taxonID	type	description	language	source
03F3861CFFC1FF93FF6CC289C422CCDC.taxon	description	About 4.7 – 8.2 m below the base of sandstone level:	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFC1FF92FF6CC4F4C5B1C96E.taxon	description	diverse mollusc fauna was described by ANDRI et al. (2005), while some brachiopods were mentioned by SACCO (1902).	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFC0FF91FE93C0CCC5F3CD7E.taxon	description	Terebratula ampulla (Brocchi, 1814), 2 fragmentary pedicle valves and 1 brachial valve, RGM. 1309900 Terebratulina retusa (Linnaeus, 1758), 1 complete specimen, RGM. 1309901 North Sea Basin England (Fig. 2) Gedgrave (Suffolk), Butley River cliff outcrop, 52.084167 N; 1.497500 E. Pliocene, Zanclean, Coralline Crag Formation, collected by A. W. Janssen, 14 September 1987. Lithostratigraphy of the Coralline Crag was given by BALSON et al. (1993) on the basis of seven boreholes between Gedgrave and Aldeburgh. Brachiopods of the Coralline Crag were discussed by classical papers as WOOD (1872) and DAVIDSON (1874 a). 0.55 – 0.85 m below top of Pliocene deposits:	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFC0FF91FE93C0CCC5F3CD7E.taxon	description	Glottidia? sp., 24 fragments, RGM. 793911	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFC0FF91FE93C0CCC5F3CD7E.taxon	description	1.00 – 1.25 m below top of Pliocene deposits:	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFCDFF9FFF6CC5F0C559CE55.taxon	description	The Netherlands (Fig. 4) Goes (Zeeland), unspecified borehole (Pouwer, pers. comm.). 68 m below surface, Pliocene: Terebratulida indet sp. (not Terebratella cf. spitzbergensis as indicated on the label), 4 indeterminable fragments, RGM. 29657 Terebratulida indet sp. (not Terebratulina caputserpentis as indicated on the label), 1 indeterminable fragment, RGM. 29672 Pliothyrina? indet. sp. (not Argiope cf. cistellula as indicated on the label), 1 fragment, RGM. 29679 74 m below surface, Pliocene:	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFCDFF9FFF6CC307C532CBB2.taxon	description	(Fig. 3) Cricqueville-en-Bessin (Basse-Normandy, Calvados), temporary excavation in meadow, Pliocene, yellowish sands with shells, collected by A. W. Janssen, September 1988. The fossiliferous locality of Cricqueville-en-Bessin was studied in detail by PAREYN et al. (1983).	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFCCFF9EFE93C0CBC54DCA4A.taxon	description	Terebratulida indet. sp. (not Terebratula grandis as indicated on the label), 4 indeterminable fragments, RGM. 29671	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFCEFF9BFE93C07FC439CC91.taxon	description	Delden Member: 3.0 – 3.5 m: Glottidia cf. dumortieri (Nyst, 1843),> 30 fragments, RGM. 793801 3.5 – 4.0 m: Glottidia cf. dumortieri (Nyst, 1843),> 30 fragments, RGM. 793802 4.0 – 4.5 m: Glottidia cf. dumortieri (Nyst, 1843),> 30 fragments, RGM. 793803 5.0 – 5.5 m: Glottidia cf. dumortieri (Nyst, 1843),> 30 fragments, RGM. 793804 5.5 – 6.0 m: Glottidia cf. dumortieri (Nyst, 1843),> 30 fragments, RGM. 793805 6.0 – 6.5 m: Glottidia cf. dumortieri (Nyst, 1843),> 30 fragments, RGM. 793806 6.5 – 7.0 m: Glottidia cf. dumortieri (Nyst, 1843),> 30 fragments, RGM. 793807 7.0 – 7.5 m: Glottidia? sp., 22 fragments, RGM. 793808 Zenderen Member: 7.5 – 8.0 m: Glottidia? sp.,> 30 fragments, RGM. 793809 8.0 – 8.5 m: Glottidia? sp., 5 fragments, RGM. 793810 8.5 – 9.0 m: Glottidia? sp., 5 fragments, RGM. 793811 9.0 – 9.5 m: Glottidia? sp.,> 30 fragments, RGM. 793812 9.5 – 10.0 m: Glottidia? sp., 12 fragments, RGM. 793813 10.0 – 10.5 m: Glottidia? sp., 6 fragments, RGM. 793814 10.5 – 11.0 m: Glottidia? sp., 7 fragments, RGM. 793815 12.0 – 12.5 m: Glottidia cf. dumortieri (Nyst, 1843),> 30 fragments, RGM. 793816 Neede (Gelderland), borehole Gelselaarsbrug (B 34 B 0175) (old name Neede II, 34 B. 3 – 1), 52.164278 N, 6.549397 E, Early Pliocene, Breda Formation, Delden and Zenderen Members, collected by M. van den Bosch, 30 October 1970. Borehole 34 B. 3 – 1 was described in detail and the section of the borehole was illustrated by VAN DEN BOSCH et al. (1975, Fig. 12); Lingula specimens and Lingula fragments were mentioned along the section. Delden Member: 32.5 – 33.5 m: Glottidia? sp., 6 fragments, RGM. 793769 33.5 – 34.5 m: Glottidia? sp., 8 fragments, RGM. 793770 34.5 – 35.5 m: Glottidia? sp.,> 10 fragments, RGM. 793771 35.5 – 36.5 m: Glottidia? sp., 1 fragment, RGM. 793772 36.5 – 37.5 m: Glottidia? sp., 4 fragments, RGM. 793773 37.5 – 38.5 m: Glottidia? sp.,> 10 fragments, RGM. 793774 38.5 – 39.5 m: Glottidia? sp.,> 10 fragments, RGM. 793775 39.5 – 40.5 m: Glottidia? sp.,> 10 fragments, RGM. 793776 40.5 – 41.5 m: Glottidia? sp.,> 10 fragments, RGM. 793777 41.5 – 42.5 m: Glottidia? sp.,> 10 fragments, RGM. 793778 42.5 – 43.5 m: Glottidia? sp.,> 10 fragments, RGM. 793779 43.5 – 45.5 m: Glottidia? sp.,> 10 fragments, RGM. 793780 45.5 – 46.5 m: Glottidia? sp.,> 10 fragments, RGM. 793781 46.5 – 47.5 m: Glottidia? sp.,> 10 fragments, RGM. 793782 47.5 – 48.5 m: Glottidia? sp., 3 fragments, RGM. 793783 48.5 – 49.5 m: Glottidia? sp., 2 fragments, RGM. 793784 Zenderen Member: 49.5 – 50.5 m: Glottidia? sp., 2 fragments, RGM. 793785 50.5 – 51.5 m: Glottidia? sp., 1 fragment, RGM. 793786 51.5 – 52.5 m: Glottidia? sp., 1 fragment, RGM. 793787 52.5 – 53.5 m: Glottidia? sp., 3 fragments, RGM. 793788 53.5 – 54.5 m: Glottidia? sp., 1 fragment, RGM. 793789 54.5 – 55.5 m: Glottidia? sp., 4 fragments, RGM. 793790 55.5 – 56.5 m: Glottidia? sp., 3 fragments, RGM. 793791 56.5 – 58.0 m: Glottidia? sp., 2 fragments, RGM. 793792 58.0 – 59.0 m: Glottidia? sp., 2 fragments, RGM. 793793 59.0 – 60.0 m: Glottidia? sp., 2 fragments, RGM. 793794 60.0 – 61.5 m: Glottidia? sp., 3 fragments, RGM. 793795 61.5 – 62.5 m: Glottidia? sp., 6 fragments, RGM. 793796 62.5 – 63.5 m: Glottidia? sp., 2 fragments, RGM. 793797 64.5 – 66.5 m: Glottidia? sp., 1 fragment, RGM. 793798 66.5 – 67.5 m: Glottidia? sp., 1 fragment, RGM. 793799 67.5 – 68.5 m: Glottidia? sp., 1 fragment, RGM. 793800	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFC9FF9BFE63C755C466CE78.taxon	materials_examined	Type species – Lingula albida Hinds, 1844	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFC8FF99FE58C07EC5DCCDC5.taxon	description	(Figs 5 – 9) 1843 Lingula Dumortieri – NYST, pp. 337 – 338, Pl. 34, Figs 4 a-c. 1872 Lingula Dumortieri Nyst – WOOD, pp. 172 – 173, Pl. 11, Figs 1 a-c. 1874 b Lingula Dumortieri Nyst – DAVIDSON, p. 153, Pl. 7, Figs 1 – 3. 1881 Lingula Dumortieri Nyst – NYST, p. 252, Pl. 28, Figs 4 a-c. 1893 Lingula Dumortieri Nyst – VINCENT, pp. 41 – 42. 1964 Glottidia dumortieri (Nyst) – CHUANG, pp. 155 – 157, Text-Fig. 1. 2004 Glottidia dumortieri (Nyst) – VOSKUIL, p. 47, Text-Fig. 1. 2013 Glottidia dumortieri (Nyst) – DULAI, pp. 25 – 26, Figs 6 – 14. Material – Gedgrave (159 fragmentary specimens); Delden (Delden Member:> 210 fragmentary specimens; Zenderen Member:> 30 fragmentary specimens). Remarks – Although their fossilization potential is rather low (EMIG 1990), remains of lingulid brachiopods are common in the Miocene and Pliocene deposits of the North Sea Basin. In several cases these are the dominant fossils, as all or most of the calcium carbonate shells dissolved from the sediments. Species dumortieri was described from the Pliocene of Belgium by NYST (1843) as Lingula. This generic assumption was accepted until the mid- 1960 s, when CHUANG (1964) recognised that this species should be assigned to the genus Glottidia. This classification was confirmed recently by DULAI (2013) on the basis of Miocene samples from the Netherlands (Beugen). Lingula sp. was mentioned several times even recently from the North Sea Basin, but up to now, the only confirmed lingulid brachiopod is Glottidia. Lingula is known from the Miocene of the Central Paratethys (EMIG & BITNER 2005), the Atlantic Ocean in France (EMIG et al. 2007) and from the Mediterranean (DREGER 1911). The studied samples contain rather fragmentary lingulids, but some of the specimens can be identified clearly as G. dumortieri. Lingulids are especially frequent in the two boreholes (Delden and Neede), which yielded samples from the Delden and Zenderen Members of the Breda Formation. The identifiable lingulid specimens are from the Delden Member of Twickel borehole at Delden (Gelderland), while the older Zenderen Member yielded very fragmentary lingulids which were not identifiable at species level. In the case of Gelselaarsbrug borehole at Neede (Gelderland) both members of the Breda Formation contain only very fragmentary lingulids. All of the Coralline Crag samples from Gedgrave contain more or less fragmentary lingulid specimens. Species dumortieri was mentioned from the Coralline Crag as early as WOOD (1872) and he realised that it is not rare at Sutton, but all specimens are mutilated. Glottidia? sp. Material – Gedgrave (24 fragments); Cricqueville-en-Bessin (2 fragments); Kreekrak (7 fragments); Delden (Delden Member: 22 fragments, Zenderen Member:> 95 fragments); Neede (Delden Member:> 124 fragments; Zenderen Member: 35 fragments). Remarks – Several samples from the Pliocene of England, France and the Netherlands yielded indeterminable Lingulidae fragments. The small size of the fragments makes it impossible to decide whether they belong to Lingula or Glottidia. Until now, only Glottidia has surely been confirmed from the Neogene of the North Sea Basin (e. g. CHUANG 1964; DULAI 2013), therefore, Glottidia is used for these fragments with question mark.	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFCAFF87FE62C0B8C4E7CAED.taxon	description	(Figs 10 – 13) 1944 Hemithiris bipartita (Brocchi) – MEZNERICS, p. 22, Pl. 5, Figs 12, 16. 1944 Hemithiris acuta n. sp. – MEZNERICS, pp. 22 – 23, Pl. 3, Figs 7 – 10. 1985 a Aphelesia bipartita (Brocchi) – GAETANI & SACCÀ, p. 5, Text-Fig. 2, Pl. 7, Figs 1 – 4 (cum. syn.). 1985 b Aphelesia bipartita (Brocchi) – GAETANI & SACCÀ, pp. 363 – 365, Text-Figs 2 – 3, Pl. 17, Figs 1 – 3, Pl. 19, Figs 1 – 3. 2001 Aphelesia bipartita (Brocchi) – BORGHI, pp. 49 – 50, Pl. 2, Figs 1 – 5. 2001 Aphelesia bipartita (Brocchi) – BITNER & MARTINELL, pp. 179 – 181, Figs 3 A-H. 2003 Aphelesia bipartita (Brocchi) – BITNER & MOISSETTE, p. 466, Figs 2 A-M. 2005 Aphelesia bipartita (Brocchi) – GARCÍA RAMOS, pp. 28 – 29, Pl. 1, Figs 9, 11, Pl. 2, Figs 5, 7. Material – Ceriale (2 complete specimens, 1 pedicle and 1 brachial valve). Remarks – The few studied specimens from Ceriale are easily recognizable as A. bipartita. This species was described in detail by COOPER (1959), GAETANI & SACCÀ (1985 a, b), BITNER & MARTINELL (2001) and BITNER & MOISSETTE (2003). It is a very significant component of the Terebratula - Aphelesia Bed in Malta (PEDLEY 1976; DULAI et al. in prep.).	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFCAFF98FD85C000C4A2C997.taxon	materials_examined	Type species – Anomia bipartita Brocchi, 1814	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFD5FF87FE55C24AC4B1CB5D.taxon	materials_examined	Type species – Terebratula nigricans Sowerby, 1846	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFD5FF86FE4AC2E2C35DCB71.taxon	description	(Figs 14 – 18) 1874 b Rhynchonella Nysti n. sp. – DAVIDSON, p. 157, Pl. 7, Fig. 17. 1881 Rhynchonella Nysti Davidson – NYST, p. 250, Pl. 28, Figs 2 a-c. 1927 Tegulorhynchia nysti (Davidson) – THOMSON, p. 153. 1959 Notosaria nysti (Davidson) – COOPER, p. 49. 1979 Notosaria nysti (Davidson) – LEE & WILSON, p. 439. 1987 Tegulorhynchia nysti (Davidson) – OTTEMA & IN’T HOUT, p. 76, Fig. 4. Material – Kreekrak (1 complete specimen, 3 brachial valves; 6 fragments). Remarks – N. nysti was originally described by DAVIDSON (1874 b) from the “ Scaldisien ” (= Pliocene, Late Zanclean to Piacenzian; LAGA & LOUWYE 2006) of Antwerp, Belgium. Unfortunately, DAVIDSON (1874 b) had very limited material and illustrated only the ventral valve, but mentioned its close similarity to Rh. nigricans. CHAPMAN & CRESPIN (1923) erected a new genus Tegulorhynchia for ribbed rhynchonellids of the southern hemisphere, and some years later THOMSON (1927) attributed species nysti to Tegulorhynchia. COOPER (1959) described a new genus, Notosaria with type species Terebratula nigricans, and distinguished his new genus from Tegulorhynchia on the basis of ornamentation, beak characters and cardinalia. Cooper also studied a pedicle and a brachial valve of Rh. nysti confirming DAVIDSON’s (1874 b) original observation and included this species also to his new genus, Notosaria. Later LEE & WILSON (1979) accepted and followed this opinion. The new, revised Treatise also mentioned Notosaria from Europe (Middle Miocene of Poland and Pliocene of Belgium) (MANCEÑIDO et al. 2002) and erected the new family Notosariidae. The only Miocene record of Notosaria from Europe is an eroded and very uncertain pedicle valve from Poland (POPIEL-BARCZYK & BARCZYK 1990). This species has not been illustrated since DAVIDSON’s (1874 b) original description (his figures were re-published by NYST 1881), although it is not rare in some Neogene assemblages. OTTEMA & IN’T HOUT (1987) probably also redrew DAVIDSON’s (1874 b) pedicle valve illustrations, which is rather strange, as this species is common in the studied Kallo assemblage. VINCENT (1893) mentioned species nysti from Belgium, while DOLFUSS & DAUTZENBERG (1901) and DE MORGAN (1915) reported it from the Miocene of France (Savigné, Saint- Saturnin, Saint-Emy) without illustrations. In the studied material this species occurs in the Kreekraksluizen sample only with some valves, but it is common in some Pliocene (“ Redonian ”) samples of France (e. g. St. Clément-de-la Place) or in the Pliocene of Belgium (e. g. Kallo). The detailed description of both of these faunas is in progress and emended diagnosis of N. nysti will be given there.	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFD7FF85FE4FC0CEC4BBC9DE.taxon	materials_examined	Type species – Anomia terebratula Linnaeus, 1758	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFD7FF83FE2AC366C337CA81.taxon	description	(Figs 19 – 26, 27 – 34) 1983 Terebratula ampulla (Brocchi) – COOPER, Pl. 4, Figs 8 – 16. 2001 Terebratula ampulla (Brocchi) – BORGHI, pp. 51 – 52, Pl. 3, Figs 2, 3, 4, 6, 7, Pl. 4, Fig. 1. 2004 Terebratula ampulla (Brocchi) – GARCÍA RAMOS, pp. 21 – 23, Pl. 5, Figs 3, 7, 8, 9, 10, Pl. 6, Fig. 3, Pl. 7, Figs 3, 4, 5, 6, 7, 9. 2008 Terebratula ampulla (Brocchi) – TADDEI RUGGIERO et al., p. 211, Figs 1 H-M. Material – Valle Andona (1 complete specimen, 4 pedicle valves); Lugagnano (4 pedicle valves); San Nicomede (7 pedicle and 8 brachial valves); Ceriale (3 complete specimens, 9 pedicle and 9 brachial valves); Apricena (2 pedicle valves and 1 brachial valve). Remarks – Terebratula and its type species T. terebratula were revised and discussed in detail by LEE et al. (2001). In this paper several species were synonymized with T. terebratula, however, the validity of T. ampulla as a medium-sized, strongly bisulcate species from the Pliocene was confirmed. TADDEI RUGGIERO et al. (2008) have accomplished a morphometric study on Late Cenozoic Terebratula from Italy, in which T. terebratula, T. ampulla, T. scillae and T. sinuosa were compared. According to their results, T. ampulla is distinct from T. scillae / T. terebratula in anterior view, because it is much more compressed dorsoventrally. In dorsal view T. ampulla is readily distinguished because of its pentagonal shape (TADDEI RUGGIERO et al. 2008). T. ampulla is widespread in the Pliocene of the Mediterranean (e. g. BORGHI 2001; GARCÍA RAMOS 2004; TADDEI RUGGIERO et al. 2008). Some papers (e. g. BOSSELAERS et al. 2004) mentioned this species also from the Neogene of the North Sea Basin, but until now, these records have not been confirmed. The beak area of several San Nicomede pedicle valves is strongly eroded and the foramen is significantly widened. In the case of Ceriale samples, all complete specimens are strongly compressed, the separated pedicle valves are intact but sometimes fragmentary. The middle part of the smallest complete specimen and edge of two pedicle valves show small-sized traces of predatory organisms. A small worm tube encrustation can be seen on the internal surface of a pedicle valve.	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFD1FF83FE41C3A0C4BBCB37.taxon	materials_examined	Type species – Terebratula sowerbyana Nyst, 1843	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFD1FF81FE35C258C6A3CC46.taxon	description	(Figs 35 – 48) 1843 Terebratula Sowerbyana – NYST, pp. 335 – 336, Pl. 27, Figs 3 a-b. 1980 Pliothyrina sowerbyana (Nyst) – VAN ROY, pp. 3 – 7, Pl. 1, Figs 1 – 7, Pl. 2, Figs 1 – 7. 1983 Pliothyrina sowerbyana (Nyst) – COOPER, pp. 237 – 238, Pl. 5, Figs 1 – 4. 2004 Pliothyrina sowerbyana (Nyst) – VOSKUIL, pp. 50 – 53, Text-Figs 7 A-L. Material – Gedgrave (3 complete specimens, 50 pedicle and 50 brachial valves); Brightwell (1 brachial valve); Cricqueville-en-Bessin (1 complete specimen, 6 pedicle and 5 brachial valves); Goes (2 pedicle valves and 1 brachial valve, 6 fragments). Remarks – Large-sized Miocene and Pliocene terebratulids have a long and complex nomenclatural history in the North Sea Basin. Several names were introduced in the literature both in the eastern (Belgium – the Netherlands) and in the western (Great Britain) part of the Basin. It seems that nearly all, or at least most of the Oligocene-Pliocene large terebratulids belong to the genus Pliothyrina described by VAN ROY (1980) with type species P. sowerbyana. LACOURT (1983) identified 30 names for Terebratulidae species from the Neogene of the Netherlands (mentioning even some Mesozoic species). Later VOSKUIL (2004) revised LACOURT’s (1983) work, and correctly synonymized nearly all of his terebratulids with P. sowerbyana. The only exception was “ Terebratula ” distinguenda Lacourt, 1984, which was separated with question mark also by VOSKUIL (2004). On the basis of some Dutch private collections (Freddie van Nieulande, Peter Mordijk, Harry Raad) this separate species with very small pedicle opening really exists, and its internal morphology is also significantly different from Pliothyrina, and refers to a separate, new genus. This new taxon hopefully will be described in the near future, in the framework of the cooperation with the above mentioned private collectors. After VAN ROY’s (1980) description, the name Pliothyrina sowerbyana became widely accepted and well-known, and several papers used this name for large terebratulids in the North Sea Basin (e. g. VOSKUIL 2004; MOERDIJK 2007; RAAD 2004 a, b, 2008; VAN NIEULANDE 2009). However, recently MOERDIJK (2016) discussed in detail the nomenclatural problems of Neogene Pliothyrina assemblages of the North Sea Basin. The correct name of the very common and wide-spread form is under discussion, and until the final decision the well-known name P. sowerbyana is used in this paper. HERMAN in BOSSELAERS et al. (2004) indicated Terebratula ampulla together with P. sowerbyana and Lingula sp. (= Glottidia) in the Late Miocene of Belgium, illustrated by the dorsal view of a fragmentary specimen (p. 32, Fig. 5 c). VAN NIEULANDE (2009) also mentioned P. sowerbyana and Terebratula sp. from the “ seashore strands ”. However, until now, the presence of Terebratula or specifically T. ampulla has not been confirmed in the North Sea Basin; it is a typical Mediterranean taxon. Pliothyrina? sp. Material – Gedgrave (177 fragments); Goes (1 fragment). Remarks – The Coralline Crag samples from Gedgrave contain several small-sized, unidentifiable smooth Terebratulidae fragments. As Pliothyrina sowerbyana specimens are common in these beds, these fragments are regarded as Pliothyrina sp. with question mark.	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFD3FF81FE2FC582C497CD12.taxon	materials_examined	Type species – Anomia retusa Linnaeus, 1758	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFD2FF8FFE50C07EC330CA8E.taxon	description	(Figs 49 – 54) 1852 Terebratulina caput-serpentis Linnaeus – DAVIDSON, pp. 12 – 14, Pl. 1, Figs 3 – 6, 14 – 15. 1979 Terebratulina retusa (Linnaeus) – BRUNTON & CURRY, p. 38, Text-Figs 17 A-C. 1985 a Terebratulina retusa (Linnaeus) – GAETANI & SACCÀ, pp. 15 – 16, Pl. 7, Figs 5 – 10, Pl. 9, Figs 6 – 9 (cum syn.). 2001 Terebratulina retusa (Linnaeus) – BORGHI, p. 52, Pl. 4, Figs 4, 5, 6, 7, 8. 2003 Terebratulina retusa (Linnaeus) – BITNER & MOISSETTE, p. 472, Figs 6 A-F. 2004 Terebratulina retusa (Linnaeus) – VOSKUIL, p. 49, Text-Figs 4 A-G. 2004 Terebratulina retusa (Linnaeus) – GARCÍA RAMOS, p. 28, Pl. 1, Figs 1 – 3. 2007 Terebratulina retusa (Linnaeus) – KOSKERIDOU, pp. 124 – 125, Pl. 1, Figs 5 – 6. Material – Apricena (1 complete specimen); Kreekrak (1 complete specimen, 1 brachial valve). Remarks – T. retusa is a common member of Neogene and Recent benthic assemblages, however, much more frequent in deeper water environments. Depth range of Recent T. retusa is very wide (18 – 2157 m; LOGAN 2007), but it is the most common between 100 and 500 m (CURRY 1982). According to LOGAN (1979) and LOGAN et al. (2004) Terebratulina (together with Gryphus, Platidia and Megerlia) belongs to the eurybathic species which are more typical of the bathyal zone in the Recent Mediterranean. Probably this is the reason that this species is very rare in the studied samples, both in the Mediterranean and the North Sea Basin. T. retusa is consistently cited from the Neogene of the Mediterranean (e. g. GAETANI & SACCÀ 1985 a; TADDEI RUGGIERO 1994; BORGHI 2001; BITNER & MOISSETTE 2003; GARCÍA RAMOS 2004; KOSKERIDOU 2007) and known from the Central Paratethys (e. g. BITNER & DULAI 2004). More rarely it was also mentioned from the North Sea Basin Neogene (DAVIDSON 1852; WOOD 1872) and Recent (BRUNTON & CURRY 1979; CURRY 1982) assemblages, sometimes under the name T. caputserpentis. The complex nomenclatural history of the retusa – caputserpentis problem was discussed recently in detail by EMIG et al. (2015).	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFDDFF8EFE29C2CDC324CD96.taxon	description	(Figs 55 – 60) 1886 Waldheimia (Macandrevia) cranium (Müller) – DAVIDSON, pp. 61 – 66, Pl. 12, Figs 11 – 23, Pl. 13, Figs 1 – 2. 1979 Macandrevia cranium (Müller) – BRUNTON & CURRY, p. 58, Figs 30 A-C. 1990 Macandrevia cranium (Müller) – THOMSEN, p. 25, Text-Fig. 1. 2001 Macandrevia cranium (Müller) – THOMSEN, pp. 121 – 123, Text-Fig. 2004 Macandrevia cranium (Müller) – VOSKUIL, pp. 49 – 50, Text-Figs 5 A-G. 2012 Macandrevia cranium (Müller) – ZEZINA, p. 86. Material – Gedgrave (2 complete specimens, 3 pedicle valves); Cricqueville-en-Bessin (2 pedicle valves); Kreekrak (1 pedicle valve, 1 brachial valve). Remarks – Today Macandrevia cranium is known mainly from the Arctic region (Western Arctic, Greenland, Norway, Northern Atlantic, Spitzbergen, Greenland, Rhode Island, and Western Sahara) but may occur also in the Mediterranean (Lion Bay, France) (ZEZINA 2012). M. cranium has a long nomenclatural history, and earlier it was attributed to different genera: Terebratula (MÜLLER 1776; JEFFREYS 1878), Waldheimia (DAVIDSON 1886; FISCHER & OEHLERT 1891; POSSELT 1898); Waldheimiathyris (WESENBERG-LUND 1941). DALL (1920) described a new species, M. novangliae which is generally synonymized with M. cranium (e. g. ZEZINA 2012). THOMSEN (1990) studied M. cranium and its usefulness in palaeoceanographic reconstructions. It belongs to the deep-water forms, the recent representatives are known from 9 – 2951 m (ZEZINA 2012). In the studied North Sea Basin samples it consistently occurs in small specimen numbers.	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFDDFF8FFE4FC215C483CBB8.taxon	materials_examined	Type species – Terebratula cranium Müller, 1776	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFDCFF8EFD86C757C4DCCE46.taxon	materials_examined	Type species – Terebratula cuneata Risso, 1826	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
03F3861CFFDFFF8CFE11C07EC343C961.taxon	description	(Figs 61 – 64) Material – Cricqueville-en-Bessin (1 pedicle valve). Remarks – Unfortunately, the available material is very limited, contains only a pedicle valve. It is very similar to DE MORGAN’s (1915) species, which was described from the Middle Miocene (Langhian) of Pontlevoy as a rare new species. This species was also listed by PACAUD (2014) among brachiopod type specimens of the Natural History Museum in Paris, without any additional information. The main difference between them that de Morgan’s specimen has rounded outline, while the Cricqueville-en-Bessin specimen is more subtrigonal. The other characters are the same (few weak and rounded ribs, lack of tubercles along the internal margin of the valve, short and low median septum in the pedicle valve). Reasonable interpretation of this form requires much more available material.	en	Dulai, Alfréd (2016): Sporadic Pliocene and Pleistocene brachiopods in Naturalis Biodiversity Center (Leiden, the Netherlands): Records from the Mediterranean, and the North Sea Basin. Fragmenta Palaeontologica Hungarica 33: 65-98, DOI: 10.17111/FragmPalHung.2016.33.65, URL: https://doi.org/10.17111/fragmpalhung.2016.33.65
