identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03FA87F4FE4FFFEBFF6DEF10FE56FAF8.text	03FA87F4FE4FFFEBFF6DEF10FE56FAF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterotarsus ceylonicus Kaszab 1976	<div><p>Heterotarsus ceylonicus Kaszab, 1976</p><p>(Figs 1 i, 5 1–7)</p><p>Heterotarsus ceylonicus Kaszab 1976: 48 .</p><p>Holotype: ♂, Sri Lanka, Ratnapura district, Uda Walawe (MNHN) .</p><p>Materials examined: 8 ♂♂, 7 ♀♀, India, Kerala, Palakkad, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=76.77&amp;materialsCitation.latitude=10.409" title="Search Plazi for locations around (long 76.77/lat 10.409)">Parambikulam Tiger Reserve, Vengoli road</a>, 10.409 N, 76.770 E, 628 m, 29.X.2022, Bindu. L &amp; party, ZSI/WGRC/IR/INV. 30342–30356 [ZSIK] ; 7 ♂♂, India, Kerala, Palakkad, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=76.755&amp;materialsCitation.latitude=10.401" title="Search Plazi for locations around (long 76.755/lat 10.401)">Parambikulam Tiger Reserve, Tunnel entry Kurukku</a>, 10.401 N, 76.755 E, 607 m, 3.XI.2022, Bindu. L &amp; party, ZSI/WGRC/IR/INV. 30357–30363 [ZSIK] .</p><p>Diagnosis. Length 10–12 mm. Body black; head shiny, coarsely and densely punctured, punctures confluent forming coarse rugae; antennae short with distinct four segmented club, the seventh segment only slightly dilated attached to the club; mentum broad trapezoid, distinctly keeled in the middle, both sides of the keel depressed. Pronotum shiny, coarsely and densely punctured, punctures medially sparse, laterally rugose; anterior margin straight without border, posterior margin finely bordered, lateral margins slightly curved, posteriorly straightened; anterior corners obtuse, posterior corners slightly emarginate; scutellum triangular, shiny, finely punctured.Prosternal process linguiform, coarsely punctured, blunt, rounded at apex; mesosternum rough, coarsely punctured; metasternum in the middle almost impunctate, laterally with very fine wrinkles and punctures. Elytra matt, interspaces convex finely punctured, elytral punctures with striae. First three abdominal segments laterally finely rugose, fourth and fifth segments finely punctured without rugae.Apicale of the aedeagus with broad parameres, parameres slightly notched apico-laterally, thereafter rounded, then slightly constricted.</p><p>Distribution. India (New record): Kerala.</p><p>Elsewhere: Sri Lanka.</p></div>	https://treatment.plazi.org/id/03FA87F4FE4FFFEBFF6DEF10FE56FAF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	T. K, Viswanath;Hegde, Vishwanath D.;K. M., Remia	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4FFFEBFF6DEDA4FE45FD95.text	03FA87F4FE4FFFEBFF6DEDA4FE45FD95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterotarsus Latreille 1829	<div><p>Genus Heterotarsus Latreille, 1829</p><p>Diagnosis. Body oblong, sides sub-parallel. Head broad, eyes usually large somewhat flattened, reniform. Antennae may or may not project beyond the pronotal base, distal four or five segments forming distinct club. Mentum broad, trapezoid either keeled or raised in the middle. Pronotum broad, transversely arched or disc shaped, lateral margins slightly or distinctly rounded. Elytral interspaces may be flat, keeled or slightly raised; glabrous or with hairs; with or without raised areas. Elytral epipleura broad, does not reach the apex. First three tarsi of forelegs and midlegs as well as first two tarsi of hindlegs expanded, densely hairy below; tarsi before the penultimate one deeply lobed (based on Kaszab 1976).</p></div>	https://treatment.plazi.org/id/03FA87F4FE4FFFEBFF6DEDA4FE45FD95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	T. K, Viswanath;Hegde, Vishwanath D.;K. M., Remia	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4FFFE8FF6DE8BDFF1FFEE9.text	03FA87F4FE4FFFE8FF6DE8BDFF1FFEE9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterotarsus crenatus Fairmaire 1894	<div><p>Heterotarsus crenatus Fairmaire, 1894</p><p>(Figs 1 ii, 6 8–15)</p><p>Heterotarsus crenatus Fairmaire 1894: 23 .</p><p>Holotype: ♀, India, Jharkhand, Konbir (MNHN) .</p><p>Material examined: 1 ♂, 4 ♀♀, India, Tamil Nadu, Coimbatore, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=76.972&amp;materialsCitation.latitude=10.459" title="Search Plazi for locations around (long 76.972/lat 10.459)">Chitrar</a>, 10.459 N, 76.972 E, 326 m, 8.III.2024, R. Babu &amp; party, ZSI/WGRC/IR/INV. 28758–28762 [ZSIK] ; 1 ♀, India, Tamil Nadu, Erode, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.074&amp;materialsCitation.latitude=11.506" title="Search Plazi for locations around (long 77.074/lat 11.506)">Sujjil Kuttai</a>, 11.506 N, 77.074 E, 293.8 m, 16.II.2024, R. Sakthivel &amp; party, ZSI/WGRC/IR/INV. 27298 [ZSIK] ; 1 ♀, India, Tamil Nadu, Thoothukudi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.851&amp;materialsCitation.latitude=8.719" title="Search Plazi for locations around (long 77.851/lat 8.719)">Vallanadu</a>, 8.719 N, 77.851 E, 210 m, 23.IX.2018, Ranjana &amp; party, ZSI/WGRC/IR/INV. 26959 [ZSIK] .</p><p>Diagnosis. Larger species. Body length 12–14 mm. Body glossy black; head broad coarsely rugose and punctate; antenna with second segment significantly shorter than the fourth; mentum trapezoid, keeled in the middle, sides of the keel finely punctured, not depressed. Pronotum broad, coarsely and densely punctured, punctures almost confluent; anterior margin straight, lateral margins broadly rounded; anterior corner blunt, posterior corner slightly protruding. Prosternal process linguiform, basal half coarsely punctured, apical half rugose, dilated at the middle, blunt, rounded at apex; mesosternum in the middle narrowly impunctate, laterally with dense coarse punctures; metasternum smooth with very fine punctures, laterally finely wrinkled. Elytral punctures with striae, interspaces strongly convex. Abdomen finely punctured in the middle, all segments except fifth, laterally with coarse rugae. Apicale of the aedeagus with long, narrow parameres, parameres apico-laterally with distinct notch.</p><p>Distribution. India: Jharkhand, Uttar Pradesh, Chhattisgarh, Maharashtra, West Bengal, Tamil Nadu (New record).</p></div>	https://treatment.plazi.org/id/03FA87F4FE4FFFE8FF6DE8BDFF1FFEE9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	T. K, Viswanath;Hegde, Vishwanath D.;K. M., Remia	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4CFFE8FF6DE941FD0DF8EA.text	03FA87F4FE4CFFE8FF6DE941FD0DF8EA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterotarsus foveoseriatus Fairmaire 1894	<div><p>Heterotarsus foveoseriatus Fairmaire, 1894</p><p>(Figs 3 v, 7 16–23)</p><p>Heterotarsus foveoseriatus Fairmaire 1894: 24 .</p><p>Lectotype: ♂, India, Bengal, Barway (designated by Kaszab, 1976) (MNHN) .</p><p>Material examined: 1 ♀, India, Tamil Nadu, Salem, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=78.051&amp;materialsCitation.latitude=11.581" title="Search Plazi for locations around (long 78.051/lat 11.581)">Seeragapadi</a>, 11.581 N, 78.051 E, 279 m, 12. V .2024, Balaji, ZSI/ WGRC /IR/INV. 26819 [ZSIK] ; 1 ♀, India, Jharkhand, Simdega, Kolebira dam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=84.71&amp;materialsCitation.latitude=22.7" title="Search Plazi for locations around (long 84.71/lat 22.7)">Palkot Wildlife Sanctuary</a>, 22.7 N, 84.71 E, 412 m, 24.VII.2021, Chitra. J &amp; party, ZSI/ WGRC /IR/INV. 23627 [ZSIK] ; 1 ♀, India, Chhattisgarh, Bilaspur, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=81.609&amp;materialsCitation.latitude=22.348" title="Search Plazi for locations around (long 81.609/lat 22.348)">Karidongri</a> FRH, 22.348 N, 81.609 E, 380 m, 19.IX.2012, Raha. A &amp; party, ZSI/ WGRC /IR/INV. 23626 [ZSIK] ; 1 ♂, India, Odisha, Raygada, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=83.404&amp;materialsCitation.latitude=19.175" title="Search Plazi for locations around (long 83.404/lat 19.175)">Shastri Nagar</a>, 19.175 N, 83.404 E, 229 m, 18. V.2025, Debora, ZSI/ WGRC /IR/INV. 30422 [ZSIK] .</p><p>Diagnosis. Length 10–11 mm. Body matt black; head coarsely and densely punctured, slightly rugose; antennae reaching the pronotal base, four segmented club, second segment almost as long as fourth; mentum trapezoid, raised in the middle, without distinct keel, rugose. Pronotum broad, widest beyond the middle; coarsely and densely punctured, partially rugose; anterior margin straight, lateral margins broadly curved irregular; anterior corner obtuse, posterior corner slightly emarginate. Protsernal process cuneiform, dilated in the middle, apex rounded. Elytral interspaces completely flat, punctural rows without striae, each puncture large, elongated. First four abdominal segments punctured medially, rugose laterally, fifth abdominal segment with only punctures, without rugae. Apicale of the aedeagus with elongated, narrow parameres obscurely notched apico-laterally, thereafter vaguely round.</p><p>Distribution. India: Chhattisgarh, Jharkhand (New record), Kerala, Maharashtra, Odisha (New record), Tamil Nadu (New record), Madhya Pradesh, West Bengal.</p></div>	https://treatment.plazi.org/id/03FA87F4FE4CFFE8FF6DE941FD0DF8EA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	T. K, Viswanath;Hegde, Vishwanath D.;K. M., Remia	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE4CFFE8FF6DECCCFDC6FC6B.text	03FA87F4FE4CFFE8FF6DECCCFDC6FC6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterotarsus indicus subsp. indicus Marseul 1876	<div><p>Heterotarsus indicus indicus Marseul, 1876</p><p>(Fig. 2 iii)</p><p>Heterotarsus indicus indicus Marseul 1876: 128 .</p><p>Lectotype: ♀, India, Madras (designated by Kaszab, 1976) (MNHN) .</p><p>Materials examined: None.</p><p>Diagnosis. Elongated black glossy body, slightly convex. Head coarsely and densely punctate, dense punctures in part forming rugae; antennae long, thin projecting beyond the pronotal base. Pronotum coarsely and densely punctured, widest at the middle; anterior margin straight, lateral margin slightly rounded. Prosternal process broad, U- shaped, apex broadly rounded, smooth; mesosternum rough; metasternum almost smooth with very fine punctures; scutellum triangular coarsely and sparsely punctured. Elytral base shagreened, oily, and shiny, elytral punctures with striae, striae deep; elytral interspaces only lightly raised, interspaces with fine punctures. Abdomen coarsely punctured. Apicale of the aedeagus with elongated, narrow parameres, parameres obscurely notched apico-laterally, thereafter slightly rounded.</p><p>Distribution: India: Tamil Nadu, Odisha, West Bengal, Karnataka.</p><p>Elsewhere. Sri Lanka, Thailand.</p></div>	https://treatment.plazi.org/id/03FA87F4FE4CFFE8FF6DECCCFDC6FC6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	T. K, Viswanath;Hegde, Vishwanath D.;K. M., Remia	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFECFF6DEDECFE0DFD6D.text	03FA87F4FE48FFECFF6DEDECFE0DFD6D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterotarsus auriculatus Marseul 1876	<div><p>Heterotarsus auriculatus Marseul, 1876</p><p>(Fig. 3 vi)</p><p>Heterotarsus auriculatus Marseul 1876: 128 .</p><p>Lectotype: ♀, India, Uttar Pradesh, Moradabad (designated by Kaszab, 1976) (MNHN) .</p><p>Material examined: None.</p><p>Diagnosis. Body black, moderately shiny; head coarsely and densely punctured, punctures confluent forming rugae laterally and at the base; frons convex; mentum trapezoid, rugose, in the middle the rugae prominently raised, but does not form distinct keel. Pronotum densely and coarsely punctured, punctures not confluent; anterior margin convex, posterior margin concave, lateral margin broadly curved; anterior and posterior corners rounded. Prosternl process coarsely punctured, elongated, narrow, broadly rounded at apex; mesosternum coarsely wrinkled with coarse punctures; metasternum with fine and sparse punctures. Elytral base shiny; punctures with striae, striae deeply furrowed, interspaces convex. Abdomen coarsely punctured. Apicale of the aedeagus with narrow, elongated parameres, notched apico-laterally, thereafter rounded, constricted at the middle.</p><p>Distribution: India: Uttar Pradesh.</p><p>Elsewhere. Nepal, China.</p></div>	https://treatment.plazi.org/id/03FA87F4FE48FFECFF6DEDECFE0DFD6D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	T. K, Viswanath;Hegde, Vishwanath D.;K. M., Remia	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFECFF6DEE48FCADFA2A.text	03FA87F4FE48FFECFF6DEE48FCADFA2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterotarsus carinula Marseul 1876	<div><p>Heterotarsus carinula Marseul, 1876</p><p>(Fig. 4 vii)</p><p>Heterotarsus carinula Marseul 1876: 127 .</p><p>Syntype: ♀, Japan, Osaka (MNHN) .</p><p>Materials examined: None.</p><p>Diagnosis. Body black, dull colored above glossy below; head coarsely rugose and punctate, frons slightly depressed; mentum trapezoid, coarsely rugose, without distinct keel, but only raised in the middle. Pronotum broad; coarse and dense confluent punctures forming rugae in pronotal disc; anterior margin straight, posterior margin slightly bisinuate, lateral margin broadly rounded; anterior corners acutely produced forward, posterior corners acute protruding; anterior and posterior margins bordered laterally, medially unbordered, lateral margins distinctly bordered. Prosternal process coarsely rugose and punctate, broad cuneiform converging to a rounded point at apex; mesosternum roughly textured with coarse rugae; metasternum coarsely punctured, rugose basally; scutellum slightly convex, rugose and punctate. Elytra slightly wider than pronotum, humeri slightly swollen; all elytral interspaces convex, eighth interspace keeled. Abdomen densely punctate. Apicale of the aedeagus with relatively narrow, elongated parameres, parameres entire, not notched apico-laterally.</p><p>Distribution: India: Kashmir.</p><p>Elsewhere. Japan, Korea, China, Vietnam, Laos, Taiwan.</p></div>	https://treatment.plazi.org/id/03FA87F4FE48FFECFF6DEE48FCADFA2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	T. K, Viswanath;Hegde, Vishwanath D.;K. M., Remia	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE48FFEDFF6DE882FCA9FF75.text	03FA87F4FE48FFEDFF6DE882FCA9FF75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterotarsus topali Kaszab 1976	<div><p>Heterotarsus topali Kaszab, 1976</p><p>(Fig. 4 viii; Fig. 8 24–31)</p><p>Heterotarsus topali Kaszab 1976: 48 .</p><p>Holotype: ♂, India, Maharashtra, Kanheri (NHMUS) .</p><p>Material examined: 1 ♂, India, Kerala, Idukki, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.158&amp;materialsCitation.latitude=10.277" title="Search Plazi for locations around (long 77.158/lat 10.277)">Marayoor</a> FRH, 10.277 N, 77.158 E, 1035 m, 6.XI.2023, R. Babu &amp; party, ZSI/ WGRC /IR/INV. 29051 [ZSIK] .</p><p>Diagnosis. Length 11 mm. Body shining black; head coarsely punctured, frons slightly convex laterally rugose longitudinally; antennae just reaching the pronotal base, second segment almost as long as fourth, with four segmented club, seventh segment only slightly dilated; mentum trapezoid, riased in the middle, with coarse punctures. Pronotum broad, coarsely punctured, punctures separated; anterior margin straight, lateral margins only slightly curved; anterior and posterior angles obtuse. Prosternal process narrow cuneiform, coarsely rugose, narrowly rounded at apex; mesosternum rough with coarse and dense punctures; metasternum with fine wrinkles and punctures. Elytral interspaces convex, punctural rows with striae. First four abdominal segments medially punctured, coarsely rugose laterally, fifth segment without rugae. Apicale of the aedeagus with slightly broadened parameres, parameres slightly notched apico-laterally, thereafter only slightly rounded.</p><p>Distribution. India: Maharashtra, Kerala (New record).</p></div>	https://treatment.plazi.org/id/03FA87F4FE48FFEDFF6DE882FCA9FF75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	T. K, Viswanath;Hegde, Vishwanath D.;K. M., Remia	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
03FA87F4FE42FFE6FF6DEDECFA75FC61.text	03FA87F4FE42FFE6FF6DEDECFA75FC61.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterotarsus Latreille 1829	<div><p>Identification key to the species of Heterotarsus from India (based on Kaszab 1976)</p><p>1. Anterior corners of pronotum produced forward, acute................................ H. carinula Marseul, 1876</p><p>– Anterior pronotal corners not produced forward.......................................................... 2</p><p>2. Mentum medially with distinct keel .................................................................... 3</p><p>– Mentum only raised in the middle, not forming distinct keel................................................ 4</p><p>3. 2 nd antennal segment significantly smaller than 4 th segment; lateral pronotal margin broadly curved; sides of keel in mentum not depressed; prosternal process linguiform basal half coarsely punctured, apical half rugose, dilated at the middle; aedeagus with long, narrow parameres, parameres apico-laterally with distinct notch....... H. crenatus Fairmaire, 1894</p><p>– 2 nd antennal segment almost as long as 4 th segment; lateral pronotal margin only slightly curved; sides of the keel in mentum depressed; prosternal process linguiform, not dilated at the middle, coarsely punctured; aedeagus with broad parameres, parameres only slightly notched apico-laterally.................................... H. ceylonicus Kaszab, 1976</p><p>4. Elytral interspaces completely flat; punctural rows without striae, individual punctures large, elongated.................................................................................... H. foveoseriatus Fairmaire, 1894</p><p>– Elytral interspaces raised; punctural rows with striae...................................................... 5</p><p>5. Antennae just reach or slightly project beyond pronotal base; parameres very slightly broadened.................................................................................................. H. topali Kaszab, 1976</p><p>– Antennae long; projecting beyond the pronotal base; parameres elongated, narrow............................ ... 6</p><p>6. Anterior pronotal margin convex; elytral striae deeply furrowed; posterior corner of pronotum rounded....................................................................................... H. auriculatus Marseul, 1876</p><p>– Anterior pronotal margin straight; elytral striae deeply or shallowly furrowed; posterior corner of pronotum emarginated or almot straight without emargination................................................................... 7</p><p>7. Pronotum coarsely punctured, punctures almost confluent; lateral sides of pronotum only gently curved, near to the base almost straight; scutellum triangular coarsely punctured;interspaces near the base roughened; parameres elongated, narrow, obscurely notched apico-laterally, thereafter slightly rounded..................... H. indicus indicus Marseul, 1876</p><p>– Pronotum coarsely punctured, punctures separate; lateral sides of pronotum comparatively more curved, near to the base slightly emarginated; scutellum triangular with rounded sides, almost smooth; interspaces near the base not roughened; parameres elongated narrow, entire,slightly emarginated apico-laterally, thereafter not rounded, straight............................................................... H. indicus besucheti Kaszab, 1976 [extra limital] (Fig. 2 iv)</p></div>	https://treatment.plazi.org/id/03FA87F4FE42FFE6FF6DEDECFA75FC61	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	T. K, Viswanath;Hegde, Vishwanath D.;K. M., Remia	T. K, Viswanath, Hegde, Vishwanath D., K. M., Remia (2026): The genus Heterotarsus Latreille (Coleoptera: Tenebrionidae: Opatrini) in India. Zootaxa 5782 (2): 347-360, DOI: 10.11646/zootaxa.5782.2.6, URL: http://dx.doi.org/10.7717/peerj.20736
