taxonID	type	description	language	source
03FD8327FF86FFE5FEF5637B1340F855.taxon	type_taxon	TYPE SPECIES. — Macrourites modestiformis Schlotheim, 1822, by subsequent designation by Glaessner (1929: 150).	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF86FFE5FEF5637B1340F855.taxon	discussion	PRELIMINARY REMARKS Eryma was previously assigned to the infraorder Glypheidea Winkler, 1881 (Karasawa et al. 2013; Feldmann et al. 2015). However, the phylogenetic study provided by Charbonnier et al. (2015) on the glypheid lobsters resulted in the exclusion of Eryma from this infraorder. The following phylogenetic study by Devillez et al. (2019), focused on erymid lobsters, concurs with the result of Charbonnier et al. (2015), resulting in the assignation of Eryma into the infraorder Astacidea Latreille, 1802. Despite these studies, the recent additions to the Treatise online (Schweitzer & Feldmann 2024: 11 - 12) maintain the erymid lobsters (including Eryma) within Glypheidea. Considering that erymid lobsters, and so Eryma, are true chelate lobsters (glypheid pereiopod terminations are subchelate), we follow herein the systematic framework proposed in Devillez et al. (2019): Eryma is assigned to Astacidea, like other lobsters bearing true chelae.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF87FFE4FEB365F71645F852.taxon	description	(Fig. 2)	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF87FFE4FEB365F71645F852.taxon	materials_examined	TYPE MATERIAL. — Holotype. France • 1 specimen (carapace; adult); Rhône-Alpes Auvergne, Hautes-Alpes, Rosans; Early Cretaceous, Albian; Arnaud Clément leg.; MNHN. F. A 57457. Paratype. France • 1 specimen (P 1 chela; adult); Rhône-Alpes Auvergne, Hautes-Alpes, Rosans; Early Cretaceous, Albian; Arnaud Clément leg.; MNHN. F. A 57458. EXAMINED MATERIAL. — France • 3 specimens (carapaces; adults); Grand-Est, Haute-Marne, Lac du Der, Presqu’Île de Nemours; Early Cretaceous, middle Albian, Hoplites dentatus zone; Richard Brochet leg.; MNHN. F. A 98126, A 98127, A 98128.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF87FFE4FEB365F71645F852.taxon	discussion	DISCUSSION The carapace groove pattern of the three specimens is typical of erymid lobsters: well-developed cervical, postcervical and branchiocardiac grooves, short gastro orbital groove and concavo-convex hepatic groove. Moreover, the junction between postcervical and branchiocardiac grooves at carapace midheight is diagnostic of Eryma. The assignation to Eryma vocontii is supported by: 1) the elongated “ V ” formed by the convergent postcervical and branchiocardiac grooves; 2) the very short ventral extension of the postcervical groove; 3) the ornamentation made of tubercles preceded by depressions; 4) with depressions deeper in branchial region; 5) the strongly inflated attachment sites of both mandibular and adductor testis muscles; and 6) the elongated, sub-rectangular outline of the attachment site of adductor testis muscles which is dorsally delimited by a narrow groove running between hepatic and branchiocardiac grooves (this last feature is typical of this species). Early Cretaceous fossils attributed to Eryma are very uncommon and only four species are known to date (Karasawa et al. 2008; Devillez et al. 2016, 2021; Devillez & Charbonnier 2022; Ossó et al. 2024). E. vocontii was described based on only two specimens, a crushed carapace and a first pereiopod chela, both from the type locality (Fig. 2 A-D). The three new specimens found about 360 km northward suggest a geographic distribution significantly broader than the French South-East Basin where the type locality is located (Devillez et al. 2016: fig. 2). These specimens are also more complete and less deformed than the holotype, with their attachment site of mandibular muscle and pterygostomial region entirely preserved. In addition, the specimen MNHN. F. A 98128 (Fig. 2 G) shows the basis of a strong antennal spine. This specimen also carries a fragment of an epibiontic bivalve shell fixed on its right flank, above the junction between postcervical and branchiocardiac grooves. Such an association is reported for the first time in Early Cretaceous erymid lobsters, but is well known, for instance, in mecochirid lobsters from the Aptian of the Isle of Wight, United Kingdom (see Robin et al. 2016).	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF87FFE4FC75649813F0FC14.taxon	type_taxon	TYPE SPECIES. — Rathbunopon polyakron Stenzel, 1945, by original designation.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF87FFE8FC1C66DA1640FB90.taxon	description	(Fig. 3) urn: lsid: zoobank. org: act: E 082 AB 6 A- 0 C 52 - 40 AF- 8 D 0 C- 27 DEEFCE 02 AF	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF87FFE8FC1C66DA1640FB90.taxon	diagnosis	DIAGNOSIS. — Carapace, subovoid almost pyriform in outline, longer than wide with maximum width in posterior third of carapace at level of metabranchial region; dorsal surface strongly sculptured, regions well defined by strong inflated tubercles and delimited by deep grooves; rostrum subtriangular, spatulate, with rounded tip; orbits large, deep, and complete; supraorbital margin with two tubercles, one bigger proximally and one smaller distally; infraorbital spine strong, dorsally visible; hook-shaped outer-orbital spine directed forward; anterolateral margins short, converging anteriorly; posterolateral margins strongly convex, converging posteriorly; posterior margin straight and rimmed; epi- and protogastric regions with one tubercle respectively; mesogastric region subtriangular elongate, with three tubercles; urogastric region with two parallel narrow transverse lobes; cardiac region inverted triangular, swollen; branchial regions separated by well-marked grooves; epibranchial region with one tubercle; meso- and metabranchial regions weakly inflated finely tuberculate; intestinal region narrow and weakly depressed; well-marked cervical, branchial, and branchiocardiac grooves.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF87FFE8FC1C66DA1640FB90.taxon	etymology	ETYMOLOGY. — The specific epithet honours Patrick Brisac, collector of fossils and friend of one of the authors (RB).	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF87FFE8FC1C66DA1640FB90.taxon	materials_examined	TYPE MATERIAL. — Holotype. France • 1 specimen (carapace; adult); Grand-Est, Haute-Marne, Éclaron-Braucourt-Sainte Livière, Lac du Der, Presqu’Île de Nemours; Early Cretaceous, middle Albian, Hoplites dentatus ammonite biozone; Richard Brochet leg.; MNHN. F. A 98122. Paratype. France • 1 specimen (carapace; adult); Grand-Est, Haute-Marne, Éclaron-Braucourt-Sainte Livière, Lac du Der, Presqu’Île de Nemours; Early Cretaceous, middle Albian, Hoplites dentatus zone; Richard Brochet leg.; MNHN. F. A 98121. TYPE LOCALITY. — Éclaron-Braucourt-Sainte-Livière, Presqu’Île de Nemours, Lac du Der (Haute-Marne, Grand-Est, France). TYPE AGE. — Early Cretaceous (middle Albian), Hoplites dentatus ammonite zone. ADDITIONAL EXAMINED MATERIAL. — Rathbunopon tuberculatum (Van Straelen, 1936): Holotype. France • 1 specimen (carapace; adult); Bourgogne Franche-Comté, Yonne, Auxerre; Early Cretaceous, Hauterivian; MHNAux. 2012.0 - 11.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF87FFE8FC1C66DA1640FB90.taxon	description	DESCRIPTION Carapace, subovoid almost pyriform in outline, longer than wide with maximum width in posterior third of carapace at level of metabranchial region (holotype: carapace length: 16 mm; width: 12 mm; paratype: carapace length without rostrum: 13 mm; width: 12 mm); dorsal surface strongly sculptured, regions well defined by strong inflated tubercles and delimited by deep grooves; dorsal surface covered with small tubercles uniformly arranged; anterolateral margins short, converging anteriorly, medially divided by cervical groove and limited by a rounded epibranchial spine directed outward; hook-shaped outer-orbital spine directed forward; posterolateral margins strongly convex, converging posteriorly twice long anterolateral margins; posterior margin straight and rimmed; front with lateral folds forming inner-orbital spines; rostrum subtriangular, spatulate, with rounded tip, downturned, strongly sulcate axially, narrowing distally; lateral margins of axial groove, reinforced with a line of aligned small tubercles; orbits large, deep, and complete; supraorbital margin with two rounded and upwardly directed tubercles, bigger proximally and smaller distally; strong hook-shaped outer-orbital spine, directed forward, closing laterally the orbit; infraorbital margin with a strong subtriangular tooth, dorsally visible; hepatic regions small, defined by a transverse inflation, contiguous to the rearmost protogastric tubercle and ending below the outer-orbital spine; epigastric region with a pair of strong tubercles located at level of rostral base where the axial depression begins; protogastric regions, with a strong posterior tubercle close to median mesogastric tubercle; mesogastric region subtriangular elongate anteriorly, longer than wide, with three tubercles, two transversely contiguous at base and one bigger in the anterior part of the region; metagastric region not distinguishable from mesogastric region; urogastric region defined by two narrow transverse parallel lobes, bounded by grooves; first urogastric lobe, strongly inflated, connecting with epibranchial lobe laterally; second urogastric lobe, strongly flat, limited by branchiocardiac grooves laterally and separated by a depression from cardiac region posteriorly; cardiac region inverted triangular elongate, weakly inflated and well bounded laterally by deep branchiocardiac groove; epibranchial regions defined by a transverse lobe with an inner single tubercle; meso- and metabranchial regions weakly inflated covered with small tubercles uniformly arranged; intestinal region narrow and flat; cervical, postcervical, branchial, and branchiocardiac grooves well marked, deep, smooth; ventral carapace and appendages not preserved.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF87FFE8FC1C66DA1640FB90.taxon	discussion	DISCUSSION According toKlompmaker et al. (2011) andOssó et al. (2018), the studied specimens can be assigned to Rathbunopon Stenzel, 1945 based on the following morphological characters: the subtriangular rostrum downturned, axially sulcate, the supraorbital margin with two tubercles, the hook-shaped outer-orbital spine directed forward, the mesogastric region with three tubercles, the epibranchial region with one tubercle, and the subtriangular cardiac region. According toOssó et al. (2018), Rathbunopon includes eight species (see included species paragraph), of which six are dated to the Early Cretaceous (Hauterivian-Albian). Klompmaker et al. (2011) revised Rathbunopon obesum based on additional specimens from Koskobilo and Monte Orobe (northern Spain) since Van Straelen’s original specimen was not encountered in the collection of Museo Geológico del Seminario de Barcelona nor in the collection of the Institut Royal des Sciences naturelles de Belgique in Brussels. The subtriangular, spatulate rostrum with rimmed axial depression of R. brisaci n. sp. (vs triangular rostrum with smooth axial depression in R. obesum), the strongly tuberculated dorsal surface (vs finely pustulate in R. obesum), and the strongly convex posterolateral margins (vs nearly straight posterolateral margins in R. obesum), and the mesogastric region subtriangular elongate anteriorly, longer than wide, with three tubercles, two transversely contiguous at base and one bigger in the anterior part of the region (vs mesogastric region divided in three raised parts in R. obesum) rule out the belonging of the studied specimens to the species from Navarra. The poor state of preservation of the holotype of R. schrattenkalkensis, above all in the frontal region, does not allow a detailed comparison with the present specimens. However, based on the description provided byKlompmaker et al. (2011), the studied specimens differ from this species by the strongly tuberculated dorsal ornamentation (vs indented pustules in R. schrattenkalkensis) and by the mesogastric region which is subtriangular, elongate anteriorly, longer than wide, and with three tubercles (vs mesogastric region with two raised areas posteriorly, and one anteriorly instead of distinct tubercles in R. schrattenkalkensis). The epigastric regions with one pair of strong tubercles and the epibranchial regions with one tubercle (vs epigastric regions with two pairs of tubercles and epibranchial regions with two tubercles in R. tarraconensis), the urogastric region with first lobe, strongly inflated, and second one, strongly flat (vs urogastric region with two short, narrow lobes similar to each other in R. tarraconensis), and dorsal regions with strong inflated tubercles uniformly arranged (vs strong tubercles limited to the branchial regions in R. tarraconensis) rules out the belonging of the studied specimens to R. tarraconensis. Rathbunopon viai exhibits sinuous supraorbital margins, finely granulated, and bi-fissured, protogastric regions with two tubercles, and epibranchial regions without tubercles. This combination of characters rules out the belonging of the studied specimens to this species. Rathbunopon tuberculatum is the oldest representative of the genus known to date. The holotype preserves only the anterior portion of the carapace, making difficult a detailed comparison with the studied specimens. However, the smooth dorsal surface and the protogastric and epibranchial regions with two tubercles are clearly different from the studied specimens. Three species are restricted to the Cenomanian (Late Cretaceous; see above-reported list). The smooth cardiac region (vs cardiac region with a pair of transverse median pits in R. polyakron) and the mesobranchial region with small tubercles uniformly arranged (vs mesobranchial region with one strong tubercle in R. polyakron) exclude the belonging of the studied specimens to the type species. The dorsal regions covered with tubercles uniformly arranged (vs tubercles limited to the branchial regions in R. oblitum) excludes the belonging of the studied specimens to R. oblitum. Finally, the dorsal regions with tubercles (vs smooth dorsal regions in R. woodsi) rules out the belonging of the studied specimens to R. woodsi. In conclusion, based on these observations, we can attest that the subtriangular spatulate rostrum with rimmed axial depression, the epibranchial regions with an inner tubercle, and the dorsal surface strongly tuberculate are morphological characters justifying the description of R. brisaci n. sp. to accommodate the studied specimens. The recovery of R. brisaci n. sp. further confirms the stratigraphic range of Rathbunopon limited to the Early Cretaceous – early Late Cretaceous.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF8BFFE8FF19607C163CF962.taxon	type_taxon	TYPE SPECIES. — Paranecrocarcinus hexagonalis Van Straelen, 1936, by monotypy.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF8BFFEBFED2623D1233FA6E.taxon	description	(Fig. 4) urn: lsid: zoobank. org: act: 70529 EDC-FE 01 - 45 A 7 - 9 EE 8 - 0 EDFCD 450 FC 6	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF8BFFEBFED2623D1233FA6E.taxon	diagnosis	DIAGNOSIS. — Carapace subhexagonal, moderately arched in both directions; spatulate-shaped rostrum, depressed axially, protruded beyond orbits, and strongly downturned; rostral lateral margins with one median strong swelling; supraorbital margin with two narrow fissures; anterolateral margins slightly convex, armed with small spines (4 - 5 as preserved); epibranchial angle strongly upturned with one well-developed rounded swelling; epigastric region flat with two post-frontal slits; protogastric regions with a pair of strong swellings and one contiguous smaller swelling, aligned transversely; cardiac region diamond shaped, slightly inflated, without tubercles; epibranchial regions with one inner small swelling close to branchiocardiac groove; mesobranchial regions with one marginal small swelling; cervical groove deep with gastric pits on either side of narrow medial gap; dorsal carapace surface from cervical groove to posterior margin covered with small granules uniformly arranged, gastric and hepatic regions scarcely granulate; granules more condensed on rostral, protogastric, epibranchial, and mesobranchial swellings.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF8BFFEBFED2623D1233FA6E.taxon	etymology	ETYMOLOGY. — The specific epithet honours Mikael Perchat, president of the amateur palaeontological and geological club CMPG (Club Marnais en Paléontologie et Géologie).	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF8BFFEBFED2623D1233FA6E.taxon	materials_examined	TYPE MATERIAL. — Holotype. France • 1 specimen (carapace; adult); Grand-Est, Haute-Marne, Éclaron-Braucourt-Sainte Livière, Lac du Der, Presqu’Île de Nemours; Early Cretaceous, middle Albian, Hoplites dentatus ammonite biozone; Richard Brochet leg.; MNHN. F. A 98123. TYPE LOCALITY. — Éclaron-Braucourt-Sainte Livière, Presqu’Île de Nemours, Lac du Der (Haute-Marne, Grand-Est, France). TYPE AGE. — Early Cretaceous (middle Albian), Hoplites dentatus ammonite zone.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF8BFFEBFED2623D1233FA6E.taxon	description	DESCRIPTION Carapace subhexagonal (length: c. 13 mm; width: c. 14 mm), moderately arched in both directions; front produced as spatulate-shaped rostrum, depressed axially, protruded beyond orbits, and strongly downturned; rostral lateral margins with one median strong swelling; orbits close together directed forwards; supraorbital margin with two narrow fissures; anterolateral margins slightly convex, armed with small spines (4 - 5 as preserved); posterolateral margins straight, backwards converging, defined by a fine rim of granules; epibranchial angle strongly upturned with a well-developed rounded swelling; posterior margin strongly concave, rimmed; regions moderately defined, generally swollen; hepatic regions small, flat; epigastric region flat with two post-frontal slits; protogastric regions with a pair of strong swellings and one contiguous smaller swelling, aligned transversely; meso- and metagastric regions indistinct; urogastric region slightly inflated; cardiac region diamond shaped, slightly inflated, without tubercles; intestinal region flat; epibranchial regions with one inner small swelling close to branchiocardiac groove; meso- and metabranchial regions fused, smooth; mesobranchial regions with one marginal small swelling; cervical groove deep with gastric pits on either side of narrow medial gap; branchiocardiac grooves deep; marked cardiac region laterally; dorsal carapace surface from cervical groove to posterior margin covered with small granules uniformly arranged, gastric and hepatic regions scarcely granulated; granules more condensed on rostral, protogastric, epibranchial, and mesobranchial swellings.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF8BFFEBFED2623D1233FA6E.taxon	discussion	DISCUSSION Based on Van Bakel et al. (2012) and Schweitzer et al. (2018), some characters support the assignment of the studied specimen to Paranecrocarcinus, such as the carapace with protogastric and branchial swellings, orbits directed anteriorly, rostrum axially sulcate, and epigastric regions with a pair of elongate slits. Ossó et al. (2024) provided an updated list of the species of Paranecrocarcinus. Among these, only three species are known to date from the Early – Late Cretaceous of Europe: P. hexagonalis (Hauterivian, France), P. xivertensis (Aptian, Spain), and P. balla (Cenomanian, France). As reported by Van Straelen (1936), the urogastric region with one median tubercle and the cardiac region with a pair of medial tubercles exclude the specimen from belonging to P. hexagonalis. The cardiac region with three tubercles (vs smooth cardiac region in the studied specimen) and the mesobranchial regions smooth (vs mesobranchial regions with one marginal small swelling in the studied specimen) exclude the belonging of the studied specimen to P. xivertensis. We exclude the belonging of the studied specimen to P. balla for the different dorsal carapace ornamentation finely pitted between small and large tubercles in P. balla (vs small granules uniformly arranged posteriorly, gastric and hepatic regions scarcely granulated in the studied specimen). The same observations on the dorsal carapace (location and number of tubercles and ornamentation with pits or granules) can be made for P. mozambiquensis and P. libanoticus. The spatulate-shaped rostrum strongly downturned, the rostral lateral margins with one median inflated swelling, the epibranchial angle strongly upturned, the cardiac region without swellings, the mesobranchial regions with one marginal swelling, the dorsal carapace surface covered posteriorly with small granules uniformly arranged, the gastric and hepatic regions scarcely granulated, and granules more thickened on rostral, protogastric epibranchial, and mesobranchial swellings are enough characters to justify the description of P. perchati n. sp. for the studied specimen. This is the second report of Paranecrocarcinus from the Early Cretaceous of France.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF88FFEAFBD6635A1672F855.taxon	type_taxon	TYPE SPECIES. — Corystes broderipii Mantell, 1844, original designation by Van Bakel et al. (2012).	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF89FFEDFC2D637D131CFCCE.taxon	description	(Fig. 5) urn: lsid: zoobank. org: act: BD 1690 E 5 - B 429 - 494 D- 96 CF- 80 CD 344 D 02 A 6	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF89FFEDFC2D637D131CFCCE.taxon	diagnosis	DIAGNOSIS. — Small carapace, longitudinally ovate, strongly convex transversely, flat longitudinally, longer than wide; fronto-orbital margin broad; orbits wide, deep; supraorbital margin with two narrow fissures, outer-orbital spines conical, acute, forwardly directed; anterolateral margins short with two spines; posterolateral margins longer, straight, rimmed; posterior margin very short, weakly concave; hepatic regions with a pair of small suborbital protuberances; cervical groove broadly U-shaped, deeper medially, almost indistinct laterally; gastric pits close together; branchiocardiac grooves deep; axial carina weakly marked for the entire length of the carapace; dorsal surface covered with small granules uniformly arranged.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF89FFEDFC2D637D131CFCCE.taxon	etymology	ETYMOLOGY. — The specific epithet honours Martial Scheitz, vice-president of the amateur palaeontological and geological club CMPG (Club Marnais en Paléontologie et Géologie).	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF89FFEDFC2D637D131CFCCE.taxon	materials_examined	TYPE MATERIAL. — Holotype. France • 1 specimen (carapace; adult); Grand-Est, Haute-Marne, Éclaron-Braucourt-Sainte Livière, Lac du Der, Presqu’Île de Nemours; Early Cretaceous, middle Albian, Hoplites dentatus ammonite biozone; Richard Brochet leg.; MNHN. F. A 98124. Paratype. France • 1 specimen (carapace; adult); Grand-Est, Haute-Marne, Éclaron-Braucourt-Sainte Livière, Lac du Der, Presqu’Île de Nemours; Early Cretaceous, middle Albian, Hoplites dentatus ammonite biozone; Richard Brochet leg.; MNHN. F. A 98125. TYPE LOCALITY. — Éclaron-Braucourt-Sainte Livière, Presqu’Île de Nemours, Lac du Der (Haute-Marne, Grand-Est, France). TYPE AGE. — Early Cretaceous (middle Albian), Hoplites dentatus ammonite zone.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF89FFEDFC2D637D131CFCCE.taxon	description	DESCRIPTION Subhexagonal carapace, longitudinally ovate, strongly convex transversely, flat longitudinally, longer than wide (holotype, length: 15 mm, width: 12 mm; paratype, length: 13 mm, width: 9 mm); fronto-orbital margin broad; rostrum no preserved; orbits broad and deep; supraorbital margins with two narrow fissures; outer-orbital spines conical, acute, forwardly directed; anterolateral margin short, slightly convex, with two spines, one at hepatic level and one at epibranchial level; posterolateral margins longer, straight, rimmed and narrowing posteriorly; posterior margin very short, weakly concave; dorsal regions indistinct; hepatic regions with a pair of small protuberances aligned transversely; cardiac region wide, well defined by deep branchiocardiac groove laterally; cervical groove, deeper medially and broadly U-shaped, laterally less marked; gastric pits distinct, close together; axial carina weakly marked; dorsal carapace surface covered with small granules uniformly arranged.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
03FD8327FF89FFEDFC2D637D131CFCCE.taxon	discussion	DISCUSSION Based on Van Bakel et al. (2012) the assignment of the studied specimens to Joeranina rather than other palaeocorystid genera is supported by the following morphological characters: the presence of hepatic suborbital protuberances, a complete cervical groove, and a cuticle microstructure with granules. Ossó et al. (2024) provided an updated list of the species of Joeranina. Among these, only four species are known to date from the Early-Late Cretaceous of Europe: J. broderipii (middle-late Albian-? Cenomanian, United Kingdom, France, Switzerland), J. gaspari (Albian, Spain), J. tausi (Albian, Spain), and J. houssineaui (Cenomanian, France) to which the studied specimens are compared. The well-developed axial carina, the anterolateral margins with one spine, and the cervical groove well marked medially and laterally rule out the belonging of the studied specimens to J. broderipii (see Charbonnier & Garassino 2022). The anterolateral margins with three spines exclude the belonging of the studied specimens to J. gaspari and J. tausi (Ossó et al. 2024). Although J. houssineaui has the anterolateral margins with two spines, we exclude the belonging of the studied specimens to this species which has gastric pits well separated (vs close together in the studied specimens), hepatic region with one protuberance (vs a pair of protuberances in the studied specimens), and dorsal carapace surface with granules most prominent on axial carina, postfrontal area, anterolateral area, and posterolateral margins (vs small granules uniformly arranged in the studied specimens) (Van Bakel 2013). In conclusion, based on the above comparisons restricted to the European species, some characters (anterolateral margins short with two spines, cervical groove broadly U-shaped, deeper medially, almost indistinct laterally, hepatic regions with a pair of small protuberances, gastric pits close together, and dorsal surface covered with small granules uniformly arranged) justify the description of J. scheitzi n. sp. to accommodate the studied specimens. It represents the second species of Joeranina in the Early Cretaceous of France.	en	Charbonnier, Sylvain, Garassino, Alessandro, Devillez, Julien, Brochet, Richard (2026): New record of crabs and lobsters from the Early Cretaceous (Albian) of Haute-Marne, France. Geodiversitas 48 (8): 145-157, DOI: 10.5252/geodiversitas2026v48a8
