taxonID	type	description	language	source
0384A6263A48E16F197CA2BB7D443CD2.taxon	description	urn: lsid: zoobank. org: act: EC 5381 E 8 - 71 BE- 49 F 9 - 92 AA- 5 E 11 B 3 BBE 41 D Figs 1 – 2, Tables 2 – 5	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A48E16F197CA2BB7D443CD2.taxon	type_taxon	Type species Miosterpa flammarubra Crispolon & Soulier-Perkins gen. et sp. nov.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A48E16F197CA2BB7D443CD2.taxon	diagnosis	Diagnosis Miosterpa Crsipolon & Yap gen. nov. possesses characters similar to Mioscarta and Poeciloterpa but the following characters taken together allow it to be distinguished clearly from both genera: habitus slightly dorso-ventrally flattened, 2 – 4 mm shorter than Mioscarta, with small ocelli, postclypeus with two prominent lateral carinae, widest part before midlength of tegmina when viewed dorsally, apical cells concave; male terminalia with subgenital plate with equal length relative to height of pygofer without a slender apical appendage extension (as in Mioscarta). Additional character diagnoses are presented in Table 3 (modified from Crispolon et al. 2021), comparing the four closely related genera of the tribe Rhinaulacini Kirkaldy, 1906 from the Philippines	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A48E16F197CA2BB7D443CD2.taxon	etymology	Etymology The generic name is derived from the two most closely related genera, Mioscarta and Poeciloterpa, with the first four letters of the former genus are concatenated with the last five letters of the latter genus. The gender of the genus is feminine.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A48E16F197CA2BB7D443CD2.taxon	description	Description Body length 6 – 7 mm (tegmina included), width 3 – 3.5 mm. GENERAL SHAPE. In dorsal view, eyes slightly elongated, small ocelli, distance between eyes 9 – 10.5 × ocellus diameter, distance between ocelli equals 1 – 1.5 × ocellus diameter, distance between ocellus and compound eye 3 – 3.5 × ocellus diameter, ocelli closer to each other than to compound eyes, eyes not prominent, 1.5 – 2 × as long as wide, vertex and frons with longitudinal median carina present (Figs 1, 2 B). Postclypeus with longitudinal furrow forming two prominent parallel longitudinal carinae (Figs 1, 2 B), slightly swollen, ovoid shape in frontal view, widest part at mid-height, not receding before anteclypleus where it bends forming obtuse triangle in lateral view (Figs 1, 2 A). Rostrum very long, reaching but not surpassing metacoxae. Tegmina opaque, R bifurcates on apical half, M bifurcates on basal third, veins not prominent on first ⅔, becoming prominent on apical third, forming concave apical cells (Figs 1, 2 A, C). MALE TERMINALIA. Subgenital plates finger-shaped, with equal length relative to height of pygofer without any elongated, thin apical appendage extension (as in Mioscarta), (Figs 1, 2 – F). Paramere (Figs 1, 2 G) midlength broad without protrusion on lateral side (as in Trigonoschema), with process on dorsal margin located apically, ventral margin bearing apically two processes. Aedeagus (Figs 1, 2 H) S-shaped with posterior protrusion (pp) and apical extension.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A48E16F197CA2BB7D443CD2.taxon	distribution	Distribution Philippines.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A48E16F197CA2BB7D443CD2.taxon	diagnosis	A new genus, distinct from Mioscarta Breddin, 1901 and Poeciloterpa Stål, 1870 At first glance, specimens of Miosterpa Crsipolon & Yap gen. nov. appear similar to Mioscarta in overall morphology, size, and general habitus. However, closer examination of detailed characters reveals clear distinctions between the two genera. When examining the male terminalia, this genus doesn’t possess the elongated, thin appendage on the subgenital plate that distinguishes Mioscarta from Poeciloterpa and other genera within the tribe. On the other hand, the absence of this appendage on the subgenital plate and other characters such as smaller ocelli, bulging pronotum, presence of thick veins in the apex of the tegmina, and presence of concave apical cells, could lead us to identify this species as belonging to the genus Poeciloterpa. However, Miosterpa doesn’t exhibit the dome-shaped general habitus that clearly distinguishes Poeciloterpa from the other cercopid genera (Crispolon et al. 2019, 2021). To support our decision to describe Miosterpa as a new genus, we used the species for which we had molecular samples to do a phylogenetic and percentage genetic distance / variation analyses using four genes (COX 1, Histone 3, 18 S, and 28 S) on top of the morphological evidence.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A48E16F197CA2BB7D443CD2.taxon	discussion	Phylogenetic support and genetic distance between genera As discussed by Soulier-Perkins & Stroiński (2017), the intra- or interspecific delimitation through genetic distance can be variable depending on the group of organisms. Even with the advancement of molecular science, there is still a huge gap in terms of our knowledge on generic and species delimitation using molecular data for the whole superfamily Cercopoidea, or at least for the family Cercopidae. This is proven by the non-inclusion of Cercopidae or Cercopoidea in some of the studies before the works of Castanhole et al. in 2013 and Crispolon et al. in 2024 (e. g., Gopurenko et al. 2013). Although their works report the intra- and interspecific genetic distances of some species of Cercopidae, these works do not suggest what percentage of genetic variation should be the standard value for the generic and species delimitation, at least for the family. Even without a clear standard value for species delimitation, the work of Crispolon et al. (2024) gives more straightforward results, clearly separating three genera from the Philippines. In their results using 522 COX 1 sites, the p-distance between species in Jacobsoniella Melchar, 1914 was 6.32 %. Unfortunately, the p-distance was not compared between species of Phantagma and Vinpietri as only one species for each genus has COX 1 data. Further, the p-distance they obtained between the three genera (Jacobsoniella vs Phantagma vs Vinpietri) was 12.07 % to 16.28 %. In the current study, together with the support of our morphological evidence and our phylogenetic analyses, we also determined the percentage genetic distance between genera and species using the COX 1 gene sites to further support that Miosterpa Crsipolon & Yap gen. nov., Poeciloterpa, Mioscarta, and Trigonoschema are distinct genera. According to the previous studies, the average percentage genetic distance within and between species of Hemiptera ranges from 0.31 % to as high as 23.4 % (Hebert et al. 2003; Bressan et al. 2009; Gitau et al. 2011; Gopurenko et al. 2013; Picciau et al. 2016; Soulier-Perkins & Stroiński 2017; Zheng et al. 2018). Here, based on 312 COX 1 sites (Table 4), the p-distance within and between species of Miosterpa ranges from 0.32 % to 12.5 %. Further, the p-distance between the species of Miosterpa and all species of Poeciloterpa with the COX 1 gene ranges from 12.18 % to 15.71 %. In addition, between the species of Miosterpa and Mioscarta, the p-distance ranges from 12.82 to 15.38 %, while between the species of Miosterpa and Trigonoschema, the p-distance ranges from 11.54 to 12.82 %. With the strong morphological evidence and phylogenetic analysis, we conclude that Miosterpa Crsipolon & Yap gen. nov. is a distinct genus from Poeciloterpa, Mioscarta and Trigonoschema. Despite the heterogeneity in the ranges of p-distance within and between species, and between genera (especially between species of Miosterpa, some species of Poeciloterpa, and Trigonoschema), this analysis still aids in the delimitation of the genera in question. Although the result of the percentage genetic distance analysis of the current study does not directly suggest a standard value of p-distance for species delimitation. They do suggest the value of a wider and more comprehensive analysis with more exemplars from the family Cercopidae for a standard value of p-distance for generic and species delimitation. Tribal and subtribal placement of Miosterpa Crsipolon & Yap gen. nov. Miosterpa Crsipolon & Yap gen. nov. possesses characters that allow its clear identification from the other Rhinaulacini genera found in the Philippines, specifically Mioscarta, Poeciloterpa, and even Trigonoschema, the genera belonging to subtribe Poeciloterpina Schmidt, 1920. The characters listed by Lallemand (1949) and Liang & Webb (2002), especially the presence of a sterno-lateral plate between the pygofer and subgenital plate on the male genitalia (Liang & Webb 2002), define the tribe Rhinaulacini. These characters were further discussed by Crispolon et al. (2021, 2024). This genitalic character was also characteristic of all the ingroups used in the current study in the phylogenetic analysis. Miosterpa fits with the definition of the tribe; we are therefore certain in placing Miosterpa in Rhinaulacini. This makes eight genera belonging to the tribe Rhinaulacini present in the Philippines: Miosterpa, Eoscarta, Jacobsoniella, Mioscarta, Phantagma, Poeciloterpa, Trigonoschema, and Vinpietri. As Miosterpa is close to Mioscarta and Poeciloterpa in the subtribe Poeciloterpina, we further place it in the subtribe Poeciloterpina. This is also supported and confirmed by the characters on the posterior wing (listed by Breddin (1901) and further elaborated by Lallemand (1949 )). Molecular phylogenetic analysis presented in the current study also supports the decision (Fig. 4).	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A48E16F197CA2BB7D443CD2.taxon	biology_ecology	Distribution and some notes on ecology Two new species are currently described, belonging to Miosterpa Crsipolon & Yap gen. nov., a new endemic genus from the Philippines. The description of a new genus and two new species brings the total number of genera endemic to the Philippines to seven out of 22 total and brings the total number of Philippine cercopid species to 75, with around 88 % endemicity. Miosterpa and the two included species are recorded in the islands of Luzon, Negros, and Mindanao (Fig. 5). The habitats of the two species are a mixed forest landscape, a combination of disturbed and native vegetation. Both species were collected in both primary and secondary forest areas with patches of agricultural lands nearby. Unfortunately, only a single female specimen of one species, Miosterpa flammarubra Crispolon & Soulier-Perkins gen. et sp. nov. from Mindanao, can be associated with an identified host plant, a species in the family Melastomataceae Juss .. Most of the specimens of both species were collected using light trapping or sweep netting. Light trapping confirms that Miosterpa, displays positive phototaxy. This adds an additional genus of Cercopidae with such behaviour, as only a few cases have been reported in the scientific literatures (e. g., Soulier-Perkins & Kunz 2012; Crispolon et al. 2021, 2024).	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A46E16C1AEAA51A7C703DB5.taxon	description	urn: lsid: zoobank. org: act: 32 C 5 D 6 FF-C 599 - 4 FC 1 - A 95 A-BA 3 F 52 DDCD 86 Figs 1, 4 – 5, Tables 2, 4	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A46E16C1AEAA51A7C703DB5.taxon	diagnosis	Diagnosis General shape close to M. kalanguya Crispolon & Yap gen. et sp. nov., but the colour pattern distinguishes it immediately. This species is red in colour, while M. kalanguya is yellowish brown.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A46E16C1AEAA51A7C703DB5.taxon	etymology	Etymology The name ‘ flammarubra ’ represents the colour of the tegmina, which is flaming red. It is a combination of the two Latin words ‘ flamma ’ which means ‘ flame’, and ‘ rubra ’ meaning red.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A46E16C1AEAA51A7C703DB5.taxon	materials_examined	Type material Holotype PHILIPPINES • ♂; “ Philippines / Negros volcan Canlaon / forêt / N 10 ° 25 ′ 31 ″ E 123 ° 05 ′ 43 ″ // MUSEUM PARIS / 1098 m / 30 - X- 2010 / D. Ouvrard Rec. // UPLBMNH HEM- 06923 ”; UPLBMNH. Paratypes PHILIPPINES • 3 ♂♂; “ Philippines / Negros volcan Canlaon / champ de coqs / N 10 ° 25 ′. 29 ″ E 123 ° 05 ′ 23 ″ // MUSEUM PARIS / Piège lumineux / 932 m / 28 - X- 2010 / A. Soulier-Perkins rec. // MNHN, Paris EH 31003 – 31005 ”; MNHN • 2 ♂♂; “ Philippines / Negros volcan Canlaon / forêt / N 10 ° 25 ′ 34 ″ E 123 ° 05 ′ 37 ″ // MUSEUM PARIS / 1070 m / 29 - X- 2010 / D. Ouvrard Rec. // MNHN, Paris EH 31006 – 31007 ”; MNHN • 1 ♂; “ Philippines / Negros volcan Canlaon / forêt / N 10 ° 25 ′ 31 ″ E 123 ° 05 ′ 40 ″ // MUSEUM PARIS / Piège lumineux / 1098 m / 29 - X- 2010 / A. Soulier-Perkins rec. // sequencage par Elorde Crispolon C- 00080 // MNHN, Paris EH 31008 ”; MNHN • 6 ♂♂; “ Philippines / Negros volcan Canlaon / forêt / N 10 ° 25 ′ 31 ″ E 123 ° 05 ′ 40 ″ // MUSEUM PARIS / Piège lumineux / 1098 m / 29 - X- 2010 / A. Soulier-Perkins rec. // MNHN, Paris EH 31009 – 31014 ”; MNHN • 1 ♂; Philippines / Negros volcan Canlaon / forêt / N 10 ° 25 ′ 31 ″ E 123 ° 05 ′ 43 ″ // MUSEUM PARIS / 1098 m / 30 - X- 2010 / D. Ouvrard Rec. // séquençage par Elorde Crispolon C- 00081 // MNHN, Paris EH 31015 ”; MNHN • 14 ♂♂; “ Philippines / Negros volcan Canlaon / forêt / N 10 ° 25 ′ 31 ″ E 123 ° 05 ′ 43 ″ // MUSEUM PARIS / 1098 m / 30 - X- 2010 • 6 ♂♂; “ Philippines / Negros volcan Canlaon / forêt / N 10 ° 25 ′ 36 ″ E 123 ° 05 ′ 37 ″ // MUSEUM PARIS / Piège lumineux / 1057 m / 31 - X- 2010 / D. Ouvrard Rec. // MNHN, PARIS EH 31039 – 31044 ”; MNHN • 3 ♂♂; “ Philippines / Negros volcan Canlaon / forêt / N 10 ° 25 ′ 36 ″ E 123 ° 05 ′ 37 ″ // MUSEUM PARIS / Piège lumineux / 1057 m / 31 - X- 2010 / A. Soulier-Perkins rec. // MNHN, Paris EH 31045 – 31047 ”; MNHN • 5 ♂♂; “ Philippines / Negros volcan Canlaon / forêt / N 10 ° 25 ′ 36 ″ E 123 ° 05 ′ 37 ″ // MUSEUM PARIS / Piège lumineux / 1057 m / 31 - X- 2010 / A. Soulier-Perkins rec. // UPLBMNH HEM- 06924 – 06928 ”; UPLBMNH • 1 ♂; “ Philippines / Luzon / Mount Isarog / Light trap // MUSEUM PARIS / 31 - V- 2011 / S. Yap Rec. // UPLBMNH HEM- 06929 ”; UPLBMNH • 1 ♀; “ Philippines / Mount Apo / Bongolanon / Magpet, N. Cotabato / 27 to 28 - VI- 2019 / ESCrispolon // UPLBMNH HEM- 06930 ”; UPLBMNH.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A46E16C1AEAA51A7C703DB5.taxon	description	Description Body length 6 – 7 mm (tegmina included), width 3.5 mm. HEAD (Fig. 1 C). Head including eyes, narrower than widest part of pronotum. In dorsal view, eyes slightly elongated, small ocelli, distance between eyes 10.5 × ocellus diameter, distance between ocelli equals 1.5 ocellar diameter, distance between ocellus and compound eye 3.5 × ocellus diameter, ocelli closer to each other than to compound eyes. Eyes not prominent, 1.5 × as long as wide. Vertex and frons longitudinal median carina present. Vertex slightly longer than wide with 3.5 × ocellus diameter in between the two vertex grooves outside ocelli and 4.5 × ocellus diameter between anterior and posterior vertex margins. No dimple in upper part of postclypeus just below frons (as in Amberana Distant, 1908). Postclypeus with longitudinal furrow forming two prominent parallel longitudinal carinae, slightly swollen, ovoid shape in frontal view, widest part at mid-height (Fig. 1 B), not receding prior to anteclypleus where it bends forming obtuse triangle in lateral view (Fig. 1 A). Rostrum very long, reaching but not surpassing metacoxae. THORAX. In dorsal view, pronotum with anterior concavities on each side, anterior margin of the pronotum as wide as posterior margin of the head including eyes, anterolateral margins curved, posterior margin grooved, posterolateral margins slightly concave and longer than anterolateral margin, humeral angle rounded. Pronotum angle in lateral view around 45 ° (Fig. 1 A). Scutellum slightly longer than wide with large median concavity. TEGMEN. R bifurcates on apical half, M bifurcates on basal third, veins not thickened on first ⅔, becoming thickened on apical third forming concave apical cells (Fig. 1 C). POSTERIOR WING. Rp separating from SC + RA nearly at midlength, r-m absent, M reaches ambient vein, CuA and CuP fused at base, CuA bifurcate in CuA 1 and CuA 2 near midlength, m-cu links M to CuA above CuA bifurcation, total of six longitudinal veins and five apical cells between SC + RA and CuP, angular protrusion of the costal margin near base present (Fig. 3 B). LEGS. Metafemur with apical spine on inner margin, metatibia bearing 1 lateral spine, both primary and secondary basitarsi with one row of at least five spines. MALE TERMINALIA. In lateral view, posterior margin of pygofer (Fig. 1 E) largely convex medially with a slight curvature on last third. Subgenital plates (Fig. 1 E – F) finger-shaped, equal in length to height of pygofer without any elongated, thin apical appendage extension (as in Mioscarta), ventral and dorsal margin regularly rounded, apex of the dorsal margin dented, sterno-lateral plate present and slightly elongated, intermediate plates present, subtriangular, linking internal sides of sterno-lateral plate and subgenital plate. Paramere (Fig. 1 G), dorsal margin convex then curving abruptly dorsally and finishing with a sharp process directed anteriorly, ventral margin regularly rounded with apical part bearing two spiniform processes directed antero and postero-ventrally. Aedeagus (Fig. 1 H), basal third of dorsal margin sharply bent and last ⅔ vertical and S-shaped, basal part of erected part of aedeagus enlarged, apical extension (ae) directed posteriorly, posterior protrusion (pp) axe-shaped with fine needle-shaped extension prolonging edge dorsally. COLOUR. Body covered with yellowish setae. Head, thorax red, rostrum orange, antennal scape and pedicel yellowish, legs reddish brown. Protibia without brown or black stripes on external margin. Male tegmina entirely flaming red, female darker red.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A46E16C1AEAA51A7C703DB5.taxon	materials_examined	Type locality Philippines: Visayas, Negros Occidental, Mount Kanlaon.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A46E16C1AEAA51A7C703DB5.taxon	distribution	Distribution Philippines: Luzon Island; Visayas, Negros Island; Mindanao Island (Fig. 5).	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A45E1771ABFA3F07BF83CAA.taxon	description	Fig. 2, 4 – 5, Tables, 2, 4	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A45E1771ABFA3F07BF83CAA.taxon	diagnosis	Diagnosis General shape close to M. flammarubra Crispolon & Soulier-Perkins gen. et sp. nov., but the colour pattern distinguishes it immediately. This species is yellowish-brown in colour, while M. flammarubra is red.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A45E1771ABFA3F07BF83CAA.taxon	etymology	Etymology The species name refers to the tribe ‘ Kalanguya’ inhabiting the area where the type specimens were collected. It is treated as a noun in apposition.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A45E1771ABFA3F07BF83CAA.taxon	materials_examined	Type material Holotype Philippines • ♂; “ Philippines / Luzon, Pangasinan / San Nicolas / Brgy. Malico // MUSEUM PARIS / 8 - IV- 2024 / SAYap & VThompson // sequencage par Elorde Crispolon C- 00204 // MNHN, Paris EH 31049 ”; MNHN. Paratypes Philippines • 1 ♂; Philippines, “ Mt. Polis S. E. / Elev. 2000 m / 29. V. 1948 / Celestino // UPLBMNH HEM- 04055 ”; UPLBMNH • 1 ♀; “ Philippines / Luzon, Pangasinan / San Nicolas / Brgy. Malico // MUSEUM PARIS / 8 - IV- 2024 / SAYap & VThompson // séquençage par Elorde Crispolon C- 00210 // MNHN, Paris EH 31048 ”; MNHN • 1 ♀; “ Philippines / Luzon, Pangasinan / San Nicolas / Brgy. Malico // MUSEUM PARIS / 8 - IV- 2024 / SAYap & VThompson // UPLBMNH HEM- 06931 ”; UPLBMNH.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A45E1771ABFA3F07BF83CAA.taxon	description	Description Body length 6 – 7 mm (tegmina included), width 3 – 3.5 mm. HEAD (Fig. 2 C). Head including eyes, narrower than widest part of pronotum. In dorsal view, eyes slightly elongated, small ocelli, distance between eyes 9 × ocellus diameter, distance between ocelli equals one ocellar diameter, distance between ocellus and compound eye 3 × ocellus diameter, ocelli closer to each other than to compound eyes. Eyes not prominent, 2 × as long as wide. Vertex and frons longitudinal median carina present. Vertex slightly longer than wide with 3 × ocellus diameter in between two vertex grooves outside ocelli and 4 × ocellus diameter between anterior and posterior vertex margins. No dimple in upper part of postclypeus just below frons (as in Amberana). Postclypeus with longitudinal furrow forming two prominent parallel longitudinal carinae, slightly swollen, ovoid shape in frontal view, widest part at mid-height (Fig. 2 B), not receding prior to anteclypleus where it bends forming obtuse triangle in lateral view (Fig. 2 A). Rostrum very long reaching but not surpassing metacoxae. THORAX. In dorsal view, pronotum with anterior concavities on each side, anterior margin of pronotum as wide as posterior margin of head including eyes, anterolateral margins curved, posterior margin grooved, posterolateral margins slightly concave and longer than anterolateral margin, humeral angle rounded. Pronotum angle in lateral view around 45 ° (Fig. 2 A). Scutellum slightly longer than wide with large median concavity. TEGMEN. R bifurcates on apical half, M bifurcates on basal third, veins not thickened on first ⅔, become thickened on apical third, forming concave apical cells (Fig. 2 C). POSTERIOR WING. Rp separating from SC + RA nearly at midlength, r-m absent, M reaches ambient vein, CuA and CuP fused at base, CuA bifurcate in CuA 1 and CuA 2 at apical third, m-cu links M to CuA above CuA bifurcation, total of six longitudinal veins and five apical cells between SC + RA and CuP, angular protrusion of the costal margin near base present (Fig. 3 B). LEGS. Metafemur with apical spine on inner margin, metatibia bearing 1 lateral spine, both primary and secondary basitarsi with one row of at least five spines. MALE TERMINALIA. In lateral view, posterior margin of pygofer (Fig. 2 E) convex medially with slight curvature towards last third. Subgenital plates (Fig. 2 E – F) finger-shaped, with equal length relative to height of pygofer without longated, thin apical appendage extension (as in Mioscarta), ventral and dorsal margins regularly rounded, apex of dorsal margin dented, sterno-lateral plate present and slightly elongated with apical end of dorsal margin pointed, intermediate plates present, roughly Y-shaped, linking internal sides of sterno-lateral plate and subgenital plate. Paramere (Fig. 2 G), dorsal margin slightly convex ⅔ of its length then becoming concave before finishing up with spiniform process directed dorsally, ventral margin convex and apex with one larger and two minute spiniform processes directed antero-ventrally. Aedeagus (Fig. 2 H), basal third of dorsal margin slightly bent and last ⅔ vertical and S-shaped, basal part of the erected part of aedeagus slightly enlarged, apical extension (ae) directed ventrally, posterior protrusion (pp) elongated directed dorsally, when viewed posteriorly, sub apex flattened, apical part with fine long appendages (Fig. 2 I). COLOUR. Body covered with golden yellow setae. Head, thorax, rostrum, antennal scape, pedicel, tegmen, legs yellowish brown, tegmen apical part much lighter and translucent.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A45E1771ABFA3F07BF83CAA.taxon	materials_examined	Type locality Philippines: Luzon, Pangasinan, San Nicolas.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A45E1771ABFA3F07BF83CAA.taxon	distribution	Distribution Philippines: Luzon Island (Fig. 5). Molecular phylogeny The result of the ML analysis with a likelihood score of - 18712.403 is shown in Figure 4. In the phylogeny, the ingroup appears as monophyletic, supported by an ML bootstrap value (ML BS) of 98.1 / 98 % (SH-aLRT support / standard bootstrap support). Miosterpa Crsipolon & Yap gen. nov. is recovered as a monophyletic group which is supported by an ML BS of 64.3 / 88 %. Is recovered as sister group of Poeciloterpa supported by an ML BS of 97.2 / 98 %. The genera Mioscarta and Trigonoschema, each represented in the phylogeny by only one specimen, are recovered together as monophyletic with an ML BS of 46.6 / 67 %. These four closely related genera: (( Mioscarta + Trigonoschema) + (Miosterpa + Poeciloterpa )), form a clade, supported by an ML BS of 89 / 81 %. For the other genera used as ingroup here, the relationships recovered corroborate the work of Crispolon et al. (2023), even with the much smaller number of terminals used in this study. The clade (Wawi + (Aufidus + (Euryaulax + (Phantagma + (Jacobsoniella + Vinpietri ))))) was recovered as sister to (( Trigonoschema + Mioscarta) + (Miosterpa Crsipolon & Yap gen. nov. + Poeciloterpa )), and the clade (Eoscarta + (Callitettix + (Sounama + Kanozata ))) is sister to this clade. While the clade (Paraliterna sp. + Colsa costaestriga) was recovered at the base of the ingroup.	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
0384A6263A5FE1741BD5A7D07C393EC9.taxon	discussion	We described Trigonoschema as a new genus with three new species and two new species of Mioscarta from the Philippines in 2021 (Crispolon et al. 2021). In that paper, we wrote the taxonomy specifically for Mioscarta, mentioning Poeciloterpina as its subtribe. In the discussion, we mentioned that we would not classify Trigonoschema in any subtribe, even with its proximity to Mioscarta. However, we did not write the taxonomy specific to Trigonoschema suggesting by inference that we followed the taxonomy written for Mioscarta. This caused confusion for some readers and was critiqued by Packer (2025). Here, based on the morphological characters and phylogenetic analyses (Crispolon et al. 2023, and this study), we place Trigonoschema in the subtribe Poeciloterpina together with Miosterpa Crsipolon & Yap gen. nov., Mioscarta, and Poeciloterpa. This makes four Poeciloterpine genera found in the Philippines, three of which are endemic. The following key characters distinguish the subtribe Poeciloterpina from other Cercopidae: radial veins of tegmina branching but are not fused by a transverse vein, the median vein of the posterior wing bifurcating (Schmidt 1920; Lallemand 1949).	en	Jr Crispolon, Elorde S., Yap, Sheryl A., Soulier-Perkins, Adeline (2026): New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema. European Journal of Taxonomy 1052: 82-104, DOI: 10.5852/ejt.2026.1052.3270, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443
