taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03F987CE8D694132FF52F84031634267.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20645509/files/figure.png	https://doi.org/10.5281/zenodo.20645509	FIGURE 3. Cobitis tosaensis, BSKU 119305, holotype, male, 64.2 mm SL: Okuura River, Japan.	FIGURE 3. Cobitis tosaensis, BSKU 119305, holotype, male, 64.2 mm SL: Okuura River, Japan.	2026-03-11	Takahashi, Hiroaki		Zenodo	biologists	Takahashi, Hiroaki			
03F987CE8D694132FF52F84031634267.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20645517/files/figure.png	https://doi.org/10.5281/zenodo.20645517	FIGURE 5. Cobitis tosaensis, A. BSKU 119303, paratype, male, 82.2 mm SL; B. BSKU 122720, paratype, female, 89.3 mm SL, mouth; C. TKPM-P 27320, paratype (male), 67.2 mm SL; dorsal view of pectoral fin.	FIGURE 5. Cobitis tosaensis, A. BSKU 119303, paratype, male, 82.2 mm SL; B. BSKU 122720, paratype, female, 89.3 mm SL, mouth; C. TKPM-P 27320, paratype (male), 67.2 mm SL; dorsal view of pectoral fin.	2026-03-11	Takahashi, Hiroaki		Zenodo	biologists	Takahashi, Hiroaki			
03F987CE8D694132FF52F84031634267.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20645519/files/figure.png	https://doi.org/10.5281/zenodo.20645519	FIGURE 6. Right lateral ethmoid with suborbital spines, lateral view (upper), and dorsal view (lower) of A. C. tosaensis holotype and B. C. biwae, BSKU 119317.	FIGURE 6. Right lateral ethmoid with suborbital spines, lateral view (upper), and dorsal view (lower) of A. C. tosaensis holotype and B. C. biwae, BSKU 119317.	2026-03-11	Takahashi, Hiroaki		Zenodo	biologists	Takahashi, Hiroaki			
03F987CE8D694132FF52F84031634267.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20645522/files/figure.png	https://doi.org/10.5281/zenodo.20645522	FIGURE 7. Lamina circularis of Cobitis tosaensis, Cobitis sp. BIWAE type A. and Cobitis sp. BIWAE type B. A and E: Cobitis tosaensis, holotype, B and F: Cobitis sp. BIWAE type A, BSKU 119317, C and G: Cobitis sp. BIWAE type B, BSKU 122715, D and H: Cobitis sp. BIWAE type C, BSKU 137908.	FIGURE 7. Lamina circularis of Cobitis tosaensis, Cobitis sp. BIWAE type A. and Cobitis sp. BIWAE type B. A and E: Cobitis tosaensis, holotype, B and F: Cobitis sp. BIWAE type A, BSKU 119317, C and G: Cobitis sp. BIWAE type B, BSKU 122715, D and H: Cobitis sp. BIWAE type C, BSKU 137908.	2026-03-11	Takahashi, Hiroaki		Zenodo	biologists	Takahashi, Hiroaki			
03F987CE8D694132FF52F84031634267.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20645524/files/figure.png	https://doi.org/10.5281/zenodo.20645524	FIGURE 8. Cobitis tosaensis, not preserved, during non-breeding season at the type locality on 16 June 2014. (Upper image is of a male and lower of a female).	FIGURE 8. Cobitis tosaensis, not preserved, during non-breeding season at the type locality on 16 June 2014. (Upper image is of a male and lower of a female).	2026-03-11	Takahashi, Hiroaki		Zenodo	biologists	Takahashi, Hiroaki			
03F987CE8D694132FF52F84031634267.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20645528/files/figure.png	https://doi.org/10.5281/zenodo.20645528	FIGURE 9. Live image of Cobitis tosaensis in nuptial coloration: TKPM-P27322, paratype, male (upper), 77.7 mm SL; BSKU 122873, paratype, female (lower), 76.7 mm SL.	FIGURE 9. Live image of Cobitis tosaensis in nuptial coloration: TKPM-P27322, paratype, male (upper), 77.7 mm SL; BSKU 122873, paratype, female (lower), 76.7 mm SL.	2026-03-11	Takahashi, Hiroaki		Zenodo	biologists	Takahashi, Hiroaki			
03F987CE8D694132FF52F84031634267.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20645515/files/figure.png	https://doi.org/10.5281/zenodo.20645515	FIGURE 4. Coloration around the caudal fin in Cobitis tosaensis, Cobitis sp. BIWAE type A and Cobitis sp. BIWAE type B Sanin population. A–D: Coloration of surface layer; E–H: Coloration of depth layer; A and E: Cobitis tosaensis, BSKU 119303, paratype, Shinjyo River, Susaki, Kochi; B and F: Cobitis sp. BIWAE type A, BSKU 119317, Yoshino River, Mima, Tokushima; C and G: Cobitis sp. BIWAE type A, BSKU 119320, Doki River, Marugame, Kagawa; D and H: Cobitis sp. BIWAE type B Sanin population, BSKU 122706, Shintate River, Matsue, Shimane.	FIGURE 4. Coloration around the caudal fin in Cobitis tosaensis, Cobitis sp. BIWAE type A and Cobitis sp. BIWAE type B Sanin population. A–D: Coloration of surface layer; E–H: Coloration of depth layer; A and E: Cobitis tosaensis, BSKU 119303, paratype, Shinjyo River, Susaki, Kochi; B and F: Cobitis sp. BIWAE type A, BSKU 119317, Yoshino River, Mima, Tokushima; C and G: Cobitis sp. BIWAE type A, BSKU 119320, Doki River, Marugame, Kagawa; D and H: Cobitis sp. BIWAE type B Sanin population, BSKU 122706, Shintate River, Matsue, Shimane.	2026-03-11	Takahashi, Hiroaki		Zenodo	biologists	Takahashi, Hiroaki			
03F987CE8D694132FF52F84031634267.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20645530/files/figure.png	https://doi.org/10.5281/zenodo.20645530	FIGURE 10. Type locality (black triangle) and distribution range (red dotted line) of Cobitis tosaensis.	FIGURE 10. Type locality (black triangle) and distribution range (red dotted line) of Cobitis tosaensis.	2026-03-11	Takahashi, Hiroaki		Zenodo	biologists	Takahashi, Hiroaki			
