identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03FE87FCFFF04974FF456C7AFC10E9AD.text	03FE87FCFFF04974FF456C7AFC10E9AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grewia souladethiana Phengmala & Tagane. A. Flowering 2025	<div><p>Grewia souladethiana Phengmala &amp; Tagane, sp. nov. (Figs. 2 &amp; 3)</p><p>TYPE: — LAOS. Khammouane Province: Hinboun Dist., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.675&amp;materialsCitation.latitude=17.825" title="Search Plazi for locations around (long 104.675/lat 17.825)">Ban Houana</a>, 17.825°N, 104.675°E, alt. 287 m, 7 September 2024, S. Tagane, P. Souladeth, V.S. Dang, T. Vongthavone, K. Souvannakhoummane, P. Phonepaseuth, K. Phengmala, Q.T. Pham, T. Yamamoto, K. Yamazaki Z1741 [fl.] (holotype FOF [FOF0007156!], isotypes KAG [KAG188404!, KAG188405!], VNM) .</p><p>Diagnosis: — Grewia souladethiana is similar to G. thailandica Chantaranothai &amp; Nualngam (2022: 2) of Thailand, in having acuminate leaf apex, axillary 2‒3-flowered inflorescences, and erect peduncles, but distinguished by its tree habit (vs. scandent shrubs to woody climbers in G. thailandica), longer acumen of leaf blades (up to 2.8 cm long vs. up to 1 cm long), shorter petioles (0.6–0.9 cm long vs. 1.0– 1.3 cm long), smaller sepals (18 mm long vs. 25–27 mm long), smaller and triangular petals 2.0 × 1.8 mm (vs. oblong or lanceolate, 8.0‒9.0 × 3.0‒4.0 mm), and solitary or 2-lobed fruits (vs. (1–)3–4-lobed) (Table 1).</p><p>Description:—Trees, 3‒4 m tall. Indumentum simple or stellate hairs. Young twigs yellowish green in vivo, dark greyish brown in sicco, sparsely hairy, old twigs glabrous, lenticellate. Stipules linear, 5.0–8.0 mm long, almost glabrous on both surfaces. Leaves alternate; blades elliptic-oblong, ovate-oblong, 8.7‒18.9 × 2.9–6.0 cm, length/width ratio 3–3.2, adaxial surface pale green, dull brownish yellow, sparsely hairy, abaxial surface dull yellowish green, sparsely hairy, apex acuminate, acumen up to 2.8 cm long, base attenuate or obtuse, symmetric or asymmetric, margin serrate, basal veins 3, midrib prominent on both surfaces, secondary veins 6‒7 pairs, prominent abaxially, tertiary veins scalariform, distinct abaxially; petioles 0.6‒0.9 cm long, sparsely stellate hairy. Inflorescences axillary, solitary or in cluster of 2- or 3-flowered; peduncle 0.9‒1.4 cm long at anthesis, erect, sparsely hairy; outer bracts narrowly-lanceolate, 3.0 × 15 mm, apex acuminate, margin ciliolate, caducous; inner bracts 3, whorled, narrowly oblong-lanceolate, 2.0 × 3.0 mm, glabrous adaxially, sparsely hairy abaxially, apex acute, margin ciliolate, caducous. Flowers bisexual; pedicels 1.4‒1.7 cm long; mature buds oblong-obovoid, 1.9 × 0.6 cm. Sepals free, 5, oblanceolate, 18 × 3.0 mm, apex acute, adaxial surface glabrous, abaxial surface verrucose, sparsely hairy, 2-grooved. Petals 5, free, triangular, 2.0 × 1.8 mm, glabrous on both surfaces, apex obtuse, margin entire, not ciliate, with clawed appendages, appendages glabrous on both surfaces, densely stellate hairy along margin, glands depressed globose. Androgynophore: lower part slightly grooved, 2.3 × 2.0 mm, glabrous; upper part cylindrical, 1.2 × 1.5 mm, densely hairy. Stamens numerous, filaments 2.8‒6.5 mm long, anthers 1.1‒1.3 mm long, glabrous. Ovary shallowly depressed globose, 1.8 mm in diam., hairy, 2-locular, 4 ovules per locule, style 7.8 mm long, sparsely hairy at basal 1/5, stigma 4‒5 lobed. Fruits drupaceous, 1‒2-lobed; lobes globose, 5.0‒9.0 mm in diam., sparsely hairy; stipe 1.7‒1.8 mm long. Seeds 2 per fruit, depressed obovoid-globose 4.0 × 7.0 mm, dark brown, scabrous, sparsely stellate hairy.</p><p>Additional specimen examined:— LAOS. Khammouane Province: Gnommalath District, near Ban Tat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.06574&amp;materialsCitation.latitude=17.54504" title="Search Plazi for locations around (long 105.06574/lat 17.54504)">Phou Hin Poun National Protected Area</a> (currently National Park), at the edge of evergreen forest on limestone hill, 17.54504°N, 105.06574°E, 206 m elev., 5 September 2024, S. Tagane et al. Z1545 [fl.] (FOF [FOF0006635], KAG [KAG188212!], VNM) ; Thakhek District, Ban Pha Laem (Phatong), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.73403&amp;materialsCitation.latitude=17.75929" title="Search Plazi for locations around (long 104.73403/lat 17.75929)">Phou Hin Poun National Protected Area</a> (currently National Park), in open grassland on limestone hill, 17.5874°N, 104.824644°E, 190 m elev., 22 November 2024, P. Souladeth et al. Z1959 [fl. &amp; fr.] (FOF [FOF0007028], KAG [KAG200707]), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.73403&amp;materialsCitation.latitude=17.75929" title="Search Plazi for locations around (long 104.73403/lat 17.75929)">Hinboun District</a>, Ban Bouamlou, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.73403&amp;materialsCitation.latitude=17.75929" title="Search Plazi for locations around (long 104.73403/lat 17.75929)">Phou Hin Poun National Park</a>, at edge of disturbed semi-evergreen forest, 17.75929°N, 104.73403°E, 190 m elev., 29 June 2025, S. Tagane et al. Z2740 [fl.] (FOF0008005, KAG202309, VNM) .</p><p>Distribution:— LAOS (Khammouane Province) (Fig. 1).</p><p>Habitat and Ecology: — Grewia souladethiana grows at the edges of evergreen broad-leaved forests and in open grasslands on limestone hills, at elevations of 190– 287 m.</p><p>Phenology: —Flowering specimens were collected in June, September and November, and fruiting in November.</p><p>Etymology: —The specific epithet honors Dr. Phetlasy Souladeth, a curator of FOF herbarium, Faculty of Forest Science, National University of Laos, who led our field surveys in Laos and encouraged the first author to intensively study on the Lao flora. Thanks to her, we could discover this new species.</p><p>Vernacular name:—UໍເƜ̰ືƟƆfflຸÐະເ00 (Por Leaung Souladeth) (suggested here).</p><p>Preliminary conservation assessment: —Vulnerable (VU). Grewia souladethiana is endmic to Khammouane Province and currently known from four locations, all within the protected areas in Phou Hin Poun National Park (Fig. 1). In each population, we observed 10–50 individuals. The area of occupancy (AOO) and the extent of occurence (EOO) are calculated as 16 km 2 and 314 km 2, repsectively, using GeoCAT (Bachman et al. 2011). Based on these information, it may satisfy the criteria of Endangered (EN). However, considering G. souladethiana is locally common in forest edges and open grassland, which are frequently affected by human disturbance, it is likely to be more widely distributed within the national park (a total area 2,388.2 km 2) and its margins, we suggest its status as Vulnerable (VU) according to the critera B1ab(iii), 2ab(iii) (IUCN 2024).</p><p>Notes:— Among ten species of Grewia previously known from Laos, G. souladethiana might resemble G. abutilifolia . However, it can be easily distinguished by its tree habit (vs. shrubs in G. abutilifolia), shorter petioles 0.6–0.9 cm long (vs. 1.0–2.0 cm long), elliptic-oblong to ovate-oblong leaf shape (vs. ovate to nearly orbicular), longer pedicels 1.4‒1.7 cm (vs. 0.1–0.8 cm), more secondary veins (6‒7 pairs in G. souladethiana vs. 3–4 pairs), and triangular petals ca. 2.0 × 1.8 mm (vs. oblong, 2–3 × 1 mm) (Jussieu 1804, Phengklai 1993, Tang et al. 2007).</p></div>	https://treatment.plazi.org/id/03FE87FCFFF04974FF456C7AFC10E9AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Phengmala, Kajonesuk;Tagane, Shuichiro;Dang, Van-Son;Kongxaisavath, Deuanta;Yamamoto, Takenori;Souvannakhoummane, Keooudone;Phonepaseuth, Phongphayboun;Vongthavone, Thyraphon;Sengthong, Anousone;Trong, Pham Quoc;Yamazaki, Kaito	Phengmala, Kajonesuk, Tagane, Shuichiro, Dang, Van-Son, Kongxaisavath, Deuanta, Yamamoto, Takenori, Souvannakhoummane, Keooudone, Phonepaseuth, Phongphayboun, Vongthavone, Thyraphon, Sengthong, Anousone, Trong, Pham Quoc, Yamazaki, Kaito (2025): Grewia souladethiana, a new species of Malvaceae from Laos. Phytotaxa 728 (1): 61-68, DOI: 10.11646/phytotaxa.728.1.6, URL: https://doi.org/10.11646/phytotaxa.728.1.6
03FE87FCFFF74977FF456BD8FCECEC8B.text	03FE87FCFFF74977FF456BD8FCECEC8B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grewia Linnaeus 1753	<div><p>Key to species of Grewia in Laos</p><p>1. Climbing shrubs or scrambling shrubs...............................................................................................................................................2</p><p>2. Leaf blades ovate to ovate-oblong, 4–6 cm wide; secondary veins ca. 10 pairs............................................................ G. acuminata</p><p>2. Leaf blades lanceolate, to 3 cm wide; secondary veins 5–6 pairs................................................................................. G. astropetala</p><p>1. Erect shrubs or trees ...........................................................................................................................................................................3</p><p>3. Inflorescence pendulous ................................................................................................................................................. G. sessilifolia</p><p>3. Inflorescence erect..............................................................................................................................................................................4</p><p>4. Leaves narrowly lanceolate to oblong-lanceolate ..............................................................................................................................5</p><p>5. Small shrubs to 1 m tall; secondary veins 3–4 pairs ...................................................................................................... G. retusifolia</p><p>5. Shrubs to trees, 1.5–10 m tall; secondary veins 4–11 pairs................................................................................................................6</p><p>6. Leaves glabrous or glabrescent ........................................................................................................................................ G. laevigata</p><p>6. Leaves hairy, especially densely tomentose on abaxial surfaces........................................................................................................7</p><p>7. Leaf margin obscurely serrulate; secondary veins 7–11 pairs................................................................................................. G. lacei</p><p>7. Leaf margin serrate; secondary veins 3–7 hairs .................................................................................................................. G. hirsuta</p><p>4. Leaves ovate, orbicular or elliptic, oblong-elliptic.............................................................................................................................8</p><p>8. Leaves ovate to orbicular, base symmetric ................................................................................................................... G. abutilifolia</p><p>8. Leaves elliptic, oblong-elliptic, base asymmetric or symmetric ........................................................................................................9</p><p>9. Branches sparsely hairy to glabrous; leaves dull yellowish green abaxially............................................................ G. souladethiana</p><p>9. Branches softly tomentose; leaves glaucous abaxially.....................................................................................................................10</p><p>10. Fruits globose, not lobed. ............................................................................................................................................... G. eriocapra</p><p>10. Fruits ovoid, 4-lobed .................................................................................................................................................... G. celtidifolia</p></div>	https://treatment.plazi.org/id/03FE87FCFFF74977FF456BD8FCECEC8B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Phengmala, Kajonesuk;Tagane, Shuichiro;Dang, Van-Son;Kongxaisavath, Deuanta;Yamamoto, Takenori;Souvannakhoummane, Keooudone;Phonepaseuth, Phongphayboun;Vongthavone, Thyraphon;Sengthong, Anousone;Trong, Pham Quoc;Yamazaki, Kaito	Phengmala, Kajonesuk, Tagane, Shuichiro, Dang, Van-Son, Kongxaisavath, Deuanta, Yamamoto, Takenori, Souvannakhoummane, Keooudone, Phonepaseuth, Phongphayboun, Vongthavone, Thyraphon, Sengthong, Anousone, Trong, Pham Quoc, Yamazaki, Kaito (2025): Grewia souladethiana, a new species of Malvaceae from Laos. Phytotaxa 728 (1): 61-68, DOI: 10.11646/phytotaxa.728.1.6, URL: https://doi.org/10.11646/phytotaxa.728.1.6
