identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03FA922DFFC2FF9966B7F9FBB09C5068.text	03FA922DFFC2FF9966B7F9FBB09C5068.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eustigmaeus specialis Ikeda & Imada & Nakano 2026	<div><p>Eustigmaeus specialis Ikeda &amp; Nakano sp. nov.</p><p>urn:lsid:zoobank.org:act: E65890F8-3DCF-4DA6-B76F-F22BF0CE8043</p><p>Figs 1–6; Table 1</p><p>Eustigmaeus sp. 2 – Ikeda et al. 2024b: 734, fig. 4.</p><p>Diagnosis</p><p>The new species shares the following character states with E. kauaiensis, E. parakauaiensis Kapaxidi, Stathakis &amp; Papadoulis, 2013, and E. zhengyii Hu &amp; Zhu, 1994: idiosomal lengths of 300 to 400 μm; an elongate major callosity is situated laterally to the prodorsal shield; dorsal seta sci is minute; seta e1 is longer than f1; and aggenital shield bears two pairs of setae. However, the new species can be distinguished from E. parakauaiensis by the presence of enlarged, irregularly shaped dimples on the anterior part of the prodorsal shield (Figs 1–2). It also differs from E. zhengyii in having a genu II with four setae and tarsus IV with eight setae (Table 1). The new species is most similar to E. kauaiensis in overall dorsal seta lengths. However, the dimples on the hysterosomal shield are medium-sized with sharply defined edges in the new species, whereas they are small with ill-defined edges in E. kauaiensis (Figs 1–2, 7). It also differs from E. kauaiensis in the number of setae on trochanter III (Table 1).</p><p>Etymology</p><p>The specific epithet ‘ specialis ’ (Latin adjective meaning ‘specialized’) refers to the apparent high host specificity of the species, associated with mosses of the genus Philonotis Brid. (Ikeda et al. 2024b). We propose the Japanese name ‘Sawagoke-maruhishidani’, derived from the Japanese name Philonotis .</p><p>Type material</p><p>Holotype JAPAN – Honshu • ♀; Wakayama, Arita, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=135.39355&amp;materialsCitation.latitude=34.032917" title="Search Plazi for locations around (long 135.39355/lat 34.032917)">Aritagawa</a>; 34°01′58.5″ N, 135°23′36.8″ E; 270 m a.s.l.; 10 Oct. 2021; S. Ikeda leg.; from moss on wet concrete wall along the mountain path; locality code 24 in Ikeda et al. (2024b); KUZ Z4626.</p><p>Paratype JAPAN – Honshu • 1 ♀; same data as for holotype; KUZ Z4628 .</p><p>Other material examined</p><p>JAPAN – Honshu • 2 ♀♀; Hiroshima, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.22614&amp;materialsCitation.latitude=34.45175" title="Search Plazi for locations around (long 133.22614/lat 34.45175)">Onomichi</a>; 34°27′06.3″ N, 133°13′34.1″ E; 10 m a.s.l.; 26 Mar. 2021; S. Ikeda leg.; from moss on a wet stone by the river; locality code 14 in Ikeda et al. (2024b); GenBank no. LC789962 (COI); KUZ Z4661, KUZ Z4829 • 1 ♀; Wakayama, Shingu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=135.8292&amp;materialsCitation.latitude=33.76289" title="Search Plazi for locations around (long 135.8292/lat 33.76289)">Kumanogawanishi</a>; 33°45′46.4″ N, 135°49′45.1″ E; 80 m a.s.l.; 11 Oct. 2021; S. Ikeda leg.; from moss on dripping rock face along the pavement road; locality code 23 in Ikeda et al. (2024b); KUZ Z4711 • 1 ♀; same data as for holotype; GenBank no. LC789943 (COI); KUZ Z4802 • 2 ♀♀; Shizuoka, Hamamatsu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=137.81259&amp;materialsCitation.latitude=34.96989" title="Search Plazi for locations around (long 137.81259/lat 34.96989)">Tenryu</a>; 34°58′11.6″ N, 137°48′45.3″ E; 180 m a.s.l.; 22 Apr. 2022; S. Ikeda leg.; from moss on wet concrete wall along the pavement road; locality code 27 in Ikeda et al. (2024b); GenBank no. LC789967 (COI); KUZ Z4703, KUZ Z4811 • 2 ♀♀; Shizuoka, Shizuoka, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.33028&amp;materialsCitation.latitude=35.14253" title="Search Plazi for locations around (long 138.33028/lat 35.14253)">Aoi</a>; 35°08′33.1″ N, 138°19′49.0″ E; 250 m a.s.l.; 21 Apr. 2022; S. Ikeda leg.; from moss on wet rock face along the pavement road; locality code 28 in Ikeda et al. (2024b); GenBank no. LC789963 (COI); KUZ Z4709, KUZ Z4830 . – Shikoku • 1 ♀; Kochi, Takaoka, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.89105&amp;materialsCitation.latitude=33.465836" title="Search Plazi for locations around (long 132.89105/lat 33.465836)">Yusuhara</a>; 33°27′57.0″ N, 132°53′27.8″ E; 860 m a.s.l.; 27 Sep. 2021; S. Ikeda leg.; from moss on dripping concrete wall along the pavement road; locality code 2 in Ikeda et al. (2024b); KUZ Z4719 • 2 ♀♀; Ehime, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.93475&amp;materialsCitation.latitude=33.743332" title="Search Plazi for locations around (long 132.93475/lat 33.743332)">Toon</a>; 33°44′36.0″ N, 132°56′05.1″ E; 630 m a.s.l.; 1 Mar. 2021; S. Ikeda leg.; from moss on dripping concrete wall along the pavement road; locality code 5 in Ikeda et al. (2024b); GenBank no. LC789948 (COI); KUZ Z4527, KUZ Z4804 • 1 ♀; Tokushima, Naka, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.15517&amp;materialsCitation.latitude=33.783028" title="Search Plazi for locations around (long 134.15517/lat 33.783028)">Naka</a>; 33°46′58.9″ N, 134°09′18.6″ E; 470 m a.s.l.; 31 Jul. 2021; S. Ikeda leg.; from moss on dripping rock face along the mountain path; locality code 6 in Ikeda et al. (2024b); GenBank no. LC789965 (COI); KUZ Z4809 • 3 ♀♀; Tokushima, Naka, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.30217&amp;materialsCitation.latitude=33.813442" title="Search Plazi for locations around (long 134.30217/lat 33.813442)">Naka</a>; 33°48′48.4″ N, 134°18′07.8″ E; 270 m a.s.l.; 31 Jul. 2021; S. Ikeda leg.; from moss on wet rock face along the mountain path; locality code 7 in Ikeda et al. (2024b); GenBank no. LC789950 (COI); KUZ Z4704, KUZ Z4807, KUZ Z4844 • 2 ♀♀; Tokushima, Myozai, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.25111&amp;materialsCitation.latitude=33.933002" title="Search Plazi for locations around (long 134.25111/lat 33.933002)">Kamiyama</a>; 33°55′58.8″ N, 134°15′04.0″ E; 600 m a.s.l.; 11 Sep. 2021; S. Ikeda leg.; from moss on wet soil along the pavement road; locality code 10 in Ikeda et al. (2024b); KUZ Z4708, KUZ Z4709 • 1 ♀; Tokushima, Myozai, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.24234&amp;materialsCitation.latitude=33.942944" title="Search Plazi for locations around (long 134.24234/lat 33.942944)">Kamiyama</a>; 33°56′34.6″ N, 134°14′32.4″ E; 420 m a.s.l.; 7 May 2022; R. Miyoshi leg.; from moss on a wet boulder by the stream; locality code 11 in Ikeda et al. (2024b); KUZ Z4697 • 1 ♀; Tokushima, Mima, Anabuki, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.15053&amp;materialsCitation.latitude=34.02353" title="Search Plazi for locations around (long 134.15053/lat 34.02353)">Anabukigawa</a>; 34°01′24.7″ N, 134°09′01.9″ E; 50 m a.s.l.; 11 Sep. 2021; S. Ikeda leg.; from mosses on a wet boulder and dripping rock face by the river; locality code 13 in Ikeda et al. (2024b); KUZ Z4652 .</p><p>Description of females (n = 15)</p><p>MEASUREMENTS. Idiosomal length 320 (293–360), idiosomal width 266 (237–276).</p><p>IDIOSOMAL DORSUM (Figs 1–3). Two large shields almost completely cover idiosomal dorsum. Dorsal shields sclerotized. Dimples oval or nearly round; small or absent on central part of prodorsal shield but enlarged, irregularly shaped on its anterior part (Fig. 3). Distances between dimples greater than their diameters. Shield surface minutely punctate (Fig. 1B). Fossettes as illustrated (Fig. 2A). Ocelli present. Lateral margin of hysterosomal shield weakly sclerotized. One pair of oval and large callosity positioned on pleural membrane close to prodorsal shield, length at longest part 65 (64–94). Suranal shield located ventrally, and sclerotized with punctate surface. Suranal setae pointed with minute barbs. Dorsal setae rod-like with bluntly pointed tips and sparsely barbed, hyaline sheaths evident near tips. Lengths of dorsal setae: vi 43 (39–56), ve 61 (59–78), sci 8 (9–11), sce 53 (51–66), c1 58 (49–70), c2 41 (39–51), d1 73 (63–81), d2 58 (52–69), e1 80 (70–92), e2 68 (61–81), f1 45 (47–59), h1 39 (37–45), h2 29 (29–36). Distances between setae: vi–vi 40 (32–41), ve–ve 91 (84–95), sci–sci 147 (142–161), sce–sce 188 (178–212), vi–ve 53 (48–60), ve–sci 30 (30–37), sci–sce 38 (29–40), c1–c1 89 (78–94), d1–d1 73 (66–90), d2–d2 221 (214–242), e1–e1 110 (97–114), e2–e2 181 (168–190), f1–f1 68 (63–82), h1–h1 29 (22–32), h2–h2 63 (65–77), c1–d1 63 (55–73), c1–d2 81 (78–91), e1–d2 129 (121–146), d1– d2 74 (72–91), d1–e1 100 (88–112), d1–e2 78 (73–84), d2–e2 78 (72–89), e1– e2 52 (49–60), e1–f1 39 (22–47), h1–h2 26 (23–31).</p><p>IDIOSOMAL VENTER (Fig. 2B). Humeral shield punctate with amorphous shallow dimples. Coxisternal plates of legs minutely punctate, posterior part of leg II coxisternal plate faintly reticulated or not. Endopodal plates punctate and reticulated, and separated medially. Aggenital plate punctate and faintly reticulated, with two pairs of aggenital setae. Anal cover with three pairs of simple pseudanal setae. Lengths of setae: ag1 21 (21–27), ag2 23 (20–26), ps1 24 (18–27), ps2 25 (19–26), ps3 22 (21–27).</p><p>GNATHOSOMA (Fig. 4). Dorsal surface of chelicera punctate. Chelicera and its short movable digit 98 (89–101) and 23 (21–28) in length, respectively. Seta l ′ on palpal tibia short, finger-like. The other palpal setae of femur, genu, and tibia sparsely barbed and pointed. For palptarsal setae, acm ζ, (ul ζ), and sul ζ smooth with blunt tips, ω finger-shaped, the other setae smooth and pointed. Counts of palpal setae from trochanter to tarsus: 0, 3, 2, 3, 6 + ω. Supracoxal seta ep short, needle-like, located laterally. Ventral surface of subcapitulum punctate. Rostrum short. Adoral seta or2 smooth, thickened, and curved with a blunt tip. Setae m, n, and or1 pointed, smooth or sparsely barbed. Lengths of setae: m 21 (17–23), n 23 (21–26), or1 5 (7–9), or2 11 (10–14). Distances between subcapitular setae: m–m 26 (25–29), n–n 38 (33–40), m–n 17 (13–19).</p><p>LEGS (Figs 5–6). Leg lengths: leg I 197 (185–218), leg II 163 (150–172), leg III 156 (146–181), leg IV 190 (176–203). Empodium with three pairs of tenant hairs. Counts of setae on leg I–IV: coxae 2 + el, 2, 2, 2; trochanters 1, 1, 2, 1; femora 6, 5, 3, 2; genua 3 + 1 k, 3 + 1 k, 1, 1; tibiae 5 + 1 φ + 1 φp, 5 + 1 φp, 5 + 1 φp, 5 + 1 φp; tarsi 13 + 1 ω, 9 + 1 ω, 7 + 1 ω, 7 + 1 ω. Leg I (Fig. 5A). Supracoxal seta el short, spine-like. Leg setae d of tibia, (ft), (tc), and (p) of tarsus eupathid-like. Seta d of femur, d and (l) of genu sparsely barbed and pointed (or bluntly pointed), with (or without) hyaline sheaths. Setae v ′ of trochanter, bv ″, (v), and (l) of femur, (v) and (l) of tibia, vs and (pl) of tarsus sparsely barbed and pointed. Setae (a) and (u) of tarsus slightly serrated or smooth. Solenidion ω and φ baculiform with round tips, φp attenuate. Seta k needle-shaped. Seta lengths: k 10 (10–14), φp 18 (12–21), φ 11 (9–15), ω 37 (33–40). Leg II (Fig. 5B). Leg setae tc ′ and p ′ of tarsus eupathid-like. Setae d of genu sparsely barbed and pointed (or bluntly pointed), with (or without) hyaline sheaths. Setae v ′ of trochanter, bv ″, v ′, (l), and d of femur, (l) of genu, (v), (l), and d of tibia, vs, pl ′, and tc ″ of tarsus sparsely barbed and pointed. Setae (a) and (u) of tarsus slightly serrated or smooth. Solenidion ω baculiform with round tip, φp attenuate. Seta k needle-shaped. Seta lengths: k 5 (4–9), φp 13 (10–16), ω 23 (21–28). Leg III (Fig. 6A). Setae d of genu and tibia sparsely barbed and pointed (or bluntly pointed), with (or without) hyaline sheaths. Setae v ′ and l ′ of trochanter, ev ″, l ′, and d of femur, (v) and (l) of tibia, vs and (tc) of tarsus sparsely barbed and pointed. Setae (a) and (u) of tarsus slightly serrated or smooth. Solenidion ω baculiform with round tip, φp attenuate. Seta lengths: φp 7 (5–11), ω 7 (5–11). Leg IV (Fig. 6B). Setae d of genu and tibia sparsely barbed and pointed (or bluntly pointed), with (or without) hyaline sheaths. Setae v ′ of trochanter, ev ′ and d of femur, (l) and (v) of tibia, (tc) and vs of tarsus sparsely barbed and pointed. Setae (a) and (u) of tarsus slightly serrated or smooth. Solenidion ω finger-shaped, φp attenuate. Seta lengths: φp 7 (4–8), ω 5 (3–7).</p><p>a) Calculation based on the seta values of the holotype in the reference.</p><p>b) Data derived from the descriptions in the references.</p><p>c) Calculation based on the maximum and minimum values of each seta.</p><p>d) Calculation based on the seta values in Doǧan (2005).</p><p>e) Data based on the illustrations in the reference.</p><p>f) Revised after examination of the holotype.</p><p>g) The original description reported tarsus IV as 7, which probably indicated lack of solenidion ω.</p><p>Host plants</p><p>Four species of Bartramiaceae Schwägr. and one species of Mniaceae Schwägr. have been recorded (Ikeda et al. 2024b): Bartramiaceae, Philonotis falcata (Hook.) Mitt., P. thwaitesii Mitt., P. hastata (Duby) Wijk &amp; Margad., and P. turneriana (Schwägr.) Mitt.; Mniaceae, Pohlia sp.</p><p>Distribution</p><p>Known from Honshu and Shikoku in the Japanese Archipelago.</p><p>Remarks</p><p>The new species belongs to a species complex including three morphologically similar members, E. kauaiensis, E. parakauaiensis, and E. zhengyii (see Diagnosis). Although the dorsal seta length ratio of ve / sci has been used as a key diagnostic character for distinguishing these species (Hu &amp; Zhu 1994; Fan 2002; Table 1), it showed substantial variation within the new species, even between two specimens from the same locality (KUZ Z4704: 5.9; KUZ Z4844: 7.3). Therefore, the utility of this diagnostic character should be reassessed in the light of intraspecific variation. The examined specimens also showed considerable intraspecific variation in prodorsal shield ornamentation: dimples on the prodorsal shield disappear middorsally in KUZ Z4652, Z4704, and Z4711, whereas evident in KUZ Z4661 and Z4708 (Fig. 3). However, pairwise genetic distances based on COI sequences among populations from seven localities were relatively small (&lt;3.7% difference using the K2P model; Ikeda et al. 2024b), indicating that this morphological variation was not correlated with genetic distance.</p></div>	https://treatment.plazi.org/id/03FA922DFFC2FF9966B7F9FBB09C5068	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ikeda, Satsuki;Imada, Yume;Nakano, Takafumi	Ikeda, Satsuki, Imada, Yume, Nakano, Takafumi (2026): Two new species and new country records of Eustigmaeus (Trombidiformes: Prostigmata: Stigmaeidae) in Japan. European Journal of Taxonomy 1082: 1-40, DOI: 10.5852/ejt.2026.1082.3312, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3312/14560
03FA922DFFCBFF9F6697FA38B00257A6.text	03FA922DFFCBFF9F6697FA38B00257A6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eustigmaeus kauaiensis Swift, Gerson & Goff 1985	<div><p>Eustigmaeus kauaiensis Swift, Gerson &amp; Goff, 1985</p><p>Fig. 7</p><p>Eustigmaeus kauaiensis Swift, Gerson &amp; Goff, 1985: 375, figs 1–4.</p><p>Material examined</p><p>Holotype USA – Hawaii • ♀; Kaua’i, Kapaa; 26 Sep. 1980; M.L. Coff leg.; moss on hau tree ( Hibiscus tiliaceus L.); 1 m above ground; BPBM 13,430.</p><p>Supplementary redescription</p><p>IDIOSOMAL DORSUM (Fig. 7). Dorsal dimples oval or nearly round; minute with ill-defined edges in central areas of shields; enlarged and irregularly shaped on anterior part of prodorsum, remaining dimples slightly enlarged toward idiosomal margin. Distances between dimples greater than their diameters.</p><p>LEGS. Counts of setae on tarsi I–IV: 13 + 1 ω, 9 + 1 ω, 7 + 1 ω, 7 + 1 ω.</p><p>Distribution</p><p>Known from Kauai Island, Hawaii, USA, and Guizhou, Jiangxi, Hainan, and Hubei provinces in China (Swift et al. 1985; Fan 2002).</p><p>Remarks</p><p>Although the original description of E. kauaiensis stated that its dorsal dimples were “prominent, oval or nearly round, distances between dimples greater than their diameters, small in central areas of propodosoma and hysterosoma”, the illustration did not clearly depict this character (Swift et al. 1985). Re-examination of the holotype revealed that the dimples on the hysterosomal shield are minute relative to those of E. specialis sp. nov. (Fig. 7). In addition, the number of setae on tarsus IV is eight (including solenidion ω), not seven as originally reported (Swift et al. 1985), and is amended here. The remaining features of the original description are coincident with those of the holotype.</p></div>	https://treatment.plazi.org/id/03FA922DFFCBFF9F6697FA38B00257A6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ikeda, Satsuki;Imada, Yume;Nakano, Takafumi	Ikeda, Satsuki, Imada, Yume, Nakano, Takafumi (2026): Two new species and new country records of Eustigmaeus (Trombidiformes: Prostigmata: Stigmaeidae) in Japan. European Journal of Taxonomy 1082: 1-40, DOI: 10.5852/ejt.2026.1082.3312, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3312/14560
03FA922DFFCDFF816549FD46B6845171.text	03FA922DFFCDFF816549FD46B6845171.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eustigmaeus saxicola Ikeda & Imada & Nakano 2026	<div><p>Eustigmaeus saxicola Ikeda &amp; Nakano sp. nov.</p><p>urn:lsid:zoobank.org:act: 3EC403CA-4EAC-4036-8674-C5AD764CDC68</p><p>Figs 8–12</p><p>Eustigmaeus sp. 3 – Ikeda et al. 2024b: 734, fig. 4.</p><p>Diagnosis</p><p>The new species shares the following characteristics with E. manus Rimando &amp; Corpuz-Raros, 1997, E. brevirostris (Wood, 1971), and E. capellus Chaudhri, 1965: dorsal shields bear dimples each enclosed in a reticulum, dorsal seta sci is minute, callosity is absent, tarsus IV is without solenidion ω, and aggenital shield bears three pairs of setae (Chaudhri 1965; Wood 1971; Rimando &amp; Corpuz-Raros 1997). However, the new species is distinguishable from the three congeners by several characteristics. Dorsal setae are relatively short and the distance between setae d1 is much longer than that of c1, whereas dorsal setae are elongate in E. capellus and E. manus and setae c1, d1, and e1 are arranged in a straight line in E. capellus according to the description and fig. 15 in Chaudhri (1965). Clavate dorsal setae are rounded and smooth in outline, frequently bearing pronounced spines only at the terminal tip, never deeply branched (vs with heavy barbs at the distal part in E. capellus; with deeply trifurcate projections in E. brevirostris and E. manus). Trochanter III has one seta (vs two setae in E. brevirostris).</p><p>Etymology</p><p>The specific name of the new species is a noun in apposition to the generic name, combination of the Latin words ‘ saxum ’ (‘rock face’) and ‘ cola ’ (‘inhabitant’), derived from its ecology. We propose the Japanese name “Kougyoku-maruhishidani” for this species. ‘Kougyoku’ means ‘ruby’ in Japanese and refers to the appearance of the species when alive, which exhibits a shiny red color due to light reflecting from the dimples on the dorsal shields.</p><p>Type material</p><p>Holotype JAPAN – Honshu • ♀; Hyogo, Shiso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.66167&amp;materialsCitation.latitude=35.153835" title="Search Plazi for locations around (long 134.66167/lat 35.153835)">Ichinomiya</a>, 35°09′13.8″ N, 134°39′42.0″ E; 550 m a.s.l.; 29 Mar. 2021; S. Ikeda leg.; from moss on dripping rock face along the pavement road; locality code 17 in Ikeda et al. (2024b); KUZ Z4553.</p><p>Paratypes JAPAN – Honshu • 2 ♀♀; same data as for holotype; KUZ Z4581, KUZ Z4554 .</p><p>Other material examined</p><p>JAPAN – Honshu • 3 ♀♀; Hiroshima, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.20769&amp;materialsCitation.latitude=34.599804" title="Search Plazi for locations around (long 132.20769/lat 34.599804)">Akita</a>; 34°35′59.3″ N, 132°12′27.7″ E; 370 m a.s.l.; 30 Mar. 2021; S. Ikeda leg.; from moss on wet rock face along the mountain path; locality code 16 in Ikeda et al. (2024b); KUZ Z4723, KUZ Z4547, KUZ Z4724 • 1 ♀; same data as for holotype; GenBank no. LC789949 (COI); KUZ Z4805 • 3 ♀♀; Hyogo, Yabu, Oya, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.61401&amp;materialsCitation.latitude=35.33428" title="Search Plazi for locations around (long 134.61401/lat 35.33428)">Tendaki</a>; 35°20′03.4″ N, 134°36′50.4″ E; 430 m a.s.l.; 29 Mar. 2021; S. Ikeda leg.; from moss on wet rock face along the mountain path; locality code 19 in Ikeda et al. (2024b); KUZ Z4548, KUZ Z4549, KUZ Z4552 • 3 ♀♀; Nara, Yoshino, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=136.01411&amp;materialsCitation.latitude=34.41353" title="Search Plazi for locations around (long 136.01411/lat 34.41353)">Higashiyosihno</a>; 34°24′48.7″ N, 136°00′50.8″ E; 350 m a.s.l.; 13 Oct. 2021; S. Ikeda leg.; from moss on wet soil along the pavement road; locality code 26 in Ikeda et al. (2024b); GenBank no. LC789944 (COI); KUZ Z4728, KUZ Z4825, KUZ Z4863 • 2 ♀♀; Shizuoka, Shizuoka, Aoi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.33688&amp;materialsCitation.latitude=35.308693" title="Search Plazi for locations around (long 138.33688/lat 35.308693)">Umegashima</a>; 35°18′31.3″ N, 138°20′12.8″ E; 1000 m a.s.l.; 21 Apr. 2022; S. Ikeda leg.; from moss on a wet rock face along the pavement road; locality code 30 in Ikeda et al. (2024b); KUZ Z4731, KUZ Z4732 • 2 ♀♀; Saitama, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.76334&amp;materialsCitation.latitude=35.994804" title="Search Plazi for locations around (long 138.76334/lat 35.994804)">Chichibu</a>; 35°59′41.3″ N, 138°45′48.0″ E; 900 m a.s.l.; 27 Oct. 2021; S. Ikeda leg.; from moss on wet rock face along the pavement road; locality code 33 in Ikeda et al. (2024b); GenBank no. LC789972 (COI); KUZ Z4702, KUZ Z4803 . – Shikoku • 3 ♀♀; Kochi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.87201&amp;materialsCitation.latitude=33.035557" title="Search Plazi for locations around (long 132.87201/lat 33.035557)">Shimanto</a>; 33°02′08.0″ N, 132°52′19.2″ E; 100 m a.s.l.; 18 Apr. 2021; S. Ikeda leg.; from moss on a wet rock in the stream; locality code 1 in Ikeda et al. (2024b); GenBank no. LC789974 (COI); KUZ Z4560, KUZ Z4725, KUZ Z4812 • 4 ♀♀; Tokushima, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.184&amp;materialsCitation.latitude=33.965084" title="Search Plazi for locations around (long 134.184/lat 33.965084)">Mima</a>; 33°57′54.3″ N, 134°11′02.4″ E; 280 m a.s.l.; 22 May 2021; S. Ikeda leg.; from moss on wet rock face along the pavement road; locality code 12 in Ikeda et al. (2024b); GenBank no. LC789973 (COI); KUZ Z4564, KUZ Z4566, KUZ Z4727, KUZ Z4806 .</p><p>Description of females (n = 19)</p><p>MEASUREMENTS. Idiosomal length 434 (407–457), idiosomal width 338 (305–338).</p><p>IDIOSOMAL DORSUM (Figs 8–9). Two large shields almost completely cover idiosomal dorsum. Dorsal shields sclerotized; circular dimples bear crinkled rims and disappear around seta insertions; thick subcuticular reticulum entirely evident except for less-sclerotized lateral margins. Shield surface minutely punctate (Fig. 8B). Fossettes as illustrated (Fig. 9A). Ocelli present. Lateral margin of prodorsal and hysterosomal shield weakly sclerotized with shallow dimples. Clavate dorsal setae frequently with a few pronounced barbs near tips, with evident hyaline sheaths; seta f1 with minute barbs; seta sci minute. Suranal shield located ventrally, with amorphous dimples enclosed in subcuticular reticulum. Suranal setae bluntly pointed with minute barbs; seta h1 with hyaline sheath near tips; seta h2 with (KUZ Z4581) or without (KUZ Z4553 and Z4554) the sheath. Lengths of dorsal setae: vi 31 (30–43), ve 42 (41–49), sci 5 (5–8), sce 31 (27–40), c1 24 (24–31), c2 23 (23–32), d1 27 (24–33), d2 24 (23–30), e1 27 (27–37), e2 30 (26–33), f1 49 (47–58), h1 47 (37–47), h2 34 (26–35). Distances between setae: vi–vi 41 (30–44), ve–ve 134 (114–138), sci–sci 209 (196–219), sce–sce 251 (246–275), vi–ve 84 (69–86), ve–sci 45 (45– 55), sci–sce 49 (45–66), c1–c1 87 (87–102), d1–d1 172 (146–172), d2–d2 295 (269–306), e1–e1 130 (117–132), e2–e2 224 (205–234), f1–f1 84 (78–94), h1–h1 39 (33–45), h2–h2 84 (77–94), c1–d1 114 (100–125), c1–d2 124 (108–126), e1–d2 164 (158–173), d1–d2 73 (69–86), d1–e1 98 (93–108), d1– e2 75 (62–78), d2–e2 117 (106–117), e1– e2 58 (45–68), e1–f1 47 (40–61), h1–h2 30 (25–36).</p><p>IDIOSOMAL VENTER (Fig. 9B). Callosity absent. Humeral shield with amorphous dimples enclosed in subcuticular reticulum. Coxisternal plates of legs minutely punctate and reticulated. Endopodal plates punctate and reticulated with setae 1a and 3a, separated medially. Aggenital plate punctate and reticulated, with three pairs of simple aggenital setae; seta ag3 longer than the others. Anal cover with three pairs of simple pseudanal setae; seta ps2 shorter than the others. Lengths of setae: ag1 10 (9–12), ag2 10 (10–12), ag3 15 (15–19), ps1 15 (15–20), ps2 5 (5–10), ps3 12 (12–16).</p><p>GNATHOSOMA (Fig. 10). Dorsal surface of chelicera punctate and reticulated. Chelicera and its short movable digit 113 (102–119) and 32 (29–34) in length, respectively. Palpal setae of d and l ′ on femur barbed and bluntly pointed. Seta l ′ on tibia short, Y-shaped. The other palpal setae of femur, genu, and tibia barbed and pointed. For palptarsal setae, acm ζ, (ul ζ), and sul ζ smooth with blunt tips, ω finger-shaped, va pointed with sparse barbs, other setae smooth and pointed. Counts of palpal setae from trochanter to tarsus: 0, 3, 2, 3, 6 + ω. Supracoxal seta ep short, corn-like, located dorsolaterally. Ventral surface of subcapitulum punctate with thick reticulum. Rostrum short. Adoral seta or2 smooth, thickened, and curved with a blunt tip. Setae m, n, and or1 smooth and pointed. Lengths of setae: m 20 (17–26), n 18 (16–21) or1 13 (13–16), or2 12 (12–14). Distances between subcapitular setae: m–m 28 (26–28), n–n 33 (28–35), m–n 16 (16–21).</p><p>LEGS (Figs 11–12). Leg lengths: leg I 218 (199–224), leg II 193 (184–197), leg III 199 (186–204), leg IV 227 (209–227). Empodium with three pairs of tenant hairs. Counts of setae on leg I–IV: coxae 2 + el, 1 (2 for right coxa in KUZ Z4553 and KUZ Z4702), 2, 2; trochanters 1, 1, 1, 1; femora 6, 5, 3, 2; genua 3 + 1 k, 3 + 1 k, 1, 1; tibiae 5 + 1 φ + 1 φp, 5 + 1 φp, 5 + 1 φp, 5 + 1 φp; tarsi 13 + 1 ω, 9 + 1 ω, 7 + 1 ω, 7. Leg I (Fig. 11A). Supracoxal seta el short, corn-like. Leg setae d of tibia, (ft), (tc), and (p) of tarsus eupathid-like. Setae d and l ″ of femur, d and (l) of genu, and (l) of tibia sparsely barbed and bluntly pointed, hyaline sheaths evident near tips; setae bv ″ and v ″ of femur with or without hyaline sheath. Setae v ′ of trochanter, v ′ and l ′ of femur, (v) of tibia, and (pl) and vs of tarsus sparsely barbed and pointed. Setae (a) and (u) of tarsus smooth or slightly serrated. Solenidia ω and φ baculiform with rounded tips, φp attenuate. Seta k long needle-shaped. Seta lengths: k 6 (6–8), φp 12 (12–14), φ 8 (7–8), ω 21 (18–23). Leg II (Fig. 11B). Leg setae tc ′ and p ′ of tarsus and d of tibia eupathid-like. Setae bv ″, d, and l ″ of femur, d and (l) of genu, (l) of tibia sparsely barbed and bluntly pointed, with or without hyaline sheath. Setae v ′ of trochanter, v ′ and l ′ of femur, (v) of tibia, tc ″, pl ′, and vs of tarsus sparsely barbed and pointed. Setae (a) and (u) of tarsus smooth or slightly serrated. Solenidion ω baculiform with round tip, φp attenuate. Seta k needle-shaped. Seta lengths: k 4 (4–6), φp 8 (6–11), ω 14 (14–17). Leg III (Fig. 12A). Setae d and l ′ of femur, d of genu, d and (l) of tibia sparsely barbed and bluntly pointed, with hyaline sheath. Setae v ′ of trochanter, ev ″ of femur, (v) of tibia, (tc) and vs of tarsus sparsely barbed and pointed. Setae (a) and (u) of tarsus smooth or slightly serrated. Solenidion ω finger-shaped, φp attenuate. Seta lengths: φp 7 (5–8), ω 6 (5–7). Leg IV (Fig. 12B). Setae d of femur, d of genu, d and (l) of tibia sparsely barbed and bluntly pointed, with hyaline sheath. Setae v ′ of trochanter, ev ′ of femur, (v) of tibia, (tc) and vs of tarsus sparsely barbed and pointed. Setae (a) and (u) of tarsus smooth or slightly serrated. Solenidion ω absent. Seta lengths: φp 5 (5–7).</p><p>Host plants</p><p>Five species of Pottiaceae Hampe, one species of Amphidiaceae M.Stech, one species of Ditrichaceae Limpr., one species of Brachytheciaceae Schimp., and one species of Bartramiaceae ? have been recorded (Ikeda et al. 2024b) as host plants. Pottiaceae, Chionoloma subduriusculum (Müll. Hal.) M.Menzel., C. orthodontum (Müll.Hal.) M.Alonso, M.J.Cano &amp; J.A.Jiménez, C. tenuirostre (Hook. &amp; Taylor) M.Alonso, M.J.Cano &amp; J.A.Jiménez, Trichostomum brachydontium Bruch, Anoectangium thomsonii Mitt.; Amphidiaceae, Amphidium mougeotii (Bruch &amp; Schimp.) Schimp.; Ditrichaceae, Ceratodon purpureus (Hedw.) Brid.; Brachytheciaceae, Brachythecium helminthocladum Broth. &amp; Paris; Bartramiaceae ? Philonotis ? sp.</p><p>Distribution</p><p>Known from Honshu and Shikoku in the Japanese Archipelago.</p><p>Remarks</p><p>This species was collected from Honshu and Shikoku Islands, Japan. Despite the geographical distances, little morphological variation and low COI genetic divergence (&lt;2.4% difference using the K2P model) were observed among specimens from eight localities (Ikeda et al. 2024b). A phylogenetic analysis in Ikeda et al. (2024b) showed that the five sequences of this species represent monophyly. This species tends to be collected from rock faces along paved roads or mountain paths and uses dicranid moss lineages as host plants (Ikeda et al. 2024b).</p></div>	https://treatment.plazi.org/id/03FA922DFFCDFF816549FD46B6845171	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ikeda, Satsuki;Imada, Yume;Nakano, Takafumi	Ikeda, Satsuki, Imada, Yume, Nakano, Takafumi (2026): Two new species and new country records of Eustigmaeus (Trombidiformes: Prostigmata: Stigmaeidae) in Japan. European Journal of Taxonomy 1082: 1-40, DOI: 10.5852/ejt.2026.1082.3312, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3312/14560
03FA922DFFD3FF84656CFB1EB171567D.text	03FA922DFFD3FF84656CFB1EB171567D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eustigmaeus etruscus (Berlese 1910)	<div><p>Eustigmaeus cf. etruscus (Berlese, 1910)</p><p>Figs 13–17</p><p>Eustigmaeus sp. 5 – Ikeda et al. 2024b: 734, fig. 4.</p><p>Material examined</p><p>JAPAN – Shikoku • 5 ♀♀; Ehime, Kamiukena, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.938&amp;materialsCitation.latitude=33.731304" title="Search Plazi for locations around (long 132.938/lat 33.731304)">Kumakogen</a>; 33°43′52.7″ N, 132°56′16.8″ E; 1020 m a.s.l.; 16 Mar. 2021; S. Ikeda leg.; from moss colony on wet concrete wall along the paved road; locality code 4 in Ikeda et al. (2024b); KUZ Z4591, KUZ Z4739, KUZ Z4743, KUZ Z4779, KUZ Z4795 .</p><p>Description of females (n = 5)</p><p>MEASUREMENTS. Idiosomal length 473–526, idiosomal width 366–413.</p><p>IDIOSOMAL DORSUM (Figs 13–14). Two large shields almost completely cover idiosomal dorsum. Dorsal shields heavily sclerotized; prominent circular dimples enclosed in subcuticular reticulum. Shield surface minutely punctate (Fig. 13B). Fossettes as illustrated (Fig. 14A). Ocelli present. Lateral margin of prodorsal and hysterosomal shield weakly sclerotized, bumpy. Suranal shield located ventrally, with amorphous dimples and thick a reticulum. Suranal seta h2 pointed with minute barbs. Dorsal setae rod-like with bluntly pointed tips, sparsely barbed, hyaline sheaths evident near tips. Lengths of dorsal setae: vi 50–61, ve 76–87, sci 17–30, sce 45–49, c1 52–57, c2 30–35, d1 58–70, d2 44–48, e1 70–84, e2 51 –59, f1 69–77, h1 45–54, h2 36–41. Distances between setae: vi–vi 27–37, ve–ve 132–149, sci–sci 250–283, sce–sce 309–339, vi–ve 83–89, ve–sci 65–81, sci–sce 49–58, c1–c1 105–124, d1–d1 73–95, d2–d2 327– 355, e1–e1 139–147, e2–e2 255–279, f1–f1 107–126, h1–h1 45–52, h2–h2 104–110, c1–d1 89–95, c1–d2 130–147, e1–d2 171–180, d1–d2 127–132, d1–e1 129–140, d1–e2 122–132, d2–e2 109–114, e1–e2 68–78, e1–f1 56–69, h1–h2 38–41.</p><p>IDIOSOMAL VENTER (Fig. 14). One pair of oval and large callosity positioned on pleural membrane near prodorsal shield, length at longest part 70–86. Humeral shield with amorphous dimples and a thick reticulum. Coxisternal plates of legs minutely punctate and reticulated. Endopodal plates reticulated and separated medially. Aggenital plate punctate and reticulated, with two pairs of simple aggenital setae. Anal cover with three pairs of simple pseudanal setae. Lengths of setae: ag1 20–26, ag2 19–24, ps1 17–21, ps2 18–22, ps3 23–24.</p><p>GNATHOSOMA (Fig. 15). Dorsal surface of chelicera punctate. Chelicera and its short movable digit 91–106 and 29–33 in length, respectively. Seta l ′ on palpal tibia short, cone-like. The other palpal setae of femur, genu, and tibia sparsely barbed and pointed. For palptarsal setae, acm ζ, (ul ζ), and sul ζ smooth with blunt tips, ω finger-shaped, va sparsely barbed, other setae smooth and pointed. Counts of palpal setae from trochanter to tarsus: 0, 3, 2, 3, 6 + ω. Supracoxal seta ep short, needle-like, located dorsolaterally. Ventral surface of subcapitulum punctate. Rostrum short. Adoral seta or2 smooth, thickened, and curved with a blunt tip. Setae m, n, and or1 smooth and pointed. Lengths of setae: m 18–24, n 19–25, or1 11–14, or2 12–13. Distances between subcapitular setae: m–m 31–34, n–n 36–44, m–n 17–20.</p><p>LEGS (Figs 16–17). Leg lengths: leg I 236–257, leg II 193–214, leg III 213–236, leg IV 246–273. Empodium with three pairs of tenant hairs. Counts of setae on leg I–IV: coxae 2 + el, 2, 2, 2; trochanters 1, 1, 1, 1; femora 6, 5, 3, 2; genua 3 + 1 k, 3 + 1 k, 1, 1; tibiae 5 + 1 φ + 1 φp, 5 + 1 φp, 5 + 1 φp, 5 + 1 φp; tarsi 13 + 1 ω, 9 + 1 ω, 7 + 1 ω, 7 + 1 ω. Leg I (Fig. 16A). Supracoxal seta el short, needle-like. Leg setae d of tibia, (ft), (tc), and (p) of tarsus eupathid-like. Setae d of femur and genu sparsely barbed and bluntly pointed, hyaline sheaths evident near tips. Setae l ″ of femur, (l) of genu, l ′ of tibia sparsely barbed and bluntly pointed (setae l ″ on genu and l ′ on tibia also with hyaline sheath in KUZ Z4795). Setae v ′ of trochanter, bv ″, (v), and l ′ of femur, l ″ and (v) of tibia, and (pl) and vs of tarsus barbed and pointed. Setae (a) and (u) of tarsus slightly serrated. Solenidia ω and φ baculiform with rounded tips, φp attenuate. Seta k long needle-shaped. Seta lengths: k 15–17, φp 17–19, φ 10–11, ω 25–30. Leg II (Fig. 16B). Leg setae tc ′ and p ′ of tarsus and d of tibia eupathid-like. Setae d of genu sparsely barbed and bluntly pointed, with or without hyaline sheath. Setae d and l ″ of femur, (l) of genu sparsely barbed and bluntly pointed (setae l ″ on genu also with hyaline sheath in KUZ Z4795). Setae v ′ of trochanter, v ′, bv ″, and l ′ of femur, (l) and (v) of tibia, tc ″, pl ′, and vs of tarsus sparsely barbed and pointed. Setae (a) and (u) of tarsus slightly serrated. Solenidion ω baculiform with round tip, φp attenuate. Seta k needle-shaped. Seta lengths: k 7–8, φp 17, ω 15–22. Leg III (Fig. 17A). Setae d of tibia sparsely barbed and bluntly pointed, with or without hyaline sheath. Setae d of femur and genu sparsely barbed and bluntly pointed. Setae v ′ of trochanter, ev ″ and l ′ of femur, (l) and (v) of tibia, and (tc) and vs of tarsus sparsely barbed and pointed. Setae (a) and (u) of tarsus slightly serrated. Solenidion ω finger-shaped, φp attenuate. Seta lengths: φp 10–13, ω 6–8. Leg IV (Fig. 17B). Seta d of tibia sparsely barbed and pointed, with hyaline sheath. Seta d of genu sparsely barbed and bluntly pointed, with hyaline sheath. Seta d of femur sparsely barbed and bluntly pointed, with or without hyaline sheath. Setae v ′ of trochanter, ev ′ of femur, (l) and (v) of tibia, (tc) and vs of tarsus sparsely barbed and pointed. Setae (a) and (u) of tarsus slightly serrated. Solenidion ω spoonshaped, φp attenuate. Seta lengths: φp 5–6, ω 4–5.</p><p>Host plants</p><p>Two species of Bryaceae Rchb. and one species of Hypnales W.R.Buck &amp; Vitt have been recorded (Ikeda et al. 2024b): Bryaceae, Rosulabryum capillare (Hedw.) J.R.Spence and Imbribryum paradoxum (Schwägr.) Ochyra &amp; Bedn.-Ochyra; Hypnales fam. et gen. unespecified.</p><p>Distribution</p><p>Known from Shikoku in the Japanese Archipelago.</p><p>Remarks</p><p>The examined specimens resemble E. granulosus (Wood, 1966), E. erzincanensis Doğan, 2005, E. baguioensis (Rimando &amp; Corpuz-Raros, 1997), and E. etruscus (Berlese, 1910) in having a large body size, rod-like dorsal setae, dorsal shields with circular, prominent dimples, a single pair of callosities, presence of seta 4a, and two pairs of aggenital setae (Berlese 1910; Wood 1966, 1973; Rimando &amp; Corpuz-Raros 1997; Doğan 2005). The examined specimens are most similar to E. etruscus based on the dorsal ornamentation, dorsal seta lengths, and shape and position of callosities. However, the examined species and E. etruscus differ in certain morphological characters. The ratio of the length of dorsal setae to body dimensions of E. etruscus is slightly greater than that in the examined specimens (according to Wood 1973: fig. 9). In addition, E. etruscus has thick-walled cells of distinct reticulum on the dorsal shields, while the reticulum appears faint in the present species (Wood 1973; Fig. 13). Lastly, the trochanter of leg III bears two setae in E. etruscus but bears only one in the present species (Wood 1973; Fig. 17). However, the redescription lacks detailed measurements and photographs (Wood 1973); therefore, revisiting its type specimens is essential to resolve the taxonomic issue.</p><p>One or several vacuoles were observed within individual dimples on the dorsal shields of the examined specimens (Fig. 13). This characteristic has been emphasized in the descriptions of E. erzincanensis, E. erciyesiensis Doğan, Ayyildiz &amp; Fan, 2003, E. baguioensis, and E. parvisetus (Chaudhri, 1965) (Chaudhri 1965; Rimando &amp; Corpuz-Raros 1997; Doğan et al. 2003; Doğan 2005). The examined species may share the same character state with these species. However, droplets with a similar appearance to vacuoles were also observed around the specimens, raising the possibility that those vacuoles are artifacts (Fig. 13A). Although the identity and taxonomic significance of those vacuoles and droplets remain uncertain, those features do not affect the taxonomic conclusion of the examined species at present.</p></div>	https://treatment.plazi.org/id/03FA922DFFD3FF84656CFB1EB171567D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ikeda, Satsuki;Imada, Yume;Nakano, Takafumi	Ikeda, Satsuki, Imada, Yume, Nakano, Takafumi (2026): Two new species and new country records of Eustigmaeus (Trombidiformes: Prostigmata: Stigmaeidae) in Japan. European Journal of Taxonomy 1082: 1-40, DOI: 10.5852/ejt.2026.1082.3312, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3312/14560
03FA922DFFD9FF8C6573FE90B7425729.text	03FA922DFFD9FF8C6573FE90B7425729.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eustigmaeus grandis Khaustov 2019	<div><p>Eustigmaeus grandis Khaustov, 2019</p><p>Figs 18–19; Table 2</p><p>Eustigmaeus grandis Khaustov, 2019: 164, figs 14–17.</p><p>Eustigmaeus sp. 6 – Ikeda et al. 2024b: 734, fig. 4.</p><p>Material examined</p><p>JAPAN – Honshu • 1 ♀; Nagano, Nagano, Mt Togakushi; 21 May 1999; M. Shiba leg.; from moss on bark of Aesculus turbinata Blume; KUZ Z5166 . – Shikoku • 3 ♀♀; Tokushima, Mima, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.09619&amp;materialsCitation.latitude=33.875416" title="Search Plazi for locations around (long 134.09619/lat 33.875416)">Koyadaira</a>; 33°52′31.5″ N, 134°05′46.3″ E; 1360 m a.s.l.; 22 May 2021; S. Ikeda leg.; from mosses on the wet concrete wall and nearby along a pavement road; locality code 8 in Ikeda et al. (2024b); GenBank no. LC789954 (COI); KUZ Z4729, KUZ Z4561, KUZ Z4817 • 1 ♀; Ehime, Mt Higashi-akaishi; 5 Oct. 2002; M. Shiba leg.; from moss on rock by the stream; KUZ Z5167 • 1 ♀; Ehime, Mt Ishizuchi; 14 Nov. 2011; M. Shiba leg.; from moss on rock; KUZ Z5168 .</p><p>Description of females (n = 5)</p><p>MEASUREMENTS. Idiosomal length 428–514, idiosomal width 327–418.</p><p>IDIOSOMAL DORSUM (Figs 18–19). Two large shields almost completely cover idiosomal dorsum. Dorsal shields heavily sclerotized; circular dimples bear crinkled rims and disappear around seta insertions; thick subcuticular reticulum entirely evident. Shield surface minutely punctate. Fossettes as illustrated (Fig. 18A). Ocelli present. Lateral margin of prodorsal and hysterosomal shield weakly sclerotized, bumpy. Suranal shield located ventrally, with amorphous dimples enclosed in subcuticular reticulum. Suranal setae bluntly pointed with minute barbs. Dorsal setae brush-like distally. Lengths of dorsal setae: vi 34–62, ve 46–78, sci 37–55, sce 34–48, c1 39–62, c2 29–39, d1 40–63, d2 36–51, e1 48 –71, e2 42 –59, f1 54–75, h1 42–53, h2 33–36. Distance between setae: vi–vi 24–39, ve–ve 124–140, sci–sci 215–260, sce–sce 304–359, vi–ve 75–90, ve–sci 59–75, sci–sce 56–69, c1–c1 98–123, d1–d1 174–189, d2–d2 325– 378, e1–e1 131–159, e2–e2 236–285, f1–f1 103–128, h1–h1 37–50, h2–h2 86–117, c1–d1 102–128, c1–d2 119–147, e1–d2 180–193, d1–d2 81–103, d1–e1 110–116, d1– e2 58–68, d2–e2 103–111, e1– e2 71–87, e1–f1 29–43, h1–h2 30–47.</p><p>IDIOSOMAL VENTER (Fig. 18). Two pairs of callosities positioned on lateral margins of idiosomal shields, major callosity bigger than minor one, longest part 30–34 and 21–26 in length, respectively. Humeral shield with amorphous dimples enclosed in subcuticular reticulum. Humeral setae distally brush-like (KUZ Z5167) or barbed and bluntly pointed. Coxisternal plates of legs minutely punctate and faintly reticulated. Endopodal plates reticulated and separated medially. Aggenital plate punctate and reticulated, with two pairs of aggenital setae. Anal cover with three pairs of pseudanal setae. Lengths of setae: ag1 16–20, ag2 14–20, ps1 11–17, ps2 9–14, ps3 13–17.</p><p>GNATHOSOMA. Dorsal surface of chelicera punctate. Chelicera and its short movable digit 103–121 and 23–27 in length, respectively. Seta l ′ on palpal tibia short with widened end. Palpfemoral seta d deeply barbed and bluntly pointed. The other palpal setae of femur, genu, and tibia barbed and pointed. For palptarsal setae, acm ζ, (ul ζ), and sul ζ smooth with blunt tips, ω finger-shaped, va pointed with sparse barbs, other setae smooth and pointed. Counts of palpal setae from trochanter to tarsus: 0, 3, 2, 3, 6 + ω. Supracoxal seta ep short, cone-like, located dorsolaterally. Ventral surface of subcapitulum punctate and reticulum present in KUZ Z4729 and Z4561. Rostrum short. Adoral seta or2 smooth, thickened, and curved with a blunt tip. Setae m, n, and or1 smooth and pointed. Lengths of setae: m 16–22, n 15–23, or1 10–13, or2 15–18. Distances between subcapitular setae: m–m 27–33, n–n 33–41, m–n 13–19.</p><p>LEGS. Leg lengths: leg I 191–238, leg II 168–207, leg III 165–211, leg IV 202–243. Empodium with three pairs of tenant hairs. Counts of setae on leg I–IV: coxae 2 + el, 2, 2, 2; trochanters 1, 1, 2, 1 or 1, 1, 1, 1; femora 6, 5, 3, 2; genua 3 + 1 k, 3 + 1 k, 1, 1; tibiae 5 + 1 φ + 1 φp, 5 + 1 φp, 5 + 1 φp, 5 + 1 φp; tarsi 13 + 1 ω, 9 + 1 ω, 7 + 1 ω, 7 + 1 ω. Leg I. Supracoxal seta el short, spine-like. Leg setae (ft), (tc), and (p) of tarsus eupathid-like. Seta lengths: k 7–10, φp 16–21, φ 7–10, ω 24–30. Leg II. Leg setae tc ′ and p ′ of tarsus eupathid-like. Seta lengths: k 5–6, φp 13–17, ω 15–19. Leg III. Seta lengths: φp 8–10, ω 3–6. Leg IV. Seta lengths: φp 6–9, ω 3–5.</p><p>Host plants</p><p>One species of Brachytheciaceae, Brachythecium rivulare Schimp., has been recorded (Ikeda et al. 2024b).</p><p>Distribution</p><p>Known from the Russian Far East (Khaustov 2019) as well as Honshu and Shikoku in the Japanese Archipelago.</p><p>Remarks</p><p>The examined specimens share the following combination of characteristics with E. grandis: dorsal shields with dimples, each enclosed by a reticulum, brush-like dorsal setae, the presence of seta e2, two pairs of callosities, and two pairs of aggenital setae. Although this combination of character states restricts the examined specimens to E. grandis in the genus, considerable morphological variation was observed among populations (Table 2). The variation, including variation in idiosomal dimensions, dorsal seta lengths, and the number of setae on trochanter III (Table 2; Fig. 19), was continuous, making it difficult to delineate the species boundary. The genetic distance based on COI sequences between specimens from the type locality and the Tokushima population was 6.1% (Ikeda et al. 2024b); however, comparable data for specimens from other localities are lacking. Obtaining molecular data from additional localities may help to delineate species boundaries. Here, Eustigmaeus sp. 6 is identified as E. grandis based on the shared major morphological characteristics.</p><p>We propose the Japanese name ‘Oo-maruhishidani’ for this species. ‘Oo’ means ‘large’ in Japanese and corresponds to the meaning of the specific epithet ( grandis).</p></div>	https://treatment.plazi.org/id/03FA922DFFD9FF8C6573FE90B7425729	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ikeda, Satsuki;Imada, Yume;Nakano, Takafumi	Ikeda, Satsuki, Imada, Yume, Nakano, Takafumi (2026): Two new species and new country records of Eustigmaeus (Trombidiformes: Prostigmata: Stigmaeidae) in Japan. European Journal of Taxonomy 1082: 1-40, DOI: 10.5852/ejt.2026.1082.3312, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3312/14560
03FA922DFFDEFFB16699FDC6B5855353.text	03FA922DFFDEFFB16699FDC6B5855353.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eustigmaeus clavatus (Canestrini & Fanzago 1876)	<div><p>Eustigmaeus cf. clavatus (Canestrini &amp; Fanzago, 1876)</p><p>Figs 20–24</p><p>Eustigmaeus sp. 10 – Ikeda et al. 2024b: 734, fig. 4.</p><p>Material examined</p><p>JAPAN – Honshu • 5 ♀♀; Mie, Watarai, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=136.69186&amp;materialsCitation.latitude=34.36525" title="Search Plazi for locations around (long 136.69186/lat 34.36525)">Minamiise</a>; 34°21′54.9″ N, 136°41′30.7″ E; 100 m a.s.l.; 14 Oct. 2021; S. Ikeda leg.; from moss on a wet soil along the mountain path; locality code 25 in Ikeda et al. (2024b); GenBank no. LC789970 (COI); KUZ Z4721, KUZ Z4722, KUZ Z5711, KUZ Z5712, KUZ Z4824 .</p><p>Description of females (n = 4)</p><p>MEASUREMENTS. Idiosomal length 364–411, idiosomal width 292–327.</p><p>IDIOSOMAL DORSUM (Figs 20–21). Two large shields almost completely cover idiosomal dorsum. Dorsal shields sclerotized; dimples varied in size and disappear around seta insertions; thick subcuticular reticulum entirely evident except for less-sclerotized lateral margins. Shield surface minutely punctate (Fig. 20B). Fossettes as illustrated (Fig. 21A). Ocelli present. Dorsal setae slightly clavate with bluntly pointed tips, sparsely barbed, hyaline sheath evident near tips. Suranal shield located ventrally, with shallow dimples enclosed in subcuticular reticulum. Suranal setae rod-like with bluntly pointed tips, minutely barbed, with hyaline sheath. Lengths of dorsal setae: vi 48–58, ve 61–73, sci 37–49, sce 53–58, c1 45–57, c2 39–42, d1 55–60, d2 48–54, e1 62–72, e2 52 –60, f1 63–72, h1 48–55, h2 35–40. Distances between setae: vi–vi 30–36, ve–ve 105–109, sci–sci 185–190, sce–sce 231–239, vi–ve 61–72, ve–sci 44– 50, sci–sce 44–51, c1–c1 81–95, d1–d1 123–133, d2–d2 245–260, e1–e1 127–133, e2–e2 199–206, f1–f1 89–94, h1–h1 37–39, h2–h2 84–91, c1–d1 96–105, c1–d2 99–114, e1–d2 131–143, d1–d2 61–67, d1–e1 95–98, d1–e2 65–72, d2–e2 80–93, e1– e2 48–53, e1–f1 36–43, h1–h2 25–31.</p><p>IDIOSOMAL VENTER (Fig. 21). Callosity absent. Humeral shield sub-triangular, punctate, shallow dimples enclosed in subcuticular reticulum. Coxisternal plates of legs minutely punctate and faintly reticulated. Endopodal plates reticulated and separated medially. Aggenital plate punctate and reticulated, with three pairs of sparsely barbed setae, ag3 slightly longer than others. Anal cover sclerotized and reticulated. Three pairs of pseudanal setae sparsely barbed and pointed, ps1 also thickened. Lengths of setae: ag1 13– 14, ag2 14–17, ag3 23–25, ps1 21–28, ps2 18–23, ps3 23–27.</p><p>GNATHOSOMA (Fig. 22). Dorsal surface of chelicera punctate. Chelicera and its short movable digit 89–98 and 28–33 in length, respectively. Seta l ′ on palpal tibia short, heart-shaped. Palpfemoral seta d sparsely barbed and bluntly pointed. The other palpal setae of femur, genu, and tibia barbed and pointed. For palptarsal setae, acm ζ, (ul ζ), and sul ζ smooth with blunt tips, ω finger-shaped, va sparsely barbed, other setae smooth and pointed. Counts of palpal setae from trochanter to tarsus: 0, 3, 2, 3, 6 + ω. Supracoxal seta ep short, cone-shaped, located dorsolaterally. Ventral surface of subcapitulum punctate with or without subcuticular reticulum. Rostrum short. Adoral seta or2 smooth, thickened and curved with blunt tip. Setae m and or1 smooth and pointed. Seta n pointed with sparse barbs. Lengths of setae: m 20–22, n 22–24, or1 12–14, or2 11–13. Distances between subcapitular setae: m–m 27–29, n–n 31–36, m–n 15–17.</p><p>LEGS (Figs 23–24). Leg lengths: leg I 200–210, leg II 172–179, leg III 174–179, leg IV 204–213. Empodium with three pairs of tenant hairs. Counts of setae on leg I–IV: coxae 2 + el, 2, 2, 2; trochanters 1, 1, 2, 1; femora 6, 5, 3, 2; genua 3 + 1 k, 3 + 1 k, 1, 1; tibiae 5 + 1 φ + 1 φp, 5 + 1 φp, 5 + 1 φp, 5 + 1 φp; tarsi 13 + 1 ω, 9 + 1 ω, 7 + 1 ω, 7. Leg I (Fig. 23A). Supracoxal seta el short, needle-like. Leg setae d of tibia, (ft), (tc), and (p) of tarsus eupathid-like. Setae d of femur, d and (l) of genu, sparsely barbed and bluntly pointed, with hyaline sheath. Setae (l) of tibia sparsely barbed and pointed, with hyaline sheath. Setae l ″ of femur sparsely barbed and bluntly pointed (also with hyaline sheath in KUZ Z5711 and Z5712). Setae v ′ of trochanter, bv ″, (v), and l ′ of femur, (v) of tibia, and (pl) and vs of tarsus barbed and pointed. Setae (a) and (u) of tarsus slightly serrated. Solenidia ω and φ baculiform with rounded tips, φp attenuate. Seta k needle-shaped. Seta lengths: k 8, φp 14–16, φ 8–10, ω 18–20. Leg II (Fig. 23B). Leg setae tc ′ and p ′ of tarsus and d of tibia eupathid-like. Setae bv ″, d, and l ″ of femur, d and (l) of genu sparsely barbed and bluntly pointed, with hyaline sheath. Setae (l) of tibia sparsely barbed and pointed, with hyaline sheath. Setae v ′ of trochanter, v ′ and l ′ of femur, (v) of tibia, tc ″, pl ′, and vs of tarsus sparsely barbed and pointed. Setae (a) and (u) of tarsus slightly serrated. Solenidion ω baculiform with round tip, φp attenuate. Seta k needle-shaped. Seta lengths: k 5–8, φp 10–13, ω 12–13. Leg III (Fig. 24A). Setae l ′ of trochanter, d and l ′ of femur, d of genu, d and (l) of tibia sparsely barbed and bluntly pointed, with hyaline sheath. Setae v ′ of trochanter, ev ″ of femur, (v) of tibia, (tc) and vs of tarsus sparsely barbed and pointed. Setae (a) and (u) of tarsus slightly serrated. Solenidion ω finger-shaped, φp attenuate. Seta lengths: φp 11–12, ω 5–7. Leg IV (Fig. 24B). Seta d of femur, d of genu, d and (l) of tibia sparsely barbed and bluntly pointed, with hyaline sheath. Setae v ′ of trochanter, ev ′ of femur, (v) of tibia, (tc) and vs of tarsus sparsely barbed and pointed. Setae (a) and (u) of tarsus slightly serrated. Solenidion φp attenuate. Seta lengths: φp 12.</p><p>Host plants</p><p>One species of Pottiaceae and one species of Leskeaceae Schimp. have been recorded (Ikeda et al. 2024b): Pottiaceae, Weissia sp.; Leskeaceae, Haplocladium microphyllum (Sw. ex Hedw.) Broth.</p><p>Distribution</p><p>Known from Honshu in the Japanese Archipelago.</p><p>Remarks</p><p>Eustigmaeus clavatus was originally described as Caligonus clavatus from Maser and Padova, Italy (Canestrini &amp; Fanzago 1876). Subsequently, this species was widely recorded in Italy (Berlese 1883, 1885; Canestrini 1889; Wood 1973; Bernini et al. 1995). Although the morphological features of these Italian specimens were only superficially described, detailed descriptions were later provided based on North American material (Summers 1957; Summers &amp; Price 1961). Wood (1973) re-examined Italian specimens housed in the Berlese collection and, by comparisons with the descriptions given in those publications, confirmed the identification of the North American material.</p><p>Eustigmaeus clavatus has been recorded in Austria, Canada, China, Italy, Poland, Russia, Slovakia, Switzerland, Ukraine, and the USA (Beron 2020) and its identification has largely relied on the descriptions by Summers &amp; Price (1961) or Wainstein &amp; Kuznetsov (1978) based on the specimen(s) from Russia (Khaustov et al. 2023). However, the identity of North American species described by Summers &amp; Price (1961) and true E. clavatus remains uncertain because a redescription based on the type material is lacking (Khaustov et al. 2023). In addition, Khaustov et al. (2023) pointed out that E. clavatus sensu Wainstein &amp; Kuznetsov (1978) is conspecific with E. jiangxiensis Hu, Chen &amp; Huang, 1996 based on characters such as the lack of seta 4a.</p><p>The present specimens are most similar to E. clavatus sensu Summers &amp; Price (1961), sharing prominent circular dimples enclosed in evident subcuticular reticulum on dorsal shields, slightly clavate dorsal setae with sheath, a medium-length seta sci, the absence of callosity, three pairs of aggenital setae, and the absence of ω on tarsus IV. However, the description in Summers &amp; Price (1961) lacks some measurements and details of the leg structures, leaving open the possibility that the Japanese species is a distinct taxon. Therefore, we tentatively identified the examined specimens as E. clavatus sensu Summers &amp; Price (1961) . Revisiting the type specimen of E. clavatus, as well as the specimens examined in Summers (1957) and Summers &amp; Price (1961), is necessary to determine the taxonomic status of the examined specimens.</p></div>	https://treatment.plazi.org/id/03FA922DFFDEFFB16699FDC6B5855353	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ikeda, Satsuki;Imada, Yume;Nakano, Takafumi	Ikeda, Satsuki, Imada, Yume, Nakano, Takafumi (2026): Two new species and new country records of Eustigmaeus (Trombidiformes: Prostigmata: Stigmaeidae) in Japan. European Journal of Taxonomy 1082: 1-40, DOI: 10.5852/ejt.2026.1082.3312, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3312/14560
