taxonID	type	description	language	source
03FA87F4FE4FFFEBFF6DEF10FE56FAF8.taxon	description	(Figs 1 i, 5 1 – 7)	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4FFFEBFF6DEF10FE56FAF8.taxon	materials_examined	Holotype: ♂, Sri Lanka, Ratnapura district, Uda Walawe (MNHN). Materials examined: 8 ♂♂, 7 ♀♀, India, Kerala, Palakkad, Parambikulam Tiger Reserve, Vengoli road, 10.409 N, 76.770 E, 628 m, 29. X. 2022, Bindu. L & party, ZSI / WGRC / IR / INV. 30342 – 30356 [ZSIK]; 7 ♂♂, India, Kerala, Palakkad, Parambikulam Tiger Reserve, Tunnel entry Kurukku, 10.401 N, 76.755 E, 607 m, 3. XI. 2022, Bindu. L & party, ZSI / WGRC / IR / INV. 30357 – 30363 [ZSIK].	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4FFFEBFF6DEF10FE56FAF8.taxon	diagnosis	Diagnosis. Length 10 – 12 mm. Body black; head shiny, coarsely and densely punctured, punctures confluent forming coarse rugae; antennae short with distinct four segmented club, the seventh segment only slightly dilated attached to the club; mentum broad trapezoid, distinctly keeled in the middle, both sides of the keel depressed. Pronotum shiny, coarsely and densely punctured, punctures medially sparse, laterally rugose; anterior margin straight without border, posterior margin finely bordered, lateral margins slightly curved, posteriorly straightened; anterior corners obtuse, posterior corners slightly emarginate; scutellum triangular, shiny, finely punctured. Prosternal process linguiform, coarsely punctured, blunt, rounded at apex; mesosternum rough, coarsely punctured; metasternum in the middle almost impunctate, laterally with very fine wrinkles and punctures. Elytra matt, interspaces convex finely punctured, elytral punctures with striae. First three abdominal segments laterally finely rugose, fourth and fifth segments finely punctured without rugae. Apicale of the aedeagus with broad parameres, parameres slightly notched apico-laterally, thereafter rounded, then slightly constricted.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4FFFEBFF6DEF10FE56FAF8.taxon	distribution	Distribution. India (New record): Kerala. Elsewhere: Sri Lanka.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4FFFEBFF6DEDA4FE45FD95.taxon	diagnosis	Diagnosis. Body oblong, sides sub-parallel. Head broad, eyes usually large somewhat flattened, reniform. Antennae may or may not project beyond the pronotal base, distal four or five segments forming distinct club. Mentum broad, trapezoid either keeled or raised in the middle. Pronotum broad, transversely arched or disc shaped, lateral margins slightly or distinctly rounded. Elytral interspaces may be flat, keeled or slightly raised; glabrous or with hairs; with or without raised areas. Elytral epipleura broad, does not reach the apex. First three tarsi of forelegs and midlegs as well as first two tarsi of hindlegs expanded, densely hairy below; tarsi before the penultimate one deeply lobed (based on Kaszab 1976).	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4FFFE8FF6DE8BDFF1FFEE9.taxon	description	(Figs 1 ii, 6 8 – 15)	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4FFFE8FF6DE8BDFF1FFEE9.taxon	materials_examined	Holotype: ♀, India, Jharkhand, Konbir (MNHN). Material examined: 1 ♂, 4 ♀♀, India, Tamil Nadu, Coimbatore, Chitrar, 10.459 N, 76.972 E, 326 m, 8. III. 2024, R. Babu & party, ZSI / WGRC / IR / INV. 28758 – 28762 [ZSIK]; 1 ♀, India, Tamil Nadu, Erode, Sujjil Kuttai, 11.506 N, 77.074 E, 293.8 m, 16. II. 2024, R. Sakthivel & party, ZSI / WGRC / IR / INV. 27298 [ZSIK]; 1 ♀, India, Tamil Nadu, Thoothukudi, Vallanadu, 8.719 N, 77.851 E, 210 m, 23. IX. 2018, Ranjana & party, ZSI / WGRC / IR / INV. 26959 [ZSIK].	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4FFFE8FF6DE8BDFF1FFEE9.taxon	diagnosis	Diagnosis. Larger species. Body length 12 – 14 mm. Body glossy black; head broad coarsely rugose and punctate; antenna with second segment significantly shorter than the fourth; mentum trapezoid, keeled in the middle, sides of the keel finely punctured, not depressed. Pronotum broad, coarsely and densely punctured, punctures almost confluent; anterior margin straight, lateral margins broadly rounded; anterior corner blunt, posterior corner slightly protruding. Prosternal process linguiform, basal half coarsely punctured, apical half rugose, dilated at the middle, blunt, rounded at apex; mesosternum in the middle narrowly impunctate, laterally with dense coarse punctures; metasternum smooth with very fine punctures, laterally finely wrinkled. Elytral punctures with striae, interspaces strongly convex. Abdomen finely punctured in the middle, all segments except fifth, laterally with coarse rugae. Apicale of the aedeagus with long, narrow parameres, parameres apico-laterally with distinct notch.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4FFFE8FF6DE8BDFF1FFEE9.taxon	distribution	Distribution. India: Jharkhand, Uttar Pradesh, Chhattisgarh, Maharashtra, West Bengal, Tamil Nadu (New record).	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4CFFE8FF6DE941FD0DF8EA.taxon	description	(Figs 3 v, 7 16 – 23)	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4CFFE8FF6DE941FD0DF8EA.taxon	materials_examined	Lectotype: ♂, India, Bengal, Barway (designated by Kaszab, 1976) (MNHN). Material examined: 1 ♀, India, Tamil Nadu, Salem, Seeragapadi, 11.581 N, 78.051 E, 279 m, 12. V. 2024, Balaji, ZSI / WGRC / IR / INV. 26819 [ZSIK]; 1 ♀, India, Jharkhand, Simdega, Kolebira dam, Palkot Wildlife Sanctuary, 22.7 N, 84.71 E, 412 m, 24. VII. 2021, Chitra. J & party, ZSI / WGRC / IR / INV. 23627 [ZSIK]; 1 ♀, India, Chhattisgarh, Bilaspur, Karidongri FRH, 22.348 N, 81.609 E, 380 m, 19. IX. 2012, Raha. A & party, ZSI / WGRC / IR / INV. 23626 [ZSIK]; 1 ♂, India, Odisha, Raygada, Shastri Nagar, 19.175 N, 83.404 E, 229 m, 18. V. 2025, Debora, ZSI / WGRC / IR / INV. 30422 [ZSIK].	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4CFFE8FF6DE941FD0DF8EA.taxon	diagnosis	Diagnosis. Length 10 – 11 mm. Body matt black; head coarsely and densely punctured, slightly rugose; antennae reaching the pronotal base, four segmented club, second segment almost as long as fourth; mentum trapezoid, raised in the middle, without distinct keel, rugose. Pronotum broad, widest beyond the middle; coarsely and densely punctured, partially rugose; anterior margin straight, lateral margins broadly curved irregular; anterior corner obtuse, posterior corner slightly emarginate. Protsernal process cuneiform, dilated in the middle, apex rounded. Elytral interspaces completely flat, punctural rows without striae, each puncture large, elongated. First four abdominal segments punctured medially, rugose laterally, fifth abdominal segment with only punctures, without rugae. Apicale of the aedeagus with elongated, narrow parameres obscurely notched apico-laterally, thereafter vaguely round.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4CFFE8FF6DE941FD0DF8EA.taxon	distribution	Distribution. India: Chhattisgarh, Jharkhand (New record), Kerala, Maharashtra, Odisha (New record), Tamil Nadu (New record), Madhya Pradesh, West Bengal.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4CFFE8FF6DECCCFDC6FC6B.taxon	description	(Fig. 2 iii)	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4CFFE8FF6DECCCFDC6FC6B.taxon	materials_examined	Lectotype: ♀, India, Madras (designated by Kaszab, 1976) (MNHN). Materials examined: None.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4CFFE8FF6DECCCFDC6FC6B.taxon	diagnosis	Diagnosis. Elongated black glossy body, slightly convex. Head coarsely and densely punctate, dense punctures in part forming rugae; antennae long, thin projecting beyond the pronotal base. Pronotum coarsely and densely punctured, widest at the middle; anterior margin straight, lateral margin slightly rounded. Prosternal process broad, U- shaped, apex broadly rounded, smooth; mesosternum rough; metasternum almost smooth with very fine punctures; scutellum triangular coarsely and sparsely punctured. Elytral base shagreened, oily, and shiny, elytral punctures with striae, striae deep; elytral interspaces only lightly raised, interspaces with fine punctures. Abdomen coarsely punctured. Apicale of the aedeagus with elongated, narrow parameres, parameres obscurely notched apico-laterally, thereafter slightly rounded.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4CFFE8FF6DECCCFDC6FC6B.taxon	distribution	Distribution: India: Tamil Nadu, Odisha, West Bengal, Karnataka. Elsewhere. Sri Lanka, Thailand.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFECFF6DEDECFE0DFD6D.taxon	description	(Fig. 3 vi)	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFECFF6DEDECFE0DFD6D.taxon	materials_examined	Lectotype: ♀, India, Uttar Pradesh, Moradabad (designated by Kaszab, 1976) (MNHN). Material examined: None.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFECFF6DEDECFE0DFD6D.taxon	diagnosis	Diagnosis. Body black, moderately shiny; head coarsely and densely punctured, punctures confluent forming rugae laterally and at the base; frons convex; mentum trapezoid, rugose, in the middle the rugae prominently raised, but does not form distinct keel. Pronotum densely and coarsely punctured, punctures not confluent; anterior margin convex, posterior margin concave, lateral margin broadly curved; anterior and posterior corners rounded. Prosternl process coarsely punctured, elongated, narrow, broadly rounded at apex; mesosternum coarsely wrinkled with coarse punctures; metasternum with fine and sparse punctures. Elytral base shiny; punctures with striae, striae deeply furrowed, interspaces convex. Abdomen coarsely punctured. Apicale of the aedeagus with narrow, elongated parameres, notched apico-laterally, thereafter rounded, constricted at the middle.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFECFF6DEDECFE0DFD6D.taxon	distribution	Distribution: India: Uttar Pradesh. Elsewhere. Nepal, China.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFECFF6DEE48FCADFA2A.taxon	description	(Fig. 4 vii)	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFECFF6DEE48FCADFA2A.taxon	materials_examined	Syntype: ♀, Japan, Osaka (MNHN). Materials examined: None.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFECFF6DEE48FCADFA2A.taxon	diagnosis	Diagnosis. Body black, dull colored above glossy below; head coarsely rugose and punctate, frons slightly depressed; mentum trapezoid, coarsely rugose, without distinct keel, but only raised in the middle. Pronotum broad; coarse and dense confluent punctures forming rugae in pronotal disc; anterior margin straight, posterior margin slightly bisinuate, lateral margin broadly rounded; anterior corners acutely produced forward, posterior corners acute protruding; anterior and posterior margins bordered laterally, medially unbordered, lateral margins distinctly bordered. Prosternal process coarsely rugose and punctate, broad cuneiform converging to a rounded point at apex; mesosternum roughly textured with coarse rugae; metasternum coarsely punctured, rugose basally; scutellum slightly convex, rugose and punctate. Elytra slightly wider than pronotum, humeri slightly swollen; all elytral interspaces convex, eighth interspace keeled. Abdomen densely punctate. Apicale of the aedeagus with relatively narrow, elongated parameres, parameres entire, not notched apico-laterally.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFECFF6DEE48FCADFA2A.taxon	distribution	Distribution: India: Kashmir. Elsewhere. Japan, Korea, China, Vietnam, Laos, Taiwan.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFEDFF6DE882FCA9FF75.taxon	description	(Fig. 4 viii; Fig. 8 24 – 31)	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFEDFF6DE882FCA9FF75.taxon	materials_examined	Material examined: 1 ♂, India, Kerala, Idukki, Marayoor FRH, 10.277 N, 77.158 E, 1035 m, 6. XI. 2023, R. Babu & party, ZSI / WGRC / IR / INV. 29051 [ZSIK].	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFEDFF6DE882FCA9FF75.taxon	diagnosis	Diagnosis. Length 11 mm. Body shining black; head coarsely punctured, frons slightly convex laterally rugose longitudinally; antennae just reaching the pronotal base, second segment almost as long as fourth, with four segmented club, seventh segment only slightly dilated; mentum trapezoid, riased in the middle, with coarse punctures. Pronotum broad, coarsely punctured, punctures separated; anterior margin straight, lateral margins only slightly curved; anterior and posterior angles obtuse. Prosternal process narrow cuneiform, coarsely rugose, narrowly rounded at apex; mesosternum rough with coarse and dense punctures; metasternum with fine wrinkles and punctures. Elytral interspaces convex, punctural rows with striae. First four abdominal segments medially punctured, coarsely rugose laterally, fifth segment without rugae. Apicale of the aedeagus with slightly broadened parameres, parameres slightly notched apico-laterally, thereafter only slightly rounded.	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFEDFF6DE882FCA9FF75.taxon	distribution	Distribution. India: Maharashtra, Kerala (New record).	en	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
