identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0382BC4B99111A29FF11F913FB83FCB2.text	0382BC4B99111A29FF11F913FB83FCB2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atlantapseudes Bacescu 1978	<div><p>Genus Atlantapseudes Băcescu, 1978</p><p>[New Japanese name: Atoranta-apuseudesu]</p><p>Diagnosis. Carapace length subequal to or longer than wide. Eye lobes acute. Pairs of spines in front of branchial chamber. Antennal squama reduced. Cheliped and pereopod-1 without exopod.</p></div>	https://treatment.plazi.org/id/0382BC4B99111A29FF11F913FB83FCB2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matsushima, Yoshinobu;Bâlcu, Maxim-Jean;Kakui, Keiichi	Matsushima, Yoshinobu, Bâlcu, Maxim-Jean, Kakui, Keiichi (2025): Atlantapseudes tridens sp. nov. (Tanaidacea: Apseudidae) from the Okinawa Trough, Japan, with a note on the taxonomic status of Atlantapseudes curvatus. Zootaxa 5679 (4): 484-500, DOI: 10.11646/zootaxa.5679.4.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
0382BC4B99101A2DFF11FF6EFB0DF8EE.text	0382BC4B99101A2DFF11FF6EFB0DF8EE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atlantapseudes nigrichela , Bacescu 1978	<div><p>Atlantapseudes nigrichela Băcescu, 1978</p><p>(Figs 2, 3)</p><p>Atlantapseudes nigrichela Băcescu, 1978, 317–322 [in part], fig. 1B–D, K.</p><p>Atlantapseudes curvatus Esquete &amp; Cunha, 2017, 61, 65–72, 87, figs 2–6</p><p>Atlantapseudes curvatus: García-Herrero et al. 2021a, 66; García-Herrero et al. 2021b, suppl. 2, 3.</p><p>Material examined. Holotype: ovigerous female (393/30686, Muzeul Național de Istorie Naturală “Grigore Antipa,” TBL = 5.85 mm), Stn 396, Atlantic Ocean, western Portugal (41°17'4"N, 09°15'7"W), 900 m depth, silty or sandy-silty bottom, cruise “Thalassa,” September 1972 . Paratypes: three females (TAN 032-1–3, TBL = 5.60 mm, 5.20 mm, 4.20 mm, respectively, Muzeul Național de Istorie Naturală “Grigore Antipa”), Atlantic Ocean, western Portugal, without other data; three males (49217-1–3, TBL = 4.50 mm, 3.75 mm, 4.10 mm, respectively, Muzeul Național de Istorie Naturală “Grigore Antipa”), Stn 407, Atlantic Ocean, western Portugal (40°33'5"N, 09°24'0"W), 740 m depth, silty or sandy-silty bottom, cruise “Thalassa,” 1 September 1972 .</p><p>Amended diagnosis. Rostrum tridentate. Pereonite-1 lateral margin with posterior spine but without anterior spines. Lateral margin of pereonites 2 and 3 with one anterior spine and one posterior spine. Lateral margin of pereonites 4–6 with anterior spine but without posterior spines. Squama more than 2.5 times longer than antennal article-3. Labial palp with three spiniform setae. Maxillipedal palp article-2 with two outer distal setae. Cheliped with carpus bearing two ventral long setae.</p><p>*, We treated as Atlantapseudes sp. in this study. †, We synonymized with A. nigrichela in this study.</p><p>Supplemental description of holotype. The holotype was strongly deformed and had lost several parts, but we observed the following character states: tridentate rostrum (left part partly broken) (Fig. 2A), a spine present on mid-lateral margin of cephalothorax (Fig. 2B), each eye lobe with anterior spine (Fig. 2A), pereonite-1 with spine on posterolateral margin, a spine present on anterolateral and posterolateral margin of pereonites 2 and 3 (Fig. 2C–F), a spine present on subanterior lateral margin of pereonites 4–6 (Fig. 2F–H), the squama about 3.3 times longer than antennal article-3 (Fig. 2I) and the cheliped with basis and merus each bearing ventro-subdistal spine and carpus bearing two long ventral simple setae (Fig. 2B).</p><p>Supplemental description of paratypes. Cephalothorax and pereonites 1–6 (Fig. 3A–D) similar to those in holotype; squama about 3.1 times longer than antennal article-3; labial palp with three spiniform setae (Fig. 3E); maxillipedal palp article-2 with two outer distal setae (Fig. 3F); pereopod-3 coxa with spine (Fig. 3D) (see also Supplementary File S1).</p><p>Remarks. The holotype of A. nigrichela had (1) a tridentate rostrum; (2) pereonite-1 with a posterolateral spine; and (3) pereonites 2 and 3 with one anterolateral spine and one posterolateral spine. All three states were also found in three female and three male paratypes. We did not detect states (2) and (3) in the monodentate individuals of “ A. nigrichela ” (Băcescu 1978, fig. 1A; Błażewicz-Paszkowycz et al. 2011, fig. 6), but did find them in A. curvatus, which has a tridentate rostrum (Esquete &amp; Cunha 2017, fig. 2A). Those character states were summarized in Table 1. The type locality of A. nigrichela is near the area where A. curvatus has been reported (Fig. 1B). The holotype of A. nigrichela was collected at 900 m depth, within the known depth range of A. curvatus (497–1001 m) (Supplementary Table S1). Based on these morphological and distributional similarities, we concluded that A. curvatus is a junior synonym of A. nigrichela .</p><p>We found that the specimen designated as the allotype and one designated paratype had a monodentate rostrum. Based on this morphology, we identified them as Atlantapseudes sp. (see below). One female designated as a paratype (TAN 032-4) was badly deteriorated and could not be identified to species level.</p></div>	https://treatment.plazi.org/id/0382BC4B99101A2DFF11FF6EFB0DF8EE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matsushima, Yoshinobu;Bâlcu, Maxim-Jean;Kakui, Keiichi	Matsushima, Yoshinobu, Bâlcu, Maxim-Jean, Kakui, Keiichi (2025): Atlantapseudes tridens sp. nov. (Tanaidacea: Apseudidae) from the Okinawa Trough, Japan, with a note on the taxonomic status of Atlantapseudes curvatus. Zootaxa 5679 (4): 484-500, DOI: 10.11646/zootaxa.5679.4.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
0382BC4B991B1A25FF11FF6EFDBDF946.text	0382BC4B991B1A25FF11FF6EFDBDF946.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atlantapseudes tridens Matsushima & Bâlcu & Kakui 2025	<div><p>Atlantapseudes tridens sp. nov.</p><p>[New Japanese name: Sanbonduno-apuseudesu]</p><p>(Figs 5–8)</p><p>Atlantapseudes sp.: Kakui et al. 2011, table 1, figs 2–5; Matsushima &amp; Kakui 2024, fig. 7.</p><p>Type material. Holotype: preparatory female (ICHUM4002, TBL = 7.10 mm, CW = 1.39 mm, 11 slides and one vial), R/V Soyo-maru, Stn Kago-7, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.01334&amp;materialsCitation.latitude=28.495" title="Search Plazi for locations around (long 128.01334/lat 28.495)">Okinawa Trough</a>, East China Sea (28°29'24.0"N, 127°59'30.0"E to 28°29'42.0"N, 128°00'48.0"E), 1076–1079 m depth, plankton net attached inside beam trawl, 29 July 2010, collected by K. Kakui. Paratype: one preparatory female (ICHUM8985, TBL = 5.66 mm, CW = 1.05 mm, 11 slides and one vial, R/V Soyo-maru, Stn Kago-9, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.96778&amp;materialsCitation.latitude=27.513277" title="Search Plazi for locations around (long 126.96778/lat 27.513277)">Okinawa Trough</a>, East China Sea (27°30'27.5"N, 126°58'10.9"E to 27°30'47.8"N, 126°58'04.0"E), 1595– 1557 m depth, plankton net attached inside beam trawl, 26 July 2014, collected by K. Kakui.</p><p>Representative DNA Sequences. One 18S (INSD accession number AB618176; 2189 bp long; Kakui et al. 2011) and one COI (LC860315; 658 bp long, encoding 219 amino acids; this study) sequences were determined from the holotype.</p><p>Etymology. The specific name tridens (Latin: trident) is a noun in the nominative case, referring to the shape of the tridentate rostrum.</p><p>Diagnosis. Rostrum tridentate. Epistome strongly curved ventrally. Pereonites 1 and 2 without lateral spines. Lateral margin of pereonites 3–6 with anterior spine (tiny in pereonite-6) but without posterior spines. Squama more than twice longer than antennal article-3. Labial palp with two distal spiniform setae. Maxillipedal palp article-2 with outer distal seta. Chelipedal carpus with two ventral long setae.</p><p>Description of female (based on holotype unless noted otherwise). Body (Figs 5, 6A 1, 2, B [holotype ICHUM4002]; 6C1–4, D [paratype ICHUM 8985]) well calcified, dorsoventrally flattened, 5.1 times longer than CW, whitish in specimens before fixation (Fig. 5). Cephalothorax length 0.2 times TBL, 1.1 times as long as wide; rostrum tridentate; eye lobes distinct, with anterior spine; mid-lateral margin with spine; acute ventral keel (hyposphenium) present, posterior to maxillipeds. Epistome (Fig. 6E) strongly curved ventrally. Pereonites 1–6 with length ratio 1.00: 1.16: 1.59: 1.84: 1.79: 1.25; with width/length ratio 3.09: 2.31: 1.93: 1.89: 1.59: 1.70 and with acute hyposphenium; lateral margin of pereonites 3–6 with anterior spine, with the spine of pereonite-6 tiny. Pleon length 0.2 times TBL. Pleonites distinctly narrower than pereonites; each pleonite with width/length ratio 3.60: 3.01: 3.05: 3.06: 2.48, similar in shape, with hyposphenium (acute in holotype; obtuse and rounded on pleonites 1–4 but acute on pleonite- 5 in paratype; Fig. 6B, D); epimera (Fig. 6A 2, B, C 4, D) projecting ventroposteriorly, with pointed apex. Pleotelson (Fig. 6A 2, B, C 4, D) 1.1 times as long as wide, narrower than pleonites, with posterior projection.</p><p>Antennule (Fig. 7A) with eight-articled outer flagellum and three-articled inner flagellum. Combined length of peduncle and outer flagellum 1.26 times longer than cephalothorax. Peduncular articles 1–4 with length ratio 1.00: 0.41: 0.11: 0.11; article-1 with two mid-outer, one mid-inner, and one inner subdistal simple setae and ten outer (four broken) PSS; article-2 with one mid-outer, two outer distal (one long), one inner subproximal, and one inner subdistal simple setae and three outer distal (one broken) and two inner distal PSS; article-3 with one outer distal and three inner distal simple setae; article-4 with two PSS at insertion of outer flagellum. Outer flagellum 0.5 times as long as peduncle; articles 1 and 2 naked; articles 3–8 with three distal, one distal, three distal (one long), one distal, two distal (one long), and two middle and four distal simple setae, respectively; article-8 with one PSS (broken); article-7 with distal long aesthetasc. Inner flagellum 0.5 as long as outer flagellum; articles 1–3 with two, one (broken), and three (one broken) distal simple setae, respectively; articles 1–3 each with distal PSS.</p><p>Antenna (Fig. 7B) with 11 articles, 0.9 times as long as antennule; articles 1–11 with length ratio 1.00: 1.80: 0.67: 1.56: 2.43: 0.91: 0.64: 0.63: 0.46: 0.47: 0.45. Article-1 naked. Article-2 with one outer subproximal, one outer distal, and one inner subproximal simple setae. Article-3 with inner distal simple seta. Article-4 with inner subproximal simple seta and two inner subdistal PSS (one broken). Article-5 with one inner subproximal and one inner subdistal simple setae and one outer subproximal, one mid-outer, two inner subdistal, and two dorsal subdistal PSS. Article-6 with one mid-outer, one distal long, and one inner subdistal simple setae. Articles 7–11 with three distal (one long), one inner subdistal and three distal (one long, one broken), two distal (one long, one broken), and five distal simple setae, respectively; article-9 with inner distal PSS. Squama 0.2 as long as antenna, 2.4 times longer than antennal article-3, and 1.0 times as long as antennal article-4; narrow, with two simple setae (one long).</p><p>Labrum (Fig. 7C) bipartite with numerous fine setae.</p><p>Mandibles (Fig. 7D–G) with well-developed molar process bearing six tufts of fine setae (Fig. 7D). Left mandibular incisor (Fig. 7E) with five rounded processes; setiferous lobe with two setulate and serrate setae, two setulate bifurcate setae, two spiniform setae, and fine setae; lacinia mobilis with five rounded processes. Right mandibular incisor (Fig. 7F) with four rounded processes; setiferous lobe with five round-tipped setae and fine setae. Palp (Fig. 7G) with three articles; articles 1–3 with length ratio 1.00: 2.67: 0.79; article-1 with middle simple seta; article-2 with seven bipinnate setae in distal region; article-3 with 10 distal bipinnate setae.</p><p>Labium (Fig. 7H) with lobe bearing several outer simple setae and inner setation. Palp with several short and long simple setae on lateral margins, and two distal bifurcate spiniform setae.</p><p>Maxillule (Fig. 7I) with biarticulate palp; palp article-1 with three fine setae; palp article-2 with five pinnatetipped setae (one broken) in distal half. Outer endite with numerous simple setae in middle region, outer proximal and inner fine setation, 10 distal spiniform setae, and two subdistal plumose setae. Inner endite with outer and inner distal fine setation and three distal setulate spiniform setae.</p><p>Maxilla (Fig. 7J) with body bearing inner distal fine setae, short simple setae in inner distal region, 13 outer distal simple setae, and outer proximal serration. Inner lobe of fixed endite with several short simple setae, three biserrate setae, and 17 basally swollen setae. Outer lobe of fixed endite with outer fine setae, eight inner short simple setae, six distal simple setae, one distal and one subdistal plumose setae, and three distal trifurcate setae. Inner lobe of movable endite with inner fine setae, nine ventral short simple setae, three plumose setae (one broken), furcate seta, and four distal simple setae. Outer lobe of movable endite with two ventral short simple setae and seven setulate serrate setae.</p><p>Maxilliped (Fig. 7L, l 1, K) with naked coxa. Basis with outer fine setae and two mid-inner simple setae. Endite (Fig. 7K, paratype) with several fine setae in middle region; inner region with four setulate serrate setae and two coupling hooks; distal region with fine setae, two plumose setae, and eight setulate serrate setae. Palp article-1 with one outer distal seta and one inner distal simple seta; article-2 with one outer distal and eleven inner simple setae; article-3 with eleven inner simple setae; article-4 with nine inner simple setae and two inner serrate setae (broken) (Fig. 7l 1). Epignath (Fig. 7M, paratype) cup-shaped, with setulate terminal seta.</p><p>Cheliped (Fig. 8A) without exopod. Basis 2.6 times as long as wide, with one mid-dorsal and four ventral simple setae and one dorsoproximal and one ventro-subdistal spines; inner surface smooth. Merus with ventrodistal simple seta and ventrodistal spine. Carpus 0.64 times as long as basis, 2.5 times as long as wide, with one dorso-subproximal, one dorsodistal, two ventro-subdistal short simple setae and two ventral long simple setae. Propodus with one mid-outer and four ventral simple setae; dorsal region of palm with two distal simple setae; cutting surface with one outer proximal, two inner proximal, and two inner subdistal simple setae, eleven outer round-tipped setae, and several denticles; unguis curved and pointed. Dactylus longer than fixed finger, with three outer and three inner simple setae and several denticles; unguis curved and pointed, longer than fixed-finger unguis.</p><p>Pereopods 1–6 with length ratio 1.00: 0.94: 0.82: 0.84: 0.88: 0.82. Pereopod-1 (Fig. 8B) 0.5 times as long as TBL, with length ratio of basis, ischium, merus, carpus, propodus, and dactylus–unguis 1.00: 0.07: 0.43: 0.39: 0.28: 0.27. Coxa with anterior spine and six plumose setae. Basis 4.83 times as long as wide, with one dorso-subdistal, one mid-ventral, one ventro-subdistal, and two ventrodistal (one broken) simple setae and two mid-dorsal (one long) PSS. Ischium with two ventrodistal simple setae. Merus with ventro-subproximal row of five simple setae, four dorsodistal and three ventrodistal simple setae, and ventrodistal spiniform seta. Carpus with two mid-dorsal, six dorsodistal, four ventral, and three ventrodistal simple setae and one dorsodistal and two ventrodistal spiniform setae. Propodus with one mid-dorsal, one dorsodistal, and two ventrodistal simple setae, two dorsal and five ventral spiniform setae, inner dorso-subdistal PSS, and inner ventrodistal pinnate seta. Dactylus with two dorso-subproximal simple setae and three ventral spines; unguis half as long as dactylus, naked.</p><p>Pereopod-2 (Figs. 6A 1, B, C1, D, 8C) with length ratio of articles from basis to dactylus–unguis 1.00: 0.13: 0.36: 0.38: 0.32: 0.34. Coxa with spine (broken in left pereopod-2 of holotype, Figs. 6A 1, B, C1, D) and three small simple setae. Basis 6.67 times as long as wide, with four setae, one mid-dorsal, one ventro-subproximal, one ventro-subdistal, and one ventrodistal simple, and one dorso-subproximal and one mid-dorsal PSS. Ischium with ventrodistal simple seta. Merus with three dorsodistal, one ventro-subproximal (broken), one mid-ventral, and three ventrodistal simple setae. Carpus with seven dorsal and nine ventral (four broken) simple setae. Propodus with six dorsal and ten ventral simple setae, dorso-subdistal PSS, and inner distal pinnate seta. Dactylus with two dorso-subproximal and two ventrodistal simple setae and ventral spine; unguis more than half as long as dactylus, naked (Fig. 8C).</p><p>Pereopod-3 (Fig. 8D) with length ratio of articles from basis to dactylus–unguis 1.00: 0.10: 0.31: 0.33: 0.27: 0.15: 0.14; coxa without spine; other characters similar to pereopod-2 except in number of setae.</p><p>Pereopod-4 (Fig. 8E) with length ratio of articles from basis to dactylus–unguis 1.00: 0.09: 0.25: 0.38: 0.30: 0.23. Coxa naked. Basis 4.7 times as long as wide, with one ventro-subdistal seta and one ventrodistal simple seta, and three dorso-subproximal (one broken), one ventroproximal, and one mid-ventral (long) PSS. Ischium with ventrodistal simple seta. Merus with one dorsodistal, one mid-ventral, and two ventrodistal (one broken) simple setae. Carpus with one mid-ventral, one ventro-subdistal, and seven distal simple setae. Propodus with mid-ventral simple seta, mid-dorsal PSS, nine distal pinnate setae, and two distal long serrate setae. Dactylus with ventrodistal simple seta; unguis half as long as dactylus, naked.</p><p>Pereopod-5 (Fig. 8F) with length ratio of articles from basis to dactylus–unguis 1.00: 0.07: 0.22: 0.37: 0.26: 0.18. Coxa with simple seta. Basis 7.1 times as long as wide, with one dorsoproximal, one dorso-subproximal, and one ventrodistal (broken) small simple, setae, and two dorso-subproximal (broken) and one mid-ventral (broken) PSS. Ischium with ventrodistal simple seta. Merus with one dorso-subdistal and two ventrodistal simple setae. Carpus with one dorso-subdistal, two dorsodistal, one mid-ventral, and three ventro-subdistal simple setae. Propodus with one dorso-subdistal, one dorsodistal, one mid-ventral, three ventro-subdistal, and one ventrodistal simple, setae, dorso-subdistal PSS, and two distal pinnate setae (one broken). Dactylus with three ventrodistal simple setae; unguis half as long as dactylus, naked.</p><p>Pereopod-6 (Fig. 8G) with length ratio of articles from basis to dactylus–unguis 1.00: 0.09: 0.22: 0.37: 0.26: 0.29. Basis 6.8 times as long as wide, with one ventroproximal, one mid-ventral, and one ventrodistal small simple, setae, and three dorso-subproximal (two broken) and one mid-ventral (long) PSS. Ischium with ventrodistal simple seta. Merus with one dorsodistal seta and one ventrodistal (broken) simple seta. Carpus with three dorsodistal, one ventro-subdistal, and two ventrodistal simple setae. Propodus with one ventro-subproximal, three mid-outer (one broken), and four distal (one broken) simple setae and mid-dorsal PSS. Dactylus with two mid-dorsal setae and one ventrodistal simple seta; unguis half as long as dactylus, naked.</p><p>Pleopods absent.</p><p>Uropod (Fig. 8H, paratype) 4.21 times longer than pleotelson (Fig. 6D), with protopod bearing two outer distal, one mid-inner, one inner subproximal (long), and inner distal simple setae. Exopod with four articles; article-3 naked; articles 1, 2 and 4 with one subproximal, one distal, and four distal (one broken) simple setae, respectively. Left endopod with 18 articles; articles 1, 3, 5, 7, 9, 14, 16, 17 naked; articles 4, 6, 8, 10–13, 15, 18 with two, one (broken), two, one (broken), one, two, one, two, and four distal simple setae, respectively; articles 2, 4, 8, 12, 15, 18 with one, three, three (two long), two (one long), two (one long), and two (one long) distal PSS, respectively. Right endopod with 19 articles (for setation see Supplementary Table S2).</p><p>Variation and stability. In addition to the holotype, one paratype female (ICHUM8985) was dissected and selected characters were observed (see Supplementary Table S2). Both specimens shared the same state for the characters listed in the Diagnosis section for this species (for the other shared characters, see Supplementary Table S2).</p><p>Remarks. Atlantapseudes tridens sp. nov. resembles A. nigrichela in having a tridentate rostrum but differs from the latter in having (1) the posterolateral margin of pereonites 1–3 without spines (one spine in A. nigrichela); (2) a tiny anterolateral spine on pereonite-6 (large in A. nigrichela); (3) two distal spiniform setae on labial palp (three in A. nigrichela); and (4) one outer distal seta on maxillipedal palp article-2 (two in A. nigrichela).</p><p>A partial COI sequence (658 bp, encoding 219 amino acids; INSD accession number LC860315) was determined from the holotype specimen. The sequence in the INSD most similar to our sequence, determined by a BLAST search, was from the insect Forcipomyia sp. (MG475921.1; identity score 76.02%, query cover 100%; deWaard et al. 2019). This was likely because our COI sequence was the first from any Atlantapseudes species in the INSD; a similar result has been reported for several other crustaceans (e.g., Tanabe &amp; Kakui 2019; Kakui &amp; Hiruta 2022; Shiraki &amp; Kakui 2024; Kakui 2024).</p></div>	https://treatment.plazi.org/id/0382BC4B991B1A25FF11FF6EFDBDF946	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matsushima, Yoshinobu;Bâlcu, Maxim-Jean;Kakui, Keiichi	Matsushima, Yoshinobu, Bâlcu, Maxim-Jean, Kakui, Keiichi (2025): Atlantapseudes tridens sp. nov. (Tanaidacea: Apseudidae) from the Okinawa Trough, Japan, with a note on the taxonomic status of Atlantapseudes curvatus. Zootaxa 5679 (4): 484-500, DOI: 10.11646/zootaxa.5679.4.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
