taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03871F323B59FE1F223C1BA02F8887FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501198/files/figure.png	https://doi.org/10.5281/zenodo.19501198	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B59FE1F223C1BA02F8887FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501202/files/figure.png	https://doi.org/10.5281/zenodo.19501202	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B59FE1F223C1BA02F8887FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501210/files/figure.png	https://doi.org/10.5281/zenodo.19501210	FIG. 3. — Anterior ventral views: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂, Brazil, Cabo Frio, Tavares det., USU-295; B, Gecarcoidea lalandii H. Milne Edwards,1837,♂, Costa Rica, Bat Island, AMS-P.11710; C, Johngarthia lagostoma (H. Milne Edwards,1837),♂, Trindade Island, Tavares det., MZUSP-9583; D, Grapsodes notatus Heller, 1865, ♂, Sumatra, Poeloe Babi Island, det. Epigrapsus notatus, RMNH-2131. Scale bars: 10 mm. Courtesy Marcos Tavares.	FIG. 3. — Anterior ventral views: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂, Brazil, Cabo Frio, Tavares det., USU-295; B, Gecarcoidea lalandii H. Milne Edwards,1837,♂, Costa Rica, Bat Island, AMS-P.11710; C, Johngarthia lagostoma (H. Milne Edwards,1837),♂, Trindade Island, Tavares det., MZUSP-9583; D, Grapsodes notatus Heller, 1865, ♂, Sumatra, Poeloe Babi Island, det. Epigrapsus notatus, RMNH-2131. Scale bars: 10 mm. Courtesy Marcos Tavares.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B59FE1F223C1BA02F8887FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19560844/files/figure.png	https://doi.org/10.5281/zenodo.19560844	FIG. 4. — Mxp3,internal view,palp and exopod:A, Cardisoma guanhumi Latreille,in Latreille,Le Peletier,Serville & Guérin,1828,♀, Brazil,Niterõi,Lagoa de Itaipu, USU-294; B, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, Montego Bay, ♂, MZUSP 18501 (ex USU-279); C, Grapsodes notatus Heller, 1865, ♂, Bismarck Archipelago, New Britain, det. Epigrapsus notatus, MHa-K 5715. Scale bars: A, B, 5 mm; C, 1 mm. Courtesy Marcos Tavares.	FIG. 4. — Mxp3,internal view,palp and exopod:A, Cardisoma guanhumi Latreille,in Latreille,Le Peletier,Serville & Guérin,1828,♀, Brazil,Niterõi,Lagoa de Itaipu, USU-294; B, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, Montego Bay, ♂, MZUSP 18501 (ex USU-279); C, Grapsodes notatus Heller, 1865, ♂, Bismarck Archipelago, New Britain, det. Epigrapsus notatus, MHa-K 5715. Scale bars: A, B, 5 mm; C, 1 mm. Courtesy Marcos Tavares.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B59FE1F223C1BA02F8887FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501227/files/figure.png	https://doi.org/10.5281/zenodo.19501227	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B59FE1F223C1BA02F8887FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501232/files/figure.png	https://doi.org/10.5281/zenodo.19501232	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B59FE1F223C1BA02F8887FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501213/files/figure.png	https://doi.org/10.5281/zenodo.19501213	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B59FE1F223C1BA02F8887FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501225/files/figure.png	https://doi.org/10.5281/zenodo.19501225	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B5BFE1A20A81D062AD680D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501198/files/figure.png	https://doi.org/10.5281/zenodo.19501198	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B5BFE1A20A81D062AD680D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501202/files/figure.png	https://doi.org/10.5281/zenodo.19501202	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B5BFE1A20A81D062AD680D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501213/files/figure.png	https://doi.org/10.5281/zenodo.19501213	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B5BFE1A20A81D062AD680D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501219/files/figure.png	https://doi.org/10.5281/zenodo.19501219	FIG. 6. — Ventral view,with male pleon:A, Discoplax longipes A.Milne Edwards,1867,♂ 32.4 × 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), 1 ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 6. — Ventral view,with male pleon:A, Discoplax longipes A.Milne Edwards,1867,♂ 32.4 × 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), 1 ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B5BFE1A20A81D062AD680D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501225/files/figure.png	https://doi.org/10.5281/zenodo.19501225	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B5BFE1A20A81D062AD680D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501227/files/figure.png	https://doi.org/10.5281/zenodo.19501227	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B5BFE1A20A81D062AD680D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501210/files/figure.png	https://doi.org/10.5281/zenodo.19501210	FIG. 3. — Anterior ventral views: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂, Brazil, Cabo Frio, Tavares det., USU-295; B, Gecarcoidea lalandii H. Milne Edwards,1837,♂, Costa Rica, Bat Island, AMS-P.11710; C, Johngarthia lagostoma (H. Milne Edwards,1837),♂, Trindade Island, Tavares det., MZUSP-9583; D, Grapsodes notatus Heller, 1865, ♂, Sumatra, Poeloe Babi Island, det. Epigrapsus notatus, RMNH-2131. Scale bars: 10 mm. Courtesy Marcos Tavares.	FIG. 3. — Anterior ventral views: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂, Brazil, Cabo Frio, Tavares det., USU-295; B, Gecarcoidea lalandii H. Milne Edwards,1837,♂, Costa Rica, Bat Island, AMS-P.11710; C, Johngarthia lagostoma (H. Milne Edwards,1837),♂, Trindade Island, Tavares det., MZUSP-9583; D, Grapsodes notatus Heller, 1865, ♂, Sumatra, Poeloe Babi Island, det. Epigrapsus notatus, RMNH-2131. Scale bars: 10 mm. Courtesy Marcos Tavares.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B5EFE0523FF1820291584C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501198/files/figure.png	https://doi.org/10.5281/zenodo.19501198	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B5EFE0523FF1820291584C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501256/files/figure.png	https://doi.org/10.5281/zenodo.19501256	FIG. 15. — Cardisomatidae n. fam., living and in the field:A-C, Cardisoma crassum Smith, 1870, Coiba I., Panama; D-F, Cardisoma guanhumi Latreille,in Latreille, Le Peletier, Serville & Guérin, 1828; D-E, Bocas del Toro, Panama; F, Ceará, Brazil. Credits: A-E, A. Anker; F, P.P.G. Pachelle.	FIG. 15. — Cardisomatidae n. fam., living and in the field:A-C, Cardisoma crassum Smith, 1870, Coiba I., Panama; D-F, Cardisoma guanhumi Latreille,in Latreille, Le Peletier, Serville & Guérin, 1828; D-E, Bocas del Toro, Panama; F, Ceará, Brazil. Credits: A-E, A. Anker; F, P.P.G. Pachelle.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B5EFE0523FF1820291584C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501210/files/figure.png	https://doi.org/10.5281/zenodo.19501210	FIG. 3. — Anterior ventral views: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂, Brazil, Cabo Frio, Tavares det., USU-295; B, Gecarcoidea lalandii H. Milne Edwards,1837,♂, Costa Rica, Bat Island, AMS-P.11710; C, Johngarthia lagostoma (H. Milne Edwards,1837),♂, Trindade Island, Tavares det., MZUSP-9583; D, Grapsodes notatus Heller, 1865, ♂, Sumatra, Poeloe Babi Island, det. Epigrapsus notatus, RMNH-2131. Scale bars: 10 mm. Courtesy Marcos Tavares.	FIG. 3. — Anterior ventral views: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂, Brazil, Cabo Frio, Tavares det., USU-295; B, Gecarcoidea lalandii H. Milne Edwards,1837,♂, Costa Rica, Bat Island, AMS-P.11710; C, Johngarthia lagostoma (H. Milne Edwards,1837),♂, Trindade Island, Tavares det., MZUSP-9583; D, Grapsodes notatus Heller, 1865, ♂, Sumatra, Poeloe Babi Island, det. Epigrapsus notatus, RMNH-2131. Scale bars: 10 mm. Courtesy Marcos Tavares.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B5EFE0523FF1820291584C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501213/files/figure.png	https://doi.org/10.5281/zenodo.19501213	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B5EFE0523FF1820291584C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501227/files/figure.png	https://doi.org/10.5281/zenodo.19501227	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B41FE0723F01C252E078745.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501198/files/figure.png	https://doi.org/10.5281/zenodo.19501198	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B41FE0723F01C252E078745.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501202/files/figure.png	https://doi.org/10.5281/zenodo.19501202	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B41FE0723F01C252E078745.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501219/files/figure.png	https://doi.org/10.5281/zenodo.19501219	FIG. 6. — Ventral view,with male pleon:A, Discoplax longipes A.Milne Edwards,1867,♂ 32.4 × 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), 1 ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 6. — Ventral view,with male pleon:A, Discoplax longipes A.Milne Edwards,1867,♂ 32.4 × 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), 1 ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B41FE0723F01C252E078745.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501225/files/figure.png	https://doi.org/10.5281/zenodo.19501225	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B41FE0723F01C252E078745.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501227/files/figure.png	https://doi.org/10.5281/zenodo.19501227	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B43FE01271F1CA52E6184A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501198/files/figure.png	https://doi.org/10.5281/zenodo.19501198	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B43FE01271F1CA52E6184A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501202/files/figure.png	https://doi.org/10.5281/zenodo.19501202	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B43FE01271F1CA52E6184A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501213/files/figure.png	https://doi.org/10.5281/zenodo.19501213	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B43FE01271F1CA52E6184A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501219/files/figure.png	https://doi.org/10.5281/zenodo.19501219	FIG. 6. — Ventral view,with male pleon:A, Discoplax longipes A.Milne Edwards,1867,♂ 32.4 × 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), 1 ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 6. — Ventral view,with male pleon:A, Discoplax longipes A.Milne Edwards,1867,♂ 32.4 × 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), 1 ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B43FE01271F1CA52E6184A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501225/files/figure.png	https://doi.org/10.5281/zenodo.19501225	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B43FE01271F1CA52E6184A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501227/files/figure.png	https://doi.org/10.5281/zenodo.19501227	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B45FE0D20551FC529D181BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501198/files/figure.png	https://doi.org/10.5281/zenodo.19501198	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B45FE0D20551FC529D181BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501202/files/figure.png	https://doi.org/10.5281/zenodo.19501202	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B45FE0D20551FC529D181BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501254/files/figure.png	https://doi.org/10.5281/zenodo.19501254	FIG. 14. — Gecarcinidae restrict., living and in the field: A, B, Hartnollius quadratus (Saussure, 1853), Coiba I., Panama; C-F, Johngarthia weileri (Sendler, 1912), São Tomé I., São Tomé and Principe. Credits: A. Anker.	FIG. 14. — Gecarcinidae restrict., living and in the field: A, B, Hartnollius quadratus (Saussure, 1853), Coiba I., Panama; C-F, Johngarthia weileri (Sendler, 1912), São Tomé I., São Tomé and Principe. Credits: A. Anker.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B45FE0D20551FC529D181BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501213/files/figure.png	https://doi.org/10.5281/zenodo.19501213	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B45FE0D20551FC529D181BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501219/files/figure.png	https://doi.org/10.5281/zenodo.19501219	FIG. 6. — Ventral view,with male pleon:A, Discoplax longipes A.Milne Edwards,1867,♂ 32.4 × 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), 1 ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 6. — Ventral view,with male pleon:A, Discoplax longipes A.Milne Edwards,1867,♂ 32.4 × 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), 1 ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B45FE0D20551FC529D181BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501225/files/figure.png	https://doi.org/10.5281/zenodo.19501225	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B45FE0D20551FC529D181BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501232/files/figure.png	https://doi.org/10.5281/zenodo.19501232	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B49FE0E22351AC72EF68704.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501198/files/figure.png	https://doi.org/10.5281/zenodo.19501198	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B49FE0E22351AC72EF68704.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501210/files/figure.png	https://doi.org/10.5281/zenodo.19501210	FIG. 3. — Anterior ventral views: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂, Brazil, Cabo Frio, Tavares det., USU-295; B, Gecarcoidea lalandii H. Milne Edwards,1837,♂, Costa Rica, Bat Island, AMS-P.11710; C, Johngarthia lagostoma (H. Milne Edwards,1837),♂, Trindade Island, Tavares det., MZUSP-9583; D, Grapsodes notatus Heller, 1865, ♂, Sumatra, Poeloe Babi Island, det. Epigrapsus notatus, RMNH-2131. Scale bars: 10 mm. Courtesy Marcos Tavares.	FIG. 3. — Anterior ventral views: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂, Brazil, Cabo Frio, Tavares det., USU-295; B, Gecarcoidea lalandii H. Milne Edwards,1837,♂, Costa Rica, Bat Island, AMS-P.11710; C, Johngarthia lagostoma (H. Milne Edwards,1837),♂, Trindade Island, Tavares det., MZUSP-9583; D, Grapsodes notatus Heller, 1865, ♂, Sumatra, Poeloe Babi Island, det. Epigrapsus notatus, RMNH-2131. Scale bars: 10 mm. Courtesy Marcos Tavares.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B49FE0E22351AC72EF68704.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19560844/files/figure.png	https://doi.org/10.5281/zenodo.19560844	FIG. 4. — Mxp3,internal view,palp and exopod:A, Cardisoma guanhumi Latreille,in Latreille,Le Peletier,Serville & Guérin,1828,♀, Brazil,Niterõi,Lagoa de Itaipu, USU-294; B, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, Montego Bay, ♂, MZUSP 18501 (ex USU-279); C, Grapsodes notatus Heller, 1865, ♂, Bismarck Archipelago, New Britain, det. Epigrapsus notatus, MHa-K 5715. Scale bars: A, B, 5 mm; C, 1 mm. Courtesy Marcos Tavares.	FIG. 4. — Mxp3,internal view,palp and exopod:A, Cardisoma guanhumi Latreille,in Latreille,Le Peletier,Serville & Guérin,1828,♀, Brazil,Niterõi,Lagoa de Itaipu, USU-294; B, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, Montego Bay, ♂, MZUSP 18501 (ex USU-279); C, Grapsodes notatus Heller, 1865, ♂, Bismarck Archipelago, New Britain, det. Epigrapsus notatus, MHa-K 5715. Scale bars: A, B, 5 mm; C, 1 mm. Courtesy Marcos Tavares.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B49FE0E22351AC72EF68704.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501213/files/figure.png	https://doi.org/10.5281/zenodo.19501213	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B49FE0E22351AC72EF68704.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501232/files/figure.png	https://doi.org/10.5281/zenodo.19501232	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4AFE08206A1C652EFA82DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501202/files/figure.png	https://doi.org/10.5281/zenodo.19501202	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4AFE08206A1C652EFA82DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501210/files/figure.png	https://doi.org/10.5281/zenodo.19501210	FIG. 3. — Anterior ventral views: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂, Brazil, Cabo Frio, Tavares det., USU-295; B, Gecarcoidea lalandii H. Milne Edwards,1837,♂, Costa Rica, Bat Island, AMS-P.11710; C, Johngarthia lagostoma (H. Milne Edwards,1837),♂, Trindade Island, Tavares det., MZUSP-9583; D, Grapsodes notatus Heller, 1865, ♂, Sumatra, Poeloe Babi Island, det. Epigrapsus notatus, RMNH-2131. Scale bars: 10 mm. Courtesy Marcos Tavares.	FIG. 3. — Anterior ventral views: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂, Brazil, Cabo Frio, Tavares det., USU-295; B, Gecarcoidea lalandii H. Milne Edwards,1837,♂, Costa Rica, Bat Island, AMS-P.11710; C, Johngarthia lagostoma (H. Milne Edwards,1837),♂, Trindade Island, Tavares det., MZUSP-9583; D, Grapsodes notatus Heller, 1865, ♂, Sumatra, Poeloe Babi Island, det. Epigrapsus notatus, RMNH-2131. Scale bars: 10 mm. Courtesy Marcos Tavares.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4AFE08206A1C652EFA82DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501219/files/figure.png	https://doi.org/10.5281/zenodo.19501219	FIG. 6. — Ventral view,with male pleon:A, Discoplax longipes A.Milne Edwards,1867,♂ 32.4 × 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), 1 ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 6. — Ventral view,with male pleon:A, Discoplax longipes A.Milne Edwards,1867,♂ 32.4 × 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), 1 ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4AFE08206A1C652EFA82DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501225/files/figure.png	https://doi.org/10.5281/zenodo.19501225	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4AFE08206A1C652EFA82DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501232/files/figure.png	https://doi.org/10.5281/zenodo.19501232	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4DFE0A23A81B802E6D82D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501198/files/figure.png	https://doi.org/10.5281/zenodo.19501198	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4DFE0A23A81B802E6D82D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501254/files/figure.png	https://doi.org/10.5281/zenodo.19501254	FIG. 14. — Gecarcinidae restrict., living and in the field: A, B, Hartnollius quadratus (Saussure, 1853), Coiba I., Panama; C-F, Johngarthia weileri (Sendler, 1912), São Tomé I., São Tomé and Principe. Credits: A. Anker.	FIG. 14. — Gecarcinidae restrict., living and in the field: A, B, Hartnollius quadratus (Saussure, 1853), Coiba I., Panama; C-F, Johngarthia weileri (Sendler, 1912), São Tomé I., São Tomé and Principe. Credits: A. Anker.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4DFE0A23A81B802E6D82D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501202/files/figure.png	https://doi.org/10.5281/zenodo.19501202	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4DFE0A23A81B802E6D82D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501232/files/figure.png	https://doi.org/10.5281/zenodo.19501232	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4DFE0A23A81B802E6D82D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501213/files/figure.png	https://doi.org/10.5281/zenodo.19501213	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4FFE3722031EC42AF382DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501198/files/figure.png	https://doi.org/10.5281/zenodo.19501198	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4FFE3722031EC42AF382DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501202/files/figure.png	https://doi.org/10.5281/zenodo.19501202	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4FFE3722031EC42AF382DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501219/files/figure.png	https://doi.org/10.5281/zenodo.19501219	FIG. 6. — Ventral view,with male pleon:A, Discoplax longipes A.Milne Edwards,1867,♂ 32.4 × 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), 1 ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 6. — Ventral view,with male pleon:A, Discoplax longipes A.Milne Edwards,1867,♂ 32.4 × 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), 1 ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4FFE3722031EC42AF382DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501225/files/figure.png	https://doi.org/10.5281/zenodo.19501225	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4FFE3722031EC42AF382DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501232/files/figure.png	https://doi.org/10.5281/zenodo.19501232	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B4FFE3722031EC42AF382DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501254/files/figure.png	https://doi.org/10.5281/zenodo.19501254	FIG. 14. — Gecarcinidae restrict., living and in the field: A, B, Hartnollius quadratus (Saussure, 1853), Coiba I., Panama; C-F, Johngarthia weileri (Sendler, 1912), São Tomé I., São Tomé and Principe. Credits: A. Anker.	FIG. 14. — Gecarcinidae restrict., living and in the field: A, B, Hartnollius quadratus (Saussure, 1853), Coiba I., Panama; C-F, Johngarthia weileri (Sendler, 1912), São Tomé I., São Tomé and Principe. Credits: A. Anker.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B73FE3320391EC42A8983D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501213/files/figure.png	https://doi.org/10.5281/zenodo.19501213	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B73FE3320391EC42A8983D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501225/files/figure.png	https://doi.org/10.5281/zenodo.19501225	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B73FE3320391EC42A8983D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501227/files/figure.png	https://doi.org/10.5281/zenodo.19501227	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B73FE3320391EC42A8983D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501232/files/figure.png	https://doi.org/10.5281/zenodo.19501232	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B73FE3320391EC42A8983D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501202/files/figure.png	https://doi.org/10.5281/zenodo.19501202	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B77FE3D23901901291F86BA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501202/files/figure.png	https://doi.org/10.5281/zenodo.19501202	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 2. — Frontal view. A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN- B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Tuerkayana aff. hirtipes (Dana, 1851), ♂ 61.0 × 79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (= MNHN-B24811); D, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; E, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). F, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHNB24656). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B77FE3D23901901291F86BA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501198/files/figure.png	https://doi.org/10.5281/zenodo.19501198	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 1. — Dorsal views of carapace: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN- IU-2013-14983 (= MNHN-B12270); B, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands,Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);C, Tuerkayana rotundum (Quoy & Gaimard,1824), ♀ 32.7 × 40.1 mm, Loyalty Islands,MNHN-IU-2013-3740 (= MNHN-B13141);D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155);E, Hartnollius lateralis (Fréminville in Guérin, 1832), ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656); F, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B77FE3D23901901291F86BA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501219/files/figure.png	https://doi.org/10.5281/zenodo.19501219	FIG. 6. — Ventral view,with male pleon:A, Discoplax longipes A.Milne Edwards,1867,♂ 32.4 × 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), 1 ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 6. — Ventral view,with male pleon:A, Discoplax longipes A.Milne Edwards,1867,♂ 32.4 × 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), 1 ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B77FE3D23901901291F86BA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501225/files/figure.png	https://doi.org/10.5281/zenodo.19501225	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B77FE3D23901901291F86BA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501227/files/figure.png	https://doi.org/10.5281/zenodo.19501227	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B79FE3F23901DA62927865A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501213/files/figure.png	https://doi.org/10.5281/zenodo.19501213	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B79FE3F23901DA62927865A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501225/files/figure.png	https://doi.org/10.5281/zenodo.19501225	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 7. — Ventral view of males, without pleon: A, Discoplax longipes A. Milne Edwards, 1867, ♂ 32.4 × 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN- IU-2008-11402 (= MNHN-B24815); B, Tuerkayana magnum (Ng & Shih, 2014), ♂ 52.8 × 64.9 mm, Indonesia, Java, MNHN-IU-2018-5239 (ex ZRC 2017.1393); C, Gecarcoidea lalandii H. Milne Edwards, 1837, ♂ 52.0 × 73.3 mm, Papua New Guinea, MNHN-IU-2013-13254; D, Johngarthia weileri (Sendler, 1912), ♂ 54.5 × 71.4 mm, Gulf of Guinea, Principe Island, MNHN-IU-2018-5230 (= MNHN-B13151). Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B79FE3F23901DA62927865A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501227/files/figure.png	https://doi.org/10.5281/zenodo.19501227	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 8. — Female pleon of Cardisomatidae n. fam.: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♀ 60.0 × 75.6 mm, Guadeloupe, îles des Saintes, MNHN-IU-2024-4621; B, Cardisoma carnifex (Herbst, 1796), ♀ 56.5 × 66.3 mm, Wallis, MNHN-IU-2013-2201; C, Tuerkayana celeste (Ng & Davie, 2012), paratype, ♀ 67.3 × 82.0 mm, Christmas Island; Hosnie’s Springs, MNHN-IU-2018-5238 (ex ZRC 2012.0015); D, Discoplax longipes A. Milne Edwards, 1867, ♀ 32.6 × 37 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815), the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B79FE3F23901DA62927865A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501232/files/figure.png	https://doi.org/10.5281/zenodo.19501232	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B79FE3F23901DA62927865A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501235/files/figure.png	https://doi.org/10.5281/zenodo.19501235	FIG. 10. — Female pleopods of mature Cardisoma armatum (Herklots, 1851),♀ cw 60.4 mm,Liberia (NMU 2008a).Anterior view:A, PL 2; B, PL 3; C, PL 4; D, PL 5. Posterior view: E, PL 2; F, PL 3; G, PL 4; H, PL 5. Scale bar: 10 mm. (After Cumberlidge et al. 2021: fig. 13).	FIG. 10. — Female pleopods of mature Cardisoma armatum (Herklots, 1851),♀ cw 60.4 mm,Liberia (NMU 2008a).Anterior view:A, PL 2; B, PL 3; C, PL 4; D, PL 5. Posterior view: E, PL 2; F, PL 3; G, PL 4; H, PL 5. Scale bar: 10 mm. (After Cumberlidge et al. 2021: fig. 13).	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B79FE3F23901DA62927865A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501256/files/figure.png	https://doi.org/10.5281/zenodo.19501256	FIG. 15. — Cardisomatidae n. fam., living and in the field:A-C, Cardisoma crassum Smith, 1870, Coiba I., Panama; D-F, Cardisoma guanhumi Latreille,in Latreille, Le Peletier, Serville & Guérin, 1828; D-E, Bocas del Toro, Panama; F, Ceará, Brazil. Credits: A-E, A. Anker; F, P.P.G. Pachelle.	FIG. 15. — Cardisomatidae n. fam., living and in the field:A-C, Cardisoma crassum Smith, 1870, Coiba I., Panama; D-F, Cardisoma guanhumi Latreille,in Latreille, Le Peletier, Serville & Guérin, 1828; D-E, Bocas del Toro, Panama; F, Ceará, Brazil. Credits: A-E, A. Anker; F, P.P.G. Pachelle.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B7CFE3A22081AA72BB8865A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501237/files/figure.png	https://doi.org/10.5281/zenodo.19501237	FIG. 11. — Epigrapsus politus Heller, 1862, ♂ 23.0 × 29.1 mm, Sumatra,Atjeh, MNHN-IU-2017-10018 (= MNHN-B12585):A, dorsal view; B, anterior ventral view; C, ventral view, with pleon; D, G1 and G2; E, hydrophilic setal tufts; F, thoracic sternum, with penis. Scale bars: A, C, 10 mm; B, F, 5 mm; D, E, 1 mm. Credits: MNHN-Soubzmaigne.	FIG. 11. — Epigrapsus politus Heller, 1862, ♂ 23.0 × 29.1 mm, Sumatra,Atjeh, MNHN-IU-2017-10018 (= MNHN-B12585):A, dorsal view; B, anterior ventral view; C, ventral view, with pleon; D, G1 and G2; E, hydrophilic setal tufts; F, thoracic sternum, with penis. Scale bars: A, C, 10 mm; B, F, 5 mm; D, E, 1 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B7EFE2620AF18E22E7782DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501210/files/figure.png	https://doi.org/10.5281/zenodo.19501210	FIG. 3. — Anterior ventral views: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂, Brazil, Cabo Frio, Tavares det., USU-295; B, Gecarcoidea lalandii H. Milne Edwards,1837,♂, Costa Rica, Bat Island, AMS-P.11710; C, Johngarthia lagostoma (H. Milne Edwards,1837),♂, Trindade Island, Tavares det., MZUSP-9583; D, Grapsodes notatus Heller, 1865, ♂, Sumatra, Poeloe Babi Island, det. Epigrapsus notatus, RMNH-2131. Scale bars: 10 mm. Courtesy Marcos Tavares.	FIG. 3. — Anterior ventral views: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂, Brazil, Cabo Frio, Tavares det., USU-295; B, Gecarcoidea lalandii H. Milne Edwards,1837,♂, Costa Rica, Bat Island, AMS-P.11710; C, Johngarthia lagostoma (H. Milne Edwards,1837),♂, Trindade Island, Tavares det., MZUSP-9583; D, Grapsodes notatus Heller, 1865, ♂, Sumatra, Poeloe Babi Island, det. Epigrapsus notatus, RMNH-2131. Scale bars: 10 mm. Courtesy Marcos Tavares.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B7EFE2620AF18E22E7782DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19560844/files/figure.png	https://doi.org/10.5281/zenodo.19560844	FIG. 4. — Mxp3,internal view,palp and exopod:A, Cardisoma guanhumi Latreille,in Latreille,Le Peletier,Serville & Guérin,1828,♀, Brazil,Niterõi,Lagoa de Itaipu, USU-294; B, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, Montego Bay, ♂, MZUSP 18501 (ex USU-279); C, Grapsodes notatus Heller, 1865, ♂, Bismarck Archipelago, New Britain, det. Epigrapsus notatus, MHa-K 5715. Scale bars: A, B, 5 mm; C, 1 mm. Courtesy Marcos Tavares.	FIG. 4. — Mxp3,internal view,palp and exopod:A, Cardisoma guanhumi Latreille,in Latreille,Le Peletier,Serville & Guérin,1828,♀, Brazil,Niterõi,Lagoa de Itaipu, USU-294; B, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, Montego Bay, ♂, MZUSP 18501 (ex USU-279); C, Grapsodes notatus Heller, 1865, ♂, Bismarck Archipelago, New Britain, det. Epigrapsus notatus, MHa-K 5715. Scale bars: A, B, 5 mm; C, 1 mm. Courtesy Marcos Tavares.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B7EFE2620AF18E22E7782DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501232/files/figure.png	https://doi.org/10.5281/zenodo.19501232	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 9. — Female pleon of Gecarcinidae restrict. (A-E) and of Grapsodes Heller, 1865 (F): A, Gecarcinus ruricola (Linnaeus, 1758), ovigerous ♀ 52.8 × 69.9 mm, Cuba, MNHN-IU-2024-6554; B, Johngarthia oceanica (Perger, 2019), ♀ 54.6 × 69.1 mm, Clipperton, MNHN-IU-2025-8003 (= MNHN-B28823); C, Hartnollius lateralis (Fréminville in Guérin,1832), ♀ 32.6 × 41.8 mm, Guadeloupe,MNHN-IU-2017-8390 (= MNHN-B24656);D, Hartnollius nobilii (Perger & Wall, 2014),paratype, ♀ 36.6 × 44.3 mm, Ecuador, St Helena, MNHN-IU-2014-11211 (= MNHN-B12314); E, Gecarcoidea natalis (Pocock, 1888), ♀ 58.2 × 78.7 mm, Christmas Island, MNHN-IU-2025-8001 (ex ZRC 2012.0177); F, Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241. In A, B, the setose pleopods (see arrows) are visible on each side of the pleon. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B7EFE2620AF18E22E7782DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501242/files/figure.png	https://doi.org/10.5281/zenodo.19501242	FIG. 12. — Grapsodes notatus Heller,1865,♂ 22.7 × 26.9 mm,Taiwan,MNHN-IU-2018-5241 (ex ZRC 2013.003):A, dorsal view;B, anterior ventral view;C, pouch of hydrophilic setae between P3/P4;D, thoracic sternum and G1;E, ventral view,with pleon.Scale bars:A,E, 10 mm;B-D,2 mm.Credits:MNHN-Soubzmaigne.	FIG. 12. — Grapsodes notatus Heller,1865,♂ 22.7 × 26.9 mm,Taiwan,MNHN-IU-2018-5241 (ex ZRC 2013.003):A, dorsal view;B, anterior ventral view;C, pouch of hydrophilic setae between P3/P4;D, thoracic sternum and G1;E, ventral view,with pleon.Scale bars:A,E, 10 mm;B-D,2 mm.Credits:MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B7EFE2620AF18E22E7782DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501252/files/figure.png	https://doi.org/10.5281/zenodo.19501252	FIG. 13. — Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241: A, dorsal view; B, ventral view; C, thoracic sternum and vulvae. Scale bars: A, B, 10 mm; C, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 13. — Grapsodes notatus Heller, 1865, ♀ 18.1 × 21.8 mm, Taiwan, MNHN-IU-2018-5241: A, dorsal view; B, ventral view; C, thoracic sternum and vulvae. Scale bars: A, B, 10 mm; C, 2 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
03871F323B7EFE2620AF18E22E7782DE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19501213/files/figure.png	https://doi.org/10.5281/zenodo.19501213	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 5. — Male pleon, thoracic sternum, median line, male gonopore with penis, and gonopods: A, Cardisoma guanhumi Latreille, in Latreille, Le Peletier, Serville & Guérin, 1828, ♂ 56.5 × 68.0 mm, Guadeloupe, MNHN-IU-2013-14983 (= MNHN-B12270): pleon; B, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm,MNHN-IU-2017-8392 (= MNHN-B13155): pleon; C, C. guahnumi Latreille,in Latreille,Le Peletier,Serville & Guérin, 1828, MNHN-IU-2013-14983 (= MNHN-B12270): thoracic sternum; D, G. ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, MNHN-IU-2017-8392 (= MNHN-B13155): thoracic sternum; E, Tuerkeyana celeste (Ng & Davie, 2012), ♂ 44.4 × 52.2 mm, Australia, Christmas Island, det. Discoplax celeste, ZRC 2012.0171: gonopods in situ; F, Hartnollius quadratus (Saussure, 1853), ♂ 50.0 × 64.0 mm, Mexico, Estero El Verde, MNHN-IU-2017-8391 (= MNHN-B20900): G1 and G2. Scale bars: A-D, 10 mm; E, 5 mm; F, 2 mm. Credits: MNHN-Soubzmaigne.	2026-03-31	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez Moreno, Paula A.			
