially depressed; eyes wide, equal to half the length between them. Pronotum with punctures large and small; small punctures regularly spaced on the pronotal surface; large punctures closer and more abundant near lateral margins. Elytra shagreened; striae fine, finely punctate; interstriae flat, with two rows of small punctures.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBBFFD1FDD6F90EFCF7FA45.taxon	materials_examined	Type material examined Lectotype BRAZIL • ♀; “ [white paper, handwritten] São Paulo, Brasilien. // [white paper, handwritten] Laxepunct. / m. // [orange paper, typewritten] TYPUS. // [brown paper, typewritten] 121. // [red paper printed] Aphodius laxepunctatus / Schmidt A., 1910 / LECTOTYPUS / Dellacasa M. & G. des., 2002. // [green paper, printed] Naturhistoriska / Riksmuseet / Stockholm / Loan no 630 / 00. // [white paper, printed] Blackburneus / laxepunctatus / (Schmidt A., 1910) / Dellacasa M. & G. det., 2002. // [white paper, printed] NHRS-ALJB / 0000000507 ”; NRM. Paralectotypes BRAZIL • 1 ♂; “ [white paper, handwritten] São Paulo, Brasilian. // [red paper printed] Aphodius laxepunctatus / Schmidt A., 1910 / LECTOTYPUS / Dellacasa M. & G. des., 2002. // [green paper, printed] Naturhistoriska / Riksmuseet / Stockholm / Loan no 633 / 00. // [white paper, printed] NHRS-ALJB / 0000000508 ”; NRM • 1 ♂; “ [white paper, handwritten] São Paulo, Brasilian. // [red paper printed] Aphodius laxepunctatus / Schmidt A., 1910 / LECTOTYPUS / Dellacasa M. & G. des., 2002. // [green paper, printed] Naturhistoriska / Riksmuseet / Stockholm / Loan no 635 / 00. // [white paper, printed] NHRS-ALJB / 0000000509 ”; NRM. Other material examined BRAZIL – Acre • 1 spec.; Assis Brasil; 10 ° 35 ′ 44 ″ S, 69 ° 37 ′ 12 ″ W; 20 Sep. 2016; Otriz and Borges leg.; CDC light trap; CEMT • 1 spec.; same data as for preceding; 23 Sep. 2016; CEMT • 1 spec.; Mâncio Lima, P. N. Serra do Divisor; 7 ° 27 ′ 04 ″ S, 73 ° 38 ′ 53 ″ W; elev. 225 m; 6 – 8 Dec. 2019; R. A. Azevedo leg.; human feces; CEMT • 6 specs; same data as for preceding; 7 ° 27 ′ 04 ″ S, 73 ° 38 ′ 57 ″ W; CEMT • 2 specs; same data as for preceding; 7 ° 24 ′ 60 ″ S, 73 ° 37 ′ 59 ″ W; elev. 254 m; CEMT • 16 specs; same data as for preceding; 7 ° 26 ′ 46 ″ S, 79 ° 39 ′ 28 ″ W; elev. 236 m; 7 – 9 Dec. 2019; CEMT • 1 spec.; Rio Branco, Fazenda Catuaba; Mar. 1997; F. Z. Vaz-de-Mello et al. leg.; primary forest; CEMT. – Amapá • 2 specs; Laranjal do Jari; 0 ° 58 ′ 06.4 ″ S, 52 ° 44 ′ 40.5 ″ W; 4 – 6 Jun. 2017; S. Milheiras leg.; pitfall feces; CEMT • 1 spec.; same data as for preceding; 0 ° 35 ′ 29 ″ S, 52 ° 13 ′ 48 ″ W; 23 Jun. 2017; CEMT. – Amazonas • 1 spec.; 7 Feb. 1978; B. C. Ratcliffe leg.; INPA • 3 specs; 90 km N of Manaus; Oct. – Nov. 1996; E. Andresen leg.; CEMT • 1 spec.; Borba, BR- 319, Km 220; 4 ° 23 ′ 26 ″ S, 60 ° 56 ′ 11 ″ W; 30 Nov. 2015; D. Pires leg.; human feces; CEMT • 1 spec.; Caseiro, BR- 319, Km 168; 4 ° 08 ′ 58 ″ S, 60 ° 43 ′ 44 ″ W; 2 Dec. 2015; D. Pires leg.; human feces; CEMT • 3 specs; same data as for preceding; 4 ° 07 ′ 58 ″ S, 60 ° 44 ′ 09 ″ W; CEMT • 1 spec.; same data as for preceding; 4 ° 08 ′ 28 ″ S, 60 ° 43 ′ 57 ″ W; CEMT • 1 ♀, 1 spec.; same data as for preceding; 4 ° 09 ′ 28 ″ S, 60 ° 43 ′ 32 ″ W; CEMT • 1 ♂, 2 specs; same data as for preceding; 04 ° 07 ′ 39 ″ S, 60 ° 43 ′ 43 ″ W; 3 Dec. 2015; CEMT • 2 specs; same data as for preceding; 4 ° 08 ′ 09 ″ S, 60 ° 43 ′ 31 ″ W; CEMT • 2 specs; same data as for preceding; 4 ° 08 ′ 40 ″ S, 60 ° 43 ′ 20 ″ W; CEMT • 2 specs; same data as for preceding; 4 ° 09 ′ 10 ″ S, 60 ° 43 ′ 08 ″ W; CEMT • 1 ♂, 7 specs; Manaquiri, BR- 319, Km 100; 3 ° 40 ′ 34 ″ S, 60 ° 17 ′ 46 ″ W; 6 Dec. 2015; D. Pires leg.; human feces; CEMT • 1 spec.; same data as for preceding; 3 ° 41 ′ 08 ″ S, 60 ° 18 ′ 40 ″ W; CEMT • 2 specs; same data as for preceding; 3 ° 40 ′ 57 ″ S, 60 ° 19 ′ 24 ″ W; 7 Dec. 2015; CEMT • 2 specs; same data as for preceding; 3 ° 40 ′ 07 ″ S, 60 ° 18 ′ 03 ″ W; CEMT • 1 spec.; same data as for preceding; 3 ° 40 ′ 23 ″ S, 60 ° 18 ′ 29 ″ W; CEMT • 3 specs; Manaus ZF- 03, Km 41; 1996 – 1997; B. C. Ratcliffe leg.; in feces of Alouatta seniculus; CEMT • 4 specs; same data as for preceding; INPA • 3 specs; Manaus, Reserva Adolfo Ducke; 2 ° 55 ′ S, 59 ° 58 ′ W; 1 Mar. 2005; M. Radtke leg.; dung-pitfall; CEMT, INPA • 3 specs; same data as for preceding; 3 ° 00 ′ 20 ″ S, 59 ° 56 ′ 25 ″ W; 16 May 2012; J. Bananomi leg.; CEMT • 1 spec.; same data as for preceding; Reserva Ducke, BR 010, Km 26; 17 Jan. 1978; B. C. Ratcliffe leg.; INPA • 1 spec.; same data as for preceding; 26 km NW of Manaus; 13 Sep. 1977; B. C. Ratcliffe leg.; dung-pitfall trap; INPA • 1 spec.; same data as for preceding; 25 Oct. 1977; INPA • 1 spec.; same data as for preceding; 15 Nov. 1977; INPA • 1 spec.; same data as for preceding; 20 Nov. 1977; CEMT • 1 spec.; same data as for preceding; INPA • 1 spec.; same data as for preceding; 22 Nov. 1977; CEMT • 1 spec.; same data as for preceding; 6 Dec. 1977; INPA • 1 spec.; same data as for preceding; 13 Dec. 1977; INPA • 1 spec.; same data as for preceding; 7 Mar. 1978; INPA • 1 spec.; same data as for preceding; 14 Mar. 1978; INPA • 1 spec.; same data as for preceding; 21 Mar. 1978; INPA • 1 spec.; same data as for preceding; 4 Apr. 1978; INPA • 1 spec.; same data as for preceding; 11 Apr. 1978; CEMT • 4 specs; same data as for preceding; INPA • 1 spec.; same data as for preceding; 2 May 1978; INPA • 1 spec.; same data as for preceding; 30 May 1978; CEMT • 2 specs; same data as for preceding; INPA • 1 spec.; same data as for preceding; 4 Jul. 1978; INPA • 1 spec.; same data as for preceding; 13 Dec. 1977; INPA • 1 spec.; same data as for preceding; 24 Jan. 1978; INPA • 1 spec.; ResEx Lago Capanã Grande, 5 ° 54 ′ 33 ″ S, 62 ° 23 ′ 56 ″ W; elev. 750 m; 30 Oct. – 1 Nov. 2019; R. A. Azevedo leg.; human feces; CEMT • 3 specs; same data as for preceding; elev. 850 m; CEMT • 7 specs; same data as for preceding; elev. 1050 m; CEMT • 1 spec.; same data as for preceding; elev. 950 m; M. A. P. A. Silveira leg.; CEMT • 6 specs; same data as for preceding; 5 ° 58 ′ 52 ″ S, 62 ° 27 ′ 38 ″ W; elev. 800 m; CEMT • 3 specs; same data as for preceding; 5 ° 54 ′ 33 ″ S, 62 ° 23 ′ 56 ″ W; elev. 850 m; CEMT • 8 specs; same data as for preceding; elev. 1050 m; CEMT • 1 spec.; same data as for preceding; elev. 600 m; 31 Oct. 2 Nov. 2019; human feces; R. A. Azevedo leg.; CEMT • 2 specs; same data as for preceding; 6 ° 01 ′ 05 ″ S, 62 ° 29 ′ 24 ″ W; elev. 700 m; CEMT • 1 spec.; same data as for preceding; elev. 1000 m; CEMT • 2 specs; same data as for preceding; elev. 1200 m; CEMT • 1 spec.; same data as for preceding; elev. 1300 m; CEMT • 8 specs; same data as for preceding; 5 ° 42 ′ 33 ″ S, 62 ° 23 ′ 56 ″ W; elev. 1050 m CEMT • 1 spec.; same data as for preceding; 5 ° 58 ′ 52 ″ S, 62 ° 27 ′ 38 ″ W; elev. 1200 m; CEMT • 2 specs; same data as for preceding; 5 ° 58 ′ 52 ″ S, 62 ° 29 ′ 24 ″ W; elev. 1400 m; 2 CEMT • 4 specs; same data as for preceding; 6 ° 1 ′ 5 ″ S, 62 ° 29 ′ 24 ″ W; CEMT • 1 spec.; same data as for preceding; 5 ° 58 ′ 52 ″ S, 62 ° 27 ′ 38 ″ W; elev. 1200 m; M. A. P. A. Silveira leg.; CEMT. – Distrito Federal • 1 spec.; Gama-Cabeça-de-Veado; 15 ° 56 ′ 49.60 ″ S, 47 ° 56 ′ 58.50 ″ W; 15 Dec. 2018; W. L. Cunha; Cerrado s. str.; feces; CEMT. – Mato Grosso • 4 specs; Alta Floresta, P. E. Cristalino; 9 ° 32 ′ 45 ″ S, 55 ° 54 ′ 56 ″ W; elev. 248 m; 15 – 17 Nov. 2019; R. A. Azevedo leg.; human feces; CEMT • 3 specs; same data as for preceding; 9 ° 40 ′ 59 ″ S, 55 ° 54 ′ 57 ″ W; elev. 260 m; CEMT • 19 specs; same data as for preceding; 9 ° 35 ′ 50 ″ S, 55 ° 53 ′ 40 ″ W; elev. 287 m; CEMT • 1 spec.; same data as for preceding; 7 ° 24 ′ 60 ″ S, 73 ° 37 ′ 56 ″ W; elev. 254 m; 6 – 8 Dec. 2019; CEMT • 1 spec.; Brasnorte, Faz Hermínia; 11 ° 56 ′ 59 ″ S, 58 ° 14 ′ 47 ″ W; elev. 304 m; 29 Nov. – 1 Dec. 2017; U. F. Santos leg.; borda; human dung; CEMT • 3 specs; same data as for preceding; 11 ° 57 ′ 2 ″ S, 58 ° 14 ′ 49 ″ W; elev. 309 m; CEMT • 1 ♂, 2 specs; same data as for preceding; elev. 311 m; 29 – 30 Nov. 2017; A. F Machado leg.; forest; pitfall; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 9 ° 50 ′ 23 ″ S, 58 ° 15 ′ 13 ″ W; 28 – 30 Oct. 2021; J. Mariano et al. leg.; matinha; human feces; CEMT • 4 specs; same data as for preceding; 9 ° 50 ′ 27 ″ S, 58 ° 15 ′ 8 ″ W; CEMT • 2 specs; same data as for preceding; 9 ° 51 ′ 39 ″ S, 58 ° 13 ′ 13 ″ W; 2 – 4 Nov. 2021; J. Mariano et al. leg.; human feces; CEMT • 1 spec.; same data as for preceding; 9 ° 50 ′ 47 ″ S, 58 ° 14 ′ 9 ″ W; Talhão 46 b; CEMT • 1 spec.; same data as for preceding; 9 ° 51 ′ 45 ″ S, 58 ° 15 ′ 50 ″ W; 5 – 7 Nov. 2021; Talhão 14 b; CEMT • 1 spec.; Novo Mundo, P. E. Cristalino; 9 ° 28 ′ 23 ″ S, 55 ° 49 ′ 19 ″ W; Nov. 2012; V. Magalhães leg.; pitfall; CEMT • 1 spec.; São José do Rio Claro; 13 ° 53 ′ 10 ″ S, 56 ° 40 ′ 42 ″ W; 29 Jun. 2018; L. G Nunes and E. Carvalho leg.; human feces; CEMT • 1 spec.; same data as for preceding; 30 Jun. 2018; CEMT. – Minas Gerais • 1 spec.; Brumadinho; 20 ° 08 ′ 7 ″ S, 44 ° 08 ′ 7 ″ W; May 2021; P. G. da Silva leg.; human dung; CEMT • 1 ♂, 5 specs; same data as for preceding; 20 ° 8 ′ 9 ″ S, 44 ° 7 ′ 52 ″ W; Nov. 2021; P. G. da Silva leg.; human dung; CEMT • 1 ♀, 1 spec.; same data as for preceding; 20 ° 8 ′ 7 ″ S, 44 ° 8 ′ 45 ″ W; CEMT • 1 spec.; same data as for preceding; 20 ° 9 ′ 24 ″ S, 44 ° 9 ′ 10 ″ W; CEMT. – Pará • 1 spec.; Belém, IPEAN; May 1985; N. Degallier leg.; CEMT • 1 spec.; Belterra; 3 ° 15 ′ 25 ″ S, 54 ° 53 ′ 18 ″ W; elev. 113 m; 15 Jul. 2016; F. França leg.; sec forest; human and pig dung; CEMT • 1 spec.; same data as for preceding; FLONA-Tapajós; 3 ° 01 ′ 43 ″ S, 55 ° 00 ′ 40 ″ W; 21 Mar. 2019; primary forest; CEMT • 2 specs; same data as for preceding; 3 ° 19 ′ 12 ″ S, 54 ° 57 ′ 37 ″ W; 22 Mar. 2019; CEMT • 1 spec.; Bragança; 1 ° 10 ′ 46 ″ S, 46 ° 40 ′ 54 ″ W; 5 Mar. 2019; CEMT • 2 specs; Santarém, Flona-Tapajós; 3 ° 19 ′ 12 ″ S, 54 ° 57 ′ 37 ″ W; 22 Mar. 2019; F. França leg.; human and pig dung; CEMT • 1 spec.; Almeirim, Jari; 2 Feb. 2011; F. M. França leg.; pitfall feces; CEMT • 2 specs; same data as for preceding; 5 Feb. 2011; CEMT • 1 ♀, 12 specs; same data as for preceding; 16 Feb. 2011; CEMT • 3 specs; same data as for preceding; 19 Feb. 2011; CEMT • 4 specs; same data as for preceding; 23 Feb. 2011; CEMT • 10 specs; same data as for preceding; 26 Feb. 2011; CEMT • 1 spec.; same data as for preceding; 2 Mar. 2011; CEMT • 2 specs; same data as for preceding; 12 Mar. 2011; CEMT • 1 spec.; same data as for preceding; 16 Mar. 2011; CEMT • 1 spec.; same data as for preceding; 16 Mar. 2012; CEMT • 1 spec.; same data as for preceding; 19 Mar. 2011; CEMT • 2 specs; Marabá; 5 ° 47 ′ 04 ″ S, 48 ° 54 ′ 55 ″ W; 21 Feb. 2019; F. França leg.; logging forest; human and pig dung; CEMT • 1 spec.; same data as for preceding; 5 ° 38 ′ 01 ″ S, 49 ° 12 ′ 39 ″ W; 22 Feb. 2019; secondary forest; CEMT • 1 spec.; same data as for preceding; 5 ° 39 ′ 35 ″ S, 49 ° 10 ′ 47 ″ W; CEMT. – Rondônia • 6 specs; Parq Estadual Guajará-Mirim; 10 ° 18 ′ S, 64 ° 32 ′ W; 25 Nov. 2016; M. A. P. A. Silveira leg.; human dung; CEMT • 1 spec.; same data as for preceding; 10 ° 09 ′ S, 64 ° 33 ′ W; 8 Mar. 2017; CEMT • 1 spec.; same data as for preceding; 23 Sep. 2017; CEMT • 6 specs; Pimenta Bueno; 11 ° 43 ′ 05 ″ S, 61 ° 28 ′ 21 ″ W; elev. 249 m; 9 – 11 Dec. 2015; D. C. Castro leg.; human dung; CEMT. – Roraima • 3 specs; Cantá; Sep. 1996; Vaz-de-Mello and Ribeiro leg.; CEMT • 4 specs; Ilha de Maracá; Sep. 1996; Ribeiro leg.; CEMT. COLOMBIA – Amazonas • 1 spec.; Letícia, BTF; Oct. 2005; L. Escobar leg.; pitfall; CEMT. FRENCH GUIANA • 2 specs; Cayenne, Montagne des Chevaux (MCV); 4 ° 44 ′ 56 ″ N, 52 ° 26 ′ 28 ″ W; 13 Apr. 2013; SEAG leg.; light (PSA); CEMT • 1 spec.; Mont Proche de Mont (BAR); 4 ° 18 ′ 28 ″ N, 53 ° 17 ′ 10 ″ W; 2 Nov. 2013; SEAG leg.; FIT (V); CEMT • 1 spec.; Nouragues; Apr. 2001; F. Feer leg.; forêt primaire; CEMT • 1 spec.; Sain Eugéne, Barrage de Petit Saut; 4 ° 51 ′ N, 53 ° 4 ′ W; elev. 90 m; May 1999; F. Feer leg.; CEMT • 1 spec.; Sain Laurent du Maroni, Belvedère Saül; 3 ° 37 ′ 22 ″ N, 53 ° 12 ′ 57 ″ W; elev. 326 m; 14 Mar. 2011; SEAG leg.; FIT; CEMT • 1 spec.; same data as for preceding; 30 Mar. 2011; FIT-V 6; CEMT • 1 spec.; same data as for preceding; 20 May 2011; FIT (V) leg.; CEMT • 1 spec.; same data as for preceding; 27 May 2011; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBBFFD1FDD6F90EFCF7FA45.taxon	description	Sexual dimorphism Males with head and pronotum less densely punctate, protibial apical spine outward and downward bent.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFBBFFD1FDD6F90EFCF7FA45.taxon	discussion	Collecting information Blackburneus laxepunctatus was found at altitudes ranging from 90 to 1400 metres. Specimens were captured with pitfall traps baited with human or cow dung, with flight intercept or were attracted by light traps in regions of primary and secondary forests.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF86FFD0FDEEF980FB07FABC.taxon	description	Figs 2 E – G, 34 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF86FFD0FDEEF980FB07FABC.taxon	diagnosis	Diagnosis Length 2.5 – 3.0 mm. Body shagreened and testaceous. Head with moderated punctures, these progressively larger near frons; clypeus emarginate, medially weakly gibbous; frontoclypeal suture present, finely impressed. Pronotum dense fine and coarse punctures. Elytra with interstriae convex, with a singe row of punctures.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF86FFD0FDEEF980FB07FABC.taxon	materials_examined	Material examined BOLIVIA • 4 specs; 2 Feb. 1960; R. B. Cumming leg.; CEMT. – Santa Cruz • 2 specs; Sta 60 mi. N of Saavedra Exp.; CEMT. BRAZIL – Distrito Federal • 1 spec.; Planaltina, Embrapa Cerrados; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 18 Nov. 2005; O. Oliveira leg.; agricultural area; light; CEMT • 1 spec.; same data as for preceding; 2 Dec. 2005; CEMT • 3 specs; same data as for preceding; 5 Jan. 2006; CEMT • 1 spec.; same data as for preceding; 15 ° 36 ′ 16 ″ S, 47 ° 44 ′ 16 ″ W; 24 Feb. 2006; native cerrado; CEMT • 1 spec.; same data as for preceding; 15 ° 36 ′ 20 ″ S, 47 ° 42 ′ 16 ″ W; 3 Nov. 2006; CEMT • 2 specs; same data as for preceding; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 24 Mar. 2006; agricultural area; CEMT. – Maranhão • 3 specs; Carolina, Fazenda Cincorá; 17 – 22 Oct. 2009; F. Limeira-de-Oliveira et al. leg.; light; CEMT. – Mato Grosso do Sul • 1 ♂; Selvíra, Santa Ofélia farm; 1 May 2004; C. A. H. Flechtmann leg.; Brachiaria decumbens pasture; bovine dropping; CEMT • 3 ♂♂; same data as for preceding; 12 Jun. 2004; CEMT • 1 ♀; same data as for preceding; 26 Feb. 2005; CEMT • 1 ♀; same data as for preceding; UNESP farm; 16 Jan. 2005; Brachiaria brizantha pasture; CEMT • 1 ♂; same data as for preceding; 20 ° 22 ′ 41 ″ S, 51 ° 25 ′ 10 ″ W; 29 Oct. 2005; F. Oikawa leg.; Brachiaria decumben s pasture; black light; CEMT • 1 ♀; same data as for preceding; 12 Nov. 2009; CEMT • 1 ♀; same data as for preceding; 5 May 2010; CEMT • 1 ♀; same data as for preceding; 20 ° 53 ′ 41 ″ S, 51 ° 24 ′ 39 ″ W; 6 Feb. 2007; F. W. Mesquita leg.; CEMT • 1 ♂; same data as for preceding; 20 ° 22 ′ 53 ″ S, 51 ° 24 ′ 39 ″ W; 7 Nov. 2006; CEMT • 1 ♀; same data as for preceding; 30 Jan. 2007; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF86FFD0FDEEF980FB07FABC.taxon	description	Sexual dimorphism Male protibial apical spine downwardly and outwardly bent; males with distal teeth of protibia with proximal margin dilated and angled.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF86FFD0FDEEF980FB07FABC.taxon	discussion	Collecting information Specimens can be found in pastures of Urochloa brizantha and Urochloa decumbens and in areas of native Cerrado. Specimens are attracted to cattle dung, light and can be captured by flight intercept. The species can be found at altitudes up to 1600 m (Dellacasa et al. 2011; Mesquita et al. 2017).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF87FFD7FE4CFAABFD8BFE64.taxon	description	Figs 2 D, 34 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF87FFD7FE4CFAABFD8BFE64.taxon	diagnosis	Diagnosis Length 3.5 – 4.0 mm. Body wide, light brown. Head with clypeus sinuate with punctures of two sizes, fine and coarse; coarse punctures closer near to posterior half of clypeus, near the frontoclypeal suture; frontoclypeal suture fine, almost imperceptible; frons coarsely punctate. Pronotum with punctures of two sizes fine and coarse; coarse punctures distant on disc, closer laterally. Elytra with interstriae flat; minutely punctured. First tarsomere of metatarsus longer than superior apical spur.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF87FFD7FE4CFAABFD8BFE64.taxon	materials_examined	Material examined BRAZIL – Amazonas • 3 specs; PDBFF [Biological Dynamics of Forest Fragments Project], Km 41; 02 ° 27 ′ S, 59 ° 46 ′ W; 12 Feb. 2005; M. Radtke leg.; dung pitfall; INPA • 1 spec.; same data as for preceding, dimona; 2 ° 20 ′ S, 60 ° 07 ′ W; 17 Feb. 2005; CEMT • 1 spec.; Reserva Ducke, 26 km NE of Manaus; 14 Mar. 1978; B. C. Ratcliffe leg.; CEMT • 1 spec.; same data as for preceding; 30 May 1978; CEMT • 1 spec.; same data as for preceding; 20 Jun. 1978; CEMT • 2 specs; same data as for preceding; 22 Nov. 1977; CEMT • 1 spec.; same data as for preceding; BR 010, Km 26; 31 Jan. 1978; B. C. Ratcliffe leg.; CEMT • 2 specs; Manaus, ZF- 03, Km 41; 1996 – 1997; Andresen and Ellen leg.; in Alouatta seniculus feces; CEMT • 5 specs; same data as for preceding; INPA.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF87FFD7FE4CFAABFD8BFE64.taxon	description	Sexual dimorphism Males with head and pronotum less densely punctured than those of female. Males with metaventrite plate longitudinally grooved medially.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF80FFD7FE6DFE63FD3EFA06.taxon	description	Figs 3 A, 35 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF80FFD7FE6DFE63FD3EFA06.taxon	diagnosis	Diagnosis Length 2.0 – 2.5 mm. Body oval shagreened, testaceous. Head with clypeus with punctures of similar sizes and equidistant; clypeus discretely emarginate, trapezoidal, weakly gibbous; frontoclypeal suture present although weak. Pronotum with punctures of two sizes; fine punctures spread throughout pronotal surface, equidistant on disc, closer near lateral margins; coarse punctures absent on pronotal disc, closer near lateral margins. Elytra shagreened; interstriae convex, with two rows of fine punctures.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF80FFD7FE6DFE63FD3EFA06.taxon	materials_examined	Material examined BRAZIL – Distrito Federal • 1 spec.; Planaltina, Embrapa Cerrados; 15 ° 36 ′ 16 ″ S, 47 ° 44 ′ 16 ″ W; 3 Nov. 2006; Oliveira leg.; light; CEMT. – Mato Grosso do Sul • 1 ♂; Selvíria, Santa Ofélia farm; 1 May 2004; C. A. H. Flechtmann leg.; Brachiaria brizantha pasture; bovine dropping; CEMT • 1 ♀; same data as for preceding; 22 May 2004; CEMT • 1 ♀; same data as for preceding; 12 Feb. 2005; CEMT • 1 ♂; same data as for preceding; UNESP farm; 12 Nov. 2009; Urochloa decubens; black light; CEMT • 1 ♂; same data as for preceding; 20 ° 22 ′ 41 ″ S, 51 ° 25 ′ 10 ″ W; 29 Oct. 2005; F. W. Mesquita leg.; Brachiaria decumbens pasture; black light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF80FFD7FE6DFE63FD3EFA06.taxon	description	Sexual dimorphism Males with head and pronotum less densely and less strongly punctured than those of females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF80FFD7FE6DFE63FD3EFA06.taxon	discussion	Collecting information Specimens were collected with black light traps and in cattle dung in pastures of Urochloa brizantha and Urochloa decumbens (Mesquita et al. 2017).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF80FFD6FE71FA41FA82FE08.taxon	diagnosis	Diagnosis Length 3.5 – 4.5 mm. Body oblong, convex, glabrous, glossy, dark testaceous or light reddish testaceous. Head strongly, densely punctate; clypeus emarginate, microreticulate, lateral angles obtusely rounded; frontoclypeal suture developed, evident. Pronotum wide, everywhere punctate, punctures closer near lateral margins; lateral margins and posterior angles with marginal line; posterior angles sinuate. Elytra without humeral denticle; interstriae glossy, lateral margins microreticulate, strongly microreticulate near posterior margins, dull. Protibia dorsally punctate. Metatibia with weakly developed set of transverse carinae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF80FFD6FE71FA41FA82FE08.taxon	discussion	Remarks Gonaphodioides is a genus comprised of nine species found in Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Mexico, Panama, Peru and Venezuela (Dellacasa et al. 2012). For more information about the diversity of this genus and keys for all species see Dellacasa et al (2012).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF81FFD6FDF7FE7FFB1BFA71.taxon	description	Figs 4 A, C, 35 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF81FFD6FDF7FE7FFB1BFA71.taxon	diagnosis	Diagnosis Length 4.0 – 4.5 mm. Body dark testaceous. Pronotum with punctures moderate, somewhat equally distant; punctures laterally closer. Elytral interstriae convex. Superior apical spur of metatibiae 1 / 4 shorter than first tarsomere of metatarsus.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF81FFD6FDF7FE7FFB1BFA71.taxon	materials_examined	Material examined BRAZIL – Acre • 1 spec.; Assis Brasil; 10 ° 35 ′ 44 ″ S, 69 ° 37 ′ 12 ″ W; 23 Sep. 2016; Ortiz and Borges leg.; CDC light trap; CEMT • 1 spec.; same data as for preceding; 10 ° 35 ′ 44 ″ S, 69 ° 37 ′ 11 ″ W; 18 Nov. 2016; wild environment; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF81FFD6FDF7FE7FFB1BFA71.taxon	description	Sexual dimorphism Female with apical spur of protibia slender, elongate and outward curved.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF81FFD6FDF7FE7FFB1BFA71.taxon	discussion	Collecting information This species can be found at altitudes up to 1300 metres. Specimens were collected with CDC light traps, flight intercept traps and are attracted to pig, human and horse dung (Dellacasa et al. 2012).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF81FFD4FDC6FA74FEB8FC76.taxon	description	Figs 4 B, 35 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF81FFD4FDC6FA74FEB8FC76.taxon	diagnosis	Diagnosis Length 3.5 – 4.0 mm. Body light reddish testaceous. Pronotum scarcely and finely punctate on disc, lateral punctures closer and larger. Elytra with interstriae on disc flat. Superior apical spur of metatibia 1 / 3 shorter than first tarsomere of metatarsus.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF81FFD4FDC6FA74FEB8FC76.taxon	materials_examined	Material examined BRAZIL – Amazonas • 1 spec.; 54 km of Leste de Manaus; 26 Apr. 1979; Arias et al. leg.; CEMT. – Distrito Federal • 1 spec.; Brasília, Fazenda Água Limpa; 15 ° 56 ′ 40 ″ S, 47 ° 55 ′ 37 ″ W; 10 Dec. 2017; Y. Ferreira leg.; pasture; pig dung; CEMT • 2 specs; Planaltina, Embrapa Cerrados; 15 ° 36 ′ 20 ″ S, 47 ° 42 ′ 16 ″ W; 3 Nov. 2006; C. Oliveira leg.; native cerrado; light; CEMT. – Mato Grosso • 1 spec.; Chapada dos Guimarães, R. Mutuca; 15 ° 18 ′ 21 ″ S, 55 ° 58 ′ 6 ″ W; 10 – 20 Nov. 2016; A. Frolov leg.; FIT; CEMT • 2 specs; Porto Estrela, E. E. Serra das Araras, Trilha Boca do José Cerrado; Nov. 2017; T. F. Conceição leg.; pitfall; CEMT. – Rio de Janeiro • 1 spec.; Nova Friburgo; Nov. 2002; E. J. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Rio Bonito de Lumiar; 22 ° 24 ′ 32 ″ S, 42 ° 20 ′ 10 ″ W; elev. 658 m; 6 Mar. 2009; J. Nissimian et al. leg.; light; CEMT. – Rondônia • 4 specs; 62 km SW of Ariquemes, Fazenda Rancho Grande; 3 – 15 Dec. 1996; J. E. Eger leg.; black light; CEMT. – São Paulo • 2 specs; São Paulo, Descalvado, Fazenda Itaúna; 3 Nov. 2005; N. W. Perioto leg.; forest Cerrado; pitfall; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF81FFD4FDC6FA74FEB8FC76.taxon	description	Sexual dimorphism Female with apical spur of protibia slender, curved outward.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF81FFD4FDC6FA74FEB8FC76.taxon	discussion	Collecting information Specimens were collected with black light traps or with flight intercept traps and appear to be attracted to pig dung.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF83FFD4FDE8FC73FA83F856.taxon	diagnosis	Diagnosis Length 3.2 – 5.1 mm. Body oblong, subparallel-sided, glossy and dorsally glabrous, yellow with dark spots on the head, disk of pronotum and elytra. Head with frontoclypeal suture tuberculate. Pronotum without posterior marginal line. Dorsal surface of protibia not punctate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF83FFD4FDE8FC73FA83F856.taxon	discussion	Remarks Labarrus is a subcosmopolitan genus comprised of 19 species, three of which are found in the New World (Dellacasa et al. 2001 a, 2002; Schoolmeesters 2023). Woodruff (1973) suggested that Labarrus was introduced to the American continent by ships transporting cattle and slaves from the Old World. Recently, Bordat (2016) showed that the New World species of Labarrus are widely distributed on the African continent along with other species not present in the Americas. These facts lead us to the same conclusion. But further studies need to be conducted to verify this assumption. For another key of the species present in the New World see Dellacasa et al. (2002). A key for Afrotropical species can be found in Bordat (2016).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8DFFDAFD85FE95FE74FAAB.taxon	description	Figs 6 E – G, 36 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8DFFDAFD85FE95FE74FAAB.taxon	diagnosis	Diagnosis Length 3.0 – 4.5 mm. Head with clypeus clearly emarginate, anteriorly rugosely punctured; frontoclypeal suture tuberculate medially; suture not visible at tubercle. Pronotum strongly convex, with punctures of two sizes; larger punctures scarce. Inferior apical spur of mesotibia obliquely truncate apically. Meso- and metatibiae dark, contrasting with femur.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8DFFDAFD85FE95FE74FAAB.taxon	materials_examined	Material examined BRAZIL – Pernambuco • 2 ♂♂; Fernando de Noronha, Sancho; 20 – 28 Feb. 2020; Grossi and Rafael leg.; light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8DFFDAFD85FE95FE74FAAB.taxon	description	Sexual dimorphism Males with frontoclypeal tubercle more developed, acuminate. Females with frontoclypeal tubercle weakly developed.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8DFFDAFD85FE95FE74FAAB.taxon	discussion	Collecting information Specimens were collected at light. Remarks After the description of Labarrus pseudolividus, the records of L. lividus on the American continent have been considered dubious since many of the specimens previously determined as the former were in fact misidentified L. pseudolividus (Dellacasa 2002; Gordon & Skelley 2007). Today, only a few reliable records of L. lividus are known from Central and North America (Dellacasa et al. 2002; Gordon & Skelley 2007; Bordat 2016). Here, to our knowledge, we present the first reliable record of Labarrus lividus in Brazil.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8DFFDEFDC2FAA2FBA8F87C.taxon	description	Figs 6 A – C, 36 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8DFFDEFDC2FAA2FBA8F87C.taxon	diagnosis	Diagnosis Length 3.2 – 3.9 mm. Clypeus clearly emarginate; with coarse punctures extending to frons; frontoclypeal suture medially tuberculate; suture distinct, excavating posterior half of median tubercle. Tarsomeres of metatarsus shortened, sub-triangularly widened. Meso- and metatibiae pale, not contrasting with femur.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8DFFDEFDC2FAA2FBA8F87C.taxon	materials_examined	Material examined ARGENTINA – Desaguadero • 13 specs; 5 Feb. 2017; G. Arriágada leg.; light; CEMT. – Córdoba • 1 spec.; 10 km NW of Alpa Corral; 8 – 9 Feb. 2009; F. C. Ocampo leg.; CEMT. – Corrientes • 6 specs; Barrio Víctor Colas; 27 ° 28 ′ 50 ″ S, 58 ° 47 ′ 18 ″ W; elev. 64 m; 21 Oct. 2012; M. I. Polesel et al. leg.; light; CEMT. – Entre Ríos • 2 specs; Liebig, Pueblo; Feb. 1994; A. Ugarte-Penã leg.; CEMT. – Mendoza • 1 spec.; Lavalle, Res. Bio. Telteca; 9 Dec. 2002; Arriágada and Diéguez leg.; light; CEMT. – San Luis • 1 spec.; Belgrano; 14 – 18 Feb. 2018; J. Jensen leg.; light; CEMT • 10 specs; same data as for preceding; Fundo el Molle; 33 ° 02 ′ 23 ″ S, 66 ° 30 ′ 48 ″ W; elev. 622 m; 7 Feb. 2017; G. Arriágada leg.; CEMT. BOLIVIA – Santa Cruz • 1 spec.; Chiquitos, 1.6 km E of Santiago de Chiquitos; 18 ° 12 ′ 04 ″ S, 59 ° 21 ′ 00 ″ W; elev. 622 m; Nov. 2008; W. D. Edmonds and T. Vidaurre leg.; light; CEMT. BRAZIL – Amazonas • 1 spec.; Itacoatiara, Fazenda Aruana, Am 010, Km 215; 16 – 19 Dec. 1990; C. S. Morra et al. leg.; luz mista de mercúrio [mistmixed mercury light]; INPA. – Bahia • 4 specs; Camacan, Rio Panelão; 15 ° 24 ′ 28 ″ S, 39 ° 32 ′ 4 ″ W; elev. 176 m; 12 Aug. 2016; A. P. M. Santos et al. leg.; light; CEMT. – Ceará • 1 spec.; Ubajara, P. N. Ubajara, Trilha Araticum, Rio Das Minas; elev. 480 m; 14 Feb. 2013; D. M. Takiya and A. P. M. Santos leg.; Pennsylvania trap; CEMT • 13 specs; same data as for preceding; Portão Neblina; 3 ° 50 ′ 18 ″ S, 40 ° 53 ′ 54 ″ W; elev. 849 m; 14 Feb. 2013; light; CEMT • 1 spec.; same data as for preceding; Cachoeira Cafundó; 3 ° 50 ′ 12 ″ S, 40 ° 54 ′ 35 ″ W; elev. 783 m; 15 Feb. 2013; Pennsylvania trap; CEMT • 1 spec.; same data as for preceding; bridge across Rio Miranda; 3 ° 50 ′ 07 ″ S, 40 ° 54 ′ 48 ″ W; elev. 792 m; CEMT • 1 spec.; same data as for preceding; Trilha Samambaia; 3 ° 50 ′ 20 ″ S, 40 ° 53 ′ 55 ″ W; 850 m; 17 Feb. 2013; D. M. Takiya leg.; CEMT. – Distrito Federal • 2 specs; Brasília; elev. 1000 m; Feb. 2001; N. Degallier leg.; light; CEMT • 1 spec.; same data as for preceding; Mar. 2001; CEMT • 1 spec.; same data as for preceding; elev. 1100 m; Oct. 2000; CEMT • 2 specs; same data as for preceding; elev. 1100 m; Jan. 2001; CEMT • 2 specs; same data as for preceding; Feb. 2001; Felipe and Rodrigues leg.; light; CEMT • 1 spec.; same data as for preceding; Fazenda Água Limpa (University of Brasilia); Nov. 2010; C. Suinaga leg.; CEMT • 1 spec.; same data as for preceding; 15 ° 56 ′ 40 ″ S, 47 ° 55 ′ 38 ″ W; 10 Dec. 2017; Y. Ferreira leg.; pig dung; CEMT • 1 spec.; Pq Nacional de Brasília; 15 ° 42 ′ 08 ″ S, 47 ° 56 ′ 03 ″ W; 15 Dec. 2018; W. L. Cunha leg.; cerrado s. str.; CEMT • 1 spec.; Planaltina, Embrapa Cerrados; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 18 Nov. 2005; C. Oliveira leg.; área agrícola [agricultural area]; light; CEMT • 1 spec.; same data as for preceding; 2 Dec. 2005; CEMT • 2 specs; same data as for preceding; 17 Dec. 2005; CEMT • 1 spec.; same data as for preceding; 19 Jan. 2006; CEMT • 1 spec.; same data as for preceding; 24 Mar. 2006; CEMT • 6 specs; same data as for preceding; 15 ° 35 ′ 52 ″ S, 47 ° 43 ′ 10 ″ W; 4 Dec. 2017; Y. Ferreira leg.; soy plantation; human feces; CEMT. – Goiás • 3 specs; Anápolis; 16 ° 16 ′ 03 ″ S, 48 ° 50 ′ 37 ″ W; 2 Jan. 2016; E. C. C. Santana leg.; manually; CEMT • 1 spec.; Rio Verde; Oct. 1995; J. Carlos leg.; CEMT. – Maranhão • 11 specs; Carolina, Fazenda Cincorá; 17 – 22 Oct. 2009; F. Lima-de-Oliveira et al. leg.; light; CEMT. – Mato Grosso • 1 spec.; Água Boa, Rancho Araguaia; 13 ° 56 ′ 40 ″ S, 52 ° 6 ′ 10 ″ W; 24 Dec. 2017; Mota and Schutz leg.; CEMT • 5 specs; Alta Floresta, CEPLAC; 4 Sep. 2009; V. Gonçalves leg.; Tapirus feces; CEMT • 1 spec.; Brasnorte, Fazenda Hermínia; 11 ° 57 ′ 3 ″ S, 58 ° 14 ′ 43 ″ W; elev. 300 m; 29 Nov. – 1 Dec. 2017; U. F. Santos leg.; soy plantation; human feces; CEMT • 2 specs; Chapada dos Guimarães; 11 Dec. 2000; A. F. Ramos leg.; light; CEMT • 1 spec.; same data as for preceding; 25 Mar. 2001; CEMT • 1 spec.; same data as for preceding; Santuário dos Elefantes; 2 Nov. 2019; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; 15 ° 18 ′ 54 ″ S, 55 ° 24 ′ 03 ″ W; 4 Nov. 2019; H. Sehn et al. leg.; elephant feces; CEMT • 5 specs; same data as for preceding; 15 ° 18 ′ 51 ″ S, 55 ° 24 ′ 10 ″ W; CEMT • 5 specs; same data as for preceding; 5 ° 18 ′ 55 ″ S, 55 ° 24 ′ 07 ″ W; horse feces; CEMT • 1 spec.; same data as for preceding; Tapirus feces; CEMT • 2 specs; same data as for preceding; cow feces; CEMT • 1 spec.; same data as for preceding; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 14 ″ W; CEMT • 1 spec.; same data as for preceding; 15 ° 18 ′ 50 ″ S, 55 ° 24 ′ 14 ″ W; CEMT • 2 specs; same data as for preceding; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 05 ″ W; 22 Jan. 2020; CEMT • 1 spec.; same data as for preceding; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 10 ″ W; Tapirus feces; CEMT • 1 spec.; same data as for preceding; elephant feces; CEMT • 1 spec.; same data as for preceding; 5 ° 18 ′ 55 ″ S, 55 ° 24 ′ 14 ″ W; human feces; CEMT • 2 specs; same data as for preceding; 23 Jan. 2020; CEMT • 1 spec.; Cuiabá, Campus Universidade Federal de Mato Grosso; Oct. 2008; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Cotriguaçu, Assentamento INCRA; 1 Nov. 2017; R. Stofel leg.; CEMT • 1 spec.; same data as for preceding; Fazenda São Nicolau; 14 Feb. 2007; O. Peres-Filho leg.; light; CEMT • 1 spec.; same data as for preceding; 1 Oct. 2007; CEMT • 1 spec.; same data as for preceding; 16 Oct. 2007; CEMT • 3 specs; same data as for preceding; 31 Oct. 2007; CEMT • 1 spec.; same data as for preceding; Nov. 2007; CEMT • 1 spec.; same data as for preceding; 31 Nov. 2007; CEMT • 1 spec.; same data as for preceding; 14 Dec. 2007; CEMT • 4 specs; same data as for preceding; 15 Dec. 2007; CEMT • 7 specs; same data as for preceding; 16 Jan. 2008; CEMT • 3 specs; same data as for preceding; 9 ° 50 ′ 19 ″ S, 58 ° 15 ′ 03 ″ W; 25 Oct. 2014; A. Asenjo leg.; CEMT • 1 spec.; same data as for preceding 9 ° 51 ′ 20 ″ S, 58 ° 14 ′ 55 ″ W; Nov. 2014; R. V. Nunes and A. Asenjo leg.; CEMT • 1 spec.; same data as for preceding; Fazenda São Nicolau; 23 Oct. 2014; W. Chamorro leg.; light; CEMT • 2 specs; same data as for preceding; Oct. 2014; S. Stofel leg.; CEMT • 3 specs; same data as for preceding; 9 ° 51 ′ 52 ″ S, 58 ° 13 ′ 45 ″ W; 2 Oct. 2017; Vaz-de-Mello et al. leg.; CEMT • 6 specs; same data as for preceding; 9 ° 51 ′ 52 ″ S, 58 ° 13 ′ 54 ″ W; airport; light; CEMT • 1 spec.; Coxambu, Parque das Águas; 30 Dec. 1979; Bello leg.; CEMT • 1 spec.; Cuiabá, Recanto Siriena; 15 ° 35 ′ 32 ″ S, 56 ° 01 ′ 38 ″ W; 4 Nov. 2014; R. V. Nunes and B. F. Becker leg.; light; CEMT • 1 spec.; Chapada dos Guimarães, Salgadeira; 12 Oct. 1990; M. Serrano leg.; CEMT • 3 specs; Diamantino, Reserva Vale da Solidão; 14 ° 22 ′ S, 56 ° 07 ′ W; elev. 450 m; 26 Oct. 2013; E. Furtado leg.; CEMT • 1 spec.; same data as for preceding; 14 ° 22 ′ 30 ″ S, 56 ° 7 ′ 30 ″ W; elev. 500 m; 26 – 31 Oct. 2013; E. Furtado leg.; CEMT • 1 spec.; Jaciara; 16 Sep. 1993; W. Mathias leg.; light; CEMT • 1 spec.; Nobres; 30 Sep. 2018; R. Segalla and M. Cassiano leg.; CEMT • 2 specs; Nova Ubiratã, ESEC Ronuro; 25 Feb. 2017; L. G. O. A. Nunes et al. leg.; CEMT • 3 specs; same data as for preceding; 13 ° 06 ′ 43 ″ S, 54 ° 26 ′ 37 ″ W; 2 Mar. 2017; L. G. O. A. Nunes leg.; light; CEMT • 1 spec.; Poconé; 28 Jan. 1998; forest; light trap; CEMT • 1 spec.; same data as for preceding; 28 Mar. 1998; light; CEMT • 1 spec.; same data as for preceding; 29 Dec. 1997; light; CEMT • 2 specs; same data as for preceding; Pousada das Araras, Pantanal; 3 Dec. 1998; J. L. Moreto and T. Mestre leg.; CEMT • 1 spec.; Santo Antônio do Leverger, São Vicente da Serra; 15 ° 49 ′ 42 ″ S, 55 ° 25 ′ 11 ″ W; 16 – 18 Feb. 2011; A. S. O. Tissiani leg.; human feces; CEMT • 1 spec.; same data as for preceding; Fazenda Experimental da UFMT [Expermental farm UFMT]; Dec. 2020; F. Z. Vaz-de-Mello and M. E. Maldaner leg.; CEMT • 1 spec.; Sesc-Pantanal; 15 Nov. 2001; light; CEMT • 1 spec.; same data as for preceding; 15 Sep. 2004; light; CEMT • 1 spec.; same data as for preceding; 5 Jul. 2003; CEMT • 1 spec.; Tangará da Serra; 14 ° 38 ′ 33 ″ S, 57 ° 33 ′ 18 ″ W; 19 – 22 Feb. 2012; A. S. Tissiani leg.; CEMT. – Mato Grosso do Sul • 1 spec.; Corumbá, Fazenda São Bento, Prox. Porto Jofre; 14 Apr. 2011; R. V. Nunes leg.; light; CEMT • 1 spec.; Dourados; 27 May 2006; M. Miloca leg.; CEMT • 1 spec.; Guia Lopes da Laguna; 15 Feb. 2007; A. Abot leg.; light; CEMT • 1 spec.; Nova Andradina, Usina Santa Helena; 21 ° 59 ′ 32 ″ S, 53 ° 26 ′ 28 ″ W; elev. 360 m; 18 Oct. 2012; G. Coutinho leg.; CEMT • 2 specs; Inocência, Fazenda Moinho; 19 ° 13 ′ 12 ″ S, 52 ° 9 ′ 34 ″ W; elev. 502 m; 21 Oct. 2018; H. L. Rainho leg.; light; CEMT • 1 spec.; Santo Antônido do Leverger, São Vicente da Cerra; 15 ° 49 ′ 42 ″ S, 55 ° 25 ′ 11 ″ W; 16 – 18 Feb. 2011; S. O. Tissiani leg.; human feces; CEMT • 2 specs; Selvíria, UNESP farm; 12 Oct. 1993; C. A. H. Flechtmann leg.; pasture area; cattle dropping; CEMT • 1 spec.; same data as for preceding; 6 Jul. 1991; Brachiaria decumbens pasture; guzera bovine dropping; CEMT • 1 spec.; same data as for preceding; 10 Oct. 1991; CEMT • 1 spec.; same data as for preceding; 1 Dec. 1991; CEMT • 1 spec.; same data as for preceding; 20 Jan. 1999; black light; CEMT. – Minas Gerais • 1 spec.; Conceição do Mato Dentro, 23 Km; elev. 711 m; 16 Sep. 2012; Uceli and Borges leg.; light; CEMT • 2 specs; same data as for preceding; elev. 734 m; 22 Sep. 2012; CEMT • 1 spec.; same data as for preceding; elev. 727 m; 1 Dec. 2012; CEMT • 1 spec.; Cordisburgo, Fazenda Pontinha; Dec. 1993; F. Z. Vaz-de-Mello leg.; CEMT • 2 specs; same data as for preceding; Jan. 1996; Vaz-de-Mello leg.; CEMT • 2 specs; Frei Gaspar, Sítio Recanto Paraíso; 18 ° 02 ′ 06 ″ S, 41 ° 29 ′ 10 ″ W; 28 Dec. 2007; M. R. Santos leg.; light; CEMT • 3 specs; same data as for preceding; elev. 571 m; CEMT • 3 specs; same data as for preceding; 30 Dec. 2007; CEMT • 5 specs; same data as for preceding; 3 Jan. 2008; CEMT • 2 specs; Ipatinga; Sep. 1993; E. Grossi leg.; CEMT • 1 spec.; Lavras; 21 ° 18 ′ 52 ″ S, 44 ° 59 ′ 33 ″ W; 20 Jan. 2008; M. S. Rocha and D. L. H. Takahashi leg.; CEMT • 1 spec.; Lima Duarte, Parque Estadual Ibitipoca; 20 Feb. 1997; C. R. M. Abreu leg.; CEMT • 1 spec.; Uberlândia, Campus UFU; 10 Sep. 2005; J. Mendes leg.; CEMT • 1 spec.; Viçosa; 1994 – 1995; Louzada et al. leg.; pastures; CEMT. – Paraíba • 7 specs; São José dos Cordeitos, Fazenda Almas; 4 – 5 May 2003; Valderez and Rothéa leg.; light trap; CEMT. – Paraná • 1 spec.; Entre Rios do Oeste; 26 Dec. 2017; M. E. Maldaner leg.; light; CEMT • 1 spec.; Icaraíma, Estância Felipe; 23 ° 20 ′ 32 ″ S, 53 ° 39 ′ 09 ″ W; 2019; Felipe and Rodrigues leg.; CEMT • 5 specs; same data as for preceding; elev. 300 m; CEMT • 1 spec.; same data as for preceding; 23 ° 23 ′ S, 53 ° 37 ′ W; elev. 357 m; CEMT • 1 spec.; Rancho Alegre, Fazenda Congonhas; 22 ° 47 ′ 45 ″ S, 51 ° 00 ′ 12 ″ W; elev. 580 m; 27 Dec. 2012; P. M. Felix leg.; feces; CEMT. – Piauí • 1 spec.; Piripiri, PN Sete Cidades; 4 ° 05 ′ 57 ″ S, 41 ° 42 ′ 34 ″ W; elev. 200 m; 7 – 12 Feb. 2013; Vaz de Mello and Grossi leg.; light; CEMT. – Rio de Janeiro • 1 spec.; Cachoeiras de Macacu, Guapiaçu ER; 22 ° 27 ′ 05 ″ S, 42 ° 46 ′ 23 ″ W; elev. 33 m; 25 Oct. 2019; L. R. Martínez leg.; horse feces; CEMT • 3 specs; Miguel Pereira; Jan. 1998; J. Carlos leg.; CEMT • 4 specs; same data as for preceding; Mar. 1998; CEMT • 2 specs; Nova Friburgo; Nov. 2002; E. J. Grossi leg.; CEMT. – Rio Grande do Sul • 1 spec.; Aceguá, Fazenda Sentinela; 31 ° 27 ′ 30 ″ S, 54 ° 21 ′ 18 ″ W; 8 – 14 Nov. 2011; R. M. Moraes leg.; FIT; CEMT • 1 spec.; Eldorado do Sul, EEA-UFRGS; 11 Oct. 1996; Carvalho and Silva leg.; light; CEMT • 2 specs; same data as for preceding; Estação Experimental UFRGS; 30 Mar. – 5 Apr. 2012; R. M. Moraes and J. Podkowa leg.; pitfall; CEMT • 1 spec.; Estrada da Ferreira; 30 ° 00 ′ 26 ″ S, 52 ° 57 ′ 22 ″ W; Dec. 2020; V. Costa-Silva leg.; CEMT • 6 specs; Santa Cruz do Sul, Cerro Alegre; 29 ° 48 ′ 18 ″ S, 52 ° 19 ′ 55 ″ W; 17 Mar. 2012; R. Bohn leg.; open area; bovine feces; CEMT • 3 specs; same data as for preceding; 24 Mar. 2012; CEMT • 2 specs; same data as for preceding; 29 ° 48 ′ 17 ″ S, 52 ° 19 ′ 59 ″ W; CEMT • 4 specs; same data as for preceding; forest; CEMT • 2 specs; same data as for preceding; 29 ° 48 ′ 11 ″ S, 52 ° 19 ′ 51 ″ W; CEMT • 3 specs; same data as for preceding; annual crop; CEMT • 4 specs; same data as for preceding; 28 ° 48 ′ 05 ″ S, 52 ° 19 ′ 39 ″ W; grass field; CEMT. – Santa Catarina • 1 spec.; Santa Rosa de Lima; 28 ° 04 ′ S, 49 ° 09 ′ W; elev. 270 m; 17 Jan. 2018; C. M. Rover leg.; silvi pastoril com núcleos [pastoral silvi with nuclei]; pitfall; CEMT. – São Paulo • 2 specs; Anhembi ESALQ / USP; E. N. L. Ferreira leg.; CEMT • 1 spec.; Porto Ferreira; 21 ° 52 ′ 11 ″ S, 47 ° 28 ′ 05 ″ W; 19 Oct. 2019; G. Biffi leg.; CEMT. – Tocantins • 4 specs; Palmas, Faz Recanto Valadares; 20 Sep. 2005; M. S. Matos leg.; CEMT. ECUADOR – El Oro • 1 spec.; Piñas; elev. 1250 m; 2 Sep. 1997; C. Carpio leg.; human feces; CEMT. MEXICO – Hidalgo • 2 specs; Cuautepec, San Juan Hueyepan; 20 ° 12 ′ 55 ″ N, 98 ° 16 ′ 50 ″ W; elev. 2265 m; Aug. 2007; F. Z. Vaz-de-Mello leg.; CEMT. – Oaxaca • 1 spec.; Pinotepa Nacional, Ixcapa; 7 Apr. 2004; Rámirez-Ponce leg.; selva media subperenifolia [subevergreen forest]; light; CEMT. PARAGUAY – Alto Paraguay • 30 km W of Bahía Negra; Jun. 1988; Aguilar C. leg.; CEMT. – Alto Paraná • 1 spec.; 13 – 15 Nov. 1990; G. Arriágada leg.; CEMT. PERU – Piura • 1 spec.; Sullama; 16 Mar. 2017; D. Saavedra leg.; dry forest; light; CEMT. URUGUAY – Rivera • 1 spec.; Rivera, Bodegas Carrau farm; 30 ° 58 ′ 19 ″ S, 55 ° 26 ′ 40 ″ W; 3 – 12 Oct. 2011; R. M. Moraes leg.; FIT; CEMT • 1 spec.; same data as for preceding; COFUSA; 30 ° 57 ′ 36 ″ S, 55 ° 29 ′ 53 ″ W; 3 – 11 Oct. 2011; human and pig dung; CEMT. VENEZUELA – Yaracuy • Bolívar Aroa; 10 ° 00 ′ 00 ″ N, 68 ° 00 ′ 00 ″ W; elev. 463 m; 19 Jul. 2009; M. Asmüssen et al. leg.; human feces; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8DFFDEFDC2FAA2FBA8F87C.taxon	description	Sexual dimorphism Male with median tubercle more developed than that of female.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8DFFDEFDC2FAA2FBA8F87C.taxon	discussion	Collecting information Specimens were collected with Pennsylvania trap net, black light and flight intercept traps and are attracted to bovine, human, pig and Tapirus feces. Specimens were found in grasslands of Urochloa brizantha, soy crops, dry forests and in the wetlands regions of the pantanal. Labarrus parnaguaensis was found at altitudes ranging from 150 to 1250 metres (Mesquita et al. 2017).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8AFFC2FDFFFE95FEFDFEF5.taxon	description	Figs 6 B, D, 36 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8AFFC2FDFFFE95FEFDFEF5.taxon	diagnosis	Diagnosis Length 3.5 – 5.2 mm. Head with clypeus and frons punctate with small and close punctures; clypeus with weak emargination; frontoclypeal suture medially tuberculate; suture distinct although not excavating posterior half of median tubercle. Tarsomeres of metatarsus sub-cylindrical, elongate. Meso- and metatibiae pale, not contrasting with femur.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8AFFC2FDFFFE95FEFDFEF5.taxon	materials_examined	Material examined ARGENTINA – Córdoba • 1 spec.; Apla Corral, 10 km NW of Sitio; 8 – 9 Feb. 2009; F. C. Ocampo leg.; CEMT. – Corrientes • 1 ♂, 2 ♀ ♀, 73 specs; Corrientes, Barrio Víctor Colas; 27 ° 28 ′ 50 ″ S, 58 ° 47 ′ 18 ″ W; elev. 64 m; 21 Oct. 2012; M. I. Polesel et al. leg.; light; CEMT. – Entre Ríos • 4 specs; Liebig, Pueblo; Feb. 1994; A. U. Peña leg.; CEMT. – Formosa • 1 spec.; Laguna, Yema, RP 37 Km 2; 22 ° 24 ′ 42 ″ S, 61 ° 17 ′ 26 ″ W; 16 Dec. 2008; F. C. Ocampo leg.; CEMT. – Mendoza • 66 specs; Desaguadero; 5 Feb. 2017; G. Arriágada leg.; CEMT • 1 spec.; Lavalle, Res. Bio. Telteca; 9 Dec. 2002; Arriágada and Diéguez leg.; light trap; CEMT. – San Luis • 11 specs; Belgrano, Fundo El Molle; 33 ° 02 ′ 23 ″ S, 66 ° 30 ′ 48 ″ W; elev. 622 m; 7 Feb. 2017; G. Arriágada leg.; CEMT • 2 specs; same data as for preceding; 14 – 18 Feb. 2018; J. Jensen leg.; light; CEMT • 3 specs; Desaguadero; 20 Feb. 2000; G. Arriágada leg.; CEMT. AUSTRALIA – Western Australia • 2 specs; Moore River; 12 Oct. 1992; H. Z. Demor leg.; CEMT. BOLIVIA – Santa Cruz • 3 specs; Chiquitos, 1.6 km E of Santiago de Chiquitos; 18 ° 12 ′ 04 ″ S, 59 ° 21 ′ 00 ″ W; elev. 622 m; Nov. 2008; W. D. Edmonds and T. Vidaurre leg.; light; CEMT. BRAZIL – Alagoas • 1 spec.; APA Marituba Peixe; 19 – 25 Aug. 2019; A. B. Farias and LASDA leg.; CEMT. – Bahia • 1 spec.; Camacan, Rio Panelão; 15 ° 24 ′ 28 ″ S, 39 ° 32 ′ 04 ″ W; elev. 176 m; 12 Aug. 2016; A. P. M. Santos and D. M. Takiyama leg.; light; CEMT • 1 spec.; Santa Rita de Cássia; 14 Jan. 2020; M. S. Cardoso leg.; CEMT. – Distrito Federal • 1 spec.; Brasília; elev. 1100 m; Jan. 2001; N. Degallier leg.; light; CEMT • 1 spec.; same data as for preceding; Fazenda Água Limpa (UNB); Nov. 2010; C. Suinaga leg.; CEMT • 1 spec.; Brasilia, Planaltina, Embrapa cerrados; 15 ° 36 ′ 16 ″ S, 47 ° 44 ′ 16 ″ W; 7 Sep. 2005; C. Oliveira leg.; agricultural area; light; CEMT • 23 specs; same data as for preceding; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 17 Dec. 2005; CEMT • 11 specs; same data as for preceding; 18 Nov. 2005; CEMT • 54 specs; same data as for preceding; 2 Dec. 2005; CEMT • 17 specs; same data as for preceding; 5 Jan. 2006; CEMT • 11 specs; same data as for preceding; 19 Jan. 2006; CEMT • 10 specs; same data as for preceding; 6 Feb. 2006; CEMT • 26 specs; same data as for preceding; CEMT • 5 specs; same data as for preceding; 11 May 2006; CEMT • 2 specs; same data as for preceding; 26 May 2006; CEMT • 23 specs; same data as for preceding; 7 Apr. 2006; CEMT • 3 specs; same data as for preceding; CEMT • 26 specs; same data as for preceding; 20 Jun. 2006; CEMT • 4 specs; same data as for preceding; 2 Jul. 2006; CEMT. – Goiás • 1 spec.; Rio Verde; 12 Nov. 1983; A. Bello leg.; CEMT • 1 spec.; same data as for preceding; Nov. 1983; CEMT • 1 spec.; same data as for preceding; Apr. 1995; S. Carlos leg.; CEMT. – Maranhão • 1 spec.; Carolina, Fazenda Cincorá; 17 – 22 Oct. 2009; F. Lima-de-Oliveira et al. leg.; light; CEMT. – Mato Grosso • 2 specs; Res. Papagaio, BR 364 Km 555; 2 Oct. 1984; E. Binda; light trap; CEMT • 2 specs. same data as for preceding; INPA • 3 specs; Água Boa, Rancho Araguaia; 13 ° 56 ′ 40 ″ S, 52 ° 06 ′ 10 ″ W; 24 Dec. 2007; Mota and Schutz leg.; CEMT • 2 specs; Alta Floresta, CEPLAC; 4 Sep. 2009; V. Gonçalves leg.; Tapirus feces; CEMT • 3 specs; same data as for preceding; 4 Sep. 2010; human feces; CEMT • 1 spec.; Brasnorte, Fazenda Hermínia; 11 ° 57 ′ 03 ″ S, 58 ° 14 ′ 43 ″ W; elev. 300 m; 29 Nov. – 1 Dec. 2017; U. F. Santos leg.; soybean plantation; human dung; CEMT • 1 spec.; same data as for preceding; CEMT • 3 specs; Campo Novo do Parecis, Rio Membeca, Sete Lagoas, Fazenda Santa Teresinha; 13 ° 32 ′ 00 ″ S, 57 ° 47 ′ 57 ″ W; 14 Nov. 2017; D. G. Arruda leg.; FIT; CEMT • 1 spec.; Chapada dos Guimarães; 12 Dec. 2000; A. F. Ramos leg.; CEMT • 1 spec.; same data as for preceding; 15 ° 24 ′ 31 ″ S, 55 ° 45 ′ 53 ″ W; 2 Dec. 2015; A. Frolov and L. Akhemetova leg.; light; CEMT • 2 specs; Chapada dos Guimarães, Santuário dos Elefantes; 2 Nov. 2019; F. Z. Vaz-de-Mello leg.; CEMT • 2 specs; same data as for preceding; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 07 ″ W; 4 Nov. 2019; H. Sehn et al. leg.; horse feces; CEMT • 1 spec.; same data as for preceding; 15 ° 18 ′ 54 ″ S, 54 ° 24 ′ 03 ″ W; elephant feces; CEMT • 9 specs; same data as for preceding; 15 ° 18 ′ 51 ″ S, 55 ° 24 ′ 10 ″ W; CEMT • 2 specs; same data as for preceding; horse feces; CEMT • 1 spec.; same data as for preceding; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 14 ″ W; 23 Jan. 2020; Sehn et al. leg.; horse feces; CEMT • 2 specs; same data as for preceding; cow feces; CEMT • 1 spec.; same data as for preceding; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 10 ″ W; elephant feces; CEMT • 2 specs; same data as for preceding; 15 ° 18 ′ 50 ″ S, 55 ° 24 ′ 12 ″ W; 24 Jan. 2020; H. Sehn et al. leg.; elephant feces; CEMT • 1 spec.; Cotriguaçu; 31 Oct. 2007; O. Peres-Filho leg.; CEMT • 4 specs; same data as for preceding; 15 Dec. 2007; light; CEMT • 1 spec.; same data as for preceding; 16 Jan. 2008; CEMT • 2 specs; same data as for preceding; Assentamento INCRA; 1 Nov. 2017; R. Stofel leg.; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 15 Aug. 2007; O. P. Filho leg.; CEMT • 3 specs; same data as for preceding; 31 Oct. 2007; O. Peres-Filho leg.; light; CEMT • 1 spec.; same data as for preceding; 15 Dec. 2007; CEMT • 8 specs; same data as for preceding; 9 ° 51 ′ 52 ″ S, 58 ° 13 ′ 45 ″ W; 2 Oct. 2017; Vaz-de-Mello et al. leg.; CEMT • 23 specs; same data as for preceding; airport; light trap; CEMT • 1 spec.; same data as for preceding; 9 ° 51 ′ 41 ″ S, 58 ° 13 ′ 51 ″ W; 2 Nov. 2017; light; CEMT • 2 specs; same data as for preceding; airport; CEMT • 1 spec.; same data as for preceding; 9 ° 49 ′ 23 ″ S, 58 ° 17 ′ 14 ″ W; 3 Nov. 2017; light trap; CEMT • 1 spec.; same data as for preceding; 9 ° 50 ′ 19 ″ S, 58 ° 15 ′ 15 ″ W; 29 Oct. 2017; Costa-Silva et al. leg.; CEMT • 2 specs; same data as for preceding; 9 ° 49 ′ 9 ″ S, 58 ° 15 ′ 31 ″ W; 21 Oct. 2018; CEMT • 1 spec.; Nobres; 30 Sep. 2018; R. Segalla and M. Cassiano leg.; CEMT • 6 specs; Diamantino, Reserva Vale da Solidão; 14 ° 22 ′ S, 56 ° 7 ′ W; elev. 450 m; 30 Oct. 2013; E. Furtado leg.; CEMT • 4 specs; Nova Ubiratã, ESEC Rio Ronuro; 25 Feb. 2017; L. G. O. A. Nunes et al leg.; CEMT • 1 spec.; same data as for preceding; 13 ° 06 ′ 43 ″ S, 54 ° 26 ′ 37 ″ W; 2 Mar. 2017; L. G. O. A. Nunes leg.; light; CEMT • 3 specs; Santo Antônio do Leverger, Porto de Fora; 16 ° 05 ′ 17 ″ S, 55 ° 48 ′ 41 ″ W; 29 – 31 Dec. 2019; B. R. Bordin and B. P. Queiroz leg.; cow dung; CEMT • 1 spec.; Sesc Pantanal; 7 Jul. 2003; light; CEMT. – Mato Grosso do Sul • 1 spec.; Inocência, Fazenda Moinho; 19 ° 13 ′ 12 ″ S, 52 ° 09 ′ 34 ″ W; elev. 502 m; 15 Oct. 2018; H. L. Rainho leg.; light; CEMT • 26 specs; same data as for preceding; 21 Oct. 2018; CEMT • 1 spec.; Guia Lopes da Laguna; 15 Feb. 2007; A. Abot leg.; light; CEMT • 1 spec.; same data as for preceding; 29 Jul. 2008; CEMT • 1 spec.; Selvíria, UNESP farm; 22 Jan. 1991, C. A. H. Flechtmann; Brachiaria decumbens pasture; Bubalus bubalis dropping; CEMT • 1 spec.; same data as for preceding; 16 Mar. 1991; CEMT • 1 spec.; same data as for preceding; 8 Jun. 1991; CEMT • 1 spec.; same data as for preceding; 10 Aug. 1991; CEMT • 3 specs; same data as for preceding; 6 Jul. 1991; CEMT • 1 spec.; same data as for preceding; 1 Jun. 1991; S. R. Rodrigues leg.; INPA. – Minas Gerais • 2 specs; Campanha, Rod. Vital Brazil; 21 ° 47 ′ 49 ″ S, 45 ° 25 ′ 40 ″ W; 5 Jun. 2016; F. França leg.; CEMT • 1 spec.; Conceição do Mato Dentro, 23 K; elev. 734 m; 22 Sep. 2012; Uceli and Borges leg.; CEMT • 1 spec.; Cordisburgo, Fazenda Pontinha; Jan. 1996; Vaz-de-Mello leg.; CEMT • 21 specs; Frei Gaspar, Sítio Recanto Paraíso; 18 ° 02 ′ 06 ″ S, 41 ° 29 ′ 10 ″ W; elev. 571 m; 28 Dec. 2007; M. R. Santos leg.; CEMT • 10 specs; same data as for preceding; 30 Dec. 2007; light; CEMT • 24 specs; same data as for preceding; 3 Jan. 2008; CEMT • 2 specs; Ipatinga; Sep. 1993; E. Grossi leg.; CEMT • 2 specs; Uberaba; Oct. 1995; Vaz-de-Mello leg.; CEMT • 4 specs; Viçosa; 20 Sep. 1993; Louzada and Silva leg.; CEMT. – Paraná • 8 specs; Icaraíma; 23 ° 23 ′ S, 53 ° 37 ′ W; elev. 357 m; 2019; Felipe and Rodrigues leg.; CEMT • 22 specs; same data as for preceding; Estância Felipe; 23 ° 20 ′ 32 ″ S, 53 ° 39 ′ 09 ″ W; elev. 300 m; CEMT • 1 spec.; Rancho Alegre, Fazenda Congonhas; 22 ° 47 ′ 45 ″ S, 51 ° 00 ′ 12 ″ W; elev. 580 m; 27 Dec. 2012; P. M. Pelix leg.; canopy; feces; CEMT. – Rio de Janeiro • 3 specs; Cachoeiras de Macacu, Guapiaçu ER; 22 ° 27 ′ 05 ″ S, 42 ° 46 ′ 23 ″ W; elev. 33 m; 25 Oct. 2019; L. R. Martínez leg.; horse feces; CEMT • 46 specs; Jan. 1998; J. Carlos leg.; CEMT • 1 spec.; same data as for preceding; Feb. 1998; CEMT • 28 specs; same data as for preceding; Mar. 1998; CEMT • 1 spec.; Nova Friburgo; Nov. 2002; E. J. Grossi leg.; CEMT. – Rio Grande do Sul • 1 spec.; Aceguá, Fazenda Sentinela; 31 ° 27 ′ 30 ″ S, 54 ° 21 ′ 18 ″ W; 8 – 14 Nov. 2011; R. M. Moraes leg.; FIT; CEMT • 1 spec.; Bagé, Santo Afonso; 7 – 13 Jan. 2012; R. M. Moraes leg.; soy farm; human and pig dung; CEMT • 1 spec.; Eldorado do Sul; 4 Oct. 1996; Carvalho and Silva leg.; CEMT • 1 spec.; same data as for preceding, Estação Experimental UFRGS; 30 Mar. – 5 Apr. 2012; R. M. Moraes and J. Podkowa; pitfall; CEMT • 1 spec.; Santa Cruz do Sul, Cerro Alegre; 29 ° 48 ′ 02 ″ S, 51 ° 19 ′ 39 ″ W; 10 Dec. 2010; R. Bohn; pitfall; CEMT • 18 specs; same data as for preceding; 29 ° 48 ′ 05 ″ S, 52 ° 19 ′ 39 ″ W; 24 Mar. 2012; R. Bohn leg.; grassfield; bovine feces; CEMT • 7 specs; same data as for preceding; 29 ° 48 ′ 11 ″ S, 52 ° 19 ′ 51 ″ W; annual crop; CEMT • 7 specs; same data as for preceding; 29 ° 48 ′ 17 ″ S, 52 ° 19 ′ 59 ″ W; forest; CEMT • 8 specs; same data as for preceding; 29 ° 48 ′ 19 ″ S, 52 ° 19 ′ 53 ″ W; secondary forest; CEMT • 29 specs; same data as for preceding; 29 ° 48 ′ 18 ″ S, 52 ° 19 ′ 55 ″ W; open area 29; CEMT. – São Paulo • 2 specs; Porto Ferreira; 21 ° 52 ′ 11 ″ S, 47 ° 28 ′ 5 ″ W; 19 Oct. 2019; G. Biffi leg.; CEMT • 1 spec.; São Paulo, Jardim Marilda; 23 ° 47 ′ 19 ″ S, 46 ° 40 ′ 34 ″ W; elev. 754 m; Nov. – Dec. 2020; H. Marota leg.; CEMT. – Tocantins • 3 specs; Palmas, Fazenda Recanto Valadares; 20 Sep. 2005; M. S. Matos leg.; CEMT. CHILE – Santiago • 3 specs; El Peñón; 28 Jan. 1995; V. M. M. Diéguez leg.; CEMT. ECUADOR – Loja • 1 spec.; Gonzanama; elev. 2000 m; 28 Aug. 1997; C. Carpio leg.; CEMT. PARAGUAY – Amambay • 1 spec.; Pedro Juan Caballero; Nov. 1998; M. A. Ruíz-Dias leg.; CEMT. MEXICO – Chiapas • 3 specs; Cintalapa, Ejido Gustavo Díaz Ordaz; 16 ° 40 ′ 15 ″ N, 94 ° 11 ′ 27 ″ W; elev. 1035 m; 4 Jul. 2006; Reyes-Castillo and Vaz-de-Mello leg.; UV light; CEMT • 1 spec.; Palenque; Jun. 2004; F. Nicolalde leg.; CEMT. – Michoacán • 1 spec.; Tzintzuntzita; 19 ° 37 ′ 34 ″ N, 101 ° 34 ′ 43 ″ W; elev. 2050 m; Jul. 2007; F. Z. Vaz-de-Mello leg.; CEMT. – Veracruz • Perote, El Conejo; 19 ° 31 ′ 20 ″ N, 97 ° 09 ′ 39 ″ W; elev. 3500 m; Jul. 2007; F. Z. Vaz-de-Mello and Louzada leg.; CEMT. UNITED STATES OF AMERICA – California • 1 spec.; Co. Riverside; 13 Feb. 1971; K. R. Cardin leg.; INPA. URUGUAY – Cerro Largo • 1 spec.; Meio, La Invernada farm; 13 – 20 Dec. 2011; R. M. Moraes leg.; human and pig dung; CEMT. – Rivera • 1 spec.; Rivera, COFUSA; 30 ° 57 ′ 36 ″ S, 55 ° 29 ′ 53 ″ W; 3 – 11 Oct. 2011; human and pig dung; R. M. Moraes leg.; CEMT • 2 specs; Rivera, Bodegas Carrau farm; 30 ° 58 ′ 19 ″ S, 55 ° 26 ′ 40 ″ W; FIT; CEMT • 2 specs; same data as for preceding; human and pig dung; CEMT • 4 specs; same data as for preceding; 30 ° 58 ′ 16 ″ S, 55 ° 26 ′ 38 ″ W; human and pig dung; CEMT • 2 specs; Vichadero, Adelaida farm; 31 ° 41 ′ 5.62 ″ S, 54 ° 39 ′ 42.76 ″ W; 3 – 11 Oct. 2011; R. M. Moraes leg.; human and pig dung; CEMT • 4 specs; same data as for preceding; 31 ° 41 ′ 04 ″ S, 54 ° 37 ′ 25 ″ W; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8AFFC2FDFFFE95FEFDFEF5.taxon	description	Sexual dimorphism Male with median tubercle more developed than that of female.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF8AFFC2FDFFFE95FEFDFEF5.taxon	discussion	Collecting information Labarrus pseudolividus specimens were collected in bovine, human, pig, horse and Tapirus feces and with black light and flight intercept traps. This species can be found at altitudes ranging from 64 to 2050 metres in open areas such as grasslands and soy crops or in areas of secondary forest (Mesquita et al. 2017).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF95FFC2FDE1FEF3FE79FCED.taxon	diagnosis	Diagnosis Length 7.0 – 9.0 mm. Body strongly convex, glabrous dorsally, piceous, elytra testaceous posteriorly. Head with moderate punctures, frontoclypeal suture weakly impressed, faint; clypeus sinuate. Pronotum without lateral and posterior marginal line, with punctures of two sizes. Mesoscutellum large, almost 1 / 3 the length of elytral suture. Protibia punctate dorsally, meso- and metatibiae with strong set of transverse carinae; metatibiae apically fringed by setae of similar sizes.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF95FFC2FDE1FEF3FE79FCED.taxon	discussion	Remarks Neodiapterna comprises two species. Neodiapterna erichsoni is distributed across Argentina, Brazil and Paraguay (Dellacasa & Dellacasa 1997). Neodiapterna tognoni Dellacasa & Dellacasa, 1997 is only known from Bolivia (Dellacasa & Dellacasa 1997). A key for the species can be found in Dellacasa & Dellacasa (1997).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF95FFC2FDEDFC18FB6DF83A.taxon	description	Figs 7 B, 37 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF95FFC2FDEDFC18FB6DF83A.taxon	diagnosis	Diagnosis Length 7.0 – 9.0 mm. Pronotum with large punctures amply distributed on disc and lateral margins; posterior angles obtuse, ending in a subtle but evident backwards projection.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF95FFC2FDEDFC18FB6DF83A.taxon	materials_examined	Material examined BRAZIL – Paraná • 1 spec.; Castro Socavão, Cachoeira Roncadora; 24 Jun. 2006; Grossi and Parizorro leg.; CEMT. – Rio Grande do Sul • 2 specs; Bagé; May 2006; L. D. Audino leg.; CEMT • 1 spec.; same data as for preceding; Embrapa CPP Sul; 31 Jul. 2006; L. D. Audino leg.; Eucalyptus; FIT; CEMT. – Santa Catarina • 1 spec.; Urubici (Parna S. JQ); 49 ° 37 ′ 04.80 ″ S, 28 ° 8 ′ 43.88 ″ W; 14 Dec. 2015; P. G. da Silva leg.; campo; horse feces; CEMT. – São Paulo • 1 spec.; São Miguel Arcanjo, P. E. Carlos Botelho; 24 ° 3 ′ 59 ″ S, 47 ° 59 ′ 39 ″ W; elev. 580 m; 15 May 2012; M. Boutefeu leg.; Tapirus feces; CEMT • 1 spec.; same data as for preceding; 11 Jun. 2012; CEMT • 1 spec.; same data as for preceding; 6 Jul. 2012; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF95FFC2FDEDFC18FB6DF83A.taxon	description	Sexual dimorphism Females with apical spur of protibia more slender and downwardly bent when compared with males. Pronotum more densely punctate in females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF95FFC2FDEDFC18FB6DF83A.taxon	discussion	Collecting information Specimens were collected in Tapirus and horse feces and with a flight intercept trap.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF97FFC0FE48FE94FA82FC44.taxon	diagnosis	Diagnosis Length 3.5 – 6.0 mm. Body, oblong, dorsally glabrous, testaceous with brownish areas or dark testaceous. Head wide, finely punctate; clypeus weakly sinuate or not sinuate at all; frontoclypeal suture evident; gena strongly obtuse. Pronotum wide, with punctures of two sizes, without marginal line posteriorly; posterior angles of pronotum truncate, weakly inwardly sinuate. Elytra with interstriae convex, microreticulate and weakly punctate. Protibiae punctate dorsally, metatibiae weakly broadened apically. Males with second labial palpomere enlarged, wider and longer than first and third palpomeres.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF97FFC0FE48FE94FA82FC44.taxon	discussion	Remarks Neotrichaphodioides is a genus with five species known from Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Peru and Venezuela (Dellacasa et al. 2010 b). A key for all its species can be found in Dellacasa et al. (2010 b).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF97FFC7FE2DFB80FD1FFDB6.taxon	description	Figs 5 C – D, 37 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF97FFC7FE2DFB80FD1FFDB6.taxon	diagnosis	Diagnosis Length 3.5 – 4.5 mm. Body light testaceous. Head with clypeus weakly sinuate, rounded. Pronotum with punctures of two sizes; posterior margin weakly sinuate. Elytra lighter than head and pronotum with darkened rounded spot; interstriae weakly convex.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF97FFC7FE2DFB80FD1FFDB6.taxon	materials_examined	Material examined BRAZIL – Distrito Federal • 1 ♂; Planaltina, Embrapa Cerrados; 15 ° 36 ′ 20 ″ S, 47 ° 42 ′ 26 ″ W; 7 Oct. 2005; C. Oliveira leg.; cerrado fragment; light; CEMT • 1 ♂; same data as for preceding; 20 Oct. 2005; native cerrado; CEMT • 8 ♂♂, 1 ♀; same data as for preceding; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 2 Dec. 2005; agricultural area; CEMT • 9 ♂♂, 1 ♀; same data as for preceding; 15 ° 36 ′ 16 ″ S, 47 ° 44 ′ 16 ″ W; native cerrado; CEMT • 4 ♀♀; same data as for preceding; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 18 Nov. 2005; agricultural area; CEMT • 1 ♂, 2 ♀♀; same data as for preceding; 15 ° 36 ′ 16 ″ S, 47 ° 44 ′ 16 ″ W; native cerrado; CEMT • 2 ♂♂, 1 ♀; same data as for preceding; 5 Jan. 2006; CEMT • 1 ♀; same data as for preceding; 5 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 19 Jan. 2006; agricultural area; CEMT • 1 ♀; same data as for preceding; 19 Jun. 2006; CEMT • 42 ♂♂, 91 ♀♀; same data as for preceding; 15 ° 36 ′ 20 ″ S, 47 ° 42 ′ 16 ″ W; 3 Nov. 2006; CEMT. – Mato Grosso • 1 ♂; Chapada dos Guimarães; 11 Dec. 2000; A. F. Ramos leg.; light; CEMT • 2 ♂♂; same data as for preceding; Santuário dos Elefantes; 2 Nov. 2019; F. Z. Vaz-de-Mello leg.; CEMT. – Minas Gerais • 1 ♂; Ijací, Fazenda FAEPE; Nov. 2002; J. Louzada leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF97FFC7FE2DFB80FD1FFDB6.taxon	description	Sexual dimorphism Males with labial palpomere enlarged, head wider and clypeus less sinuate than that of females; females with head and pronotum more densely punctate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF97FFC7FE2DFB80FD1FFDB6.taxon	discussion	Collecting information Specimens are easily collected at light in Cerrado and agricultural areas. This species was found at altitudes up to 1380 m (Dellacasa et al. 2010 b).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF90FFC7FDD2FDB6FD92F8A5.taxon	description	Figs 5 A – B, 37 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF90FFC7FDD2FDB6FD92F8A5.taxon	diagnosis	Diagnosis Length 5.0 – 6.0 mm. Body dark testaceous. Head with clypeus trapezoidal. Pronotum with punctures of two sizes; posterior margin strongly sinuate. Elytra as dark as disc of pronotum and head, with a rounded lighter area posteriorly located; interstriae strongly convex.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF90FFC7FDD2FDB6FD92F8A5.taxon	materials_examined	Material examined BRAZIL – Distrito Federal • 1 ♀; Brasília, Planaltina; 15 ° 36 ′ 24.52 ″ S, 47 ° 44 ′ 42.45 ″ W; 6 Nov. 2015; A. Specht leg.; CEMT. – Minas Gerais • 1 ♂; Corinto; 16 Jan. 1998; CEMT • 5 ♂♂, 2 ♀♀; Ingaí-Lavras, Próx Poço Bonito; Nov. 2002; F. Z. Vaz-de-Mello leg.; CEMT • 2 ♂♂, 3 ♀♀; Ijací, Fazenda FAEPE; Nov. 2002; J. Louzada leg.; CEMT • 1 ♂, 1 ♀; Lavras, Campus da Ufla; Nov – Dec. 2001; F. Z. Vaz-de-Mello leg.; light; CEMT • 1 ♂; same data as for preceding; 15 Nov. 1997; CEMT • 1 ♀; Lavras, Prox. Poço Bonito; Dec. 2001; Vaz-de-Mello leg.; CEMT • 3 ♂♂, 1 ♀; Santa Barbará; 17 Feb. 1996; J. C. Zanuncio leg.; CEMT • 1 ♀; Sete Lagoas, Embrapa; 19 ° 26 ′ 49 ″ S, 44 ° 10 ′ 19 ″ W; elev. 715 m; 25 Nov. 2011; Mendes leg.; light; CEMT • 1 ♂, 1 ♀; Teixeiras; Nov. 1996; Hardy and Harrison leg.; CEMT • 1 ♂; Viçosa; Oct. 1998; Vaz-de-Mello leg.; FIT; CEMT • 1 ♂; same data as for preceding; 9 Nov. 2000; M. F. G. Mata leg.; CEMT • 1 ♀; same data as for preceding; elev. 780 m; 7 – 14 Nov. 1996; M. Hardy leg.; CEMT • 1 ♀; same data as for preceding; Córrego do Paraíso, Mata da Prefeitura; 11 Dec. 1984; Fiuza and Martins leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF90FFC7FDD2FDB6FD92F8A5.taxon	description	Sexual dimorphism Males with second palpomere of labial palpi enlarged, head wider and clypeus less sinuate than that of females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF90FFC7FDD2FDB6FD92F8A5.taxon	discussion	Collecting information Specimens collected at light and with flight intercept traps. This species can be found at altitudes of 780 metres (Dellacasa et al. 2010 b).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF90FFC6FD8FF8A0FBC2FDAC.taxon	diagnosis	Diagnosis Length 3.5 – 5.5 mm. Body elongate, piceous. Head, with clypeus emarginate; gena prominent; frontoclypeal suture tuberculate. Pronotum convex, punctate with punctures of two sizes. Elytral interstriae weakly convex; punctate with fine punctures. Protibiae not punctate in dorsal view. Meso- and metatibiae with evident set of transversal carinae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF90FFC6FD8FF8A0FBC2FDAC.taxon	discussion	Remarks Similar to Labarrus, Nialaphodius may have been introduced in the New World. The genus is currently comprised of two species Nialaphodius calidus (Harold, 1871) and N. nigrita (Schoolmeesters 2023). Both species are widely distributed across the African continent but only N. nigrita is currently recorded in the New Word (Dellacasa et al. 2001 a; Skelley et al. 2007; Bordat 2011).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF91FFC5FDEFFDDBFD1EF921.taxon	description	Figs 7 C, 37 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF91FFC5FDEFFDDBFD1EF921.taxon	diagnosis	Diagnosis Length 3.5 – 5.5 mm. Body strongly convex. Head densely punctate; frontoclypeal suture elevate. Pronotum with punctures of two sizes, fine and coarse; posterior margin without posterior marginal line.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF91FFC5FDEFFDDBFD1EF921.taxon	materials_examined	Material examined BOLIVIA – Santa Cruz • 1 spec.; Chiquitos, 1.6 km E of Santiago de Chiquitos; 18 ° 12 ′ 4 ″ S, 59 ° 21 ′ 0 ″ W; elev. 622 m; Nov. 2008; W. D. Edmonds and T. Vidaurre leg.; light; CEMT. BRAZIL – Amazonas • 1 spec.; Lago de Janauoá; 30 Aug. 1962; Júlio and Delono leg.; CEMT • 1 spec.; Itacoatiara, Fazenda Aruana, Am 010, Km 215; 12 – 15 Jan. 1991; C. S. Motta et al. leg.; CEMT. – Bahia • 1 spec.; Caravelas; Mar. 1996; R. Diniz leg.; CEMT • 1 spec.; same data as for preceding; Feb. 1997; S. R. Silveira leg.; CEMT • 1 spec.; Macuri City; 4 Oct. 1997; Anjos and Silveira leg.; CEMT. – Distrito Federal • 1 spec.; Brasília; elev. 1100 m; Feb. 2001; N. Degallier leg.; CEMT. – Espírito Santo • 1 spec.; Conceição do Castelo, Ribeirão do Meio, Posto Pinga Fogo BR 262; Feb. 2010; R. Falqueto leg.; CEMT. – Maranhão • 3 specs; Carolina, Fazenda Cincorá; 17 – 22 Oct. 2009; F. Limeira-de-Oliveira et al. leg.; light; CEMT. – Mato Grosso • 1 spec.; Res. Papagaio, BR- 364 Km 555; 2 Oct. 1984; E. Binda leg.; light; INPA • 1 spec.; Chapada dos Guimarães, Santuário dos Elefantes; 15 ° 18 ′ 51 ″ S, 55 ° 24 ′ 10 ″ W; 4 Sep. 2019; H. Sehn leg.; elephant feces; CEMT • 1 spec.; same data as for preceding; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 7 ″ W; 23 Jan. 2020; cow feces; CEMT • 2 specs; Cotriguaçu, Fazenda São Nicolau; 16 Oct. 2007; O. P. Peres-Filho leg.; light; CEMT • 2 specs; same data as for preceding; 31 Nov. 2007; CEMT • 3 specs; same data as for preceding; light; CEMT • 2 specs; same data as for preceding; Nov. 2007; CEMT • 1 spec.; same data as for preceding; 15 Dec. 2007; CEMT • 9 specs; same data as for preceding; 9 ° 51 ′ 52 ″ S, 58 ° 13 ′ 45 ″ W; 2 Oct. 2017; Vaz-de-Mello et al. leg.; airport; light trap; CEMT • 1 spec.; same data as for preceding; 9 ° 49 ′ 23 ″ S, 58 ° 17 ′ 14 ″ W; 3 Oct. 2017; CEMT • 1 spec.; same data as for preceding; 9 ° 50 ′ 19 ″ S, 58 ° 15 ′ 15 ″ W; Matinha do Fernando; CEMT • 1 spec.; same data as for preceding; 9 ° 51 ′ 20 ″ S, 58 ° 14 ′ 55 ″ W; Nov. 2014; R. V. Nunes and A. Asenjo leg.; light; CEMT • 5 specs; same data as for preceding; 9 ° 50 ′ 19 ″ S, 58 ° 15 ′ 3 ″ W; 25 Oct. 2014; A. Asenjo leg.; light; CEMT • 1 spec.; Diamantino; Jan. 2001; A. M. Bello leg.; CEMT • 1 spec.; Sesc Pantanal; 13 Apr. 2002; CEMT. – Mato Grosso do Sul • 1 spec.; Dourados; 6 Feb. 2006; M. Milo leg.; CEMT • 2 specs; Guia Lopes da Laguna; 25 May 2008; A. Abot leg.; CEMT • 1 spec.; Selvíria, UNESP farm; 1 Jun. 1991; C. A. H. Flechtmann leg.; Brachiaria decumbens pasture; Guzera bovine dropping; CEMT • 1 spec.; same data as for preceding; 4 Jul. 1990; CEMT • 1 spec.; same data as for preceding; 11 Jul. 1990; CEMT • 2 specs; same data as for preceding; 22 Jan. 1991; CEMT • 1 spec.; same data as for preceding; 29 Jan. 1991; CEMT • 1 spec.; same data as for preceding; 1 Jun. 1991; CEMT • 2 specs; same data as for preceding; 20 Jul. 1991; CEMT • 1 spec.; same data as for preceding; 10 Aug. 1991; CEMT • 1 spec.; same data as for preceding; 20 Aug. 1991; CEMT • 1 spec.; same data as for preceding; 27 Oct. 1991; CEMT • 1 spec.; same data as for preceding; 30 May 1992; CEMT • 1 spec.; same data as for preceding; 30 May 1992; CEMT • 2 specs; same data as for preceding; 25 Jul. 1992; CEMT • 2 specs; same data as for preceding; 30 May 1993; CEMT • 1 spec.; same data as for preceding; 1 Jun. 1991; S. R. Rodrigues leg.; INPA. – Minas Gerais • 1 spec.; Conceição do Mato Dentro, 23 k; 66 ° 41 ′ 32 ″ S, 79 ° 11 ′ 853 ″ W; elev. 734 m; 5 Dec. 2012; Uceli and Borges leg.; light; CEMT • 1 spec.; Cordisburgo, Fazenda Pontinha; Jan. 1994; CEMT • 1 spec.; same data as for preceding; Jul. 1994; de Mello leg.; CEMT • 2 specs; same data as for preceding; Jan. 1996; Vaz-de-Mello leg.; CEMT • 3 specs; same data as for preceding; Jan. 1999; CEMT • 1 spec.; Ipatinga; Sep. 1993; E. Grossi leg.; CEMT • 1 spec.; Mariana; 7 Dec. 1998; J. L. Moreno and T. Mestre leg.; CEMT • 3 specs; Paraopeba; Feb. 1997; Lourenço S. leg.; CEMT • 1 spec.; Viçosa; 24 May 1997; CEMT • 1 spec.; Visconde do Rio Branco; Jan. 1998; W. Zikan leg.; CEMT • 1 spec.; Recreio; Jan. 2000; Lopes-Andrade leg.; CEMT. – Rio de Janeiro • 1 spec.; Nova Friburgo, Macaé de Cima; Nov. 1998; F. Z. Vaz-de-Mello and E. Grossi leg.; CEMT. – Rondônia • 1 spec.; Anhembi, ESALQ / USP; E. N. L. Ferreira leg.; CEMT • 1 spec.; 62 km S of Ariquemes, Fazenda Rancho Grande; Feb. 1997; Vaz-de-Mello et al. leg.; CEMT • 1 spec.; Peto Velho, Rio das Garças; 8 ° 49 ′ 56 ″ S, 63 ° 46 ′ 47 ″ W; 27 Aug. 2017; D. C. Santos and K. K. G. Silva leg.; FIT; CEMT. – Roraima • 1 spec.; Amajari, Eec Maracá, Uraricoera River; 3 ° 20 ′ 59 ″ N, 61 ° 25 ′ 22 ″ W; elev. 83 m; 24 Mar. 2016; A. P. M. Santos et al. leg.; CEMT. – São Paulo • 1 spec.; Beira Pereiro Barreto, Vale do Paraíso Farm; 5 May 1993; M. A. F. Aio leg.; Brachiaria decumbens pasture; Nelore dropping; CEMT. COLOMBIA – Cundinamarca • 1 spec.; Fusa; Sep. 2000; J. Noriega leg.; cow feces; CEMT. ECUADOR – Manabi • 1 spec.; El Carmen; 0 ° 16 ′ 08 ″ S, 79 ° 28 ′ 20 ″ W; elev. 260 m; 28 Dec. 1995; T. Jaramillo leg.; CEMT. – Zamora • 1 spec.; Ch. Las Penãs; elev. 1000 m; 24 Aug. 1997; C. Carpio leg.; CEMT. MEXICO – Chiapas • 2 specs; Palenque; Jun. 2004; F. Nicolalde leg.; CEMT • 2 specs; Cintalapa, Ejido, Gustavo Díaz Ordaz; 16 ° 40 ′ 15 ″ N, 94 ° 11 ′ 27 ″ W; elev. 1035 m; 4 Jul. 2006; Reyes-Castillo and Vaz-de-Mello leg.; UV light; CEMT. PARAGUAY – Paraguary • 1 spec.; Naranjo; 5 Nov. 2000; G. Arriágada leg.; CEMT. PERU – Corongo • 3 specs; La Pampa; 5 – 12 Feb. 2012; P. Castilla leg.; CEMT. UNITED STATES OF AMERICA – Florida • 1 spec.; W. S. B. Dunedin leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF91FFC5FDEFFDDBFD1EF921.taxon	description	Sexual dimorphism Males with frontoclypeal tubercle more developed than that of females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF91FFC5FDEFFDDBFD1EF921.taxon	discussion	Collecting information Specimens were collected in Urochloa brizantha pasture regions with UV light and flight intercept trap as well as in cattle dung (Mesquita et al. 2017).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF92FFC5FDE3F927FCDCF839.taxon	diagnosis	Diagnosis Length 3.0 – 5.0 mm. Body oval-elongate, glabrous. Head with clypeus sinuate medially; frontoclypeal suture evident, not tuberculate. Pronotum transverse; posterior angles obliquely truncate. Elytra with humeral tooth developed; interstriae of disc carinate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF93FFC4FDC3FE95FC64FBA5.taxon	diagnosis	Diagnosis Length 3.5 mm. Body narrow, weakly arched. Head densely punctured with anterior margin somewhat elevate, clypeus faintly sinuate. Pronotum with posterior marginal line. Elytra with humerus lacking teeth; interstriae somewhat raised, third, fifth and sixth interstriae more elevate than others.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF93FFC4FDC3FE95FC64FBA5.taxon	description	Sexual dimorphism No information available in literature.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF93FFC4FDC3FE95FC64FBA5.taxon	discussion	Collecting information No information available in literature. Remarks Pleuraphodius is a genus considered to be restricted to the Old World (Dellacasa et al. 2001 a). Pleuraphodius vespuccii was described from Serra de Paranapiacaba, São Paulo State, Brazil, but Skelley et al. (2007) comment on how this species may have been described based on a mislabeled specimen and should probably be removed from the list of New World taxa.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF93FFC4FDECFBA0FBCFFA54.taxon	diagnosis	Diagnosis Length 6.0 – 8.0 mm. Body oblong, dark testaceous. Head with small and sparse punctures; clypeus sinuate medially, with medial tubercle anterior to frontoclypeal suture; frontoclypeal suture evident. Pronotum with sparse punctation, punctures of two sizes; pronotum without posterior marginal line. Dorsal surface of protibia with dense punctation. Meso- and metatibia with two distinct transverse carinae on lateral margin. Apical and lateral margin of metatibia crowned with setae; setae progressively longer closer to margins.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF93FFC4FDECFBA0FBCFFA54.taxon	discussion	Remarks Trichaphodiellus is a Neotropical monotypic genus (Dellacasa et al. 2001 a).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF93FFCAFE23F993FAF8FA03.taxon	description	Figs 4 D – F, 38 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF93FFCAFE23F993FAF8FA03.taxon	diagnosis	Diagnosis Same as for the genus.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF93FFCAFE23F993FAF8FA03.taxon	materials_examined	Material examined BRAZIL – Acre • 2 specs; Rio Branco; CEMT • 1 spec.; same data as for preceding; 10 ° 01 ′ 58.09 ″ S, 67 ° 42 ′ 12.74 ″ W; 9 Aug. 2015; A. Specht leg.; light; CEMT. – Amazonas • 1 spec.; Presidente Figueiredo, Ilha Pantanal; 1 ° 47 ′ 49 ″ S, 59 ° 26 ′ 53 ″ W; 22 – 24 Jun. 2016; L. Sawaris leg.; submontane dense ombrophilous forest; human and pig dung; CEMT. – Espírito Santo • 1 spec.; Linhares, AFCVRD; Oct. 1988; J. S. Santos leg.; CEMT • 2 specs; same data as for preceding; Reserva Natural Vale; 19 ° 09 ′ 04 ″ S, 40 ° 04 ′ 14 ″ W; Dec. 1987; J. S. Santos leg.; light; CEMT. – Mato Grosso • 1 spec.; Alta Floresta, CEPLAC; 4 Sep. 2009; V. Gonçalves leg.; Tapirus feces; CEMT • 1 spec.; same data as for preceding; CEMT • 3 specs; same data as for preceding; CEMT • 1 spec.; Apiacás, Parna Juruena; 10 Mar. 2011; M. Lima leg.; CEMT • 1 spec.; Chapada dos Guimarães, Santuário dos Elefantes; 15 ° 18 ′ 50 ″ S, 55 ° 24 ′ 19 ″ W; 21 Jan. 2020; H. Sehn et al. leg.; elephant feces; CEMT • 4 specs; same data as for preceding; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 14 ″ W; 22 Jan. 2020; CEMT • 1 spec.; same data as for preceding; 15 ° 18 ′ 22 ″ S, 55 ° 24 ′ 14 ″ W; 23 Jan. 2020; CEMT • 1 spec.; same data as for preceding; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 07 ″ W; CEMT • 1 spec.; same data as for preceding; 15 ° 18 ′ 50 ″ S, 55 ° 24 ′ 12 ″ W; 24 Jan. 2020; CEMT • 3 specs; Poconé; 26 Apr. 1998; light; CEMT • 1 spec.; Corumbá, Porto Jofre, Fazenda São Bento; 13 Apr. 2011; R. V. Nunes leg.; CEMT • 1 spec.; Cotriguaçú, Fazenda São Nicolau; 10 Dec. 2009; M. R. Barreto leg.; forest; CEMT • 1 spec.; Poconé, Base UFMT; 16 ° 29 ′ 56 ″ S, 56 ° 24 ′ 49 ″ W; Mar. 2012; M. Rossini leg.; CEMT. – Mato Grosso do Sul • 1 spec.; Chapadão do Sul; 18 ° 46 ′ 30.09 ″ S, 52 ° 31 ′ 04.98 ″ W; 5 Jan. 2016; A. Specht leg.; light; CEMT. – Minas Gerais • 1 spec.; Conceição do Mato Dentro, 23 K; 67 ° 69 ′ 53 ″ S, 79 ° 11 ′ 57 ″ W; elev. 711 m; 16 Sep. 2012; Uceli and Borges leg.; CEMT • 1 spec.; same data as for preceding; 66 ° 31 ′ 39 ″ S, 79 ° 11 ′ 235 ″ W; elev. 704 m; 20 Sep. 2012; Uceli and Borges leg.; light; CEMT • 1 spec.; Cordisburgo; Jul. 1992; F. Zagury leg.; CEMT • 1 spec.; same data as for preceding; Fazenda Pontinha; Jan. 1996; Vaz-de-Mello leg.; CEMT • 1 spec.; Marlieria; Jul. 1994; Mello leg.; CEMT • 1 spec.; Viçosa; 11 Feb. 1996; J. N. C. Louzada leg.; CEMT • 4 specs; Pains, ISLA G. do Zé Serafim III; 25 Aug. 2009; R. A. Zampauto leg.; CEMT • 6 specs; Paraopeba; Mar. 1997; S. Lourenço jr leg.; CEMT. – Pará • 1 spec.; Parauapebas; 5 ° 56 ′ 46 ″ S, 50 ° 12 ′ 17 ″ W; 26 Feb. 2019; F. França leg.; secondary forest; human and pig dung; CEMT. – Paraná • 2 specs; Guaratuba, Estrada dos Castelhanos; elev. 650 m; 25 Nov. 2009; Grossi et al. leg.; light; CEMT. – Paraíba • 1 spec.; Areia, Mata de Pau Ferro; 2 May – 1 Jun. 2000; Avany and Maria leg.; light; CEMT. – Rio de Janeiro • 9 specs; Cachoeira de Macacu, Guapiaçu ER; 22 ° 27 ′ 5 ″ S, 42 ° 46 ′ 23 ″ W; elev. 33 m; 22 Sep. 2019; L. R. Martínez leg.; Tapirus feces; CEMT • 1 spec.; same data as for preceding; 25 Oct. 2019; horse feces • 7 specs; same data as for preceding; 10 Jan. 2020; CEMT • 1 spec.; Miguel Pereira; Mar. 1998; J. Carlos leg.; CEMT • 2 specs; Nova Friburgo; Nov. 1993; E. J. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Dec. 1995; P. Grossi and E. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Nov. 1997; CEMT • 2 specs; same data as for preceding; Macaé de Cima; Nov. 1998; F. Z. Vaz-de-Mello and E. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Dec. 2000; P. Grossi leg.; CEMT • 1 spec.; Santa Maria Madalena, Pousada Verbicaro, RPPN, MAD 03; 29 Oct. – 4 Nov. 2015; Mermudes and Mattos leg.; Pennsylvania trap; CEMT. – São Paulo • 1 spec.; São Miguel Arcanjo, P. E. Carlos Botelho; 24 ° 03 ′ 58 ″ S, 47 ° 59 ′ 53 ″ W; elev. 743 m; 6 Mar. 2012; M. Boutefeu leg.; Tapirus feces; CEMT • 1 spec.; same data as for preceding; 24 ° 03 ′ 59 ″ S, 47 ° 59 ′ 39 ″ W; elev. 580 m; 8 Apr. 2012; CEMT • 1 spec.; same data as for preceding; 4 ° 40 ′ 01 ″ S, 47 ° 59 ′ 40 ″ W; elev. 595 m; 9 Apr. 2012; CEMT • 1 spec.; same data as for preceding; 24 ° 03 ′ 50 ″ S, 47 ° 59 ′ 23 ″ W; elev. 747 m; CEMT • 4 specs; same data as for preceding; 11 Jun. 2012; CEMT • 1 spec.; same data as for preceding; 13 Jun. 2012; CEMT • 1 spec.; same data as for preceding; 15 May 2012; CEMT. – Sergipe • 2 specs; São Cristóvão; 24 Aug. 2016; M. F. Santos leg.; CEMT. COLOMBIA – Antioquia • 3 specs; Río Negro, Capiro; Abr. 1995; F. Vallejo leg.; CEMT. – Cundinamarca • 1 spec.; La Calera; Jul. 2000; J. Noriega leg.; CEMT • 1 spec.; Bogotá; Sep. 1996; J. Noriega leg.; CEMT • 1 spec.; same data as for preceding; Sep. 1998; CEMT • 1 spec.; Nocaima; 5 ° 04 ′ 16 ″ N, 74 ° 22 ′ 61 ″ W; elev. 1100 m; Sep. 2005; J. Noriega leg.; culture of sugar cane; CEMT. – Meta • 1 spec.; San. Martín, Tocancipa; elev. 330 m; Abr. 2006; P. López leg.; sabama forest; CEMT • 1 spec.; same data as for preceding; N. Montoya leg.; pitfall; CEMT. ECUADOR – Chimborazo • 1 spec.; Huigra, Chimborazo; elev. 1850 m; 31 Dec. 1997; G. Onore leg.; CEMT • 3 specs; same data as for preceding; 1 Jan. 1998; CEMT. – El Oro • 23 specs; Piñas; elev. 1250 m; 1 Sep. 1997; C. Carpio leg.; cow feces; CEMT • 5 specs; same data as for preceding; 2 Sep. 1997; human feces CEMT. – Loja • 1 spec.; Río Catamayo; elev. 1500 m; 29 Aug. 1997; C. Carpio leg.; CEMT. – Morona-Santiago • 2 specs; Bosque Domono; elev. 1650 m; 22 Jul. 1997; C. Carpio leg.; CEMT • 1 spec.; Vía Plan Milagro-Gral Gutiérrez Km 5; elev. 1500 m; 19 Aug. 1997; C. Carpio leg.; CEMT • 4 specs; same data as for preceding; feces; CEMT • 3 specs; Yunganza; elev. 1500 m; 28 Aug. 1997; C. Carpio leg.; cow feces; CEMT • 1 spec.; Palmeras; elev. 1800 m; 19 Nov. 1994; P. Guardenas leg.; forest; CEMT. – Napo • 4 specs; Cosanga; 0 ° 35 ′ 36 ″ S, 77 ° 56 ′ 18 ″ W; elev. 1750 m; 25 Mar. 1996; T. Jaramillo leg.; CEMT • 1 spec.; Vía Hollín-Loreto Km 3; elev. 1100 m; 6 Dec. 1987; A. Cordova leg.; CEMT • 1 spec.; same data as for preceding; Km 25; elev. 1100 m; P. Mendoza leg.; CEMT • 2 specs; same data as for preceding, Km 3; A. Cordova leg.; CEMT • 1 spec.; same data as for preceding; Km 60; R. Figuerda leg.; CEMT • 1 spec.; same data as for preceding; Km 17; M. Peñaherrera leg.; CEMT. – Pichincha • 3 specs; Alangasi; 19 Jan. 1988; M. E. Sánchez leg.; CEMT • 2 specs; El Tingo; 5 May 1997; P. Leone leg.; CEMT • 1 spec.; same data as for preceding; 30 Jan. 1988; CEMT • 1 spec.; Mindo; elev. 1500 m; 30 Dec. 1994; P. Salazar leg.; CEMT • 2 specs; same data as for preceding; elev. 1400 m; 8 Jun. 1995; F. Sancho leg.; CEMT • 1 spec.; Quito, Alangasi; 19 Jan. 1988; P. Merdoxa leg.; CEMT • 1 spec.; Río Silanchi; elev. 640 m; 30 Apr. 1991; E. Baus leg.; CEMT • 5 specs; same data as for preceding; Mindo; elev. 1200 m; 17 Jun. 1995; M. Lascano leg.; CEMT • 3 specs; same data as for preceding; elev. 1250 m; 8 Jul. 1995; M. Vallejo leg.; CEMT • 1 spec.; same data as for preceding; 10 Jul. 1995; CEMT • 1 spec.; Pam. Argentin; elev. 1500 m; Apr. 1988; M. Griialua leg.; CEMT. – Zamora • 6 specs; Ch Las Peñas; elev. 1000 m; 24 Aug. 1997; C. Carpio leg.; CEMT. PERU – Madre de Dios • 1 spec.; 6 km NE of Mazuko; elev. 380 m; 19 Aug. 2012; Cavichioli et al. leg.; CEMT. VENEZUELA – Mérida • 2 specs; Bailadores; Jan. 1984; B. Joffre leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF93FFCAFE23F993FAF8FA03.taxon	description	Sexual dimorphism Males with clypeal tubercle larger than that of female; females with metaventrite plate nearly flat.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF93FFCAFE23F993FAF8FA03.taxon	discussion	Collecting information This species can be collected with light traps and is attracted to equine, bovine, Tapirus and human feces (Mesquita et al. 2017). Specimens were collected at altitudes ranging from 33 to 1850 metres.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9DFFCAFDE3FA4AFE3DF979.taxon	diagnosis	Diagnosis Length 2.5 – 6.0 mm. Body oval, elongate, testaceous. Head punctate, clypeus faintly sinuate; frontoclypeal suture finely impressed. Pronotum transverse, punctate; laterally with marginal marginal line; posteriorly with or without marginal line, marginal line might be medially interrupted. Elytra without humeral teeth.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9DFFC9FDDCF96CFEFDFBEE.taxon	description	Figs 7 A, 38 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9DFFC9FDDCF96CFEFDFBEE.taxon	diagnosis	Diagnosis Length 6.0 mm. Body convex, almost glabrous. Head punctate with clypeus truncate, longitudinally gibbous medially; frontoclypeal suture finely impressed. Pronotum convex, without posterior marginal line with punctures of two sizes; larger punctures concentrated on sides, smaller finer punctures densely distributed across surface. Elytra strongly alutaceous with lighter spot at posterior declivity; interstriae weakly convex, alutaceous, finely punctured. Metatibiae with apical and lateral margin fringed by setae progressively longer closer to lateral margin.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9DFFC9FDDCF96CFEFDFBEE.taxon	materials_examined	Material examined BRAZIL – Paraná • 3 specs; São José dos Linhares, Ser. Mar Br 277, Km 54; 31 Dec. 1986; light; CEMT • 2 specs; same data as for preceding; 30 Jan. 1986; light; CEMT. – Rio Grande do Sul • 1 spec.; São Francisco de Paula, Promata; 14 – 18 Dec. 2009; Grossi et al. leg.; CEMT. Collecting information Specimens were collected at light. Sexual dimorphism Male unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9DFFC9FDDCF96CFEFDFBEE.taxon	discussion	Remarks Trichaphodius is possibly restricted to the Old World (Dellacasa et al. 2001 a). Trichaphodius quadripartitus may be based on a mislabeled specimen (Skelley et al. 2007). Until further studies are conducted with the type material, the species is provisionally retained in Trichaphodius (Skelley et al. 2007).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9EFFC9FDE0FB59FA82F853.taxon	diagnosis	Diagnosis Length 1.6 – 1.9 mm. Body oblong-oval, strongly convex, piceous. Head with clypeus denticulate or obtusely rounded angled on each side of medial emargination, anterior margin of clypeus thickened on emargination. Pronotum variably punctate, not fringed by setae. Meso- and metatibia two with distinct transverse carinae and with lateral apical margin with three dentiform processes.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9EFFC9FDE0FB59FA82F853.taxon	discussion	Remarks Aphotaenius currently has 8 described species distributed across the eastern USA, Brazil, Bolivia, Colombia, Mexico and Venezuela (Skelley & Vaz-de-Mello 2020). For a key to all species of Aphotaenius see Skelley & Vaz-de-Mello (2020).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFC8FE38FE95FEA3FC0D.taxon	description	Figs 20 D, 38 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFC8FE38FE95FEA3FC0D.taxon	diagnosis	Diagnosis Length 1.6 – 2.7 mm. Head with clypeus emarginate, angles lateral to emargination denticulate; punctures of frons larger than those of clypeus. Pronotum with punctures of two sizes uniformly distributed across its surface. Elytral interstriae with two marginal rows of small punctures.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFC8FE38FE95FEA3FC0D.taxon	materials_examined	Material examined BRAZIL – Maranhão • 1 spec.; Carolina, Fazenda Cincorá; 17 – 22 Oct. 2009; F. Limeira-de-Oliveira et al. leg.; light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFC8FE38FE95FEA3FC0D.taxon	description	Sexual dimorphism Males with penultimate ventrite of abdomen narrower than those of females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFC8FE38FE95FEA3FC0D.taxon	discussion	Collecting information Specimens were collected associated with feces of Mazama sp. and with light traps (Skelley & Vaz-de-Mello 2020).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFC8FE39FC7EFBFBF91F.taxon	description	Figs 20 E, 38 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFC8FE39FC7EFBFBF91F.taxon	diagnosis	Diagnosis Length 2.5 mm. Head with scarce and fine punctures; lateral angles to clypeal emargination rounded. Pronotum with punctures of two sizes; small and big punctures uniformly and densely distributed across pronotal surface. Elytral interstriae with fine punctures.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFC8FE39FC7EFBFBF91F.taxon	materials_examined	Material examined BRAZIL – Rio Grande do Sul • 1 spec.; Santa Cruz do Sul, Cerro Alegre; 29 ° 48 ′ 19 ″ S, 52 ° 19 ′ 53 ″ W; 11 Feb. 2012; R. Bohn leg.; secondary forest; bovine feces; CEMT • 2 specs; same data as for preceding; 29 ° 42 ′ 17 ″ S, 52 ° 19 ′ 59 ″ W; 17 Dec. 2022; bovine feces; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFC8FE39FC7EFBFBF91F.taxon	description	Sexual dimorphism Males with penultimate ventrites of abdomen narrower than those of females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFC8FE39FC7EFBFBF91F.taxon	discussion	Collecting information Specimens are attracted to bovine feces (Skelley & Vaz-de-Mello 2020).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFCFFDFCF94EFADEFC9D.taxon	description	Figs 20 F, 38 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFCFFDFCF94EFADEFC9D.taxon	diagnosis	Diagnosis Length 2.6 – 2.7 mm. Head with large punctures; clypeus emarginate, margin obtusely angled on each side of emargination. Pronotum with punctures of two sizes; coarse punctures restricted to posterior and lateral margins; smaller punctures distributed across pronotal surface. Elytral interstriae with discrete punctures.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFCFFDFCF94EFADEFC9D.taxon	materials_examined	Material examined BRAZIL – Rio Grande do Sul • 1 spec.; Santa Cruz do Sul, Cerro Alegre; 29 ° 48 ′ 17 ″ S, 52 ° 19 ′ 29 ″ W; 24 Dec. 2011; R. Bohn leg.; forest (trap p 9); pitfall; bovine feces; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFCFFDFCF94EFADEFC9D.taxon	description	Sexual dimorphism Males with penultimate ventrite of abdomen narrower than those of females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9FFFCFFDFCF94EFADEFC9D.taxon	discussion	Collecting information Specimens were collected with pitfalls baited with bovine feces (Skelley & Vaz-de-Mello 2020).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF98FFCFFD81FCC8FB7DFB6F.taxon	diagnosis	Diagnosis Length 5.5 – 6.0 mm. Body elongate, dark reddish brown. Head wide; clypeus longitudinally wrinkled, strongly gibbous; frontoclypeal suture absent; frons punctate. Pronotum wide; with large and oval punctures; lateral margins straight, explanate and furrowed; posterior margin with deep groove. Elytra with humerus forwardly expanded, apically glabrous; fifth interstriae with anterior tubercle; fifth stria grooved anteriorly. Legs elongate and slender.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF98FFCFFD81FCC8FB7DFB6F.taxon	discussion	Remarks Arupaia is a monotypic genus currently only known from Brazil (Stebnicka 2009).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF98FFCEFDE7FA9EFE2CFE51.taxon	description	Figs 14 A – C, 39 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF98FFCEFDE7FA9EFE2CFE51.taxon	diagnosis	Diagnosis Same as for the genus.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF98FFCEFDE7FA9EFE2CFE51.taxon	materials_examined	Material examined BRAZIL – Distrito Federal • 1 spec.; Planaltina, Embrapa Cerrados; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 18 Nov. 2005; C. Oliveira leg.; área agrícola; light; CEMT • 1 spec.; same data as for preceding; 2 Nov. 2005; CEMT. – Minas Gerais • 2 specs; Ijaci, Fazenda FAEPE; Nov. 2002; Louzada leg.; light; CEMT • 1 spec.; Viçosa; 28 Mar. 2023; E. J. Hambleton leg.; CEMT. – Rio de Janeiro • 1 spec.; Nova Friburgo; Jan. 1999; P. Grossi; CEMT • 1 spec.; same data as for preceding; Jan. 2002; E. Grossi and P. Grossi leg.; CEMT. – Rio Grande do Sul • 1 spec.; Bagé; 31 ° 18 ′ 57 ″ S, 53 ° 59 ′ 53 ″ W; elev. 242 m; 21 Feb. 2017; R. N. Sisti and J. U. P. Corrêa leg.; glycine; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF98FFCEFDE7FA9EFE2CFE51.taxon	description	Sexual dimorphism Males with apical spur acuminate, big, inwardly bent, hook like.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF98FFCEFDE7FA9EFE2CFE51.taxon	discussion	Collecting information Specimens were collected with UV light and in nests of Solenopsis geminata (Fabricius, 1804) (Stebnicka 1999 add letter, 2007 a).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF99FFCEFD91FD97FA9BFA6B.taxon	diagnosis	Diagnosis Length 2.8 – 4.0 mm. Body elongate, subparallel-sided, dorsally glabrous, glossy, light reddish testaceous to piceous. Head with transverse wrinkles weakly developed; clypeus with anterior margin slightly sinuate, angles delimiting sinuate area denticulate or obtuse; frons with coarse punctures variably spaced. Pronotum with punctures of two sizes, variably distributed. Elytra glabrous; striae and interstriae variably punctate. Profemur without anterior and posterior lines; anterior margin ends in a translucent process. Abdominal ventrites glossy, strigate or not. Pygidium near anterior margin, near uro-genital opening, glossy and convex.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF99FFCEFD91FD97FA9BFA6B.taxon	discussion	Remarks Ataeniopsis comprises 15 species distributed across the Neartic and Neotropical regions (Stebnicka 2003 b). For a key to all species see Stebnicka (2003 b).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF99FFCDFE33FA62FE7AFBB5.taxon	description	Figs 8 C – D, 39 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF99FFCDFE33FA62FE7AFBB5.taxon	diagnosis	Diagnosis Length 4.2 – 4.8 mm. Body piceous. Head with clypeus denticulate; weakly emarginate medially; emarginate area with nearly half the length of frons; frontoclypeal suture not visible. Pronotum with posterior marginal line preset; punctures of two sizes, fine and large; fine punctures inconspicuous among the large ones; large punctures closer near lateral margins. Elytra glabrous, glossy; length more than three times that of pronotum. Pygidium convex, glossy with fine punctures.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF99FFCDFE33FA62FE7AFBB5.taxon	materials_examined	Material examined ARGENTINA – Chaco • 1 spec.; Parque Nacional Chaco; 12 Jan. 2008; CEMT. – Mendoza • 1 spec.; Res. Ñacuñán; Mar. 2002; C. A. Medina leg.; CEMT • 3 specs; Mendoza-San Luis, Desaguadero; 11 Jan. 2005; C. Fortino leg.; light trap; CEMT • 10 specs; Desaguadero; 5 Feb. 2017; G. Arriágada leg.; light; CEMT • 68 specs; Lavalle, Res. Bio. Telteca; 9 Dec. 2002; Arriágada and Diéguez leg.; light trap; CEMT • 1 spec.; Res. Biológica Ñacuñán; Feb. 2012; G. Arriágada leg.; light; CEMT. – San Luis • 4 specs; Belgrano, Fundo El Molle; 33 ° 2 ′ 23 ″ S, 66 ° 30 ′ 48 ″ W; elev. 622 m; 7 Feb. 2017; G. Arriágada leg.; CEMT. BRAZIL – Mato Grosso • 1 spec.; Poconé; 23 Jun. 1998; acuri palm tree; light trap; CEMT • 3 specs; Sesc Pantanal; 15 Sep. 2004; light trap; CEMT • 1 spec.; Vila Bela, Margem Guporé; 1 Oct. 1984; Sebastião and Marcolina leg.; light; CEMT. – Mato Grosso do Sul • 1 spec.; Aquidauana; S. R. Rodrigues leg.; CEMT • 2 specs; Guia Lopes da Laguna; 25 May 2008; A. Abot leg.; light trap; CEMT. – Piauí • 2 specs; Teresina; Nov. 1952 – Jan. 1953; A. K. Oliveira leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF99FFCDFE33FA62FE7AFBB5.taxon	description	Sexual dimorphism Males with third abdominal ventrite longer than fourth. Females with body longer than males, with second and fourth ventrite with the same length.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF99FFCDFE33FA62FE7AFBB5.taxon	discussion	Collecting information Specimens are attracted to light and can be found in flooded plains and seasonal subtropical forests (Stebnicka 2009).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9AFFCDFDE2FBB3FBBFF839.taxon	description	Figs 7 D – F, 39 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9AFFCDFDE2FBB3FBBFF839.taxon	diagnosis	Diagnosis Length 3.2 – 4.8 mm. Body reddish testaceous. Head alutaceous; clypeus subtly, but rather widely sinuate; frontoclypeal suture inconspicuous. Pronotum without posterior marginal line; punctate with punctures of two sizes irregularly spaced. Elytra twice as long as pronotum. Abdominal ventrites with extremely small punctures and weakly strigate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9AFFCDFDE2FBB3FBBFF839.taxon	materials_examined	Material examined BRAZIL – Piauí • 4 specs; Teresina, Embrapa CPAMN; 12 Nov. 2011; M. Bevilaqua and R. Querino leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9AFFCDFDE2FBB3FBBFF839.taxon	description	Sexual dimorphism Males with third abdominal ventrite wider than fourth. Females with body longer than males, with second and fourth ventrite of the same width.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9AFFCDFDE2FBB3FBBFF839.taxon	discussion	Collecting information Collecting methodology not specified in the literature and material examined.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9BFFCCFDF0FE94FD57FB3A.taxon	description	Figs 8 A – B, 39 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9BFFCCFDF0FE94FD57FB3A.taxon	diagnosis	Diagnosis Length 2.8 – 3.0 mm. Body dark red. Head alutaceous, clypeus slightly sinuate with somewhat large denticles; sinuate area less than 1 / 3 the length of frons; frontoclypeal suture inconspicuous, present as a faint elevated line. Pronotum densely punctate with large and moderate punctures; posteriorly marginate. Elytra length approximately twice as long as pronotum; interstriae punctured by fine and inconspicuous punctures. Pygidium with three to four fine punctures along median carina; anterior margin glossy and convex.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9BFFCCFDF0FE94FD57FB3A.taxon	materials_examined	Material examined BRAZIL – Amazonas • Castanho-Careiro, AM- 359 Km 39; 3 ° 43 ′ 59 ″ S, 60 ° 20 ′ 09 ″ W; 6 – 7 Nov. 2010; Rafael et al. leg.; light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9BFFCCFDF0FE94FD57FB3A.taxon	description	Sexual dimorphism Males with third abdominal ventrite wider than fourth. Females with body longer than males, with second and fourth ventrite of the same width.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9BFFCCFDF0FE94FD57FB3A.taxon	discussion	Collecting information Specimens were collected with light trap.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9BFFB2FDE5FB32FC93FD64.taxon	description	Figs 8 E – F, 39 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9BFFB2FDE5FB32FC93FD64.taxon	diagnosis	Diagnosis Length 3.2 – 4.0 mm. Body piceous. Head alutaceous, clypeus with two minute denticles; weakly emarginate medially; emarginate area with approximately half the length of frons. Pronotum with punctures of two sizes, large and minute; large punctures everywhere distributed, intermixed with inconspicuously minute ones; pronotum with posterior marginal line. Elytral length approximately twice that of pronotum; interstriae with small, irregularly spaced punctures. Pygidium disc eroded by group of large coalescent punctures.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9BFFB2FDE5FB32FC93FD64.taxon	materials_examined	Material examined BRAZIL – Mato Grosso • 1 spec.; Trindade, U. B. S.; 1 Oct. 1984; E. Binda leg.; light; CEMT • 1 spec.; same data as for preceding; INPA. – Roraima • 5 specs; Alto Alegre, Lavrado; 2 ° 53 ′ 45 ″ N, 60 ° 39 ′ 42 ″ W; elev. 81 m; 5 May 2015; E. G. F. Morais leg.; CEMT • 13 specs; Alto Alegre, Rio Cauamé, Fazenda Jarbas Melo; 3 ° 13 ′ 34 ″ N, 61 ° 09 ′ 54 ″ W; 16 Oct. 2020; R. C. L. Costa and R. B. Silva leg.; light; CEMT • 1 specs. Amajari, EsEc Maracá, Uraricoera River; 3 ° 20 ′ 59 ″ N, 61 ° 25 ′ 22 ″ W; elev. 83 m; 24 Mar. 2016; A. P. M. Santos et al. leg.; CEMT • 3 specs; Ilha de Maracá, Rio Uraricoera, Ilha de Maraca; 2 – 13 May 1987; J. A. Rafael et al. leg.; light trap; CEMT • 19 specs; same date as for preceding; INPA • 25 specs; same data as for preceding; armadilha suspensa [suspended trap]; CEMT INPA • 3 specs; same data as for preceding; 21 – 30 Nov. 1987; Malaise trap; INPA.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9BFFB2FDE5FB32FC93FD64.taxon	description	Sexual dimorphism Males with third abdominal ventrite wider than fourth. Females with body longer than males, with second and fourth ventrite of the same width.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF9BFFB2FDE5FB32FC93FD64.taxon	discussion	Collecting information Specimens were collected with UV light traps in open areas, near ponds and coconut groves (Stebnicka 2009). Specimens were found at altitudes up to 81 metres.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFE5FFB1FDB6FD63FA83F971.taxon	diagnosis	Diagnosis Length 2.0 – 7.0 mm. Body elongate, cylindrical, glabrous or not, light reddish testaceous to piceous. Head variably sculptured, may present wrinkles, punctures or argillaceous coating; clypeus always sinuate medially, margin may be rounded, denticulate or obtuse. Pronotum transverse, glossy, shagreened or with argillaceous coating, variably punctate. Elytra variable, interstriae may be glossy or opaque, glabrous or pilose, convex, carinate or flat, tuberculate or not, shagreened or with argillaceous coating.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFE5FFB1FDB6FD63FA83F971.taxon	discussion	Remarks Ataenius is a cosmopolitan genus comprised of more than 300 species (Stebnicka 2007 b). Stebnicka (2007 b) presents dorsal habitus images of 187 of the 190 species of Ataenius the New World. For more information on the morphological diversity in Ataenius and keys for almost all of the Western Hemisphere species the following publications are recommended: Stebnicka (2001 c, 2003 e, 2004, 2005 b, 2006 b, 2007 b, 2007 c) and Stebnicka & Lago (2005).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFE6FFB0FD93F974FAE2FE3F.taxon	description	Figs 9 A – D, 40 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFE6FFB0FD93F974FAE2FE3F.taxon	diagnosis	Diagnosis Length 2.1 – 6.0 mm. Body cylindrical or oval, weakly convex, glossy, or shagreened, piceous. Head variably punctate; clypeus sinuate, lateral angles rounded or obtuse. Pronotum punctate, puncture sizes variable, normally with two distinct sizes; lateral margins normally fringed with evident setae. Elytra with striae not deep, finely punctate; interstriae glossy, flat or convex. Ventral surface with setaceous regions, mostly glabrous and glossy. Apical accessory spine of metatibiae fine or absent, if present, always shorter than apical fringe of setae. Abdominal ventrites punctate and weakly strigate anteriorly; punctures coarse, normally near lateral margins, but these can be found across ventrite surface.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFE7FFB7FF61FE7FFC4BFA10.taxon	materials_examined	BRAZIL – Acre • 2 specs; Rio Branco; Nov. 2000; M. A. Oliveira leg.; CEMT. – Amazonas • 1 spec.; Itacoatiara Fazenda Aruana, Am 010, Km 215; 12 – 15 Jan. 1991; C. S. Motta leg.; UV light; INPA. – Bahia • 13 specs; Itamarajú; 16 ° 59 ′ 27 ″ S, 39 ° 25 ′ 55 ″ W; Abr. 2011; C. M. P. Leite leg.; pitfall; CEMT • 1 spec.; same data as for preceding; 17 ° 01 ′ 09 ″ S, 39 ° 28 ′ 15 ″ W; CEMT. – Distrito Federal • 1 spec.; Brasília, Fazenda Água Limpa; Nov. 2010; C. Suinaga leg.; CEMT. – Espírito Santo • 1 spec.; Venda Nova; Dec. 1999; F. Z. Vaz-de-Mello leg.; CEMT. – Goiás • 1 spec.; Rio Verde; Nov. 1995; J. Carlos leg.; CEMT. – Mato Grosso • 1 spec.; Res. Papagaio, BR- 364 Km 555; 1 Oct. 1984; E. Binda leg.; polo noroeste [northwest pole]; light; INPA • 3 specs; Água Boa, Rancho Araguaia; 13 ° 56 ′ 40 ″ S, 52 ° 6 ′ 10 ″ W; 24 Dec. 2017; Mota leg.; CEMT • 10 specs; Alta Floresta, CEPLAC; 4 Sep. 2012; V. Gonçalvez leg.; human feces; CEMT • 4 specs; same data as for preceding; 4 Jan. 2009; Tapirus feces; CEMT • 2 specs; same data as for preceding; 7 Aug. 2009; CEMT • 1 spec.; Alta Floresta; Chácara Recanto das Orquídeas; 3 Oct. 2011; A. B. Petini leg.; CEMT • 8 specs; Chapada dos Guimarães; 11 Dec. 2000; A. F. Ramos leg.; light trap; CEMT • 1 spec.; same data as for preceding; 25 Feb. 2001; CEMT • 1 spec.; same data as for preceding; 15 ° 24 ′ 31 ″ S, 55 ° 45 ′ 53 ″ W; 2 Dec. 2015; A. Frolov and L. Akhmetova leg.; light; CEMT • 1 spec.; same data as for preceding; Casa do Mel; 15 ° 22 ′ 53 ″ S, 55 ° 50 ′ 35 ″ W; elev. 520 m; Dec. 2015; R. V. Nunes et al. leg.; light; CEMT • 2 specs; Cotriguaçu, Fazenda São Nicolau; 14 Feb. 2007; O. Peres-Filho leg.; CEMT • 22 specs; same data as for preceding; 15 Dec. 2007; light trap; CEMT • 3 specs; same data as for preceding; 16 Oct. 2007; CEMT • 15 specs; same data as for preceding; 31 Oct. 2007; CEMT • 2 specs; same data as for preceding; 14 Nov. 2007; CEMT • 6 specs; same data as for preceding; 15 Dec. 2007; CEMT • 5 specs; same data as for preceding; Nov. 2007 • 1 spec.; same data as for preceding; 14 Apr. 2008 • CEMT • 1 spec.; same data as for preceding; 9 ° 49 ′ 32 ″ S, 58 ° 15 ′ 46 ″ W; elev. 220 m; 10 Dec. 2009; Vaz-de-Mello leg.; Ficus plantation; CEMT • 1 spec.; same data as for preceding; 9 ° 49 ′ 09 ″ S, 58 ° 15 ′ 47 ″ W; CEMT • 5 specs; same data as for preceding; 9 ° 50 ′ 19 ″ S, 58 ° 15 ′ 03 ″ W; 25 Oct. 2014; A. Asenjo leg.; light; CEMT • 1 spec.; same data as for preceding; 28 Oct. 2014; UV light; CEMT • 1 spec.; same data as for preceding; 9 ° 51 ′ 18 ″ S, 58 ° 14 ′ 57 ″ W; elev. 220 m; base camp; 14 Dec. 2009; Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; 9 ° 51 ′ 20 ″ S, 58 ° 14 ′ 55 ″ W; Nov. 2014; R. V. Nunes and A. Asenjo leg.; light; CEMT • 2 specs; same data as for preceding; 30 Oct. 2014; W. Chamorro leg.; black light trap; CEMT • 1 spec.; Cuiabá; 15 ° 35 ′ 21 ″ S, 56 ° 01 ′ 46 ″ W; elev. 183 m; Sep. 2015; R. V. Nunes and B. F. Becker leg.; CEMT • 7 specs; same data as for preceding; Recanto Siriema; 15 ° 35 ′ 32 ″ S, 56 ° 01 ′ 38 ″ W; 4 Nov. 2014; light; CEMT • 1 spec.; Diamantino; Jan. 2001; A. M. Bello leg.; CEMT • 1 spec.; same data as for preceding; Reserve Vale da Solidão; 14 ° 21 ′ 29 ″ S, 56 ° 08 ′ 05 ″ W; 31 Jan. 2009; D. C. T. Oliveira leg.; CEMT • 1 spec.; Alto Rio Arinos; Jan. 2001; E. Furtado leg.; CEMT • 1 spec.; Farinópolis, Fazenda Sudan; 15 ° 15 ′ 14 ″ S, 58 ° 27 ′ 20 ″ W; Jul. 2004; M. Santos-Filhos leg.; watershed Jauru; CEMT • 1 spec.; Porto Estrela, ESEC Serra das Araras; 15 ° 39 ′ 20 ″ S, 57 ° 12 ′ 51 ″ W; elev. 225 m; 12 Oct. 2011; F. Z. Vaz-de-Mello leg.; Cerrado; pitfall; CEMT • 2 specs; same data as for preceding; helipad trail; CEMT • 20 specs; same data as for preceding; Parque Boca do José; 15 Oct. 2011; final trail; Cerrado; CEMT • 1 spec.; same data as for preceding; Estação Ecológica Cerra das Aráras; Nov. 2011; Souza and Gigliotti leg.; CEMT • 1 spec.; Santo Antônio do Leverger, São Vincente da Serra; 15 ° 49 ′ 42 ″ S, 55 ° 25 ′ 11 ″ W; 15 – 17 Dec. 2010; Tissiani and Vaz-de-Mello leg.; pasture; human dung; CEMT • 4 specs; Tangará da Serra; 14 ° 38 ′ 33 ″ S, 57 ° 33 ′ 18 ″ W; 19 – 22 Feb. 2012; A. S. Tissiani leg.; CEMT • 7 specs; same data as for preceding; Fazenda Bahía; 14 ° 37 ′ 13 ″ S, 57 ° 24 ′ 50 ″ W; elev. 428 m; 26 – 28 Jan. 2012; R. J. Silva leg.; semidecidual forest; pitfall-hum-pig dung; CEMT • 3 specs; same data as for preceding; 14 ° 37 ′ 14 ″ S, 57 ° 25 ′ 15 ″ W; elev. 419 m; 12 – 14 Jan. 2011; pasture; CEMT • 1 spec.; same data as for preceding; 14 ° 37 ′ 4 ″ S, 57 ° 25 ′ 0 ″ W; elev. 428 m; 26 – 28 Jan. 2012; CEMT. – Mato Grosso do Sul • 1 spec.; Corumbá, Fazenda São Judas, Rio Vermelho; 19 ° 36 ′ 43 ″ S, 56 ° 57 ′ 50 ″ W; Jun. 1999; L. Vieira leg.; CEMT • 1 spec.; Corumbá, RPPN Acurizal; 17 ° 50 ′ 13 ″ S, 57 ° 33 ′ 03 ″ W; 24 – 26 Jul. 2012; F. R. Tortato leg.; CEMT • 1 spec.; Poconé, Bom Pastor; 15 Dec. 2011; J. Figueiredo leg.; manually; CEMT. – Minas Gerais • 1 spec.; Águas Vermelhas; Dec. 1997; A. Bello leg.; CEMT • 1 spec.; same data as for preceding; A. Bello and F. Z. Vaz-de-Mello leg.; CEMT • 2 specs; Bocaiuva, Fazenda Corredor, Arredores do Alojamento; 5 Nov. 1998; M. F. Vasconcelos and Azevedo leg.; CEMT • 2 specs; Cordisburgo, Fazenda Pontinha; Jul. 1994; de Mello leg.; CEMT • 4 specs; Ijací, Faz FAEPE; Nov. 2002; J. Louzada leg.; CEMT • 1 spec.; Ipatinga; Nov. 1991; E. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Sep. 1993; CEMT • 1 spec.; Lavras, Próx Poço Bonito; Dec. 2001; Vaz-de-Mello leg.; CEMT • 3 specs; Paracatu; Mar. 1997; Vaz-de-Mello leg.; CEMT • 4 specs; Recreio; Jan. 2000; Lopes-Andrade and Vaz-de-Mello leg.; CEMT • 6 specs; Três Marias; Jan. 1993; CEMT • 1 spec.; Uberlândia, Campus UFU; 28 Sep. 2005; J. Mendes leg.; CEMT • 1 spec.; Vespasioano; Nov. 1963; A. Machado leg.; CEMT • 3 specs; Viçosa, Campus UFV; Nov. 1997; F. Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; 1994 – 1995; Louzada et al. leg.; pasture; CEMT. – Pará • 2 specs. Santarém; fruit culture; CEMT • 1 spec.; Vigia, Fazenda Mangue Seco; 20 – 22 Jun. 2010; I. S. Araújo and N. Moura leg.; CEMT. – Paraná • 2 specs; Entre Rio do Oeste; 26 Dec. 2017; M. E. Maldaner leg.; light; CEMT. – Pernambuco • 1 spec.; Salgueiro; May 2012; R. V. Nunes leg.; CEMT • 1 spec.; Miguel Pereira; Jan. 1998; J. Carlos leg.; CEMT • 1 spec.; same data as for preceding; Dec. 2000; CEMT. – Rondônia • 1 spec.; Cacaulândia, Fazenda Rancho Grande; Feb. 1997; Vaz-de-Mello leg.; CEMT • 1 spec.; Porto Velho, Rio das Garças, Bom Jesus; 8 ° 49 ′ 48 ″ S, 63 ° 46 ′ 45 ″ W; 11 Apr. 2017; D. C. Santos and K. K. G. Silva leg.; FIT; CEMT. – Roraima • 1 spec.; Rio Uraricoera, Ilha de Maraca; 2 – 13 May 1987; J. A. Rafael et al. leg.; suspended trap; CEMT • 1 spec.; same data as for preceding; INPA • 1 spec.; same data as for preceding; 30 Nov. 1987; Rafael et al. leg; Malaise trap; INPA. MEXICO – Chiapas • 1 spec.; Palenque; Jun. 2004; F. Nicolalde leg.; CEMT. – Veracruz • 3 specs; Los Tuxlas; Oct. 1997; F. Z. Vaz-de-Mello leg.; CEMT. NICARAGUA – Masaya • 1 spec.; Las Flores; J. M. Maes leg.; light trap; CEMT. – Tachira • 1 spec.; San Critóbal, Barrio El Lobo; 7 Aug. 1984; B. Joffre leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFE0FFB7FF61FA57FD25F8BA.taxon	materials_examined	BRAZIL – • 1 spec.; Xingu; Oct. 1947; CEMT. – Mato Grosso • 1 spec.; Cuiabá, Recanto Siriema; 15 ° 35 ′ 32 ″ S, 56 ° 01 ′ 38 ″ W; 4 Nov. 2014; R. V. Nunes and B. F. Becker leg.; light; CEMT. – Mato Grosso do Sul • 1 spec.; Aquidauana; S. R. Rodrigues leg.; CEMT. – Minas Gerais • 1 spec.; Águas Vermelhas; Dec. 1998; A. Bello and F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Itumirim; 21 ° 20 ′ 24 ″ S, 44 ° 48 ′ 06 ″ W; 19 Oct. 2016; Vierra et al. leg.; light; CEMT. – São Paulo • 1 spec.; Barueri; 29 Feb. 1956; K. Lenko leg.; CEMT • 1 spec.; Cerqueira César; Feb. 1995; J. Carlos leg.; CEMT • 1 spec.; Piracicaba; 30 Dec. 1966; C. A. Triplehorn leg.; UV light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFE0FFB7FF61F8ADFC2AF80C.taxon	materials_examined	BRAZIL – São Paulo • 1 spec.; Agudos, Duraflora S. A.; 12 Oct. 1993; C. A. H. Flechtmann leg.; Pinus caribaensis var. bahamensis stand; control-tent trap (blank); CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFE1FFB6FF61FE95FB17FD4B.taxon	materials_examined	BRAZIL – Bahia • 1 spec.; Presidente Tancredo Neves; 13 ° 22 ′ S, 39 ° 19 ′ W; CEMT. – Distrito Federal • 2 specs; Brasilia; elev. 1100 m; Mar. 2001; N. Degallier leg.; light; CEMT. – Goiás • 1 spec.; Rio Verde; Nov. 1995; J. Carlos leg.; CEMT. – Mato Grosso • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 9 ° 50 ′ 19 ″ S, 58 ° 15 ′ 03 ″ W; 25 Oct. 2014; A. Asenjo leg.; light; CEMT • 2 specs; same data as for preceding; UV light; CEMT • 1 spec.; Cuiabá, Campus UFMT; Nov. 2009; R. Miyazaki leg.; CEMT • 1 spec.; Novo Mundo, P. E. Cristalino; 9 ° 27 ′ 46 ″ S, 55 ° 48 ′ 58 ″ W; Nov. 2012; V. Magalhães leg.; pitfall; CEMT • 2 specs; Santo Antônio do Leverger, Águas quentes; 23 Nov. 2016; A. Frolov and M. Cupello leg.; UV light; CEMT • 1 spec.; São Vincente da Serra, Campus UFMT; 15 ° 49 ′ 42 ″ S, 55 ° 25 ′ 11 ″ W; elev. 900 m; 21 – 23 May 2012; Tissiani and Vaz-de-Mello leg.; human dung; CEMT • 1 spec.; Tangará da Serra; 14 ° 38 ′ 33 ″ S, 57 ° 33 ′ 18 ″ W; 19 – 22 Feb. 2012; A. S. Tissiani leg.; CEMT. – Pernambuco • 2 specs; Igarassu-RECD; 11 May 2007; Silva et al. leg.; CEMT. – Rondônia • 4 specs; 62 km SW of Ariquemes nr Fznd Rancho Grande; 8 – 20 Nov. 1994; J. Eger and C. O’Brien leg.; black light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFE1FFB4FF61FC82FB7AFBB2.taxon	materials_examined	BRAZIL – Bahia • 1 spec.; Barreiras; Oct. 1991; light trap; CEMT • 1 spec.; Ubaíra; 13 ° 8 ′ 47 ″ S, 39 ° 40 ′ 34 ″ W; Nov. 2011; C. M. P. Leite leg.; pitfall; CEMT. – Distrito Federal • 2 specs; Brasília; elev. 1100 m; Jan. 2001; N. Degallier leg.; CEMT • 1 spec.; same data as for preceding; Feb. 2001; light; CEMT • 2 specs; same data as for preceding; Jan. 2003; CEMT • 2 specs; Planaltina, Fazenda Sem Três Pinheiros; 15 ° 36 ′ 38 ″ S, 47 ° 30 ′ 16 ″ W; 9 Dec. 2017; Y. Ferreira leg.; soybean plantation; human feces; pitfall; CEMT. – Goiás • 5 specs; Niquelândia; Oct. 1993; light trap; CEMT • 1 spec.; Rio Verde; Nov. 1995; J. Carlos leg.; CEMT • 3 specs; same data as for preceding; Jun. 1997; CEMT. – Maranhão • 2 specs; Caxias, Reserva Ecol. Inhamun, Povoado Coités; 4 ° 54 ′ 43 ″ S, 43 ° 25 ′ 30 ″ W; 28 – 29 Dec. 2012; M. R. A. Vieira-Neta leg.; pitfall; CEMT • 2 specs; Mirador, P. E. Mirador, Base Geraldina; 20 – 24 Dec. 2006; R. O. Souza and J. C. Silva leg.; light; CEMT • 1 spec.; same data as for preceding; 27 Nov. 2008; M. B. Aguiar-Nelto and A. L. Costa leg.; light; CEMT. – Mato Grosso • 3 specs; Chapada dos Guimarães; 12 Dec. 2000; A. F. Ramos leg.; light trap; CEMT • 2 specs; same data as for preceding; 15 ° 24 ′ 31 ″ S, 55 ° 45 ′ 53 ″ W; 2 Dec. 2015; A. Frolov and L. Akhmetova leg.; light; CEMT • 1 spec.; Casa do Mel; 15 ° 22 ′ 53 ″ S, 55 ° 50 ′ 35 ″ W; elev. 520 m; Dec. 2015; R. V. Nunes et al. leg.; CEMT • 1 spec.; Fazenda Jardim; 2 Feb. 2013; V. C. Jorge leg.; CEMT • 1 spec.; same data as for preceding; 10 Nov. 2012; CEMT • 4 specs; same data as for preceding; 13 Oct. 2012; CEMT • 1 spec.; same data as for preceding; 26 May 2012; CEMT • 1 spec.; Fazenda Jardim; 28 Apr. 2012; CEMT • 5 specs; same data as for preceding; 29 Sep. 2012; CEMT • 1 spec.; same data as for preceding; Fazenda Morro das Lajes; 10 Apr. 1983; S. Marcolino leg.; CEMT • 1 spec.; Fazenda Toa-Toa; 15 ° 20 ′ 31 ″ S, 55 ° 51 ′ 17 ″ W; elev. 300 m; 20 Dec. 2012; M. D. Gimo leg.; pitfall; CEMT • 1 spec.; same data as for preceding; Sep. 2011; R. V. Nunes leg.; pasto (próx mirante); CEMT • 1 spec.; Conquista do Oeste; Nov. 2012; M. A. Carvalho leg.; CEMT • 1 spec.; Cotriguaçú, Fazenda São Nicolau; Nov. 2007; O. Peres-Filho leg.; light; CEMT • 1 spec.; Diamantino; Jan. 2001; A. M. Bello leg.; CEMT • 1 spec.; Poconé; 28 Feb. 1998; forest; kight trap; CEMT • 1 spec.; same data as for preceding; Fazenda Conceição; 16 ° 19 ′ 41 ″ S, 56 ° 30 ′ 21 ″ W; 23 – 31 Mar. 2011; M. B. Pessôa leg.; human feces; CEMT • 1 spec.; Porto Estrela, E. E. Serra das Araras; 15 ° 39 ′ 19 ″ S, 57 ° 12 ′ 50 ″ W; 21 Nov. 2017; R. J. P. Machados and R. Nunes leg.; heliporto; light; CEMT • 1 spec.; Primavera do Leste; 10 Jun. 2010; CEMT • 1 spec.; same data as for preceding; Ilha Grande Farm; 15 ° 49 ′ 83 ″ S, 54 ° 12 ′ 82 ″ W; Feb. 2013; A. S. Tissiani leg.; manually; CEMT • 8 specs; Tangará da Serra; 14 ° 38 ′ 33 ″ S, 57 ° 33 ′ 18 ″ W; 19 – 22 Feb. 2012; A. S. Tissiani leg.; CEMT • 1 spec.; same data as for preceding; Fazenda Bahía; 14 ° 37 ′ 04 ″ S, 57 ° 25 ′ 00 ″ W; elev. 480 m; 26 – 28 Jan. 2012; R. J. Silva leg.; pasture; pitfall cow; CEMT • 1 spec.; Trindade, U. B. S.; 1 Oct. 1984; E. Binda leg.; polo noroeste; light; INPA. – Mato Grosso do Sul • 3 specs; Corquinho Quinta do Sol; 19 ° 49 ′ 57 ″ S, 54 ° 49 ′ 45 ″ W; Feb. 2011; L. O. Bavutii leg.; CEMT • 3 specs; Guia Lopes da Laguna; 15 Nov. 2007; A. Abot leg.; light trap; CEMT • 1 spec.; Selvíria, UNESP farm.; C. A. H. Flechtmann leg.; Brachiaria decumbens pasture; ex Guzerá bovine dropping; CEMT. – Minas Gerais • 2 specs; Águas Vermelhas; Dec. 1997; A. Bello and Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; Dec. 1998; CEMT • 1 spec.; Buenópolis, P. E. Serra Cabral; 17 ° 54 ′ 48 ″ S, 44 ° 12 ′ 50 ″ W; 17 Apr. 2008; M. F. Souza leg.; human dung; CEMT • 2 specs; same data as for preceding; 17 ° 55 ′ 13 ″ S, 44 ° 14 ′ 21 ″ W; CEMT • 2 specs; same data as for preceding; 17 ° 55 ′ 18 ″ S, 44 ° 17 ′ 48 ″ W; CEMT • 1 spec.; Conceição dos Ouros, Rio Sapucaí; Jan. 2003; A. G. P Neto leg.; CEMT • 1 spec.; Ingaí, Reserva Boqueirão, UNILAVRAS; 22 Oct. 2016; J. Mariano leg.; CEMT • 1 spec.; Itumirim; 21 ° 20 ′ 24 ″ S, 44 ° 48 ′ 06 ″ W; 19 Oct. 2016; Vieira et al. leg.; light; CEMT • 2 specs; Lavras-Ingaí, Próx Quedas do Rio Bonito; 21 Oct. 2016; CEMT • 2 specs; Lavras; 21 Oct. 2016; Silva et al. leg.; manually CEMT • 1 spec.; same data as for preceding; 21 ° 18 ′ 16 ″ S, 44 ° 58 ′ 01 ″ W; P 19; CEMT • 1 spec.; same data as for preceding; 21 ° 20 ′ 21 ″ S, 44 ° 58 ′ 42 ″ W; 22 Mar. 2008; M. R. Santos and D. L. H. Takahashi leg.; CEMT • 1 spec.; same data as for preceding; 21 ° 20 ′ 21 ″ S, 44 ° 58 ′ 43 ″ W; CEMT • 2 specs; same data as for preceding; 21 ° 20 ′ 03 ″ S, 44 ° 58 ′ 30 ″ W; C 28; CEMT • 1 spec.; same data as for preceding; 21 ° 20 ′ 6 ″ S, 44 ° 58 ′ 20 ″ W; C 54; CEMT • 1 spec.; Lavras, Próx Poço Bonito; Dec. 2001; Vaz-de-Mello leg.; CEMT • 3 specs; Paracatu; Mar. 1997; S. Lourengo jr leg.; CEMT • 1 spec.; Santa Barbara; 21 Oct. 1994; J. C. Zanuncio leg.; CEMT • 1 spec.; Santana do Riacho, PN Serra do Cipó, base camp; elev. 1500 m; Jan. 2001; F. Z. Vaz de Mello leg.; CEMT • 1 spec.; Três Marias; Dec. 1991; CEMT • 1 spec.; Uberlândia, Campus UFU; 26 Jun. 2005; CEMT. – Tocantins • 1 spec.; Porto Nacional; Jun. 2003; J. C. Oliveira leg; Col 1 / 1 n 1 S; CEMT. PARAGUAY – Gran Asunción • 1 spec.; 30 Nov. 1989; G. Arriágada leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFE3FFBFFF62FBB5FCA0FCEC.taxon	materials_examined	ARGENTINA – San Luis • 1 spec.; Dpto. Belgrano, Fdo. El Molle; 18 Feb. – 8 Mar. 2013; G. Arriágada leg.; human feces; pitfall; CEMT. – Tucumán • 2 specs; Tucumán; Feb. 1959; L. Golbach leg.; CEMT. BOLIVIA – Santa Cruz • 1 spec.; Buena Vista env.; 27 – 29 Nov. 1998; V. Tichy leg.; CEMT • 1 spec.; Chiquitos, 20 km NE of Santiago de Chiquitos; 18 ° 08 ′ 45 ″ S, 59 ° 16 ′ 45 ″ W; elev. 215 m; Nov. 2008; W. D. Edmonds and T. Vidaurre leg.; human feces; CEMT. BRAZIL – Acre • Assis Brasil; 10 ° 35 ′ 44 ″ S, 69 ° 57 ′ 11 ″ W; elev. 170 m; 21 Sep. 2016; Ortiz and Borges leg.; wild environment; light trap (CDC); CEMT • 4 specs; Rio Branco, Universidade Federal de Rio Branco, Parque Zoobotânico; Feb. 1997; Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; Nov. 2000; M. A. Oliveira leg.; CEMT. – Amazonas • 4 specs; Itacoatiara, Fazenda Aruana, Am 010, Km 215; 12 – 15 Jan. 1991; C. S. Motta et al. leg.; UV light; INPA • 8 specs; same data as for preceding; C. S. Motta, H. Mesquita and R. Andreazze leg.; INPA • 23 specs; same data as for preceding; 13 – 16 Feb. 1991; C. S. Motta, R. Andreazze and V. Kawazoe leg.; UV light; INPA • 1 spec.; same data as for preceding; 13 – 16 Mar. 1991; C. S. Motta, F. J. A. Peralta and B. Rochin-Teles leg.; CEMT • 8 specs; same data as for preceding; INPA • 4 specs; same data as for preceding; 13 – 15 May 1991; C. S. Motta et al. leg.; UV light; INPA • 4 specs; Manaus; 11 Apr. 1962; Corq leg.; INPA • 7 specs; same data as for preceding; 22 Aug. 1977; INPA • 1 spec.; same data as for preceding; Sítio São José, C. Nova V.; 27 Sep. 1992; M. G. V. A. O. Barvosa leg.; CEMT • 1 spec.; Manaus, Reserva Ducke; 4 Jun. 1976; Eduardo leg.; INPA • 1 spec.; Manaus, B. Raiz; 2 May 1976; Eduardo leg.; INPA • 4 specs; Manaus, B. de S. Francisco; 16 Jul. 1962; Eduardo leg.; INPA • 1 spec.; Lago de Janaucá; 30 Aug. 1962; INPA • 1 spec.; São Gabriel da Cachoeira, Querari Pelotão; 1 ° 05 ′ N, 69 ° 51 ′ W; 7 – 23 Apr. 1993; C. S. Motta et al. leg.; CEMT. – Bahia • 3 specs; Barreiras, 150 km W of Barreiras; Jan. 2003; P. A. Schmidt leg.; CEMT • 7 specs; same data as for preceding; Aug. 2002; CEMT • 1 spec.; Barreiras, Dist Placas; 11 ° 49 ′ S, 46 ° 07 ′ W; Jan. 2004; P. A. Schmidt leg.; CEMT • 2 specs; same data as for preceding; Fazenda Girassol; 11 ° 50 ′ S, 46 ° 10 ′ W; Feb. 2010; R. V. Nunes leg.; human feces; pitfall; CEMT • 1 spec.; Caetité, C. Urbn Lagoa Real-INB; 8 – 16 Jan. 2000; Nessian and Baptista leg.; light trap; CEMT • 1 spec.; Caravelas; 20 Jan. 1997; Anjos and Silveira leg.; CEMT • 4 specs; same data as for preceding; 25 Mar. 1997; CEMT • 1 spec.; same data as for preceding; Mar. 1996; R. Diniz leg.; CEMT • 5 specs; Cruz das Almas; Jan. 2005; A. F. Brito leg.; CEMT • 1 spec.; Itamaraju; 17 ° 00 ′ 04 ″ S, 39 ° 26 ′ 01 ″ W; Apr. 2011; C. M. P. Leite leg.; pitfall; CEMT • 9 specs; same data as for preceding; 17 ° 01 ′ 09 ″ S, 39 ° 28 ′ 15 ″ W; Apr. 2011; CEMT • 1 spec.; same data as for preceding; 15 ° 15 ′ 29 ″ S, 39 ° 55 ′ 31 ″ W; Nov. 2011; CEMT • 1 spec.; same data as for preceding; 15 ° 15 ′ 51 ″ S, 39 ° 57 ′ 52 ″ W; CEMT • 8 specs; same data as for preceding; Matriz; 2011; CEMT • 2 specs; Mucuri; 21 Jan. 1997; Anjos and Silveira leg.; CEMT • 3 specs; Porto Seguro, RPPN Veracel Mussununga; 11 Nov. 2004; J. N. C. Louzada leg.; CEMT • 7 specs; Presidente Tancredo Neves; 13 ° 22 ′ 56 ″ S, 30 ° 20 ′ 27 ″ W; Jan. 2011; C. M. P. Leite leg.; pitfall; CEMT • 1 spec.; same data as for preceding; 13 ° 22 ′ S, 39 ° 19 ′ W; CEMT • 2 specs; same data as for preceding; 13 ° 24 ′ 32 ″ S, 39 ° 20 ′ 05 ″ W; CEMT • 7 specs; Ubaíra; 13 ° 08 ′ 47 ″ S, 39 ° 40 ′ 34 ″ W; Nov. 2011; CEMT • 3 specs; Wenceslau Guimarães, Matriz; 13 ° 41 ′ 21 ″ S, 9 ° 21 ′ 52 ″ W; elev. 146 m; CEMT. – Ceará • 2 specs; Ubajara, P. N. Ubajara, Cachoeira Cafundó; 3 ° 50 ′ 12 ″ S, 40 ° 54 ′ 35 ″ W; elev. 783 m; 15 Feb. 2013; D. M. Takiya and A. P. M. Santos leg.; Pennsylvania trap; CEMT • 1 spec.; Ubajara, P. N. Ubajara, Portão Neblina; 3 ° 50 ′ 18 ″ S, 40 ° 53 ′ 54 ″ W; elev. 849 m; 14 Feb. 2013; D. M. Takiya and A. P. M. Santos leg.; light; CEMT • 7 specs; Ubajara, P. N. Ubajara, Trilha da Samambaia; 3 ° 50 ′ 20 ″ S, 40 ° 53 ′ 55 ″ W; elev. 850 m; 17 Feb. 2013; D. M. Takiya leg.; light; CEMT • 14 specs; same data as for preceding; Vaz-de-Mello and Grossi leg.; CEMT • 1 spec.; same data as for preceding; Trilha Araticum, Rio Das Mina; 3 ° 50 ′ 21 ″ S, 40 ° 54 ′ 23 ″ W; elev. 880 m; 13 Feb. 2013; D. M. Takiya and A. P. M. Santos leg.; plus light; CEMT. – Distrito Federal • 1 spec.; Brasília, Lago Norte; 1 Mar. 1999; Fábio leg.; CEMT • 1 spec.; Brasília; elev. 1100 m; Jan. 2001; N. Degallier leg.; CEMT • 3 specs; same data as for preceding; light; CEMT • 2 specs; same data as for preceding; Feb. 2001; CEMT • 89 specs; Brasília, Faz Água Limpa; Nov. 2010; C. Suinaga leg.; CEMT • 2 specs; same data as for preceding; 15 ° 56 ′ 40.2 ″ S, 47 ° 55 ′ 38.9 ″ W; 10 Dec. 2017; Y. Ferreira leg.; pasture; pitfall fezes de porco; CEMT • 4 specs; Planaltina, Embrapa Cerrados; 15 ° 35 ′ 52 ″ S, 47 ° 43 ′ 10 ″ W; 4 Dec. 2017; Y. Ferreira leg.; soy; human feces; pitfall; CEMT. – Espírito Santo • 4 specs; Linhares, R. F. C. V. R. D.; Jun. 1988; J. S. Santos leg.; CEMT • 6 specs; same data as for preceding; Reserva Natural da Vale; Nov. 1988; J. S. Santos leg.; CEMT • 12 specs; same data as for preceding; Sooretama; Dec. 1988; CEMT • 1 spec.; Venda Nova do Imigrante, Clube; 20 ° 20 ′ 05 ″ S, 41 ° 08 ′ 18 ″ W; Jan. 2010; Vaz-de-Mello leg.; CEMT • 2 specs; Venda Nova do Imigrante, Vila Santa Cruz; 4 – 12 Jan. 2018; Falqueto and Vaz-de-Mello leg.; CEMT • 1 spec.; Venda Nova do Imigrante; Oct. 1998; Falqueto and Vaz-de-Mello leg.; CEMT. – Goiás • 1 spec.; Joanópolis, Chácara São José; 27 Feb. 2015; E. Santana leg.; CEMT • 1 spec.; Luziânia; Nov. 1975; A. Bello leg.; CEMT • 13 specs; Rio Verde; Nov. 1995; J. Carlos leg.; CEMT • 1 spec.; same data as for preceding; Nov. 1997; CEMT. – Maranhão • 2 specs; Chapadinha; 4 ° 51 ′ 50 ″ S, 43 ° 20 ′ 51 ″ W; 12 May 2012; C. G. Silva leg.; CEMT • 2 specs; Maranhão, Mirador, P. E. Mirador, Base Geraldina; 20 – 24 Dec. 2006; R. O. Souza and A. L. Silva leg.; CEMT • 1 spec.; same data as for preceding; 27 Nov. 2008; M. B. Aguiar-Neto and A. L. Costa leg.; light; CEMT • 4 specs; São Luís, UEMA, Aléia; Feb. 2009; M. C. Batista leg.; CEMT. – Mato Grosso • 1 spec.; Água Boa, Bairro Operário; Dec. 2017; Mota and Schutz leg.; CEMT • 7 specs; same data as for preceding; Rancho Araguaia; 13 ° 56 ′ 40 ″ S, 52 ° 06 ′ 10 ″ W; 24 Dec. 2017; Mota and Schutz leg.; CEMT • 1 spec.; Alta Floresta; 28 Nov. 1988; Bello leg.; CEMT • 6 specs; same data as for preceding; CEPLAC; 4 Sep. 2009; V. Gonçalves leg.; inside Tapirus feces; CEMT • 2 specs; same data as for preceding; human feces; CEMT • 1 spec.; same data as for preceding; Inter. Vôo; CEMT • 1 spec.; same data as for preceding; pitfall; CEMT • 3 specs; same data as for preceding; 4 Sep. 2010; human feces; CEMT • 12 specs; Chacara Recanto das Orquídeas; 3 Oct. 2011; A. Petini-Bevell leg.; CEMT • 1 spec.; same data as for preceding; 7 Aug. 2012; CEMT • 1 spec.; Barão de Melgaço, Centro; 1 Apr. 2012; R. Marco leg.; manually; CEMT • 1 spec.; Campo Verde, Fazenda Santa Luzia; 15 ° 19 ′ 18 ″ S, 54 ° 52 ′ 23 ″ W; elev. 668 m; 9 May 2012; K. Peña leg.; CEMT • 1 spec.; Chapada dos Guimarães; 11 Dec. 2000; A. F. Ramos leg.; light trap; CEMT • 1 spec.; same data as for preceding; Fazenda Jardim; 7 Jul. 2012; V. C. Jorge leg.; CEMT • 1 spec.; same data as for preceding; 21 Jul. 2012; CEMT • 1 spec.; same data as for preceding; 29 Sep. 2012; CEMT • 1 spec.; same data as for preceding; 19 Jan. 2013; CEMT • 1 spec.; Chapada dos Parecis, 30 km N of Uirapuru; 14 ° 17 ′ S, 59 ° 15 ′ W; Dec. 2002; A. Foucart leg.; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 15 Dec. 2007; Peres-Filho leg.; light trap 03; CEMT • 2 specs; same data as for preceding; 15 Dec. 2007; CEMT • 3 specs; same data as for preceding; 31 Oct. 2007; light trap; CEMT • 1 spec.; same data as for preceding; 9 ° 49 ′ 08 ″ S, 58 ° 15 ′ 40 ″ W; 19 – 24 Oct. 2012; F. Z. Vaz-de-Mello leg.; PPBio P 1; CEMT • 1 spec.; same data as for preceding; 9 ° 51 ′ 18 ″ S, 58 ° 14 ′ 57 ″ W; elev. 220 m; 14 Dec. 2009; F. Z. Vaz-de-Mello leg.; base camp; light; CEMT • 2 specs; same data as for preceding; 9 ° 51 ′ S, 58 ° 15 ′ W; Nov. 2010; F. Z. Vaz-de-Mello leg.; CEMT • 2 specs; same data as for preceding; 9 ° 51 ′ 20 ″ S, 58 ° 14 ′ 55 ″ W; Nov. 2014; R. V. Nunes and A. Asenjo leg.; light; CEMT • 1 spec.; same data as for preceding; Fazenda São Nicolau; Nov. 2007; O. Peres-Filho leg.; light; CEMT • 1 spec.; same data as for preceding; 15 Oct. 2007; light trap; CEMT • 2 specs; same data as for preceding; 31 Oct. 2007; white light trap 05; CEMT • 3 specs; same data as for preceding; 31 Oct. 2014; W. Chamorro leg.; white light trap; CEMT • 1 spec.; same data as for preceding; 9 ° 50 ′ 24 ″ S, 58 ° 15 ′ 10 ″ W; E. Sanhudo leg.; primary forest; CEMT • 2 specs; Cotriguaçu, Fazenda São Nicolau; Oct. 2014; R. Stofel leg.; CEMT • 1 spec.; Cuiabá; 15 ° 16 ′ 45 ″ S, 56 ° 00 ′ 06 ″ W; 7 Dec. 2015; A. Frolov and L. Akhmetova leg.; CEMT • 2 specs; Cuiabá; Boa Esperança; Dec. 2009; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; Oct. 2016; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; CEMT • 11 specs; Cuiabá; Campus UFMT; Nov. 2009; R. Miyazaki leg.; CEMT • 7 specs; same data as for preceding; 9 Oct. 2016; A. Frolov leg.; UV light; CEMT • 1 spec.; Cuiabá, Fazenda Santhidi; 15 ° 23 ′ 06 ″ S, 56 ° 06 ′ 53 ″ W; A. F. Oliveira and L. R. Silva leg.; pitfall-banana; CEMT • 1 spec.; same data as for preceding; Jardim Brasil; 15 May 2012; B. S. Neves leg.; manually; CEMT • 3 specs; same data as for preceding; Recanto Siriema; 15 ° 35 ′ 32 ″ S, 56 ° 01 ′ 38 ″ W; 4 Nov. 2014; R. V. Nunes and B. F. Becker leg.; light; CEMT • 1 spec.; same data as for preceding; Terro Nova; 3 Jan. 2012; V. Henrique leg.; manually; CEMT • 4 specs; Diamantino, Alto Rio Arinos; Jan. 2001; E. Furtado leg.; CEMT • 1 spec.; Farinópolis, Fazenda Água Limpa; 15 ° 16 ′ 14 ″ S, 58 ° 32 ′ 11 ″ W; Dec. 2004; M. Santos-Filho leg.; Watershed Jauru; CEMT • 1 spec.; same data as for preceding; Fazenda Sudan; 5 ° 15 ′ 14 ″ S, 58 ° 27 ′ 20 ″ W; CEMT • 1 spec.; Indiavai, Fazenda Nova Canaã; 15 ° 15 ′ 52 ″ S, 58 ° 42 ′ 59 ″ W; Jan. 2004; M. Santos-Filho leg.; Watershed Jauru; CEMT • 1 spec.; Lucas do Rio Verde; 2 Nov. 2011; B. F. Camera leg.; CEMT • 8 specs; Nova Ubitatã, ESEC Ronuro; 13 ° 06 ′ 43 ″ S, 54 ° 26 ′ 37 ″ W; 2 Mar. 2017: L. O. F. A. Nunes leg.; Watershed Jauru; CEMT • 2 specs; Poconé, Base UFMT; 16 ° 29 ′ 58 ″ S, 56 ° 24 ′ 49 ″ W; Mar. 2012; M. Rossini leg.; CEMT • 1 spec.; Poconé, Centro; 19 Dec. 2011; V. Henrique leg.; manually; CEMT • 2 specs; Poconé, Mata; 24 Jun. 1998; light trap; CEMT • 1 spec.; Poconé; Jul. 2000; J. Lopes leg.; CEMT • 4 specs; Recanto Siriema; 15 ° 35 ′ 32 ″ S, 56 ° 01 ′ 38 ″ W; 4 Nov. 2014; R. V. Nunes and B. F. Becker leg.; light; CEMT • 1 spec.; Santa Carmem, Fazenda São Vicente; 11 ° 58 ′ 12 ″ S, 55 ° 16 ′ 39 ″ W; elev. 349 m; 3 May 2014; K. Peña leg.; field; CEMT • 1 spec.; Sesc Pantanal; 14 May 2002; light trap; CEMT • 1 spec.; same data as for preceding; 4 Jul. 2002; CEMT • 1 spec.; same data as for preceding; 5 Jul. 2003; CEMT • 6 specs; Tangara da Serra; 12 Feb. 2012; B. F. Camera leg.; CEMT • 4 specs; same data as for preceding; 14 ° 38 ′ 33 ″ S, 57 ° 33 ′ 18 ″ W; 19 – 22 Feb. 2012; A. S. Tissiani leg.; CEMT • 4 specs; same data as for preceding; Fazenda Três Rios; 14 ° 21 ′ S, 57 ° 42 ′ W; elev. 592 m; Feb. 2008; R. J. Silva leg.; Cerrado; pitfall sem isca; CEMT. – Mato Grosso do Sul • 1 spec.; Aquidauana; S. R. R. Rodrigues leg.; CEMT • 6 specs; Bodoquena, Fazenda Californea; 20 ° 41 ′ 02 ″ S, 56 ° 51 ′ 40 ″ W; Mar. 2001; L. O. Bavutti leg.; fligth intercept; CEMT • 2 specs; Campo Grande; 1990 – 1992; I. Biachin leg.; CEMT • 3 specs; Cassilândia; Jan. 2014; R. V. Nunes and L. Sawaris leg.; semidecidual forest; human feces; pitfall; CEMT • 1 spec.; Corquinho Quinta do Sol; 19 ° 49 ′ 57 ″ S, 54 ° 49 ′ 45 ″ W; Feb. 2011; L. O. Bavutti leg.; flight intercept; CEMT • 1 spec.; Corumbá, Serra do Amolar; Apr. 2011; C. Aoki leg.; CEMT • 1 spec.; Dourados, Mata do Azulão; 22 ° 12 ′ 40 ″ S, 54 ° 55 ′ 06 ″ W; elev. 450 m; 13 – 16 Dec. 2010; F. Z. Vaz-de-Mello leg.; manually; CEMT • 1 spec.; Nova Andradina, Usina Santa Helena; 21 ° 59 ′ 32 ″ S, 53 ° 26 ′ 28 ″ W; elev. 360 m; 18 Oct. 2012; G. Coutinho leg.; CEMT. – Minas Gerais • 1 spec.; Águas Vermelhas; Dec. 1997; A. Bello; CEMT • 10 specs; same data as for preceding; Dec.; 1997; A. Bello and F. Z. Vaz-de-Mello leg.; CEMT • 2 specs; Araponga, Pico do Boné; 28 – 29 Dec. 2000; E. Stehling leg.; CEMT • 2 specs; Conceição dos Ouros, Rio Sapucaí; Jan. 2003; Almeida and G. P. Neto leg.; CEMT • 2 specs; Cordisburgo, Fazenda Pontinha; Jul. 1994; De Mello leg.; CEMT • 1 spec.; same data as for preceding; Jan. 1996; CEMT • 1 spec.; same data as for preceding; Jan. 1999; CEMT • 1 spec.; Coxambu; Dec. 1979; A. Bello leg.; CEMT • 1 spec.; same data as for preceding; Dec. 1990; CEMT • 1 spec.; Diamantina; 23 Feb. – 3 Mar. 2001; N. Degallier leg.; CEMT • 11 specs; Ijací, Faz FAEPE; Nov. 2002; J. Louzada leg.; CEMT • 6 specs; Ipatinga; 30 Apr. 1991; light trap; CEMT • 2 specs; same data as for preceding; Nov. 1991; E. Grossi; CEMT • 1 spec.; same data as for preceding; Sep. 1993; E. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Dec. 1993; P. Grossi and E. Grossi leg.; CEMT • 15 specs; Itumurim; 21 ° 20 ′ 24 ″ S, 44 ° 48 ′ 06 ″ W; 19 Oct. 2016; Viera, F. Z. Vaz-de-Mello and J. Louzada leg.; CEMT • 1 spec.; Lavras; 21 ° 18 ′ 16 ″ S, 44 ° 58 ′ 01 ″ W; CEMT • 1 spec.; same data as for preceding; 21 ° 18 ′ 52 ″ S, 44 ° 59 ′ 33 ″ W; 20 Jan. 2008; M. Rocha and D. L. H. Takahashi leg.; CEMT • 13 specs; Lavras, Próx Poço Bonito; Dec. 2001; Vaz-de-Mello leg.; CEMT • 1 spec.; Lavras, Ponte Nova; 23 Oct. 1979; Planosulcar leg.; CEMT • 1 spec.; same data as for preceding; 30 Oct. 1979; CEMT • 1 spec.; same data as for preceding; 6 Nov. 1979; CEMT • 1 spec.; same data as for preceding; 1 Apr. 1980; CEMT • 1 spec.; Lavras UFLA; Sep. 2002; Vaz-de-Mello leg.; CEMT • 1 spec.; Lima Duarte, Parque Estadual Ibitipoca; 20 Feb. 1997; C. R. M. Abreu leg.; CEMT • 1 spec.; Montes Claros; Nov. 1990; CEMT • 1 spec.; same data as for preceding; Jan. 1991; CEMT • 9 specs; Nova Era; Aug. 1987; CEMT • 9 specs; Paracatu; Mar. 1997; S. Lourengo leg.; CEMT • 1 spec.; same data as for preceding; Nov. 1997; CEMT • 6 specs; same data as for preceding; Mar. 1998; CEMT • 4 specs; Recreio; Jan. 2000; Lopes-Andrade leg.; CEMT • 1 spec.; Santa Barbara; 11 Jan. 1995; J. C. Zanuncio leg.; CEMT • 5 specs; same data as for preceding; 17 Feb. 1994; CEMT • 1 spec.; same data as for preceding; 27 Oct. 1994; CEMT • 2 specs; same data as for preceding; 26 Jan. 1995; CEMT • 1 spec.; Santana do Riacho, PARNA Serra do Cipó; 11 Nov. 2004; G. Schiffler leg.; Cerrado; CEMT • 6 specs; Serra Bocaina, Campo Sujo; 21 ° 19 ′ 47 ″ S, 44 ° 59 ′ 46 ″ W; 6 Sep. 2012; A. Diaz-Rojas leg.; pig dung; CEMT • 1 spec.; Sete Lagoas; Jan. 1991; F. Z. Vaz-de-Mello leg.; CEMT • 4 specs; Teixeiras; Nov. 1996; Hardy and Harrison leg.; CEMT • 1 spec.; Viçosa; 7 Feb. 1996; J. N. C. Louzada leg.; CEMT • 1 spec.; same data as for preceding; 10 Apr. 1933; CEMT • 2 specs; same data as for preceding; 20 Feb. 1986; P. S. F. Fiuza leg.; UV trap; CEMT • 1 spec.; same data as for preceding; 20 Apr. 1990; G. A. R. Melo leg.; attracted by light; CEMT • 4 specs; same data as for preceding; 20 Sep. 1993; Louzada and Silva leg.; CEMT • 6 specs; same data as for preceding; 23 Sep. 1993; CEMT • 7 specs; same data as for preceding; Campus UFV; Nov. 1997; F. Vaz-de-Mello leg.; CEMT • 4 specs; Viçosa, Córrego do Paraíso, Mata da Prefeitura; 11 Dec. 1988; Fiuza and Martins leg.; CEMT • 3 specs; same data as for preceding; 13 Dec. 1984; CEMT • 2 specs; Viçosa; 1994 – 1995; Louzada, Carvalho and Silva leg.; pasture; CEMT • 2 specs; Viçosa, UFV; Dec. 1999; Lopes-Andrade and Vaz-de-Mello leg.; CEMT • 4 specs; Viçosa; Nov. 1996; Vaz-de-Mello, Hardy and Harrion leg.; CEMT. – Pará • 1 spec.; Paragominas; 2.9372 ° S, 47.210 ° W; elev. 114 m; Feb. 2001; R. Solar leg.; agric; CEMT • 1 spec.; same data as for preceding; 2.9123 ° S, 47.674 ° W; Nov. 2011; R. Solar leg.; perturbed primary forest; CEMT • 4 specs; Santarém; Agric. M, B 129 T 9 P 2 ARM 1; CEMT • 1 spec.; Santarém, Alter do Chão; 2 Aug. 2003; savana; CEMT • 8 specs; Santarém; fruit culture; CEMT • 1 spec.; same data as for preceding; B 112 T 9 P 3 ARM 1; CEMT. – Paraíba • 1 spec.; Areia, Mata Pau de Ferro; 3 – 4 May 2000; M. Avany leg.; light trap; CEMT • 2 specs; Cacimba de Dentro, Fazenda Cachoeira de Capivara; 6 ° 40 ′ S, 35 ° 45 ′ W; elev. 340 m; May 2003; M. I. M. Hernández leg.; caatinga; pitfall; CEMT • 1 spec.; Estua do Rio Mamanguape; 1 – 7 Jun. 2000; D. Araújo leg.; Malaise trap; CEMT • 1 spec.; same data as for preceding; 25 – 27 Sep. 2000; light trap; CEMT • 1 spec.; Rio Tinto, APA Barra do Rio Mamanguape; 6 ° 46 ′ 41 ″ S, 64 ° 35 ′ 37 ″ W; 13 Nov. 2012; D. Dal-Bó leg.; Malaise trap; CEMT. – Paraná • 4 specs; Entre Rio do Oeste; 26 Dec. 2017; M. E. Maldaner leg.; manually (light); CEMT • 1 spec.; Jaguariaíva, Parque Estadual do Cerrado; 3 Feb. 2011; Grossi and Parizotto leg.; CEMT • 1 spec.; Londrina; Nov. – Dec. 1998; M. Medri leg.; CEMT. – Pernambuco • 1 spec.; Igarassu, RECD; 11 May 2007; Costa et al. leg.; CEMT • 1 spec.; same data as for preceding; 27 Apr. 2007; Costa et al. leg.; CEMT • 1 spec.; same data as for preceding; Silva et al. leg.; CEMT. – Rio de Janeiro • 2 specs; Angra dos Reis, Bracuí, lowland; 10 – 11 May 2012; Nessimian leg.; UV light; CEMT • 14 specs; Miguel Pereira; Jan. 1998; J. Carlos leg.; CEMT • 4 specs; same data as for preceding; Feb. 1998; CEMT • 3 specs; same data as for preceding; Mar. 1998; CEMT • 2 specs; Niteroi, Pendotiba; Feb. 2000; B. Schott leg.; CEMT • 8 specs; Nova Friburgo; Nov. 1996; F. Z. Vaz-de-Mello leg.; CEMT • 8 specs; same data as for preceding; Nov. 1998, P. Grossi and E. Grossi leg; CEMT • 12 specs; Feb. 2001; P. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Dec. 2001; CEMT • 1 spec.; same data as for preceding; Mar. 2003; E. Grossi and P. Grossi leg.; CEMT • 3 specs; same data as for preceding; Mar. 2012; E. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Nov. 2016; E. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Macaé de Cima; Nov. 1998; F. Z. Vaz-de-Mello and E. Grossi leg.; CEMT • 1 spec.; Rio de Janeiro, Deodoro; Jan. 1946; W. Zikán leg.; CEMT • 1 spec.; Rio de Janeiro, Parque da Cidade; 30 Jan. 1991; A. Bello leg.; CEMT • 1 spec.; Saquarema; Dec. 1980; A. Bello leg.; CEMT • 1 spec.; Teresópolis, Granja Comary; 10 Jan. 1979; A. Bello leg.; CEMT. – Rio Grande do Sul • 1 spec.; Bagé, Fazenda Querências da Pedreira; 30 ° 58 ′ 58 ″ S, 54 ° 20 ′ 12 ″ W; 20 – 26 Oct. 2011; R. M. Moraes leg.; pitfall; CEMT • 1 spec.; Bento Gonçalves; Nov. 2001; C. Arioli leg.; CEMT • 1 spec.; Dom Pedrito; 30 ° 58 ′ 41,19 ″ S, 54 ° 40 ′ 38,78 ″ W; 27 Dec. 2015; P. G. da Silva leg.; manually collect (light); CEMT • 9 specs; El Dorado do Sul, E. E. A- U. F. R. G. S.; 11 Oct. 1996; Carvalho and Silva leg.; arm. lum; CEMT • 2 specs; same data as for preceding; Oct. 1996; Silva and Carvalho leg.; CEMT • 1 spec.; Santa Cruz do Sul, Cerro Alegre; 29 ° 48 ′ 05 ″ S, 52 ° 19 ′ 39 ″ W; 10 Dec. 2011; R. Bohn leg.; grass land field (P 1); pitfall bovine feces; CEMT • 1 spec.; Santa Maria; Nov. 2014 – May 2015; J. Boscardin leg.; CEMT • 16 specs; Silveira Martins; 29 ° 38 ′ S, 53 ° 37 ′ W; Jan. 2011; G. Viegas leg.; human feces; pitfall; CEMT. – Rondônia • 1 spec.; Porto Velho, Rio das Garças; 8 ° 49 ′ 47 ″ S, 63 ° 46 ′ 51 ″ W; Bom Jesus; 30 Mar. 2017; D. C. Santos and K. K. G. Silva leg.; FIT; CEMT • 1 spec.; Rolim de Moura; 11 ° 44 ′ 24 ″ S, 61 ° 55 ′ 26 ″ W; elev. 274 m; 7 – 9 Dec. 2015; D. C. Castro leg.; SAF, pitfall human dung; CEMT • 1 spec.; same data as for preceding; 11 ° 44 ′ 25 ″ S, 61 ° 55 ′ 26 ″ W; elev. 273 m; CEMT. – Roraima • 6 specs; Caracaraí; Jun. 1997; Vaz-de-Mello leg.; CEMT. – Santa Catarina • 2 specs; Santa Rosa de Lima; 28 ° 02 ′ S, 49 ° 08 ′ W; elev. 365 m; 18 Jan. 2018; C. M. Rover leg.; pasto aberto; pitfall; CEMT • 1 spec.; São Francisco do Sul; 17 – 23 Dec. 2016; J. V. Korgut leg.; manually; CEMT. – São Paulo • 2 specs; Descalvado, Fazenda Itaúnas; 6 Apr. 2006; N. W. Perioto leg.; mata-cerrado; pitfall; CEMT • 1 spec.; same data as for preceding; 21 Sep. 2006; forest + Citrus plantation; CEMT • 1 spec.; same data as for preceding; 24 Jul. 2006; CEMT. – Tocantins • 1 spec.; Palmas; 22 Nov. 2001; U. H. E. Lajeado leg.; Equipe de Resgate; CEMT • 4 specs; Porto Nacional; 5 Oct. 2005; M. S. Matos leg.; agrotins; CEMT. ECUADOR – El Oro • 5 specs; Santa Rosa; elev. 230 m; 1 Sep. 1997; C. Carpio leg.; excremento de vaca; CEMT. – Esmeraldas • 1 spec.; Camaroes; Nov. 1987; N. Silva leg.; CEMT. EL SALVADOR – Ahuachapán • 1 spec.; Cara Sucia, Parq. Nacional El Imposible; 13 ° 49 ′ 41 ″ N, 89 ° 56 ′ 37 ″ W; 18 – 20 Jul. 2004; Q. Santiago leg.; FIT; CEMT. MEXICO – Oaxaca • 3 specs; San José, Del Chilar, S. baja caducufolia; 20 Jan. 2005; Ramírez and Ponce leg.; directa en excremento de vaca [extracted from cow dung]; CEMT. PARAGUAY – Alto Paraná • 2 specs; Limoy; 23 Nov. 1990; G. Arriágada; CEMT. – Boquerón • 6 specs; Loma Plata; 25 – 28 Jan. 1991; R. A. Matavelli leg; CEMT. – Gran Asunción • Asunción; 30 Nov. 1989; G. Arriágada leg.; CEMT. – Paraguari • 1 spec.; Narajo; 5 Nov. 2000; CEMT. URUGUAY – Rivera • 2 specs; Rivera, Bodegas Carrau farm.; 30 ° 58 ′ 16 ″ S, 55 ° 26 ′ 38 ″ W; 3 – 11 Oct. 2011; R. M. Moraes leg.; pitfall trap baited (human + pig dung); CEMT • 2 specs; same data as for preceding; 31 ° 41 ′ 5 ″ S, 54 ° 39 ′ 42 ″ W; CEMT • 4 specs; same data as for preceding; 31 ° 41 ′ 4 ″ S, 54 ° 37 ′ 25 ″ W; CEMT. UNITED STATES OF AMERICA – Florida • 1 spec.; Martin, Jonathan State Park; Jun. 1995; P. Bélanger leg.; CEMT. – Texas • 1 spec.; Live Oak, Choke Canyon S. P., Three Rivers; 24 Jun. 1995; B. Buchli leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFE3FFBFFF62FBB5FCA0FCEC.taxon	discussion	Collecting information Specimens can be found in various locations at altitudes ranging 10 to 1100 metres and can be collected with different traps. Specimens were collected with light and flight intercept traps, with pitfalls baited with human and bovine feces, or directly on the faecal mass and in forest litter (Stebnicka 2005 b). Remarks Ataenius aequalis-platensis group comprises 27 species, nine of these are found in Brazil. A key for species of this group can be found in Stebnicka (2005 b).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFE8FFBFFDAEFC18FDBCFA9A.taxon	description	Figs 10 D, 40 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFE8FFBFFDAEFC18FDBCFA9A.taxon	diagnosis	Diagnosis Length 2.8 – 7.0 mm. Body elongate, shagreened, glabrous or not, piceous. Head coarsely punctate, punctures larger as they approach frons; clypeus always sinuate; lateral angles obtuse or denticulate; ventral edge of clypeus medially downward bent and acuminate. Pronotum glabrous or not, always covered by large and close punctures. Scutellum foveate. Elytra with distinct sculpture; striae always with coarse, deep punctures reaching lateral margins of interstriae; interstriae distinctively elevated carinate or tectiform; lateral margins crenulate by strial punctures. Ventral surface shagreened, covered with coarse punctures. Metatibiae with apical accessory spine always present, large and acuminate, longer than apical fringe of setae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFE8FFBDFF61FAE2FD89FA72.taxon	materials_examined	ARGENTINA – San Luis • 1 spec.; Dept. Belgrano, Fundo. El Molle; 18 Feb. – 8 Mar. 2012; A. Gerardo leg.; human feces; pitfall; CEMT. BRAZIL – Amazonas • 3 specs; Autazes, Urucumtuba; 8 Feb. 2013; P. M. leg.; CEMT • 1 spec.; Itacoatiara, Fazenda Aruana, Am 010, Km 215; 13 – 14 Apr. 1991; C. S. Motta and F. J. A. Peralta leg.; INPA • 1 spec.; Lago de Janaucá; 30 Aug. 1962; J. Delona leg.; INPA. – Bahia • 2 specs; Caravelas; 20 Feb. 1997; Anjos and Silveira leg; CEMT • 1 spec.; Cruz das Almas; Jan. 2005; A. F. Brito and Rebeiro leg.; CEMT • 3 specs; Feira de Santana, Fazenda Cruzeiro do Mocó; 12 ° 12 ′ S, 39 ° 02 ′ W; 11 – 13 Jul. 2013; pitfall human feces; CEMT • 1 spec.; Presidente Tancredo Neves; 13 ° 22 ′ S, 39 ° 19 ′ W; Jan. 2011; C. P. M. Leite leg.; pasture; CEMT • 1 spec.; same data as for preceding; 13 ° 22 ′ 56 ″ S, 39 ° 20 ′ 27 ″ W; Jan. 2011; pitfall; CEMT. – Distrito Federal • 1 spec.; Brasília, Lago Norte; Sep. 2013; R. V. Nunes leg.; CEMT • 1 spec.; Chapada dos Guimarães, Fazenda Jardim; 29 Sep. 2012; V. C. Jorge leg.; CEMT • 1 spec.; same data as for preceding; 10 Nov. 2012; CEMT. – Mato Grosso • 2 specs; Chapada dos Guimarães, Cachoeira Geladeira; 15 ° 25 ′ 26 ″ S, 55 ° 42 ′ 53 ″ W; elev. 620 m; 9 Jan. 2013; M. D. Gimo leg.; pitfall; CEMT • 1 spec.; Poconé, Fazenda Conceição; 16 ° 19 ′ 41 ″ S, 56 ° 30 ′ 11 ″ W; 23 – 31 Mar. 2011; M. B. Pessôa leg.; hum fec; CEMT. – Mato Grosso do Sul • 1 spec.; Guia Lopes da Laguna; 10 Dec. 2007; A. R. Abot leg.; light trap; CEMT • 1 spec.; same data as for preceding; 20 Dec. 2007: CEMT. – Minas Gerais • 8 specs; Águas Vermelhas; Dec. 1997; A. Bello leg.; CEMT • 1 spec.; Buenópolis, Parque Estadual da Serra do Cabral, Almirante; 15 Apr. 2008; M. F. Souza leg.; campo limpo [clean field]; CEMT • 1 spec.; Caminho de São Thomé das Letras para Cruzilia; Sep. 1997; F. Frieiro-Costa leg.; CEMT • 2 specs; Cordisburgo; Jan. 1999; Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; Fazenda Pontinha; Jul. 1994; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; Jan. 1998; CEMT • 1 spec.; same data as for preceding; Jan. 1999; CEMT • 1 spec.; Diamantina; 23 Feb. – 3 Mar. 2001; N. Degallier leg.; CEMT • 1 spec.; Lavras; 21 ° 20 ′ 19 ″ S, 44 ° 58 ′ 35 ″ W; CEMT • 1 spec.; same data as for preceding; 21 ° 20 ′ 19 ″ S, 44 ° 58 ′ 37 ″ W; 22 Mar. 2008; M. R. Santos and D. L. H. Takahashi leg.; CEMT • 1 spec.; same data as for preceding; 21 ° 20 ′ 19 ″ S, 44 ° 58 ′ 34 ″ W; CEMT • 1 spec.; Lavras, Próx. Poço Bonito; Dec. 2001; Vaz-de-Mello leg.; CEMT • 1 spec.; Uberlândia, Campus UFU; 18 Feb. 2006; J. Mendes leg.; CEMT • 2 specs; Paracatu; Mar. 1997; S. Lourengo leg.; CEMT • 2 specs; Três Marias; Dec. 1991; CEMT • 2 specs; Vespasiano; Nov. 1953; A. Machado leg.; CEMT • 1 spec.; Viçosa; 10 Jan. 1994; Louzada and Silva leg.; CEMT • 1 spec.; same data as for preceding; 13 Jan. 1994; CEMT. – Pará • 1 spec.; Alter do Chão; 24 Jul. 2003; Savana A. R. Matavelli leg.; CEMT • 6 specs; Monte Dourado, Jari Cellulose; 0 ° 49 ′ S, 52 ° 35 ′ W; Mar. – Apr. 2005; T. A. Gardner and Hernández leg.; Eucalyptus; dung pitfall trap; CEMT • 1 spec.; Santarém; pasture; CEMT. – Paraíba • 1 spec.; Cacimba de dentro, Fazenda Cachoeira de Capivara; 30 May 2003; Olivia and Rembrandt leg.; CEMT • 1 spec.; Piracuruca, P. N. Sete Cidades; 4 ° 06 ′ 28 ″ S, 41 ° 40 ′ 13 ″ W; Cachoeira do Riachão; elev. 171 m; 8 Feb. 2013; D. M. Takiya and Santos leg.; Pennsylvania trap; CEMT • 3 specs; São José dos Cordeiros, Fazenda Almas; 8 – 9 Mar. 2003; Rothéa and Creão leg.; light trap; CEMT. – Piauí • 1 spec.; S. R. Nonato, P. N. S. da Capivara; Jan. 1999; C. A. Matrangolo leg.; CEMT. – Rio de Janeiro • 1 spec.; Angra dos Reis, Bracuí, Baixada; 10 – 11 May 2002; Nessimini leg.; UV light; CEMT • 1 spec.; Miguel Pereira; Jan. 1998; J. Carlos leg.; CEMT • 1 spec.; Nova Friburgo; Oct. 1997; P. Grossi leg.; CEMT • 1 spec.; Nova Friburgo, Macae de Cima; 22 ° 22 ′ S, 42 ° 29 ′ W; E. Grossi and P. Grossi leg.; CEMT. COLOMBIA – Cundinamarca • 1 spec.; Fusa; Sep. 1962; CEMT. MEXICO – Veracruz • 1 spec.; Apazapan; 19 ° 19 ′ 34 ″ S, 96 ° 43 ′ 34 ″ W; elev. 3300 m; Jul. 1999; E. Montes de Oca and Q. Santiago leg.; mango and banana plantation; cow dung trap; CEMT. NICARAGUA – León • 1 spec.; Jul. 1989; B. Barcete leg.; CEMT. – Masaya • 2 specs; Las Flores; May 1993; Cantamesa leg.; lecocq; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFEAFFA3FF61FA75FCE7FD50.taxon	materials_examined	BRAZIL – Amazonas • 4 specs; Autazes, Urucumbuta; P. M. leg.; C. D. C; CEMT. – Bahia • 2 specs; Mucuri; 21 Jan. 1997; Anjos and Silveira leg.; CEMT. – Distrito Federal • 1 spec.; Brasília, Fazenda Água Limpa; Nov. 2010; C. Suinaga leg.; CEMT. – Goiás • 1 spec.; Rio Verde; Sep. 1997; J. Carlos leg.; CEMT. – Maranhão • 1 spec.; Ipecuru-Mirim; 3 ° 28 ′ 01 ″ S, 14 ° 20 ′ 47 ″ W; 31 Aug. 2010; R. Matavelli and A. Campos leg; pasto II; pitfall human feces FH 2; CEMT. – Mato Grosso • 1 spec.; Alta Floresta, CEPLAC; 7 May 2010; V. Gonçalvez leg.; under Tapirus feces; CEMT • 1 spec.; Alta Floresta; 9 ° 49 ′ 44 ″ S, 56 ° 19 ′ 51 ″ W; Jun. 2008; E. Berenguer leg.; Frag 27; pitfall with human feces; CEMT • 1 spec.; Chapada dos Guimarães, Santuário de Elefantes; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 14 ″ W; 22 Jan. 2020; H. Sehn et al. leg.; horse feces; CEMT • 2 specs; same data as for preceding; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 14 Oct. 2007; CEMT • 2 specs; same data as for preceding; 31 Oct. 2007; CEMT • 4 specs; same data as for preceding; 15 Dec. 2007; O. Peres-Filho leg.; light trap; CEMT • 22 specs; same data as for preceding; 14 Apr. 2008; CEMT • 1 spec.; Diamantino; Jan. 2001; A. M. Bello leg.; CEMT • 5 specs; Poconé, Fazenda Alvorada; 16 ° 27 ′ 10 ″ S, 56 ° 24 ′ 53 ″ W; 23 – 31 Mar. 2011; M. B. Pessoa leg.; human feces; CEMT • 2 specs; Poconé, Pousada das Aráras; 3 Dec. 1998; J. L. Moreno and T. Mestre leg.; Pantanal; CEMT • 1 spec.; Sesc Pantanal; 16 Nov. 2001; CEMT • 3 specs; same data as for preceding; 5 Jul. 2003; light trap; CEMT • 2 specs; same data as for preceding; 5 Aug. 2003; CEMT • 1 spec.; same data as for preceding; 13 Nov. 2003; CEMT • 2 specs; same data as for preceding; 16 Nov. 2003; CEMT • 1 spec.; Paracatu; Mar. 1997; S. Lourenço leg.; CEMT • 3 specs; same data as for preceding; Mar. 1998; S. Lourenço jr leg.; CEMT. – Mato Grosso do Sul • 1 spec.; Corumbá, Fazenda São Bento; Apr. 2012; M. Rossini leg.; manually; bovine feces; CEMT • 3 specs; same data as for preceding; 7 ° 19 ′ 14 ″ S, 56 ° 41 ′ 55 ″ W; 28 Apr. 2012; M. Rossini leg.; riparian forest of Rio Piquiri; CEMT • 1 spec.; same data as for preceding; 7 ° 19 ′ 14 ″ S, 56 ° 41 ′ 59 ″ W; CEMT • 2 specs; same data as for preceding; 17 ° 19 ′ 16 ″ S, 56 ° 41 ′ 51 ″ W; CEMT • 1 spec.; same data as for preceding; 17 ° 19 ′ 16 ″ S, 56 ° 42 ′ 13 ″ W; CEMT • 1 spec.; same data as for preceding; 17 ° 19 ′ 17 ″ S, 56 ° 42 ′ 15 ″ W; CEMT • 1 spec.; same data as for preceding; 17 ° 19 ′ 18 ″ S, 56 ° 41 ′ 51 ″ W; CEMT • 2 specs; same data as for preceding; 17 ° 21 ′ 23 ″ S, 56 ° 41 ′ 47 ″ W; Vochysia divergens for.; CEMT • 1 spec.; same data as for preceding; 17 ° 21 ′ 29 ″ S, 56 ° 41 ′ 51 ″ W; CEMT • 2 specs; same data as for preceding; 17 ° 21 ′ 31 ″ S, 56 ° 41 ′ 52 ″ W; CEMT • 2 specs; same data as for preceding; 17 ° 21 ′ 31 ″ S, 56 ° 41 ′ 53 ″ W; CEMT • 5 specs; same data as for preceding; 17 ° 21 ′ 32 ″ S, 56 ° 41 ′ 55 ″ W; CEMT • 5 specs; same data as for preceding; 17 ° 21 ′ 33 ″ S, 56 ° 41 ′ 54 ″ W; CEMT • 12 specs; Corumbá, Passo do Lontra; Feb. 1996; Louzada and Vaz-de-Mello leg.; CEMT • 1 spec.; Dourados; 10 Dec. 2005; M. Milo leg.; CEMT • 2 specs; Guia Lopes da Laguna; 15 Nov. 2007; A. Abot leg.; light trap; CEMT • 1 spec.; Selviria, UNESP Farm; 6 Jul. 1991; Brachiaria decumbens pasture; ex Guzera cattle dropping; INPA. – Pará • 1 spec.; Paragominas; elev. 114 m; Feb. 2011; R. Solar leg.; agric; CEMT. – Paraíba • 1 spec.; Cacimba de Dentro, Fazenda Cachoeira de Capivara; 6 ° 40 ′ S, 35 ° 45 ′ W; elev. 340 m; May 2003; M. I. M. Hernández leg.; caatinga; pitfall; CEMT. – Paraná • 1 spec.; Entre Rio do Oeste; 26 Dec. 2017; M. E. Maldaner leg.; manually (light); CEMT. – Rio Grande do Sul • 4 specs; Bagé, Fazenda Querência da Pedreira; 30 ° 58 ′ 58 ″ S, 54 ° 20 ′ 12 ″ W; 26 Oct. – 4 Nov. 2011; R. M. Moraes leg.; pitfall; CEMT • 1 spec.; Bagé, Leões farm; 30 ° 59 ′ 12 ″ S, 54 ° 29 ′ 47 ″ W; 20 – 26 Oct. 2011; R. M. Moraes leg.; pitfall trap (human + pig dung); CEMT • 2 specs; Eldorado Do Sul, Estação Experimental Agronomica-UFRGS; 30 Mar. – 5 Apr. 2012; R. M. Moraes and J. Podkowa leg.; pitfall; CEMT • 1 spec.; Santa Cruz do Sul, Cerro Alegre; 29 ° 48 ′ 19 ″ S, 52 ° 19 ′ 53 ″ W; 17 Dec. 2011; R. Bohn leg.; secondary forest; pitfall bovine feces; CEMT • 1 spec.; Silveira Martins; 29 ° 38 ′ S, 53 ° 37 ′ W; Jan. 2011; G. Viegas leg.; CEMT. – Santa Catarina • 1 spec.; Campos Novos; 27 ° 23 ′ S, 51 ° 12 ′ W; Feb. 2011; R. C. Campos leg.; baited pitfall; CEMT • 1 spec.; Santa Rosa de Lima; 28 ° 3 ′ S, 49 ° 8 ′ W; elev. 278 m; 18 Jan. 2018; C. M. Rover leg.; silvopastoral with nuclei; CEMT. – Tocantins • 1 spec.; Distrito de Pium, Fazenda Quero-Quero; 10 Oct. 2016; L. A. Medeiros leg.; pitfall; CEMT • 5 specs; Porto Nacional; 5 Oct. 2005; M. S. Matos leg.; agrotins; CEMT. COLOMBIA – Cesar • 1 spec.; Valledupar, La Arcadia; 10 ° 21 ′ N, 73 ° 23 ′ W; elev. 160.6 m; 13 Aug. 2012; S. Montoya leg.; forest; t. exc; CEMT. – Meta • 2 specs; San Martín, Hacienda Tocancipa, Sabana Bosque; elev. 330 m; Jan. 2000, J. Noriega leg.; CEMT. MEXICO – Veracruz • 1 spec.; 11 km W of Palma Sola, Ravine of Río Platanar; 19 ° 46 ′ N, 96 ° 25 ′ W; elev. 120 m; 23 Aug. – 3 Sep. 1976; Edmonds and Kohlmann leg.; on road to plan de las Hayas; CEMT • 11 specs; Los Tuxtlas; Oct. 1997; Vaz-de-Mello leg.; CEMT • 1 spec.; Plan de Hidalgo; elev. 200 m; 30 Jul. 1997; R. Sánchez and A. Díaz leg.; maizal, trampa excremento de vaca [cornfiel, trap with cow feces]; CEMT. PARAGUAY – Gran Asunción • 1 spec.; Asunción; 30 Nov. 1989; G. Arriágada leg.; CEMT. URUGUAY – Cerro Largo • 1 spec.; Banãdo de Medina, Piedras Sueltas Farm; 32 ° 21 ′ 03 ″ S, 53 ° 58 ′ 54 ″ W; 13 – 20 Dec. 2011; R. M. Moraes leg.; preserved area; pitfall; CEMT • 1 spec.; Melo, La Invernada farm; 13 – 20 Dec. 2011; R. M. Moraes leg.; pitfall trap (human + pig dung); CEMT. – Rivera • 1 spec.; Rivera, COFUSA-Compañia Florestal Uruguaya S. A.; 3 – 11 Oct. 2011; R. M. Moraes leg.; pitfall baited (human + pig dung); CEMT • 3 specs; same data as for preceding; 30 ° 57 ′ 36 ″ S, 55 ° 29 ′ 53 ″ W; CEMT • 6 specs; Rivera, Fazenda Bodegas Cacau; 30 ° 58 ′ 19 ″ S, 55 ° 26 ′ 40 ″ W; 3 – 11 Oct. 2011; R. M. Moraes leg.; FIT; CEMT • 1 spec.; Vichadero, El Pino Farm.; 21 – 27 Nov. 2011; R. M. Moraes leg.; pitfall trap baited (human + pig dung); CEMT • 1 spec.; Vichadero, Santa Adelaida Farm.; 31 ° 41 ′ 05 ″ S, 54 ° 39 ′ 42 ″ W; 3 – 11 Oct. 2011; R. M. Moraes leg.; pitfall trap (human + pig dung); CEMT. VENEZUELA – Yaracuy • 3 specs; Bolívar Aroa; 10 ° 00 ′ 00 ″ N, 68 ° 00 ′ 00 ″ W; elev. 459 m; 19 Jul. 2009; M. Asmüssen and P. Colmenares leg.; human feces; CEMT. UNKNOWN COUNTRY • 1 spec.; no data; INPA.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF4FFA2FF61FC97FEBAFCA3.taxon	materials_examined	BRAZIL – Distrito Federal • 2 specs; Brasilia, Fazenda água limpa; 15 ° 56 ′ 40 ″ S, 47 ° 55 ′ 38 ″ W; 10 Dec. 2017; Y. Ferreira leg.; pasture; pitfall with cow dung; CEMT. – Espírito Santo • 1 spec.; Venda Nova do Imigrante; 14 Jan. 2000; Lopes-Andrade and Vaz-de-Mello leg.; CEMT • 3 specs; same data as for preceding; Feb. 2010; F. Falqueto leg.; CEMT. – Goiás • 1 spec.; Bela Vista de Goiás, Cristianópolis, Fazenda Arapuca Velha; 21 Sep. 1993; Bankovics leg.; Cerrado; at light; CEMT • 1 spec.; Rio Verde; Sep. 1997; J. Carlos leg.; CEMT. – Maranhão • 2 specs; Mirador, P. E. Mirador, Base Geraldina; 24 Dec. 2006; R. O. Souza and J. C. Silva leg.; light; CEMT. – Mato Grosso • 1 spec.; Água Boa, Rancho Araguaia; 13 ° 56 ′ 40 ″ S, 52 ° 06 ′ 10 ″ W; 24 Dec. 2017; Mota and Schutz leg.; CEMT • 1 spec.; Alta Floresta, CEPLAC; 4 Sep. 2009; V. Gonçalves leg.; CEMT • 1 spec.; Alta Floresta, Chacara Recanto das Orquídeas; 3 Oct. 2011; A. Petini and Bevell leg.; CEMT • 2 specs; Chapada dos Guimarães; 15 ° 24 ′ 31 ″ S, 55 ° 45 ′ 53 ″ W; 2 Dec. 2015; A. Frolov and L. Akhemetova leg.; CEMT • 7 specs; Cotriguaçu, Fazenda São Nicolau; 14 Nov. 2007; O. Peres-Filho leg.; light trap; CEMT • 6 specs; same data as for preceding; 15 Nov. 2007; CEMT • 1 spec.; same data as for preceding; 15 Dec. 2007; light trap 01; CEMT • 2 specs; same data as for preceding; Oct. 2014; R. Stofel leg.; CEMT • 2 specs; same data as for preceding; Nov. 2007; O. Peres-Filho leg.; light; CEMT • 2 specs; Cotriguaçu, Fazenda São Nicolau; 9 ° 51 ′ 18 ″ S, 58 ° 14 ′ 57 ″ W; elev. 220 m; 14 Dec. 2009; Vaz-de-Mello leg.; light, manual; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 14 Dec. 2007; O. Peres-Filho leg.; CEMT • 1 spec.; same data as for preceding; light trap; CEMT • 1 spec.; same data as for preceding; 15 Dec. 2007; CEMT • 11 specs; same data as for preceding; light trap; CEMT • 1 spec.; same data as for preceding; 31 Oct. 2007; CEMT • 8 specs; same data as for preceding; 18 Oct. 2014; W. Chamorro leg.; CEMT • 2 specs; Cuiabá; 15 ° 16 ′ 45 ″ S, 56 ° 00 ′ 06 ″ W; 7 Dec. 2015; A. Frolov and L. Akhemetova leg.; CEMT • 2 specs; same data as for preceding; Fazenda Mutuca; 15 ° 19 ′ 31 ″ S, 55 ° 58 ′ 3 ″ W; elev. 285 m; Nov. 2015; R. V. Nunes and A. Frolov leg.; light; CEMT • 5 specs; same data as for preceding; Recanto Siriema; 15 ° 35 ′ 32 ″ S, 56 ° 01 ′ 38 ″ W; 4 Nov. 2014; R. V. Nunes and Becker leg.; light; CEMT • 2 specs; Diamantino; Jan. 2001; A. M. Bello leg.; CEMT • 2 specs; Diamantino, Alto Rio Arinos; Sep. 2001; E. Furtado leg.; CEMT • 1 spec.; Poconé, Pousada das Aráras; 3 Dec. 1998; J. L. Moreno and T. Mestre leg.; Pantanal; CEMT • 2 specs; Porto Estrela, EES das Araras; 15 ° 39 ′ 9 ″ S, 57 ° 12 ′ 53 ″ W; elev. 217 m; 8 – 12 Oct. 2011; Vaz-de-Mello leg.; light; CEMT. – Mato Grosso do Sul • 2 specs; Campo Grande; 1990 – 1992; I. Biachin leg.; CEMT • 1 spec.; Corumbá, Fazenda São Bento, Próx porto Jofre; 14 Apr. 2011; R. V. Nunes leg.; light; CEMT • 1 spec.; Guia lopes da Laguna; 15 Nov. 2007; A. Abot leg.; light trap; CEMT. – Minas Gerais • 2 specs; Caminho de São Thomé das Letras para Cruzilia; Sep. 1997; F. Frieiro-Costa leg.; CEMT • 3 specs; Cordisburgo; Jan. 1999; F. Z. Vaz Mello leg.; CEMT • 3 specs; Ijaci, Fazenda FAEPE; Nov. 2002; J. Louzada leg.; CEMT • 3 specs; Ipatinga; Nov. 1994; A. Bello leg.; CEMT • 1 spec.; Lavras-Ingaí, Próx Quedas do Rio Bonito; 21 Oct. 2016; J. Mariano leg.; CEMT • 2 specs; Lavras; 21 ° 19 ′ 55 ″ S, 44 ° 58 ′ 08 ″ W; 22 Mar. 2008; M. R. Santos and D. L. H. Takahashi leg.; CEMT • 2 specs; same data as for preceding; 21 ° 20 ′ 19 ″ S, 44 ° 58 ′ 35 ″ W; CEMT • 3 specs; Paracatu; Mar. 1997; S. Lourengo jr leg.; CEMT • 2 specs; Patos de Minas; Jan. 1970; P. S. F. Fiuza leg.; CEMT • 1 spec.; Santa Barbara; 8 Dec. 1994; J. C. Zanuncio leg.; CEMT • 1 spec.; same data as for preceding; 26 Jan. 1995; J. C. Zanuncio leg.; CEMT • 1 spec.; Viçosa; 20 Sep. 1993; Louzada and Silva leg.; CEMT • 1 spec.; same data as for preceding; 23 Sep. 1993; CEMT • 1 spec.; Viçosa; Universidade Federal de Viçosa; 14 Dec. 1999; Lopes-Andrade and Vaz-de-Mello leg.; CEMT. – Paraíba • 1 spec.; Rio Tinto; APA Barra do Rio Mamanguape; 6 ° 46 ′ 41 ″ S, 34 ° 55 ′ 37 ″ W; 23 Mar. 2012; Da-Bó leg.; CEMT. – Piauí • 1 spec.; Piripiri, P. N. Sete Cidades; 4 ° 06 ′ 27 ″ S, 41 ° 40 ′ 12 ″ W; elev. 165 m; 7 – 12 Feb. 2013; Vaz-de-Mello and Grossi leg.; Kerodon dung; CEMT. – Rio de Janeiro • 1 spec.; Angra dos Reis, Bracuí, lowland; 10 – 11 May 2002; Nessiamian leg.; UV light; CEMT • 1 spec.; Miguel Pereira; Nov. 1997; CEMT. – Rio Grande do Sul • 1 spec.; Santa Maria; Nov. 2014 – May 2015; J. Boscardin leg.; CEMT • 1 spec.; São Geronimo; 17 Dec. 1984; Frace leg.; CEMT. – São Paulo • 1 spec.; São Carlos; 11 Nov. 1993; J. Mendes leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF5FFA1FF61FCAAFD23FEC7.taxon	materials_examined	BRAZIL – Mato Grosso • 1 spec.; Água Boa, Rancho Araguaia; 13 ° 56 ′ 40 ″ S, 52 ° 06 ′ 10 ″ W; 24 Dec. 2007; Mota and Schutz leg.; CEMT • 2 specs; Alta Floresta, Ceplac; 4 Sep. 2009; V. Gonçalves leg.; Tapirus feces; CEMT • 1 spec.; Chapada dos Guimarães; 15 ° 24 ′ 31 ″ S, 55 ° 45 ′ 53 ″ W; 2 Dec. 2015; A. Frolov and L. Akhmetova leg.; light; CEMT • 5 specs; Cotriguaçu, Fazenda São Nicolau; 15 Jul. 2007; O. Peres-Filho leg.; light trap; CEMT • 2 specs; same data as for preceding; 15 Dec. 2007; CEMT • 2 specs; same data as for preceding; Nov. 2007; CEMT • 1 spec.; same data as for preceding; 9 ° 49 ′ 32 ″ S, 58 ° 15 ′ 46 ″ W; elev. 220 m; 10 Dec. 2009; Vaz-de-Mello leg.; Ficus plantation; CEMT • 1 spec.; same data as for preceding; 9 ° 50 ′ 19 ″ S, 58 ° 15 ′ 03 ″ W; 25 Oct. 2014; A. Asenjo leg.; light; CEMT • 1 spec.; same data as for preceding; 9 ° 51 ′ 20 ″ S, 58 ° 14 ′ 55 ″ W; Nov. 2014; R. V. Nunes and Asenjo leg.; light; CEMT • 6 specs; Cotriguaçu, Fazenda São Nicolau; 18 Oct. 2014; W. Chamorro leg.; CEMT • 2 specs; same data as for preceding; 95 ° 12 ′ 01 ″ S, 58 ° 14 ′ 55 ″ W; elev. 220 m; 16 – 20 Oct. 2012; F. Z. Vaz-de-Mello leg.; base camp, manually; CEMT • 1 spec.; Cuiabá, Pari; 6 Oct. 1988; auxiliadora [unreadable]; CEMT • 1 spec.; same data as for preceding; Recanto Siriema; 15 ° 35 ′ 32 ″ S, 56 ° 01 ′ 38 ″ W; 4 Nov. 2014; R. V. Nunes and Becker leg.; CEMT • 3 specs; Diamantino; Jan. 2001; A. M. Bello leg.; CEMT • 2 specs; Paranaita, Teles Pires; 9 ° 19 ′ 47 ″ S, 56 ° 47 ′ 49 ″ W; A. P. Benelli leg.; left margin, 33; CEMT. – Mato Grosso do Sul • 1 spec.; Dourados; 22 Oct. 2005; M. Milo leg.; CEMT. – Minas Gerais • 1 spec.; Buenópolis, Parque Estadual da Serra do Cabral, Almirante; 15 Apr. 2008, M. F. Sulza leg.; clean field; CEMT • 2 specs; same data as for preceding; Buriti; 14 Apr. 2008; CEMT • 2 specs; Ijaci; Nov. 2002; L. Julio leg.; CEMT • 1 spec.; Lavras, Próx. Poço Bonito; Dec. 2001; Vaz-de-Mello leg.; CEMT • 2 specs; Paracatu; Mar. 1998; S. Lourengo jr leg.; CEMT • 2 specs; Recreio; Jan. 2000; Lopes-Andrade leg.; CEMT. URUGUAY – Rivera • 1 spec.; Rivera, Bañado de Medina, Piedras Sueltas Farm; 32 ° 18 ′ 21 ″ S, 53 ° 57 ′ 30 ″ W; 13 – 20 Dec. 2011; R. M. Moraes leg.; pitfall human + pig dung; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF5FFA1FF61FCAAFD23FEC7.taxon	discussion	Collecting information Specimens were collected at altitudes from 60 to 950 metres. The collecting methods vary. Flight and light traps are effective, as well as traps baited with human, pig, bovine and Tapirus feces. Species belonging to the group can be found in dry and wet forests under dead trees and forest litter (Stebnicka 2006 b). Remarks Ataenius complicatus group comprises 16 species, five of these are found in Brazil. A key for species of this group can be found in Stebnicka (2006 b).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF6FFA1FF61F9E8FC28F894.taxon	materials_examined	BRAZIL – Amazonas • 1 spec.; Manaus, Taruma-Mirim, nr Manaus; 28 Oct. 1981; INPA. – Roraima • 1 spec.; Rio Uraricoera, Ilha de Maraca; 2 – 13 May 1987; J. A. Rafael, J. E. B. Brasil and L. S. Aquino leg.; suspended trap; CEMT • 3 specs; same data as for preceding; INPA.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF6FFA1FF61FCD2FCA5FB0D.taxon	materials_examined	BRAZIL – Minas Gerais • 1 spec.; Cordisburgo, Fazenda Pontinha; Jul. 1994; De Mello leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF6FFA1FF61FCD2FCA5FB0D.taxon	discussion	Collecting information Specimens were collected under logs in rainforest, and at light (Stebnicka 2001 a). Remarks Ataenius crenator group comprises five species, two are found in Brazil (Stebnicka 2001 c). A key for species of this group can be found in Stebnicka (2001 c).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF6FFA1FD5BFE01FE8AFD4A.taxon	description	Figs 9 E – F, 41 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF6FFA1FD5BFE01FE8AFD4A.taxon	diagnosis	Diagnosis Length 5.0 – 7.0 mm. Body large, broad, elongate, glossy piceous. Head with clypeus punctate, punctures finer than those of frons; emarginate, with weak but distinct transverse wrinkles. Pronotum laterally and posteriorly with marginal line; lateral margins fringed with setae; surface with punctures fine and coarse, densely distributed. Elytra with striae deeply punctate; punctures circular, crenating lateral margins of interstriae; interstriae convex. Metafemur with incomplete posterior line; metatibiae with acuminate accessory spine.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF6FFA1FDA6FB78FCC4F9AC.taxon	description	Figs 10 E, 41 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF6FFA1FDA6FB78FCC4F9AC.taxon	diagnosis	Diagnosis Length 3.0 – 6.0 mm. Body cylindrical, glabrous or not, shagreened or not, normally covered with argillaceous coating which may difficult observing some structures. Head with coarse longitudinally elongated punctures. Pronotum with punctures of variable sizes; lateral margins always fringed with setae. Elytra glabrous or not, shagreened or not. Ventral surface shagreened covered with coarse punctures. Metatibiae without apical accessory spine.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF6FFA0FF61F8D2FEE0FE3E.taxon	materials_examined	BRAZIL – Paraíba • 1 spec.; São José dos Cordeiros, Fazenda Almas; 4 – 5 May 2003; Valderez and Rothéa leg.; light trap; CEMT. – Piauí • 1 spec.; Piracuruca, Parna Sete Cidades, Posto ICMBio; 4 ° 05 ′ 57 ″ S, 41 ° 42 ′ 34 ″ W; 11 – 14 Dec. 2012; F. Limeira-de-Oliveira and J. S. Junior Pinto leg.; light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF7FFA7FF61FE29FB6AFD12.taxon	materials_examined	BRAZIL – 1 spec.; Est. rio Mamangua; 25 – 27 Oct. 2000; CEMT. – Acre • 1 spec.; Rio Branco; CEMT • 1 spec.; same data as for preceding; Embrapa, Jun. 1987; M. Fazolin leg.; CEMT. – Alagoas • 2 specs; Penedo, APA Murituba Peixe; 19 – 25 Jul. 2019; A. B. Farias and Lasda leg.; CEMT. – Bahia • 1 spec.; Camaru, 13 ° 58 ′ S, 39 ° 10 ′ W; Dec. 2011; C. M. P. Leite leg.; pasture; CEMT • 2 specs; Caravelas; 25 Mar. 1997; Anjos and Silveira leg.; CEMT • 2 specs; Mucuri; 21 Jan. 1997; Anjos and Silveira leg.; CEMT • 1 spec.; same data as for preceding; 24 Nov. 1997; CEMT • 1 spec.; Presidente Tancredo Neves; 13 ° 23 ′ 42 ″ S, 39 ° 18 ′ 35 ″ W; Jan. 2011; C. M. P. Leite leg.; 253 mols. parc 57; pitfall; CEMT • 1 spec.; Una, REBIO de Una; 15 ° 10 ′ 19 ″ S, 39 ° 3 ′ 40 ″ W; 44 m; 10 Aug. 2016; APMS, DMT and APP leg.; light; CEMT • 1 sec.; same data as for preceding; 15 ° 10 ′ 32 ″ S, 39 ° 03 ′ 22 ″ W; elev. 17 m; 11 Aug. 2016; Rio Maruim; CEMT. – Espírito Santo • 1 spec.; Linhares; Jan. 2005; J. Santos leg.; CEMT. – Ceará • 1 spec.; Pacatuba, Caracanga; 2 Feb. 1986; P. M. B. Carvalho leg.; CEMT. – Maranhão • 1 spec.; Carolina, Fazenda Cincorá; 17 – 22 Oct. 2009; F. Limeira-de-Oliveira, R. O. Souza and M. B. Aguiar Neto leg.; light trap; CEMT. – Mato Grosso • 1 spec.; Alta Floresta, Chacará Recanto das Orquídeas; 3 Oct. 2011; P. A. Bevel; CEMT • 1 spec.; Cuiabá, Boa Esperança, 15 ° 36 ′ 31 ″ S, 56 ° 03 ′ 19 ″ W; 28 Oct. 2021; F. Z. Vaz-de-Mello leg.; light; CEMT • 1 spec.; Cuiabá, 15 ° 36 ′ 39 ″ S, 56 ° 03 ′ 33 ″ W; 3 Dec. 2008; Mota and Schutz leg.; CEMT • 1 spec.; Cuiabá, Distrito Guia, Fazenda Santhidi; 15 ° 28 ′ 47 ″ S, 56 ° 7 ′ 33 ″ W; elev. 180 m; 11 Dec. 2009; L. R. Silva leg.; CEMT • 1 spec.; Cuiabá, Morada do Ouro; 15 ° 34 ′ 02 ″ S, 26 ° 03 ′ 25 ″ W; May 2021 – Nov. 2021; L. A. C. S. Santos leg.; CEMT • 1 spec.; Nobres, Distrito coqueiral; 26 Jun. 2010; D. Nonato leg.; manually; CEMT • 1 spec.; Poconé; Jul. 2000; J. Lopes leg.; CEMT • 1 spec.; Poconé, Parque Nacional do Pantanal, Sede do Parque; 17 ° 50739 ′ S, 57 ° 24170 ′ W; 15 Nov. 2011; J. L. Silva leg.; light; CEMT • 1 spec.; Poconé, RPPN SESC, Base UFMT; 16 ° 29 ′ 52 ″ S, 56 ° 24 ′ 45 ″ W; 19 – 23 Jul. 2011; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; SESC Pantanal; 28 Jan. 1998; forest; light trap; CEMT • 1 spec.; same data as for preceding; 29 Jan. 1998; acuri palm tree; light trap; CEMT • 1 spec.; same data as for preceding; 28 Feb. 1998; forest; CEMT • 1 spec.; same data as for preceding; 28 Mar. 1998; CEMT • 1 spec.; same data as for preceding; 23 Apr. 1998; CEMT • 1 spec.; same data as for preceding; 26 Apr. 1998; CEMT • 1 spec.; same data as for preceding; 29 Dec. 1997; CEMT • 1 spec.; Sesc Pantanal; 14 May 2002; light trap; CEMT • 1 spec.; same data as for preceding; 4 Jul. 2002; CEMT • 1 spec.; same data as for preceding; 6 Sep. 2002; CEMT • 1 spec.; same data as for preceding; 5 Jul. 2003; CEMT • 1 spec.; same data as for preceding; 7 Jul. 2003; CEMT • 1 spec.; same data as for preceding; 5 Aug. 2003; CEMT • 1 spec.; same data as for preceding; 15 Sep. 2004; CEMT • 1 spec.; Poconé, Pousada Araras; 3 Dec. 1998; J. L. Moreno and T. Mestre leg.; CEMT • 1 spec.; Santo Antônio do Leverger, Porto de fora; 16 ° 05 ′ 17 ″ S, 55 ° 48 ′ 41 ″ W; 29 – 31 Dec. 2019; B. Bordin, B. Queiroz and M. Pereira leg.; CEMT • 1 spec.; Várzea Grande, Sítio Toca da Coruja; 15 ° 33 ′ 07 ″ S, 56 ° 19 ′ 07 ″ W; elev. 230 m; 18 Oct. 2020; A. Petini-Belini leg.; CEMT • 1 spec.; Várzea Grande; 15 ° 40 ′ S, 56 ° 10 ′ W; Nov. 2021 – Feb. 2022; J. Arias leg.; light CEMT. – Mato Grosso do Sul • 1 spec.; Corumbá, Fazenda São Bento, Prox Porto Jofre; 14 Apr. 2011; R. V. Nunes leg.; light; CEMT • 1 spec.; Guia Lopes da Laguna; 5 Mar. 2008; A. R. Abot leg.; light trap; CEMT. – Minas Gerais • 4 specs; Águas Vermelhas; Dec. 1997; A. Bello and F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; Dec. 1997; A. Bello leg.; CEMT. – Pernambuco • 1 spec.; Igarassu; 27 Apr. 2007; Costa leg.; CEMT • 1 spec.; Carolina; 9 ° 08 ′ 14 ″ S, 40 ° 18 ′ 7 ″ W; elev. 365 m; 11 Nov. 2015; J. B. G. Santos leg.; CEMT. – Pará • 1 spec.; Vigia, Fazenda Mangue Seco; 20 – 22 Jun. 2010; CEMT. – Paraíba • 1 spec.; São José dos Cordeiros, Fazenda Almas; 4 – 5 May 2003; Valderez and Rothéa leg.; light; CEMT. – Roraima • 1 spec.; Amajari, EsEc Maracá; 3 ° 20 ′ 59 ″ S, 61 ° 25 ′ 22 ″ W; elev. 83 m; 24 Mar. 2016; A. P. M. Santos et al. leg.; CEMT • 1 spec.; Alto Alegre, lavrado; 2 ° 53 ′ 45 ″ S, 60 ° 39 ′ 42 ″ W; elev. 81 m; 5 May 2005; E. G. F. Morais leg.; CEMT • 1 spec.; Rio Uraricoera, Ilha de Maraca; 2 – 13 May 1987; J. A. Rafael, J. E. B Brasil and L. S. Aquino leg.; suspended trap; CEMT • 2 specs; same data as for the preceding; INPA • 1 spec.; same data as for preceding; 21 – 30 Nov. 1987; J. A. Rafael et al. leg.; Malaise trap; INPA. – Santa Catarina • 1 spec.; Florianópolis; 27 ° 40 ′ S, 48 ° 31 ′ W; Jan. 2009; Hernández and Heusi leg.; frontal dune; pitfall; CEMT. – São Paulo • 1 spec.; Anhembi, ESALQ / USP; E. N. L. Ferreira leg.; CEMT • 1 spec.; Cerqueira César; Dec. 1992; J. Carlos leg.; CEMT. MEXICO – Veracruz • 1 spec.; Los Tuxtlas; Oct. 1997; Vaz-de-Mello leg.; CEMT • 1 spec.; Acajete, Plan de Sedeño; elev. 1950 m; 28 May 2004; Ordano and Vaz-de-Mello leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF0FFA7FF61FD54FB31FBB5.taxon	materials_examined	BRAZIL – Mato Grosso • 3 specs; Diamantino, Alto Rio Arinos; Nov. 2001; E. Furtado leg.; CEMT • 3 specs; Sesc Pantanal, 15 Nov. 2001; light trap; CEMT • 11 specs; same data as for preceding; 14 May 2002; CEMT • 5 specs; same data as for preceding; 9 Aug. 2002; CEMT • 1 spec.; same data as for preceding; 7 Jul. 2003; CEMT • 1 spec.; same data as for preceding; 8 Jul. 2002; CEMT • 15 specs; same data as for preceding; 5 Aug. 2003; CEMT. – Minas Gerais • 1 spec.; Cordisburgo; Dec. 1993; Zagury leg.; CEMT • 1 spec.; same data as for preceding; Jan. 1997; Vaz-de-Mello leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF0FFA5FF61FBB0FAC1FC61.taxon	materials_examined	BRAZIL – Acre • 3 specs; Rio Branco, PZ-UFAC; Feb. 1997; F. Vaz-de-Mello leg.; CEMT • 3 specs; Rio Branco; Nov. 2000; M. A. Oliveira leg.; CEMT. – Amazonas • 1 spec.; Barcelos; 15 Jan. 1978; N. D. Penny leg.; CEMT • 1 spec,; same data as for preceding; INPA • 1 spec.; Benjamin Constant, Capim imperial; 9 Mar. 2004; P. H. Silva leg.; CEMT • 1 spec.; Itacoatiara, Fazenda Aruana, Am 010, Km 215; Mar. 1997; INPA • 7 specs; same data as for preceding; 13 – 16 Feb. 1991; C. S. Motta, R. Andreazze and W. Kawazoe leg.; UV light; INPA • 1 spec.; same data as for preceding; 13 – 16 Mar. 1991; C. S. Motta, F. J. A. Peralta and Ronchi-Teles leg,; INPA • 17 specs; same data as for preceding; 14 – 15 May 1991; C. S. Motta et al. leg.; CEMT • 1 spec.; same data as for preceding; 16 – 17 May 1991; INPA • 1 spec.; Manaus; 6 Dec. 1977; B. C. Ratcliffe leg.; INPA • 1 spec.; Manaus, Bairro Aleixo; 3 ° 04 ′ 40 ″ S, 59 ° 59 ′ 13 ″ W; Res. Espanha; 8 Sep. 2015; F. S. Godoi leg.; light; CEMT • 65 specs; same data as for preceding; 24 Apr. 2017; CEMT • 52 specs; same data as for preceding; 1 May 2017; CEMT • 11 specs; same data as for preceding; 8 Sep. 2017; CEMT • 1 spec.; São Gabriel da Cachoeira, Querari Pelotão; 1 ° 05 ′ N, 69 ° 51 ′ W, 7 – 23 Apr. 1993; C. S. Motta et al leg.; INPA • 1 spec.; same data as for preceding; 3 – 4 May 1993; INPA. – Bahia • 1 spec.; Caravelas; 20 Jan. 1997; Anjos and Silveir leg.; CEMT • 1 spec.; Ilhéus; 14 ° 44 ′ 32 ″ S, 39 ° 05 ′ 14 ″ W; Oct. 2011; C. M. P. Leite leg.; 52 mosl Parc 68; pitfall; CEMT • 2 specs; Ilhéus; 14 ° 43 ′ 23 ″ S, 39 ° 07 ′ 37 ″ W; Oct. 2011; C. M. P. Leite leg.; 52. mols. Parc 21; pitfall; CEMT • 4 specs; Pres. Tancredo Neves; 13 ° 22 ′ 17 ″ S, 39 ° 17 ′ 41 ″ W; Jan. 2011; C. M. P. Leite leg.; 253 mols Parc 20; pitfall; CEMT • 1 spec.; same data as for preceding; 13 ° 22 ′ S, 39 ° 19 ′ W; pasture; CEMT • 1 spec.; Mucuri; 24 Oct. 1997; Anjos and Oliveira leg.; CEMT. – Ceará • 1 spec.; Ubajara, P. N. Ubajara, Trilha Aracitum, Rio das Minas; 3 ° 50 ′ 21 ″ S, 40 ° 54 ′ 23 ″ W; elev. 880 m; 13 Feb. 2013; D. M. Takiya and A. P. M. Santos leg.; plus light; CEMT. – Maranhão • 1 spec.; Bom Jardim, REBIO-Res. Biol. Gurupi, Basee; 5 – 15 Jun. 2010; J. A. Silva et al. leg.; light trap; CEMT. – Mato Grosso • 1 spec.; Res. Papagaio, BR- 364, Km 555; 2 Oct. 1984; E. Binda leg.; INPA • 26 specs; Alta Floresta, Chacará Recanto das Oquideas; 3 Oct. 2011; A. Petini-Bevell leg.; CEMT • 1 spec.; Chapada dos Guimarães, Casa do Mel; 15 ° 22 ′ 53 ″ S, 55 ° 50 ′ 35 ″ W; elev. 520 m; Dec. 2015; R. V. Nunes, A. Frolov and L. G. Nunes leg.; light; CEMT • 1 spec.; Chapada dos Guimarães, Rio Claro; 20 Apr. 1984; Nelsina and Costa leg.; CEMT • 3 specs; Chapada dos Guimarães, Comunidade Jão Carro; 3 Nov. 2015; M. A. S. Serrano leg.; CEMT • 2 specs; Cotriguaçú, Fazenda São Nicolau; 14 Feb. 2007; O. Peres-Filho leg.; CEMT • 23 specs; same data as for preceding; 31 Oct. 2007; light trap; CEMT • 3 specs; same data as for preceding; 4 Nov. 2007; CEMT • 2 specs; same data as for preceding; Nov. 2007; CEMT • 15 specs; same data as for preceding; 15 Dec. 2007; CEMT • 2 specs; same data as for preceding; 14 Apr. 2008; CEMT • 2 specs; same data as for preceding; 9 ° 51 ′ 20 ″ S, 58 ° 14 ′ 55 ″ W; Nov. 2014; R. V. Nunes and A. Asenjo leg.; light; CEMT • 9 specs; Cotriguaçú, Fazenda São Nicolau; 30 Oct. 2014; W. Chamorro leg.; black light trap; CEMT • 1 spec.; same data as for preceding; Oct. 2014; R. V. Nunes leg.; CEMT • 1 spec.; same data as for preceding; Oct. 2014; R. Stofel leg.; CEMT • 1 spec.; Cotriguaçú, Fazenda São Nicolau, base camp; 9 ° 12 ′ 01 ″ S, 58 ° 14 ′ 55 ″ W; elev. 220 m; 16 – 20 Oct. 2012; F. Z. Vaz-de-Mello leg.; manually; CEMT • 1 spec.; Cuiabá; May 2010; Vaz-de-Mello leg.; CEMT • 1 spec.; Cuiabá, Fazenda Mutuca; 15 ° 19 ′ 31 ″ S, 55 ° 58 ′ 3 ″ W; elev. 285 m; Nov. 2015; R. V. Nunes and A. Frolov leg.; light; CEMT • 1 spec.; Diamantino; Jan. 2001; A. M. Bello leg.; CEMT • 1 spec.; Juara; 11 ° 06 ′ 33 ″ S, 57 ° 45 ′ 20 ″ W; R. Santos leg.; M 2 l 4; CEMT • 4 specs; Pixaim, Pantanal; 3 Dec. 1998; J. L. Moreno and T. Mestre leg.; CEMT • 1 spec.; Poconé; 29 Dec. 1997; forest; light trap; CEMT • 11 specs; same data as for preceding; Pousada Araras, Pantanal; 3 Dec. 1998; J. L. Moreno and T. Mestre leg.; CEMT • 1 spec.; Sesc Pantanal; 13 Oct. 2001; light trap; CEMT • 2 specs; same data as for preceding; 16 Oct. 2001; CEMT • 1 spec.; same data as for preceding; 16 Dec. 2001; CEMT • 7 specs; same data as for preceding; 14 May 2002; CEMT • 1 spec.; same data as for preceding; 4 Jul. 2002; CEMT • 2 specs; same data as for preceding; 5 Jul. 2002; CEMT • 1 spec.; same data as for preceding; 11 Jul. 2002; CEMT • 2 specs; same data as for preceding; 6 Sep. 2002; CEMT • 5 specs; same data as for preceding; 5 Aug. 2003; CEMT • 1 spec.; Poconé, Acuri palme tree; 28 Feb. 1998; light trap; CEMT • 1 spec.; Porto Estrela, EES das Araras, base camp; 15 ° 39 ′ 09 ″ S, 57 ° 12 ′ 53 ″ W; elev. 217 m; 8 – 12 Oct. 2011; Vaz-de-Mello leg.; light; CEMT • 1 spec.; Santo Antônio do Leverger, Pesquiro Casa Branca; 6 Apr. 2015; H. D. Bianchini leg.; CEMT • 1 spec.; Trindade, Polo noroeste; 1 Oct. 1984; E. Binda leg.; light; CEMT • 5 specs; same data of the preceding; INPA • 1 spec.; same data as for preceding; J. Elias leg.; CEMT • 2 specs; same data as for preceding; INPA • 1 spec.; Vila Bela, Margem Guaporé; 1 Oct. 1984; Mancolino and Sebastião leg.; light; CEMT. – Mato Grosso do Sul • 1 spec.; Corumbá; Fazenda são Bento; CEMT • 3 specs; Corumbá, Fazenda são Bento, Pox Porto Jofre; 14 Apr. 2011; R. V. Nunes leg.; light trap; CEMT • 3 specs; Corumbá, Fazenda São Bento; 2010. F. R. Tortato leg.; CEMT • 2 specs; Passo do Lontra; Nov. 2000; J. Raizer leg.; CEMT • 1 spec.; Santo Antônio do Leverger, São Vicente da Serra; 15 ° 49 ′ 42 ″ S, 55 ° 25 ′ 11 ″ W; 15 – 17 Dec. 2010; Tissiani and Vaz-de-Mello leg.; pasture; human dung; CEMT • 1 spec.; Três Lagoas, Horto Moeda; Feb. 2008; Uehara-Prado leg.; CEMT. – Minas Gerais • 2 specs; Cordisburgo, Fazenda Pontinha; Dec. 1998; Vaz-de-Mello leg.; CEMT • 1 spec.; Ipatinga; Nov. 1991; E. Grossi leg.; CEMT • 1 spec.; [Coronel] Pacheco, água limpa; 2 Mar. 1945; Gomes and Vasco leg.; CEMT • 1 spec.; Itumirim; 21 ° 20 ′ 24 ″ S, 44 ° 48 ′ 6 ″ W; 19 Oct. 2016; Vieira, F. Z. Vaz-de-Mello and J. Louzada leg.; light; CEMT • 1 spec.; Lavras, Campus UFLA; 15 – 21 Oct. 2016; P. Grossi and F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Lavras, Ponte Nova; 30 Oct. 1979; Planalsucar leg.; CEMT • 1 spec.; Viçosa; Nov. 1996; Vaz-de-Mello, Hardy and Harrison leg.; CEMT • 2 specs; Teixeiras; Nov. 1996; Hardy and Harrison leg.; CEMT. – Pará • 1 spec.; Cachoeira do Arari; 20 Jul. 1983; A. I. Henrique leg.; INPA • 1 spec.; Ilha Marajo; 23 Jan. 1978; INPA • 2 specs; same data as for preceding; 23 May 1978; CEMT • 17 specs; same data as for preceding; INPA • 19 specs; same data as for preceding; D. Domingo leg.; INPA • 2 specs; Jacareacanga, UHET Teles Pires; 9 ° 20 ′ 43 ″ S, 56 ° 46 ′ 31 ″ W; 28 Oct. 2011; Petini-Benelli leg.; CEMT • 1 spec.; same data as for preceding; Agric. M. B 199 T 1 P 3 ARM 1; CEMT • 2 specs; Tailandia; 7 Aug. 2014; J. Malta and G. Santos leg.; trap 02 - D 21; CEMT. – Paraíba • Areia, Mata Pau de Ferro; 2 Mar. 2000; M. Avany leg.; light trap; CEMT. – Rio de Janeiro • 9 spec.; Angra dos Reis, Bracuí, lowland; 10 – 11 May 2002; Nessimian leg.; UV light; CEMT • 1 spec.; Guapimirim; 20 Oct. 1988; CEMT • 1 spec.; Miguel Pereira; Jan. 1998; J. Carlos leg.; CEMT • 3 specs; same data as for preceding; Mar. 1998; J. Carlos leg.; CEMT • 1 spec.; Nova Friburgo; Nov. 2002; E. Grossi leg.; CEMT • 1 spec.; Nova Friburgo, Macaé de Cima; Jan. 1999; P. Grossi leg.; CEMT. – Rondônia • 6 specs; 62 km S of Ariquemes; Feb. 1997; Vulinec and Mello leg.; CEMT • 1 spec.; Ariquemes, Fazenda Rancho Grande, 60 km SW of Cidade; Feb. 1997; F. Z. Vaz-de-Mello leg.; CEMT • 3 specs; Cacaulandia, Fazenda Rancho Grande; Feb. 1997; Vaz-de-Mello leg.; CEMT • 1 spec.; Porto Velho, Rio das Garças, Bom Jesus; 8 ° 49 ′ 48 ″ S, 63 ° 46 ′ 45 ″ W; 22 Oct. 2017; D. C. Santos and K. K. C. Silva leg.; FIT; CEMT. – Roraima • 3 specs; Caracaraí; Jul. 1997; Vaz-de-Mello leg.; CEMT • 2 specs; Rio Uraricoera, Ilha de Maraca; 2 – 13 May 1987; J. A. Rafael, J. E. B. Brasil and L. S. Aquino leg.; suspended trap; CEMT • 4 specs; same data as for preceding; INPA • 1 spec.; same data as for preceding; 18 – 28 Aug. 1987; J. A. Rafael et al. leg.; light trap; INPA • 1 spec.; same data as for preceding; 21 – 30 Nov. 1987; Malaise trap; INPA. – Santa Catarina • 1 spec.; Florianópolis, Praia Pantano do Sul; 27 ° 47 ′ S, 48 ° 31 ′ W; Jan. 2009; Hernandez and Heusi leg.; CEMT. – São Paulo • 2 specs; Cerqueira César; Dec. 1993; J. L. Carlos leg.; CEMT • 1 spec.; same data as for preceding; Dec. 1996; CEMT. – Tocantins • 1 spec.; Araguaína, Cimbá, BR- 70; Dec. 2015; V. E. Sandoval leg.; CEMT. FRENCH GUIANA – 3 specs; Cayenne, Montagne des Chevaux (MCV); 4 ° 44 ′ 56 ″ N, 52 ° 26 ′ 28 ″ W; elev. 75 m; 27 Jan. 2013; SAEG leg.; light (GEML); CEMT • 1 spec.; same data as for preceding; 16 Feb. 2013; SAEG leg.; CEMT. PERU – Loreto • 1 spec.; 21 km W of Iquitos; 10 Jul. 2001; M. Curoe and Callegari leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF2FFA5FF61FC64FB20FA73.taxon	materials_examined	BRAZIL – Amazonas • 5 specs; Iranduba, Rio Solimões, Ilha de Marchantaria, lowland 3 ° 15 ′ S, 59 ° 58 ′ W; 12 – 14 Nov. 1980; J. Adis leg.; standing crop; CEMT • 12 specs. same data as for preceding; INPA • 4 specs; same data as for preceding; 24 Feb. 1981; light; CEMT • 25 specs; same data as for preceding; INPA • 1 spec.; same data as for preceding; 29 Apr. 1981; CEMT • 2 specs; same data as for preceding; INPA • 2 specs; same data as for preceding; 16 Jul. 1981; INPA • 1 spec.; same data as for preceding; 14 Aug. 1981; INPA • 1 spec.; same data as for preceding; 18 May 1981; CEMT. – Mato Grosso • 1 spec.; Sesc Pantanal; 5 Jul. 2003; light trap; CEMT. – Mato Grosso do Sul • 3 specs; Corumbá, Fazenda São Bento, Prox Porto Jofre; 14 Apr. 2011; R. V. Nunes leg.; light; CEMT. ECUADOR – Napo • 1 spec.; elev. 250 m; 8 Jun. 1997; G. Onore leg.; light trap; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF2FFA5FF61FA75FBC1F9C4.taxon	materials_examined	BRAZIL – Mato Grosso • 1 spec.; Cuiabá, Boa Esperança; 24 – 27 Oct. 2016; F. Z. Vaz de Mello leg.; manually; CEMT • 1 spec.; Sesc Pantanal; 14 May 2002; light trap; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF2FFA5FF61F903FCF5F8E3.taxon	materials_examined	BRAZIL – Bahia • 5 specs; Caravelas; Mar. 1996; R. Diniz leg.; CEMT. – Espírito Santo • 2 specs; Pedro Canario; May 1997; R. Diniz; leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF2FFA4FF61F8E5FCF9FD8E.taxon	materials_examined	BRAZIL – Amazonas • 1 spec.; Manaus, Feb. 1992; B. Nelson leg.; INPA. – Mato Grosso • 3 specs; Cuiabá; 15 ° 16 ′ 45 ″ S, 56 ° 0 ′ 6 ″ W; 7 Dec. 2015; A. Frolov and L. Akhmetova leg.; CEMT • 1 spec.; Varzea Grande, Bairro Almeida, urban area next to Rio Cuiabá; 29 Nov. 2015; R. E. Vicente leg.; CEMT • 1 spec.; Sesc Pantanal; 14 May 2002; light trap; CEMT • 1 spec.; Poconé; 29 Jan. 1998; acuri palm tree; light trap; CEMT • 1 spec.; same data as for preceding; Jul. 2000; J. Lopes leg.; CEMT. – Mato Grosso do Sul • 3 specs; Corumbá, Fazenda São Bento, Prox Porto Jofre; 14 Apr. 2011; R. V. Nunes leg.; light; CEMT. – Roraima • 1 spec.; Rio Uraricoera, Ilha de Maraca; 2 – 13 May 1987; J. A. Rafael, J. E. B Brasil and L. S. Aquino leg.; suspended trap; INPA • 2 specs; same data as for preceding; 21 – 30 Nov. 1987; J. A. Rafael et al. leg.; Malaise trap; CEMT, INPA.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF3FFA4FF61FDF9FDAAF985.taxon	materials_examined	BRAZIL – Amapá • 2 specs; Macapa; 16 May 1980; Elias and Penny leg.; CEMT • 2 specs; same data as for preceding; INPA. – Amazonas • 1 spec.; Manaus, Est. Aml, Km 134; 10 Aug. 1968; E. V. Silva and A. Faustino leg.; INPA. – Bahia • 2 specs; Caravelas; Mar. 1997; R. D. Silveira leg.; CEMT. – Espírito Santo • 2 specs; São Mateus; Dec. 1996; J. N. C. Louzada leg.; CEMT. – Mato Grosso • 1 spec.; Chapada dos Guimarães; 15 ° 24 ′ 31 ″ S, 55 ° 45 ′ 53 ″ W; 2 Dec. 2015; A. Frolov and L. Akhmetova leg.; light; CEMT • 1 spec.; Sesc Pantanal; 13 Apr. 2002; light trap; CEMT • 1 spec.; same data as for preceding 8 Jul. 2003; armadilha pano; CEMT • 1 spec.; same data as for preceding; 5 Aug. 2003; light trap; CEMT. – Mato Grosso do Sul • 1 spec.; Guia Lopes da Laguna; 15 Feb. 2007; A. Abot leg.; light trap; CEMT. – Minas Gerais • 1 spec.; 20 km E of Viçosa; 5 Dec. 1998; J. L. Moreno and T. Mestre leg.; CEMT • 1 spec.; Cordisburgo, F. Pontinha; Jul. 1994; de Mello leg.; CEMT • 1 spec.; Ipatinga; Sep. 1998; E. Grossi leg.; CEMT • 1 spec.; Mariana; 7 Dec. 1998; J. L. Moreno and T. Mestre leg.; CEMT • 3 specs; Teixeiras; Nov. 1996; Hardy and Herrison leg.; CEMT. – Pará • 1 spec.; Ilha Marajo; 23 May 1978; INPA. – Paraná • 1 spec.; São Jose dos Pinhais, Bairro São Marcos; 25 ° 36 ′ 40 ″ S, 49 ° 10 ′ 03 ″ W; elev. 894 m; 26 Oct. 2014; F. M. Neves leg.; CEMT. – Rio Grande do Sul • 2 specs; Bagé, Embrapa Pecuaria; Jan. 2006; L. D. Audino leg.; pitfall; CEMT • 1 spec.; Leões farm; 30 ° 59 ′ 12 ″ S, 54 ° 29 ′ 47 ″ W; 20 – 26 Oct. 2011; R. M. Moraes leg.; pitfall trap (human + pig dung); CEMT • 1 spec.; Santa Cruz do Sul, Cerro Alegre; 29 ° 48 ′ 02 ″ S, 52 ° 19 ′ 39 ″ W; 14 Jan. 2012; R. Bohn leg.; grass field (P 2); pitfall; CEMT. – Santa Catarina • 1 spec.; Itapema; Jan. 1993; J. Carlos leg.; CEMT • 1 spec.; Florianopolis; Oct. 2001; P. Grossi leg.; CEMT • 3 specs; same data as for preceding; Nov. 2001; CEMT. – São Paulo • Cujuina Cia; Feb. 1995; J. Carlos leg.; CEMT. – Tocantins • 1 spec.; Palmas; 13 Nov. 2001; P 1 D COL. Not. Equipe de Resgate UHE Lajeado leg.; CEMT. URUGUAY – Florida • 1 spec.; Florida, Cerro Colorado, Estacion Esperimental del Sul; 19 Feb. 1993; CEMT. – Rivera • 1 spec.; Vichadero, El. Pino farm; 21 – 27 Nov. 2011; R. M. Moraes leg.; pitfall trap baited (human + pig dung); CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF3FFABFF61F9C1FBD1FD92.taxon	materials_examined	BRAZIL – Mato Grosso • 1 spec.; Estação Ecológica de Taiamã; 11 Dec. 1981; M. Serrano leg.; CEMT • 1 spec.; Poconé; Jul. 2000; J. Lopes leg.; CEMT • 1 spec.; Poconé, Pousada Araras; 3 Dec. 1998; J. L. Moreno and T. Mestre leg.; Pantanal; CEMT • 1 spec.; Poconé, RPPN SESC, Base UFMT; 16 ° 29 ′ 52 ″ S, 56 ° 24 ′ 45 ″ W; 19 – 23 Jul. 2011; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Sesc Pantanal; 13 Apr. 2002; light trap; CEMT • 1 spec.; Varzea Grande, Cuiabá, Parque de Exposição; 20 Jun. 1972; Williams and Roger leg.; blacklight trap; CEMT • 3 specs; same data as for preceding; 1 Jul. 1972; CEMT • 1 spec.; Trindade, U. B. S; 1 Oct. 1984; E. Binda leg.; light trap; northwestern Polo; INPA • 1 spec.; same data as for preceding; J. Elias leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF3FFABFF61F9C1FBD1FD92.taxon	discussion	Collecting information Specimens were collected at altitudes ranging from 220 to 950 metres. Specimens are commonly collected with UV light traps, with traps baited with equine and bovine dung and with decomposing fruits. Specimens can be found in mangrove areas, pastures and forests (Stebnicka 2003 e). Remarks The Ataenius imbricatu s group has 19 recognized species and 12 are known to occur in Brazil (Stebnicka 2003 e). A key for species of this group can be found in Stebnicka (2003 e).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFCFFABFD56FDDAFE0AFC2F.taxon	description	Fig. 12 A – C, 41 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFCFFABFD56FDDAFE0AFC2F.taxon	diagnosis	Diagnosis Length 3.0 – 4.8 mm. Body punctate, glossy, piceous. Head punctate, may present transverse wrinkles; clypeus emarginate. Pronotum with fine and coarse punctures; coarse punctures scattered on disc, extremely close near anterior and posterior angles; laterally and posteriorly with marginal line; posterior angles sinuate. Elytra with interstriae near posterior margin and laterally convex; interstriae 7 – 10 with closer and larger punctures. Metaventrite plate bound by lines of coarse punctures. Metatibia with acuminate accessory spine.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFCFFABFF61FC6DFAFDFA53.taxon	materials_examined	BRAZIL – Amazonas • 2 specs; Benjamin Constant, Guanabara II; 10 Mar. 2004; P. H. Silva leg.; CEMT. – Mato Grosso • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 15 Dec. 2007; O. Peres-Filho leg.; CEMT. – Minas Gerais • 1 spec.; Ijací, Faz FAEPE; Dec. 2002; J. Louzada leg.; CEMT. – Santa Catarina • 6 specs; Florianópolis, Praia Pantano do Sul; 27 ° 47 ′ S, 48 ° 31 ′ W; Oct. 2008; M. I. M. Hernandez et al. leg.; frontal dune; pitfall; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFCFFABFF61FC6DFAFDFA53.taxon	discussion	Collecting information Specimens can be collected with light traps (Stebnicka 2001 c). Remarks The Ataenius nugator group has 15 recognized species but Ataenius nugator Harold, 1880 is the only species of this group in Brazil (Stebnicka 2001 c). Ataenius nugator was first recorded by Stebnicka (2001 c) in Rondônia State. A key for species of this group can be found in Stebnicka (2001 c).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFCFFABFDA0F99BFC8CF8CE.taxon	description	Figs 9 E – F, 42 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFCFFABFDA0F99BFC8CF8CE.taxon	diagnosis	Diagnosis Length 5.0 – 7.0 mm. Body large, elongate, dorsally glabrous, glossy, piceous. Head with punctures of two sizes, fine punctures inconspicuous in between coarse ones; coarser punctures closer on frons; clypeus sinuate; ventral anterior edge of clypeus downward bent, triangularly acuminate; lateral angles rounded, weakly wrinkled near margin. Pronotum with punctures of two sizes; fine punctures inconspicuous amidst coarser punctures; coarse punctures concentrated laterally; lateral margins fringed or not by setae; posterior angles sinuate or not. Scutellum foveate anteriorly. Elytral interstriae convex. Accessory spine of metatibiae present or not, if present, long and acuminate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFDFFAAFF61FEABFEE0FE77.taxon	materials_examined	BRAZIL – Rio de Janeiro • 2 specs; Macaé de Cima; Nov. 1998; F. Z. Vaz-de-Mello and E. Grossi leg,; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFDFFAAFF61FE76FA83FAF4.taxon	materials_examined	BRAZIL – Espírito Santo • 1 spec.; Venda nova do Imigrante; Dec. 2001; F. Z. Vaz-de-Mello leg.; CEMT. – Minas Gerais • 1 spec.; Conceição dos Ouros, Rio Sapucaí; Jan. 2003; Neto and G. P. Almeida leg.; CEMT • 1 spec.; Itamonte; Sep. 1999; D. Takyia leg.; CEMT. – Rio de Janeiro • 3 specs; Nova Friburgo; Dec. 1996; E. Grossi and P. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Jan. 1999; P. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Nov. 2002; E. Grossi leg.; CEMT • 1 spec.; Nova Friburgo, Conquista; Jan. 2004; E. Grossi leg.; CEMT. – Rio Grande do Sul • 1 spec.; Bagé, Leões farm; 30 ° 59 ′ 12 ″ S, 54 ° 29 ′ 47 ″ W; 20 – 26 Oct. 2011; R. M. Moraes leg.; pitfall trap baited (human + pig dung); CEMT • 1 spec.; same data as for preceding; 30 ° 59 ′ 13 ″ S, 54 ° 29 ′ 47 ″ W; CEMT. – Santa Catarina • 1 spec.; Urubici (Parna S. JQ); 28 ° 09 ′ 35 ″ S, 49 ° 37 ′ 52 ″ W; 21 Jul. 2015; P. G. da Silva leg.; cow dung; CEMT • 1 spec.; Urubici, P. N. São Joaquim; 16 – 19 Mar. 2012; Grossi, Parizotto and Leivas leg.; CEMT. – São Paulo • 2 specs; São Miguel Arcanjo, P. E. Carlos Botelho; 24 ° 03 ′ 59 ″ S, 47 ° 59 ′ 39 ″ W; elev. 580 m; 11 Jun. 2012; M. Boutefeu leg.; Tapirus feces; CEMT • 1 spec.; Serra da Bocaina, Fazenda Lageado; elev. 1700 m; May 1951; Delcy and Machado leg.; CEMT. MEXICO – Chiapas • 1 spec.; Palenque; Jun. 2004; F. Nicolalde leg.; CEMT. URUGUAY – Rivera • 1 spec.; Rivera, Richadero, El Cerro Farm; 21 – 27 Nov. 2011; R. M. Moraes leg.; pitfall trap baited (human + pig dung); CEMT • 1 spec.; Rivera, COFUSA; 30 ° 57 ′ 36 ″ S, 55 ° 29 ′ 53 ″ W; 3 – 11 Oct. 2011; R. M. Moraes leg.; pitfall trap baited (human + pig dung); CEMT • 1 spec.; Rivera, Bodegas Carrau farm; 3 – 11 Oct. 2011; R. M. Moraes leg.; pitfall trap baited (human + pig dung); CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFDFFAAFF61FAF3FB5EF855.taxon	materials_examined	BRAZIL – Minas Gerais • 1 spec.; Guanhães; 25 May 1991; C. Z. [unreadable] leg.; CEMT. PANAMA – Chiriquí • 3 specs; 2 km S of Boque; elev. 1400 m; 24 Jul. 1998; M. Hardy leg.; CEMT. VENEZUELA – Yaracuy • 1 spec.; Bolívar, Aroa; 10 ° 20 ′ 24 ″ N, 68 ° 50 ′ 06 ″ W; elev. 1373 m; 19 Jul. 2009; M. Asmüssen, P. Colomenares and H. Martínez leg.; human feces; CEMT • 2 specs; same data as for preceding; 10 ° 23 ′ 11 ″ N, 68 ° 50 ′ 49 ″ W; elev. 1380 m; CEMT • 4 specs; same data as for preceding; 10 ° 23 ′ 07 ″ N, 68 ° 50 ′ 45 ″ W; elev. 1410 m; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFDFFAAFF61FAF3FB5EF855.taxon	discussion	Collecting information Members of this group can be found in grasslands and forests. Specimens can be collected with light traps and are attracted to human, bovine, equine and Tapirus feces (Stebnicka 2001 c). Remarks The Ataenius perforatus group has six recognized species, but only three species are registered in Brazil (Stebnicka 2001 c). A key for species of this group can be found in Stebnicka (2001 c).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFEFFA9FF61FD21FB5BFCF8.taxon	materials_examined	BRAZIL – Pernambuco • 1 spec.; Fernando de Noronha, Sancho; 20 – 28 Feb. 2020; Grossi and Rafael leg.; light; CEMT • 1 spec.; same data as for preceding; southeast mangroove; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFEFFA9FF61FCEFFEE0F9A1.taxon	materials_examined	BRAZIL – Distrito Federal • 4 specs; Brasília; elev. 1100 m; Jan. 2001; N. Degallier leg.; light; CEMT • 1 spec.; same data as for preceding; Feb. 2001; CEMT • 1 spec.; same data as for preceding; Oct. 2000; CEMT. – Goiás • 1 spec.; Bom Jesus; Nov. 1996; J. Carlos leg.; CEMT. – Mato Grosso • 1 spec.; Cotriguaçu, Fazenda São Nicolau; Nov. 2007; O. Peres-Filho leg.; CEMT • 1 spec.; Diamantino, Vale da Solidão; 7 Feb. 2011; E. Furtado leg.; CEMT • 1 spec.; Poconé; 22 Feb. 1998; light trap; CEMT. – Mato Grosso do Sul • 1 spec.; Campo Grande; 1990 – 1992; I. Bianchin leg.; CEMT. – Minas Gerais • 2 specs; Águas Vermelhas; Dec. 1997; A. Bello and Vaz-de-Mello leg.; CEMT • 8 specs; Cordisburgo, Fazenda Pontinha; Dec. 1993; F. Z. Vaz-de-Mello leg.; CEMT • 8 specs; same data as for preceding; Jan. 1996; CEMT • 1 spec.; same data as for preceding; Jan. 1998; CEMT • 1 spec.; same data as for preceding; Jan. 1999; Falqueto and Vaz-de-Mello leg.; CEMT • 1 spec.; Lavras; 21 ° 19 ′ 56 ″ S, 44 ° 58 ′ 08 ″ W; 22 Mar. 2008; M. R. Santos and D. L. H. Takahashi leg.; CEMT • 1 spec.; same data as for preceding; 21 ° 19 ′ 59 ″ S, 44 ° 58 ′ 17 ″ W; CEMT • 2 specs; same data as for preceding; 21 ° 20 ′ 03 ″ S, 44 ° 58 ′ 30 ″ W; CEMT • 1 spec.; Monte Claro; Jan. 2000; Louzada and Louzada leg.; Cerrado; CEMT • 6 specs; Santa Barbara; 5 Jan. 1994; J. C. Zanuncio leg.; light trap; CEMT • 1 spec.; Três Marias; Jan. 1993; CEMT. – Rio de Janeiro • 1 spec.; Nova Friburgo; Dec. 1998; P. Grossi leg.; CEMT. – Tocantins • 2 specs; Santa Isabel do Morro, Ilha do Bananal; 5 Dec. 1978; Bello leg.; CEMT • 1 spec.; same data as for preceding; 16 Apr. 1979; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFEFFA9FDA0FE94FE88FDD5.taxon	description	Figs 10 C, 42 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFEFFA9FDA0FE94FE88FDD5.taxon	diagnosis	Diagnosis Length 2.5 – 5.3 mm. Body oblong, dorsally glabrous, light reddish testaceous to piceous. Head punctate; clypeus sinuate, margin medially bent downward, acuminate, lateral angles rounded. Pronotum densely and coarsely punctate, punctures larger and closer laterally; lateral margins may or may not be fringed by setae. Scutellum anteriorly foveolate. Elytral interstriae convex or not, variably punctured. Ventral surface glabrous with coarse punctures. Abdominal ventrites weakly strigate. Metatibiae without accessory spine.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFEFFA8FF61F9A4FADBFB14.taxon	materials_examined	BRAZIL – Acre • 1 spec.; Rio Branco; CEMT • 1 spec.; Rio Branco, PZ-UFAC; Feb. 1997; Vaz-de-Mello leg.; CEMT. – Amazonas • 1 spec.; Itacoatiara, Fazenda Aruana, Am 010, Km 215; 20 – 23 Aug. 1990; C. S. Motta, F. J. A. Peralta and B. Ronchi-Teles leg.; UV light; CEMT • 13 specs; same data as for preceding; 18 – 21 Sep. 1990; C. S. Motta, R. L. M. Ferreira and R. Andreazze leg.; CEMT • 1 spec.; same data as for preceding; INPA • 7 specs; same data as for preceding; 12 – 15 Jan. 1991; C. S. Motta, H. Misquita and R. Andreazze leg.; INPA • 9 specs; same data as for preceding; 13 – 16 Feb. 1991; UV light; C. S. Motta, R. Andreazze and U. Kawazoe leg.; INPA • 8 specs; same data as for preceding; Mar. 1991; C. S. Motta, F. J. A. Peralta and B. Rochin-Teles leg.; CEMT • 15 specs; same data as for preceding; 14 – 15 May 1991; C. S. Motta et al. leg.; INPA • 15 specs; same data as for preceding; 16 – 17 May 1991; INPA • 2 specs; same data as for preceding; 22 – 23 Jul. 1990; C. S. Motta et al. leg.; CEMT • 1 spec.; Manaus, INPA-base camp; 8 Jun. 1976; A. P. A Luna Dias leg.; CEMT. – Mato Grosso • 1 spec.; Alta Floresta, CEPLAC; 2 Oct. 2009; V. Gonçalvez leg.; fezes antas, por cima; CEMT • 1 spec.; Chapada dos Guimarães; 15 ° 24 ′ 31 ″ S, 55 ° 45 ′ 53 ″ W; 2 Dec. 2015; A. Frolov and L. Akhemetova leg.; light; CEMT • 1 spec.; Chapada dos Guimarães; 27 Apr. 2001; A. F. Ramos leg.; light trap; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 14 Feb. 2007; O. Peres-Filho leg.; CEMT • 1 spec.; same data as for preceding; 15 Dec. 2007; CEMT • 2 specs; Cotriguaçu, Fazenda São Nicolau; 9 ° 49 ′ 08 ″ S, 58 ° 15 ′ 40 ″ W; 19 – 24 Oct. 2012; F. Z. Vaz-de-Mello leg.; PPBIO P 1, FIT; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 18 Oct. 2014; W. Chamorro leg.; CEMT • 1 spec.; Poconé, Pousada das Araras; 3 Dec. 1998; J. L. Moreno and T. Mestre leg.; Pantanal; CEMT. – Minas Gerais • 1 spec.; Santa Barbara; 17 Feb. 1994; J. C. Zanuncio leg.; CEMT. – Pará • 2 specs; Jacarecanga; 9 ° 12 ′ 21 ″ S, 56 ° 57 ′ 34 ″ W; Feb. 2015; M. A. L. Bragaça leg.; pitfall; CEMT • 1 spec.; same data as for preceding; INTERD 1; 9 ° 15 ′ 30 ″ S, 56 ° 48 ′ 20 ″ W; Jan. 2016; M. A. L. Bragaça leg.; pitfall; CEMT. – Rondônia • 3 specs; 62 km S of Ariquemes; Feb. 1997; Vulinec-Mello leg.; CEMT. ECUADOR – Manabi • 1 spec.; Manta; elev. 5 m; 23 Jan. 1999; A. Pérez leg.; CEMT. – Napo • 1 spec.; Yasuni, Bloque; 16 Aug. 1996; F. Bersosa leg.; light trap; CEMT. – Pastaza • 1 spec.; Mera, 24 May [locality]; 1 ° 20 ′ S, 78 ° 10 ′ W; elev. 1140 m; 6 Apr. 1996; M. López leg.; CEMT. FRENCH GUIANA • 1 spec.; Cayenne, Nouragues, Inselberg; 4 ° 05 ′ N, 52 ° 41 ′ W; elev. 411 m; 9 Sep. 2010; SEAG leg.; FIT; CEMT • 1 spec.; 15 Nov. 2011; SEAG leg.; FIT (V); CEMT • 1 spec.; St Laurent de Maroni, Bélvédère de Saül; 3 ° 37 ′ 22 ″ S, 53 ° 12 ′ 57 ″ W; elev. 326 m; 22 Mar. 2011; SEAG leg.; CEMT • 1 spec.; same data as for preceding; 30 Jun. 2011; CEMT • 1 spec.; same data as for preceding; 13 May 2011; CEMT • 3 specs; same data as for preceding; elev. 360 m; 15 Sep. 2011; FIT (V); CEMT. PERU – Loreto • 1 spec.; 21 km W of Iquitos; 10 Jul. 2001; Curoeand M. Callegari leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFFFFA8FF61FB52FC27FA74.taxon	materials_examined	Paratype BRAZIL • 1 spec.; “ [yellow paper, printed] PARATYPE / Ataenius / skelleyi m. / Dt. Z. Stebnicka. // [white paper, printed] BRAZIL: Rondônia, 62 / km. SW. Ariquemes, nr. Fdza Rancho Grande / 8 - 20 - XI- 1994; J. Eger, / C. O’ Brien; black light // [white paper, printed] Ataenius / skelleyi / Stebnicka. // [white paper, printed] Q. R code to the left, CEMT / CUIABÁ / 00122412 ”; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFFFFA8FF61FA72FC58F817.taxon	materials_examined	BRAZIL – Amazonas • 1 spec.; Rio Tarumã-mirim, NW Manus; 8 Mar. 1976; J. Adis leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFFFFA8FF61FA72FC58F817.taxon	discussion	Collecting information Specimens were found at altitudes from 5 to 1100 metres. Members of this group are attracted to light and feces of humans, Bos Linnaeus, 1758 and Tapirus. Some species outside of Brazil were collected in bat guano and in forest litter (Stebnicka 2007 c). Remarks The Ataenius scutellaris group has 15 recognized species, five are found in Brazil (Stebnicka 2007 c). A key for species of this group can be found in Stebnicka (2007 c).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF8FFAFFF62FD04FE20F875.taxon	materials_examined	ARGENTINA – Formosa • 1 spec.; 50 km NW of Clorinda, PN Río Pilcomavo; 19 Dec. 1990; S. Peck and J. Peck leg.; UV; CEMT. BRAZIL – Amazonas • 1 spec.; Careiro da Várzea, Ilha Curarí S. Manaus; 5 Feb. 1976 J. Adis leg.; CEMT • 1 spec.; Iranduba, Ilha de Marchantaria, Rio Solimões; 3 ° 15 ′ S, 59 ° 58 ′ W; 6 Jan. 1981; J. Adis leg.; várzea [lowland]; INPA • 1 spec.; same data as for preceding; 2 Sep. 1981; INPA • 1 spec.; same data as for preceding; 14 Sep. 1981; CEMT • 4 specs; same data as for preceding; INPA • 1 spec.; Manaus; 6 Dec. 1977; B. C. Ratcliffe leg.; CEMT • 1 spec.; same data as for preceding; INPA • 1 spec.; 31 ° 05 ′ S, 59 ° 58 ′ W; 2 – 4 Mar. 1998; S. Golovatch leg.; CEMT • 20 specs; same data as for preceding; INPA • 1 spec.; Itacoatiara, Fazenda Aruana, Am 010, Km 215; 20 – 23 Aug. 1990; C. S. Motta, F. J. A. Peralth and Roncai-Teles leg.; INPA • 1 spec.; Manaus, Japiim – Cohabam; 14 Jun. 1976; J. Adis leg.; INPA • 1 spec.; Manaus, Rio Tarumá Mirim; 3 ° 02 ′ S, 60 ° 17 ′ W; 26 May 1983; terra firme [solid land]; INPA. – Distrito Federal • 1 spec.; Brasilia, Plano Piloto; 10 Oct. 1988; Bello leg.; CEMT. – Mato Grosso • 2 specs; Chapada dos Guimarães, Trilha Casa do Mel; 15 ° 22 ′ 53 ″ S, 55 ° 50 ′ 35 ″ W; elev. 520 m; Dec. 2015; R. N. Nunes, J. Cabra and M. Rossini leg.; light; CEMT • 1 spec.; Cuiabá, Boa Esperança; Dec. 2009; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Cuiabá; 15 ° 35 ′ 21 ″ S, 56 ° 01 ′ 46 ″ W; elev. 183 m; Sep. 2015; R. V. Nunes and B. F. Becker leg.; CEMT • 1 spec.; same data as for preceding; Nov. 2009; F. Z. Vaz-de-Mello leg.; CEMT • 5 specs; Poconé, RPPN SESC, Base UFMT; 16 ° 29 ′ 52 ″ S, 56 ° 24 ′ 45 ″ W; 19 – 23 Jul. 2011; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Porto Estrela, Heliporto; 15 ° 39 ′ 19 ″ S, 57 ° 12 ′ 50 ″ W; 21 Oct. 2017; R. J. P. Machado and R. Nunes leg.; CEMT • 2 specs; Santo Antônio do Leverger, Serra de São Vicente; 15 ° 52 ′ 39 ″ S, 55 ° 22 ′ 28 ″ W; 22 Feb. 2013; A. Petin and Benell leg.; CEMT • 1 spec.; Sesc Pantanal; 16 Nov. 2001; light trap; CEMT. – Mato Grosso do Sul • 1 spec.; Corumbá, Fazenda São Bento, Prox. Porto Jofre; 14 Apr. 2011; R. N. Nunes leg.; light; CEMT • 1 spec.; Guia Lopes da Laguna; 15 Nov. 2007; A. Abot leg.; light trap; CEMT. – Minas Gerais • 1 spec.; Águas Vermelhas; Dec. 1997; A. Bello; CEMT • 1 spec.; Cordisburgo, Fazenda Pontinha; Dec. 1998; Vaz-de-Mello leg.; CEMT. – Rio Grande do Sul • 1 spec.; Eldorado do Sul, EEA-UFRGS; Oct. 1996; Silva and Carvalho leg.; CEMT. – Rio de Janeiro • 1 spec.; Parque Nacional Itatiaia; 12 Dec. 1989; Bello leg.; CEMT • 1 spec.; Nova Friburgo; Nov. 2002; E. J. Grossi leg.; CEMT • 1 spec.; Macaé, PN Jurubatiba; 22 ° 16 ′ 40 ″ S, 41 ° 41 ′ 35 ″ W; elev. 3 m; 1 May 2007; J. A. Rafael and F. F. Xavier leg.; restinga vegetation; light; INPA. COLOMBIA – Antioquia • 1 spec.; Medellín; CEMT. PARAGUAY – Concepción • 1 spec.; San Carlos del Apa; 22 ° 14 ′ 04 ″ S, 5 ° 17 ′ 48 ″ W; 30 Oct. 2002; B. Garcete leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF8FFAFFD5BFE94FDB9FDF0.taxon	description	Figs 10 F, 43 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF8FFAFFD5BFE94FDB9FDF0.taxon	diagnosis	Diagnosis Length 3.4 – 6.0 mm. Body elongate, glabrous, glossy and piceous. Head with fine and coarse punctures, transverse wrinkles variably developed; clypeus sinuate, lateral angles rounded. Pronotum wide, with variable punctures. Elytra with variable sculpture, shagreened or glossy; interstriae convex or flat. Ventral surface glossy or shagreened. Anterior margin of metafemur normally with a fringe of setae. Accessory spine of metatibiae present or not.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF9FFAEFF61FE95FBE8F946.taxon	materials_examined	BRAZIL – Amazonas • 1 spec.; Autazes; 2 – 8 Aug. 1981; E. Brasil leg.; CEMT • 1 spec.; Iranbudá, Ilha de Marchantaria; 20 – 23 Feb. 1990; C. R. V. Fonseca leg.; CEMT • 1 spec.; same data as for preceding; 3 ° 15 ′ S, 59 ° 58 ′ W; 2 Sep. 1981; J. Adis leg.; INPA • 1 spec.; same data as for preceding; 2 Oct. 1981; J. Adis leg.; INPA • 4 specs; same data as for preceding; 4 Nov. 1981; CEMT • 19 specs; same data as for preceding; INPA • 10 specs; same data as for preceding; 17 Nov. 1981; CEMT • 298 specs; same data as for preceding; INPA • 3 specs; same data as for preceding; 2 Dec. 1981; CEMT • 29 specs; same data as for preceding; INPA • 1 spec.; same data as for preceding; 21 Dec. 1981; CEMT • 3 specs; same data as for preceding; INPA • 1 spec.; same data as for preceding; 22 Jan. 1982; CEMT. – Mato Grosso • 1 spec.; Cuiabá, Boa Esperança; 24 – 27 Oct. 2016; F. Z. Vaz-de-Mello leg.; manually; CEMT • 1 spec.; Cuiabá, CPA IV; 11 May 2010; Marcondes and Eliane leg.; manually; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 30 Oct. 2014; W. Chamorro leg.; blacklight trap; CEMT • 13 specs; Pixaim, Pantanal; 3 Dec. 1998; J. L. Moreno and T. Mestre leg.; CEMT • 1 spec.; Poconé, Serra das Araras; 3 Dec. 1998, J. L. Moreno and T. Mestre leg.; Pantanal; CEMT • 1 spec.; Poconé, RPPN SESC, Base UFMT; 16 ° 29 ′ 52 ″ S, 56 ° 24 ′ 45 ″ W; 19 – 23 Jul. 2011; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Primavera do Leste; 10 Jun. 2007; Benelli leg.; CEMT • 2 specs; Sesc Pantanal; 14 May 2002; light trap; CEMT • 7 specs; same data as for preceding; 8 Aug. 2002; CEMT • 1 spec.; same data as for preceding; 6 Sep. 2002; CEMT • 1 spec.; same data as for preceding; 4 Jul. 2002; CEMT • 1 spec.; same data as for preceding; 5 Aug. 2003; CEMT • 2 specs; same data as for preceding; 15 Sep. 2004; CEMT. – Minas Gerais • 1 spec.; Guanhães; 7 Dec. 1993; J. C. Zanuncio leg.; CEMT • 1 spec.; Ijací, Faz FAEPE; Nov. 2002; J. Louzada leg.; CEMT • 1 spec.; Visconde Rio Branco; Jan. 1998; W. Zikan leg.; CEMT. – Roraima • 9 specs; Amajari, Rio Uraricoera, Ilha de Maraca; 2 – 13 May 1987; J. A. Rafael, J. E. B Brasil and L. S. Aquino leg.; suspended trap; CEMT, INPA • 1 spec.; same data as for preceding; 18 – 28 Aug. 1987; J. A. Rafael et al. leg.; light trap; INPA • 1 spec.; same data as for preceding; 21 – 30 Nov. 1987; Malaise trap; INPA. – Rio de Janeiro • 2 specs; Itaguaí, Km 47 Estr. Rio; 30 Oct. 1945; Wygod leg.; CEMT. – Rio Grande do Sul • 1 spec.; Eldorado do Sul, EEA-UFRGS; 11 Oct. 1996; Carvalho and Silva leg.; arm. lum; CEMT. – Santa Catarina • 1 spec.; Urubici; 28 ° 08 ′ S, 49 ° 38 ′ W; 6 Feb. 2008; M. Wuerges leg.; pitfall bait; CEMT. – São Paulo • 1 spec.; 2 Nov. 1914; Melxei leg.; CEMT. COLOMBIA – Meta • 1 spec.; San Martín, San Francisco; elev. 335 m; Apr. 2006; J. Noriega leg.; directa; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF9FFAEFF61FE95FBE8F946.taxon	discussion	Collecting information The group can be found at altitudes ranging from 183 to 1100 metres. Specimens were collected with UV and are attracted to cow dung. Specimens can be found in savanna regions, pastures, flooded plains and forest (Stebnicka & Lago 2005). Remarks The Ataenius strigatus group has 17 recognized species, two occur in Brazil (Stebnicka & Lago 2005). A key for species of this group can be found in Stebnicka & Lago (2005).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF9FFADFDABF886FC03FE1F.taxon	description	Figs 11 A – D, 43 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFF9FFADFDABF886FC03FE1F.taxon	diagnosis	Diagnosis Length 4.5 – 6.0 mm. Body elongate, piceous. Head with punctures variable, punctures normally coarse and distinctly grouped on frons; clypeus with transverse wrinkles variably developed, medially sinuate, lateral angles rounded. Pronotum with punctures fine and coarse; lateral margins fringed with setae. Elytra with interstriae 8 – 10, at least near lateral margins, with larger, more evident and coarse punctures, compared with interstriae of disc. Ventral surface glossy, shining. Metaventral plate with group of large and coarse punctures near posterior margin of mesocoxa (except for Ataenius picinus). Metatibiae with acuminate accessory spine, its length surpassing apical fringe of setae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFAFFADFF61FD9DFDFBF971.taxon	materials_examined	ARGENTINA – Tucumán • 1 spec.; Sierra de San Javier, Horco Molle; Mar. 2002; F. Z. Vaz-de-Mello leg.; CEMT. BRAZIL – Bahia • 3 specs; Jan. 2005; A. F. Brito and G. R. Ribeiro leg.; CEMT • 2 specs; Presidente Tancredo Neves; 13 ° 24 ′ 32 ″ S, 39 ° 20 ′ 05 ″ W; Jan. 2011; C. M. P. Leite leg.; pitfall; CEMT. – Distrito Federal • 1 spec.; Brasilia; elev. 1100 m; Jan. 2001; N. Degallier leg.; CEMT • 11 specs; Brasilia, Fazenda Água Limpa (UNB); Nov. 2010; C. Suinaga leg.; CEMT • 1 spec.; Planaltina, Embrapa Cerrados; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 19 Jan. 2006; C. Oliveira leg.; agricultural area; light; CEMT. – Espírito Santo • 1 spec.; Corrego Itá; Oct. 1954; W. Zikan leg.; CEMT. – Mato Grosso • 1 spec.; Alta Floresta, Chacará Recanto das Oquideas; 3 Oct. 2011; A. Petini-Bevell leg.; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 15 Dec. 2007; O. Peres-Filho leg.; CEMT • 1 spec.; Poconé, Base UFMT; 16 ° 29 ′ 58 ″ S, 56 ° 24 ′ 49 ″ W; Mar. 2012; M. Rossini leg.; CEMT • 1 spec.; Viçosa; 10 Nov. 1982; Fiuza and Martina leg.; CEMT • 1 spec.; same data as for preceding; Feb. 2000; Lopes-Andrade and Vaz-de-Mello leg.; CEMT • 1 spec.; Viçosa, Corrego do Paraiso, Mata da Prefeitura; 1 Dec. 1982; P. S. F. Fiuza leg.; UV trap; CEMT • 1 spec.; Sesc Pantanal; 5 Jul. 2003; light trap; CEMT. – Paraná • 1 spec.; Cornélio Procópio, Parque Estadual Mata São Francisco; 23 ° 09 ′ 03 ″ S, 50 ° 33 ′ 51 ″ W; 19 Sep. 2009; N. Cipola leg.; CEMT • 1 spec.; Rancho Alegre, Fazenda Congonhas; 22 ° 47 ′ 45 ″ S, 51 ° 00 ′ 12 ″ W; elev. 580 m; 27 Sep. 2012; P. M. Felix leg.; pitfall fish; CEMT • 1 spec.; Rio Negro; Jan. 1924; M. Witti leg.; CEMT. – Rio Grande do Sul • 1 spec.; Bagé, Embrapa pecuárea; Jan. 2006; L. D. Audino leg.; pitfall; CEMT • 1 spec.; Canela, Flona; 8 Nov. 1998; J. Soledar leg.; CEMT. – Santa Catarina • 1 spec.; Painel-BAP IBAMA; 27 ° 50 ′ S, 50 ° 13 ′ W; 29 Sep. 2010; M. I. M. Hernández leg.; pitfall; meat atractive; CEMT. – São Paulo • 1 spec.; Luiza Antônio; 21 ° 34 ′ 35 ″ S, 47 ° 45 ′ 29 ″ W; 30 Jan. – 1 Feb. 2009; S. P. Rosa and M. Landenthin leg.; CEMT • 1 spec.; São Miguel Arcanjo, P. E. Carlos Botelho; 24 ° 04 ′ 01 ″ S, 47 ° 59 ′ 08 ″ W; elev. 730 m; 15 Oct. 2011; E. Bovy leg.; Brachyteles feces; CEMT. COLOMBIA – Quindío • 1 spec.; Filandia, Estacion Bremen; elev. 1800 m; Apr. 1998; J. Noriega leg.; bosque nativo; CEMT. URUGUAY – Cerro Largo • 1 spec.; Melo, La Invernada farm; 13 – 20 Dec. 2011; pitfall trap baited (human + pig dung); CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFAFFACFF61F975FD3EF9D2.taxon	materials_examined	ARGENTINA – Formosa • 1 spec.; Las Lomitas., Fortíns Soledad, Banhado la Estrella; 8 Jun. 2015; C. Strusmann leg.; rotten wood; CEMT. AUSTRALIA – Queensland • 2 specs; Dunmore State Forest, 12 km W of Cecil Plains, near Dalby; 22 Jan. 1999; CEMT. BRAZIL – Amazonas • 1 spec.; Itacoatiara, Fazenda Aruana, Am 010, Km 215; 16 – 19 Dec. 1990; C. S. Motta, R. Andreazze and V. Kawazoe leg.; UV light; CEMT • 1 spec.; same data as for preceding; INPA • 1 spec.; same data as for preceding; 12 – 15 Jan. 1991; INPA • 1 spec.; same data as for preceding; 13 – 16 Feb. 1991; INPA. – Distrito Federal • 1 spec.; Brasilia, Fazenda Água Limpa (UNB); Nov. 2010; C. Suinaga leg.; CEMT. – Mato Grosso • 1 spec.; Alta Floresta, Chácara Recanto das Orquídeas; 3 Oct. 2011; A. Petinni-Bevell leg.; CEMT • 2 specs; Chapada dos Guimarães, Trilha Casa do Mel; 15 ° 22 ′ 53 ″ S, 55 ° 50 ′ 35 ″ W; elev. 520 m; Feb. 2015; R. V. Nunes, J. Cabra and M. Rossini leg.; light; CEMT • 1 spec.; Diamantino; 18 Sep. 2010; M. E. Maldaner leg.; manually; CEMT • 1 spec.; Pousada das Araras; 3 Dec. 1998; J. L. Moreno and T. Mestre leg.; Pantanal; CEMT. – Mato Grosso do Sul • 1 spec.; Corumbá, Fazenda São Bento; 2010; F. R. Tortato leg.: CEMT • 1 spec.; Corumbá; Prox. Porto Jofre; 14 Apr. 2011; R. V. Nunes leg.; light; CEMT • 2 specs; Porto Murtinho, Margem do Rio Paraguai, Zona Urbana; 6 Apr. 2015; F. Tortato leg.; light; CEMT • 1 spec.; Selvíria, UNESP. Farm; 15 Jun. 1991; S. R. Rodrigues leg.; ex Guzera cattle dropping; Brachiaria decumbens pasture; INPA. – Minas Gerais • 1 spec.; Mariana; 7 Nov. 1998; J. L. Moreno and T. Mestre leg.; CEMT • 1 spec.; Ponte Nova; 18 Oct. 1979; Planalsucar leg.; CEMT • 1 spec.; Viçosa; 4 Aug. 1982; Fiuza and Martins leg.; CEMT • 1 spec.; Viçosa; 20 Feb. 1986; P. S. F. Fiuza leg.; UV light; CEMT • 1 spec.; Viçosa, Córrego do Paraíso, Mata da Prefeitura; 12 Feb. 1984; Fiuza and Martins leg.; CEMT • 1 spec.; Viçosa, Campus UFV; Nov. 1997; F. Vaz-de-Mello leg.; CEMT. – Rio Grande do Sul • 7 specs; Eldorado do Sul, Estação Experimental Agronomica-UFRGS; 11 Oct. 1996; Carvalho and Silva leg.; arm. lum; CEMT. – Rio de Janeiro • 1 spec.; Macaé, P. N. Jurubatiba; 22 ° 16 ′ 44 ″ S, 41 ° 41 ′ 35 ″ W; 1 May 2007; J. A. Rafael and F. F. Xavier leg.; restinga vegetation; light; CEMT • 1 spec.; Miguel Pereira; Jan. 1998; J. Carlos leg.; CEMT • 1 spec.; Nova Friburgo; Dec. 1994; F. Z. Vaz-de-Mello leg.; CEMT. – Santa Catarina • 1 spec.; Florianópolis; 27 ° 36 ′ 15 ″ S, 48 ° 30 ′ 12 ″ W; 31 Jan. 2016; P. G. Silva leg.; at light; CEMT. – Tocantins • Porto Nacional, Agrotins; 5 Oct. 2005; M. S. Matos leg.; CEMT. FIJI – Viti levu • 3 specs; Suva; 20 – 28 Oct. 1985; G. Borrea and Issza leg.; CEMT. MEXICO – Chiapas • 2 specs; Palenque; Jun. 2004; F. Nicolalde leg.; CEMT. UNITED STATES OF AMERICA – Texas • 1 spec.; Bosque, Whitney, Lake Wihtney State Park; 26 Jul. 1997; B. Buchli leg.; blacklight trap; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFBFF91FF61F915FD1DFA08.taxon	materials_examined	BRAZIL – Acre • 3 specs; Rio Branco; CEMT • 1 spec.; Rio Branco, PZ-UFAC; Feb. 1997; CEMT • 1 spec.; same data as for preceding; Nov. 2000; M. A. Oliveira leg.; CEMT. – Amazonas • 1 spec.; Manaus, Est. Aml, Km 134; May 1968; INPA. – Bahia • 1 spec.; Reserva ALOAO; Jun. 1998; CEMT • 1 spec.; Caravelas; 8 Aug. 1997; CEMT • 1 spec.; Ilhéus; 14 ° 42 ′ 23 ″ S, 39 ° 07 ′ 37 ″ W; 21 Oct. 2011; C. M. P. Leite leg.; pitfall; CEMT • 2 specs; Itamarajú; 16 ° 57 ′ 58 ″ S, 39 ° 26 ′ 01 ″ W; Apr. 2011; C. M. P. Leite leg.; pitfall; CEMT • 20 specs; Mucugê, Chapada Diamantina, Carrasco; 27 Feb. 2010; T. Vasconcelos leg.; CEMT • 3 specs; Porto Seguro, RPPN, Veracel, Mussununga; 11 Nov. 2004; J. N. C. Louzada leg.; CEMT • 1 spec.; Presidente Tancredo Neves; 13 ° 22 ′ 56 ″ S, 39 ° 20 ′ 27 ″ W; Jan. 2011; C. M. P. Leite leg.; pitfall; CEMT • 3 specs; same data as for preceding; 13 ° 24 ′ 32 ″ S, 39 ° 20 ′ 05 ″ W; Jan. 2011; CEMT • 1 spec.; Ubaíra; 13 ° 08 ′ 47 ″ S, 39 ° 40 ′ 34 ″ W; Oct. – Jan. 2011; CEMT. – Espírito Santo • 1 spec.; P. E. Pedra Azul; elev. 1500 m; Jan. 2000; Lopes-Andrade and Vaz-de-Mello leg.; CEMT • 2 specs; Venda Nova do Imigrante, Clube; 20 ° 20 ′ 5 ″ S, 41 ° 08 ′ 18 ″ W; Jan. 2010; Vaz-de-Mello leg.; F 2; CEMT • 2 specs; Venda Nova do Imigrante, Vila Santa Cruz; 4 – 12 Jan. 2018; Falqueto and Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; Dec. 1999; F. Z. Vaz-de-Mello leg.; CEMT. – Mato Grosso • 1 spec.; Alta Floresta, Chácara Recanto das Orquídeas; 3 Oct. 2011; A. Petini-Bevell leg.; CEMT • 1 spec.; Campo Verde; 10 May 2012; K. Peña leg.; CEMT • 4 specs; Cotriguaçu, Faz São Nicolau; 9 ° 50 ′ 19 ″ S, 58 ° 15 ′ 03 ″ W; 25 Oct. 2014; A. Asenjo leg.; CEMT • 2 specs; same data as for preceding; 26 Oct. 2014; CEMT • 1 spec.; same data as for preceding; 28 Oct. 2014; UV light; CEMT • 1 spec.; same data as for preceding; 9 ° 51 ′ 20 ″ S, 58 ° 14 ′ 55 ″ W; Nov. 2014; R. V. Nunes and A. Asenjo leg.; light; CEMT • 2 specs; same data as for preceding; 15 Dec. 2007; O. Peres-Filho leg.; CEMT • 1 spec.; same data as for preceding; 30 Oct. 2014; W. Chamorro leg.; blacklight trap; CEMT • 4 specs; same data as for preceding; 95 ° 12 ′ 01 ″ S, 58 ° 14 ′ 55 ″ W; elev. 220 m; 16 – 20 Oct. 2012; F. Z. Vaz-de-Mello leg.; base camp; manually collected; CEMT • 1 spec.; same data as for preceding; 9 ° 51 ′ S, 58 ° 15 ′ W; Nov. 2010; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Cuiabá, Recanto Siriema; 15 ° 35 ′ 32 ″ S, 56 ° 1 ′ 38 ″ W; Sep. 2014; R. V. Nunes and B. F. Becker leg.; light; CEMT • 1 spec.; Curvelândia, Fazenda Luizinho, Bacia, Cabaçal; Feb. 2004; M. Santos-Filho leg.; CEMT • 2 specs; Diamantino, Alto Rio Arinos; Oct. 2001; E. Furtado; CEMT • 1 spec.; Nobres, Jd. Petrópolis; Nov. 2017; K. N. Pereira leg.; CEMT • 3 specs; Poconé, Pousada das Araras, Pantanal; 3 Dec. 1998; J. L. Moreno and T. Mestre leg.; CEMT • 1 spec.; Primavera do Leste; 10 Jun. 2007; P. Benelli leg.; CEMT • 1 spec.; same data as for preceding; 10 Nov. 2010; CEMT • 1 spec.; Santo Antônio do Leverger, Faz exp da UFMT; 15 ° 51 ′ 17 ″ S, 56 ° 04 ′ 17 ″ W; 17 – 19 Mar. 2011; A. S. Tissiani leg.; hum dung; CEMT • 1 spec.; Sesc Pantanal; 15 Apr. 2002; light trap; CEMT. – Mato Grosso do Sul • 1 spec.; Corumbá, Fazenda São Bento; 17 ° 19 ′ 16 ″ S, 56 ° 41 ′ 51 ″ W; 28 Apr. 2012; Rossini leg.; riparian forest of Piquiri River; CEMT • 1 spec.; Corumbá, Passo da Lontra; Nov. 2000; J. Raizer leg.; CEMT • 1 spec.; same data as for preceding; Dec. 2000; J. Raizer leg.; CEMT • 1 spec.; same data as for preceding; Feb. 2001; CEMT • 1 spec.; Poconé; 23 Jun. 1998; light trap on acuri palm tree; CEMT. – Minas Gerais • 1 spec.; Caxambu; Dec. 1990; A. Bello leg.; CEMT • 1 spec.; Cordisburgo; Jan. 1994; Vaz Mello leg.; CEMT • 1 spec.; same data as for preceding; Dec. 1997; CEMT • 1 spec.; same data as for preceding; Jan. 1998; CEMT • 1 spec.; Cordisburgo, F. Pontinha; Jul. 1994; de Mello leg.; CEMT • 2 specs; same data as for preceding; Jan. 1999; F. Z. Vaz-de-Mello leg.; CEMT • 2 specs; Ijací, Faz FAEPE Nov. 2002; J. Louzada leg.; CEMT • 2 specs; Ipatinga; Sep. 1993; E. Grossi leg.; CEMT • 1 spec.; Lavras; Nov. 2002; Vaz-de-Mello leg.; CEMT • 1 spec.; Lima Duarte, Parque Estadual Ibitipoca; 20 Feb. 1997; C. R. M. Abreu leg.; CEMT • 1 spec.; Ponte Nova; 23 Oct. 1979; Planalsucar leg.; CEMT • 1 spec.; São Gonçalo do Sapucaí; 28 Jan. 2017; L. F. Maia leg.; CEMT • 1 spec.; Viçosa; 13 Dec. 1982; D. S. Martins leg.; CEMT • 1 spec.; same data as for preceding; 26 Nov. 1986; P. S. F. Fiuza leg.; CEMT • 3 specs; same data as for preceding; 23 Sep. 1993; Louzada and Silva leg.; CEMT • 1 spec.; same data as for preceding; Oct. 1997; Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; Campus UFV; Nov. 1997; CEMT. – Pará • 7 specs; Curionópolis, Caverna SL 65 ES; 5 ° 58 ′ 53 ″ S, 49 ° 37 ′ 12 ″ W; 5 Jun. 2010; R. A. Zampaulo leg.; CEMT • 2 specs; Santarém; pasture; CEMT • 3 specs; Tailândia; 7 Aug. 2014; J. Malta and G. Santos leg.; CEMT. – Rio de Janeiro • 3 specs; Balneáreo São Pedro da Aldeia; 3 Jan. 1979; Bello leg.; CEMT • 1 spec.; same data as for preceding; 9 Feb. 1980; CEMT • 1 spec.; Itatiaia; Jan. 1995; CEMT • 1 spec.; Miguel Pereira; Feb. 1997; J. Carlos leg.; CEMT • 1 spec.; same data as for preceding; Feb. 1998; CEMT • 7 specs; same data as for preceding; Jan. 1998; CEMT • 1 spec.; same data as for preceding; Mar. 1998; CEMT • 1 spec.; same data as for preceding; Jan. 2002; CEMT • 1 spec.; Nova Friburgo; Nov. 1999; P. Grossi leg.; CEMT • 3 specs; same data as for preceding; Nov. 2016; E. Grossi leg.; CEMT • 1 spec.; Rio de Janeiro, Copacabana; 11 Nov. 1990; Bello leg.; CEMT • 1 spec.; same data as for preceding; 6 Oct. 1992; CEMT • 1 spec.; Rio de Janeiro, Ipanema; Nov. 1991; Zagury leg.; CEMT. – Rio Grande do Sul • 5 specs; Eldorado do Sul, EEA-UFRGS; Oct. 1996; Silva and Carvalho leg.; CEMT • 1 spec.; São Gabriel; 30 ° 20 ′ 28 ″ S, 54 ° 15 ′ 56 ″ W; 17 Oct. 2013, J. M. Flores leg.; floresta; pitfall (egg); CEMT. – Rondônia • 2 specs; 62 km S of Ariquemes; Feb. 1997; Vulinec and Mello leg.; CEMT • 2 specs; Cacaulândia, Fazenda Rancho Grande; Feb. 1997; Vaz-de-Mello leg.; CEMT • 1 spec.; Porto Velho, Caiçara; 9 ° 26 ′ 28 ″ S, 64 ° 49 ′ 00 ″ W; 20 Jun. 2011; M. F. Souza leg.; FIT; CEMT • 1 spec.; Porto Velho, Mutum-paraná, Rio Madeira; 12 Jun. 2004; margem direita interior; light trap; CEMT. – Roraima • 1 spec.; Rio Uraricoera, Ilha de Maraca; 2 – 13 May 1987; J. A. Rafael, J. E. B. Brasil and L. S. A. Aquino leg.; suspended trap; INPA. – São Paulo • 1 spec.; Cerqueira César; Nov. 2002; J. Carlos leg.; CEMT • 1 spec.; same data as for preceding; Dec. 1992; J. Carlos leg.; CEMT. – Tocantins • 1 specs; Porto Nacional, Agrotins; 5 Oct. 2005; M. S. Matos leg.; CEMT. COSTA RICA • 3 specs; Hac. Guachipelin, Vólcan Rincón de la Vieja; 13 Aug. 1999; J. L Moreno and T. Mestre leg.; CEMT. MEXICO – Veracruz • 2 specs; Km 22 Carr. Palma sola, Plan de Las Hayas; elev. 800 m; 12 Oct. 1982; Villalobos leg.; en suelo de pastizal inducido [in induced pastil soil]; CEMT • 3 specs; Km 14.5 Xico – Oxllapa, Mpio. Xico; elev. 2050 m; 18 Nov. 1999; E. Montes de Oca leg.; snm potrero [stable]; directa en excremento de vaca [directly on cow dung]; CEMT. NICARAGUA – León • 1 spec.; Las Marías; 9 Feb. 1995; Maes et al. leg.; CEMT. PANAMA – Chiriqui • 2 specs; 10 Km N of Santa Clara; elev. 1600 m; 26 Jul. 1998; M. Hardy leg.; CEMT. – Cocl e • 4 specs; La Mesa; elev. 850 m; 5 – 10 May 2007; Curoe leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFFBFF91FF61F915FD1DFA08.taxon	discussion	Collecting information Members of this group can be found at altitudes ranging from 220 to 1100 metres. Species of this group can be found on a variety of substrates. The species Ataenius picinus was found in forest litter, rotten wood, associated with fungus, in feces of different mammals and carcasses in regions of forest and grasslands. In general, these insects can be collected with UV light traps and with traps baited with dung and rotten flesh (Stebnicka 2004). Remarks The Ataenius strigicauda group has six recognized species, four are found in Brazil. A key for species of this group can be found in Stebnicka (2004).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC6FF91FD92FA7FFC40F888.taxon	description	Figs 10 A – B, 44 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC6FF91FD92FA7FFC40F888.taxon	diagnosis	Diagnosis Length 2.5 – 4.5 mm. Body elongate sub-cylindrical, opaque, piceous. Head with coarse longitudinally elongate punctures; clypeus emarginate, lateral angles can be rounded, obtuse or denticulate; surface behind clypeal margin may present granules. Pronotum with coarse punctures across surface; lateral margins fringed with setae. Scutellum foveate anteriorly. Elytra shagreened with humeral tooth present; interstriae with variable sculpture can be granulate, carinate or only punctate, glabrous or not. Ventral surface coarsely punctured. Accessory spine of metatibia absent.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC0FF96FF61FAB2FBC2FE42.taxon	materials_examined	ECUADOR – Esmeraldas • 2 specs; Quinindé; May 1987; M. H. López leg.; CEMT. – Imbabura • 1 spec.; Barcelona; 12 – 20 Sep. 1995; A. Endara leg.; CEMT • 1 spec.; Los Ríos, CCRP; 2 Jan. 1980; S. Sandoval leg.; CEMT • 1 spec.; same data as for preceding; 4 Jan. 1980; CEMT • 1 spec.; same data as for preceding; 11 Jan. 1981; CEMT • 1 spec.; same data as for preceding; 5 Jun. 1980; CEMT • 2 specs; same data as for preceding; ex cacao abierto [extracted from opened cocoa bean]; CEMT • 3 specs; same data as for preceding; 8 Jun. 1980; CEMT • 1 spec.; same data as for preceding; ex palma abierta [extracted from plam tree]; CEMT • 1 spec.; same data as for preceding; 11 [unreadable] 1980; CEMT • 3 specs; same data as for preceding; CCRP; 11 Jun. 1980; CEMT • 1 spec.; same data as for preceding; 12 Jun. 1980; CEMT • 2 specs; same data as for preceding; 17 Jun. 1980; CEMT • 1 spec.; same data as for preceding; 13 Jun. 1980; ex cacao abierto [extracted from opened cocoa bean]; CEMT. – Pichincha • 5 specs; CCRP; 14 Jun. 1980; S. Sandoval leg.; ex palma cerrada; CEMT • 1 spec.; same data as for preceding; 17 Jun. 1980; ex cacao abierto [extracted from opened cocoa bean]; CEMT • 2 specs; same data as for preceding; 21 Jul. 1980; ex palma cerrada [extracted from plam tree]; CEMT • 1 spec.; same data as for preceding; 30 May 1980; ex cacao abierto [extracted from opened cocoa bean]; CEMT. MEXICO – Veracruz • 2 specs; Actopan, Laguna La Mancha; Sep. 2003; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Actopan Dunas; 17 Sep. 2003; F. Z. Vaz-de-Mello leg.; CEMT. SAINT VINCENT AND THE GRENADINES • 1 spec.; [Saint Andrew], Emerald Valley Hotel; 13 ° 12 ′ N, 61 ° 14 ′ W; elev. 20 m; 27 – 29 Aug. 2006; S. Peck and J. Peck leg.; E of Layou, streamside; UV trap; CEMT • 1 spec.; [Saint Patrick], Hermitage Forest; 13 ° 14 ′ N, 61 ° 12 ′ W; elev. 350 m; 23 Aug. 2006; J. Peck and S. Peck leg.; E of Spring Village, forest; UV trap; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC1FF96FF61FD84FC2AFD1B.taxon	materials_examined	BRAZIL – Mato Grosso do Sul • 1 spec.; Aquidauana; S. R. Rodrigues leg.; CEMT. – Minas Gerais • 1 spec.; Lavras, Ponte Nova; 29 Apr. 1980; Planalsucar leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC1FF96FF61FD52FC8EFC70.taxon	materials_examined	BRAZIL – Acre • 1 spec.; Rio Branco, PZ-UFAC; Feb. 1997; F. Z. Vaz-de-Mello leg.; CEMT. VENEZUELA – Yaracuy • 1 spec.; Bolívar, Aroa; 10 ° 00 ′ 00 ″ N, 68 ° 00 ′ 00 ″ W; elev. 459 m; 19 Jul. 2009; M. Asmüssen and P. Colmenares leg.; human feces; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC1FF96FF61FC77FC2AFBCA.taxon	materials_examined	BRAZIL – Rio de Janeiro • 1 spec.; Itatiaia; elev. 2400 m; Feb. 2004; P. Grossi leg.; CEMT. – Santa Catarina • 1 spec.; Florianópolis; Nov. 2001; P. Grossi leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC1FF95FF61FB3DFEBAFDCA.taxon	materials_examined	BRAZIL – Amazonas • 1 spec.; Ilha da Marcantaria; 20 – 29 Nov. 1999; C. R. V. Fonseca leg.; CEMT. – Bahia • 1 spec.; Lençóis, Rio Santo Antonio; 12 ° 29 ′ 24 ″ S, 41 ° 19 ′ 46 ″ W; elev. 350 m; 5 Jun. 2007; Rafael and Xavier leg.; light; CEMT. – Distrito Federal • 1 spec.; Brasília; Feb. 2001; N. Degallier leg.; CEMT • 1 spec.; same data as for preceding; elev. 1100 m; CEMT • 1 spec.; same data as for preceding; Nov. 1989; A. Bello leg.; CEMT • 2 specs; Brasília, Aeronáutica; 5 Nov. 1981; Bello; CEMT. – Goiás • 1 spec.; Joánapolis, Chacara São José; 27 Feb. 2015; E. Santana leg.; CEMT. – Mato Grosso • 2 specs; Cuiabá, Campus UFMT; Nov. 2009; R. Miyazaki leg.; CEMT. – Minas Gerais • 1 spec.; Astolfo Dutra; Jul. 1974; CEMT • 2 specs; Cordisburgo, F. Pontina; Jul. 1994; de Mello leg.; CEMT • 2 specs; same data as for preceding; Jan. 1999; Falqueto and Vaz-de-Mello leg.; CEMT • 1 spec.; Teixeiras; Nov. 1996; Hardy and Harrison leg.; CEMT • 1 spec.; Viçosa; 22 Feb. 1997; A. P. Lobo leg.; CEMT • 1 spec.; same data as for preceding; 10 May 1997; H. Siqueira leg.; CEMT • 1 spec.; same data as for preceding; 24 May 1997; H. Siqueira leg.; CEMT. – Pará • 1 spec.; São José dos Pinhais; 25 ° 33 ′ S, 48 ° 58 ′ W; elev. 970 m; 13 Oct. 2012; P. B. Santos leg.; CEMT. – Rio de Janeiro • 1 spec.; Angra dos Reis, Bracuí; 10 – 11 May 2002; Nissimian leg.; baixada [lowland]; UV light; CEMT • 1 spec.; Nova Friburgo; Dec. 2002; E. Grossi leg.; CEMT. – São Paulo • 1 spec.; [unreadable]; 23 Oct. 2010; Bello leg.; CEMT. ECUADOR – Napo • 7 specs; SC Yasuní; elev. 250 m; 6 – 14 Sep. 1997; F. Maza leg.; light trap; CEMT • 6 specs; Yasuní; elev. 250 m; 8 Apr. 1997; G. Onore leg.; light trap; CEMT. MEXICO – Sonora • 3 specs; Sierra Madres Occid, Yaqui River Valley; 3 Sep. 1999; Mathison and Singer leg.; CEMT. – Veracruz • 1 spec.; Veracruz-Actopan, La Mancha, Dunas; 27 Sep. 2003; F. Z. Vaz-de-Mello leg.; light; CEMT. NICARAGUA – Carazo • 1 spec.; Bio. Chococente; 11 ° 30 ′ N, 86 ° 10 ′ W; 11 – 13 Sep. 1992; J. M. Maes and López.; CEMT • 1 spec.; same data as for preceding; J. M. Maes and Martínez leg; bosque tropical seco (denso) [tropical dry forest (dense)]. – Leon • 1 spec.; 11 ° 30 ′ N, 86 ° 10 ′ W; 10 Feb. 1993; M. Pogatshnik leg.; UV 220 V; CEMT. UNITED STATES OF AMERICA – Virginia • 1 spec.; Louisa Co., Mineral; 6 Jul. 1990; D. C. Hildebrant leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC2FF95FF61FD3DFCE6FA35.taxon	materials_examined	BRAZIL – Bahia • 1 spec.; Itamaraju; 16 ° 57 ′ 58 ″ S, 39 ° 26 ′ 01 ″ W; Apr. 2011; C. M. P. Leite leg.; pitfall; CEMT • 2 specs; same data as for preceding; 17 ° 01 ′ 09 ″ S, 39 ° 28 ′ 15 ″ W; Apr. 2011; CEMT • 1 spec.; Wenceslau Guimarães, Matriz; 13 ° 41 ′ 21 ″ S, 39 ° 21 ′ 52 ″ W; elev. 146 m; C. M. P. Leite leg.; pitfall; CEMT. – Mato Grosso • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 18 Oct. 2014; V. Koppe leg.; CEMT • 2 specs; Cotriguaçu, Fazenda São Nicolau, PPBioP 1; 9 ° 49 ′ 08 ″ S, 58 ° 15 ′ 40 ″ W; 19 – 24 Oct. 2012; F. Z. Vaz-de-Mello leg.; FIT; CEMT. – Minas Gerais • 2 specs; Ipatinga; Nov. 1994; A. Bello leg.; CEMT • 1 spec.; Santa Barbara; 17 Nov. 1994; J. C. Zanuncio leg.; CEMT • 1 spec.; Viçosa; Feb. 1995; J. N. C. Louzada leg.; forest; CEMT • 1 spec.; Viçosa; 23 Jan. 1995; J. N. C. Louzada leg.; CEMT • 1 spec.; same data as for preceding; 4 Feb. 1995; J. N. C. Louzada leg.; CEMT • 1 spec.; same data as for preceding; 9 Feb. 1996; CEMT • 1 spec.; Viçosa, M. B. U.; 5 Feb. 1996; J. N. C. Louzada leg.; CEMT • 1 spec.; Viçosa, Mata do Paraiso; Jan. 1995; Louzada and Lopes leg.; CEMT • 2 specs; Viçosa, UFV; Dec. 1999; Lopes-Andrade and Vaz-de-Mello leg.; CEMT • 2 specs; same data as for preceding; Nov. 1997; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Viçosa, Mata do Paraiso; Jan. 1994; Louzada and Lopes leg.; CEMT. – Pará • 1 spec.; Belém, IPEAN; Oct. 1984; N. Degallier leg.; FIT; CEMT • 1 spec.; Redenção, Fazenda Marajoara; 7 ° 30 ′ S, 50 ° 16 ′ W; 12 Oct. 1998; P. Y. Scheffler leg.; CEMT • 2 specs; Serra Norte, Carajás; 18 – 20 Oct. 1984; N. Degallier leg.; CEMT. – Paraná • 1 spec.; Rancho Alegre, Fazenda Congonhas; 22 ° 47 ′ 45 ″ S, 51 ° 00 ′ 12 ″ W; 25 Oct. 2012; P. M. Felix leg.; canopy control; CEMT. – Rondônia • 3 specs; Rolim de Moura; 11 ° 44 ′ 25 ″ S, 61 ° 55 ′ 26 ″ W; elev. 273 m; 7 – 9 Dec. 2015; D. C. Castro leg.; SAF, pitfall human dung; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC2FF94FF61FA30FB6AFC80.taxon	materials_examined	BRAZIL – Goiás • 1 spec.; Rio Verde; Feb. 1998; J. Carlos leg.; CEMT. – Maranhão • 3 specs; C. N. Maranhão Rebio, Res. Biol. Gurupi; 3 ° 14 ′ 01 ″ S, 46 ° 40 ′ 52 ″ W; 1 – 5 Jul. 2011; M. M. Abreu and J. A. Silva leg.; light trap; CEMT • 2 specs; Corumbá, Fazenda São Bento; CEMT. – Mato Grosso • 1 spec.; Chapada dos Guimarães, Manso; 14 Sep. 2016; M. Debuss leg.; manually; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 30 Oct. 2014; W. Chamorro leg.; black light trap; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 9 ° 50 ′ 19 ″ S, 58 ° 15 ′ 03 ″ W; 25 Oct. 2014; light; CEMT • 1 spec.; same data as for preceding; 9 ° 51 ′ 20 ″ S, 58 ° 14 ′ 55 ″ W; Oct. 2014; R. V. Nunes leg.; base camp; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 9 Dec. 2009; M. R. Barreto leg.; platio de caixeta [caixeta tree plantation]; luz branca [white light]; CEMT • 1 spec.; Cuiabá; 15 ° 16 ′ 45 ″ S, 56 ° 00 ′ 06 ″ W; 7 Dec. 2015; A. Frolov and L. Akhmenotova leg.; CEMT • 1 spec.; Cuiabá, Boa Esperança; Oct. 2012; F. Z. Vaz-de-Mello leg.; CEMT • 5 specs; Estação Ecológica de Taiamã; 11 Dec. 1981; M. Serrano leg.; CEMT • 1 spec.; Nobres, Distrito Coqueiral; 26 May 2010; D. Nonato leg.; manually; CEMT • 3 specs; Poconé; 29 Jan. 1998; acuri palm tree; light trap; CEMT • 1 spec.; same data as for preceding; 27 Feb. 1998; light trap; CEMT • 25 specs; same data as for preceding; 28 Feb. 1998; light trap; acuri palm tree; CEMT • 8 specs; same data as for preceding; 28 Mar. 1998; light trap; CEMT • 3 specs; same data as for preceding; 23 Jun. 1998; CEMT • 4 specs; same data as for preceding; 24 Jun. 1998; CEMT • 4 specs; same data as for preceding; 26 Apr. 1998; light trap; acuri palm tree; CEMT • 2 specs; same data as for preceding; 28 Apr. 1998; light trap; CEMT • 1 spec.; Santo Antônio do Leverger, Pesqueiro Casa Branca; 23 May 2014; H. D. Biachini leg.; manually; CEMT • 1 spec.; Sesc-Pantanal; 1 Nov. 2001; light trap; CEMT • 4 specs; same data as for preceding; 14 May 2002; CEMT • 2 specs; same data as for preceding; 9 Aug. 2002; CEMT • 2 specs; same data as for preceding; 15 Apr. 2002; CEMT • 1 spec.; same data as for preceding; 11 Jul. 2002; CEMT • 4 specs; same data as for preceding; 5 Aug. 2003; CEMT • 3 specs; same data as for preceding; 6 Sep. 2002; CEMT • 1 spec.; same data as for preceding; 17 Jul. 2004; CEMT • 75 specs; same data as for preceding; 15 Sep. 2004; CEMT • 1 spec.; same data as for preceding; 15 Oct. 2004; CEMT • 2 specs; Varzea Grande, Cuiabá, Parque de Exposição; 10 May 1972; Roger and Williams leg.; blacklight trap; CEMT • 9 specs; same data as for preceding; 30 May 1972; CEMT. – Mato Grosso do Sul • 12 specs; Corumbá, Faz São Bento, Prox Porto Jofre; 14 Apr. 2011; R. V. Nunes leg.; CEMT • 2 specs; Guia Lopes da Laguna; 15 Feb. 2007; A. Abot leg.; light trap; CEMT. – Piauí • 1 spec.; Piracuruca, P. N. Sete Cidades, Olho d’Água dos Milagres; 4 ° 05 ′ 32 ″ S, 41 ° 40 ′ 48 ″ W; elev. 180 m; 12 Feb. 2013; D. M. Takiyama and A. P. M. Santos leg.; Pennsylvania trap; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC3FF9AFF61FCC7FE6FFD9A.taxon	materials_examined	BRAZIL – Bahia • 3 specs; Caravelas; Mar. 1996; R. Dizin leg.; CEMT • 8 specs; same data as for preceding; 25 Mar. 1997; Anjos and Silveira leg.; CEMT • 1 spec.; same data as for preceding; Mar. 1997; R. D. Silveira leg.; CEMT • 1 spec.; Presidente Tancredo Neves; 13 ° 22 ′ 56 ″ S, 39 ° 20 ′ 27 ″ W; Jan. 2011; C. M. P. Leite leg.; pitfall; CEMT • 1 spec.; Wenceslau Guimarães Matriz; 13 ° 41 ′ 21 ″ S, 39 ° 21 ′ 52 ″ W; elev. 146 m; C. M. P. Leite leg.; CEMT. – Ceará • 1 spec.; Ubajara, PN Ubajara, Trilha da Samambaia; 3 ° 50 ′ 30 ″ S, 40 ° 53 ′ 55 ″ W; elev. 850 m; 17 Feb. 2013; D. M. Takiya leg.; light; CEMT. – Distrito Federal • 1 spec.; Brasília; elev. 1100 m; Feb. 2001; N. Degallier leg.; CEMT • 65 specs; Brasília, Fazenda Água Limpa (UNB); Nov. 2010; C. Suinaga leg.; CEMT. – Goiás • 1 spec.; Rio Verde; Sep. 1997; J. Carlos leg.; CEMT. – Maranhão • 7 specs; Mirador, P. E. Mirador, Base Geraldina; 20 – 24 Dec. 2006; R. O. Souza and J. C. Silva leg.; light; CEMT • 1 spec.; São Luís, UEMA, Aléia; Feb. 2009; M. C. Batista leg.; CEMT. – Mato Grosso • 1 spec.; Chapada dos Guimarães; 15 ° 24 ′ 31 ″ S, 55 ° 45 ′ 53 ″ W; 2 Dec. 2015; A. Frolov and L. Akhmetova leg.; light; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 30 Oct. 2014; W. Chamorro leg.; black light; CEMT • 1 spec.; Cuiabá; 15 ° 16 ′ 45 ″ S, 56 ° 06 ′ 06 ″ W; 7 Dec. 2015; A. Frolov and L. Akhmetova leg.; CEMT • 4 specs; Cuiabá, Campus UFMT; 9 Oct. 2016; A. Frolov leg.; UV light; CEMT • Cuiabá, Fazenda Mutuca; 15 ° 19 ′ 31 ″ S, 55 ° 08 ′ 03 ″ W; elev. 285 m; Nov. 2015; R. V. Nunes and A. Frolov leg.; light; CEMT • 4 specs; Diamantino; Jan. 2001; A. Bello leg.; CEMT • 3 specs; Diamantino, Alto Rio Arinos; Oct. 2001; E. Furtado leg.; CEMT • 1 spec.; Paranaita, Margem Esquerda Teles Pieres; 9 ° 19 ′ 47 ″ S, 56 ° 47 ′ 49 ″ W; Sep. 2011; A. P. Benelli leg.; CEMT • 1 spec.; Porto Estrela, EES das Araras, base camp; 15 ° 39 ′ 09 ″ S, 57 ° 12 ′ 53 ″ W; elev. 217 m; 8 – 12 Oct. 2011; Vaz-de-Mello leg.; light; CEMT • 1 spec.; Santo Antônio do Leverger, São Vicente da Cerra; 15 ° 49 ′ 42 ″ S, 55 ° 25 ′ 11 ″ W; 16 – 18 Feb. 2011; A. S. O. Tissiani leg.; forest; pitfall human dung; CEMT • 2 specs; Sesc-Pantanal; 6 Sep. 2002; light trap; CEMT • 1 spec.; Sinop; R. L. Silva leg.; manually; CEMT • 1 spec.; Sinop, Campus UFMT; R. L. Silva leg.; manually; CEMT • 1 spec.; Vila Bela, Margem Guaporé; 1 Oct. 1984; S. Marcolina leg.; light; CEMT. – Minas Gerais • 2 specs; Águas Vermelhas; Dec. 1997; A. Bello leg.; CEMT • 1 spec.; same data as for preceding; A. Bello and F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Ijací, Faz FAEPE; Nov. 2002; J. Louzada leg.; CEMT • 2 specs; Ipatinga; Nov. 1994; Bello leg.; CEMT • 1 spec.; Juiz de Fora; Nov. 1974; G. S. Andrade leg.; CEMT • 2 specs; Santana do Riacho, PARNA Serra do Cipó; 11 Dec. 2004; G. S. Schiffler leg.; forest; CEMT • 1 spec.; Teixeiras; Nov. 1996; Hardy and Harrison leg.; CEMT • 2 specs; Viçosa; Jan. 1995; J. Louzada leg.; CEMT. – Paraíba • 1 spec.; Est. Rio Mamanguape; 29 – 31 Jul. 2000; Deyse and Araujo leg.; light trap; CEMT • 1 spec.; same data as for preceding; 1 – 7 Jun. 2000; Malaise trap; CEMT • 1 spec.; João Pessoa; Campus UFPB; 17 May 1980; R. Baltasar leg.; CEMT • 1 spec.; Rio Tinto, APA Barra do Rio Mamanguape; 6 ° 46 ′ 41 ″ S, 34 ° 55 ′ 37 ″ W; 27 Mar. 2012; D. Dal-Bó leg.; CEMT. – Pernambuco • 1 spec.; Fernando de Noronha, Sancho; 20 – 28 Feb. 2020; Grossi and Rafael leg.; light; CEMT • 1 spec.; Igarassu, RECD; 11 May 2007; Silva leg.; CEMT. – Rio Grande do Norte • 1 spec.; Natal, EMPARN; 5 ° 45 ′ 56 ″ S, 35 ° 11 ′ 21 ″ W; elev. 32 m; 23 May 2007; J. A. Rafael and F. Xavier leg.; light; CEMT. – São Paulo • 1 spec.; Descalvado, Fazenda Itaúnas; 24 Aug. 2006: N. W. Perioto leg.; forest + Citrus plantation; pitfall; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC3FF9AFF61FCC7FE6FFD9A.taxon	discussion	Collecting information Specimens were collected at altitudes ranging from 146 to 2400 metres in regions of Cerrado, Atlantic and Amazon forest. These insects can be collected with UV light and flight intercept traps and are attracted to human and cow dung (Stebnicka 2007 c). Remarks The Ataenius texanus-carinator group has over 40 recognized species, 11 are registered in Brazil (Stebnicka 2007 c).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCDFF9AFD95FDCDFA83FB19.taxon	diagnosis	Diagnosis Length 5.5 – 6.5 mm. Body oval, piceous. Head broad, strongly punctate. Pronotum transverse, lateral margins strongly explanate, surface densely and coarsely punctate. Elytra widened posteriorly, with tubercles variably tuberculate. Meso- and metatibiae, as well as tarsus, densely setaceous ventrally; apical spurs sometimes hidden by setae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCDFF9AFD95FDCDFA83FB19.taxon	discussion	Remarks Batesiana currently has two known species: B. chamorroi and B. tuberculata (Skelley & Vaz-de-Mello 2023).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCDFF9AFE3FFB4CFD26F80B.taxon	description	Figs 17 F, 44 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCDFF9AFE3FFB4CFD26F80B.taxon	diagnosis	Diagnosis Length 7.5 mm. Head, with coarse and wide punctures; clypeus anteriorly bent at an angle of nearly 90 ° degrees before median gibbosity. Pronotum coarsely and densely punctate, with a large lateral fovea; lateral margin explanate, with posterior angles strongly projected. Elytra tuberculate; tubercles distended anteriorly, rounded posteriorly. Meso- and metatibiae, as well as tarsi densely setaceous ventrally; apical spur visible amidst setae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCDFF9AFE3FFB4CFD26F80B.taxon	materials_examined	Material examined BRAZIL – Roraima • 1 spec.; P. N. Monte Roraiman; 5 ° 07 ′ 31 ″ N, 60 ° 35 ′ 50 ″ W; elev. 1000 m; Dec. 2019; F. Oliveira, F. Xavier and S. Lima leg.; em igarapé [in stream]; light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCDFF9AFE3FFB4CFD26F80B.taxon	description	Sexual dimorphism Unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCDFF9AFE3FFB4CFD26F80B.taxon	discussion	Collecting information This species can be collected with light traps.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCEFF99FD9EFE95FB80F8C3.taxon	description	Figs 17 E, 44 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCEFF99FD9EFE95FB80F8C3.taxon	diagnosis	Diagnosis Length 5.3 – 6.6 mm. Head strongly punctate, gibbous medially; clypeus not sinuate, with glabrous band near anterior margin. Pronotum with lateral margins explanate, posterior angles right-angled; punctate with coarse punctures, punctures larger and closer near lateral margins. Elytra with interstriae tuberculate. Meso- and metatibiae, as well tarsi, densely setaceous ventrally; apical spurs sometimes hidden by setae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCEFF99FD9EFE95FB80F8C3.taxon	materials_examined	Material examined BRAZIL – Acre • 1 spec.; Assis Brasil; 10 ° 35 ′ 44 ″ S, 69 ° 37 ′ 11 ″ W; 18 Sep. 2016; Ortiz and Borges leg.; light trap (CDC); CEMT • 1 spec.; same data as for preceding; 23 Sep. 2016; CEMT. – Amazonas • 1 spec.; Manaus; Jun. 1955; Martínez leg.; FSCA. – Mato Grosso • 1 spec.; Chapada dos Guimarães; 15 ° 01 ′ 18 ″ S, 55 ° 15 ′ 04 ″ W; 27 Oct. 2006; R. L. Ferreira leg.; CEMT • 2 specs; Cotriguaçu, Fazenda São Nicolau, PPBIO; Sep. 2010; F. P. Florêncio leg.; Orbignya phalerata; CEMT. – Pará • 1 spec.; Itaituba, Mina do Palito; 6 ° 20 ′ 16 ″ S, 55 ° 47 ′ 45 ″ W; 27 Jan. 2018; K. K. G. Silva and L. P. Prado leg.; CEMT • 2 specs; Novo Progresso, Fazenda Florentino; 7 ° 08 ′ 11 ″ S, 55 ° 23 ′ 42 ″ W; elev. 230 m; Oct. – Nov. 2010; D. Krinski leg.; Amazon forest; light; CEMT. – Rondônia • 1 spec.; Nova Mamoré; 10 ° 19 ′ 47 ″ S, 64 ° 45 ′ 12 ″ W; elev. 190 m; Jan. – Feb. 2010; FIT; F. Coletti leg.; Mata 3; CEMT • 1 spec.; Porto Velho, Rio das Graças, Bom Jesus; 8 ° 49 ′ 47 ″ S, 63 ° 46 ′ 51 ″ W; 22 Oct. 2017; D. C. Santos and K. K. G. Silva leg.; FIT; CEMT. COLOMBIA – Chocó • 2 specs; Jurubida; elev. 100 m; 9 Sep. 1995; P. Duque leg.; litter; CEMT. FRENCH GUIANA • 1 spec.; Cayenne, Kourou, Forêt de Wayabo; 1 Dec. 2012; M. Duranton leg.; CEMT • 2 specs; Montagne des Chavaux; 4 ° 44 ′ 56 ″ N, 52 ° 26 ′ 28 ″ W; elev. 75 m; 4 Apr. 2014; SAEG leg.; CEMT • 1 spec.; same data as for preceding; 11 Feb. 2013; FIT (V); CEMT • 3 specs; same data as for preceding; 21 Dec. 2013; CEMT • 1 spec.; same data as for preceding; 25 May 2013; light (GEML); CEMT. PANAMA – Colón • 1 spec.; Gamboa; elev. 50 m; 4 May 2007; Curoe leg.; FIT; CEMT. UNKNOWN COUNTRY – 1 spec.; [unknown location]; A 2 FIT; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCEFF99FD9EFE95FB80F8C3.taxon	diagnosis	Sexual dimorphism Males with fifth abdominal ventrite shorter and disc of pygidium broader than that of female.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCEFF99FD9EFE95FB80F8C3.taxon	discussion	Collecting information Specimens can be collected in lights and with flight intercept traps. Some specimens were found associated with Attalea speciosa Mart. (as Orbignya phalerata on the label of the examined material) palm trees. Skelley & Vaz-de-Mello (2022) discussed the association of B. tuberculata with Asterocaryum paramaca Mart., since adults and larvae were found nesting in the palm’s crown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCFFF98FDC2FCFAFDC6F802.taxon	description	Figs 27 F, 58 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCFFF98FDC2FCFAFDC6F802.taxon	diagnosis	Diagnosis Length 3.0 mm. Head with clypeus sinuate; frontoclypeal and genal suture elevated. Pronotal lateral margins sinuated; disc with one transverse, almost completely glossy and strongly punctate anterior carina with three longitudinal carinae parting from it; lateral carinae interrupted by oblique furrow; middle carina bifurcate near posterior margin; all carinae glabrous and glossy. Elytra with humeral denticle fine but evident.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCFFF98FDC2FCFAFDC6F802.taxon	materials_examined	Material examined ARGENTINA – Corrientes • 4 specs; Corrientes, Barrio Víctor Colas; 27 ° 28 ′ 50 ″ S, 58 ° 47 ′ 18 ″ W; elev. 64 m; 21 Oct. 2012; M. I. Polesel et al. leg.; CEMT. BRAZIL – Goiás • 1 spec.; Goiânia; 12 Jan. 2002; S. S. Silva leg.; CEMT. – Mato Grosso • 1 spec.; Poconé, Parque Sesc Baixa das Pedra; 13 Sep. 2018; G. B. Silva and L. G. P. Dantas leg.; pitfall; CEMT • 1 spec.; Poconé, Sesc Pantanal; Nov. 2016; S. Montoya leg.; CEMT. – Mato Grosso do Sul • 2 specs; Corumbá, Passo do Lontra; Feb. 1996; J. Louzada and F. Z. Vaz de Mello leg.; human dung; CEMT. – Paraná • 1 spec.; Londrina; Nov. – Dec. 1998; I. M. Medri leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCFFF98FDC2FCFAFDC6F802.taxon	description	Sexual dimorphism Males with apical spur of protibiae long, downwardly and inwardly bent. Penultimate abdominal ventrite shortened. Female spur almost straight, weakly downwardly bent. Penultimate abdominal ventrite similar to the others.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCFFF98FDC2FCFAFDC6F802.taxon	discussion	Collecting information Cartwrightia cartwrighti were collected at altitudes ranging from 64 to 1000 metres. Specimens were collected with flight intercept and black light traps, in coffee forest litter, and appear to be attracted to human dung (Cartwright 1967).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCFFF98FDBCFE94FEE6FCF3.taxon	diagnosis	Diagnosis Length 3.0 – 3.4 mm. Body elongate, surface partially covered with argillaceous coating. Head with frontoclypeal and genal suture evident; clypeus sinuate, medially with longitudinal carina; anterior margin thickened, downwardly bent and acuminate medially. Pronotum coarsely punctate, punctures scattered; lateral margin sinuate; five incomplete longitudinal carinae present on disc, medial carina bifurcate medially. Elytra with odd interstriae carinate; even interstriae flat; striae not excavate, evidenced by row of coarse punctures lateral to interstriae; posterior declivity with a single big tubercle near posterior margin. Protibia with two to three teeth, first and second teeth closer than third; first tooth positioned apically. Basal tarsomere of meso- and metatarsus as long as the following three combined.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCFFF98FDBCFE94FEE6FCF3.taxon	discussion	Remarks Cartwrightia comprises three recognized species C. islasi Cartwright, 1967, C. cartwrighti (Mexico, El Salvador and Guatemala) and C. intertribalis Islas, 1958 (Mexico). For keys to all species see Cartwright (1967).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC8FF9FFD93FE94FB13FD16.taxon	diagnosis	Diagnosis Length 3.2 – 3.4 mm. Body, elongate, dorsal surface shagreened, strongly punctate, dark brown. Head with clypeal margin serrate. Pronotum strongly punctate, lateral and posterior margins serrate. Elytra strongly punctate; epipleuron covering mesepimeron and metepisternum (see Fig. 32 B). Mesocoxa elongate, reaching elytral epipleuron.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC8FF9FFD93FE94FB13FD16.taxon	discussion	Remarks Euparixoides has five recognized species (Stebnicka & Skelley 2005; Stebnicka 2009). This genus has been reported from Brazil, Costa Rica, Honduras, Nicaragua, Panama, Peru and Venezuela (Stebnicka & Skelley 2005; Stebnicka 2009). For a key to all its species see Stebnicka & Skelley (2005).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC8FF9FFDF2FD57FD2DFA33.taxon	description	Figs 18 F, 45 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC8FF9FFDF2FD57FD2DFA33.taxon	diagnosis	Diagnosis Length 3.2 – 3.4 mm. Head with clypeus emarginate, shagreened, finely punctate; anterior margins serrate; frons more strongly punctate than clypeus. Pronotum with anterior margin explanate coarsely punctate; lateral and posterior margins serrate; posterior margins sinuate; posterior angles prominent, acuminate. Elytra with striae coarsely punctate, wider than interstriae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC8FF9FFDF2FD57FD2DFA33.taxon	materials_examined	Material examined BRAZIL – Rondônia • 1 spec.; Parque Nacional Guajara-Mirim; 10 ° 19 ′ S, 64 ° 32 ′ W; 22 Nov. 2016; M. A. P. A. Silveira leg.; FIT; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC8FF9FFDF2FD57FD2DFA33.taxon	description	Sexual dimorphism Unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC8FF9FFDF2FD57FD2DFA33.taxon	discussion	Collecting information Specimens collected with flight intercept trap.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC8FF9FFDF6FA38FB7DF96F.taxon	diagnosis	Diagnosis Length 5.0 – 6.0 mm. Body robust, glabrous, piceous. Head with clypeus strongly gibbous medially. Pronotum transverse; anterior angles explanate; lateral margins not fringed by setae. Elytra with interstriae glabrous and flat, except for ninth one, which is carinate and expanded, forming a pseudoepipleuron.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC8FF9FFDF6FA38FB7DF96F.taxon	discussion	Remarks Flechtmanniella is a monotypic genus reported only from Brazil (Stebnicka 2009).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC8FF9EFDCBF89CFDC9FB67.taxon	description	Figs 17 C – D, 45 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC8FF9EFDCBF89CFDC9FB67.taxon	diagnosis	Diagnosis Same as for the genus.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC8FF9EFDCBF89CFDC9FB67.taxon	materials_examined	Material examined BRAZIL – Bahia • 2 specs; Caravelas; Mar. 1997; R. Diniz leg.; CEMT. – Mato Grosso • 1 spec.; Cuiabá, Flor do Cerrado; 15 ° 29 ′ 38 ″ S, 56 ° 4 ′ 36 ″ W; 15 Nov. 2015; A. Frolov leg.; CEMT • 2 specs; Cuiabá, Res. Coxipó; Oct. 2018; F. M. Lima leg.; light; CEMT • 1 spec.; Diamantino, Reserve Vale da Solidão; 14 ° 22 ′ S, 56 ° 07 ′ W; elev. 450 m; 29 Oct. 2013; E. Furtado leg.; CEMT • 1 spec.; Porto Estrela, EE Serra das Araras, heliporto; 15 ° 39 ′ 19 ″ S, 57 ° 12 ′ 50 ″ W; 21 Nov. 2017; R. J. P. Machado and R. Nunes leg.; light; CEMT • 1 spec.; Nova Xavantina, Fazenda Abonizia; 14 ° 45 ′ 52 ″ S, 52 ° 33 ′ 17 ″ W; elev. 406 m; 14 – 16 Feb. 2019; R. J. Silva and R. S. A. Silva leg.; Cerrado Campo Limpo com área inundada; pitfall hum dung; CEMT. – Mato Grosso do Sul • 1 spec.; Guia Lopes da Laguna; 15 Nov. 2007; A. Abot leg.; light trap; CEMT • 1 spec.; Selvíria, UNESP farm; 30 Oct. 2000; C. A. H. Flechtmann leg.; Brachiaria decumbens pasture; black light trap; CEMT • 1 spec.; Três Lagoas, Horto Rio Verde, Tres Lagoas Agroflorestal; 15 Feb. 1994; C. A. H. Flechtmann leg.; Cerrado satand; black light trap; CEMT • 1 spec.; same data as for preceding; 19 Oct. 2003; CEMT. – Minas Gerais • 1 spec.; Buenopolis, PE Serrado Cabral; 17 ° 55 ′ 18 ″ S, 44 ° 14 ′ 26 ″ W; 17 Apr. 2008 M. F. Souza leg.; 1041 mosl, hum dung; CEMT. – São Paulo • 1 spec.; Anhembi, ESALQ / USP; E. N. L. Ferreira leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC8FF9EFDCBF89CFDC9FB67.taxon	description	Sexual dimorphism Males with apical spur of protibiae downward and inward curved; males with thicker tarsus when compared with females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC8FF9EFDCBF89CFDC9FB67.taxon	discussion	Collecting information Specimens were collected with light traps in pastures of Urochloa decumbens and Eucalyptus grandis W. Hill ex Maiden forests. Some specimens were also found in nests of Acromyrmex lobicornis (Emery, 1888) (Stebnicka 1999 a, 2007 a).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC9FF9DFDBFFB61FA83FC9A.taxon	diagnosis	Diagnosis Length 4.5 – 5.0 mm. Body shagreened, reddish brown to piceous. Head with longitudinal wrinkles; clypeus emarginate, amply gibbous. Pronotum convex, impunctate or with wide and coarse punctures; posterior and lateral margins excavated, with wide furrow posteriorly; anterior angles and lateral margins explanate undulate. Elytra with striae strongly punctate, crenating margin of interstriae; interstriae three, five and seven with weak tubercle anteriorly, fifth stria depressed anteriorly; humeral region prominent, crowned with group of setae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFC9FF9DFDBFFB61FA83FC9A.taxon	discussion	Remarks Iarupea has five recognized species and all of them can be found in Brazil. But this genus is not restricted to Brazil, as it has been found in Argentina, Bolivia, Suriname and Guyana (Stebnicka 2007 a, 2009).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCAFF9DFDB8FCCCFADDF98C.taxon	description	Figs 14 D, 45 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCAFF9DFDB8FCCCFADDF98C.taxon	diagnosis	Diagnosis Length 5.0 mm. Body dark reddish brown. Pronotum without punctures, instead covered with small tubercles; posterior furrow not strigate; some specimens present the posterior margin of pronotum, before furrow, medially acuminate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCAFF9DFDB8FCCCFADDF98C.taxon	materials_examined	Material examined BRAZIL – Maranhão • 2 specs; Carolina, Fazenda Cincorá; 22 Oct. 2009; F. Lima-de-Oliveira, R. O. Souza, M. B. Aguiar Neto leg.; light; CEMT. – Minas Gerais • 1 spec.; Cordisburgo; Jan. 1994; F. Z. Vaz de Mello leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCAFF9DFDB8FCCCFADDF98C.taxon	description	Sexual dimorphism Males unknown, but based on the other species of the genus, males probably have the apical spur of protibiae strongly arched when compared with the females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCAFF9DFDB8FCCCFADDF98C.taxon	discussion	Collecting information Specimens were collected with light traps and in galleries of nests of Atta sp. (Stebnicka 2007 a).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCAFF9CFE22F9FAFDE6FDCB.taxon	description	Figs 15 C – D, 45 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCAFF9CFE22F9FAFDE6FDCB.taxon	diagnosis	Diagnosis Length 4.5 – 5.0 mm. Body piceous. Pronotum everywhere punctate with large, not confluent and coarse punctures; posterior margin, near posterior angles, with a single tubercle.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCAFF9CFE22F9FAFDE6FDCB.taxon	materials_examined	Material examined BRAZIL – Mato Grosso • 2 specs; Varzea Grande, Sítio Toca da Coruja; 15 ° 33 ′ 07 ″ S, 56 ° 19 ′ 07 ″ W; elev. 230 m; 18 Oct. 2020; A. Petini-Benelli leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCAFF9CFE22F9FAFDE6FDCB.taxon	description	Sexual dimorphism Males with apical spur of protibiae strongly curved; female apical spur short, almost straight.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCAFF9CFE22F9FAFDE6FDCB.taxon	discussion	Collecting information Specimens were attracted to black light traps and found in waste piles from nests of Atta vollenweideri Forel, 1893 (Martínez 1953).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCBFF9CFDBCFD02FDCAFA02.taxon	description	Figs 14 E, 45 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCBFF9CFDBCFD02FDCAFA02.taxon	diagnosis	Diagnosis Length 4.8 – 5.0 mm. Body dark reddish brown. Head with frons coarsely punctate; punctures coalescent. Pronotum densely punctate with large punctures; punctures can be coalescent; lateral margins clearly explanate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCBFF9CFDBCFD02FDCAFA02.taxon	materials_examined	Material examined BRAZIL – Maranhão • 1 spec.; Pedrinhas, São Luís Island; 29 Aug. 1984; E. C. Bergmann leg.; alluvial ombrophylous dense forest; blacklight; FIT; CEMT • 1 spec.; same data as for preceding; 11 Mar. 1987; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCBFF9CFDBCFD02FDCAFA02.taxon	description	Sexual dimorphism Males unknown, but based on the other species of the genus, males probably have the apical spur of protibiae strongly arched when compared with the females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCBFF9CFDBCFD02FDCAFA02.taxon	discussion	Collecting information Specimens were collected with black light and flight intercept traps in mangrove areas and extracted from Atta sp. nests (Stebnicka 2007 a).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCBFF82FDE1FA45FD39FC94.taxon	description	Figs 14 F, 45 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCBFF82FDE1FA45FD39FC94.taxon	diagnosis	Diagnosis Length 4.8 – 5.0 mm. Body dark reddish brown, pronotum microreticulate, elytra strongly shagreened. Head with frons with scarce punctures. Pronotum punctate with large and oval punctures; punctures closer and smaller near anterior margin, not coalescent; near posterior margins, punctures larger and more widely spaced; lateral margins weakly explanate, strongly sinuate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCBFF82FDE1FA45FD39FC94.taxon	materials_examined	Material examined BRAZIL – Amazonas • 1 spec.; Querari Pelotão; 1 ° 05 ′ N, 69 ° 51 ′ W; 7 – 23 Apr. 1993; C. Motta et al. leg.; CEMT • 1 spec.; Rio Urubu; 2 ° 10 ′ S, 59 ° 49 ′ W; 12 – 13 Mar. 83; P. Bührnheim and N. Otaviano leg.; CEMT • 1 spec.; Manaus, Embrapa-CPAA projeto SHIFT, 010 Km 24 - 251; 17 Jun. 1993; A. Pamplona leg.; light trap; CEMT. – Maranhão • 1 spec.; São Luis Island, Pedrinhas; 6 May 1987; E. C. Bergmann leg.; mangrove area, MA 23 ENDO 13; black light trap; CEMT. – Pará • 1 spec.; Monte Dourado; Jan. 1993; CEMT • 1 spec.; same data as for preceding; Aug. 1993; CEMT • 1 spec.; same data as for preceding; Dec. 1992; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCBFF82FDE1FA45FD39FC94.taxon	description	Sexual dimorphism Males with spur of protibia inwardly hooked; females with spur straight.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFCBFF82FDE1FA45FD39FC94.taxon	discussion	Collecting information Specimens were collected in mangrove areas with black light and from detritus cavities of nests of Atta sexdens (Linnaeus, 1758) (Stebnicka 2007 a).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD5FF86FDECFCD2FB43FE2B.taxon	description	Figs 15 A – B, 46 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD5FF86FDECFCD2FB43FE2B.taxon	diagnosis	Diagnosis Length 4.5 – 5.0 mm. Body dark reddish brown. Head with longitudinal wrinkles posteriorly convergent anteriorly to frontoclypeal suture; clypeus emarginate. Pronotal surface with coarse, big and wide punctures, margins between punctures sometimes obsolete giving the pronotal surface a wrinkled appearance; anterior angles explanate; lateral margins weakly explanate, strongly sinuate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD5FF86FDECFCD2FB43FE2B.taxon	materials_examined	Material examined BRAZIL – Acre • 1 spec.; Rio Branco; CEMT. – Distrito Federal • 1 spec.; Brasília; Jan. 2001; N. Degallier leg.; CEMT • 1 spec.; same data as for preceding; elev. 1100 m; Mar. 2001; light; CEMT • 1 spec.; same data as for preceding; Jan. 2003; CEMT • 1 spec.; Planaltina; 15 ° 36 ′ 24 ″ S, 47 ° 44 ′ 42 ″ W; 7 Nov. 2015; A. Specht leg.; light; CEMT • 1 spec.; same data as for preceding; 4 Apr. 2016; CEMT. – Espírito Santo • 1 spec.; [Barra de São Francisco], Corrego Itá; Nov. 1954; W. Zikan leg.; CEMT. – Mato Grosso • 1 spec.; Diamantino, Reserve Vale da Solidão; 14 ° 22 ′ S, 56 ° 07 ′ W; elev. 450 m; 30 Oct. 2013; E. Furtado leg.; CEMT. – Mato Grosso do Sul • 1 spec.; Selvíria, UNESP farm; 18 Mar. 1999; C. A. H. Flechtmann leg.; Brachiaria decumens pasture; black light trap; CEMT • 1 spec.; same data as for preceding; 27 May 1999; C 607; CEMT • 1 spec.; same data as for preceding; Bubalus bubalis dropping; 29 Aug. 1993; CEMT. – Minas Gerais • 1 spec.; Cordisburgo; Jan. 1998; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Itumirim; 21 ° 20 ′ 24 ″ S, 44 ° 48 ′ 6 ″ W; 19 Oct. 2016; Viera, F. Z. Vaz-de-Mello and J. Louzada leg.; light; CEMT. – Piauí • 2 specs; Teresina, Embrapa CPAMN; 4 Nov. 2011; M. Bevilaqua and R. Querino leg.; CEMT • 1 spec.; same data as for preceding; 5 ° 02 ′ 15 ″ S, 42 ° 47 ′ 13 ″ W; elev. 61 m; 10 Jun. 2011; R. B. Querino leg.; CEMT. – Rio de Janeiro • 1 spec.; Nova Friburgo; Jan. 1999; P. Grossi leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD5FF86FDECFCD2FB43FE2B.taxon	description	Sexual dimorphism Males with protibial apical spine downwardly and inwardly bent.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD5FF86FDECFCD2FB43FE2B.taxon	discussion	Collecting information Specimens were collected in Urochloa decumbens pastures with black light traps and extracted from Bubalus bubalis (Linnaeus, 1758) faecal masses. Specimens were also collected in Atta sexdens (Stebnicka 2007 a). Specimens were found at altitudes ranging from 61 to 1100 metres.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD1FF86FD92FE22FA84FAC6.taxon	diagnosis	Diagnosis Length 4.8 – 5.2 mm. Body broad, oval, dorsally pilose, covered with testaceous setae, reddish brown. Head variably sculptured; clypeus sinuate. Pronotum with anterior angles forwardly explanate; lateral margins fringed with setae; posterior angles densely setaceous. Elytra wide, setaceous, striae impressed. Mesocoxal cavity partially enclosed by meso and meta ventrites, elongate, almost reaching epipleuron; mesepimeron and metepisternum not visible (see Fig. 32 B).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD1FF86FD92FE22FA84FAC6.taxon	discussion	Remarks Lomanoxia has seven recognized species and has been reported from Argentina, Brazil, Costa Rica, Paraguay and Surinam (Skelley & Howden 2003; Stebnicka 2009). A key for all species can be found in Skelley & Howden (2003).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD1FF85FE34FA00FDB3FDDE.taxon	description	Figs 18 E, 46 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD1FF85FE34FA00FDB3FDDE.taxon	diagnosis	Diagnosis Length 4.8 – 5.0 mm. Body extremely broad. Head minutely granulated; clypeus emarginate. Pronotum setaceous; anterior angles expanded forward; posterior margin of pronotum without lobe. Elytra wider than pronotum. Abdomen with only fifth ventrite strigate anteriorly.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD1FF85FE34FA00FDB3FDDE.taxon	materials_examined	Material examined BRAZIL – Rondônia • 1 spec.; Santo Antônio, Rio Madeira, Margem Direita; 1 Nov. 2004; Luminosa leg.; CEMT. FRENCH GUIANA • 1 spec.; Cayenne, Montage des Chevaux; 4 ° 44 ′ 56 ″ N, 52 ° 26 ′ 28 ″ W; elev. 75 m; 27 Jan. 2013; SEAG leg.; CEMT. TRINIDAD AND TOBAGO – Tunapuna-piarco • 2 specs; Arima Valley, Simla (Ny Zoo. Soc. Sta); 11 Jun. 1977; R. E. Woodruff leg.; Atta nest debris; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD1FF85FE34FA00FDB3FDDE.taxon	description	Sexual dimorphism Pygidium of male with sharp transverse groove and depressed opaque area; female with pygidium raised and glossy.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD1FF85FE34FA00FDB3FDDE.taxon	discussion	Collecting information Specimens were collected with light traps and inside nests of Atta sp. and Atta cephalotes (Linnaeus 1758) (Skelley & Howden 2003).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD2FF85FDE3FD09FC4CF805.taxon	description	Figs 18 A – B, 46 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD2FF85FDE3FD09FC4CF805.taxon	diagnosis	Diagnosis Length 4.5 – 5.0 mm. Head with minute granules; clypeus sinuate. Pronotum setaceous; anterior angles expanded forward; lateral margins of pronotum contiguous to posterior margin, fringed by setae of similar size of those of posterior angles; posterior margin of pronotum with medial lobe. Elytra broad, not broader than pronotum; interstriae tectiform, with minute setaceous elevations. Abdomen with ventrites four and five strigate anteriorly.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD2FF85FDE3FD09FC4CF805.taxon	materials_examined	Material examined BRAZIL – Goiás • 1 spec.; Niquelândia; Oct. 1993; CEMT. – Mato Grosso • 1 spec.; Rondonópolis, P. E. Dom Osório Stoffel; 16 ° 38 ′ S, 54 ° 45 ′ W; 6 Mar. 2021; V. Costa-Silva leg.; light; CEMT. – Mato Grosso do Sul • 1 spec.; Selvíria, UNESP Farm; 3 Oct. 1993; C. A. H. Flechtmann leg.; Brachiaria decumbens pasture; black light trap; CEMT • 1 spec.; Três Lagoas, Horto Novo Palmito Três Lagoas Agroflorestal; 4 Mar. 1993; C. A. H. Flechtmann leg.; black light trap; CEMT • 1 spec.; Três Lagoas, Horto Rio Verde, Três Lagoas Agroflorestal; 23 Nov. 1993; C. A. H. Flechtmann leg.; Cerrado stand; black light trap; CEMT • 1 spec.; Três Lagoas, Horto Barra do Moeda Três Lagoas Agroflorestal; 30 Nov. 1993; C. A. H. Flechtmann leg.; Eucalyptus grandis stand; black light trap; CEMT • 1 spec.; Três Lagoas, Horto Rio Verde, Três Lagoas Agroflorestal; 12 Jan. 1995; C. A. H. Flechtmann leg.; Eucalyptus grandis stand; CEMT. – Minas Gerais • 2 specs; Teixeiras; Nov. 1996; Hardy and Harrison leg.; CEMT. – Rio de Janeiro • 1 spec.; Rio de Janeiro, Floresta da Tijuca; J. E Celso leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD2FF85FDE3FD09FC4CF805.taxon	description	Sexual dimorphism Males and females are indistinguishable from each other.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD2FF85FDE3FD09FC4CF805.taxon	discussion	Collecting information Specimens were collected with UV light traps in Eucalyptus grandis forests and Urochloa decumbens pastures. Some specimens were extracted from nests of Atta sexdens (Stebnicka 2007 a). Remarks This species is extremely similar to Lomanoxia melloi. In addition to the characters provided in the key, Lomanoxia costulata can be identified by a subtly broader elytra.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD3FF84FD9CFE94FD4BFB1D.taxon	description	Figs 18 D, 46 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD3FF84FD9CFE94FD4BFB1D.taxon	diagnosis	Diagnosis Length 5.0 – 5.2 mm. Head with minute granules; clypeus sinuate. Pronotum setaceous; anterior angles expanded forward; posterior margin without medial lobe, semicircular. Elytra with interstriae setaceous and flattened. Abdomen with ventrites four and five strigate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD3FF84FD9CFE94FD4BFB1D.taxon	materials_examined	Type material examined Paratype BRAZIL • 1 ♀; “ [white paper, handwritten] BR-MS- Campo / Grande ex pitfall trap / Koller W. W. / [to the side] 10 - VII- 1995. // [white paper, handwritten] CODE / C- 11. // [blue paper, prited] Paratype / Lomanoxia / ituensus sp. n. / dt. Z. Stebnicka. // [white paper, printed] Q. R. code to the side / CEMT / CUIABÁ / 00111112 ”; CEMT. Other material examined BRAZIL – Minas Gerais • 1 spec.; Lavras; 3 Dec. 97; CEMT. – Rio de Janeiro • 1 spec.; Pedro do Rio; Dec. 1993; A. Bello leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD3FF84FD9CFE94FD4BFB1D.taxon	description	Sexual dimorphism Males and females are indistinguishable from each other.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD3FF84FD9CFE94FD4BFB1D.taxon	discussion	Collecting information Specimens were collected with pitfall traps.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD3FF8AFD95FB49FC66FB65.taxon	description	Figs 18 C, 47 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD3FF8AFD95FB49FC66FB65.taxon	diagnosis	Diagnosis Length 4.8 – 5.0 mm. Head with minute granules; clypeus sinuate. Pronotum setaceous; anterior angles expanded forward; lateral margin fringed with setae, sinuate before posterior angles; setae before and at sinuate area, larger than those at the posterior angles. Elytra with interstriae weakly convex, with row of setaceous granules. Abdomen with ventrites four and five strigate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD3FF8AFD95FB49FC66FB65.taxon	materials_examined	Type material examined Paratype BRAZIL • 1 ♀ “ [white paper, handwritten] BRASIL: MG / Cordisburgo / I- 1998 / Vaz-de-Mello. // [blue paper, prited] Paratype / Lomanoxia / ituensus sp. n. / dt. Z. Stebnicka. // [white paper, printed] Q. R. code to the side / CEMT / CUIABÁ / 00111115 ”; CEMT. Other material examined BRAZIL – Bahia • 1 spec.; Caravelas; 20 Jan. 1997; Anjos and Silva leg.; CEMT. – Distrito Federal • 1 spec.; Brasília; Dec. 2008; N. Degallier leg.; CEMT • 2 specs; Planaltina; 15 ° 36 ′ 24 ″ S, 47 ° 44 ′ 42 ″ W; 4 Nov. 2015; A. Specht leg.; light; CEMT • 2 specs; same data as for preceding; 5 Nov. 2015; light; CEMT • 1 spec.; same data as for preceding; 8 Nov. 2015; CEMT • 1 spec.; Planaltina, Embrapa Cerrados; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 18 Nov. 2005; C. Oliveira leg.; agricultural area; light; CEMT. – Goiás • 1 spec.; Goiânia; 13 Jan. 1997; P. B. Silva leg.; CEMT. – Maranhão • 1 spec.; Mirador, PE Mirador, Base Geraldina; 20 – 24 Dec. 2006; R. O. Sousa and J. C Silva leg.; light; CEMT. – Mato Grosso • 4 specs; Diamantino, Alto Rio Arinos; Jan. 2001; E. Furtado leg.; CEMT. – Minas Gerais • 2 specs; Cordisburgo, Fazenda Pontinha; Jan. 1998; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Guanhães; Oct. 1991; CEMT • 9 specs; Ijaci, Fazenda FAEPE; Nov. 2002; J. Louzada leg.; light; CEMT • 1 spec.; same data as for preceding; Nov. 2002; CEMT • 1 spec.; Itumirim, Serra do Janela; 2019; J. Louzada and L. Vieira leg.; CEMT • 2 specs; Lavras; UFLA; 8 Nov. 1997; CEMT • 2 specs; Montes Claros; Jan. 1991; CEMT • 2 specs; Montes Claros; Mar. 1991; CEMT • 1 spec.; Nova Era, Mar.; 1987; CEMT • 2 specs; Viçosa; 11 Dec. 1984; Fiuza and Martins leg.; CEMT • 2 specs; Visconde do Rio Branco; Jan. 1998; W. Zikan leg.; CEMT. – Rio de Janeiro • 1 spec.; Nova Friburgo; Nov. 2002; J. Louzada; CEMT • 3 specs; Nova Friburgo, Macaé de Cima; Jan. 1999; P. Grossi leg.; CEMT • 1 spec.; Nova Friburgo, Conquista; Jan. 2004; E. Grossi leg.; CEMT • 1 spec.; Nova Friburgo; Jan. 2005; E. J. Grossi leg.; CEMT • 1 spec.; Pedro do Rio; 3 Dec. 1989; Bello leg.; CEMT • 1 spec.; Pedro do Rio; Nov. 1991; A. Bello leg.; CEMT. – Rio Grande do Sul • 2 specs; El Dorado do Sul, EEA-UFRGS; 16 Dec. 1996; Carvalho and Silva leg.; light trap; CEMT. – Rondônia • 1 spec.; Vilhena; 12 Nov. 1997; O. Roppa and M. Magno leg.; CEMT. – São Paulo • 2 specs; Descalvado, Fazenda Itaúnas; 6 Oct. 2006; N. W. Perioto leg.; forest + Citrus plantation; pitall; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD3FF8AFD95FB49FC66FB65.taxon	description	Sexual dimorphism Male with fifth abdominal ventrite shorter than that of female.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD3FF8AFD95FB49FC66FB65.taxon	discussion	Collecting information Specimens are attracted to light. Remarks This species is extremely similar to Lomanoxia costulata. In addition to the characters provided, L. melloi has a slightly thinner elytra when compared with L. costulata.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDDFF89FDEAFB60FA82FEF5.taxon	diagnosis	Diagnosis Length 4.0 – 5.5 mm. Body strongly convex, oval, piceous to reddish dark brown. Head densely punctate, frons with punctures closer and stronger; clypeus emarginate, margin, in frontal view, thickened at emargination. Pronotum with tubercles and / or lateral fossae variably developed and positioned; lateral margins always fringed with long setae. Elytral interstriae convex, tectiform.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDDFF89FDEAFB60FA82FEF5.taxon	discussion	Remarks Lomanoxoides has seven recognized species known from Argentina, Brazil, Costa Rica, Guiana, Honduras, Panama, Paraguay, Peru, and Venezuela (Stebnicka 2009). For a key to all its species see Stebnicka & Skelley (2005).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDEFF89FE60FEF1FDBBFBD9.taxon	description	Figs 19 B, 47 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDEFF89FE60FEF1FDBBFBD9.taxon	diagnosis	Diagnosis Length 5.0 – 5.5 mm. Body piceous. Head punctate; larger punctures restricted on frons; clypeus emarginate. Pronotum punctate; larger punctures restricted to posterior third; pronotal surface irregular, laterally tuberculate, with shallow posteromedial fossa; lateral margins explanate, with evident, but short, spatulate setae. Elytral interstriae convex, glabrous with row of fine punctures medially.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDEFF89FE60FEF1FDBBFBD9.taxon	materials_examined	Material examined BRAZIL – Rio Grande do Sul • 1 spec.; Bagé; 31 ° 21 ′ 21 ″ S, 54 ° 02 ′ 28 ″ W; 25 – 28 Dec. 2017; P. G. Silva leg.; light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDEFF89FE60FEF1FDBBFBD9.taxon	description	Sexual dimorphism Sexual dimorphism slight, males with penultimate abdominal ventrite more slender than that of female.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDEFF89FE60FEF1FDBBFBD9.taxon	discussion	Collecting information Specimens were collected at light.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDEFF88FDCDFB0DFDC2FD1C.taxon	description	Figs 19 A, 47 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDEFF88FDCDFB0DFDC2FD1C.taxon	diagnosis	Diagnosis Length 5.0 – 5.5 mm. Body piceous. Head punctate; punctures larger and closer on frons. Pronotum with punctures of two sizes, larger punctures scattered across surface, mixed with moderate punctures; surface uneven with two lateral tubercles and large deep posteromedial fossa; lateral margins explanate. Elytra setaceous with interstriae convex, tectiform.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDEFF88FDCDFB0DFDC2FD1C.taxon	materials_examined	Type material examined Paratype BRAZIL • 1 ♂; “ [yellow paper, printed] BR-MS-Selvíria / UNESP Farm / Ex Guzerá cattle drop- / ping Brachiaria de- / cumbens pasture / Rodrigues, S. R. col / [handwritten to the side] 11 - IV- 1993. // [red paper, printed] Type / Lomanoxoides / selviriaensis / Dt. Z. Stebnicka // [yellow paper] [handwritten] Lomanoxoides / selviriaensis / [typewritten] DET. Z. Stebninicka ”; CEMT. Other material examined BRAZIL – Mato Grosso • 1 spec.; Diamantino, Reserve Vale da Solidão; 14 ° 22 ′ S, 56 ° 07 ′ W; elev. 450 m; 16 Jan. 2013; CEMT • 1 spec.; Cuiabá; May 2010; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Cuiabá, Boa Esperança; Nov. 2011; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; 15 ° 36 ′ 31 ″ S, 56 ° 03 ′ 22 ″ W; Dec. 2008; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Cuiabá, Recanto do Siriema; 15 ° 35 ′ 32 ″ S, 56 ° 01 ′ 38 ″ W; 4 Nov. 2004; R. V. Nunes leg.; light; CEMT • 1 spec.; Cuiabá, UFMT; 24 Sep. 2016; J. Soares leg.; CEMT • 1 spec.; Porto Esperidão; Nov. 1984; Magno and Alvarenga leg.; CEMT. – Mato Grosso do Sul • 1 spec.; Selvíria, UNESP Farm; 11 Apr. 1993; S. R. Rodrigues leg.; Brachiaria decumbens pasture; Guzera bovine dropping; CEMT • 2 specs; same data as for preceding; 18 Mar. 1999; C. A. H. Flechtmann leg.; black light trap; CEMT • 1 spec.; same data as for preceding; 19 Oct. 1999; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDEFF88FDCDFB0DFDC2FD1C.taxon	description	Sexual dimorphism Sexual dimorphism slight, males with penultimate abdominal ventrite more slender than females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDEFF88FDCDFB0DFDC2FD1C.taxon	discussion	Collecting information Specimens were collected in Urochloa decumbens with UV light traps and extracted from Guzera bovine fecal masses (Stebnicka 1999 a).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDFFF8EFDF5FD4BFA9BFDD7.taxon	description	Figs 19 C, 47 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDFFF8EFDF5FD4BFA9BFDD7.taxon	diagnosis	Diagnosis Length 3.9 – 4.1 mm. Body reddish brown. Head punctate; punctures coarser and closer on frons; clypeus medially emarginate. Pronotum not explanate, fringe with evident setae; posterior margin with two medial tubercles. Elytral interstriae convex, each with row of small setaceous tubercles, each tubercle with a single seta.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDFFF8EFDF5FD4BFA9BFDD7.taxon	materials_examined	Material examined BRAZIL – Amazonas • 1 spec.; “ olo ” [unknown meaning] Km 133, 15 Mar. 1979; Young and Arias leg.; CEMT. – Bahia • 1 spec.; Itamarajú; 16 ° 57 ′ 58 ″ S, 39 ° 26 ′ 01 ″ W; Apr. 2011; C. M. P. Leite leg.; CEMT • 1 spec.; Wenceslau Guimarães, Matriz; 13 ° 41 ′ 21 ″ S, 39 ° 21 ′ 52 ″ W; elev. 146 m; C. M. P. Leite leg.; pitfall; CEMT • 1 spec.; same data as for preceding; Jan. 2011; Q 6 matriz; CEMT. – Distrito Federal • 1 spec.; Brasilia; Nov. 2002; N. Degallier leg.; CEMT • 1 spec.; same data as for preceding; elev. 1000 m; Mar. 2001; N. Degallier leg.; light; CEMT • 4 specs; Planaltina, Embrapa Cerrados; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 2 Dec. 2005; C. Oliveira leg.; área agrícola; light; CEMT. – Mato Grosso • 1 spec.; Itanhangá; 18 – 22 Oct. 2016; J. V. Korgut leg.; manually; CEMT • 1 spec.; Nova Xavantina; 14 ° 62 ′ 35 ″ S, 52 ° 48 ′ 57 ″ W; 23 Oct. 2009; V. Campos leg.; Cerrado; CEMT • 1 spec.; Porto Estrela, Est. Eco Serra das Araras, Boca do José; Nov. 2013; R. V. Nunes leg.; FIT; CEMT. – Mato Grosso do Sul • 1 spec.; Guia Lopes da Laguna; 12 Dec. 2007; A. R. Abot leg.; light trap; CEMT. – Minas Gerais • 2 specs; Águas Vermelhas; Dec. 1998; A. Bello and F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Montes claros; Jan. 1991; CEMT • 1 spec.; same data as for preceding; Dec. 1992; CEMT • 1 spec.; Nova era; Jul. 1987; CEMT • 1 spec.; Santa Barbara; 8 Dec. 1994; J. C. Zanuncio leg.; CEMT • 1 spec.; Teixeiras; Nov. 1996; Hardy and Harrison leg.; CEMT • 1 spec.; Viçosa, Campus UFV; Nov. 1997; F. Z. Vaz-de-Mello leg.; CEMT. – Paraíba • 1 spec.; Areia, Mata de Pau Ferro; 3 – 4 May 2000; M. Avany leg.; light trap; CEMT. – Paraná • 1 spec.; Icaraíma, Estância Felipe; 23 ° 20 ′ 32 ″ S, 53 ° 39 ′ 09 ″ W; elev. 300 m; 2019; Felipe and Rodrigues leg.; CEMT. – Pernambuco • 3 specs; Igarassu, RECD; 25 May 2007; Cavalcanti et al. leg.; CEMT • 4 specs; same data as for preceding; 27 Apr. 2007; Costa et al. leg.; CEMT. FRENCH GUIANA • 1 spec.; Cayenne, Montage des Chevaux; 4 ° 44 ′ 56 ″ N, 52 ° 26 ′ 28 ″ W; elev. 75 m; 27 Jan. 2013; SEAG leg.; CEMT. UNKNOWN COUNTRY • 1 spec.; [without data]; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDFFF8EFDF5FD4BFA9BFDD7.taxon	description	Sexual dimorphism Slight, males with penultimate abdominal ventrite more slender than that of females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDFFF8EFDF5FD4BFA9BFDD7.taxon	discussion	Collecting information Specimens were collected with light traps and were found at altitudes ranging from 75 to 1000 metres.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD9FF8EFDEFFD14FC3CF8B3.taxon	description	Figs 19 D – E, 48 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD9FF8EFDEFFD14FC3CF8B3.taxon	diagnosis	Diagnosis Length 4.8 – 5.5 mm. Body reddish brown. Head punctate; punctures coarser and closer on frons. Pronotum not explanate, somewhat regularly convex, except by small fovea and tubercle laterally. Elytra margin posteriorly and laterally setaceous; interstriae convex, tectiform with two rows of fine punctures.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD9FF8EFDEFFD14FC3CF8B3.taxon	materials_examined	Material examined BRAZIL – Distrito Federal • 1 spec.; Planaltina, Embrapa Cerrados; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 2 Dec. 2005; C. Oliveira leg.; agricultural area; light; CEMT. – Mato Grosso do Sul • 1 spec.; Bonito; Fazenda Remanso; 20 ° 47 ′ S; 56 ° 43 ′ W; Nov. 2009; F. O. Roque leg.; CEMT. – Mato Grosso • 1 spec.; Cuiabá, Boa Esperança; 15 ° 36 ′ 31 ″ S, 56 ° 3 ′ 22 ″ W; Dec. 2008; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Diamantino, Alto Rio Arinos; Jan. 2001; E. Furtado leg.; CEMT. – Minas Gerais • 2 specs; Águas Vermelhas; Dec. 1997; A. Bello and F. Z. Vaz-de-Mello leg.; CEMT • 2 specs. Cordisburgo; Jan. 1994; F. Z. Vaz-Mello leg.; CEMT • 2 specs; same data as for preceding; Jan. 1998; CEMT • 1 spec.; Cordisburgo, Fazenda Pontinha; Jan. 1998; F. Z. Vaz-de-Mello leg.; CEMT • 2 specs; same data as for preceding; Jan. 1999; F. Z. Vaz-de-Mello leg.; CEMT • 2 specs; Ijaci, Fazenda FAEPE; Nov. 2002; J. Louzada leg.; light; CEMT • 1 spec.; Três Marias; Nov. 1991; CEMT. – Rio de Janeiro • 1 spec.; Angra dos Reis, Bracuí, Baixada; 10 – 11 May 2002; Nessimian leg.; UV light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD9FF8EFDEFFD14FC3CF8B3.taxon	description	Sexual dimorphism Slight, males with penultimate abdominal ventrite slender than that of females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD9FF8EFDEFFD14FC3CF8B3.taxon	discussion	Collecting information Specimens were collected with UV light traps, some specimens were found within nests of Atta sexdens and in bovine feces (Stebnicka 1999 a; Stebnicka & Skelley 2005).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD9FF8CFE38F8BBFA82FBDB.taxon	diagnosis	Diagnosis Length 5.0 – 6.5 mm. Body elongate, convex, dark brown or piceous. Head with large, gibbous medially; clypeus medially sinuate. Pronotum transverse, with punctures of different sizes, lateral margins fringed by large and evident setae; lateral margins of pronotum widely explanate, except in Martineziana simplex; lateral and posterior margins furrowed, posterior furrow may be strigate. Elytra glossy.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFD9FF8CFE38F8BBFA82FBDB.taxon	discussion	Remarks Martineziana has six recognized species distributed across southern United States, Central and South America and Greater Antilles (Stebnicka 2009). A key to all its species can be found in Chalumeau (1983 a).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDBFF8CFDF3FB0DFA83F873.taxon	description	Figs 15 E – F, 48 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDBFF8CFDF3FB0DFA83F873.taxon	diagnosis	Diagnosis Length 6.5 mm. Body piceous. Head with longitudinal wrinkles, strongly gibbous; clypeus emarginate. Pronotum with punctures of two sizes; minute extremely fine and inconspicuous punctures scattered; larger punctures moderate in size scattered across disc and lateral margins; lateral margins not deeply excavate. Elytra posteriorly widened.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDBFF8CFDF3FB0DFA83F873.taxon	materials_examined	Material examined BRAZIL – Minas Gerais • 6 specs; Ijaci, Fazenda FAEPE; Nov. 2002; J. Louzada leg.; light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDBFF8CFDF3FB0DFA83F873.taxon	description	Sexual dimorphism Males with spur of protibia inwardly hooked; spur of females somewhat straight.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFFDBFF8CFDF3FB0DFA83F873.taxon	discussion	Collecting information Specimens were collected at light and in nest of Solenopsis saevissima (Smith, 1855) (Stebnicka 2009).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF24FF72FDC7FE94FE24FE38.taxon	description	Figs 16 B – C, 48 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF24FF72FDC7FE94FE24FE38.taxon	diagnosis	Diagnosis Length 5.6 mm. Body thin, dark piceous. Head with longitudinal wrinkles, strongly gibbous medially. Pronotum with punctures of two sizes; fine punctures dense, mixed with larger ones; larger punctures spread across pronotum, smaller anteriorly, larger near posterior and lateral margins. Elytra long, not greatly widened posteriorly.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF24FF72FDC7FE94FE24FE38.taxon	materials_examined	Material examined ARGENTINA – Corrientes • 1 spec.; Corrientes, Barrio Víctor Colas; 27 ° 28 ′ 50 ″ S 58 ° 47 ′ 18 ″ W; elev. 64 m; 21 Oct. 2012; Polesel et al. leg.; CEMT. – Jujuy • 1 spec.; Parque Nacional Calilegua, 11 km NW of campground; elev. 1100 m; 14 Jan. 2008; D. M. Takiya leg.; light trap 1; CEMT. BRAZIL – Acre • 1 spec.; Assis Brasil; 10 ° 35 ′ 44 ″ S, 69 ° 37 ′ 11 ″ W; 25 Sep. 2016; Ortiz and Borges leg.; wild environment; CDC light trap; CEMT • 1 spec.; Rio Branco; CEMT • 1 spec.; Rio Branco; 5 Jun. 1995; S. M. F. Almeira leg.; CEMT • 1 spec.; same data as for preceding; Fazenda Catuaba; 2000; CEMT. – Amazonas • 1 spec.; Sist. Janauacá, Rio Solimoes; 9 Apr. 1977; Geraldo leg.; CEMT. – Mato Grosso • 1 spec.; Cuiabá; Nov. 2009; F. Z. Vaz-de-Mello leg.; CEMT • 2 specs. Diamantino, Alto Rio Arinos; Jan. 2001; E. Furtado leg.; CEMT. – Rio Grande do Sul • 3 specs; Bagé; 31 ° 21 ′ 05 ″ S, 54 ° 01 ′ 13 ″ W; elev. 232 m; 2 Feb. 2017; R. N. Sisti et al. leg.; forest; CEMT • 7 specs; Eldorado do Sul, EEA-UFRGS; 11 Oct. 1996; Carvalho and Silva leg.; arm. lum; CEMT. PARAGUAY – Central • 1 spec.; Asunción; 20 Apr. 2008; C. Aguiar leg.; CEMT. UNITED STATES OF AMERICA – Alabama • 2 specs; Perry Co, 6.5 mi SW of Marion; 30 Jun. 1987; S. C. Harris leg.; CEMT. – Mississippi • 1 spec.; Forrest Co., Hattiesburg, Desoto NF; 25 Jun. 1991; B. Buchli leg.; blacklight trap; CEMT • 1 spec.; Lee Co., 4.8 mi SE of Saltillo; 7 May 1995; J. Dakin leg.; CEMT. – Virginia • 1 spec.; Chesapeake, Northwest River State Park; 27 Jul. 1997; M. Hardy leg.; CEMT • 1 spec.; Chesterfield Co., Pocahontas State Park; 27 Jun. 1992; P. Bélanger leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF24FF72FDC7FE94FE24FE38.taxon	description	Sexual dimorphism Males with spur of protibia inwardly hooked; spur of females straight.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF24FF72FDC7FE94FE24FE38.taxon	discussion	Collecting information Specimens were collected with light traps. Martineziana dutertrei were reported to be found in galleries of several species of ants of the genus Solenopsis Westwood, 1840 (Wojcik et al. 1977). Remarks Martineziana dutertrei was found in nests of Solenopsis invicta Burren, 1972, S. richteri Forel, 1913, S. geminata, S. xyloni McCook, 1879 and Linepithema humile (Mayr, 1868) (referred to in the consulted literature as Iridomyrmex humilis (Mayr, 1868 )) (Wojcik et al. 1977). Of the mounds opened by Collins & Making (1971) 98 % contained 12 to 20 individuals of Martineziana dutertrei (Stebnicka 2007 a). The entire life cycle of this species possibly happens inside the nests of these ants. Wojcik et al. (1977) found several pupae and larvae in nests of S. invicta. The authors believed the species observed was Martineziana excavaticollis. Later Stebnicka (2007 a) confirmed the species found was actually Martineziana dutertrei.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF25FF71FDD9FE2FFD2BFEE3.taxon	description	Figs 16 D – F, 49 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF25FF71FDD9FE2FFD2BFEE3.taxon	diagnosis	Diagnosis Length 6.0 mm. Body broad, dark brown. Head with transverse wrinkles, medially gibbous. Pronotum transverse, punctate with punctures of two sizes; fine punctures inconspicuous among the extremely large punctures. Elytra short, broad, interstriae microreticulate with inconspicuous punctures.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF25FF71FDD9FE2FFD2BFEE3.taxon	materials_examined	Material examined ARGENTINA – San Luis • 1 spec.; Desaguadero; Feb. 2000; G. Arriágada leg.; CEMT • 1 spec.; Belgrano, Fundo El Molle; 33 ° 02 ′ 22 ″ S, 66 ° 30 ′ 47 ″ W; elev. 622 m; 15 – 16 Dec. 2015; G. Arriágada leg.; collected with light; CEMT. BRAZIL – Acre • 1 spec.; Rio Branco, Faz Catuaba; 2000; CEMT. – Distrito Federal • 1 spec.; Brasilia, Fazenda Água Limpa; Nov. 2010; C. Suinagua leg.; CEMT. – Mato Grosso • 1 spec.; Sesc Pantanal; 16 Nov. 2001; light trap; CEMT • 2 specs; Alta Floresta, Chácara recanto das Orquíedeas; 7 Aug. 2012; A. Petini-Beneli leg.; CEMT • 1 spec.; Cotriguaçú, Fazenda São Nicolau; 10 Dec. 2009; M. R. Barreto leg.; forest; blacklight trap; CEMT • 1 spec.; same data as for preceding; 30 Oct. 2014; W. Chamorro leg.; blacklight; CEMT • 1 spec.; same data as for preceding; 9 ° 49 ′ 08 ″ S, 58 ° 15 ′ 40 ″ W; 19 – 24 Oct. 2012; F. Z. Vaz-de-Mello leg.; PPBioP 1; CEMT • 1 spec.; same data as for preceding; 9 ° 49 ′ 09 ″ S, 58 ° 15 ′ 31 ″ W; 28 – 30 Oct. 2017; F. Z. Vaz de Mello et al. leg.; CEMT • 1 spec.; same data as for preceding; 9 ° 49 ′ 09 ″ S, 58 ° 15 ′ 31 ″ W; 21 Oct. 2018; V. Costa-Silva et al. leg.; CEMT • 1 spec.; same data as for preceding; 9 ° 51 ′ 20 ″ S, 58 ° 14 ′ 55 ″ W; Nov. 2014; R. V. Nunes and A. Asenjo leg.; light; CEMT • 2 specs; same data as for preceding; airport; 9 ° 51 ′ 52 ″ S, 58 ° 13 ′ 45 ″ W; 2 Oct. 2017; F. Z. Vaz-de-Mello leg.; light; CEMT • 4 specs; Cuiabá; 15 ° 35 ′ 32 ″ S, 56 ° 01 ′ 39 ″ W; elev. 80 m; Jul. 2014 – Jan. 2015; R. V. Nunes leg.; CEMT • 1 spec.; Cuiabá, Boa Esperança; Oct. 2016; F. Z. Vaz-de-Mello leg.; CEMT • 2 specs; same data as for preceding; 15 ° 36 ′ 31 ″ S, 56 ° 03 ′ 22 ″ W; Dec. 2008; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; UFMT; 20 Apr. 2009; K. F. Santos leg.; coleta ativa; CEMT • 1 spec.; Nov. 2009; R. Miyazaki leg.; CEMT • 1 spec.; Duque de Caxias; 3 Jan. 2011; F. Simioni leg.; manually; CEMT • 1 spec.; Duque de Caxias, Recanto dos Pássaros; 5 May 2009; K. A. Azevedo leg.; coleta manually; CEMT • 1 spec.; Duque de Caxias, Res. Coxipó; Oct. 2012; F. M. Lima leg.; CEMT • 1 spec.; Poconé; 21 Oct. 1953; C. R. Gonçalves leg.; CEMT • 1 spec.; Chapada dos Guimarães; 25 May 2001; A. F. Ramos leg.; light trap; CEMT • 2 specs; same data as for preceding; 22 May 2001; A. F. Ramos leg.; light trap; CEMT • 6 specs; Várzea Grande; 15 ° 40 ′ S, 56 ° 10 ′ W; Nov. 2021 – Feb. 2022; J. Arias leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF25FF71FDD9FE2FFD2BFEE3.taxon	description	Sexual dimorphism Males with spur of protibia inwardly hooked; spur of females straight.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF25FF71FDD9FE2FFD2BFEE3.taxon	discussion	Collecting information Specimens were collected with UV light traps.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF26FF71FDF2FEEBFECDFB4E.taxon	description	Figs 16 A, 48 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF26FF71FDF2FEEBFECDFB4E.taxon	diagnosis	Diagnosis Length 5.0 mm. Body piceous. Head and pronotum covered with fine setae. Head covered with small setaceous tubercles. Pronotum with punctures of two sizes; moderate, setaceous punctures mixed with wide coarse areolate punctures. Elytra with striae deep; interstriae convex, with rows of punctures on each side.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF26FF71FDF2FEEBFECDFB4E.taxon	materials_examined	Material examined BRAZIL – Mato Grosso • 1 spec.; Varzea Grande, 2 km W of Parque de Exposição; 22 Mar. 1972; W. H. Whitecomb leg.; CEMT • 2 specs; Varzea Grande, Sítio Toca da Coruja; 15 ° 33 ′ 07 ″ S, 56 ° 19 ′ 07 ″ W; elev. 230 m; 18 Oct. 2020; A. Petini-Beneli leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF26FF71FDF2FEEBFECDFB4E.taxon	description	Sexual dimorphism Unknown. Based on the other species of the genus, males probably have the spur of the protibia inwardly hooked.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF26FF71FDF2FEEBFECDFB4E.taxon	discussion	Collecting information Unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF26FF70FDF1FAB9FBAEFBC3.taxon	description	Figs 17 A, 49 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF26FF70FDF1FAB9FBAEFBC3.taxon	diagnosis	Diagnosis Length 4.30 – 4.60 mm. Body dark reddish brown. Head with weak or indistinct transverse wrinkles. Pronotum with punctures of two sizes; fine punctures everywhere densely distributed; coarser punctures scarce near anterior margin, abundant near lateral and posterior margins; lateral margins not explanate; lateral and posterior marginal furrow present. Elytra wider posteriorly.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF26FF70FDF1FAB9FBAEFBC3.taxon	materials_examined	Material examined BRAZIL – Amazonas • Amazonas • 1 spec.; Itacoatiara, Fazenda Aruana, Am 010, Km 215; 20 – 23 Aug. 1990; C. S. Motta, F. J. A. Peralta and B. Rochin-Teles leg.; UV light; CEMT • 3 specs; same data as for preceding; INPA • 1 spec.; same data as for preceding; C. S. Motta et al. leg.; CEMT • 3 specs; same data as for preceding; 14 – 17 May 1991; C. S. Motta et al. leg. CEMT • 5 specs; same data as for preceding; INPA • 1 spec.; same data as for preceding; 18 – 21 Sep. 1990; C. S. Motta, R. L. M. Ferreira and R. Andreazze leg.; CEMT • 4 specs; same data as for preceding; INPA • 3 specs; same data as for preceding; 13 – 16 Feb. 1991; C. S. Motta, R. Andreazze and V. Kawazoe leg.; CEMT • 13 specs; same data as for preceding; INPA • 1 spec.; Manaus, Reserva Ducke; 2 ° 55 ′ 51 ″ S, 59 ° 58 ′ 59 ″ W; Mar. 2004; L. Montel leg.; light; INPA. – Distrito Federal • 1 spec.; Brasilia; Dec. 2003; N. Degallier leg.; CEMT. – Goiás • 1 spec.; P. N. Emas; Nov. 1999; F. Rodrigues leg.; CEMT. – Mato Grosso • 1 spec.; Res. Papagaio, BR- 364, Km 555; 2 Oct. 1984; J. Elias leg.; light trap; CEMT • 1 spec.; Sesc Pantanal; 15 Apr. 2002; light trap; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; Oct. 2014; R. Stofel leg.; CEMT • 1 spec.; same data as for preceding; airport; 9 ° 51 ′ 52 ″ S, 58 ° 13 ′ 45 ″ W; 2 Oct. 2017; F. Z. Vaz-de-Mello et al. leg.; light trap; CEMT • 1 spec.; Cuiabá; 15 ° 35 ′ 32 ″ S, 56 ° 01 ′ 29 ″ W; elev. 80 m; Jul. 2014 – Jan. 2015; R. V. Nunes leg.; CEMT • 1 spec.; Trindade U. B. S., Nortwestern pole; 1 Oct. 1984; E. Binda leg.; CEMT. – Mato Grosso do Sul • 7 specs; MS- 080, Km 48, NW of Campo Grande; 6 Jun. 1986; J. Senatore leg.; CEMT • 1 spec.; Guia Lopes da Laguna; 22 Feb. 2008; A. Abot leg.; light trap; CEMT • 1 spec.; Selvíria, UNESP farm; 17 Jan. 1991; C. A. H. Flechtmann leg.; Brachiaria decumens pasture; black light; CEMT. – Piauí • 4 specs; Teresina, CPAMN; 5 ° 02 ′ 16 ″ S, 42 ° 47 ′ 13 ″ W; elev. 61 m; 10 May 2011; R. B. Querino leg.; light trap; CEMT. FRENCH GUIANA • 1 spec.; Cayenne, Montagne des Chevaux; 4 ° 44 ′ 56 ″ N, 52 ° 26 ′ 28 ″ W; elev. 75 m; 4 Feb. 2013; SEAG leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF26FF70FDF1FAB9FBAEFBC3.taxon	description	Sexual dimorphism Males with spur of protibia inwardly hooked; spur of females straight.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF26FF70FDF1FAB9FBAEFBC3.taxon	discussion	Collecting information Specimens were collected with UV light traps in Urochloa decumbens pasture.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF27FF77FDF5FB0BFA84FB04.taxon	diagnosis	Diagnosis Length 3.0 – 8.0 mm. Body oval or elongate, dark reddish brown to brown piceous. Head broad, almost as wide as lateral margins of pronotum, variably punctate. Pronotum with variably shaped and distributed punctures; lateral margins, explanate or not, transverse or not. Elytra with humeral denticle developed. Mesoventrite with one or two callosities, or if callosity not present, mesometaventral carina present. Mesoventrite and metatarsus with dense setae on its ventral surface or not.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF27FF77FDF5FB0BFA84FB04.taxon	discussion	Remarks Odontolytes has 19 recognized species reported from the USA (Florida), Cuba, and several countries from Central and South America (Stebnicka 2009). Stebnicka (2002) presents a key to all species described at the time. Odontolytes waoraniae Stebnicka & Skelley, 2005 and O. tectipennis Stebnicka & Skelley, 2005 were described after Stebnicka (2002) and are absent from the key. The current taxonomic status of Odontolytes was reviewed by Krell et al. (2011).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF20FF76FE2BFB40FE7BFE57.taxon	description	Figs 23 A, 49 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF20FF76FE2BFB40FE7BFE57.taxon	diagnosis	Diagnosis Length 3.0 – 3.5 mm. Body piceous. Head wide, everywhere punctate; clypeus subtruncate with an anterior glossy region with fine small tubercles. Pronotum transverse; sides weakly explanate. Scutellum with longitudinal carina. Elytra with interstriae convex, punctate. Meso-metaventral carina present; mesoventrite without callosity.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF20FF76FE2BFB40FE7BFE57.taxon	materials_examined	Material examined BRAZIL – Amazonas • 2 specs; Manaus; 2 ° 25 ′ S, 59 ° 50 ′ W; Feb. 1994; R. Didham leg.; Winkler extractor; CEMT • 9 specs; same data as for preceding; INPA • 3 specs; same data as for preceding; May 1994; CEMT • 4 specs; same data as for preceding, INPA.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF20FF76FE2BFB40FE7BFE57.taxon	description	Sexual dimorphism Unknown	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF20FF76FE2BFB40FE7BFE57.taxon	discussion	Collecting information Specimens were found in forest litter and collected using with Winkler extractor and black light traps (Stebnicka 2009).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF21FF76FDE7FD96FB4DF8E8.taxon	description	Figs 21 A, 49 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF21FF76FDE7FD96FB4DF8E8.taxon	diagnosis	Diagnosis Length 6.0 – 8.0 mm. Body oval, piceous. Head densely punctate with fine punctures; clypeus emarginate. Pronotum with punctures moderate, separated by their diameter; sides not explanate, only anterior angles; posterior angles sinuate. Metafemur with incomplete posterior line. Tarsus without dense setae on its ventral surface. Mesoventrite with double callosity.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF21FF76FDE7FD96FB4DF8E8.taxon	materials_examined	Material examined BRAZIL – Acre • 1 spec.; Rio Branco; 14 Jun. 1996; CEMT • 1 spec.; same data as for preceding; 15 Apr. 1987; Fazolin and Murilo leg.; UV; CEMT. – Espírito Santo • 1 spec.; Linhares, Flona de Goytacazes; 19 ° 26 ′ 10 ″ S, 40 ° 04 ′ 24 ″ W; 10 Nov. 2010; D. S. Martins leg.; primary forest; light; CEMT. – Mato Grosso • 1 spec.; Rio Branco; Dec. 1984; Magno and Alvarenga leg.; CEMT. ECUADOR – Cotopaxi • 1 spec.; Los Libres; elev. 1900 m; 4 Nov. 1994; N. Marchan leg.; CEMT. – Orellana • 1 spec.; Estación de Diversidad Tiputini; elev. 250 m; Sep. 1997; P. Araujo leg.; trampa de luz [light trap]; CEMT. FRENCH GUIANA • 1 spec.; Cayenne, Montagne des Chevaux; 4 ° 44 ′ 56 ″ N, 52 ° 26 ′ 28 ″ W; elev. 75 m; 27 Jan. 2013; SEAG leg.; light (GEML); CEMT. MEXICO – Oaxaca • 2 specs; Acatlán, Laguna Verde; 2 May 2004; J. V. Bueno leg.; selva baja [low land frorest] leaf litter; CEMT. PERU – Loreto • 1 spec.; 21 km W of Iquitos; 10 Jul. 2001; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF21FF76FDE7FD96FB4DF8E8.taxon	description	Sexual dimorphism Males with fifth abdominal ventrite shorter than that of females, the disc of the pygidium is longer and the metatarsals are thicker.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF21FF76FDE7FD96FB4DF8E8.taxon	discussion	Collecting information Specimens were collected at light at high altitudes from 250 to 1200 metres and on basal debris of Bactris Jacq. ex Scop. palm (Stebnicka 2002). Some specimens were also found in forest litter.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF23FF74FDC8FE95FD4DFA9C.taxon	description	Fig. 49 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF23FF74FDC8FE95FD4DFA9C.taxon	diagnosis	Diagnosis Length 6.4 – 7.0 mm. Body elongate, piceous. Head punctate; punctures elongate near frons; clypeus emarginate. Pronotum with double (fine and coarse) puncturation; lateral margins weakly explanate. Elytra convex, humeral denticle sharp; interstriae convex, visibly punctate; 10 th interstria flat with row of minute granules. Mesoventrite with fine mesometasternal carina. Sexual dimorphism Males with pronotal punctures more widely spaced and posterior marginal line deeper and wider than those of females; fifth abdominal ventrite shorter than that of female, disc of pygidium is longer and the basal tarsomere of metatarsus are thicker. Collecting information Specimens were collected with black lights.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF23FF74FDC8FE95FD4DFA9C.taxon	discussion	Remarks Specimens were unavailable for the present study, the information here presented was taken from the available literature (Stebnicka 2002, 2009).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF23FF7BFE3EFACBFC50FD94.taxon	description	Figs 22 F, 49 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF23FF7BFE3EFACBFC50FD94.taxon	diagnosis	Diagnosis Length 4.2 – 5.0 mm. Body oblong, reddish browns. Head coarsely punctate; clypeus sinuate; laterally with faint wrinkles. Pronotum with punctures fine and coarse; fine punctures mixed with coarse punctures; coarse punctures wider and confluent closer to lateral margins. Elytra with interstriae convex, crenulate by coarse strial punctures. Mesometaventral carina present. Tarsus with dense setae on its ventral surface.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF23FF7BFE3EFACBFC50FD94.taxon	materials_examined	Material examined BRAZIL – Acre • 1 spec.; Mâncio Lima, Parna Serra do Divisor; 7 ° 26 ′ 46 ″ S, 73 ° 39 ′ 28 ″ W; 11 – 15 Dec. 2019; FIT; CEMT. – Amazonas • 1 spec.; Itacoatiara, Fazenda Itapiranga; 2 ° 42 ′ 14 ″ S, 58 ° 31 ′ 49 ″ W; 29 Jan. – 6 Feb. 2015; R. S. Moura leg.; pitfall 3; CEMT. FRENCH GUIANA • 1 spec.; Cayenne, Montagne des Chevaux; 4 ° 44 ′ 56 ″ N, 52 ° 26 ′ 28 ″ W; elev. 75 m; 11 Feb. 2013; SEAG leg.; FIT (V); CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF23FF7BFE3EFACBFC50FD94.taxon	description	Sexual dimorphism Males with fifth abdominal ventrite shorter than that of females and longer disc of pygidium.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF23FF7BFE3EFACBFC50FD94.taxon	discussion	Collecting information Specimens were collected with pitfall and flight intercept traps.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2CFF7BFDEDFDD0FEFDF9E2.taxon	description	Figs 21 E – F, 50 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2CFF7BFDEDFDD0FEFDF9E2.taxon	diagnosis	Diagnosis Length 4.8 – 5.0 mm. Body piceous. Head with dense and coarse punctures; punctures elongate anterior to frontoclypeal suture. Pronotal surface everywhere densely and coarsely punctured; punctures larger and closer near lateral margins; lateral angles greatly explanate. Scutellum with two foveae. Elytra with deep and wide striae; interstriae convex and shagreened. Proventral process wide and scabrous. Mesoventrite with wide callosity. Ventral surface of meso- and metatarsi with dense setae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2CFF7BFDEDFDD0FEFDF9E2.taxon	materials_examined	Material examined BRAZIL – Acre • 1 spec.; Assis Brasil; 10 ° 35 ′ 44 ″ S, 69 ° 37 ′ 12 ″ W; 20 Sep. 2016; Ortiz and Borges leg.; CEMT. – Amazonas • 4 specs; Novo Airão; 2 ° 38 ′ 39 ″ S, 60 ° 56 ′ 07 ″ W; 27 – 28 Jul. 2011; F. Xavier and A. Agudelo leg.; CEMT • 1 spec.; same data as for preceding; 27 Mar. 2011; J. A. Rafael, D. Takiya and J. T. Câmara leg.; mobile light trap; INPA • 1 spec.; Rio Tarumã Mirim, NW of Manaus; 5 Dec. 1975; J. Adis leg.; INPA • 1 spec.; same data as for preceding; 27 Apr. 1977; J. Adis leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2CFF7BFDEDFDD0FEFDF9E2.taxon	description	Sexual dimorphism Males with fifth abdominal ventrite shorter than that of females and longer disc of pygidium.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2CFF7BFDEDFDD0FEFDF9E2.taxon	discussion	Collecting information Specimens were collected with flight intercept traps and UV light traps in meaodw regions (Stebnicka 2009).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2CFF7AFDF0F9EAFD82FC5F.taxon	description	Figs 21 B – D, 50 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2CFF7AFDF0F9EAFD82FC5F.taxon	diagnosis	Diagnosis Length 4.0 – 5.0 mm. Body oval, robust piceous. Head densely punctate with fine punctures; clypeus sinuate. Pronotum with moderate punctures, separated by their diameter; lateral margins not explanate, anterior angles explanate, posterior angles sinuate. Mesoventrite with double callosity. Metafemur with complete marginal line.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2CFF7AFDF0F9EAFD82FC5F.taxon	materials_examined	Material examined BOLIVIA – Santa Cruz • 1 spec.; 3.7 km SSE of Buena Vista, Hotel Flora & Fauna; elev. 430 m; 5 – 15 Nov. 2001; M. C. Thomas and B. K. Dozier leg.; tropical transition forest; black light trap; CEMT. BRAZIL – Acre • 2 specs; Assis Brasil; 10 ° 35 ′ 44 ″ S, 69 ° 37 ′ 12 ″ W; 20 Sep. 2016; Ortiz and Borges leg.; CEMT. – Rondônia • 1 spec.; Jaci-paraná, Rio Madeira; 4 Oct. 2004; margem direita interior [interior right margin]; light trap 34 b; CEMT. FRENCH GUIANA • 2 specs; Cayenne, Montagne des Chevaux; 4 ° 44 ′ 56 ″ N, 52 ° 26 ′ 28 ″ W; elev. 75 m; 11 Feb. 2013; SEAG leg.; FIT (V); CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2CFF7AFDF0F9EAFD82FC5F.taxon	description	Sexual dimorphism Males with fifth abdominal ventrite shorter than that of females; disc of pygidium is longer and metatarsals are thicker.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2CFF7AFDF0F9EAFD82FC5F.taxon	discussion	Collecting information Specimens were collected with flight intercept and with light traps. A specimen was found in forest litter near Ficus L. fruits (Stebnicka 2002).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2DFF7AFE0EFB8EFC6EF836.taxon	description	Figs 22 D, 49 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2DFF7AFE0EFB8EFC6EF836.taxon	diagnosis	Diagnosis Length 3.8 – 4.0 mm. Body reddish brown. Head coarsely punctate; punctures somewhat oval near frontoclypeal suture; clypeus sinuate. Pronotum punctate with punctures of two sizes, fine punctures inconspicuous among the coarse punctures; coarse punctures larger and confluent laterally; laterally not explanate. Scutellum foveate. Proventral process wide and scabrous. Mesoventrite with wide oval callosity.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2DFF7AFE0EFB8EFC6EF836.taxon	materials_examined	Material examined BRAZIL – Pará • 2 specs; Almeirim; 5 Mar. 2011; F. M. França leg.; CEMT. FRENCH GUIANA • 1 spec.; Cayenne, Montagne des Chevaux; 4 ° 44 ′ 56 ″ N, 52 ° 26 ′ 28 ″ W; elev. 75 m; 31 Mar. 2013; SEAG leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2DFF7AFE0EFB8EFC6EF836.taxon	description	Sexual dimorphism Males with fifth abdominal ventrite shorter than that of female and longer disc of pygidium.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2DFF7AFE0EFB8EFC6EF836.taxon	discussion	Collecting information No information available in literature or with material studied.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2EFF79FDEAFE95FC54FABF.taxon	description	Figs 22 C, 50 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2EFF79FDEAFE95FC54FABF.taxon	diagnosis	Diagnosis Length 3.8 – 4.0 mm. Body elongate, reddish brown. Head punctate; punctures elongate anteriorly to frontoclypeal suture; clypeus sinuate. Pronotum coarsely punctate, punctures larger and confluent near lateral margins; anterior angles explanate. Scutellum foveate. Proventral process glossy acuminate. Mesoventrite with large callosity.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2EFF79FDEAFE95FC54FABF.taxon	materials_examined	Material examined BRAZIL – Mato Grosso do Sul • 8 specs; Selvíria; 30 Apr. – 1 Dec. 2011; Caterino and Tischechkin leg.; Cerradão; FIT; CEMT. – Mato Grosso • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 8 – 14 Apr. 2011; Peres-Filho leg.; light trap; CEMT. – Rondônia • 1 spec.; 62 km SW of Ariquemes, nr Fazenda Rancho Grande; 28 Oct. 1996; K. M. V. Vulinec leg.; CEMT. – São Paulo • 1 spec.; Descalvado, Fazenda Itaunas; 22 Sep. 2005; N. W. Perioto leg.; pitfall; CEMT • 1 spec.; same data as for preceding; 6 Oct. 2005; N. W. Perioto leg.; forest + Citrus plantation; pitfall; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2EFF79FDEAFE95FC54FABF.taxon	description	Sexual dimorphism Males with fifth abdominal ventrite shorter than that of female and longer disc of pygidium.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2EFF79FDEAFE95FC54FABF.taxon	discussion	Collecting information Specimens were collected in leaf litter in the Amazon rainforest, in cerradão forest with flight intercept trap. Specimens were also collected with light and pitfall traps.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2EFF7FFDF8FAAFFC18FE6C.taxon	description	Figs 23 B, 49 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2EFF7FFDF8FAAFFC18FE6C.taxon	diagnosis	Diagnosis Length 6.5 – 7.0 mm. Body dark reddish brown. Head coarsely punctate; clypeus sinuate. Pronotum with punctures of two sizes; fine punctures mixed with coarser larger ones; large punctures larger and closer near lateral margins; laterally not explanate. Scutellum not foveolate. Mesometaventral carina present. Meso- and metatarsomeres with dense setae on ventral surface.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2EFF7FFDF8FAAFFC18FE6C.taxon	materials_examined	Material examined BRAZIL – Amazonas • 1 spec.; Manaus, Alaixo; 3 ° 04 ′ 40 ″ S, 59 ° 59 ′ 13 ″ W; 10 Oct. 2017; F. S. Godoi leg.; light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2EFF7FFDF8FAAFFC18FE6C.taxon	description	Sexual dimorphism Males with the disc of pronotum less densely punctate; posterior margin or pronotum with deeper and wider emargination; fifth abdominal ventrite shorter than that of females and longer disc of pygidium.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2EFF7FFDF8FAAFFC18FE6C.taxon	discussion	Collecting information Specimens were collected with black light trap in tropical rainforests.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF28FF7FFDF6FD99FC51F8AF.taxon	description	Figs 22 A – B, 50 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF28FF7FFDF6FD99FC51F8AF.taxon	diagnosis	Diagnosis Length 4.8 – 5.0 mm. Body glossy, reddish brown. Head punctate; clypeus, just before margin, glossy with small tubercles; posteriorly with coarse equally spaced, not confluent punctures, punctures smaller on frons. Pronotum with punctures of two sizes, fine and coarse; coarse punctures almost absent on disc, more abundant, larger and closer near lateral margins; without posterior marginal line. Elytral interstriae glossy. Mesoventrite with thin, rectangular callosity; meso- and metatarsi with dense setae ventrally.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF28FF7FFDF6FD99FC51F8AF.taxon	materials_examined	Material examined BRAZIL – Espírito Santo • 1 spec.; Sooretama, Reserva Natural da Vale; Dec. 1988; J. S. Santos leg.; CEMT. – Minas Gerais • 1 spec.; Guanhães; 19 Apr. 1994; C. Jose and I. O. Zanune leg.; CEMT • 1 spec.; Lavras; 5 Dec. 2002; Proj. Fragm. Entom leg.; FI R 10 - B; CEMT • 1 spec.; same data as for preceding; 11 Nov. 2002; FI R 7 - B; CEMT • 3 specs; same data as for preceding; 23 Nov. 2002; FI R 9 - B; CEMT • 2 specs; Lavras, Mata do Capivari; 21 ° 16 ′ 25 ″ S, 44 ° 16 ′ 57 ″ W; Dec. 2001; G. Schiffler leg.; CEMT • 1 spec.; Viçosa; 13 Mar. 1998; T. Zanuncio leg.; CEMT • 1 spec.; same data as for preceding; 30 Jan. 1995; J. N. C. Louzada leg.; CEMT • 9 specs; same data as for preceding; 6 Feb. 1996; G 3; CEMT • 3 specs; same data as for preceding; Mata do Paraiso; Sep. 1994; P. S. Fiúza leg.; CEMT. – Rio de Janeiro • 1 spec.; Nova Friburgo; elev. 1000 m; Jan. 2002; E. Grossi and P. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Jan. 1999; P. Grossi leg.; CEMT • 1 spec.; Nova Friburgo, Macané de Cima; elev. 1500 m; Mar. 2000; Lopes-Andrade, Gumier and Vaz-de-Mello leg.; CEMT • 1 spec.; Nova Friburgo, Macané de Cima; Oct. 2000; P. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Dec. 2000; P. Grossi and E. Grossi leg.; CEMT • 6 specs; same data as for preceding; Dec. 2009; E. Grossi leg.; CEMT • 2 specs; Nova Friburgo, Sítio Caturama; 22 ° 16 ′ 51 ″ S, 42 ° 30 ′ 43 ″ W; Nov. 2016; E. G. Grossi leg.; CEMT • 15 specs; Nova Friburgo; Nov. 2002; E. J. Grossi leg.; CEMT • 2 specs; same data as for preceding; Nov. 2016; E. Grossi leg.; CEMT • 1 spec.; Vila do Pião; 27 Dec. 1997; A. Y. M. Kushida leg.; CEMT. Sexual dimorphism Males with fifth abdominal ventrite shorter than that of females and longer disc of pygidium. Collecting information No information available in literature or with materials studied.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF28FF7DFDC9F8DCFABDF8EF.taxon	description	Figs 22 E, 50 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF28FF7DFDC9F8DCFABDF8EF.taxon	diagnosis	Diagnosis Length 4.8 – 5.2 mm. Body piceous. Head densely and coarse punctate; clypeus sinuate, with glossy band just before anterior margin. Pronotum everywhere densely and coarsely punctate; punctures larger and closer near lateral margins; lateral margins not widely explanate. Elytra with deep striae; interstriae flat. Proventral process wide and scabrous. Mesoventrite with large callosity. Ventral surface of meso- and metatarsi not densely setaceous.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF28FF7DFDC9F8DCFABDF8EF.taxon	materials_examined	Material examined BRAZIL – Acre • 1 spec.; Assis; 10 ° 35 ′ 44 ″ S, 69 ° 57 ′ 11 ″ W; elev. 1.70 m; 21 Sep. 2016; Ortiz and Borges leg.; light trap (CDC); CEMT. – Distrito Federal • 1 spec.; Planaltina, Embrapa Cerrados; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 2 Dec. 2005; C. Oliveira leg.; light; CEMT • 1 spec.; same data as for preceding; 15 ° 36 ′ 46 ″ S, 47 ° 44 ′ 16 ″ W; 2 Dec. 2005; C. Oliveira leg.; light; CEMT. – Mato Grosso • 1 spec.; Alta Floresta, P. E. Cristalino; 9 ° 32 ′ 45 ″ S, 55 ° 54 ′ 57 ″ W; 11 – 15 Nov. 2019; R. A. Azevedo leg.; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 9 ° 49 ′ 17 ″ S, 58 ° 16 ′ 09 ″ W; 10 Dec. 2009; F. Z. Vaz-de-Mello leg.; flor. prim. 250, Inter. Vôo; CEMT • 2 specs; same data as for preceding; 9 ° 49 ′ 08 ″ S, 58 ° 15 ′ 40 ″ W; 19 – 24 Oct. 2012; F. Z. Vaz-de-Mello leg.; PPBioP 1; FIT; CEMT • 1 spec.; same data as for preceding; 9 ° 49 ′ 17 ″ S, 58 ° 15 ′ 32 ″ W; 1 – 3 Nov. 2017; F. Z. Vaz-de-Mello leg.; FIT; CEMT • 1 spec.; Novo Mundo, P. E. Cristalino; 9 ° 28 ′ 16 ″ S, 55 ° 48 ′ 47 ″ W; Nov. 2012; V. Magalhães leg.; P 10, pitfall; CEMT. – Minas Gerais • 1 spec.; Coração de Jesus, Lapa da Maria Cobra; 16 ° 45 ′ 08 ″ S, 44 ° 23 ′ 43 ″ W; 29 Jan. 2015; L. M. Rabelo leg.; CEMT • 1 spec.; Lavras, Campus da UFLA; Nov – Dec. 2001; F. Z. Vaz-de-Mello leg.; light; CEMT • 1 spec.; Lavras, Prox. Poço Bonito; Dec. 2001; Vaz-de-Mello leg.; CEMT. – Pará • 1 spec.; Belterra, Flona-Tapajós; 3 ° 01 ′ 06 ″ S, 55 ° 00 ′ 17 ″ W; 21 Mar. 2019; F. França leg.; primary forest; CEMT • 1 spec.; Jacarecanga; 9 ° 14 ′ 04 ″ S, 56 ° 46 ′ 08 ″ W; Dec. 2015; M. A. L. Bragança leg.; SQD 3, pitfall; CEMT • 1 spec.; Novo Progresso, Faz Florentino; 7 ° 08 ′ S, 55 ° 25 ′ W; elev. 230 m; Oct. – Nov. 2010; D. Krinski leg.; Amazon forest; FIT; CEMT • 1 spec.; Redenção, Pinkaiti-Aik; 7 ° 46 ′ S, 51 ° 58 ′ W; 23 May 1999; P. Y. Scheffler leg.; CEMT. – Rondônia • 1 spec.; 62 km SW of Ariquemes, nr Fazenda Rancho Grande; 8 – 20 Nov. 1994; J. Eger and C. O’Brien leg.; black light; CEMT • 2 specs; Porto Velho, Rio das Garças; 8 ° 49 ′ 48 ″ S, 63 ° 46 ′ 45 ″ W; 4 May 2017; D. C. Santos and K. K. G. Silva leg.; FIT; CEMT. ECUADOR – Napo • 1 spec.; Yasuni Res.; 0 ° 40 ′ S, 76 ° 24 ′ W; Jul. 1999; A. Tischechkin leg.; Mid Rio Tiputini; CEMT. PERU – Loreto • 1 spec.; Allpahuayo ca 30 km S of Iquitos; 26 Mar. 1998; S. Golovatch leg.; terra firme primary tropical rainforest; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF28FF7DFDC9F8DCFABDF8EF.taxon	description	Sexual dimorphism Males with fifth abdominal ventrite shorter than that of females and longer disc of pygidium.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF28FF7DFDC9F8DCFABDF8EF.taxon	discussion	Collecting information Specimens can be collected with light and flight intercept traps. Specimens were collected at light in Pinus caribbea subsp. hondurensis (Sénécl.) Silba stands and tropical rainforests (Stebnicka 2009).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2BFF7CFE2BFC08FAC6F924.taxon	description	Figs 20 A, 51 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2BFF7CFE2BFC08FAC6F924.taxon	diagnosis	Diagnosis Length 4.2 – 5.5 mm. Body piceous some specimens with light elytra. Head punctate with fine punctures only. Pronotum with fine, dense punctures similar to those of head; larger coarser punctures mixed with fine ones; coarser punctures scarce on disc but more abundant near lateral and posterior margins; lateral and posterior marginal furrow preset; lateral margin crenulate. Elytra glabrous dorsally; interstriae with small setae near posterior margin; lateral margin fringed with setae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2BFF7CFE2BFC08FAC6F924.taxon	materials_examined	Material examined BRAZIL – Paraná • 3 specs; Piraquara, Manasiais da Serra; elev. 1100 m; 22 Mar. 2012; Grossi and Leivas leg.; nest of Acromyrmex; CEMT. – Santa Catarina • 2 specs; Nova Teutônia; 22 Nov. 1952; F. Plaumann leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2BFF7CFE2BFC08FAC6F924.taxon	description	Sexual dimorphism Males with fifth abdominal ventrite parallel-sided and shorter than that of females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2BFF7CFE2BFC08FAC6F924.taxon	discussion	Collecting information Specimens are attracted to light and were found in detritus piles of Acromyrmex Mayr, 1865 ants.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2BFF7CFDC5FE94FA84FCC2.taxon	diagnosis	Diagnosis Length 4.2 – 5.5 mm. Body oval, broad, dorsally setaceous, reddish brown to piceous. Head broad, punctate; clypeus sinuate medially, margin thickened; gena obtuse and setaceous. Pronotum with marginal line; lateral margins fringed with setae. Elytra with small humeral teeth, setaceous. Mesoventrite with mesometasternal carina. Metatibiae with lateral apical spine reduced, inconspicuous among apical fringe of setae or absent.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2BFF7CFDC5FE94FA84FCC2.taxon	discussion	Remarks Paraplesiataenius has three recognized species, P. catarinaensis, P. genieri and P. tremolerasi (Schmidt, 1911), currently only reported from Argentina and Brazil (Stebnicka 2003 c, 2009). A key for all species can be found in Stebnicka (2003 c).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2BFF63FDC1F923FBB9FD6B.taxon	description	Figs 20 B – C, 51 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2BFF63FDC1F923FBB9FD6B.taxon	diagnosis	Diagnosis Length 4.5 – 5.1 mm. Body reddish brown, opaque, shagreened. Head with clypeus more finely punctate than frons. Pronotum swollen laterally; coarsely punctate; punctures close to anterior margin, larger posteriorly and near lateral margins; surface finely setiferous. Elytra with interstriae with a row of large setae along outer margin, interstriae glabrous medially.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2BFF63FDC1F923FBB9FD6B.taxon	materials_examined	Material examined BRAZIL – Paraná • 3 specs; Piraquara, Manasiais da Serra; 24 Apr. 2012; P. Grossi leg.; Acromyrmex subterraneus nest; CEMT • 1 spec.; same data as for preceding; elev. 1100 m; 22 Mar. 2012; Grossi and Leivas leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2BFF63FDC1F923FBB9FD6B.taxon	description	Sexual dimorphism Males with pronotal sides arcuate; posterior angles are truncate; posterior margin lobed.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF2BFF63FDC1F923FBB9FD6B.taxon	discussion	Collecting information Specimens were extracted from Acromyrmex subterraneus (Forel, 1893) nest.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF34FF63FD94FD62FA9BFA50.taxon	diagnosis	Diagnosis Length 3.8 – 5.9 mm. Body strongly convex in lateral and frontal views, glossy, reddish brown. Head with strongly developed transverse wrinkles; clypeus emarginate, weakly gibbous. Pronotum convex, robust, with only moderate punctures or with moderate punctures intermixed with coarse, scarce punctures restricted to posterior and lateral margins; lateral margins crenate, fringed with large and evident setae. Elytra convex. Ventral surface glossy, variably setaceous. Abdominal ventrites weakly strigate, except the sixth one which is more strongly strigate; ventrites punctate with fine punctures and larger, not coarse, setaceous punctures. First metatarsomere discretely curved.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF34FF63FD94FD62FA9BFA50.taxon	discussion	Remarks Parataenius is comprised of five species recorded from Southeastern United States, several South American countries and was introduced in Africa, Europe and Oceania (Stebnicka 2009; Stebnicka & Skelley 2009). A key to all species was written by Stebnicka & Skelley (2009).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF34FF62FD9BF997FD6FFD6F.taxon	description	Figs 12 F, 51 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF34FF62FD9BF997FD6FFD6F.taxon	diagnosis	Diagnosis Length 4.5 – 5.0 mm. Body suboval, dorsally glabrous. Head with transverse wrinkles; clypeus impunctate, emarginate. Pronotum convex, with moderate punctures distributed across its surface; lateral margin crenulate, fringed with setae. Abdominal ventrites weakly strigate, except the sixth one; ventrites with punctures of two sizes, larger punctures setiferous.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF34FF62FD9BF997FD6FFD6F.taxon	materials_examined	Material examined ARGENTINA – Mendoza • 1 spec.; Reserva La Payunia, Valle del Saino; 36 ° 07 ′ 01 ″ S, 68 ° 48 ′ 01 ″ W; elev. 1706 m; 6 Jan. 2003; F. C. Ocampo and A. B. T. Smith leg.; guanaco dung; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF34FF62FD9BF997FD6FFD6F.taxon	description	Sexual dimorphism Without evident dimorphism.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF34FF62FD9BF997FD6FFD6F.taxon	discussion	Collecting information Specimens were collected with light traps and are attracted to cow and guanaco Lama guanicoe (Müller, 1776) feces (Stebnicka & Skelley 2007).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF35FF61FDE4FC9EFE9FFEC7.taxon	description	Figs 12 D – E, 51 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF35FF61FDE4FC9EFE9FFEC7.taxon	diagnosis	Diagnosis Length 3.8 – 5.9 mm. Body robust, elongate, glabrous dorsally. Head with strong transverse wrinkles, punctures present at frons; clypeus emarginate, impunctate. Pronotum with punctures of two sizes; larger punctures scarce, restricted to posterior and lateral margins; lateral margins fringed with setae. Elytra with interstriae punctate; punctures fine and widely spaced. Abdominal ventrites with large punctures to the sides.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF35FF61FDE4FC9EFE9FFEC7.taxon	materials_examined	Material examined ARGENTINA – Corrientes • 1 spec.; Corrientes, Barrio Víctor Colas; 27 ° 28 ′ 50 ″ S, 58 ° 47 ′ 18 ″ W; elev. 64 m; 21 Oct. 2012; M. I. Polesel leg.; light; CEMT. – Mendoza • 535 specs; Desaguadero; 5 Feb. 2017; G. Arriágada leg.; CEMT. – San Luis • 2 specs; Desaguadero; 11 Jan. 2005; C. Fortino leg.; light trap; CEMT. – San Luis • 124 specs; Belgrano, Fundo El Molle; 33 ° 02 ′ 23 ″ S, 66 ° 30 ′ 48 ″ W; elev. 622 m; 7 Feb. 2017; G. Arriágada leg.; CEMT. BRAZIL – Rio Grande do Sul • 1 spec.; Cidreira; 31 May 2003; J. Alvenir leg.; dunas em restinga [dunes in restinga vegetation]; pitfall; CEMT • 2 specs; Santa Cruz do Sul, Cerro Alegre; 28 ° 48 ′ 11 ″ S, 52 ° 19 ′ 51 ″ W; 11 Feb. 2012; R. Bohn leg.; annual crop, bovine feces; CEMT. UNITED STATES OF AMERICA – Alabama • 1 spec.; Barbour Co., Blue Springs St Pk.; 11 Mar. 1985; S. C. Harris leg.; CEMT • 1 spec.; Bibb Co., Little Cahaba Riv.; 12 Jul. 1981; S. C. Harris leg.; CEMT. – Mississippi • 1 spec.; Perry Co., Camp Shelby; 18 Mar. 2000; T. C. Lockley leg.; T- 44 site 1; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF35FF61FDE4FC9EFE9FFEC7.taxon	description	Sexual dimorphism Males with penultimate abdominal ventrite thinner than that of female.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF35FF61FDE4FC9EFE9FFEC7.taxon	discussion	Collecting information Adult are commonly attracted to light and larvae have been collected in bovine faecal masses (Verdú & Galante 1999).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF36FF61FD93FE01FA82FA9C.taxon	diagnosis	Diagnosis Length 2.8 – 4.0 mm. Body elongate, parallel-sided, dorso-ventrally flattened, reddish brown. Head variably sculptured, in some species is only punctured, while in others the clypeus may be tuberculated; clypeus sinuate; gena prominent, obtuse; frontoclypeal suture present or not. Pronotum quadrate, with fine and coarse punctures. Elytra elongate; striae deep with coarse and round punctures; interstriae variably punctured. Metatibiae with 2 – 3 additional apical spines on apical lateral margin. Abdominal ventrites coalescent, sutures visible only laterally, except second ventrite; ventrites somewhat depressed laterally; sixth ventrite foveate laterally.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF36FF61FD93FE01FA82FA9C.taxon	discussion	Remarks Passaliolla has five recognized species and it is recorded from several countries in Central and South America. For more information about the diversity of this genus and keys for all its species see Stebnicka (2000).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF36FF60FDE6FACBFB47FC90.taxon	description	Figs 23 C – E, 52 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF36FF60FDE6FACBFB47FC90.taxon	diagnosis	Diagnosis Length 3.8 – 4.0 mm. Head tuberculate, tubercle extending to frontoclypeal suture; frontoclypeal suture present as a fine impunctate carina. Pronotum with moderate and coarser punctures; coarse punctures greatly separated, moderate punctures densely distributed.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF36FF60FDE6FACBFB47FC90.taxon	materials_examined	Material examined BRAZIL – Minas Gerais • 1 spec.; Viçosa, Mata do Paraíso; 2 Dec. 2014; C. Lopes et al. leg.; CEMT • 1 spec.; Viçosa; 20 ° 48 ′ 11 ″ S, 42 ° 51 ′ 25 ″ W; elev. 725 m; 1 May 2013; J. Chaul and R. Jesus leg.; epigaeic Winkler extractor; CEMT. – Rio de Janeiro • 1 spec.; Parque Nacional do Itatiaia, Casa do pesquisador; elev. 750 m; 22 – 24 Oct. 2010; W. Beiroz and M. Cupello leg.; flight intercept trap; CEMT. – São Paulo • 1 spec.; Bertioga, Parque das Neblinas; 23 ° 44 ′ S, 46 ° 10 ′ W; elev. 790 m; Feb. 2014; R. V. Nunes leg.; human feces; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF36FF60FDE6FACBFB47FC90.taxon	description	Sexual dimorphism Males and females are indistinguishable from each other.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF36FF60FDE6FACBFB47FC90.taxon	discussion	Collecting information Specimens were found at altitudes ranging from 725 to 790 metres. Specimens were collected with flight traps and in forest litter, this species was also found in Tapirus and human dung. Remarks Information about the distribution of this species remains uncertain. In Brazil, this species was only reported from “ Amazonas ” (Stebnicka 2002). However, specimens from southeastern Brazil meeting the diagnostic character provided by Stebnicka (2002, 2009) were found in the CEMT. Regardless, further studies must be conducted to more accurately ascertain the distribution of this species.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF37FF60FD9FFCD7FA83F802.taxon	description	Figs 24 A, 52 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF37FF60FD9FFCD7FA83F802.taxon	diagnosis	Diagnosis Length 2.8 – 3.0 mm. Head punctate, anteriorly tuberculate, tubercles not extending beyond anterior gibbosity; frontoclypeal suture indistinct. Pronotum with punctures of two sizes, coarse and moderate; coarse and moderate punctures densely distributed across pronotum. Elytra with striae deep, coarsely and closely punctate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF37FF60FD9FFCD7FA83F802.taxon	materials_examined	Material examined BRAZIL – Amazonas • 1 spec.; Manaus Mu Est. AM. 01. hm 31 CEPLAc [unknown meaning]; 9 May 1976; W. Irioma and S. Serrano leg.; CEMT • 4 specs; Coari, Rio Urucu; 4 ° 55 ′ 53 ″ S, 65 ° 18 ′ 13 ″ W; 25 Feb. – 10 Mar. 1995; P. F. Bührnheim et al. leg.; RUC- 36; CEMT • 1 spec.; Tefé, Locação São Mateus; 4 ° 43 ′ 24 ″ S, 65 ° 05 ′ 06 ″ W; 7 – 16 Sep. 1994; P. F. Bührnheim et al. leg.; CEMT. – Mato Grosso • 31 specs; Cotiguaçu, Fazenda São Nicolau; 9 ° 49 ′ 22 ″ S, 58 ° 15 ′ 56 ″ W; 1 Nov. 2017; Vaz-de-Mello et al. leg.; manually; CEMT • 3 specs; Cotiguaçu, Fazenda São Nicolau, Prainha; 9 ° 51 ′ 53 ″ S, 58 ° 13 ′ 13 ″ W; 2 Nov. 2017; Vaz-de-Mello et al. leg.; manually; CEMT • 3 specs; Cotiguaçu, Fazenda São Nicolau; 9 ° 52 ′ 31 ″ S, 58 ° 16 ′ 21 ″ W; 2 Nov. 2017; Vaz-de-Mello et al. leg.; CEMT. – Rondônia • 1 spec.; Guaporé, 12 ° 16 ′ 05 ″ S, 60 ° 42 ′ 30 ″ W; 23 Apr. 2006; J. A. Rafael and F. F. Xavier leg.; CEMT. UNKNOWN COUNTRY • [Unreadable]; 18 Apr. 1976; E. Rujino leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF37FF60FD9FFCD7FA83F802.taxon	description	Sexual dimorphism Males and females are indistinguishable from each other.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF37FF60FD9FFCD7FA83F802.taxon	discussion	Collecting information Specimens were collected inside rotten logs (Vaz-de-Mello pers. com.) and forest litter (Stebnicka 2009).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF30FF67FD96FE95FB69FA69.taxon	description	Figs 23 F, 52 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF30FF67FD96FE95FB69FA69.taxon	diagnosis	Diagnosis Length 3.1 – 3.9 mm. Head with clypeus entirely granulate; frontoclypeal suture indistinct. Pronotum with punctures fine and moderate; coarse punctures spaced at least one diameter; pronotum with small foveae laterally.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF30FF67FD96FE95FB69FA69.taxon	materials_examined	Material examined BRAZIL – Espírito Santo • 5 specs; Venda Nova do Imigrante; Jan. 2000; Lopes-Andrade and Naviaux leg.; CEMT. – Mato Grosso • 8 specs; Chapada dos Guimarães, Vale Jamacá; 15 ° 27 ′ 53 ″ S, 55 ° 42 ′ 55 ″ W; elev. 760 m; 7 Nov. 2014; S. Boucher and R. V. Nunes leg.; CEMT. COLOMBIA – Vaupés • 1 spec.; R. N. Mosiro-Itajura (Caparú), Centro Ambiental; 1 ° 04 ′ S, 69 ° 31 ′ W; elev. 60 m; 20 Jan. – 1 Feb. 2003; M. Shakoy and D. Arias leg.; FIT; CEMT. FRENCH GUIANA • 1 spec.; St Laurent de Maroni, Bélvédère de Saül; 3 ° 37 ′ 22 ″ N, 53 ° 12 ′ 57 ″ W; elev. 326 m; 14 Mar. 2011; SEAG leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF30FF67FD96FE95FB69FA69.taxon	description	Sexual dimorphism Males and females are indistinguishable from each other.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF30FF67FD96FE95FB69FA69.taxon	discussion	Collecting information Specimens were collected in forest litter, inside logs and some have been captured with flight intercept traps. This species was found at altitudes ranging from 725 to 1000 metres. Remarks Passaliolla corticalis is another species with uncertain distribution. Originally described from “ Ega ” now known as Tefé (IBGE 1990 a), several specimens matching the diagnosis provided by Stebnicka (2000) have been found in forest fragments in the states of Mato Grosso and Espírito Santo.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF30FF65FDE6F99CFD5FFCAF.taxon	description	Fig. 24 B, 52 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF30FF65FDE6F99CFD5FFCAF.taxon	diagnosis	Diagnosis Length 2.8 – 3.2 mm. Head without granules; clypeus subrugosely punctate medially; frontoclypeal suture indistinct. Pronotum with punctures of two sizes, fine and coarse, everywhere densely distributed. Elytra with striae deep and coarsely punctured.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF30FF65FDE6F99CFD5FFCAF.taxon	materials_examined	Material examined BRAZIL – Espírito Santo • 1 spec.; Venda Nova do Imigrante; Feb. 1998; Falgueto and Vaz-de-Mello leg.; CEMT • 10 specs; Venda Nova do Imigrante, Vila Santa Cruz; Jan. 2011; F. Z. Vaz-de-Mello leg.; CEMT. – Minas Gerais • 1 spec.; Viçosa; Dec. 1995; F. Z. Vaz-de-Mello leg.; CEMT. ECUADOR – Napo • 1 spec.; Res. Étnica Waorani, 1 km S of Onlone Gare Camp, Trans. Ent.; 0 ° 39 ′ 10 ″ S, 76 ° 26 ′ 00 ″ W; elev. 220 m; 8 Feb. 1996; T. L Erwin. et al. leg.; fogging with insecticides; CEMT. – Pastaza • 1 spec.; Arajuno, Comuna Oglán, Río Oglán; elev. 580 m; 25 Jul. 2009; P. Araujo leg.; fumig; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF30FF65FDE6F99CFD5FFCAF.taxon	description	Sexual dimorphism Males and females are indistinguishable from each other.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF30FF65FDE6F99CFD5FFCAF.taxon	discussion	Collecting information Specimens were found inside rotting logs.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF32FF65FD94FCD9FEE8FBE4.taxon	diagnosis	Diagnosis Length 4.2 – 4.8 mm. Body elongate, cylindrical, dorsally glabrous, reddish testaceous. Head punctate; clypeus sinuate, margin not thickened medially. Pronotum not fringed with setae; laterally and posteriorly with marginal line; posterior angles sinuate. Elytra with fine humeral teeth. Tarsus of meso- and metatibiae slender and as long as tibiae. Mesometaventral carina absent. Abdominal ventrites minutely strigate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF32FF64FDD3FBE0FE49FEC6.taxon	description	Figs 13, 52 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF32FF64FDD3FBE0FE49FEC6.taxon	diagnosis	Diagnosis Length 4.2 – 4.8 mm. Head with clypeus sinuate; frons coarsely punctate. Pronotum not fringed by setae; margin entirely with marginal line; posterior angles sinuate. Elytra with interstriae glossy, punctate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF32FF64FDD3FBE0FE49FEC6.taxon	materials_examined	Material examined BRAZIL – Distrito Federal • 1 spec.; Brasília, RECOR-IBGE; Nov. 1999; M. Milhomem leg.; CEMT • 6 specs; same data as for preceding; I. Diniz leg.; CEMT • 2 specs; Planaltina, Embrapa CPAC; 15 ° 36 ′ 24 ″ S, 47 ° 42 ′ 45 ″ W; elev. 1025 m; 18 Nov. 2016; C. M. Oliveira leg.; flight intercept trap; CEMT. – Goiás • 1 spec.; Joanópolis, Chac. São José; 16 ° 16 ′ 43 ″ S, 48 ° 50 ′ 57 ″ W; 26 Jan. 2019; E. Carvalho leg.; FIT; CEMT. – Minas Gerais • 1 spec.; Viçosa; 10 Nov. 1999; F. Z. Vaz-de-Mello leg.; FIT; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF32FF64FDD3FBE0FE49FEC6.taxon	description	Sexual dimorphism Males with head densely punctate; clypeus medially gibbous; protibia with lateral teeth of unequal sizes; apical spur inwardly, strongly curved. Tarsi longer and slenderer than that of female. Females with clypeus not particularly gibbous, wrinkled; frons strongly punctate. Protibiae with lateral teeth similar in size; apical spur lanceolate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF32FF64FDD3FBE0FE49FEC6.taxon	discussion	Collecting information Specimens were collected at altitudes ranging from 700 to 1025 metres with flight intercept traps and with black light traps.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF33FF64FDE6FE01FA82F9F7.taxon	diagnosis	Diagnosis Length 3.0 – 6.0 mm. Body cylindrical, somewhat flattened, dark reddish brown. Head variably punctured, may present tubercles depending on the species observed; clypeus emarginate; frontoclypeal suture visible or not, if visible, carinate. Pronotum variably punctured, never fringed by setae. Elytra with humeral denticle fine. Tibiae with two poorly developed transversal carinae. Metatibiae with 2 – 3 additional apical spines on its outer margin. Abdominal ventrites sinuate, strigate anteriorly.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF33FF64FDE6FE01FA82F9F7.taxon	discussion	Remarks Saprosites currently has 136 recognized species distributed across the Afrotropical, Australian, Neotropical, Oriental and Palearctic regions (Stebnicka 2001 a; Schoolmeesters 2023). A key to the 16 Neotropical species was written by Stebnicka (2001 a).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF33FF6BFDE6F9F6FEE1FAC7.taxon	description	Figs 25 B – D, 53 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF33FF6BFDE6F9F6FEE1FAC7.taxon	diagnosis	Diagnosis Length 3.0 – 4.0 mm. Head punctate, punctures minute near anterior margin of clypeus, fine near and on frons; clypeus sinuate. Pronotum quadrate; densely punctate with fine and coarse punctures; posterior angles not serrate; posterior margin without marginal line. Elytra with lateral interstriae less coarsely punctate than discal interstriae. Abdominal ventrites coarsely and densely punctate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF33FF6BFDE6F9F6FEE1FAC7.taxon	materials_examined	Material examined BRAZIL – Espírito Santo • 1 spec.; Venda Nova do Imigrante; Feb. 1998; Falqueto and Vaz-de-Mello leg.; CEMT. – Mato Grosso • 1 spec.; Alta Floresta, P. E. Cristalino; 9 ° 40 ′ 59 ″ S, 55 ° 54 ′ 58 ″ W; 20 – 24 Nov. 2019; R. A. Azevedo leg.; FIT; CEMT • 1 spec.; Chapada dos Guimarães, Vale Jamacá; 15 ° 27 ′ 53 ″ S, 55 ° 42 ′ 55 ″ W; elev. 760 m; 7 Nov. 2014; S. Boucher and R. V. Nunes leg.; CEMT • 1 spec.; Cotiguaçu, Fazenda São Nicolau, PPBioP 1; 9 ° 49 ′ 08 ″ S, 58 ° 15 ′ 40 ″ W; 19 – 24 Oct. 2012; F. Z. Vaz-de-Mello leg.; PPBioP 1, FIT; CEMT. – Minas Gerais • 1 spec.; Viçosa; Nov. 1999; F. Z. Vaz-de-Mello and Serrano leg.; CEMT • 2 specs; same data as for preceding; Nov. 2000; CEMT • 1 spec.; Mata do Paraíso; 2 Dec. 2014; C. Lopes-Andrade and S. Aloquito leg.; CEMT. – Pará • 1 spec.; Belém, IPEAN; May 1985; N. Degallier leg.; CEMT. – Rio de Janeiro • 1 spec.; Angra dos Reis, Estação do Rio Jussaral; Oct. 1935; L. Trave and E. T. Lopes leg.; CEMT • 1 spec.; Macaé, Rio São Pedro; 22 ° 13 ′ 45 ″ S, 42 ° 02 ′ 39 ″ W; elev. 111 m; 22 Mar. 2009; J. Nessimian et al. leg.; light; CEMT • 2 specs; Nova Friburgo; Oct. 1996; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Nova Friburgo, Cascata; 22 ° 21 ′ 55 ″ S, 42 ° 13 ′ 03 ″ W; elev. 319 m; 8 Mar. 2009; J. Nessimian leg.; af 2 ° ord [second-order tributary] of Rio Macaé; UV light; CEMT • 1 spec.; Pedro do Rio; 4 Dec. 1966; J. R. Celso leg.; CEMT. – São Paulo • 1 spec.; Ribeirão Preto, EERP-MST; 21 ° 13 ′ 33 ″ S, 47 ° 51 ′ 21 ″ W; 22 May 2009; G. Biffi and E. A. Nascimento leg.; CEMT. ECUADOR – Pastaza • 1 spec.; Arajuno, B. P. Oglán; elev. 540 m; 9 Oct. 2009; F. Gallo and W. Chamorro leg.; collected manually on trunk; CEMT • 1 spec.; Arajuno, Río Oglán; elev. 598 m; 25 Jul. 2009; P. Araujo leg.; fumig [fumigation]; CEMT. FRENCH GUIANA • 1 spec.; St Laurent de Maroni, Bélvédère de Saül; 3 ° 37 ′ 22 ″ N, 53 ° 12 ′ 57 ″ W; elev. 326 m; 14 Mar. 2011; SEAG leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF33FF6BFDE6F9F6FEE1FAC7.taxon	description	Sexual dimorphism Males with pronotum wider and less densely punctate than that of females; fifth abdominal ventrites shorter, apical lip of pygidium longer than that of females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF33FF6BFDE6F9F6FEE1FAC7.taxon	discussion	Collecting information This species has been collected inside rotten logs, litter, sticks with fungus, flight intercept and UV light traps next to streams (Stebnicka 2009). This species was found at altitudes ranging from 111 to 760 metres.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3CFF6AFD89FA06FC2DFD47.taxon	description	Figs 24 D – F, 53 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3CFF6AFD89FA06FC2DFD47.taxon	diagnosis	Diagnosis Length 5.0 – 6.0 mm. Head with clypeus transversely carinate behind medial emargination; frons coarsely punctate. Pronotum with minute and coarse punctures; minute punctures everywhere distributed; coarse punctures closer and wider near anterior angles; posterior angles smooth. Elytra with lateral striae thinner than lateral interstriae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3CFF6AFD89FA06FC2DFD47.taxon	materials_examined	Material examined BRAZIL – Espírito Santo • 1 ♂; Venda Nova do Imigrante; Dec. 2000; F. Z. Vaz-de-Mello leg.; CEMT. – Minas Gerais • 1 ♀; Viçosa; 2 Oct. 1982; P. S. F. Fiuza leg.; CEMT • 1 ♀; same data as for preceding; Nov. 1999; Vaz-de-Mello and Serrano leg.; CEMT • 1 ♂; same data as for preceding; Nov. 2000; F. Z. Vaz-de-Mello leg.; CEMT. – Rio de Janeiro • 5 ♀♀; Macaré de Cima; Oct. 2002; P. Grossi and E. Grossi leg.; CEMT • 1 ♀; Nova Friburgo; elev. 1000 m; Nov. 2000; P. Grossi leg.; CEMT • 1 ♂; Nova Friburgo; Nov. 1996; F. Z. Vaz-de-Mello leg.; CEMT • 1 ♀; Pedro do Rio; 17 Sep. 1961; J. R. Celso leg.; CEMT • 1 ♀; Petrópolis, Alto da Serra; 10 Sep. 1961; H. Schubart leg.; CEMT. – São Paulo • 4 ♀♀; 24 Oct. 1914; Saude leg.; CEMT • 1 ♂; São Paulo; 10 Oct. 1920; Saude leg.; CEMT • 2 ♂; same data as for preceding; 4 Apr. 1920; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3CFF6AFD89FA06FC2DFD47.taxon	description	Sexual dimorphism Females with clypeus granulate, transverse carina more developed and frontoclypeal suture elevated; males with clypeus not tuberculate, instead, finely punctate; transverse carina less acutely elevated; frontoclypeal suture indistinct; frons more coarsely punctate than clypeus.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3CFF6AFD89FA06FC2DFD47.taxon	discussion	Collecting information Specimens were collected at altitudes up to 1000 m with light traps.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3DFF6AFD97FC87FB64F9D6.taxon	description	Fig. 25 E – F	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3DFF6AFD97FC87FB64F9D6.taxon	diagnosis	Diagnosis Length 4.2 – 4.8 mm. Head punctate; punctures near clypeal margin minute, punctures at and near frons slightly larger. Pronotum with punctures fine and coarse, the latter more scattered, closer near lateral margins. Elytra with discal interstriae with punctures as coarse as the lateral ones. Abdominal ventrites with minute scattered punctures.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3DFF6AFD97FC87FB64F9D6.taxon	materials_examined	Material examined COLOMBIA – Risaralda • 1 spec.; Ucumari La Suiza; elev. 2800 m; Mar. 2000; J. Noriega leg.; CEMT. ECUADOR – Pastaza • 1 spec.; Chuyayacu; elev. 810 m; May 1999; P. Araujo leg.; fumigación dosel [canopy fumagation]; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3DFF6AFD97FC87FB64F9D6.taxon	description	Sexual dimorphism Males with pronotum broader and less densely punctate than that of females; fifth abdominal ventrite shorter and apical lip longer in males.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3DFF6AFD97FC87FB64F9D6.taxon	discussion	Collecting information Specimens were collected at altitudes ranging from 810 to 3000 metres and can be found inside dead trees (Stebnicka 2009). The specimens analyzed were collected by canopy fogging.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3DFF68FD9DF911FAEBFBA9.taxon	description	Figs 24 C, 53 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3DFF68FD9DF911FAEBFBA9.taxon	diagnosis	Diagnosis Length 3.8 – 4.0 mm. Head punctate; punctures on frons larger than those on clypeus; clypeus emarginate; frontoclypeal suture faint, present as a fine, glabrous and glossy angulate line. Pronotum everywhere coarsely, densely punctate, punctures closer and coarser laterally; posterior margin with marginal line; margin evidenced by a row of coarse punctures; posterior angles serrate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3DFF68FD9DF911FAEBFBA9.taxon	materials_examined	Material examined BRAZIL – Espírito Santo • 2 specs; Domingos Martins, Pq. E. Pedra Azul; elev. 1500 m; Jan. 2000; Lopes-Andrade and Vaz-de-Mello leg.; CEMT • 1 spec.; Venda Nova do Imigrante; Nov. 1999; Falquear and Vaz-de-Mello leg.; CEMT • 1 spec.; same data as for preceding; Nov. 1999; Vaz-de-Mello and Serrano leg.; CEMT. – Minas Gerais • 2 specs; Lima Duarte, P. E. Ibitipoca; Jun. 2001; F. Z. Vaz-de-Mello leg.; CEMT • 2 specs; Viçosa; 17 Nov. 1999; A. Serrano leg.; CEMT • 1 spec.; same data as for preceding; Nov. 1999; Vaz-de-Mello and Serrano leg.; CEMT. – Paraná • 4 specs; Castro Socavão, Cachoeira Roncadura; 24 Jun. 2006; Grossi and Parizotto leg.; CEMT. – Rio de Janeiro • 1 spec.; Nova Friburgo, Macaé de Cima; Dec. 1998; Grossi and Vaz-de-Mello leg.; CEMT • 5 specs; same data as for preceding; Jan. 1999; P. Grossi leg.; CEMT • 1 spec.; same data as for preceding; Jan. 2000; E. Grossi and P. Grossi leg.; FIT; CEMT • 1 spec.; Parque Nacional do Itatiaia, Casa do Pesquisador; elev. 750 m; 22 – 24 Oct. 2010; W. Beiroz and M. Cupello leg.; flight interception trap; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3DFF68FD9DF911FAEBFBA9.taxon	description	Sexual dimorphism Males with fifth abdominal ventrite shorter than that of female; males with weakly strigate ventrites; lip of pygidium longer.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3DFF68FD9DF911FAEBFBA9.taxon	discussion	Collecting information Specimens were found at altitudes up to 1500 m and can be collected with flight intercept traps.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3FFF6FFD89FBDCFC58FEC7.taxon	description	Figs 25 A, 53 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3FFF6FFD89FBDCFC58FEC7.taxon	diagnosis	Diagnosis Length 5.0 – 5.5 mm. Head punctate; clypeus emarginate, finely and densely punctate; frontoclypeal suture impressed; frons with punctures larger and less dense than clypeus. Pronotum with minute and coarse punctures; coarse punctures scattered on disc, closer and larger near lateral margins, especially near anterior angles; posterior margin with marginal line of coarse punctures; posterior angles serrate. Elytra with lateral striae coarsely and closely punctate, as wide or wider than interstriae; strial punctures crenate interstriae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3FFF6FFD89FBDCFC58FEC7.taxon	materials_examined	Material examined BRAZIL – Ceará • 6 specs; Ubajara, PN Ubajara, Trilha da Samambaia; 3 ° 50 ′ 22 ″ S, 40 ° 53 ′ 55 ″ W; elev. 875 m; 16 – 20 Feb. 2013; Vaz-de-Mello and Grossi leg.; trilha da samambaia [fern trail]; manually; CEMT. – Distrito Federal • 1 spec.; Brasília; Nov. 2002; N. Degallier leg.; CEMT. – Mato Grosso • 6 specs; Santo Antônido do Leverger, IFMT-São Vicente da Serra; Jun. 2012; G. A. P. C. Neves leg.; CEMT. – Minas Gerais • 4 specs; Lavras, Campus da UFLA; Nov. – Dec. 2001; F. Z. Vaz-de-Mello leg.; light; CEMT • 1 spec.; Viçosa; Nov. 1996; Vaz-de-Mello and Hardy leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3FFF6FFD89FBDCFC58FEC7.taxon	description	Sexual dimorphism Males pronotum less densely punctate than that of females; fifth abdominal ventrite shorter than that of females; lip of pygidium longer.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3FFF6FFD89FBDCFC58FEC7.taxon	discussion	Collecting information Specimens were collected under wood bark and rotten logs associated with fungi. This species was found at altitudes ranging from 400 to 1280 metres (Stebnicka 2001 a).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF38FF6FFDBCFC41FC1BF936.taxon	description	Fig. 54 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF38FF6FFDBCFC41FC1BF936.taxon	diagnosis	Diagnosis Length 4.8 – 5.0 mm. Lateral margins of pronotum and margins of elytra greatly explanate. Head wide; clypeus emarginate. Pronotum with punctures of two sizes; fine and moderate. Elytra with interstriae convex, lacking humeral teeth.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF38FF6FFDBCFC41FC1BF936.taxon	description	Sexual dimorphism Males with apical spurs of protibia inwardly hooked.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF38FF6FFDBCFC41FC1BF936.taxon	discussion	Collecting information Specimens were collected at light. Remarks There were no specimens available to be examined. The Brazilian record and other information here presented were based on available literature (Stebnicka 2005 a, 2009).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF38FF6FFDBCFE01FA83FC05.taxon	diagnosis	Diagnosis Length 4.8 – 5.0 mm. Body elongated, glabrous reddish testaceous. Head wide, covered with fine punctures; clypeus emarginate. Pronotum laterally explanate, with punctures of two sizes fine and moderate; fringed by spatulate setae. Elytra with humeral denticle developed; margin of elytra explanate. Meso- and metatibiae slim, somewhat circular in cross section, almost completely glabrous, except for a few sparse setae concentrated in the apical third.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF38FF6FFDBCFE01FA83FC05.taxon	discussion	Remarks Selviria currently has only two known species from Bolivia and Brazil (Stebnicka 2009).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF38FF6EFDFEF931FD7EFC4E.taxon	description	Figs 17 B, 54 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF38FF6EFDFEF931FD7EFC4E.taxon	diagnosis	Diagnosis Length 4.8 – 5.0 mm. Head wide with finely punctate; clypeus emarginate, margin upturned. Pronotum laterally explanate; fringed by spatulate setae; posterior angles not distinctly explanate. Elytra with humeral teeth present, interstriae flat; lateral and posterior margins explanate. Meso- and metatibiae slim, somewhat circular in a cross section, almost completely glabrous, except for a few sparse setae concentrated in apical third.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF38FF6EFDFEF931FD7EFC4E.taxon	materials_examined	Type material examined Paratype BRAZIL • 1 spec.; “ [white paper, printed] Br-Ms- Três Lagoas / Horto Rio Verde / Três Lagoas Agroflorestal / Black light trap / Eucalyptus grandis stand / Flechtmann, C. A. H. Col / [hand writen to the side] 19 / X / 1993. // [red paper, printed] Types genus / Selviria / / Dt. Z. Stebnicka. // [yellow paper, printed] Selviria / matogrossoensis / D. t. Z. Stebnicka. ”; CEMT. Other material examined BRAZIL – Mato Grosso do Sul • 1 spec.; Selvíria, UNESP farm; 21 Nov. 1993; C. A. H. Flechtmann leg.; Brachiaria decumbens pasture; black light trap; CEMT. – Minas Gerais • 1 spec.; Ijaci, Fazenda FAEPE; Nov. 2002; J. Louzada leg.; light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF38FF6EFDFEF931FD7EFC4E.taxon	description	Sexual dimorphism Males with apical spur of protibiae inwardly curved.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF38FF6EFDFEF931FD7EFC4E.taxon	discussion	Collecting information Specimens were collected in pasture regions with black light traps and in nests of Solenopsis invicta Buren, 1972 (Stebnicka 1999 a, 2007 a).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF39FF6EFD86FBB9FA82F914.taxon	diagnosis	Diagnosis Length 2.0 – 2.7 mm. Body oval, dorsally glabrous, reddish brown. Head punctate; clypeus sinuate. Pronotum with lateral and posterior margins grooved, not fringed with setae; posterior margin without marginal line; posterior angles sinuate, delimited by two dentiform processes. Elytra with fine humeral teeth; striae wider close to lateral margins. Mesoventrite with V-shaped furrow parallel to the medial margin of mesocoxa; metaventrite with arched groove near the posterior margin of the mesocoxa.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF39FF6EFD86FBB9FA82F914.taxon	discussion	Remarks Tanyana currently has two recognized species recorded from Brazil, Colombia, French Guiana, Guiana, Peru and Suriname (Schoolmeesters 2023; Stebnicka 2003 a; Minkina 2022).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF39FF6DFDBAF952FBC8FCC3.taxon	description	Figs 26 C, 54 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF39FF6DFDBAF952FBC8FCC3.taxon	diagnosis	Diagnosis Length 2.5 – 2.9 mm. Head with coarse punctures scarce, scattered; clypeus weakly sinuate. Pronotum with fine and coarse punctures; coarse punctures scattered, closer near lateral margins; posterior angles weakly sinuate, posterior tooth, present as an obtuse angulation; posterior margin weakly sinuate. Elytral striae coarsely punctate. Abdominal clearly strigate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF39FF6DFDBAF952FBC8FCC3.taxon	materials_examined	Material examined FRENCH GUIANA • 1 spec.; St Laurent de Maroni, Bélvédère de Saül; 3 ° 37 ′ 22 ″ N, 53 ° 12 ′ 57 ″ W; elev. 326 m; 22 Mar. 2011; SEAG leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF39FF6DFDBAF952FBC8FCC3.taxon	description	Sexual dimorphism Unknown	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF39FF6DFDBAF952FBC8FCC3.taxon	discussion	Collecting information Specimens were collected with flight intercept traps. Remarks This species was first recorded for Brazil by Minkina (2022). The single analyzed specimens is damaged, its head is missing and the pronotum is detached from the rest of the body. Due to these facts, part of the diagnosis was made following only the original description (Minkina 2020).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3AFF6DFDF7FC0BFEE2F852.taxon	description	Figs 26 A – B, 54 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3AFF6DFDF7FC0BFEE2F852.taxon	diagnosis	Diagnosis Length 2.0 – 2.1 mm. Head with coarse punctures close and denser coarse punctures; clypeus clearly sinuate. Pronotum with punctures fine and coarse; coarse punctures dense, absent at anterior half of pronotum; posterior angles acutely sinuate, posterior tooth acuminate. Elytral striae coarsely punctate. Abdominal ventrites strigate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3AFF6DFDF7FC0BFEE2F852.taxon	materials_examined	Material examined BRAZIL – Acre • 1 spec.; Mâncio Lima, Parna Serra do Divisor; 7 ° 24 ′ 59 ″ S, 73 ° 37 ′ 56 ″ W; 10 – 14 Dec. 2019; R. A. Azevedo leg.; CEMT. – Mato Grosso • 1 spec.; Alta Floresta, P. E. Cristalino; 9 ° 32 ′ 45 ″ S, 55 ° 54 ′ 57 ″ W; 11 – 15 Nov. 2019; R. A. Azevedo leg.; FIT; CEMT • 1 spec.; same data as for preceding; 9 ° 32 ′ 59 ″ S, 55 ° 54 ′ 58 ″ W; 20 – 24 Nov. 2019; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 9 ° 49 ′ 08 ″ S, 58 ° 15 ′ 40 ″ W; 19 – 24 Oct. 2012; F. Z. Vaz-de-Mello leg.; FIT; CEMT. COLOMBIA – Vaupés • 3 specs; R. N. Mosiro Itajura (Caparú), Centro Ambiental; 1 ° 04 ′ S, 69 ° 31 ′ W; elev. 60 m; 20 Jan. – 1 Feb. 2013; M. Shakoy and D. Arias leg.; FIT; CEMT. FRENCH GUIANA • 1 spec.; Cayenne, Station Nouragues; 4 ° 05 ′ N, 52 ° 41 ′ W; 8 Mar. 2011; SEAG leg.; inselberg; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3AFF6DFDF7FC0BFEE2F852.taxon	description	Sexual dimorphism Males with fifth abdominal ventrite shorter than that of females.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF3AFF6DFDF7FC0BFEE2F852.taxon	discussion	Collecting information Specimens were collected with flight intercept traps at altitudes ranging from 30 to 480 metres (Stebnicka 2003 a).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF04FF53FD9EFC92FA85FA5D.taxon	description	Figs 26 F, 54 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF04FF53FD9EFC92FA85FA5D.taxon	diagnosis	Diagnosis Same as for the genus.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF04FF53FD9EFC92FA85FA5D.taxon	materials_examined	Material examined BRAZIL – Amazonas • 1 spec.; Manaus, Tarumá Mirim; 3 Mar. 1982; CEMT • 1 spec.; Manaus, Rio Tarumá Mirim, Igapo; 10 Mar. 1977; J. Adis leg.; CEMT • 2 specs: same data as for preceding; INPA • 2 specs; same data as for preceding; 24 Mar. 1977; CEMT • 1 spec.; same data as for preceding; 30 Mar. 1977; CEMT • 1 spec.; same data as for preceding; INPA • 1 spec.; same data as for preceding; 13 Apr. 1977; CEMT • 1 spec.; terra firme [dry land forest]; 3 ° 02 ′ S, 60 ° 17 ′ W; 26 May 1983; A. Rodrigues leg.; secondary forest; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF04FF53FD9EFC92FA85FA5D.taxon	description	Sexual dimorphism Unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF04FF53FD9EFC92FA85FA5D.taxon	discussion	Collecting information Specimens have been collected at 90 metres above sea level in the Amazon rainforest (Skelley 2007 b).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF04FF53FDDDFED4FB07FD59.taxon	diagnosis	Diagnosis Length 2.4 mm. Body elongate, cylindrical, argillaceous gray. Head wide, downward bent; clypeus sinuate, anterior margin thickened. Pronotum with sides sharply declined; dorsally with two longitudinal and parallel carinae; lateral margins strongly sinuate before posterior margin, emargination forming a small tooth; posterior margin with strong marginal line, sinuate. Elytra with humerus projecting forwards, finely setaceous. Profemur broad with only two apical teeth. Meso- and metatibia dorso-ventrally flattened, gradually wider towards apical margin.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF04FF53FDDDFED4FB07FD59.taxon	discussion	Remarks Amerilochus is a monotypic genus with previously only a record from Peru (Skelley 2007 b)	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF04FF53FE6FF988FB7AF864.taxon	diagnosis	Diagnosis Length 2.3 – 4.0 mm. Body elongate, convex, piceous. Head broad, with strongly gibbous medially, coarsely punctate posteriorly and with short transverse wrinkles anterior margin of clypeus; clypeus sinuate, anterior margin thickened medially, downward bent with big acuminate dentiform process. Pronotum tumid anteriorly, laterally declined and variably punctate. Elytra with humeral denticle and with interstriae weakly convex; striae shallow, with coarse punctures separated by short transverse carina. Mesoventrite with V-shaped furrow parallel to lateral margin of mesocoxa.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF04FF53FE6FF988FB7AF864.taxon	discussion	Remarks Saprolochus has four recognized species reported from Bolivia, Guyana, Peru and Venezuela (Skelley 2007 b; Stebnicka & Galante 2007). For a key to all its species see Skelley (2007 b).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF05FF52FDEAFE95FEABFBA8.taxon	description	Figs 26 D, 54 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF05FF52FDEAFE95FEABFBA8.taxon	diagnosis	Diagnosis Length 3.5 – 5.7 mm. Head broad, strongly gibbous medially; coarsely punctate posteriorly; clypeus sinuate, anterior margin thickened medially, downwardly bent with big acuminate dentiform process. Pronotum punctate; finer punctures mixed with larger punctures; marginal line present; posterior angles with three dentiform processes. Elytra with interstriae weakly convex, shagreened; striae shallow, with somewhat quadrate punctures.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF05FF52FDEAFE95FEABFBA8.taxon	materials_examined	Material examined BRAZIL – Acre • 2 specs; Mâncio Lima, Parna Serra do Divisor; 7 ° 24 ′ 59 ″ S, 73 ° 37 ′ 56 ″ W; 6 – 10 Dec. 2019; R. A. Azevedo leg.; FIT; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF05FF52FDEAFE95FEABFBA8.taxon	description	Sexual dimorphism Male unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF05FF52FDEAFE95FEABFBA8.taxon	discussion	Collecting information Specimens were collected from rotten palm tree flowers (Skelley 2007 a) and with flight intercept traps in rainforests.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF05FF52FD9FFBDFFA9BF8BF.taxon	diagnosis	Diagnosis Length 2.8 – 3.0 mm. Body elongate, cylindrical, convex, piceous. Head punctate, with wrinkled area just after anterior margin; clypeal margin medially thickened and acuminate, downwardly bent; margin sinuate, lateral angles with group of denticles. Pronotum tumid anteriorly, punctate, with shallow and short posteromedial longitudinal furrow and two lateral fovea; sides sharply declined; lateral margin ending in obtuse teeth. Elytra with striae coarsely punctured; interstriae convex to carinate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF05FF52FD9FFBDFFA9BF8BF.taxon	discussion	Remarks Saprositellus has five recognized species. This genus has been recorded from Bolivia, Brazil, French Guiana, Guyana, Panama and Peru (Stebnicka 2003 d; Skelley 2007 b). For a key to all its species see Skelley (2007 b).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF05FF51FDC2F8AEFD06FC91.taxon	description	Figs 27 B, 55 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF05FF51FDC2F8AEFD06FC91.taxon	diagnosis	Diagnosis Length 2.8 – 3.0 mm. Head with coarse punctures; clypeus with angles lateral to emarginate area with six to seven denticle. Pronotum coarsely and densely punctate; without posterior marginal line. Elytra with small humeral tooth; striae coarsely punctured as wide as interstriae; interstriae convex. Posterior margin of meso- and metafemora lobed medially.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF05FF51FDC2F8AEFD06FC91.taxon	materials_examined	Material examined BRAZIL – Amazonas • 1 spec.; Coari, Rio Urucu; 4 ° 55 ′ 53 ″ S, 65 ° 18 ′ 13 ″ W; 25 Feb. – 10 Mar. 1995; Bührnheim et al. leg.; RUC- 36; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF05FF51FDC2F8AEFD06FC91.taxon	description	Sexual dimorphism Unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF05FF51FDC2F8AEFD06FC91.taxon	discussion	Collecting information Specimens were collected in the Amazon forest.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF06FF51FDFBFCD5FB9DF93E.taxon	description	Figs 27 A, 55 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF06FF51FDFBFCD5FB9DF93E.taxon	diagnosis	Diagnosis Length 2.6 – 2.8 mm. Head with coarse punctures; clypeus with lateral angles with six to seven denticles. Pronotum coarsely and densely punctate; with posterior marginal line. Elytra with small humeral tooth; striae wider than interstriae; interstriae carinate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF06FF51FDFBFCD5FB9DF93E.taxon	materials_examined	Material examined BRAZIL – Acre • 2 specs; Rio Branco; 10 ° 01 ′ 58 ″ S, 67 ° 42 ′ 13 ″ W; elev. 183 m; 12 Nov. 2015; M. Fazolin leg.; forest; CEMT. – Amazonas • 1 spec.; Iranbuda, Rio Solimões, Ilha de Marchantaria; 3 ° 15 ′ S, 59 ° 58 ′ W; 22 Jan. 1982; J. Adis leg.; várzea [lowland]; CEMT • 1 spec.; same data as for preceding; 4 Feb. 1982; CEMT • 5 specs; Manaus, Res. Adolpho Ducke; 2 ° 55 ′ 49 ″ S, 59 ° 58 ′ 32 ″ W; 28 Apr. – 3 May 2013; F. Z. Vaz-de-Mello leg.; CEMT. – Mato Grosso • 10 specs; Cotriguaçu, Fazenda São Nicolau; 9 ° 51 ′ 53 ″ S, 58 ° 13 ′ 13 ″ W; 3 Nov. 2017; F. Z. Vaz-de-Mello leg.; prainha; manually; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF06FF51FDFBFCD5FB9DF93E.taxon	description	Sexual dimorphism Males with fifth abdominal ventrite shorter than that of female; disc of pygidium longer than that of female.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF06FF51FDFBFCD5FB9DF93E.taxon	discussion	Collecting information Specimens were collected manually or with black light traps (Stebnicka 2003 d).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF06FF57FD93F929FDC3FEC7.taxon	diagnosis	Diagnosis Length 3.3 mm. Body piceous, oval, strongly convex, reddish brown. Head wider than elytra, finely punctate; clypeus inconspicuously sinuate, margin thickened medially. Pronotum strongly tumid anteriorly; disc densely and finely punctate; lateral margin explanate; posterior margin with rows of coarse, wide punctures, depressed and laterally compressed. Elytra oval, convex with small humeral tooth; interstriae weakly convex, striae deep. Mesoventrite coarsely punctured, with two sinuate furrows near lateral margins of mesocoxa.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF06FF57FD93F929FDC3FEC7.taxon	discussion	Remarks Stebnickiella is a monotypic genus previously recorded from Colombia and Guyana (Skelley 2007 b; Buriticá & Vaz-de-Mello 2016).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF00FF57FDE7FB57FAEEF96C.taxon	diagnosis	Diagnosis Length 1.6 – 2.2 mm. Body elongate, cylindrical, posteriorly dilate piceous with testaceous legs. Head granulated, clypeus emarginate; frontoclypeal suture distinct; frons variably punctured. Pronotum with at least two shallow transverse furrows and two transverse ridges, third transverse furrow and ridge may not be present or weakly developed; posteromedial longitudinal furrow present, dividing second and third transverse furrows and ridges. Elytra with interstriae carinate or flat. Metatarsomere with first tarsomere cylindrical, somewhat dilated posteriorly, shorter than longer apical spur.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF00FF57FDE7FB57FAEEF96C.taxon	discussion	Remarks Afrodiastictus has seven recognized species, but only two occur in the New World: A. mimicus (Gordin & Cartwright, 1980) and A. minutus (Gordon & Cartwright 1980; Pittino & Marini 1986; Gordon & Pittino 1992). This genus is recorded from Bolivia, Brazil and Venezuela (Gordon & Cartwright 1980). For a key to the Neotropical species see Gordon & Cartwright (1980) (as Rhyssemus Mulsant, 1842).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF00FF57FDE6FE06FBD3FBD1.taxon	description	Figs 26 E, 54 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF00FF57FDE6FE06FBD3FBD1.taxon	diagnosis	Diagnosis Same as for the genus.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF00FF57FDE6FE06FBD3FBD1.taxon	materials_examined	Material examined BRAZIL – Amazonas • 2 specs; Barcelos, Parna do Jaú; 1 ° 50 ′ 35 ″ S, 61 ° 34 ′ 07 ″ W; 18 Oct. – 2 Nov. 2021; R. A. Azevedo leg.; FIT; CEMT. COLOMBIA – Vaupés • 1 spec.; R. N. Mosiro-Itajura (Caparú), Centro Ambiental; 1 ° 04 ′ S, 69 ° 31 ′ W; 20 Jan. – 1 Feb. 2003; M. Shakoy and D. Arias leg.; FIT; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF00FF57FDE6FE06FBD3FBD1.taxon	description	Sexual dimorphism Male unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF00FF57FDE6FE06FBD3FBD1.taxon	discussion	Collecting information Specimens were collected with flight intercept traps in tropical rainforests.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF00FF56FDF4F898FCECFCF8.taxon	description	Figs 28 A, 55 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF00FF56FDF4F898FCECFCF8.taxon	diagnosis	Diagnosis Length 1.6 – 1.8 mm. Head with clypeus sinuate, tuberculate; frontoclypeal suture grooved; frons coarsely punctured. Pronotum with two distinct transverse furrows and ridges, third furrow if present, weakly developed, present as a line of coarse confluent punctures; posteromedial longitudinal furrow present cutting in half the second furrow.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF00FF56FDF4F898FCECFCF8.taxon	materials_examined	Material examined BRAZIL – Acre • 13 specs; Rio Branco; 10 ° 01 ′ 58 ″ S, 67 ° 47 ′ 13 ″ W; elev. 183 m; 12 Jul. 2015; M. Fazolin leg.; forest; CEMT. – Roraima • 1 spec.; Amajari, EsEc Maracá, Uranicoera River; 3 ° 20 ′ 59 ″ N, 61 ° 25 ′ 22 ″ W; elev. 83 m; 24 Mar. 2016; A. P. M. Santos et al. leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF00FF56FDF4F898FCECFCF8.taxon	description	Sexual dimorphism Male and female indistinguishable from each other.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF00FF56FDF4F898FCECFCF8.taxon	discussion	Collecting information Specimens were at altitudes from 83 to 183 metres.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF01FF56FDEDFCEFFA85F85B.taxon	diagnosis	Diagnosis Length 2.5 – 3.8 mm. Body short, robust, dilated posteriorly. Head granulate; clypeus emarginate, not denticulate. Pronotum with at least one vestigial transversal furrow behind eyes. First tarsomere of metatarsus posteriorly dilated, somewhat triangularly shaped.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF01FF56FDEDFCEFFA85F85B.taxon	discussion	Remarks Leiopsammodius currently has 49 recognized species but only 14 are found in the New World (Rakovic 1990; Gordon & Pittino 1992; Harpootlian et al. 2000; Schoolmeesters 2023). The genus has a worldwide distribution (Rakovic 1990), in the New World this taxon has been recorded from Argentina, Bolivia, Brazil, Chile, Dominica, Guadeloupe, Paraguay and the USA (Rakovic 1990; Harpootlian et al. 2000). Identification keys for species of Leiopsammodius can be found in Cartwright (1955) (as Psammodius Fallén, 1807), Rakovic (1990) and Harpootlian et al. (2000).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF02FF55FDC9FE95FDCEFB8B.taxon	description	Fig. 55 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF02FF55FDC9FE95FDCEFB8B.taxon	diagnosis	Diagnosis Length 3.5 mm. Body piceous. Clypeus emarginate; gena notched at genal suture. Pronotum without transverse furrows; lateral margins without fringe of setae. Elytra without humeral tooth; striae strongly impressed. Protibiae with posterior outer teeth reduced, the middle teeth normally developed, the apical tooth extremely large, almost straight forward.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF02FF55FDC9FE95FDCEFB8B.taxon	description	Sexual dimorphism Unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF02FF55FDC9FE95FDCEFB8B.taxon	discussion	Collecting information No information available in literature. Remarks This species was originally described from Serra de Paranapiacaba, São Paulo State, Brazil (Petrovitz 1972). Specimens unavailable for analysis, thus diagnosis and positioning in the dichotomous key were based on the available literature.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF02FF55FDD6FBC2FE04F800.taxon	description	Fig. 55 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF02FF55FDD6FBC2FE04F800.taxon	diagnosis	Diagnosis Length 2.5 mm. Body ferruginous. Head with frons coarsely punctured; clypeus emarginate, without teeth; gena fimbriate. Pronotum with posterior angles obtusely rounded; sides not fimbriate nor crenulate; surface with three transverse furrows and a medial longitudinal furrow. Elytra with fine humeral tooth, striae somewhat shallow; punctures fine, weakly crenating interstriae. Abdominal ventrites crenulate anteriorly.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF02FF55FDD6FBC2FE04F800.taxon	description	Sexual dimorphism Unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF02FF55FDD6FBC2FE04F800.taxon	discussion	Collecting information No information available in literature. Remarks This species was originally described from Manaus, Amazonas State, Brazil (Cartwright 1955). Specimens unavailable for analysis, thus diagnosis and positioning in the dichotomous key were based on the available literature.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0CFF5BFDD9FE95FD87FA69.taxon	description	Figs 29 E – F, 55 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0CFF5BFDD9FE95FD87FA69.taxon	diagnosis	Diagnosis Length 3.8 mm Body robust, compact, piceous. Head granulate; clypeus emarginate. Pronotum with vestigial transverse furrows right behind eyes; dorsal surface everywhere densely punctured by punctures moderate to coarse.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0CFF5BFDD9FE95FD87FA69.taxon	materials_examined	Material examined BRAZIL – Distrito Federal • 4 specs; Planaltina, Embrapa CPAC; 15 ° 36 ′ 15 ″ S, 47 ° 43 ′ 09 ″ W; elev. 1030 m; 3 Oct. 2013; C. M. Oliveira leg.; molasses, alcohol and pineapple; CEMT • 1 spec.; same data as for preceding; 18 Nov. 2013; CEMT • 2 specs; Reserva Ecológica IBGE; 2009; R. Japiassu leg.; CEMT. – Mato Grosso • 1 spec.; Chapada dos Guimarães, Condomínio Ecoville; 15 ° 12 ′ 14 ″ S, 55 ° 55 ′ 38 ″ W; 23 Feb. 2019; A. Mota and Leon leg.; Cerrado; manually; CEMT • 6 specs; Cuiabá, Estância Leôncio; 15 ° 11 ′ 11 ″ S, 55 ° 58 ′ 00 ″ W; 1 Nov. 2020; A. Mota and Leon leg.; CEMT. – Minas Gerais • 1 spec.; Ingaí; Jan. 2003; S. A Falqueto leg.; CEMT • 1 spec.; Ingaí-Lavras, prox. Poço Bonito; Nov. 2002; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Ingaí, prox. Poço Bonito (Lavras); 1 Nov. 2002; S. A. Falqueto leg.; bovine feces; CEMT. – Paraná • 1 spec.; Castro, Estra. Castro-Tibagi, Km 15; elev. 1200 m; Oct. 2005; Grossi and Parizott leg.; CEMT. – São Paulo • 1 spec.; Descalvado, Fazenda Itaúnas; 19 Oct. 2006; N. W. Perioto leg.; forest + Citrus plantation; pitfall; CEMT • 2 specs; Luis Antonio, E. E. de Jataí; 20 – 23 Nov. 2008; S. P. Rosa and M. Ladenthin leg.; CEMT. PARAGUAY – Amambay • 1 spec.; Pedro Juan Caballero, Facultad de Ciencias Agrarias; 5 Nov. 1994; M. A. R. Díaz leg.; pasture area; bovine feces; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0CFF5BFDD9FE95FD87FA69.taxon	description	Sexual dimorphism Unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0CFF5BFDD9FE95FD87FA69.taxon	discussion	Collecting information Specimens were collected within bovine faecal masses and fermented fruits (pineapple). This species occurs at altitudes up to 1500 metres.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0CFF59FDDDF99CFE56FDB4.taxon	description	Fig. 55 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0CFF59FDDDF99CFE56FDB4.taxon	diagnosis	Diagnosis Length 2.5 mm. Body ferruginous. Head emarginate; frons with three to four coarse punctures on each side near frontoclypeal. Pronotum with three transverse furrows. Elytra humeral region finely denticulate; lateral margin not fringed; striae wide, deep with moderate punctures, strongly crenate interstriae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0CFF59FDDDF99CFE56FDB4.taxon	description	Sexual dimorphism Unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0CFF59FDDDF99CFE56FDB4.taxon	discussion	Collecting information No information available in literature. Remarks This species was originally described from Santarém, Pará, Brazil (Cartwright 1955). Specimens unavailable for analysis, thus diagnosis and positioning in the dichotomous key were based on the available literature.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0EFF59FDCCFDB3FBA5FAED.taxon	description	Fig. 55 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0EFF59FDCCFDB3FBA5FAED.taxon	diagnosis	Diagnosis Length 3.1 mm. Head with clypeus emarginate; frontoclypeal suture indistinct; pronotum not punctate. Pronotum with three transverse furrows; transverse ridges flat and wine with fine and scattered punctures; lateral margins finely fimbriate. Elytra with humeral region covered with few long and fine setae; interstriae strongly convex smooth and not punctate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0EFF59FDCCFDB3FBA5FAED.taxon	description	Sexual dimorphism Unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0EFF59FDCCFDB3FBA5FAED.taxon	discussion	Collecting information No information available in literature. Remarks This species was originally described from Juazeiro farm, now city of Juazeirinho, near Soledade City, Paraiba State, Brazil (Petrovitz 1961 b; IBGE 1990 b). Specimens unavailable for analysis, thus diagnosis and positioning in the dichotomous key were based on the available literature.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0EFF59FE31FA1BFE6DF82A.taxon	diagnosis	Diagnosis Length 3.75 mm. Body elongate, oval, strongly convex, piceous to light piceous. Head granulate; emarginate, lateral angles denticulate; frontoclypeal suture not visible. Pronotum dorsal surface variably punctate; fringed with setae; without evident transverse furrows or ridges except for fine furrow right behind the eyes. Elytra with fine striae; interstriae weakly convex. Metatibiae broad, robust. First metatarsomere widened apically, triangular, shorter than great apical spur and as long as the following three tarsomeres combined.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0EFF59FE31FA1BFE6DF82A.taxon	discussion	Remarks Odontopsammodius has 12 recognized species (Gordon & Pittino 1992) recorded from Argentina, Bolivia, Brazil, Cuba, Dominica, Guadeloupe, Puerto Rico, Mexico and the USA (Rakovic 1990). For keys to all its species see Cartwright (1955) (as Psammodius) and Rakovic (1990) (as Leiopsammodius).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0FFF58FDD6FBFCFA86F8DF.taxon	diagnosis	Diagnosis Length 1.7 – 2.1 mm. Body elongate, broadened posteriorly, dark testaceous. Head with clypeus granulate and sinuate medially; frontoclypeal suture visible, grooved. Pronotum with five transverse ridges; posteromedial longitudinal furrow present. Elytra with interstriae alternately carinate. First metatarsomere cylindrical, somewhat dilated posteriorly.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0FFF58FDD6FBFCFA86F8DF.taxon	discussion	Remarks Neorhyssemus has four recognized species reported from Brazil and Guyana (Gordon & Cartwright 1980). For key to all its species see Gordon & Cartwright (1980) (as Rhyssemus).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0FFF58FE20FE95FDADFB8A.taxon	description	Fig. 56 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0FFF58FE20FE95FDADFB8A.taxon	diagnosis	Diagnosis Length 3.75 mm. Body piceous. Head granulate; frons smooth. Pronotum quite densely punctured; punctures smaller near anterior margin and sides; sides crenate, fringed with setae; posteriorly without marginal line. Elytra with interstriae smooth, flat, striae fine; margins not fimbriate.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0FFF58FE20FE95FDADFB8A.taxon	description	Sexual dimorphism Unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0FFF58FE20FE95FDADFB8A.taxon	discussion	Collecting information No information available in literature. Remarks This species was originally described from Blumenau, Santa Catarina State, Brazil. Specimens unavailable for analysis, diagnosis and positioning in the dichotomous key were based on the available literature (Gordon & Pittino 1992).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF08FF5FFE16FEB6FE39FBEF.taxon	description	Fig. 56 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF08FF5FFE16FEB6FE39FBEF.taxon	diagnosis	Diagnosis Length 1.7 – 2.0 mm. Body somewhat short. Head tuberculate; tubercles sharp; clypeus emarginate; frontoclypeal suture grooved. Pronotum with four transversal ridges interrupted medially, only first ridge complete. Elytral margin rounded.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF08FF5FFE16FEB6FE39FBEF.taxon	description	Sexual dimorphism Male and female indistinguishable from each other.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF08FF5FFE16FEB6FE39FBEF.taxon	discussion	Collecting information Specimens were collected in the Amazon rainforest; collecting methodology unavailable in the consulted literature or analyzed material. Remarks This species was originally described from Jacaré, Xingu National Park, Mato Grosso State, Brazil. Specimens unavailable for analysis, diagnosis and positioning in the dichotomous key were based on the available literature.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF08FF5EFE01FB1CFDD3FEC1.taxon	description	Figs 28 C, 56 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF08FF5EFE01FB1CFDD3FEC1.taxon	diagnosis	Diagnosis Length 1.7 – 2.0 mm. Head tuberculate; clypeus emarginate; frontoclypeal suture furrowed, sinuate. Pronotum with second transverse ridge interrupted medially by longitudinal furrow; longitudinal ridges not joined anteriorly. Elytra with costate interstriae, costae rounded.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF08FF5EFE01FB1CFDD3FEC1.taxon	materials_examined	Material examined BRAZIL – Amazonas • 1 spec.; Manaus, Taruma-Mirim nr Manaus; 3 Mar. 1982; CEMT • 1 spec.; same data as for preceding; 18 Aug. 1981; CEMT • 1 spec.; same data as for preceding; 3 ° 02 ′ S, 60 ° 17 ′ W; 25 Aug. 1982; A. Rodrigues leg.; secondary forest; CEMT • 1 spec.; same data as for preceding; 26 May 1983; CEMT • 2 specs; same data as for preceding; 27 Jun. 1983; CEMT • 1 spec.; Manaus, Rio Tarumã Mirim; 3 ° 46 ′ S, 60 ° 33 ′ W; 13 Mar. 1998; S. Golovatch leg.; igapó [flooded forest]; CEMT • 1 spec.; Manaus, Rio Tarumã Mirim-igapó [flooded forest]; 1 Jun. 1976; J. Adis leg.; INPA • 1 spec.; same data as for preceding; 8 Oct. 1976; INPA • 1 spec.; same data as for preceding; 15 Dec. 1976; INPA • 1 spec.; same data as for preceding; 3 ° 02 ′ S, 60 ° 17 ′ W; 26 May 1983; CEMT • 1 spec.; same data as for preceding; 27 Jun. 1983; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF08FF5EFE01FB1CFDD3FEC1.taxon	description	Sexual dimorphism Male and female indistinguishable from each other.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF08FF5EFE01FB1CFDD3FEC1.taxon	discussion	Collecting information Specimens were collected in the Amazon rainforest; collecting methods not available in the consulted literature or analyzed material.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF09FF5EFE25FE05FD5AFAF3.taxon	description	Figs 28 B, 56 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF09FF5EFE25FE05FD5AFAF3.taxon	diagnosis	Diagnosis Length 1.9 – 2.1 mm. Head with clypeus emarginate; frontoclypeal suture furrowed; sinuate. Pronotal transverse ridges widely separated with second transverse ridge not interrupted medially; longitudinal ridges joined among them anteriorly. Elytra with thick costae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF09FF5EFE25FE05FD5AFAF3.taxon	materials_examined	Material examined BRAZIL – Bahia • 1 spec.; Caravelas; Mar. 1996; R. Diniz leg.; CEMT. – Maranhão • 8 specs; Carolina, Fazenda Cincorá; 17 – 22 Oct. 2009; F. Limeira-de-Oliveira, R. O. Souza and M. B. Aguiar Neto leg.; CEMT • 2 specs; Pedrinhas, São Luís Island; 17 Aug. 1984; E. C. Bergmann leg.; blacklight; CEMT. – Mato Grosso • 2 specs; Chapada dos Guimarães; 25 Mar. 2001; A. D. Ramos leg.; light trap; CEMT • 3 specs; same data as for preceding; 15 ° 24 ′ 31 ″ S, 55 ° 43 ′ 53 ″ W; 2 Dec. 2015; A. Frolov and L. Akhmetova leg.; CEMT • 8 specs; Diamantino, Alto Rio Arinos; Oct. 2001; E. Furtado leg.; CEMT. – Rondônia • 4 specs; Fazenda Rancho Grande, 62 km S of Ariquemes; Feb. 1997; Vaz-de-Mello and Vulinec leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF09FF5EFE25FE05FD5AFAF3.taxon	description	Sexual dimorphism Unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF09FF5EFE25FE05FD5AFAF3.taxon	discussion	Collecting information Specimens were collected with light traps.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF09FF5DFD89FAFBFA82FDC3.taxon	diagnosis	Diagnosis Length 1.8 – 3.3. Body elongate, convex, dilated posteriorly, testaceous to piceous. Head granulate; clypeus emarginate; frontoclypeal suture evident or not. Pronotum with vestiges of transverse furrows and ridges; transverse furrows normally only visible right behind eyes, evidenced by line of punctures; posteromedial longitudinal furrow evident. Elytra with striae deep; interstriae flat or weakly convex. Tarsus cylindrical.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF09FF5DFD89FAFBFA82FDC3.taxon	discussion	Remarks Platytomus is a cosmopolitan genus with 29 recognized species (Schoolmasters 2023) and at least 12 are found in the new world (Gordon & Pittino 1992; Rakovic et al. 2020 b). A key for some of the species of Platytomus in the New World was written by Cartwright (1948).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0AFF5DFE28FD08FC8CF9A7.taxon	description	Figs 29 A – B, 56 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0AFF5DFE28FD08FC8CF9A7.taxon	diagnosis	Diagnosis Length 2.1 – 2.4 mm. Body piceous. Head granulate; granules reaching anterior half of clypeus; clypeus emarginate; frontoclypeal suture finely impressed. Pronotum with coarse and fine punctures; coarse punctures scattered. Elytra with striae deep; interstriae convex. Metaventrite with posterior transverse furrows obtusely arched. Abdominal ventrites medially finely punctured; coarse punctures restricted to lateral margins. Pygidium with only two setae near uro-genital opening.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0AFF5DFE28FD08FC8CF9A7.taxon	materials_examined	Material examined BRAZIL – Bahia • 1 spec.; Lençóis Ribeirão de Baixo; 12 ° 35 ′ 12 ″ S, 41 ° 22 ′ 56 ″ W; elev. 340 m; 4 Jun. 2007; J. A. Rafael and F. F. Xavier leg.; light; CEMT. – Distrito Federal • 2 specs; Brasília, Fazenda Água (UNB); Sep. 2010; C. Suinaga leg.; CEMT • 3 specs; Brasília; elev. 1100 m; Jan. 2001; N. Degallier leg.; light; CEMT • 2 specs; same data as for preceding; Feb. 2001; CEMT • 1 spec.; same data as for preceding; Mar. 2001; CEMT • 1 spec.; Planaltina, Embrapa Cerrados; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 24 Mar. 2006; C. Oliveira leg.; agricultural area; light; CEMT • 3 specs; same data as for preceding; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 7 Apr. 2006; C. Oliveira leg.; agricultural area; light; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0AFF5DFE28FD08FC8CF9A7.taxon	description	Sexual dimorphism Females with sixth abdominal ventrite longer than males.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0AFF5DFE28FD08FC8CF9A7.taxon	discussion	Collecting information Specimens were collected at light at altitudes up to 1100 m.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0AFF5CFE07F9A4FC0BFC28.taxon	description	Figs 29 C – D, 56 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0AFF5CFE07F9A4FC0BFC28.taxon	diagnosis	Diagnosis Length 2.2 – 2.3 mm. Body piceous. Head with weakly elevated granules; granules restricted to anterior third of clypeus; clypeus emarginate; frontoclypeal suture fine. Pronotum densely punctured by fine and coarse punctures. Elytra with striae deep; interstriae weakly convex. Metaventrite with posterior transverse furrows arched. Abdominal ventrites with median row of coarse punctures. Pygidium with only two setae near uro-genital opening.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0AFF5CFE07F9A4FC0BFC28.taxon	materials_examined	Material examined BRAZIL – Mato Grosso • 1 spec.; Sesc-Pantanal; 1 Nov. 2001; Malaise trap; CEMT • 1 spec.; same data as for preceding; 15 Apr. 2002; light; CEMT • 1 spec.; same data as for preceding; 5 Jul. 2003; CEMT • 1 spec.; same data as for preceding; 9 Aug. 2002; CEMT • 1 spec.; same data as for preceding; 7 Jul. 2004; CEMT • 8 specs; Cotriguaçu, Fazenda São Nicolau; 20 Aug. 2018; R. Stofel leg.; CEMT • 10 specs; same data as for preceding; 9 ° 49 ′ 09 ″ S, 58 ° 15 ′ 31 ″ W; 21 Oct. 2018; V. Costa-Silva leg.; CEMT • 45 specs; same data as for preceding; 9 ° 51 ′ 52 ″ S, 58 ° 13 ′ 45 ″ W; 2 Oct. 2017; Vaz-de-Mello et al. leg.; CEMT • 1 spec.; Cuiabá, Fazenda Mutuca; 15 ° 19 ′ 31 ″ S, 55 ° 58 ′ 03 ″ W; elev. 285 m; Nov. 2015; R. V. Nunes and A. Frolov leg.; light; CEMT • 1 spec.; Juína, Distrito de Fontanillas; 17 May 2012; S. E. Silva leg.; FIT; CEMT • 4 specs; Tangará da Serra; 14 ° 38 ′ 33 ″ S, 57 ° 33 ′ 18 ″ W; 19 – 22 Feb. 2012; A. S. Tissiani leg.; CEMT. – Minas Gerais • 1 spec.; Diamantina; 23 Feb. – 3 Mar. 2001; N. Degallier leg.; CEMT. – Paraná • 1 spec.; Telêmaco Borda, Klabin Papel e Celulose; 13 Oct. 2006; C. A. H. Flechtmann leg.; Pinus taeda; multiple funnel trap baited with sulcato and pinene; FIT; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0AFF5CFE07F9A4FC0BFC28.taxon	description	Sexual dimorphism Female with sixth abdominal ventrite longer than that of males.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0AFF5CFE07F9A4FC0BFC28.taxon	discussion	Collecting information Specimens were collected with Malaise, light and flight intercept traps.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0BFF5CFF61FAD8FC56F95A.taxon	materials_examined	ARGENTINA – Formosa • 1 spec.; Laguna Yema, Reserva Biosfera Teuquito; 24 ° 16 ′ 02 ″ S, 61 ° 14 ′ 65 ″ W; 11 Dec. 2008; F. C. Ocampo leg.; CEMT. BRAZIL – Amazonas 1 spec.; Castanho-Careiro, AM- 359, Km 39; 3 ° 43 ′ 59 ″ S, 60 ° 20 ′ 09 ″ W; 6 – 7 Nov. 2010; J. A. Rafael et al. leg.; light; CEMT. – Mato Grosso do Sul • 1 spec.; Selvíra, UNESP Farm; 7 Nov. 1991; C. A. H. Flechtmann leg.; Brachiaria decumbens pasture; CEMT. – Mato Grosso • 4 specs; Cotriguaçu, Fazenda São Nicolau, Airport; 9 ° 51 ′ 52 ″ S, 58 ° 13 ′ 45 ″ W; 2 Oct. 2017; F. Z. Vaz-de-Mello et al. leg.; light; CEMT. – Rio Grande do Sul • 1 spec.; Santa Cruz do Sul, Cerro Alegre; 29 ° 48 ′ 05 ″ S, 52 ° 19 ′ 39 ″ W; 11 Feb. 2012; R. Bohn leg.; bovine feces CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0BFF5CFDBBFC5DFB9CFB5C.taxon	description	Figs 28 F – G, 57 A	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0BFF5CFDBBFC5DFB9CFB5C.taxon	diagnosis	Diagnosis Length 2.0 – 2.4 mm. Body light to dark testaceous. Head granulate; clypeus with its margins elevated; frontoclypeal suture fine. Pronotum with punctures of two sizes, fine and coarse. Elytra with medial darkened area near lateral margins; striae deep; interstriae flat. Pygidium with 5 – 7 long setae near uro-genital opening. Metaventrite with longitudinal furrow obsolete anteriorly. First metatarsomere cylindrical, dilated apically shorter than the following two tarsomere combined.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0BFF43FF61F88DFD25F837.taxon	materials_examined	BRAZIL – Amazonas • 4 specs; Castanho-Careiro, AM- 359 Km 39; 3 ° 43 ′ 59 ″ S, 60 ° 20 ′ 09 ″ W; 6 – 7 Nov. 2010; J. A. Rafael et al. leg.; light; CEMT • 1 spec.; Manaus, 24 km of Embrapa-CPAA; 20 Aug. 1993; A. Pamplona leg.; light; CEMT. – Distrito Federal • 1 spec.; Brasília, Plano Piloto; 31 May 1958; Bello leg.; CEMT • 3 specs; Brasília, Embrapa Cerrados; 15 ° 35 ′ 54 ″ S, 47 ° 42 ′ 51 ″ W; 24 Mar. 2006; C. Oliveira leg.; agricultural area; light; CEMT. – Espírito Santo • 1 spec.; Linhares, Res. Vale Rio Doce; 19 ° 14 ′ 46 ″ S, 39 ° 57 ′ 40 ″ W; elev. 12 m; 5 – 6 May 2007; J. A. Rafael and F. Xavier leg.; light; CEMT. – Mato Grosso • 3 specs; Chapada dos Guimarães; 27 Apr. 2001; A. F. Ramos leg.; light; CEMT • 4 specs; same data as for preceding; 28 Apr. 2001; CEMT • 1 spec.; Cotriguaçu, Fazenda São Nicolau; 16 Jan. 2008; O. Peres-Filho leg.; light; CEMT • 2 specs; same data as for preceding, airport; 9 ° 51 ′ 52 ″ S, 58 ° 13 ′ 45 ″ W; 2 Oct. 2017; F. Z. Vaz-de-Mello et al. leg.; light; CEMT • 1 spec.; Cuiabá; 15 ° 16 ′ 45 ″ S, 56 ° 00 ′ 06 ″ W; 7 Jul. 2015; A. Frolov and L. Ahkmetova leg.; CEMT • 1 spec.; Sesc Pantanal; 15 Sep. 2004; light; CEMT • 1 spec.; Tangará da Serra, Fazenda Três Rios; 14 ° 21 ′ S, 57 ° 42 ′ W; elev. 592 m; Feb. 2008; N. Degallier leg.; Cerrado; pitfall; CEMT. – Mato Grosso do Sul • 1 spec.; Corumbá, Fazenda São Bento, Prox. Porto Jofre; 14 Apr. 2011; R. V. Nunes leg.; light; CEMT • 3 specs; Selvíra, UNESP Farm; 1 Oct. 1992; C. A. H. Flechtmann leg.; Brachiaria decumbens pasture; CEMT • 2 specs; same data as for preceding; 7 Nov. 1991; CEMT. – Minas Gerais • 2 specs; Cordisburgo; Dec. 1997; F. Z. Vaz-de-Mello leg.; CEMT • 1 spec.; Viçosa; May 1998; F. Z. Vaz-de-Mello leg.; CEMT. – Paraná • 3 specs; Guaratuba, Portal do Itararé; elev. 900 m; 15 Apr. 2005; Grossi and Caron leg.; light; CEMT • 1 spec.; Icaraíma, Estância Felipe; 23 ° 20 ′ 32 ″ S, 53 ° 39 ′ 09 ″ W; elev. 300 m; 2019; Felipe and Rodrigues leg.; CEMT. – Pernambuco • 1 spec.; Afrânio; 8 ° 31 ′ 53 ″ S, 41 ° 02 ′ 59 ″ W; elev. 550 m; 16 May 2007; J. A. Rafael and F. Xavier leg.; light; CEMT. – Rio Grande do Sul • 5 specs; Bagé; 31 ° 21 ′ 05 ″ S, 54 ° 01 ′ 13 ″ W; elev. 232 m; 4 Jan. 2016; R. N. Sisti et al. leg.; forest; CEMT • 1 spec.; Santa Cruz do Sul, Cerro Alegre; 29 ° 48 ′ 11 ″ S, 52 ° 19 ′ 51 ″ W; 21 Jan. 2012; R. Bohn leg.; annual crop; bovine feces; CEMT • 1 spec.; same data as for preceding; 29 ° 48 ′ 18 ″ S, 52 ° 19 ′ 55 ″ W; CEMT • 8 specs; same data as for preceding; 29 ° 48 ′ 18 ″ S, 52 ° 19 ′ 55 ″ W; 21 Jan. 2012; open area; CEMT • 1 spec.; same data as for preceding; 29 ° 48 ′ 05 ″ S, 52 ° 19 ′ 39 ″ W; 11 Feb. 2012; CEMT. – Rondônia • 2 specs; 62 km S of Ariquemes, Fazenda Rancho Grande; Feb. 1997; F. Z. Vaz-de-Mello and Vulinec leg.; CEMT • 1 spec.; Porto Velho, Rio das Graças, Bom Jesus; 8 ° 49 ′ 47 ″ S, 63 ° 46 ′ 51 ″ W; 22 Oct. 2017; D. C. Santos and L. Silvia leg.; FIT; CEMT. – Santa Catarina • 1 spec.; Florianópolis, Lagoa do Peri; 27 ° 43 ′ S, 48 ° 32 ′ W; Feb. 2009; A. Martins and M. Hernández leg.; Atlantic forest; CEMT. – São Paulo • 1 spec.; Anhembi; ESALQ / USP; E. N. L. Ferreira leg.; CEMT • 8 specs; Anhembi, ESALQ / USP, Petrobras; 22 ° 42 ′ S, 47 ° 39 ′ W; 2 Apr. 2013; E. N. L. Ferreira leg.; light; CEMT • 6 specs; Anhembi, ESALQ / USP; 22 ° 43 ′ S, 48 ° 10 ′ W; 2 Apr. 2013; E. N. L. Ferreira leg.; E. urophylla; light; CEMT • 1 spec.; Anhembi, ESALQ / USP, R. Legal; 22 ° 43 ′ S, 48 ° 10 ′ W; E. N. L. Ferreira leg.; light; CEMT • 2 specs; Anhembi, ESALQ / USP; 22 ° 43 ′ S, 49 ° 11 ′ W; 2 Apr. 2013; E. N. L. Ferreira leg.; E. grandis; light; CEMT. UNITED STATES OF AMERICA – Mississippi • 4 specs; Lafayette, Oxford; 8 Sep. 1995; P. K. Lago leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0BFF43FF61F88DFD25F837.taxon	description	Sexual dimorphism Females with meso- and metatarsus shorter than tibiae; abdomen is more convex than in males and disc of pygidium is wider and shorter compared with that of males.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF0BFF43FF61F88DFD25F837.taxon	discussion	Collecting information Specimens have been collected with light traps in pastures of Urochloa decumbens, Cerrado regions and in tropical rainforests. Remarks Platytomus longulus and Platytomus freudei are here treated as a complex of species due to the difficulties encountered when trying to distinguish them.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF15FF42FDEDFE95FCEEF9EC.taxon	description	Figs 28 D – E, 56 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF15FF42FDEDFE95FCEEF9EC.taxon	diagnosis	Diagnosis Length 1.8 – 3.0 mm. Body brown. Head with clypeus emarginate; margin not upturned; frontoclypeal suture not visible. Pronotum with punctures of two sizes, fine and coarse. Elytra with striae deep; interstriae weakly convex. Metaventrite furrow complete. First metatarsomere cylindrical, dilated apically shorter than the following two tarsomere combined.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF15FF42FDEDFE95FCEEF9EC.taxon	materials_examined	Material examined BRAZIL – Mato Grosso • 1 spec.; Cáceres, Fazenda Baia de Pedra; 16 ° 29 ′ 30 ″ S, 58 ° 08 ′ 55 ″ W; Mar. 2009; D. D. Pinheiro leg.; CEMT • 1 spec.; Cuiabá, Morada do Ouro; 15 ° 34 ′ 02 ″ S, 56 ° 03 ′ 25 ″ W; May – Nov. 2021; L. A. C. S. Santos leg.; CEMT • 1 spec.; Poconé, Pousada das Araras; 3 Dec. 1998; J. L Moreno and T. Mestre leg.; Pantanal; CEMT • 1 spec.; Sesc Pantanal; 11 Jun. 2002; light; CEMT • 1 spec.; same data as for preceding; 14 May 2002; CEMT • 35 specs; same data as for preceding; 15 Sep. 2004; CEMT • 1 spec.; same data as for preceding; 16 Sep. 2004: CEMT • 2 specs; same data as for preceding; 17 Jul. 2004; CEMT. – Pernambuco • 2 specs; Fernando de Noronha, Sancho; 20 – 28 Feb. 2020; Rafael and Limeira-de-Oliveira leg.; light; CEMT. – Rio Grande do Norte • 1 spec.; Baía Formosa, Mata Estrela; 6 ° 23 ′ 19 ″ S, 35 ° 01 ′ 05 ″ W; elev. 44 m; 24 May 2007 J. A. Rafael and M. Xavier leg.; light; CEMT. GRENADA – St Andrew • 3 specs; Mirabeau, Agri. Lab.; 12 Sep. 1990; J. Telesford leg.; light; CEMT • 1 spec.; same data as for preceding; 20 Sep. 1990; CEMT. PARAGUAY – Presidente Hayes • 1 spec.; Cruce de los Pioneros; 11 Jan. 1999; U. Peña leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF15FF42FDEDFE95FCEEF9EC.taxon	description	Sexual dimorphism Unknown. Collecting information Specimens commonly attracted to light.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF15FF42FDEDFE95FCEEF9EC.taxon	discussion	Remarks Platytomus parvulus was tentatively reported for the first time in Brazil on Fernando de Noronha island by Rafael et al. (2020), we here confirm this report.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF15FF42FDC7F919FA8CF86E.taxon	diagnosis	Diagnosis Length 3.0 – 3.8 mm. Body elongate, oval, covered with setae, dark reddish brown or piceous. Head with clypeus granulate. Pronotum without vestiges of transverse furrows, covered with setiferous alveolate punctures. Elytra with interstriae covered with setaceous tubercles.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF15FF42FDC7F919FA8CF86E.taxon	discussion	Remarks Trichiopsammobius has two recognized species and is recorded from Brazil, Jamaica, Paraguay and Venezuela (Rakovic & Mencl 2013). A key for all its species can be found in Rakovic & Mencl (2013).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF16FF41FE20FE95FA83F821.taxon	description	Figs 29 G, 57 B	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF16FF41FE20FE95FA83F821.taxon	diagnosis	Diagnosis Length 3.6 – 3.8 mm. Head with clypeus entirely granulate; clypeus sinuate; lateral angles with small upturned teeth; frons with punctures smaller than those of pronotum, grouped. Pronotum without transverse ridges; densely punctured by setiferous punctures. Elytra with interstriae covered with setaceous tubercles.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF16FF41FE20FE95FA83F821.taxon	materials_examined	Material examined BRAZIL – Alagoas • 1 spec.; Olho d’Agua do Casado, Fazenda Talhado; 21 Dec. 2000; L. Iannuzzi leg.; CEMT. – Bahia • 8 spec.; Barreiras; Jan. 1991: CEMT • 5 specs; same data as for preceding; Oct. 1991; J. C. Zanuncio leg.; light; CEMT • 1 spec.; 150 km W of Barreiras; Jan. 2003; P. A. Schmidt leg.; CEMT. – Goiás • 3 specs; Rio Verde; Nov. 1995; J. Carlos leg.; CEMT. – Maranhão • 26 specs; Carolina, Fazenda Cincorá; 17 – 22 Oct. 2009; F. Limeira-de-Oliveira, R. O. Souza and M. B. A. Neto leg.; light; CEMT • 5 specs; Imperatriz; Jan. 1998; CEMT • 2 specs; Mirador, P. E. Mirador, Base Geraldina; 27 Nov. 2008; Aguiar-Neto and A. L. Costa leg.; light; CEMT • 1 spec.; São João do Paraíso; 6 ° 32 ′ 14 ″ S, 46 ° 54 ′ 22 ″ W; 12 Jan. 2021; M. C. Batista leg.; human feces; CEMT. – Mato Grosso • 1 spec.; Chapada dos Guimarães, F. Sadia; 15 ° 18 ′ 51 ″ S, 55 ° 58 ′ 20 ″ W; 1 Feb. 2013; G. M. Daniel leg.; pitfall; CEMT • 7 specs; Chapada dos Guimarães, Fazenda Jardim; 29 Sep. 2012; V. C. Jorge leg.; CEMT • 4 specs; same data as for preceding; 13 Oct. 2012; CEMT • 11 specs; same data as for preceding; 10 Nov. 2012; CEMT • 1 spec.; same data as for preceding; 27 Nov. 2012; CEMT • 1 spec.; same data as for preceding; 15 Jun. 2013; CEMT • 1 spec.; Chapada dos Guimarães, Rio Claro; 15 ° 19 ′ 15 ″ S, 55 ° 52 ′ 51 ″ W; elev. 293 m; 5 Dec. 2012; M. A. Carvalho; CEMT • 2 specs; Chapada dos Guimarães, Santuário dos Elefantes; 15 ° 18 ′ 55 ″ S, 55 ° 24 ′ 14 ″ W; 23 Jan. 2020; H. Sehn et al. leg.; elephant feces; CEMT • 1 spec.; same data as for preceding; Tapirus feces; CEMT • 8 specs; Chapada dos Parecis, 30 km N of Uirapuru; 14 ° 17 ′ S, 59 ° 15 ′ W; 1 – 15 Dec. 2001; A. Foucart leg.; FIT; CEMT • 7 specs; same data as for preceding; Dec. 2002; CEMT • 2 specs; Cuiabá; 15 ° 16 ′ 45 ″ S, 56 ° 00 ′ 06 ″ W; 7 Dec. 2015; A. Frolov and L. Akhmetova leg.; CEMT • 1 spec.; same data as for preceding; Estância Leôncio; 15 ° 11 ′ 11 ″ S, 55 ° 58 ′ 00 ″ W; 1 Nov. 2020; A. Mota and Leon leg.; CEMT • 3 specs; Cuiabá, Fazenda Mutuca; 15 ° 19 ′ 31 ″ S, 55 ° 58 ′ 03 ″ W; elev. 285 m; Nov. 2015; R. V. Nunes and A. Frolov leg.; light; CEMT • 1 spec.; Diamantino; Oct. 2001; E. Furtado leg.; CEMT • 6 specs; Tangará da Serra, Fazenda Três Rios; 14 ° 21 ′ S, 57 ° 42 ′ W; elev. 592 m; Feb. 2008; R. J. Silva leg.; Cerrado; pitfall; CEMT. – Mato Grosso do Sul • 1 spec.; Bodoque na Fazenda Califórnia; 20 ° 41 ′ 02 ″ S, 56 ° 51 ′ 40 ″ W; Mar. 2011; L. O. Bavutti leg.; flight intercept; CEMT • 1 spec.; Corumbá, Nhumirim Farm; Aug. 1992; A. T. M. Barros leg.; Equus caballus dropping; CEMT • 1 spec.; Guia Lopes da Laguna; 15 Dec. 2007; CEMT • 1 spec.; same data as for preceding; 14 Aug. 2008; CEMT • 1 spec.; same data as for preceding; 15 Dec. 2008; A. R. Abot leg.; light; CEMT. – Minas Gerais • 1 spec.; Corações de Jesus, Gruta Sumitumba; 16 ° 39 ′ 47 ″ S, 44 ° 22 ′ 8 ″ W; 29 Jan. 2015; L. M. Rabelo leg.; CEMT • 1 spec.; Três Marias; Nov. 1989; CEMT • 3 specs; same data as for preceding; Dec. 1991; CEMT. – Paraíba • 1 spec.; São José dos Cordeiros, Fazenda Almas; 7 ° 27 ′ 11 ″ S, 36 ° 53 ′ 57 ″ W; Jun. 2019; F. Alvarado leg.; CEMT. – Rio Grande do Norte • 4 specs; Natal; Feb. 1951; M. Alvarenga leg.; CEMT. – Tocantins • 1 spec.; Araguatins, Campus IFTO; 5 ° 37 ′ 56 ″ S, 48 ° 03 ′ 55 ″ W; 5 Nov. 2020; Marinho and D. Bomfim leg.; pitfall; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF16FF41FE20FE95FA83F821.taxon	discussion	Collecting information Specimens can be collected in pastures with light and flight intercept trap and on horse and Tapirus dung.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF17FF40FDE8FED6FB6AFCAB.taxon	diagnosis	Diagnosis Length 3.0 – 3.8 mm. Body elongate, oval, posteriorly narrowed, piceous. Head with six short longitudinal carinae; clypeus with transversal groove before anterior margin, with four teeth, two anterior to clypeal margin and two just before gena, anterior margin not emarginate inward bent. Pronotum with lateral margin sinuate, with two lateral lobes; with six medially longitudinal carinae interrupted by a row of large pits. Elytra narrowed posteriorly, with eight longitudinal carinae and two posterior setaceous tubercles. Protibia with two teeth; meso- and metatibiae inner margin grooved; metatibia laterally flattened, wide.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF17FF40FDE8FED6FB6AFCAB.taxon	discussion	Remarks Aschnarhyparus is monotypic genus recorded from Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guyana, Mexico, Nicaragua, Panama, Peru, Suriname and Trinidad (Skelley 2007 a).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF17FF47FDCAFCA3FD05FCAB.taxon	description	Figs 27 C, 58	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF17FF47FDCAFCA3FD05FCAB.taxon	diagnosis	Diagnosis Same as for the genus.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF17FF47FDCAFCA3FD05FCAB.taxon	materials_examined	Material examined BRAZIL – Acre • 1 spec.; Assis Brasil; 10 ° 35 ′ 44 ″ S, 69 ° 37 ′ 12 ″ W; 23 Sep. 2016; Ortiz and Borges leg.; light; CEMT. – Amazonas • 1 spec.; Rio Nhamundá, Lg. Areias; 1 ° 35 ′ 11 ″ S, 57 ° 37 ′ 32 ″ W; 18 – 19 May 2008; Roger H. Hutchings leg.; CDC; INPA. – Mato Grosso • 1 spec.; Cotriguaçu, Fazenda São Nicolau, PPBio 1; 9 ° 49 ′ 17 ″ S, 58 ° 15 ′ 32 ″ W; 31 Oct. 2017; Vaz-de-Mello et al. leg.; Winkler extractor; CEMT • 3 specs; same data as for preceding; 9 ° 51 ′ 53 ″ S, 58 ° 13 ′ 13 ″ W; 2 Nov. 2017; Vaz-de-Mello et al. leg.; prainha; manually; CEMT. – Pará • 1 spec.; Marabá, Reserva Biológica de Tapirapé; 27 Nov. 2003; F. Gumier-Costa leg.; legal Amazon; CEMT. COLOMBIA – Vaupés • 2 specs; R. N. Mosiro-Itajura (Caparú), Centro Ambiental; 1 ° 04 ′ S, 69 ° 31 ′ W; elev. 60 m; 20 Jan. – 1 Feb. 2003; M. Skakoy and D. Arias leg.; FIT; CEMT. ECUADOR – Pastaza • 1 spec.; Arajuno, B. P. Oglán; elev. 540 m; 25 Apr. 2009; P. A. Araujo and W. Chamorro leg.; fumigation; CEMT. FRENCH GUIANA • 1 spec.; Cayenne, Montage des Chevaux; 4 ° 44 ′ 56 ″ N, 52 ° 26 ′ 28 ″ W; elev. 75 m; 4 Apr. 2014; SAEG leg.; CEMT • 2 specs; same data as for preceding; 4 ° 18 ′ 58 ″ N, 53 ° 17 ′ 10 ″ W; 2 Nov. 2013; FIT; CEMT • 2 specs; same data as for preceding; 4 ° 44 ′ 56 ″ N, 52 ° 26 ′ 28 ″ W; elev. 75 m; 11 Feb. 2013; CEMT • 1 spec.; same data as for preceding; 30 Jan. 2013; light; CEMT • 1 spec.; same data as for preceding; 27 Jan. 2013; CEMT • 1 spec.; same data as for preceding; 4 Feb. 2013 CEMT • 1 spec.; Saint Laurent de Maroni, Belvédère de Saül; 3 ° 37 ′ 22 ″ N, 53 ° 12 ′ 57 ″ W; elev. 326 m; 7 Apr. 2011; FIT, SAEG leg.; CEMT • 3 specs; same data as for preceding; 15 Sep. 2011; CEMT • 1 spec.; same data as for preceding; 2011; CEMT • 1 spec.; same data as for preceding; Mar. 2011; CEMT • 3 specs; same data as for preceding; 22 Mar. 2010; CEMT • 2 specs; same data as for preceding; 13 May 2011; CEMT • 1 spec.; same data as for preceding; 22 Mar. 2011; SAEG; CEMT. PANAMA – Colón • 2 specs; San Lorenzo Forest; 9 ° 17 ′ N, 79 ° 58 ′ W; 25 – 26 Mar. 2004; A. Tishenchkin leg.; FIT; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF17FF47FDCAFCA3FD05FCAB.taxon	description	Sexual dimorphism Males mesotibial marginal groove ending in a prominent pre-apical tooth before notch. Females with tooth less prominent.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF17FF47FDCAFCA3FD05FCAB.taxon	discussion	Collecting information Specimens have been collected with flight intercepts and light trap. This species has also been found inside dead trees, extracted from Ochroma pyramidale (Cav. ex Lam.) Urb. roots and from termite nests of Coptotermes Wasmann, 1896 and Nasutitermes Dudley, 1890 (Skelley 2007 a). The altitudes this species was found range from 50 to 600 metres.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF10FF47FDE2FCA2FBC0FABF.taxon	diagnosis	Diagnosis Length 2.3 – 3.4 mm. Body elongate, cylindrical, red. Head punctate; clypeus weakly sinuate; middle of clypeus with two low longitudinal carinae; frons with four rounded longitudinal carinae. Pronotum with six low longitudinal ridges, the lateral ridges interrupted by large almost circular depressions; the two middle ridges complete. Elytra with sides almost parallel; four low carinae present, three of these ending abruptly before posterior tubercle; longitudinal depressions between carinae with two rows of small punctures.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF10FF47FDE2FCA2FBC0FABF.taxon	discussion	Remarks Leptorhyparus has four recognized species from Brazil, Costa Rica, Nicaragua and Panama (Minkina 2020; Skelley 2021). A key for the species can be found in Minkina (2020).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF10FF46FDC3FAAEFAC2FEC7.taxon	description	Figs 27 E, 58	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF10FF46FDC3FAAEFAC2FEC7.taxon	diagnosis	Diagnosis Length 2.3 mm. Head widely and weakly sinuate medially, bounded by obtuse angles, lateral margins sinuate before rounded gena, protruding eye line; clypeus medially with two longitudinal costae; frons with four longitudinal costae. Pronotum with six longitudinal ridges weakly elevated, only the two central ridges complete. Elytra with five elevated costae and five convex interstriae with a row of coarse punctures between them.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF10FF46FDC3FAAEFAC2FEC7.taxon	description	Sexual dimorphism Unknown.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF10FF46FDC3FAAEFAC2FEC7.taxon	discussion	Collecting information Specimens were collected with black light traps (Minkina 2020). Remarks No specimens were available for the present study, thus the diagnosis provided and positioning of the group in the dichotomous key were made based solely on the available literature (Minkina 2020).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF11FF46FDECFC38FEFDF937.taxon	description	Figs 27 D, 58	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF11FF46FDECFC38FEFDF937.taxon	diagnosis	Diagnosis Length 4.0 mm. Body with bulbous and carinate areas opaque, clypeus, elytral surface between carinae and tubercles near posterior margin, glossy. Clypeus with obsolete set of medial longitudinal costae medially. Pronotum with paramedian costae with anterior lobes elevated and circular in dorsal view, almost as big as the lobes of other pronotal costae.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF11FF46FDECFC38FEFDF937.taxon	materials_examined	Material examined BRAZIL – Acre • 1 spec.; Rio Branco, Fazenda Catuaba; Feb. 1997; F. Z. Vaz-de-Mello leg.; CEMT.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF11FF46FDECFC38FEFDF937.taxon	description	Sexual dimorphism Males with inner margin of mesotibia, with small preapical denticle; lateral margin sinuate between tooth and apical margin; apical fringe of setae complete.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF11FF46FDECFC38FEFDF937.taxon	discussion	Collecting information Specimens have been collected in nests of Nasititermes meinerti (Wasmann, 1894) termites (Wasmann 1894).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF11FF46FD9BFE06FC20FCCC.taxon	diagnosis	Diagnosis Length 3.5 – 4.0 mm. Body elongate, oval, posteriorly narrowed, covered with small setae, reddish brown. Head with anterior margin inflexed medially, acuminate; frons with four longitudinal costae. Pronotum with lateral margins sinuate; disc with six variably developed longitudinal carinae; peridiscal costae normally obsolete medially, visible only near anterior and posterior margin; pronotal costae always bulbous anteriorly; pronotum with two perimarginal costae. Elytra with large tubercle anterior to posterior margin and with six longitudinal carinae. Meso- and metatibia elongate, thin, not widening toward apex.	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
038D87BEFF11FF46FD9BFE06FC20FCCC.taxon	discussion	Remarks Termitodius has four recognized species reported from Brazil, Colombia, Guatemala and Mexico (Skelley et al. 2022). A key for the species was written Skelley et al. (2022).	en	Gama, Emanuel R. R., Skelley, Paul E., Vaz-De-Mello, Fernando Z. (2026): Identification guide to the Aphodiinae (Coleoptera: Scarabaeidae) of Brazil. European Journal of Taxonomy 1038 (1): 1-225, DOI: 10.5852/ejt.2026.1038.3175, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3175/14121
