taxonID	type	description	language	source
03F9B178B274FFBB2AE1D86E0107FE6C.taxon	type_taxon	Type species: Hilarographa zapyra Meyrick, 1886 (New Guinea)	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B274FFBB2AE1D86E0107FE6C.taxon	diagnosis	Diagnosis. Wing expanse range, 7.5 – 27.0 mm (average size: 12 – 18 mm). The genus is defined in the tribe and distinguished from Hilarographa by the following particulars: the labial palpi are distinctly dorso-ventrally flattened (tubular in Hilarographa); in the male genitalia, the uncus is basally fused to the tegumen (membranous basally in Hilarographa), the hami are prominent and usually tubular, and the socii are sclerotic and usually slender, little setose, usually basally fused to the hami (socii wide, flattened and highly setose, and not fused to the hami in Hilarographa). The female genitalia usually have a bursal diverticular accessory sac, and the signum a bilobed fan-like form of numerous long spines curved inwardly, rarely absent or reduced (usually a scobinate patch in Hilarographa). Descriptive notes. Heppner et al. (2025) gave a detailed descriptive review of the generic characters for Thaumatographa. The main distinctive features include the small head (noticeably smaller than most tortricids), the upturned, smooth-scaled and typically flattened labial palpus (mostly porrect in other tortricids, with the apical palpal segment usually somewhat decumbent, and mid-segment ventral scale tufts), and the very colorful forewing maculation (many species with orange markings) and somber hindwings (but often with a median orange diffusion going to the wing base); antennae are usually short (less than half forewing length) and with short ventral cilia (rarely longer) in males. The male genitalia have the uncus usually narrow, strongly sclerotic and bent, and basally fused with the tegumen (membranous joint in Hilarographa, all Neotropical plus two species in North America); paired hami usually sclerotic and slender (but rarely more flattened); a long pair of setose socii (but rarely short and stubby); the gnathos is usually short, ovate to quadrate, and pendulate; the valva typically simple and elongated, usually without ornamentation except for a single dense setal patch; a transtilla band or less frequently more elaborated into a complex structure (rarely stubby and incomplete); the juxta usually a small ovate plate, often slightly caudally concave and the dorsal margin truncated; the anellus usually a membranous ring; the aedeagus usually narrow and tubular (rarely short), with one long or few cornuti, phallobase usually prominent and bulbous proximally (rarely undeveloped), and the ductus ejaculatorius usually is long and with a proximal hood (rarely short); and the vinculum usually triangular with a little developed or stubby saccus. The abdomen of males typically has a pair of lateral coremata on the pregenital sternum, which is usually somewhat caudally invaginated. The female genitalia typically have a short ovipositor, and very long and thin ductus bursae (rarely short); the sterigma is usually simple and not ornamented; the bursa is usually asymmetrically ovate, with an accessory sac on a short tubular section adjacent to both the ductus bursae juncture and the ductus seminalis juncture from one end of the bursa, thus all three of their tubules approximate from the bursa (but some species have the ductus seminalis from the caudal end of the ductus bursae).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B274FFBB2AE1D86E0107FE6C.taxon	biology_ecology	Biology. Mostly unknown, but adults appear to all be diurnally active, yet may come to lights. Larvae are borers of shoots and roots as far as is known: Thaumatographa caminodes (Meyrick) feeding on roots of cardamom (Elettaria cardamomum) and also a wild Zingiberaceae in Sri Lanka (Diakonoff 1986), T. leucopyrga (Meyrick) in shoots of Ardisia sieboldi (Myrsinaceae) in Java (Indonesia), and T. oenobapta Diakonoff in calyxes of Ixora (Rubiaceae), are the only examples known, but T. leucopyrga is not a typical member of the genus. The other species with biological data that have been included among Thaumatographa (Diakonoff & Arita 1981; Heppner 1983, 2025 a), appear to all be pine trunk feeders (cambium layer) but are atypical and will be transferred to their own genus. None of the other more tropical species of the genus have any host data available (likewise for species of Hilarographa from the Neotropics), so we do not know what tropical trees they may feed on, but perhaps the other Asian species are also on Myrsinaceae, Rubiaceae, and Zingiberaceae hosts. The only larva and pupa of Hilarographini with detailed descriptions are for T. eremnotorna in Japan, with details described by Diakonoff & Arita (1981), but this species will be transferred to a related new genus of northern pine-feeding species (Heppner 2025 a). The larval characters described for this species, nonetheless, may have the main characters applicable also to the tribe, since larvae of all the species are thought to be borers. The primary distinction of this borer larva from Japan is the bisetose state of the prothoracic L-group setae, normally trisetose in all other Tortricidae and including the other group of borer larvae in the subfamily Olethreutinae (Horak 1984, 1991, 1998; MacKay 1959, 1962, 1963).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B274FFBB2AE1D86E0107FE6C.taxon	distribution	Distribution. South Asia (India and Sri Lanka) to Southeast Asia, from Thailand to Vietnam and Indonesia (including Borneo, Java, Moluccas, Sulawesi, and Sumatra, and undoubtedly the other islands as well), north to southern China (Hainan and Hong Kong), Taiwan and Japan, and in northeastern Australia (Queensland) and the South Pacific (New Guinea and Solomons, and an undescribed species from Tahiti) (Diakonoff 1986; Heppner 2025 b, c; Razowski 2009). The genus is undoubtedly also in the Philippines but has not been recorded there yet (Diakonoff 1968). Three species are recorded in North America (Heppner 1983), and one related pine-feeder from Cuba, but these are partially in a related new genus while two are in the genus Hilarographa. The pine feeders in the related genus occur in Japan, the Kuril Islands (Russian Far East) and North America (plus one in Cuba) (Diakonoff 1986; Heppner 1983, 2025 a).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B274FFBB2AE1D86E0107FE6C.taxon	discussion	Discussion. Most species of the genus are known from only single or very few specimens. The 2 - bristle frenulum of female Thaumatographa conforms to this morphology in Hilarographini and for the subfamily Chlidanotinae, versus the 3 - bristle norm for the remainder of tortricids (Yang & Brown 2009). A thin ovate dorsal flap at the middle of tergite 8 in females is present in some Thaumatographa species but the function of this structure is unknown. Likewise, some species have a small bulbous appendage sac (or pouch) ventrally at the base of the ostial funnel at the caudal end of the ductus bursae, seemingly emergent approximate to the antrum (sometimes even distally bifid), but the function of this sac or pouch is also unknown. The small head and upcurved labial palpi also are very typical of hilarographines but very atypical for Tortricidae, thus the century-long placement (from 1877 to 1977) of the genus among the so-called ‘ Glyphipterigidae’ (Diakonoff 1977 b; Heppner 1978, 1982 a, b). Adults often have the labial palpus dorsally iridescent pale blue to metallic blue-silver on the apical segment, depending on light angles. The species of Thaumatographa in Vietnam show an assemblage of the different types of male genitalia found in the genus, as noted below for each new species. With most of Vietnam going a long way south from the northern provinces, it is very likely that many more species of the genus remain undiscovered in central and southern Vietnam. These moths are certainly very rare, and no Hilarographini were reported in the extensive treatment of Vietnam tortricids in several papers by Razowski, summarized in the faunal review of Nedoshivina (2013) (the tribe was included in this review but only for two genera now known to belong to Archipini, so no true Hilarographini have been verified for Vietnam until now); and likewise, the older lists of Vietnam moths did not include any hilarographine tortricids (Candèze 1927, Joannis 1928 – 30).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B276FFBE2AE1DC8E0009FEA4.taxon	description	(Figs. 1, 5)	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B276FFBE2AE1DC8E0009FEA4.taxon	materials_examined	Type material. Holotype. [VIETNAM] ♂, Cuc Phuong N. P. (Mac Lake) (155 m), Ninh Binh Prov., 4 – 7 May 2009, J. B. Heppner (gen. slide JBH- 4648; adult photo 12691) (MGCL).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B276FFBE2AE1DC8E0009FEA4.taxon	diagnosis	Diagnosis. This species is distinctive in its large orange subapical bar, slightly curved. The male genitalia have the aedeagus with a long, thin cornutus (almost as long as the entire aedeagus) with a large apical recurved hook. The forewing maculation and male genitalia most resemble those of the Taiwan species, T. mesostigmatias Diakonoff (1977 b), but the new species in Vietnam is smaller and has the subterminal black spots diffuse (prominent in T. mesostigmatias) and the central angled striae from the dorsal margin are orange (white in T. mesostigmatias).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B276FFBE2AE1DC8E0009FEA4.taxon	description	Description. Wing expanse: 12.8 mm (n = 1). Head (Fig. 1 a): mostly yellow, with black-brown on distally on vertex and on frontal eye margins, light yellow ventrally; antenna thick, and gray-brown, with short ventral cilia in male; labial palpus white, with yellow and dark brown at base and dorsally on middle segment, with dark brown on caudal surface of apical segment; head venter light yellow. Thorax (Fig. 1): dark brown, mixed with dull yellow as lateral longitudinal line each side, and merged to yellow-suffused metathorax; tegula black-brown, margined with dull yellow; venter white; legs white, with dark brown bars on midtibia at each end, and hindtibia dark brown-gray, with tarsi white bisected by black-brown bars; legs mesally white; leg spurs white. Forewing (Fig. 1): black-brown marked with orange-tan striae from costa and dorsal margin angled to termen and three prominent striae convergent medially at mid-wing, with basal stria horizontal to wing base and costal basal stria recurved to base from basal 1 / 4 of wing; costa with two short preapical striae directed toward tornus, the most distal one almost convergent with a sinuous vertical stria from subtornus; a subapical orange bar to termen and ventrally slightly recurved; near apex and thin orange-tan line to near mid-termen and continued along termen to tornus; dorsal margin pale yellow; fringe dark brown (likely pale distally, but specimen fringe too warn to verify), with white along apex; venter dark brown with dorsal markings repeated in pale yellow, but white apical stria and on subapical costal margin, and anal sector dull white. Hindwing (Fig. 1): dark brown, with basal half of wing with a yellow linear patch to wing base and suffused on its margins; anal sector orange-brown; fringe dark brown, slightly pale distally and elongate along anal margin; venter dark brown with light gray-white speckles on distal half and dull yellow in cell to wingbase. Abdomen (Fig. 1): dark brown, with dark tan posterior sternite margins; venter tan-white; genital tufts dark brown, with pale lateral coremata; pregenital segment (Fig. 5 c), with sternite caudal margin blunt-rounded laterally and medially with a U-shaped invagination; short coremata laterally from invaginated corematal pouch each side. Male genitalia (Fig. 5): tegumen an inverted V-shape; uncus (Fig. 5 b), stout, bent caudally from wide base and tapering distally only near somewhat blunt apex; hami (Fig. 5 b), slightly shorter than uncus, strongly sclerotic and bent 90 ° from stout base, slightly curved ventrally near distal end and flattened on distal 2 / 3 with slightly recurved and stout ventral margin before acute apex; socii (Fig. 5 b), short (half uncus length) and thick, densely setose dorsally; gnathos (Fig. 5 b), with flattened arms, convergent medially; transtilla (Fig. 5 b), band-like, convergent medially; juxta (Fig. 5 b), an ovate plate (slightly concave caudally), with dorsal margin truncated and slightly concave, laterally projected narrowly into membranous anellus ring; aedeagus (Fig. 5 a), short, tubular, with spatulate caudal projection and distally acute; phallobase obsolete; cornutus a long thin spine (subequal to aedeagus length), with a distal recurved hook; ductus ejaculatorius short, with globular hood; valva (Fig. 5), elongate, oblong with rounded termen, overall setose and with dense setal patch on costa near base; sacculus slightly rounded; vinculum V-shaped, merging to short and slightly bulbous saccus. Female unknown.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B276FFBE2AE1DC8E0009FEA4.taxon	etymology	Etymology. The species is named for the type-locality, Cuc Phuong N. P., in Ninh Binh Prov.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B276FFBE2AE1DC8E0009FEA4.taxon	biology_ecology	Biology. Unknown. Flight period: May. Habitat (Fig. 10) is lowland tropical forest (Sterling et al. 2006).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B276FFBE2AE1DC8E0009FEA4.taxon	distribution	Distribution. Known only from northern Vietnam.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B276FFBE2AE1DC8E0009FEA4.taxon	discussion	Discussion. In the recent review of Thaumatographa species for Taiwan (Heppner et al. 2025), it was noted that the Taiwan endemic species, T. mesostigmatias, should have its own species-group, once such groups are organized for the genus, based mainly on the unusual features of the female genitalia. We have no female specimens available for T. cucphuongana, but based on the similar maculation and male genitalia of T. mesostigmatias, we suspect the new Vietnam species also belongs with the same species-group of the related species from Taiwan, but await discovery of the female to verify this.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B273FFBF2AE1DCC60666F91F.taxon	description	(Figs. 2, 6)	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B273FFBF2AE1DCC60666F91F.taxon	materials_examined	Type material. Holotype. [VIETNAM] ♂, Cuc Phuong N. P. (Mac Lake) (155 m), Ninh Binh Prov., 4 – 7 May 2009, J. B. Heppner (gen. slide JBH- 4649; adult photo 12689) (MGCL).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B273FFBF2AE1DCC60666F91F.taxon	diagnosis	Diagnosis. This species is distinctive in its maroon-orange forewing tips and basal sector, with the black-brown center interspersed with pale tan-yellow striae medially; distinguished from the similar T. melinhana sp. nov. by its dark brown thorax (brown-orange in T. melinhana) and brown-orange head (yellow in T. melinhana). The male genitalia are distinct in the crab-carapace-like transtilla with a large lateral spine each side (quadratic plate with caudal projections in T. melinhana).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B273FFBF2AE1DCC60666F91F.taxon	description	Description. Wing expanse: 16.6 mm (n = 1). Head (Fig. 2 a): mostly orange, with brown on vertex and frons, yellow ventrally; antenna brown with yellow band each segment, with short ventral cilia in male; labial palpus orange-yellow, with brown dorsally on apical segment; head venter yellow. Thorax (Fig. 2): dark brown, with maroon longitudinal lateral line; tegula dark brown with maroon margins; venter white; legs white-yellow, with black-brown bars, except hindtibia gray and dull white mesally; spurs gray. Forewing (Fig. 2): black-brown medially, with dark maroon basal 1 / 3 and apical 1 / 5; wing base with dark maroon area divided medially by a longitudinal black-brown line and similar line on anal vein, with same on costal margin; median dark region irrorated with curved stria of tan-yellow curved toward apex and ending midwing, all interrupted to form spots and bars; yellow spot at end of cell, basad of a subtornal series of several yellow spots curved toward apex; costa with five striae angled toward tornus, each margined basad with orange and apical three white on the costal margin and blended into silver mesally; wing apical 1 / 5 maroon, with orange at apex and subterminally; termen orange, with three black spots equidistantly spaced; fringe dark brown, with white along apex and on termen below apical 1 / 4; venter dark brown with dosal marks barely evident as dull orange marks, except bright white-yellow on costa near apex and along termen, with anal sector white. Hindwing (Fig. 2): black-brown with pale yellow longitudinal suffusion along cell area to wing base; slight pale yellow to gray along apical section of termen; anal region light tan to light brown; costal sector white from 3 / 4 to wing base; venter brown-gray with a pale yellow terminal line and a white vertical line at end of cell as margin to black cell area to wing base. Abdomen: dark brown, with tan posterior margins of tergites; venter tan-yellow; genital tufts dark brown with tan-yellow distally and ventrally; pregenital segment (Fig. 6 c), with caudal margin of sternite rounded laterally and with a U-shaped median invagination; short coremata laterally from invaginated corematal pouch each side. Male genitalia (Fig. 6): tegumen an inverted V-shape; uncus (Fig. 6 b), stout and short, slightly curved caudad, from small base and uniform to blunt-acute apex; hami (Fig. 6 b), slightly shorter than uncus, sclerotic and narrow from small base, slightly curved basally and slightly narrowing to acute apex; socii (Fig. 6 b), short (half uncus length) and thick, laterally flattened, densely setose; gnathos (Fig. 6 b), with flattened arms, barely convergent medially with knobbed ends; transtilla (Fig. 6 b), a complicated plate-like quadratic structure, with a proximal-dorsal round margin and lateral strong spine each side (like a crab carapace) to valval juncture, and caudally with dentate thorns on a dorsally indented projection, medially with an invaginated margin and a lateral spined process each side; juxta (Fig. 6 b), an ovate plate (slightly concave caudally), with dorsal margin truncated, to membranous anellus ring; aedeagus (Fig. 6 a), tubular of moderate length, with spinate and narrowed caudal projection, distally acute; phallobase large and decumbent; cornutus a long and thin flechette; vesica spinose; ductus ejaculatorius long, with elongated hood; valva (Fig. 6), elongate, oblong with round termen but medially slightly acute, overall setose with longer setae near apical dorsal margin; sacculus nearly straight; vinculum U-shaped, merging to short and blunt saccus (undeveloped). Female unknown.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B273FFBF2AE1DCC60666F91F.taxon	etymology	Etymology. The species is named for the unusual and elaborate transilla of the male genitalia, which has the appearance of the carapace of some crab species, with lateral spines (e. g., blue crabs, Callinectes sp., Decapoda: Portunidae).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B273FFBF2AE1DCC60666F91F.taxon	biology_ecology	Biology. Unknown. Flight period: May. Habitat (Fig. 10) is lowland tropical forest (Sterling et al. 2006).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B273FFBF2AE1DCC60666F91F.taxon	distribution	Distribution. Known only from northern Vietnam.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B273FFBF2AE1DCC60666F91F.taxon	discussion	Discussion. This species is somewhat similar to the following species, but the transtilla of the male genitalia is unique in its form, although the next species has a less developed version of transilla.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B272FFB12AE1DA1F0742FAD4.taxon	description	(Figs. 3, 7)	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B272FFB12AE1DA1F0742FAD4.taxon	materials_examined	Type material. Holotype. [VIETNAM] ♂, Me Linh Biol. Sta. (100 m), 20 – 21 Apr 2015, J. B. Heppner (gen. slide 4650; adult photo 12646) (MGCL).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B272FFB12AE1DA1F0742FAD4.taxon	diagnosis	Diagnosis. This species is distinctive in its maroon-orange and dark brown wing coloring, with irrorated spots and striae of pale orange, with a yellow head (orange and brown in T. carapaceana sp. nov.). The male genitalia have many features of the previous species, but is distinct in the transtilla being less developed compared to the previous species.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B272FFB12AE1DA1F0742FAD4.taxon	description	Description. Wing expanse: 14 mm (n = 1). Head (Fig. 3): yellow overall; antenna light brown with pale yellow venter, with short ventral cilia in male; labial palpus yellow, with white on basal segment and small brown dorsal area on apical segment; head venter light yellow. Thorax (Fig. 3): dull orange, with brown-gray median longitudinal line and brown-gray on dorsum of meso- and metathoraces; tegula brown-gray; venter silver-white; legs silver-white (but tan-white on midleg), with dark gray-brown markings, and hindleg dark gray (silver-white mesally) and yellow end tuft and on tarsi between dark marks brown; spurs dark gray (tan-white distally). Forewing (Fig. 3): light maroon on basal 1 / 3 and apical 1 / 5, with median region dark brown speckled with spots and striae of pale orange; basal maroon area divided by brown-gray horizontal median line and likewise on costal margin and along anal vein; basal vertical stria of pale orange at 2 / 5, straight from dorsal margin but sharply bent basad before coming to costal margin; costa with 7 pale orange (more orange near apex), angled to tornus, each margined by dark brown, with distal three mostly silvery between orange striae each; apex with light orange subapical bar; subtermen with elongated orange area, with three black spots (middle one more a bar); end of cell with pale orange-yellow spot; fringe dark brown (pale to dull white along apical margin); venter dark brown with yellow termen and apex (black dorsal marks repeated), and anal sector white. Hindwing (Fig. 3): dark brown with diffuse pale yellow in cell from midwing to wing base, and slight pale brown along cubital sector; fringe brown; venter dark brown with orange-yellow termen and as thin line at end of cell, and diffuse orange-yellow along cubital vein. Abdomen: dark brown with bronze-golden posterior tergal margins; venter tan; genital tufts brown, with tan laterally and tan-white venter; pregenital segment (Fig. 7 c), with blunt-rounded sternal margin, medially invaginated in U-shape; lateral coremata long, each with deep lateral corematal pouch. Male genitalia (Fig. 7): tegumen an inverted V-shape; uncus (Fig. 7 b), stout, from small base, evenly narrow and tapering distally only at somewhat blunt apex; hami (Fig. 7 b), long (subequal to uncus length) and narrow, strongly sclerotic, bent from stout base, slightly curved ventrally near at middle before acute narrow apex; socii (Fig. 7 b), long (3.4 uncus length) and thick, basally wide and narrowing to apex, densely setose dorsally; gnathos (Fig. 7 b), with flattened narrow arms, medially convergent to acute apex; transtilla (Fig. 7 b), a quadratic plate, with dorsal margin slightly convex and slightly divergent laterally to valval juncture, with caudal end projected and medially divided to bifurcate rounded projections, with margin of fine needle-like spinae; juxta (Fig. 7 b), an ovate plate (slightly concave caudally), with dorsal margin truncated and slightly concave; anellus a membranous ring; aedeagus (Fig. 7 a), tubular of moderate length, with blunt end; phallobase long and decumbent; cornutus undeveloped, with spinose vesica; ductus ejaculatorius long, with elongated hood; valva (Fig. 7), short, oblong with blunt-rounded termen, overall setose and with denser setal area on costa before apex; sacculus slightly rounded near base; vinculum V-shaped, merging to short ventral extension as undeveloped rounded saccus. Female unknown.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B272FFB12AE1DA1F0742FAD4.taxon	etymology	Etymology. The species is named for its type-locality site, the Me Linh Biol. Sta., near Dai Lai, at the base of Tam Dao Mountain, in Vinh Phuc Prov.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B272FFB12AE1DA1F0742FAD4.taxon	biology_ecology	Biology. Unknown. Flight period: April Habitat lowland semi-wet subtropical forest, bordering the Red River Valley of the Hanoi region (Sterling et al. 2006).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B272FFB12AE1DA1F0742FAD4.taxon	distribution	Distribution. Known only from northern Vietnam.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B272FFB12AE1DA1F0742FAD4.taxon	discussion	Discussion. This species is clearly related to the previous species, T. carapaceana, with a less developed transtilla. The maculation of both species is similar, with T. melinhana being lighter brown-orange at wingbase.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B27CFFB52AE1D8D60683FE6C.taxon	description	(Figs. 4, 8)	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B27CFFB52AE1D8D60683FE6C.taxon	materials_examined	Type material. Holotype. [VIETNAM] ♂, Tam Dao [Village] (930 m), 8 Jul 2003, Y. S. Bae et al. (gen. slide JBH- 4545; adult photos 15921) (INUC / NIBR).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B27CFFB52AE1D8D60683FE6C.taxon	diagnosis	Diagnosis. This species is distinctive in the white marks of the forewing apex and the subapical bright orange-yellow bar, as well as for the straight horizontal orange-yellow line from the wing base to 1 / 3. The male genitalia have a very long thin uncus, and the hami are also long and thin, tapering from a bulbous base to a sickle-like acute apex; and the socii are long and thin. Viewed dorsally, this species has the labial palpi approximate to each other (see Fig. 4), while in the other species the palpi are further apart (Figs. 1 – 3).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B27CFFB52AE1D8D60683FE6C.taxon	description	Description. Wing expanse: 12 mm (n = 1). Head (Fig. 4): brown-gray, with yellow laterally, but white on caudal eye margin and neck; antenna yellow and brown-gray, with amber venter, with long black ventral cilia in pairs; labial palpus mostly white, with yellow and brown-gray on basal segment, and apical segment with black laterally near apex; head venter white. Thorax (Fig. 4): brown-gray, with yellow lateral line each side; tegula brown-gray with yellow margin distad; venter white; legs white with dark brown bar marks as in the other species. Forewing (Fig. 4): black-brown, with median striae white and angled from dorsal margin toward apex; a straight light orange line along radius from base to 1 / 3, with a corresponding faint shorter pale white line on cubitus; costa with angled white marks from margin to radius, each bordered by black-brown and with orange between, the distad white mark more straight; wing apex with a large white wedge from costal margin to near termen and then sharply recurved back to near costa, margined by black-brown and bordered by orange; a large orange to orange-yellow curved wedge, widening from near costal margin toward termen; termen with thin orange line, interrupted below apex by a white black-margined fringe indentation; tornal quarter with series of white spots and short striae, all curved toward apex, and one larger yellowing-spot on cubitus below end of cell; subtornal orange area with four black spots, with basal two larger; fringe black-brown, with white along apex and inside termen invagination; venter dark brown, with very dull repetition of the dorsal markings, but bright orange-yellow and white dorsal marks repeated in apex and termen. Hindwing (Fig. 4): dark brown with orange suffused cell area from midwing to near wing base; termen with orange line from apex to tornus; anal sector lightly pale brown; costal sector white; venter dark brown, with orange-yellow on termen and apex, and pale yellow thin line at end of cell. Abdomen: lustrous bronze-brown; venter lustrous white, but sequentially less white on sternites 3 – 6; genital tufts brown with white ventrally and laterally; pregential segment (Fig. 8 c), truncated, with medial U-shaped invagination; coremata long, from deep lateral pouches. Male genitalia (Fig. 8): tegumen globular, rounded; uncus (Fig. 8 b), very long (ca. 2 / 3 valval length), sclerotic and thin, from bulbous base quickly thinned and uniform to acute-round apex, overall decumbent and evenly recurved after basal bend; hami (Fig. 8 b), long and thin sclerotic, from wide base and tapering distally gradually to sickle-like acute point; socii (Fig. 8 b), long and thin, moderately setose; gnathos (Fig. 8 b), with short lateral flattened arms, convergent medially to knobbed ends; transtilla (Fig. 8 b), a thin band, somewhat arched dorsally; juxta (Fig. 8 b), an ovate plate (slightly concave caudally), with dorsal margin truncated and slightly concave, laterally projected narrowly into anellus ring (partially as sclerotized band; aedeagus (Fig. 8 a), short, tubular, with short caudal projection and distally blunt; phallobase obsolete; cornutus a long, thin flechette; vesica spinose; ductus ejaculatorius long, with elongate-globular hood; valva (Fig. 8), elongate, oblong with rounded termen, overall setose; sacculus slightly bulging beyond base; vinculum U-shaped quadratic (Fig. 8 b has the vinculum twisted out of position and dorsad instead of ventrad); saccus obsolete. Female unknown.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B27CFFB52AE1D8D60683FE6C.taxon	etymology	Etymology. The species is named for the hill station, Tam Dao, now included also in Tam Dao N. P., Vinh Phuc Prov.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B27CFFB52AE1D8D60683FE6C.taxon	biology_ecology	Biology. Unknown. Flight period: July. Habitat (Fig. 11) is montane subtropical forest (Sterling et al. 2006).	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B27CFFB52AE1D8D60683FE6C.taxon	distribution	Distribution. Known only from northern Vietnam.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
03F9B178B27CFFB52AE1D8D60683FE6C.taxon	discussion	Discussion. This species is among those species superficially resembling the generic type-species, T. zapyra from New Guinea, but the genitalia are very different. The most similar species by maculation is one from Thailand, Thaumatographa johnbradleyi (Razowski, 2009) (n. comb., originally described in Hilarographa), however, the genitalia are very different. These species form a different species-group from the previous species described herein, but all the Thaumatographa species need further study before species-groups can be assigned.	en	Heppner, Jhon B., Bae, Yang-Seop (2025): Thaumatographa tortricids in Vietnam (Lepidoptera, Tortricidae, Chlidanotinae, Hilarographini). Zootaxa 5609 (3): 375-389, DOI: 10.11646/zootaxa.5609.3.4, URL: https://doi.org/10.11646/zootaxa.5609.3.4
