taxonID	type	description	language	source
03FE87B39E2DFFDD48DBFF4AFA2DFC6C.taxon	type_taxon	Type species. Odontomutilla abbottii Ashmead, 1899 by original designation (= Mutilla saussurei Sichel & Radoszkowski, 1869, according to André 1901: 329), Afrotropical region.	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
03FE87B39E2DFFDD48DBFF4AFA2DFC6C.taxon	diagnosis	Diagnosis. Female. Mesonotum laterally expanded to tooth or tubercle. T 1 broadly disciform. T 2 with large deep anterolateral pubescent fovea or linear felt line. MALE. Mesoscutellum bidentate posteriorly. Propodeum with posterolateral corners angulate or dentate. T 1 broadly disciform. Both sexes of Odontomutilla are larger-bodied (8 – 18 mm) than the structurally similar Yamanetilla Lelej, 1996 (6 – 9 mm). Species included and distribution. Currently this genus is composed of 100 valid species and 17 valid subspecies distributed mainly in the Afrotropical (59 species and 13 subspecies) and Oriental (35 species and three subspecies) regions, with four species and one subspecies in the Australian and two species in the Palaearctic regions (Pagliano et al. 2020, updated; Lelej et al. 2020; Lelej 2023; Terine & Girish Kumar 2024; and current data) (Table 1). Sixteen taxa have been named by Nonveiller (1975) for Afrotropical fauna, but never described (nomina nuda).	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
03FE87B39E2DFFDE48DBFBAAFEDEFD31.taxon	description	(Figs 1 – 4) Type material. Holotype — ♀, Iran, Loristan Province, Mahmudvand [Veysian, 33 ° 29 ′ 10 ″ N 48 ° 01 ′ 46 ″ E], 4 – 5. VI. 2010, R. Yakovlev [IBSS]. Paratype, SW Iran, Chaharmahal and Bakhtiari Province, Gandoman, Kuhe-Kalar, 2750 m, 13 – 14. VII. 1982, H. Brouman & A. Pazuki, 1 ♀ [IBSS].	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
03FE87B39E2DFFDE48DBFBAAFEDEFD31.taxon	diagnosis	Diagnosis. FEMALE. Genal carina strong, forming tooth at hypostomal carina and extending posteriorly to occipital carina where distinctly angulate. Head, mesosoma, legs and antennae ferruginous-red; metasoma black. T 2 with basal medial spot of white setae. T 3 with no interrupted band of white setae. MALE. Unknown.	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
03FE87B39E2DFFDE48DBFBAAFEDEFD31.taxon	description	Description. FEMALE. Body length 8.4 – 11.2 mm. Coloration and setation. Head and mesosoma ferruginous-red, while metasoma black. Legs, antennae and palps ferruginous-red. Mandible ferruginous-red with brown apex. Body setae generally sparse and whitish, except on T 1, T 2, T 4 – 6 with subappressed and brownish setae. Frons, vertex and mesosomal dorsum with sparse, suberect, brownish setae, while posterior propodeal face and anterior face of T 1 with long, dense, recumbent setae. Mesopleural vertical carina has long erect whitish setae. Legs, especially meso- and metatibia, with long, dense, recumbent, whitish setae. T 2 with basal medial spot of whitish setae. T 3 with noninterrupted band of whitish setae. S 2 – 6 with sparse, suberect, long, whitish setae, forming apical fringe of dense setae on S 2 – 5. Head. Width behind eye 0.93 × humeral mesosomal width. Frons, vertex, and gena with large sometimes confluent punctures. Mandible with dorsal carina extending from base to apex, apex unidentate. Clypeus apically not dentate, with transverse preapical carina. Antennal scrobe with arcuate lateral carina. Genal carina strong, forming large tooth at hypostomal carina and extending posteriorly to occipital carina where distinctly angulate. F 1 1.7 × F 2 and 2.0 × pedicel length, F 2 1.25 × pedicel length. Mesosoma. Length 1.08 × maximum width. Dorsum of mesosoma with large elongated coarse punctures; mesopleuron convex, punctured, vertical carina ending dorsally to tooth. Humeral carina well developed with oblique carina posterad. Ratio of width of humeral angle, anterior spiracle, lateral mesonotal tooth, propodeal spiracle, and posterolateral propodeal angle 72: 78: 82: 75: 67. Scutellar scale lacking. Posterior propodeal face abrupt, with dense punctures smaller to metasomal base; lateral propodeal face with large dense punctures. Metasoma. T 1 broad with distinct anterior and dorsal faces, dorsally with small dense punctures. T 2 with large coarse punctures, posteromedially with small dense ones; posterior border smooth. T 4 – 5 with small dense punctures, posterior half smooth. S 3 – 5 with small dense punctures, posterior border smooth. S 1 with simple longitudinal lamella. S 2 with dense large punctures, posterior border smooth. T 2 anterolaterally with linear felt line. T 6 basally and laterally coarsely punctate, setose, apical half with medial part without setae, pygidial plate not carinated laterad even apically, considerably convergent basally, with transverse striae. MALE unknown.	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
03FE87B39E2DFFDE48DBFBAAFEDEFD31.taxon	distribution	Distribution. Iran (Loristan, Chaharmahal and Bakhtiari).	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
03FE87B39E2DFFDE48DBFBAAFEDEFD31.taxon	discussion	Remarks. A new species is the second one of Odontomutilla in the Palaearctic and related with species from the Afrotropical region by the linear shape of lateral felt line on T 2. The female of Odontomutilla storozhenkoi, sp. nov. easily differs from that of O. chrysococcina (Sichel & Radoszkowski, 1869) also described from Iran by the lacking of lateral pale spots on T 2.	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
03FE87B39E2DFFDE48DBFBAAFEDEFD31.taxon	etymology	Etymology. I am pleased to name this new species in honor of Prof. Sergey Yu. Storozhenko, world authority on orthopterous insects and grilloblattids and my best colleague, on the occasion of his 70 th birthday. Treat as a noun in apposition.	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
03FE87B39E2EFFD848DBFD67FCD4FE49.taxon	description	(Figs 5 – 8)	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
03FE87B39E2EFFD848DBFD67FCD4FE49.taxon	materials_examined	Type material. Holotype — ♀, Laos, Vientiane Province, Nam Lik Eco-Village, 22. V. 2016, M. M. Omelko [IBSS]. Paratype, Vietnam [Dong Nai], Cat Tien [national park], 21. IV. 2007, V. A. Zryanin, 1 ♀ [IBSS].	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
03FE87B39E2EFFD848DBFD67FCD4FE49.taxon	diagnosis	Diagnosis. FEMALE. Genal carina strong, extending anteriorly to mandibular base and extending posteriorly to occipital carina where distinctly rounded. Hypostomal carina anteriorly with strong teeth. Head and mesosoma ferruginous-red, metasoma black. T 1 dorsally with black setae. T 2 in basal half medially with wide spot (invaginated posterad) of whitish setae and small lateral spots of whitish setae. T 3 with narrowly interrupted band of whitish setae. MALE. Unknown.	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
03FE87B39E2EFFD848DBFD67FCD4FE49.taxon	description	Description. FEMALE. Body length 10.4 – 12.0 mm. Coloration and setation. Head and mesosoma ferruginous-red, metasoma black. Head with sparse whitish setae mixed on frons, occiput and genae with sparse erect black setae. Legs black, palps brown. Body setae generally sparse and black, except posterior propodeal face and anterior face of T 1 with long dense erect whitish setae; frons, vertex, and mesosomal dorsum with short sparse erect black setae. Mesopleural vertical carina with long erect whitish setae. Legs, especially meso- and metatibia with long dense recumbent yellowish setae. T 1 dorsally with dense black setae. T 2 with medial large spot invaginated posterad of whitish setae and two lateral small spots located on level of posterior transverse line of medial spot. T 3 with narrowly interrupted band of whitish setae, interruption much wider anterad. T 6 except pygidial plate with whitish setae. S 2 – 5 with sparse sub-erect long yellowish setae, apically with fringe of dense yellowish setae, S 6 with dense yellowish setae oriented backward. Head. Width behind eye 0.93 × humeral mesosomal width. Frons, vertex, and gena with large confluent punctures. Mandible with dorsal carina extending from base to apex, apex unidentate. Clypeus with elevated anterior border, basomedial portion with two tubercles. Antennal scrobe with arcuate lateral carina. Genal carina strong, extending anteriorly to mandibular base and extending posteriorly to occipital carina where distinctly rounded. Hypostomal carina anteriorly with strong teeth. F 1 1.15 × F 2 and 1.5 × pedicel length, F 2 1.4 × pedicel length. Mesosoma. Length 1.15 × maximum width. Dorsum of mesosoma with large coarse elongated punctures; mesopleuron convex, punctured, vertical carina ending dorsally to strong tooth. Humeral carina well developed with oblique carina posterad. Ratio of width of humeral angle, anterior spiracle, lateral mesonotal tooth, propodeal spiracle, and posterolateral propodeal angle 70: 72: 80: 70: 60. Scutellar scale lacking. Posterior propodeal face abrupt, lateral propodeal face with sparse micropunctures. Metasoma. T 1 broad with distinct anterior and dorsal faces, dorsally with small dense punctures. T 2 with large coarse punctures, behind spots with small dense ones, posterior border smooth. T 4 – 5 and S 3 – 5 with small dense punctures, posterior border of T 4 – 5 smooth. S 1 with simple longitudinal lamella. S 2 with dense large punctures. T 2 anterolaterally with deep elongated pubescent fovea. T 6 basally and laterally coarsely punctate, setose, apical half with medial part without setae, pygidial plate not carinated laterad even apically, considerably convergent basally, with transverse striae. MALE unknown.	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
03FE87B39E2EFFD848DBFD67FCD4FE49.taxon	distribution	Distribution. Laos, Vietnam.	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
03FE87B39E2EFFD848DBFD67FCD4FE49.taxon	discussion	Remarks. A new species belongs to the small group of the genus with three spots on T 2 in female: O. spectra (Bingham, 1908) and O. paderua Das & Girish Kumar in Das et al., 2017 from India; O. chrysococcina (Sichel & Radoszkowski, 1869) from Iran and O. speciosa (Smith, 1855) from South-East China. Odontomutilla williamsi sp. nov. is most related with O. paderua but easily differs by the position of lateral pale spots on T 2 far from base of T 2 and by the shape of medial pale spot on T 2.	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
03FE87B39E2EFFD848DBFD67FCD4FE49.taxon	etymology	Etymology. I am delighted to name this new species in honor of Kevin A. Williams, world authority on Mutillidae and my best colleague. Treat as a noun in apposition.	en	Lelej, Arkady S. (2025): The color syndrome of species in the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae), and a description of two new red-headed species. Zootaxa 5715 (1): 236-243, DOI: 10.11646/zootaxa.5715.1.21, URL: https://doi.org/10.11646/zootaxa.5715.1.21
