taxonID	type	description	language	source
03FA87E4FF8BFFCCFF06FC0D79B43700.taxon	type_taxon	Type species: Agudus typicus Oman, 1938	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8BFFCCFF06FC0D79B43700.taxon	description	Description. Relatively large, cylindrical, somewhat elongate paralimnines (4.5 to 9.5 mm long); color stramineous, with either paired red longitudinal stripes or symmetrical black spots, or both on dorsum of head and thorax. Head with crown flat, crown texture shagreen anteriorly and laterally near eyes, postfrontal suture (ecdysial line) distinct with texture posterad of line smooth and shiny, shagreen, or some combination or grade of both, coronal suture sometimes furrowed with adjacent parts of crown slightly inflated, head produced and acutely angulate anteriorly in dorsal view, much longer than basal width between eyes, crown-face transition acutely angulate in profile; lorum not extended to genal margin; anteclypeus convex, distinctly tapered distad. Forewing variably developed, may be shortened to some extent in one or both sexes exposing up to three pregenital abdominal tergites, with three closed anteapical cells, outer cell often greatly reduced in size, appendix weakly developed or absent. Hind wing present but usually reduced in size. Front tibia dorsal macrosetae usually 4 + 4. Male pygofer much longer than tall in lateral view, often elongated, with macrosetae distributed over most of surface, lobe with or without process or tooth; valve trapezoidal; segment X well sclerotized but not extended beyond pygofer apex; subgenital plate longer than basal width, with single row of lateral or sublateral macrosetae; connective loop shaped with stem usually short and relatively broad and anterior arms convergent anterad and touching each other or fused preapically; style with preapical lobe usually angulate, apophysis variable among species, often enlarged with denticuli; aedeagus with dorsal apodeme usually well developed, shaft tubular, apex often bifid, with or without additional processes. Female sternite VII shorter than wide, with posterior margin variably shaped among species; first valvula dorsal sculpture imbricate, extended to margin; second valvulae with basal fused section toothed at base and apex, median cleft well indicated by pigment-free area, distal toothed section slightly shorter than basal section, dorsal margin slightly arcuate with 16 - 30 sparse conical teeth larger and more widely spaced toward base, then becoming smaller and more closely spaced distally; arranged somewhat asymmetrically on left and right blades.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8BFFCCFF06FC0D79B43700.taxon	discussion	Notes. Agudus is the only genus of Paralimnini recorded from South America, and species of the genus are easily distinguished from other grass specialist South American leafhoppers by their relatively large size, cylindrical body with acutely produced head, and slightly to strongly reduced forewings, as well as by the characteristic loopshaped male connective, which is articulated to the aedeagus. Available collection records suggest that this genus is restricted to the tropical and subtropical grasslands and savannas of southeastern South America, including the Cerrado, Chaco, and Pampas biomes of Argentina, Brazil and Paraguay. The genus appears to be absent from the predominantly cool season grasslands of Patagonia and the Andes Mountains.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF88FFCDFF06FAD878D532C4.taxon	description	(Figs. 2 A – B, 5 A – D, 8 A)	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF88FFCDFF06FAD878D532C4.taxon	materials_examined	Material examined. PARAGUAY: NW Asuncion, 2 km E Limpio, at light, 17 June 1975, Elzinga, Granovsky & Blocker / PARATYPE Agudus blockeri YJ Cheng / USNMENT 02110324 (1 male) [USNM]; PARAGUAY: between San Patricio y San Ramone, 7 June 1975, sweeping, Elzinga & Granovsky / PARATYPE Agudus blockeri YJ Cheng / USNMENT 02110737 (1 male) [USNM]. Other material: ARGENTINA: Entre Rios, rt. 14 km 275 ca. Concordia, 50 m, 31 ° 19 ' 15 " S 58 ° 5 ' 8 " W, 4 Jan 2008, C. H. Dietrich, vacuum, AR 1 - 1 (1 male, 4 females); ARGENTINA: Entre Rios, Federación, Sirirí Campgd., 30 m, 30 ° 58 ' 6 " S 57 ° 56 ' 38 " W, 4 Jan 2008, C. H. Dietrich, Hg vapor lights, AR 2 - 1 (1 male); ARGENTINA: Corrientes, jct rt 14 & 119, 30 ° 4 ' 35 " S 57 ° 59 ' 57 " W, 5 Jan 2008 C. H. Dietrich, vacuum, AR 3 - 1 (1 female); ARGENTINA: Corrientes, P. N. Mburucuyá, 1.8 km W campgd., 80 m, 28 ° 1 ' 11 " S 58 ° 3 ' 18 " W, 8 Jan 2008, C. H. Dietrich, vacuum, AR 9 - 4 (1 male) [MLP, INHS].	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF88FFCDFF06FAD878D532C4.taxon	discussion	Notes. Examination of a series of specimens from northern Argentina with males matching Cheng’s (1980) drawings of the male genitalia of A. blockeri and females matching DeLong & Linnavuori’s (1978) drawing of the female seventh sternite of A. amabilis indicates that the former is a junior synonym of the latter. This species is easily recognized by the red longitudinal stripes on the head and thorax, the fully developed forewings in both sexes, the broadly emarginate posterior margin of female sternite VII with a pair of indistinct, darkly pigmented submedial lobes, and the robust, posteriorly curved hooklike process on the ventral margin of the male pygofer.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF89FFCDFF06FEA37930371C.taxon	description	(Figs. 2 C, 5 E – H) Description. Body length: male 4.8 mm. Color: face dark brown with pale muscle scars; crown brown with black apical macula and paired stramineous longitudinal stripes extended posterad from midlength across pronotum and mesonotum, with additional pair of alternating light and dark stripes more laterad on pronotum and mesonotum; forewing with black macula on clavus posterad of crossvein, in inner discal cell and middle anteapical cell, with additional less distinct spots in other cells. Male crown 2 × longer than width between eyes. Forewing short, extended to but not over pygofer, outer anteapical cell petiolate apically, apical cells somewhat shortened. Male pygofer> 3 × longer than basal height in lateral view, without basal apodemes, macrosetae relatively short, scattered over most of lobe, apex broadly rounded with short, darkly sclerotized ventroapical hook, ventral margin membranous and concave. Anal tube cylindrical, well sclerotized, not extended to pygofer apex, with sclerotized ventromesal extension at base articulated to dorsal apodeme of aedeagus. Subgenital plate nearly as long as pygofer, in ventral view with lateral margin concave near base then relatively straight to obliquely tapered apex, macrosetal row not extended beyond basal two thirds. Style preapical lobe acute, apophysis robust, relatively long, clavate, evenly curved laterad, apex rounded and bearing two small teeth. Connective> 3 × longer than greatest width, anterior arms slightly divergent from stem then convergent and nearly touching near apex, stem short, constructed at base of arms, then broadened posterad with posterior margin slightly bifid. Aedeagus with dorsal apodeme narrow in lateral and posterior views, slightly broadened dorsally; shaft short, tubular, slightly depressed and curved dorsad distally; gonopore apical on posterior surface, occupying entire width of shaft. Female unknown.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF89FFCDFF06FEA37930371C.taxon	etymology	Etymology. The new species name, a Latinized noun in apposition meaning “ club style ”, refers to the clublike apex of the style.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF89FFCDFF06FEA37930371C.taxon	materials_examined	Material examined. Holotype male, ARGENTINA: Chaco, P. N. Chaco, 70 m, 26 ° 48 ' 50 " S 59 ° 36 ' 52 " W, 12 Jan 2008, T. Catanach, vacuum, AR 10 - 12 [MLP].	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF89FFCDFF06FEA37930371C.taxon	discussion	Notes. This species, described based on a single male, resembles A. typicus in external morphology and coloration but has the head more elongate and the forewings distinctly reduced, and the male genitalia are distinctly different. The coloration and proportions of the head are similar to those of A. inesae sp. nov. (described below, also from Argentina) but the latter also has distinctive male genitalia and the forewings are fully developed.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF89FFCAFF06FADB7E3F302C.taxon	description	(Figs. 2 D – E, 5 I – L, 7, 8 B) Description. Body length: male 4.8 – 5.1 mm, female 5.0 – 5.5 mm. Color: face medium brown with pale muscle scars, crown tan with pair of bright red submedial stripes extended from apex posterad, continuing over pronotum onto mesonotum, thin brown median line corresponding to coronal suture extended as thin reddish line across pronotum and mesonotum, pronotum with sublateral pair of less conspicuous red stripes; forewing veins outlined with dark brown, larger black spot in inner discal cell and inner anteapical cell. Male crown 1.4 × longer than width between eyes, female crown 1.2 × longer than width between eyes. Forewing extended to or slightly beyond apex of pygofer in male, slightly shorter than pygofer in female; outer anteapical cell truncate apically. Male pygofer 2 × longer than basal height in lateral view, without basal apodemes, macrosetae unusually long, scattered over distal two thirds, apex tapered to blunt point, ventral margin with large hook-shaped process near midlength curved posterad. Anal tube cylindrical, moderately long, completely sclerotized, not extended to pygofer apex. Subgenital plate half as long as pygofer, in ventral view narrowly triangular with lateral margin concave in distal half, macrosetal row not extended beyond basal two thirds. Style preapical lobe acutely angulate, apophysis moderately stout, tapered, curved laterad with apical tooth. Connective 2.5 × longer than maximum width, anterior arms evenly convergent anterad, fused preapically, stem broad and very short, truncate posteriorly. Aedeagus with dorsal apodeme moderately long, triangular in posterior view; shaft tubular, moderately long, curved dorsad, with pair of moderately long, stout distal processes arising laterad of gonopore, extended dorsomesad and nearly touching each other then bowed laterad and recurved to form heart-shaped space in posterior view, each process with short retrorse preapical lateral tooth; gonopore apical, occupying entire width of shaft. Female sternite VII posterior margin shallowly V-shaped with pair of very indistinct, darkly pigmented submedial lobes.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF89FFCAFF06FADB7E3F302C.taxon	etymology	Etymology. The species name is a Latin noun meaning “ heart bearer ”, referring to the heart shaped space formed by the distal processes of the aedeagus in posterior view.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF89FFCAFF06FADB7E3F302C.taxon	materials_examined	Material examined. Holotype male, BRAZIL: MG, Santuario do Caraça, Taboõs Trl. 20.08329 ºS 43.50665 ºW 1200 m 15 Mar 2015 CH Dietrich, vacuum grass, cerrado BR 15 - 03 - 5 [UFRJ]. Paratypes: 1 male, 2 females, same data as holotype [INHS]. Other material: BRAZIL: MG, Santuario do Caraça, Banho do Belchior 20.11450 ºS 43.49115 ºW 1280 m 16 Mar 2015 CH Dietrich sweep / beat BR 15 - 09 - 4 (3 males, 2 females); BRAZIL: MG, Santuario do Caraça, Lobo Guará Trl., 20.09878 ºS 43.48677 ºW, 1280 m, 13 Mar 2015, CH Dietrich, sweeping cerrado, BR 15 - 01 - 1 (4 males, 3 females) [INHS].	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8EFFCAFF06FD0B78E137F4.taxon	description	(Figs. 1 B, 2 F – G, 5 M – P, 8 C) Description. Body length: male 5.1 mm; female 5.7 – 5.9 mm. Color: face brown with pale muscle scars; crown tan with apical and medial black macula, merging together in some specimens; pronotum with three indistinct yellow longitudinal stripes extended onto mesonotum, lateral margins black; forewing with distinct black spot in clavus just distad of crossvein, in inner discal cell and in inner subapical cell. Male crown 1.8 × longer than width between eyes, female crown 2 × longer than width between eyes. Forewing extended nearly to apex of pygofer in both male and female. Male pygofer less than 2 × longer than basal height in lateral view, without basal apodemes, with darkly pigmented ovoid, denticulate sclerite on mesal surface of each lobe near apex; macrosetae, some unusually long, scattered over most of lobe; ventral margin evenly convex, without teeth or processes. Anal tube cylindrical, tapered, well-sclerotized and unusually large, extended well beyond pygofer. Style preapical lobe acute, apophysis robust, digitiform, denticulate, curved slightly laterad with apical tooth. Connective more than 3 × longer than greatest width, arms subparallel, abruptly convergent and fused preapically, stem short and broad with truncate posterior margin. Aedeagus with dorsal apodeme and shaft in lateral view forming nearly complete ellipse, dorsal apodeme long and slender, extended to dark mesal lobes of pygofer, shaft longer and more slender, with pair of long, slender distal processes extended distad, slightly bowed laterad in ventral view but with apices convergent and extended between darkly sclerotized pygofer lobes; gonopore apical. Female sternite VII posterior margin with broad, convex medial lobe flanked by pair of narrower triangular lateral lobes of same length.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8EFFCAFF06FD0B78E137F4.taxon	etymology	Etymology. The new species name, a Latin adjective, refers to the elliptical shape of the aedeagus in lateral view.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8EFFCAFF06FD0B78E137F4.taxon	materials_examined	Material examined. Holotype male, ARGENTINA: Chaco, P. N. Chaco, 70 m 26 ° 48 ' 50 " S 59 ° 36 ' 52 " W 12 Jan 2008 T. Catanach, vacuum, AR 10 - 14 [MLP]. Paratypes: ARGENTINA: Chaco, P. N. Chaco, 70 m 26 ° 48 ' 50 " S 59 ° 36 ' 52 " W 12 Jan 2008 C. H. Dietrich, vacuum, AR 10 - 5, 8, 9, 10 (2 males, 4 females) [INHS].	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8EFFCBFF06F9B37F8232C4.taxon	description	(Figs. 2 H – I, 5 Q – T, 8 D)	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8EFFCBFF06F9B37F8232C4.taxon	description	Description of female. Length 6.5 mm. Forewing at rest exposing most of pygofer in dorsal view. Abdominal sternite VII with posterior margin broadly emarginate with pair of brown obtuse submedial projections flanking shorter, acute medial lobe.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8EFFCBFF06F9B37F8232C4.taxon	materials_examined	Material examined. Male paratype, PARAGUAY: 2 km W Eusebio Ayala, 10 June 1975, Blocker, Granovsky & Elzinga / PARATYPE Agudus forficatus YJ Cheng / USNMENT 02110302 [USNM]. Other material: ARGENTINA: Entre Rios, P. N. El Palmar, Mirador Ao. El Palmar, 31.89149 ºS, 58.24071 ºW, 21 m, 15 - Feb- 2014, J. N. Zahniser, vacuum, AR 14 - 27 - 1 (1 male, 1 female) [INHS].	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8EFFCBFF06F9B37F8232C4.taxon	discussion	Notes. This species was described based on three male specimens from Paraguay. It is newly recorded from Argentina and the female is described and illustrated for the first time.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8FFFCBFF06FEA37F223419.taxon	description	(Figs. 1 A, 2 J – K, 5 U – X, 8 E) Description. Body length: male 5.8 – 7.3 mm, female 6.3 – 7.0 mm. Color: face dark brown with pale muscle scars; crown brown with black apical macula and paired stramineous longitudinal stripes extended posterad from midlength across pronotum and mesonotum, with additional pair of alternating light and dark stripes more laterad on pronotum and mesonotum; forewing with black macula on clavus posterad of crossvein, in inner discal cell and middle anteapical cell, with additional less distinct spots in other cells. Crown 1.9 to 1.8 × longer than width between eyes in male and female, respectively. Forewing extended to but not covering pygofer in male and female, outer anteapical cell not petiolate apically. Male pygofer slightly more than twice as long as maximum height, apex rounded, without processes, with elongate unpigmented submarginal area along distal two thirds of ventral margin of lobe bordered by narrow sclerotized marginal band, macrosetae scattered throughout length of lobe. Anal tube with sclerotized anteroventral extension connected by membrane to dorsal apodeme of aedeagus. Subgenital plate extended slightly more than half distance to apex of pygofer, lateral margin concave in ventral view, macrosetal row not extended beyond midlength. Style preapical lobe obtusely angulate, apophysis curved slightly laterad, with distinct preapical lateral angle, apex narrowed to blunt point. Connective 3.5 × longer than maximum width, arms slightly bowed laterad, fused preapically, stem short with posterior margin notched. Aedeagus with dorsal apodeme slender in lateral and posterior view, slightly sinuate dorsally in lateral view; shaft tubular, evenly curved dorsad, apex with pair of short lateral projections as long as shaft width extended basolaterad and pair of very slender submedial processes of same length extended distad, gonopore elongate, ovoid, occupying entire width of shaft posterior surface near apex. Female sternite VII with posterior margin broadly V-shaped, with very narrow medial notch extended to midlength.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8FFFCBFF06FEA37F223419.taxon	materials_examined	Material examined. Holotype male, ARGENTINA: Buenos Aires, R. N. Otamendi, 34.22742 ºS, 58.89921 ºW, 22 m, 17 - Feb- 2014, C. H. Dietrich, vacuum, AR 14 - 14 - 5 [MLP]. Paratypes: 2 males, same data as holotype [INHS]. Other material: ARGENTINA: Entre Rios, P. N. El Palmar, 31.86906 ºS, 58.2452 ºW, 32 m, 15 - Feb- 2014, J. N. Zahniser, vacuum, AR 14 - 26 - 2 (1 male, 1 female) [INHS]. Other material: ARGENTINA: Corrientes, jct rt 14 & 119, 30 ° 4 ' 35 " S 57 ° 59 ' 57 " W, 5 Jan 2008 C. H. Dietrich, vacuum, AR 3 - 1 (1 female) [INHS]; PARAGUAY: Chaco Expt. Sta. Cattle Res. Sta., 295 km NW Asuncion on Trans-Chaco Hwy, 20 June 1975, Blocker, Elzinga & Granovsky (2 males) [INHS]; PARAGUAY: NW Asuncion 2 km E Limpio, at light, 17 June 1975, Elzinga, Granovsky & Blocker (1 female) [INHS].	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8FFFCBFF06FEA37F223419.taxon	etymology	Etymology. The species is named in honor of Dr. Maria Inés Catalano, a leafhopper expert at Universidad Nacional del Noroeste de la Provincia de Buenos Aires, who helped facilitate collection of many of the specimens included in this study.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8FFFCBFF06FEA37F223419.taxon	discussion	Notes. This species closely resembles A. typicus in external morphology but may be distinguished by the male genitalia and the shape of the female pregenital sternite.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8FFFC8FF06F9DC7FB335CB.taxon	description	(Figs. 2 L – M, 4 A – B, 6 A – D, 8 F)	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8FFFC8FF06F9DC7FB335CB.taxon	description	Description of female. Length 7.4 – 7.6 mm (apex of head to apex of pygofer). Sternite VII posterior margin with medial triangular notch, slightly sinuate laterad of notch, posterolateral corner slightly produced; mostly strawcolored (same as body) and with diffuse brown spot laterad of medial notch on both sides.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF8FFFC8FF06F9DC7FB335CB.taxon	materials_examined	Material examined. Paratype male, PARAGUAY: 2 km W Eusebio Ayala, 10 June 1975, Blocker, Granovsky & Elzinga / PARATYPE Agudus lepidus Cheng / USNMENT 02110775. Other material: one female, ARGENTINA: Loreto, Misiones, XII- 6 - 1931, AA Ogloblin / Agudus longiceps Berg [label folded] / Agudus lepidus Cheng, Det. JN Zahniser 2026 / USNMENT 02110393 (1 female); same locality data and date (1 female); same locality, XII- 3 - 1931 (1 female) [all USNM]. Notes. This species was previously only recorded from Paraguay. Three female specimens from Loreto, Misiones Prov., Argentina collected at the same time as one of the male paratypes are identified as A. lepidus here based on their shared color pattern, dimensions of the head, and size.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF82FFC6FF06FF127D14302C.taxon	discussion	Notes. This species, originally described from Buenos Aires, Argentina, was redescribed and illustrated by Linnavuori (1959) based on the type and an additional male specimen from a locality near Buenos Aires. It is considerably larger than other known species of the genus (males 9.5 mm long) and also differs in having a darkly pigmented ventral tooth on the pygofer lobe and the apical processes of the aedeagus sharply bent laterad near the apex. The female remains unknown. Recent collecting in pampas grasslands near Buenos Aires has failed to yield additional specimens of this species. Given the extensive urbanization and agricultural intensification in the Buenos Aires region since the species was originally collected and described, it appears that this species is now either extremely rare or extinct.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF82FFC6FF06FD0A79263149.taxon	discussion	Notes. This species was described from two localities in the cerrado of central Brazil. No specimens were available for study but it is included in the above key based on the description and illustrations of Zanol (1999).	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF82FFC6FF06FC2C7E223686.taxon	discussion	Notes. This species was described from the same two localities in the cerrado of central Brazil as the previously listed species. No specimens were available for study but it is included in the above key based on the description and illustrations of Zanol (1999).	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF82FFC7FF06FB6D7F003094.taxon	description	(Figs. 2 N – O, 3 A – D, 4 C – D, 6 E – H, 8 G)	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF82FFC7FF06FB6D7F003094.taxon	description	Description of female. Length 6.5 mm. Forewing at rest covering abdomen except apex of pygofer in dorsal view. Abdominal sternite VII relatively short, posterior margin shallowly concave with pair of brown angulate submedial projections flanking short, rounded medial lobe.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF82FFC7FF06FB6D7F003094.taxon	materials_examined	Material examined. Holotype male, ARGENTINA: Loreto, Misiones, XI- 25 - 1931, AA Ogloblin / USNMENT 02110135 [USNM]. Other material: ARGENTINA: Corrientes, P. N. Mburucuyá, 1.8 km W campgd., 80 m, 28 ° 1 ' 11 " S 58 ° 3 ' 18 " W, 8 Jan 2008, C. H. Dietrich, vacuum, AR 9 - 4 (3 males) [MLP, INHS]. PARAGUAY: 95 km W. Pto Pte Stroessner, 11 Jun 1975, Blocker, Elzinga, Granovsky, sweeping (1 male, 1 female) [INHS]; ARGENTINA: Loreto, Misiones, XI- 27 - 1931, AA Ogloblin / Agudus sexmaculatus Lnv. Det. Kramer 1963 (1 male) [USNM]; same locality, XI- 29 - 1931 (1 male) [USNM]; same locality, XII- 10 - 1931 (sex undetermined, abdomen missing) [USNM]. PARAGUAY: Aregua, 20 mi. E of Asuncion, 16 Aug 1980, D. C. Lowrie (1 male) / Agudus sexmaculatus Linnavuori, Det. JN Zahniser 2026 / USNMENT 02110346 [USNM].	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF82FFC7FF06FB6D7F003094.taxon	discussion	Notes. This species is recorded from Argentina and Paraguay. The female is here described and illustrated for the first time. Linnavuori (1959) indicated that the type specimen (a male) and one female paratype were deposited at USNM and one specimen (sex not stated) was retained in his personal collection. Upon initial examination of the USNM type collection, the type was not found. However, one male specimen in the main collection in a unit tray identified as A. sexmaculatus was apparently examined by Linnavuori and the genitalia match Linnavuori’s figures. His examination of that specimen is deduced by the preparation of the genitalia which are dismembered and glued to a card, a procedure that was typical of Linnavuori for specimens he examined and illustrated, including types, and was not a technique known to be used by other New World leafhopper workers. Here, the genitalia on the card were imaged and then the parts were lifted off by soaking in water and are now stored in glycerine in a genitalia vial. This male has the locality data (Loreto, Misiones; coll. AA Ogloblin) given by Linnavuori, but the date (XI- 25 - 1931) differs slightly from what was stated for all three specimens (XII- 10 - 1931) in Linnavuori’s publication. Despite this discrepancy, this male is interpreted here as the holotype. There are many other leafhopper specimens at the USNM from the same locality with different dates of collection, and it is possible that Linnavuori incorrectly transcribed the collection dates. The female stated by Linnavuori (1959) to be deposited at USNM was not found, although it is possible the specimen listed in Material Examined (above) with the abdomen missing is this female, and the collection date of that specimen (XII- 10 - 1931) matches that reported by Linnavuori. Linnavuori (1959) stated that males of this species are 5 to 6 mm in length, and that the female he examined was 7.2 mm. However, these measurements seem to be inaccurate. The male type specimen is 4.5 mm, measured from apex of head to forewing apex (forewings surpass the abdomen in this species). The other three males at USNM are 4.5, 5.0, and 5.0 mm long. The specimen with missing abdomen is 5.2 mm.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF81FFC5FF06FF127D453395.taxon	discussion	Notes. This species was described from a male and female specimen from Paraná State, southern Brazil. No other specimens are known. It is included in the above key based on the description and illustrations of DeLong & Linnavuori (1978).	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF81FFC2FF06F8C17D9333BC.taxon	description	(Figs. 2 P, 4 E – F, 6 I – L, 8 H)	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF81FFC2FF06F8C17D9333BC.taxon	materials_examined	Material examined. ARGENTINA: Loreto, Misiones, XII- 10 - 1931, AA Ogloblin (6 males, 2 females, 1 sex undet.; one of these males with label USNMENT 02110683). Same locality, XII- 12 - 1931 (1 male); XII- 1 - 1931 (1 male); XII- 4 - 1931 / USNMENT 02110493 (1 male); XII- 18 - 1931 (1 male); XII- 6 - 1931 / allotype (1 female) [all USNM].	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF81FFC2FF06F8C17D9333BC.taxon	discussion	Notes. This species was described based on three specimens from Misiones Province, Argentina. It differs from other red-striped species of Agudus in having the scutellum almost completely red and the aedeagus with three pairs of apical processes. No recently collected specimens of this species have been found, despite recent sampling in grasslands throughout northern Argentina, suggesting that this species is now extremely rare or extinct (see above notes under A. longiceps).	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF86FFC0FF06F8AE7D663374.taxon	description	(Figs. 2 Q – R, 6 M – P, 8 I) Description. Body length: male 5.0 mm, female 5.2 mm. Color: face dark brown with pale muscle scars; crown brown with black apical macula and paired stramineous longitudinal stripes extended posterad from midlength across pronotum and mesonotum, with additional pair of alternating light and dark stripes more laterad on pronotum and mesonotum; forewing with black macula on clavus posterad of crossvein, in inner discal cell and middle anteapical cell, with additional less distinct spots in other cells. Crown 1.6 to 1.8 × longer than width between eyes in male and female, respectively. Forewing very short in both male and female, exposing three pregenital abdominal tergites, outer anteapical cell petiolate apically, apical cells very short. Male pygofer ~ 2.5 × longer than basal height in lateral view, with pair of blunt ventrally curved basal dorsolateral apodemes, macrosetae present in band extended along ventral half of lobe in distal two thirds, apex tapered to blunt point, ventral margin concave except for small round lobe near midlength, inner surface with pair of long sclerotized bars extended ventrad from base of anal tube. Anal tube short, cylindrical, well sclerotized laterally and dorsally. Subgenital plate nearly as long as pygofer, evenly tapered toward apex in ventral view, with macrosetal row well separated from lateral margin, apex curved slightly dorsad with small dorsoapical tooth. Style with preapical lobe prominent, rounded, apophysis relatively long, slender, tapered, curved laterad. Connective slender, in ventral view with anterior arms fused and indistinguishable except for small preapical gap, stem elongate, evenly broadened posterad, with rounded apex. Aedeagus relatively short, simple, preatrium and dorsal apodeme weakly developed, shaft slightly depressed, slightly curved dorsad, with arcuate dorsal preapical carina, apex tapered to point in lateral and ventral view, gonopore occupying full width of shaft on posterior surface of apex. Female sternite VII posterior margin with broad, shallow V-shaped emargination and weak median longitudinal carina.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF86FFC0FF06F8AE7D663374.taxon	etymology	Etymology. The new species name is a Latinized noun in apposition meaning “ slender style ” referring to the attenuate apex of the male style.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF86FFC0FF06F8AE7D663374.taxon	materials_examined	Material examined. Holotype male, ARGENTINA: Entre Rios, P. N. El Palmar, jct. Pref. Naval, 31.87321 ºS, 58.21846 ºW, 33 m, 14 - Feb- 14, J. N. Zahniser, vacuum, AR 14 - 22 - 2 [MLP]. Paratypes: same data as holotype (3 males, 3 females) [INHS].	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF86FFC0FF06F8AE7D663374.taxon	discussion	Notes. This species resembles A. inesae in external morphology and coloration but differs in having the wings strongly reduced, fully exposing three pregenital abdominal tergites, and in the distinctive male genitalia and female seventh sternite.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF84FFC0FF06FE337D2D34C6.taxon	description	(Figs. 2 S – T, 3 E – H, 6 Q – T, 8 J)	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF84FFC0FF06FE337D2D34C6.taxon	description	Description. Length of male 4.3 – 4.5 mm, female 4.8 – 5.1 mm. Color: face dark brown with pale muscle scars; crown brown with black apical macula and paired stramineous longitudinal stripes extended posterad from midlength across pronotum and mesonotum, with additional pair of alternating light and dark stripes more laterad on pronotum and mesonotum; forewing with black macula on clavus posterad of crossvein, in inner discal cell and middle anteapical cell, with additional less distinct spots in other cells. Crown 1.8 to 2.0 × longer than width between eyes in male and female, respectively. Forewing extended to but not covering pygofer in male and female, outer anteapical cell narrowed but not petiolate apically. Male pygofer slightly more than twice as long as maximum height, apex of lobe tapered posterodorsally, with well-sclerotized process arising preapically and extended anteroventrad with apex tapered and curved slightly laterad, macrosetae scattered throughout length of lobe. Anal tube unmodified. Subgenital plate extended slightly more than half distance to apex of pygofer, lateral margin concave, macrosetae marginal, distributed over entire length of plate. Style with preapical lobe prominent, apex digitiform strongly curved laterad. Connective constricted near midlength, stem weakly developed, posterior margin rounded. Aedeagus with dorsal apodeme relatively short and slender; shaft tubular, moderately slender, curved evenly dorsad, tapered apically in lateral view, with pair of slender dorsally curved processes extended basad along dorsal margin of shaft and pair of shorter acuminate apical processes extended distad and curved slightly laterad in ventral view; gonopore slender, preapical on dorsal surface. Female sternite VII with lateral margins evenly convergent posteriorly, posterior margin shallowly concave with pair of distinct sublateral maculae.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF84FFC0FF06FE337D2D34C6.taxon	materials_examined	Material examined. Holotype female, ARGENTINA: Loreto, Misiones, XII- 1 - 1931, AA Ogloblin / Type No. 51667 U. S. N. M. / Agudus typicus Oman, Det. PW Oman / USNMENT 02110807 [USNM]. Other material: same locality as holotype, XII- 6 - 1931 (1 female) [USNM]; PARAGUAY: 2 km W Eusebio Ayala, 10 June 1975, Blocker, Granovsky & Elzinga / PARATYPE Agudus cyrtobrachium YJ Cheng / USNMENT 02110884 (1 male) [USNM]; ARGENTINA: Corrientes, jct rt 14 & 119. 30 ° 4 ' 35 " S 57 ° 59 ' 57 " W, 5 Jan 2008 C. H. Dietrich, vacuum, AR 3 - 1 (1 male, 2 females); ARGENTINA: Corrientes, 1 km S Pte. Internacional, Santo Tome, 5 Jan 2008, C. H. Dietrich, vacuum, AR 4 - 1 (2 males) [MLP, INHS].	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF84FFC0FF06FE337D2D34C6.taxon	discussion	Notes. This species was originally described based only on females from Misiones Province, Argentina. Comparison of the holotype female to a series of recently collected males and females from northern Argentina (Corrientes) revealed that A. cyrtobrachium Cheng is a junior synonym of A. typicus. Males matching Cheng’s (1980) description and illustrations of A. cyrtobrachium were collected in association with females having the distinctive shape and markings of the pregenital abdominal segment identical to that of the female holotype of A. typicus in the U. S. National Museum. Agudus cyrtobrachium was described based on three male specimens from Paraguay.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF84FFDEFF06F8A2796F32C4.taxon	description	(Figs. 2 U – V, 6 U – X, 8 K) Description. Body length: male 4.7 mm, female 4.7 – 5.0 mm. Color: face medium brown with pale muscle scars, crown tan with pair of bright red submedial stripes extended from apex posterad, continuing over pronotum onto mesonotum, thin brown median line corresponding to coronal suture extended as thin reddish line across pronotum and mesonotum, pronotum with sublateral pair of less conspicuous red stripes; forewing with dark brown spot in clavus posterad of crossvein, in inner discal cell and in inner anteapical cell, spots extended into adjacent cells in some specimens. Male crown 1.3 × longer than width between eyes, female crown 1.5 × longer than width between eyes. Forewing short, exposing at least two pregenital abdominal tergites in both male and female; outer anteapical cell truncate apically, apical cells very short. Male pygofer 2 × longer than basal height in lateral view, without basal apodemes, macrosetae scattered over entire surface except dorsally near base, apex broadly rounded, ventral margin with broad rounded lobe near base, straight in distal two thirds, inner surface without sclerotized bars or lobes. Anal tube short, cylindrical, well sclerotized laterally and dorsally. Subgenital plate half as long as pygofer, narrowly triangular with lateral margin concave, macrosetal row near lateral margin. Connective 2.5 × longer than wide in ventral view, anterior arms evenly divergent, then abruptly convergent and fused near apex; stem broad and very short, truncate posteriorly. Style with preapical lobe short and broadly rounded, apophysis narrow and flattened, curved laterad, apex blunt. Aedeagus with dorsal apodeme in lateral view slender and moderately long, curved anterad, triangular in posterior view; shaft tubular, extended posteroventrad then curved posterodorsad, abruptly broadened with arcuate transverse carina on dorsal surface preapically, distal part tapered in ventral and lateral views with pair of slender spines extended distad from acute apex; gonopore preapical on dorsal surface. Female sternite VII posterior margin with pair of rounded lateral lobes flanking broad, shallow concave medial emargination.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF84FFDEFF06F8A2796F32C4.taxon	etymology	Etymology. The species name is a Latin noun meaning “ hook bearer ”, referring to the hooklike aedeagus.	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
03FA87E4FF84FFDEFF06F8A2796F32C4.taxon	materials_examined	Material examined. Holotype male, PARAGUAY: Missiones, 191 km N. Encenada, 174 km S. Anuncion, 31 May 1975 [INHS]. Paratypes: same data as holotype (1 male, 5 females) [INHS]. Other material: PARAGUAY: NW Asuncion, 2 km E Limpio, at light, 17 June 1975 Elzinga, Granovsky & Blocker (2 females) [INHS]; ARGENTINA: Entre Rios, P. N. El Palmar, Mirador Ao. El Palmar, 31.89149 ºS, 58.24071 ºW, 21 m, 15 - Feb- 14, J. N. Zahniser, vacuum, AR 14 - 27 - 1 (3 males, 2 females) [MLP, INHS]; ARGENTINA: Entre Rios, P. N. El Palmar, Mirador La Glorieta, 31.88713 ºS, 58.27187 ºW, 14 m, 15 - Feb- 14, J. N. Zahniser, vacuum, AR 14 - 24 - 1 (1 male) [INHS].	en	Dietrich, Christopher H., Zahniser, James N. (2026): Review of the endemic South American grassland leafhopper genus Agudus Oman (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini). Zootaxa 5845 (3): 545-562, DOI: 10.11646/zootaxa.5845.3.6, URL: https://doi.org/10.11646/zootaxa.5845.3.6
