taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03FF87FAFFEC870CFC392387D036759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965170/files/figure.png	https://doi.org/10.5281/zenodo.22965170	FIG. 1. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, outer view of chelae.	FIG. 1. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, outer view of chelae.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFEC870CFC392387D036759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965174/files/figure.png	https://doi.org/10.5281/zenodo.22965174	FIG. 2. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, dorsal view of left cheliped; B, dorsal view of right cheliped; C, right fourth ambulatory leg; D, right third maxilliped; E, sternopleonal cavity;F, anterior thoracic sternum, telson and pleonal somites 5 and 6; G, tho- racic sternum, telson and pleonal somites 2-6; H, posterior thoracic sternum and pleonal somites 1-6. A and B photographed to same scale.	FIG. 2. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, dorsal view of left cheliped; B, dorsal view of right cheliped; C, right fourth ambulatory leg; D, right third maxilliped; E, sternopleonal cavity;F, anterior thoracic sternum, telson and pleonal somites 5 and 6; G, tho- racic sternum, telson and pleonal somites 2-6; H, posterior thoracic sternum and pleonal somites 1-6. A and B photographed to same scale.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFEC870CFC392387D036759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965185/files/figure.png	https://doi.org/10.5281/zenodo.22965185	FIG. 6. — Balssiathelphusa sucki (Balss, 1937) (A-F), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”, and B. cursor Ng, 1986 (G-M), holotype, ♂ 15.4 × 13.3 mm (MZB Cru Nr. 1279a), Kalimantan. A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, distal part of right G1 (dorso-mesial view, setae not drawn); F, right G2; G, left third maxilliped; H, frontal median triangle; I, left G1 (ventral view); J, left G1 (dorsal view); K, distal part of left G1 (ventral view); L, distal part of left G1 (dorsal view); M, left G2. Scale bars: A, B, F, 0.5 mm; C, D, 0.2 mm. I-M, after Ng (1986a: fig. 2A-F).	FIG. 6. — Balssiathelphusa sucki (Balss, 1937) (A-F), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”, and B. cursor Ng, 1986 (G-M), holotype, ♂ 15.4 × 13.3 mm (MZB Cru Nr. 1279a), Kalimantan. A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, distal part of right G1 (dorso-mesial view, setae not drawn); F, right G2; G, left third maxilliped; H, frontal median triangle; I, left G1 (ventral view); J, left G1 (dorsal view); K, distal part of left G1 (ventral view); L, distal part of left G1 (dorsal view); M, left G2. Scale bars: A, B, F, 0.5 mm; C, D, 0.2 mm. I-M, after Ng (1986a: fig. 2A-F).	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFEC870CFC392387D036759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965212/files/figure.png	https://doi.org/10.5281/zenodo.22965212	FIG. 16. — Thoracic sternums of: A, Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”; B, B. cursor Ng, 1986, paratype,♂ 15.6 × 13.4 mm (ZRC 1989.3036), Kalimantan; C, Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; D, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan. Abbreviations: bp, press button tubercle of pleonal locking mechanism; br5/6, median bridge between thoracic sternites 5 and 6; br6/7, median bridge between thoracic sternites 6 and 7; st4-8, thoracic sternites 4-8, respectively.	FIG. 16. — Thoracic sternums of: A, Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”; B, B. cursor Ng, 1986, paratype,♂ 15.6 × 13.4 mm (ZRC 1989.3036), Kalimantan; C, Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; D, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan. Abbreviations: bp, press button tubercle of pleonal locking mechanism; br5/6, median bridge between thoracic sternites 5 and 6; br6/7, median bridge between thoracic sternites 6 and 7; st4-8, thoracic sternites 4-8, respectively.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFE18709FC2C2581D70276FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965177/files/figure.png	https://doi.org/10.5281/zenodo.22965177	FIG. 3. — Balssiathelphusa cursor Ng, 1986, paratype, ♂ 15.6 × 13. 4 mm (ZRC 1989.3036), Kalimantan: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, outer view of chelae.	FIG. 3. — Balssiathelphusa cursor Ng, 1986, paratype, ♂ 15.6 × 13. 4 mm (ZRC 1989.3036), Kalimantan: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, outer view of chelae.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFE18709FC2C2581D70276FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965180/files/figure.png	https://doi.org/10.5281/zenodo.22965180	FIG. 4. — Balssiathelphusa cursor Ng, 1986, paratype, ♂ 15.6 × 13.4 mm (ZRC 1989.3036), Kalimantan: A, right fourth ambulatory leg; B, right third maxilliped; C, anterior thoracic sternum, telson and pleonal somites 5 and 6; D, thoracic sternum, telson and pleonal somites 2-6; E, posterior thoracic sternum and pleonal somites 1-4; F, sternopleonal cavity.	FIG. 4. — Balssiathelphusa cursor Ng, 1986, paratype, ♂ 15.6 × 13.4 mm (ZRC 1989.3036), Kalimantan: A, right fourth ambulatory leg; B, right third maxilliped; C, anterior thoracic sternum, telson and pleonal somites 5 and 6; D, thoracic sternum, telson and pleonal somites 2-6; E, posterior thoracic sternum and pleonal somites 1-4; F, sternopleonal cavity.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFE18709FC2C2581D70276FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965183/files/figure.png	https://doi.org/10.5281/zenodo.22965183	FIG. 5. — Balssiathelphusa cursor Ng, 1986, paratype, ♀ 16.4 × 13.0 mm (ZRC 1989.3037), Kalimantan: A, overall dorsal habitus; B, female anterior thoracic sternum and pleon; C, sternopleonal cavity showing vulvae.	FIG. 5. — Balssiathelphusa cursor Ng, 1986, paratype, ♀ 16.4 × 13.0 mm (ZRC 1989.3037), Kalimantan: A, overall dorsal habitus; B, female anterior thoracic sternum and pleon; C, sternopleonal cavity showing vulvae.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFE18709FC2C2581D70276FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965185/files/figure.png	https://doi.org/10.5281/zenodo.22965185	FIG. 6. — Balssiathelphusa sucki (Balss, 1937) (A-F), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”, and B. cursor Ng, 1986 (G-M), holotype, ♂ 15.4 × 13.3 mm (MZB Cru Nr. 1279a), Kalimantan. A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, distal part of right G1 (dorso-mesial view, setae not drawn); F, right G2; G, left third maxilliped; H, frontal median triangle; I, left G1 (ventral view); J, left G1 (dorsal view); K, distal part of left G1 (ventral view); L, distal part of left G1 (dorsal view); M, left G2. Scale bars: A, B, F, 0.5 mm; C, D, 0.2 mm. I-M, after Ng (1986a: fig. 2A-F).	FIG. 6. — Balssiathelphusa sucki (Balss, 1937) (A-F), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”, and B. cursor Ng, 1986 (G-M), holotype, ♂ 15.4 × 13.3 mm (MZB Cru Nr. 1279a), Kalimantan. A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, distal part of right G1 (dorso-mesial view, setae not drawn); F, right G2; G, left third maxilliped; H, frontal median triangle; I, left G1 (ventral view); J, left G1 (dorsal view); K, distal part of left G1 (ventral view); L, distal part of left G1 (dorsal view); M, left G2. Scale bars: A, B, F, 0.5 mm; C, D, 0.2 mm. I-M, after Ng (1986a: fig. 2A-F).	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFE18709FC2C2581D70276FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965174/files/figure.png	https://doi.org/10.5281/zenodo.22965174	FIG. 2. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, dorsal view of left cheliped; B, dorsal view of right cheliped; C, right fourth ambulatory leg; D, right third maxilliped; E, sternopleonal cavity;F, anterior thoracic sternum, telson and pleonal somites 5 and 6; G, tho- racic sternum, telson and pleonal somites 2-6; H, posterior thoracic sternum and pleonal somites 1-6. A and B photographed to same scale.	FIG. 2. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, dorsal view of left cheliped; B, dorsal view of right cheliped; C, right fourth ambulatory leg; D, right third maxilliped; E, sternopleonal cavity;F, anterior thoracic sternum, telson and pleonal somites 5 and 6; G, tho- racic sternum, telson and pleonal somites 2-6; H, posterior thoracic sternum and pleonal somites 1-6. A and B photographed to same scale.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFE18709FC2C2581D70276FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965212/files/figure.png	https://doi.org/10.5281/zenodo.22965212	FIG. 16. — Thoracic sternums of: A, Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”; B, B. cursor Ng, 1986, paratype,♂ 15.6 × 13.4 mm (ZRC 1989.3036), Kalimantan; C, Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; D, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan. Abbreviations: bp, press button tubercle of pleonal locking mechanism; br5/6, median bridge between thoracic sternites 5 and 6; br6/7, median bridge between thoracic sternites 6 and 7; st4-8, thoracic sternites 4-8, respectively.	FIG. 16. — Thoracic sternums of: A, Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”; B, B. cursor Ng, 1986, paratype,♂ 15.6 × 13.4 mm (ZRC 1989.3036), Kalimantan; C, Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; D, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan. Abbreviations: bp, press button tubercle of pleonal locking mechanism; br5/6, median bridge between thoracic sternites 5 and 6; br6/7, median bridge between thoracic sternites 6 and 7; st4-8, thoracic sternites 4-8, respectively.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFE18709FC2C2581D70276FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965170/files/figure.png	https://doi.org/10.5281/zenodo.22965170	FIG. 1. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, outer view of chelae.	FIG. 1. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, outer view of chelae.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFE48714FC292783D38D745D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965187/files/figure.png	https://doi.org/10.5281/zenodo.22965187	FIG. 7. — Balssiathelphusa inopinata n. sp., holotype,♀ 23.2 × 17.6 mm (MRSN 1209), Kalimantan: A, overall dorsal habitus;B, dorsal view of carapace;C, frontal view of cephalothorax; D, E, frontal median triangle, antennules, antennae and epistome (viewed at slightly different angles).	FIG. 7. — Balssiathelphusa inopinata n. sp., holotype,♀ 23.2 × 17.6 mm (MRSN 1209), Kalimantan: A, overall dorsal habitus;B, dorsal view of carapace;C, frontal view of cephalothorax; D, E, frontal median triangle, antennules, antennae and epistome (viewed at slightly different angles).	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFE48714FC292783D38D745D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965189/files/figure.png	https://doi.org/10.5281/zenodo.22965189	FIG. 8. — Balssiathelphusa inopinata n. sp., holotype, ♀ 23.2 × 17.6 mm (MRSN 1209), Kalimantan: A, left third maxilliped; B, right fourth ambulatory leg; C, buccal cavity and outer view of chelae; D, pleonal somites 4-6 and telson; E, sternopleonal cavity showing vulvae.	FIG. 8. — Balssiathelphusa inopinata n. sp., holotype, ♀ 23.2 × 17.6 mm (MRSN 1209), Kalimantan: A, left third maxilliped; B, right fourth ambulatory leg; C, buccal cavity and outer view of chelae; D, pleonal somites 4-6 and telson; E, sternopleonal cavity showing vulvae.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFE48714FC292783D38D745D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965170/files/figure.png	https://doi.org/10.5281/zenodo.22965170	FIG. 1. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, outer view of chelae.	FIG. 1. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, outer view of chelae.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFE48714FC292783D38D745D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965174/files/figure.png	https://doi.org/10.5281/zenodo.22965174	FIG. 2. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, dorsal view of left cheliped; B, dorsal view of right cheliped; C, right fourth ambulatory leg; D, right third maxilliped; E, sternopleonal cavity;F, anterior thoracic sternum, telson and pleonal somites 5 and 6; G, tho- racic sternum, telson and pleonal somites 2-6; H, posterior thoracic sternum and pleonal somites 1-6. A and B photographed to same scale.	FIG. 2. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, dorsal view of left cheliped; B, dorsal view of right cheliped; C, right fourth ambulatory leg; D, right third maxilliped; E, sternopleonal cavity;F, anterior thoracic sternum, telson and pleonal somites 5 and 6; G, tho- racic sternum, telson and pleonal somites 2-6; H, posterior thoracic sternum and pleonal somites 1-6. A and B photographed to same scale.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFE48714FC292783D38D745D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965177/files/figure.png	https://doi.org/10.5281/zenodo.22965177	FIG. 3. — Balssiathelphusa cursor Ng, 1986, paratype, ♂ 15.6 × 13. 4 mm (ZRC 1989.3036), Kalimantan: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, outer view of chelae.	FIG. 3. — Balssiathelphusa cursor Ng, 1986, paratype, ♂ 15.6 × 13. 4 mm (ZRC 1989.3036), Kalimantan: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, outer view of chelae.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFE48714FC292783D38D745D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965183/files/figure.png	https://doi.org/10.5281/zenodo.22965183	FIG. 5. — Balssiathelphusa cursor Ng, 1986, paratype, ♀ 16.4 × 13.0 mm (ZRC 1989.3037), Kalimantan: A, overall dorsal habitus; B, female anterior thoracic sternum and pleon; C, sternopleonal cavity showing vulvae.	FIG. 5. — Balssiathelphusa cursor Ng, 1986, paratype, ♀ 16.4 × 13.0 mm (ZRC 1989.3037), Kalimantan: A, overall dorsal habitus; B, female anterior thoracic sternum and pleon; C, sternopleonal cavity showing vulvae.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFF98714FED12522D622759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965192/files/figure.png	https://doi.org/10.5281/zenodo.22965192	FIG. 9. — Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, buccal cavity, anterior thoracic sternum, telson and pleonal somites 5 and 6; F, posterior thoracic sternum and pleonal somites 1-4; G, sternopleonal cavity.	FIG. 9. — Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, buccal cavity, anterior thoracic sternum, telson and pleonal somites 5 and 6; F, posterior thoracic sternum and pleonal somites 1-4; G, sternopleonal cavity.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFF98714FED12522D622759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965170/files/figure.png	https://doi.org/10.5281/zenodo.22965170	FIG. 1. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, outer view of chelae.	FIG. 1. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, outer view of chelae.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFF98714FED12522D622759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965174/files/figure.png	https://doi.org/10.5281/zenodo.22965174	FIG. 2. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, dorsal view of left cheliped; B, dorsal view of right cheliped; C, right fourth ambulatory leg; D, right third maxilliped; E, sternopleonal cavity;F, anterior thoracic sternum, telson and pleonal somites 5 and 6; G, tho- racic sternum, telson and pleonal somites 2-6; H, posterior thoracic sternum and pleonal somites 1-6. A and B photographed to same scale.	FIG. 2. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, dorsal view of left cheliped; B, dorsal view of right cheliped; C, right fourth ambulatory leg; D, right third maxilliped; E, sternopleonal cavity;F, anterior thoracic sternum, telson and pleonal somites 5 and 6; G, tho- racic sternum, telson and pleonal somites 2-6; H, posterior thoracic sternum and pleonal somites 1-6. A and B photographed to same scale.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFF98714FED12522D622759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965212/files/figure.png	https://doi.org/10.5281/zenodo.22965212	FIG. 16. — Thoracic sternums of: A, Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”; B, B. cursor Ng, 1986, paratype,♂ 15.6 × 13.4 mm (ZRC 1989.3036), Kalimantan; C, Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; D, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan. Abbreviations: bp, press button tubercle of pleonal locking mechanism; br5/6, median bridge between thoracic sternites 5 and 6; br6/7, median bridge between thoracic sternites 6 and 7; st4-8, thoracic sternites 4-8, respectively.	FIG. 16. — Thoracic sternums of: A, Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”; B, B. cursor Ng, 1986, paratype,♂ 15.6 × 13.4 mm (ZRC 1989.3036), Kalimantan; C, Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; D, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan. Abbreviations: bp, press button tubercle of pleonal locking mechanism; br5/6, median bridge between thoracic sternites 5 and 6; br6/7, median bridge between thoracic sternites 6 and 7; st4-8, thoracic sternites 4-8, respectively.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFF98714FED12522D622759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965185/files/figure.png	https://doi.org/10.5281/zenodo.22965185	FIG. 6. — Balssiathelphusa sucki (Balss, 1937) (A-F), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”, and B. cursor Ng, 1986 (G-M), holotype, ♂ 15.4 × 13.3 mm (MZB Cru Nr. 1279a), Kalimantan. A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, distal part of right G1 (dorso-mesial view, setae not drawn); F, right G2; G, left third maxilliped; H, frontal median triangle; I, left G1 (ventral view); J, left G1 (dorsal view); K, distal part of left G1 (ventral view); L, distal part of left G1 (dorsal view); M, left G2. Scale bars: A, B, F, 0.5 mm; C, D, 0.2 mm. I-M, after Ng (1986a: fig. 2A-F).	FIG. 6. — Balssiathelphusa sucki (Balss, 1937) (A-F), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”, and B. cursor Ng, 1986 (G-M), holotype, ♂ 15.4 × 13.3 mm (MZB Cru Nr. 1279a), Kalimantan. A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, distal part of right G1 (dorso-mesial view, setae not drawn); F, right G2; G, left third maxilliped; H, frontal median triangle; I, left G1 (ventral view); J, left G1 (dorsal view); K, distal part of left G1 (ventral view); L, distal part of left G1 (dorsal view); M, left G2. Scale bars: A, B, F, 0.5 mm; C, D, 0.2 mm. I-M, after Ng (1986a: fig. 2A-F).	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFF88717FF472388D30B76FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965192/files/figure.png	https://doi.org/10.5281/zenodo.22965192	FIG. 9. — Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, buccal cavity, anterior thoracic sternum, telson and pleonal somites 5 and 6; F, posterior thoracic sternum and pleonal somites 1-4; G, sternopleonal cavity.	FIG. 9. — Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, buccal cavity, anterior thoracic sternum, telson and pleonal somites 5 and 6; F, posterior thoracic sternum and pleonal somites 1-4; G, sternopleonal cavity.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFF88717FF472388D30B76FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965198/files/figure.png	https://doi.org/10.5281/zenodo.22965198	FIG. 11. — Natunusa natunaensis (Bott, 1970) n. comb.: A-E, holotype,♂ 23.0 × 20.0 mm (RMNH D29300),Natuna Island;F-H, ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; I-L, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., holotype, ♂ 23.1 × 18.9 mm (MNHN-IU-2013-13148), Kalimantan: A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, right G2; F, J, distal part of left G1 (ventral view); G, H, distal part of left G1 (dorsal views, drawn at slightly different angles); I, left G1 (ventral view); K, distal part of left G1 (dorsal view); L, left G2. Scale bars: A-E, J, K, 0.5 mm; C, D, F-H, I, L, 0.1 mm. A-H after Ng et al. (2023: fig. 13G-N); I-L after Ng & Guinot (2014: fig. 3).	FIG. 11. — Natunusa natunaensis (Bott, 1970) n. comb.: A-E, holotype,♂ 23.0 × 20.0 mm (RMNH D29300),Natuna Island;F-H, ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; I-L, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., holotype, ♂ 23.1 × 18.9 mm (MNHN-IU-2013-13148), Kalimantan: A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, right G2; F, J, distal part of left G1 (ventral view); G, H, distal part of left G1 (dorsal views, drawn at slightly different angles); I, left G1 (ventral view); K, distal part of left G1 (dorsal view); L, left G2. Scale bars: A-E, J, K, 0.5 mm; C, D, F-H, I, L, 0.1 mm. A-H after Ng et al. (2023: fig. 13G-N); I-L after Ng & Guinot (2014: fig. 3).	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFF88717FF472388D30B76FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965212/files/figure.png	https://doi.org/10.5281/zenodo.22965212	FIG. 16. — Thoracic sternums of: A, Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”; B, B. cursor Ng, 1986, paratype,♂ 15.6 × 13.4 mm (ZRC 1989.3036), Kalimantan; C, Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; D, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan. Abbreviations: bp, press button tubercle of pleonal locking mechanism; br5/6, median bridge between thoracic sternites 5 and 6; br6/7, median bridge between thoracic sternites 6 and 7; st4-8, thoracic sternites 4-8, respectively.	FIG. 16. — Thoracic sternums of: A, Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”; B, B. cursor Ng, 1986, paratype,♂ 15.6 × 13.4 mm (ZRC 1989.3036), Kalimantan; C, Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; D, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan. Abbreviations: bp, press button tubercle of pleonal locking mechanism; br5/6, median bridge between thoracic sternites 5 and 6; br6/7, median bridge between thoracic sternites 6 and 7; st4-8, thoracic sternites 4-8, respectively.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFF88717FF472388D30B76FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965185/files/figure.png	https://doi.org/10.5281/zenodo.22965185	FIG. 6. — Balssiathelphusa sucki (Balss, 1937) (A-F), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”, and B. cursor Ng, 1986 (G-M), holotype, ♂ 15.4 × 13.3 mm (MZB Cru Nr. 1279a), Kalimantan. A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, distal part of right G1 (dorso-mesial view, setae not drawn); F, right G2; G, left third maxilliped; H, frontal median triangle; I, left G1 (ventral view); J, left G1 (dorsal view); K, distal part of left G1 (ventral view); L, distal part of left G1 (dorsal view); M, left G2. Scale bars: A, B, F, 0.5 mm; C, D, 0.2 mm. I-M, after Ng (1986a: fig. 2A-F).	FIG. 6. — Balssiathelphusa sucki (Balss, 1937) (A-F), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”, and B. cursor Ng, 1986 (G-M), holotype, ♂ 15.4 × 13.3 mm (MZB Cru Nr. 1279a), Kalimantan. A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, distal part of right G1 (dorso-mesial view, setae not drawn); F, right G2; G, left third maxilliped; H, frontal median triangle; I, left G1 (ventral view); J, left G1 (dorsal view); K, distal part of left G1 (ventral view); L, distal part of left G1 (dorsal view); M, left G2. Scale bars: A, B, F, 0.5 mm; C, D, 0.2 mm. I-M, after Ng (1986a: fig. 2A-F).	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFA8717FEA42784D68C759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965170/files/figure.png	https://doi.org/10.5281/zenodo.22965170	FIG. 1. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, outer view of chelae.	FIG. 1. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, outer view of chelae.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFA8717FEA42784D68C759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965196/files/figure.png	https://doi.org/10.5281/zenodo.22965196	FIG. 10. — Lemurophusa phasma (Ng & Guinot, 2014) n. comb.: A, D, holotype, ♂ 23.1 × 18.9 mm (MNHN-IU-2013-13148), Kalimantan; B, E, paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan; C, F, G, H, paratype ♂ 23.5 × 19.0 mm (ZRC 2014.0092), Kalimantan. A, overall dorsal habitus; B, C, dorsal view of carapace; D, frontal view of cephalothorax; E, F, frontal median triangle, antennules, antennae and epistome; G, posterior thoracic sternum and pleonal somites 1-5; H, sternopleonal cavity.	FIG. 10. — Lemurophusa phasma (Ng & Guinot, 2014) n. comb.: A, D, holotype, ♂ 23.1 × 18.9 mm (MNHN-IU-2013-13148), Kalimantan; B, E, paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan; C, F, G, H, paratype ♂ 23.5 × 19.0 mm (ZRC 2014.0092), Kalimantan. A, overall dorsal habitus; B, C, dorsal view of carapace; D, frontal view of cephalothorax; E, F, frontal median triangle, antennules, antennae and epistome; G, posterior thoracic sternum and pleonal somites 1-5; H, sternopleonal cavity.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFA8717FEA42784D68C759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965212/files/figure.png	https://doi.org/10.5281/zenodo.22965212	FIG. 16. — Thoracic sternums of: A, Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”; B, B. cursor Ng, 1986, paratype,♂ 15.6 × 13.4 mm (ZRC 1989.3036), Kalimantan; C, Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; D, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan. Abbreviations: bp, press button tubercle of pleonal locking mechanism; br5/6, median bridge between thoracic sternites 5 and 6; br6/7, median bridge between thoracic sternites 6 and 7; st4-8, thoracic sternites 4-8, respectively.	FIG. 16. — Thoracic sternums of: A, Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”; B, B. cursor Ng, 1986, paratype,♂ 15.6 × 13.4 mm (ZRC 1989.3036), Kalimantan; C, Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; D, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan. Abbreviations: bp, press button tubercle of pleonal locking mechanism; br5/6, median bridge between thoracic sternites 5 and 6; br6/7, median bridge between thoracic sternites 6 and 7; st4-8, thoracic sternites 4-8, respectively.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFA8717FEA42784D68C759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965174/files/figure.png	https://doi.org/10.5281/zenodo.22965174	FIG. 2. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, dorsal view of left cheliped; B, dorsal view of right cheliped; C, right fourth ambulatory leg; D, right third maxilliped; E, sternopleonal cavity;F, anterior thoracic sternum, telson and pleonal somites 5 and 6; G, tho- racic sternum, telson and pleonal somites 2-6; H, posterior thoracic sternum and pleonal somites 1-6. A and B photographed to same scale.	FIG. 2. — Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”: A, dorsal view of left cheliped; B, dorsal view of right cheliped; C, right fourth ambulatory leg; D, right third maxilliped; E, sternopleonal cavity;F, anterior thoracic sternum, telson and pleonal somites 5 and 6; G, tho- racic sternum, telson and pleonal somites 2-6; H, posterior thoracic sternum and pleonal somites 1-6. A and B photographed to same scale.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFA8717FEA42784D68C759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965198/files/figure.png	https://doi.org/10.5281/zenodo.22965198	FIG. 11. — Natunusa natunaensis (Bott, 1970) n. comb.: A-E, holotype,♂ 23.0 × 20.0 mm (RMNH D29300),Natuna Island;F-H, ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; I-L, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., holotype, ♂ 23.1 × 18.9 mm (MNHN-IU-2013-13148), Kalimantan: A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, right G2; F, J, distal part of left G1 (ventral view); G, H, distal part of left G1 (dorsal views, drawn at slightly different angles); I, left G1 (ventral view); K, distal part of left G1 (dorsal view); L, left G2. Scale bars: A-E, J, K, 0.5 mm; C, D, F-H, I, L, 0.1 mm. A-H after Ng et al. (2023: fig. 13G-N); I-L after Ng & Guinot (2014: fig. 3).	FIG. 11. — Natunusa natunaensis (Bott, 1970) n. comb.: A-E, holotype,♂ 23.0 × 20.0 mm (RMNH D29300),Natuna Island;F-H, ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; I-L, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., holotype, ♂ 23.1 × 18.9 mm (MNHN-IU-2013-13148), Kalimantan: A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, right G2; F, J, distal part of left G1 (ventral view); G, H, distal part of left G1 (dorsal views, drawn at slightly different angles); I, left G1 (ventral view); K, distal part of left G1 (dorsal view); L, left G2. Scale bars: A-E, J, K, 0.5 mm; C, D, F-H, I, L, 0.1 mm. A-H after Ng et al. (2023: fig. 13G-N); I-L after Ng & Guinot (2014: fig. 3).	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFA8717FEA42784D68C759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965185/files/figure.png	https://doi.org/10.5281/zenodo.22965185	FIG. 6. — Balssiathelphusa sucki (Balss, 1937) (A-F), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”, and B. cursor Ng, 1986 (G-M), holotype, ♂ 15.4 × 13.3 mm (MZB Cru Nr. 1279a), Kalimantan. A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, distal part of right G1 (dorso-mesial view, setae not drawn); F, right G2; G, left third maxilliped; H, frontal median triangle; I, left G1 (ventral view); J, left G1 (dorsal view); K, distal part of left G1 (ventral view); L, distal part of left G1 (dorsal view); M, left G2. Scale bars: A, B, F, 0.5 mm; C, D, 0.2 mm. I-M, after Ng (1986a: fig. 2A-F).	FIG. 6. — Balssiathelphusa sucki (Balss, 1937) (A-F), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”, and B. cursor Ng, 1986 (G-M), holotype, ♂ 15.4 × 13.3 mm (MZB Cru Nr. 1279a), Kalimantan. A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, distal part of right G1 (dorso-mesial view, setae not drawn); F, right G2; G, left third maxilliped; H, frontal median triangle; I, left G1 (ventral view); J, left G1 (dorsal view); K, distal part of left G1 (ventral view); L, distal part of left G1 (dorsal view); M, left G2. Scale bars: A, B, F, 0.5 mm; C, D, 0.2 mm. I-M, after Ng (1986a: fig. 2A-F).	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFA8717FEA42784D68C759B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965192/files/figure.png	https://doi.org/10.5281/zenodo.22965192	FIG. 9. — Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, buccal cavity, anterior thoracic sternum, telson and pleonal somites 5 and 6; F, posterior thoracic sternum and pleonal somites 1-4; G, sternopleonal cavity.	FIG. 9. — Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island: A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, antennules, antennae and epistome; E, buccal cavity, anterior thoracic sternum, telson and pleonal somites 5 and 6; F, posterior thoracic sternum and pleonal somites 1-4; G, sternopleonal cavity.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFC8713FF342387D3A0769E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965196/files/figure.png	https://doi.org/10.5281/zenodo.22965196	FIG. 10. — Lemurophusa phasma (Ng & Guinot, 2014) n. comb.: A, D, holotype, ♂ 23.1 × 18.9 mm (MNHN-IU-2013-13148), Kalimantan; B, E, paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan; C, F, G, H, paratype ♂ 23.5 × 19.0 mm (ZRC 2014.0092), Kalimantan. A, overall dorsal habitus; B, C, dorsal view of carapace; D, frontal view of cephalothorax; E, F, frontal median triangle, antennules, antennae and epistome; G, posterior thoracic sternum and pleonal somites 1-5; H, sternopleonal cavity.	FIG. 10. — Lemurophusa phasma (Ng & Guinot, 2014) n. comb.: A, D, holotype, ♂ 23.1 × 18.9 mm (MNHN-IU-2013-13148), Kalimantan; B, E, paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan; C, F, G, H, paratype ♂ 23.5 × 19.0 mm (ZRC 2014.0092), Kalimantan. A, overall dorsal habitus; B, C, dorsal view of carapace; D, frontal view of cephalothorax; E, F, frontal median triangle, antennules, antennae and epistome; G, posterior thoracic sternum and pleonal somites 1-5; H, sternopleonal cavity.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFC8713FF342387D3A0769E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965198/files/figure.png	https://doi.org/10.5281/zenodo.22965198	FIG. 11. — Natunusa natunaensis (Bott, 1970) n. comb.: A-E, holotype,♂ 23.0 × 20.0 mm (RMNH D29300),Natuna Island;F-H, ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; I-L, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., holotype, ♂ 23.1 × 18.9 mm (MNHN-IU-2013-13148), Kalimantan: A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, right G2; F, J, distal part of left G1 (ventral view); G, H, distal part of left G1 (dorsal views, drawn at slightly different angles); I, left G1 (ventral view); K, distal part of left G1 (dorsal view); L, left G2. Scale bars: A-E, J, K, 0.5 mm; C, D, F-H, I, L, 0.1 mm. A-H after Ng et al. (2023: fig. 13G-N); I-L after Ng & Guinot (2014: fig. 3).	FIG. 11. — Natunusa natunaensis (Bott, 1970) n. comb.: A-E, holotype,♂ 23.0 × 20.0 mm (RMNH D29300),Natuna Island;F-H, ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; I-L, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., holotype, ♂ 23.1 × 18.9 mm (MNHN-IU-2013-13148), Kalimantan: A, right G1 (dorsal view); B, right G1 (ventral view); C, distal part of right G1 (dorsal view); D, distal part of right G1 (ventral view); E, right G2; F, J, distal part of left G1 (ventral view); G, H, distal part of left G1 (dorsal views, drawn at slightly different angles); I, left G1 (ventral view); K, distal part of left G1 (dorsal view); L, left G2. Scale bars: A-E, J, K, 0.5 mm; C, D, F-H, I, L, 0.1 mm. A-H after Ng et al. (2023: fig. 13G-N); I-L after Ng & Guinot (2014: fig. 3).	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFC8713FF342387D3A0769E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965212/files/figure.png	https://doi.org/10.5281/zenodo.22965212	FIG. 16. — Thoracic sternums of: A, Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”; B, B. cursor Ng, 1986, paratype,♂ 15.6 × 13.4 mm (ZRC 1989.3036), Kalimantan; C, Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; D, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan. Abbreviations: bp, press button tubercle of pleonal locking mechanism; br5/6, median bridge between thoracic sternites 5 and 6; br6/7, median bridge between thoracic sternites 6 and 7; st4-8, thoracic sternites 4-8, respectively.	FIG. 16. — Thoracic sternums of: A, Balssiathelphusa sucki (Balss, 1937), holotype, ♂ 21.2 × 17.1 mm (ZMH-K-003545), “Sumatra”; B, B. cursor Ng, 1986, paratype,♂ 15.6 × 13.4 mm (ZRC 1989.3036), Kalimantan; C, Natunusa natunaensis (Bott, 1970) n. comb., ♂ 12.9 × 11.3 mm (ZRC 2021.0831), Natuna Island; D, Lemurophusa phasma (Ng & Guinot, 2014) n. comb., paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan. Abbreviations: bp, press button tubercle of pleonal locking mechanism; br5/6, median bridge between thoracic sternites 5 and 6; br6/7, median bridge between thoracic sternites 6 and 7; st4-8, thoracic sternites 4-8, respectively.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFC8713FF342387D3A0769E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965214/files/figure.png	https://doi.org/10.5281/zenodo.22965214	FIG. 17. — Lemurophusa empusa n. gen.,n. sp.,holotype,♀, 24.2 × 18.5 mm (MZB Cru),Kalimantan,thoracic sternum:A-C, pattern of sutures in sternopleonal cavity photographed at different angles; D, internal skeletal view showing longitudinal median plate. Abbreviations: bp, press button tubercle of pleonal locking mechanism;br5/6, median bridge between thoracic sternites 5 and 6; br6/7, median bridge between thoracic sternites 6 and 7; st4-8, thoracic sternites 4-8, respectively.	FIG. 17. — Lemurophusa empusa n. gen.,n. sp.,holotype,♀, 24.2 × 18.5 mm (MZB Cru),Kalimantan,thoracic sternum:A-C, pattern of sutures in sternopleonal cavity photographed at different angles; D, internal skeletal view showing longitudinal median plate. Abbreviations: bp, press button tubercle of pleonal locking mechanism;br5/6, median bridge between thoracic sternites 5 and 6; br6/7, median bridge between thoracic sternites 6 and 7; st4-8, thoracic sternites 4-8, respectively.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFE8713FEBA2663D1AF755C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965200/files/figure.png	https://doi.org/10.5281/zenodo.22965200	FIG. 12. — Lemurophusa empusa n. gen., n. sp.: A, C, holotype, ♀ 24.2 × 18.5 mm (MZB Cru), Kalimantan; B, D, paratype ♀ 22.2 × 17.2 mm (MZB Cru), Kalimantan. A, B, overall dorsal habitus; C, D, dorsal view of carapace.	FIG. 12. — Lemurophusa empusa n. gen., n. sp.: A, C, holotype, ♀ 24.2 × 18.5 mm (MZB Cru), Kalimantan; B, D, paratype ♀ 22.2 × 17.2 mm (MZB Cru), Kalimantan. A, B, overall dorsal habitus; C, D, dorsal view of carapace.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFE8713FEBA2663D1AF755C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965202/files/figure.png	https://doi.org/10.5281/zenodo.22965202	FIG. 13. — Lemurophusa empusa n. gen., n. sp., holotype, ♀ 24.2 × 18.5 mm (MZB Cru), Kalimantan:A, frontal view of cephalothorax; B, C, frontal median triangle, antennules, antennae and epistome;D, right third maxilliped; E, outer view of right chela; F, outer view of left chela; G, third maxillipeds and anterior thoracic sternum; H, third maxillipeds, anterior thoracic sternum and pleon; I, sternopleonal cavity showing vulvae.	FIG. 13. — Lemurophusa empusa n. gen., n. sp., holotype, ♀ 24.2 × 18.5 mm (MZB Cru), Kalimantan:A, frontal view of cephalothorax; B, C, frontal median triangle, antennules, antennae and epistome;D, right third maxilliped; E, outer view of right chela; F, outer view of left chela; G, third maxillipeds and anterior thoracic sternum; H, third maxillipeds, anterior thoracic sternum and pleon; I, sternopleonal cavity showing vulvae.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFE8713FEBA2663D1AF755C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965196/files/figure.png	https://doi.org/10.5281/zenodo.22965196	FIG. 10. — Lemurophusa phasma (Ng & Guinot, 2014) n. comb.: A, D, holotype, ♂ 23.1 × 18.9 mm (MNHN-IU-2013-13148), Kalimantan; B, E, paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan; C, F, G, H, paratype ♂ 23.5 × 19.0 mm (ZRC 2014.0092), Kalimantan. A, overall dorsal habitus; B, C, dorsal view of carapace; D, frontal view of cephalothorax; E, F, frontal median triangle, antennules, antennae and epistome; G, posterior thoracic sternum and pleonal somites 1-5; H, sternopleonal cavity.	FIG. 10. — Lemurophusa phasma (Ng & Guinot, 2014) n. comb.: A, D, holotype, ♂ 23.1 × 18.9 mm (MNHN-IU-2013-13148), Kalimantan; B, E, paratype, ♂ 18.1 × 15.1 mm (ZRC 2014.0092), Kalimantan; C, F, G, H, paratype ♂ 23.5 × 19.0 mm (ZRC 2014.0092), Kalimantan. A, overall dorsal habitus; B, C, dorsal view of carapace; D, frontal view of cephalothorax; E, F, frontal median triangle, antennules, antennae and epistome; G, posterior thoracic sternum and pleonal somites 1-5; H, sternopleonal cavity.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
03FF87FAFFFE8713FEBA2663D1AF755C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/22965217/files/figure.png	https://doi.org/10.5281/zenodo.22965217	FIG. 18. —Indonesian Borneo localities of Lemurophusa phasma (Ng & Guinot, 2014) n. comb. (red circle) and L. empusa n. gen., n. sp. (green circles) in East Kalimantan,Indonesia,Borneo:1, Gua Kambing,Mount Batu,Mangkalihat Peninsula,01°09.444’N,117°28.479’E; 2, Kecabe Cave,Sungai Baai karst,01°38.018’N, 117°24.240’E; 3, Lubang Tebot Cave, Merabu karst, 01°34.964’N, 117°21.113’E. From Google Maps.	FIG. 18. —Indonesian Borneo localities of Lemurophusa phasma (Ng & Guinot, 2014) n. comb. (red circle) and L. empusa n. gen., n. sp. (green circles) in East Kalimantan,Indonesia,Borneo:1, Gua Kambing,Mount Batu,Mangkalihat Peninsula,01°09.444’N,117°28.479’E; 2, Kecabe Cave,Sungai Baai karst,01°38.018’N, 117°24.240’E; 3, Lubang Tebot Cave, Merabu karst, 01°34.964’N, 117°21.113’E. From Google Maps.	2026-07-13	Ng, Peter K. L.;Guinot, Danièle		Zenodo	biologists	Ng, Peter K. L.;Guinot, Danièle			
