taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507247/files/figure.png	https://doi.org/10.5281/zenodo.18507247	Fig. 16. Rheocricotopus (P.) binotatus sp. nov. A, C, G. ♂ (JNU 23020507). B, D–F. Holotype, ♂ (JNU 170723009). A–B. Abdominal colour pattern, showing variation. C. Humeral pit. D. Wing. E. Setal arrangement on tergites II–IV. F. Hypopygium, dorsal view. G. Gonostylus.	Fig. 16. Rheocricotopus (P.) binotatus sp. nov. A, C, G. ♂ (JNU 23020507). B, D–F. Holotype, ♂ (JNU 170723009). A–B. Abdominal colour pattern, showing variation. C. Humeral pit. D. Wing. E. Setal arrangement on tergites II–IV. F. Hypopygium, dorsal view. G. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507266/files/figure.png	https://doi.org/10.5281/zenodo.18507266	Fig. 22. Rheocricotopus (P.) nippobifasciatus sp. nov. A–D. Holotype, ♂ (NSMT-I-Dip 36256). E. Paratype, ♂ (NSMT-I-Dip 36262). F–G. Paratype, ♀ (NSMT-I-Dip 36269). A. Abdominal colour pattern. B. Thorax with humeral pit, lateral view. C. Wing. D. Hypopygium, dorsal view. E. Gonostylus. F. Genitalia, ventral view. G. Genitalia, dorsal view, showing tergite IX.	Fig. 22. Rheocricotopus (P.) nippobifasciatus sp. nov. A–D. Holotype, ♂ (NSMT-I-Dip 36256). E. Paratype, ♂ (NSMT-I-Dip 36262). F–G. Paratype, ♀ (NSMT-I-Dip 36269). A. Abdominal colour pattern. B. Thorax with humeral pit, lateral view. C. Wing. D. Hypopygium, dorsal view. E. Gonostylus. F. Genitalia, ventral view. G. Genitalia, dorsal view, showing tergite IX.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507274/files/figure.png	https://doi.org/10.5281/zenodo.18507274	Fig. 24. Rheocricotopus (P.) yunnanensis sp. nov. A–C, E. Holotype, ♂ (JNU 170723007). D, F. Paratype, ♂ (JNU 170614009). G. Paratype, ♂ (JNU 17061406). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Setal arrangement on tergites II–IV. E. Hypopygium, dorsal view. F. Superior volsella. G. Gonostylus.	Fig. 24. Rheocricotopus (P.) yunnanensis sp. nov. A–C, E. Holotype, ♂ (JNU 170723007). D, F. Paratype, ♂ (JNU 170614009). G. Paratype, ♂ (JNU 17061406). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Setal arrangement on tergites II–IV. E. Hypopygium, dorsal view. F. Superior volsella. G. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507231/files/figure.png	https://doi.org/10.5281/zenodo.18507231	Fig. 11. Rheocricotopus (P.) valgus Chaudhuri & Sinharay, 1983. A–H. ♂ (JNU 170208001). I. Pupa (JNU 170308002). A. Abdominal colour pattern. B. Colour pattern of foreleg. C. Colour pattern of mid leg. D. Wing. E. Hypopygium, dorsal view. F–H. Gonostylus, showing variation. I. Abdomen, dorsal view.	Fig. 11. Rheocricotopus (P.) valgus Chaudhuri & Sinharay, 1983. A–H. ♂ (JNU 170208001). I. Pupa (JNU 170308002). A. Abdominal colour pattern. B. Colour pattern of foreleg. C. Colour pattern of mid leg. D. Wing. E. Hypopygium, dorsal view. F–H. Gonostylus, showing variation. I. Abdomen, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507237/files/figure.png	https://doi.org/10.5281/zenodo.18507237	Fig. 13. Rheocricotopus (P.) bannaensis sp. nov. A–D. Holotype, ♂ (JNU 170429003). E–G. Paratype, ♀ (JNU). A. Abdominal colour pattern. B. Thorax with humeral pit, lateral view. C. Wing. D. Hypopygium, dorsal view. E. Thorax, lateral view (arrow showing humeral pit). F. Genitalia, ventral view. G. Genitalia, dorsal view, showing tergite IX.	Fig. 13. Rheocricotopus (P.) bannaensis sp. nov. A–D. Holotype, ♂ (JNU 170429003). E–G. Paratype, ♀ (JNU). A. Abdominal colour pattern. B. Thorax with humeral pit, lateral view. C. Wing. D. Hypopygium, dorsal view. E. Thorax, lateral view (arrow showing humeral pit). F. Genitalia, ventral view. G. Genitalia, dorsal view, showing tergite IX.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507262/files/figure.png	https://doi.org/10.5281/zenodo.18507262	Fig. 21. Rheocricotopus (P.) monovittatus sp. nov. A–D. Holotype, ♂ (JNU 23032701). E. Paratype, ♂ (JNU 24033120). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Hypopygium, dorsal view. E. Gonostylus.	Fig. 21. Rheocricotopus (P.) monovittatus sp. nov. A–D. Holotype, ♂ (JNU 23032701). E. Paratype, ♂ (JNU 24033120). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Hypopygium, dorsal view. E. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507209/files/figure.png	https://doi.org/10.5281/zenodo.18507209	Fig. 2. Rheocricotopus (P.) bifasciatus Wang & Zheng, 1991. A, F, H. Holotype, ♂(NKU). B–E, G, I. ♂ (JNU). A–B. Abdominal colour pattern, showing variation. C. Colour pattern of foreleg. D. Colour pattern of mid leg. E. Humeral pit. F. Wing. G. Setal arrangement on tergites II–IV. H. Hypopygium, dorsal view. I. Gonostylus.	Fig. 2. Rheocricotopus (P.) bifasciatus Wang & Zheng, 1991. A, F, H. Holotype, ♂(NKU). B–E, G, I. ♂ (JNU). A–B. Abdominal colour pattern, showing variation. C. Colour pattern of foreleg. D. Colour pattern of mid leg. E. Humeral pit. F. Wing. G. Setal arrangement on tergites II–IV. H. Hypopygium, dorsal view. I. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507249/files/figure.png	https://doi.org/10.5281/zenodo.18507249	Fig. 17. Rheocricotopus (P.) chinogodavarius sp. nov. A, D–H. Holotype, ♂ (JNU 170724006). B–C, I–J. Paratype, ♂ (JNU). A–C. Abdominal colour pattern, showing variation. D. Colour pattern of foreleg. E. Colour pattern of mid leg. F. Humeral pit. G. Wing. H. Hypopygium, dorsal view. I. Inferior volsella, showing variation. J. Gonostylus, showing variation.	Fig. 17. Rheocricotopus (P.) chinogodavarius sp. nov. A, D–H. Holotype, ♂ (JNU 170724006). B–C, I–J. Paratype, ♂ (JNU). A–C. Abdominal colour pattern, showing variation. D. Colour pattern of foreleg. E. Colour pattern of mid leg. F. Humeral pit. G. Wing. H. Hypopygium, dorsal view. I. Inferior volsella, showing variation. J. Gonostylus, showing variation.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507219/files/figure.png	https://doi.org/10.5281/zenodo.18507219	Fig. 6. Rheocricotopus (P.) oiraprimus Sasa, 1991. A–E. ♂ (PCHN). F. Holotype, ♂ (NSMT-I-Dip 5989). A–B. Abdominal colour pattern, showing variation. C. Thorax with humeral pit, lateral view. D. Wing. E. Hypopygium, dorsal view. F. Gonostylus.	Fig. 6. Rheocricotopus (P.) oiraprimus Sasa, 1991. A–E. ♂ (PCHN). F. Holotype, ♂ (NSMT-I-Dip 5989). A–B. Abdominal colour pattern, showing variation. C. Thorax with humeral pit, lateral view. D. Wing. E. Hypopygium, dorsal view. F. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507241/files/figure.png	https://doi.org/10.5281/zenodo.18507241	Fig. 14. Rheocricotopus (P.) albulus sp. nov. A. ♂ (JNU). B–G. Holotype, ♂ (JNU 170512011). H–I. Paratype, pupa (JNU). A–B. Abdominal colour pattern, showing variation. C. Foreleg colour pattern. D. Humeral pit. E. Wing. F. Setal arrangement on tergites II–IV. G. Hypopygium, dorsal view. H. Frontal apotome. I. Abdomen, dorsal view.	Fig. 14. Rheocricotopus (P.) albulus sp. nov. A. ♂ (JNU). B–G. Holotype, ♂ (JNU 170512011). H–I. Paratype, pupa (JNU). A–B. Abdominal colour pattern, showing variation. C. Foreleg colour pattern. D. Humeral pit. E. Wing. F. Setal arrangement on tergites II–IV. G. Hypopygium, dorsal view. H. Frontal apotome. I. Abdomen, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507251/files/figure.png	https://doi.org/10.5281/zenodo.18507251	Fig. 18. Rheocricotopus (P.) fasciolatus sp. nov., holotype, ♂ (JNU 170509002). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Setal arrangement on tergites II–IV. E. Hypopygium, dorsal view.	Fig. 18. Rheocricotopus (P.) fasciolatus sp. nov., holotype, ♂ (JNU 170509002). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Setal arrangement on tergites II–IV. E. Hypopygium, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507233/files/figure.png	https://doi.org/10.5281/zenodo.18507233	Fig. 12. A. Rheocricotopus (P.) kongi Lin & Wang, 2020, ♂ (NKU, XL-1853). B–J. R. (P.) orientalis Wang, 1995. B–E. ♂(JNU161009005). F–G. ♀ (JNU 24040903). H–J. Pupa (JNU 24041002). A. Hypopygium, dorsal view (two red arrows showing ventral and inner projections). B. Abdominal colour pattern. C. Thorax, lateral view (arrow showing humeral pit). D. Wing. E. Hypopygium, dorsal view. F. Genitalia, ventral view. G. Genitalia, dorsal view. H. Frontal apotome. I. Thoracic horn. J. Abdomen, dorsal view.	Fig. 12. A. Rheocricotopus (P.) kongi Lin & Wang, 2020, ♂ (NKU, XL-1853). B–J. R. (P.) orientalis Wang, 1995. B–E. ♂(JNU161009005). F–G. ♀ (JNU 24040903). H–J. Pupa (JNU 24041002). A. Hypopygium, dorsal view (two red arrows showing ventral and inner projections). B. Abdominal colour pattern. C. Thorax, lateral view (arrow showing humeral pit). D. Wing. E. Hypopygium, dorsal view. F. Genitalia, ventral view. G. Genitalia, dorsal view. H. Frontal apotome. I. Thoracic horn. J. Abdomen, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507257/files/figure.png	https://doi.org/10.5281/zenodo.18507257	Fig. 19. Rheocricotopus (P.) hanmiensis sp. nov., holotype, ♂ (JNU 251215A1). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Hypopygium, dorsal view. E. Gonostylus.	Fig. 19. Rheocricotopus (P.) hanmiensis sp. nov., holotype, ♂ (JNU 251215A1). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Hypopygium, dorsal view. E. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507223/files/figure.png	https://doi.org/10.5281/zenodo.18507223	Fig. 8. Rheocricotopus (P.) togakuroasi (Sasa & Okazawa, 1992). A–B, D–F. ♂ (PCHN). C. Holotype, ♂ (NSMT-I-Dip 4757). G. ♀ (PCHN). A. Thorax with humeral pit, lateral view. B. Wing. C. Tergite IV, showing setal arrangement. D. Hypopygium, dorsal view. E. Superior volsella. F. Gonostylus, showing variation. G. Genitalia, dorsal view, showing tergite IX.	Fig. 8. Rheocricotopus (P.) togakuroasi (Sasa & Okazawa, 1992). A–B, D–F. ♂ (PCHN). C. Holotype, ♂ (NSMT-I-Dip 4757). G. ♀ (PCHN). A. Thorax with humeral pit, lateral view. B. Wing. C. Tergite IV, showing setal arrangement. D. Hypopygium, dorsal view. E. Superior volsella. F. Gonostylus, showing variation. G. Genitalia, dorsal view, showing tergite IX.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507211/files/figure.png	https://doi.org/10.5281/zenodo.18507211	Fig. 3. Rheocricotopus (P.) chalybeatus (Edwards, 1929). A. ♂ (JNU 2402). B–H. ♂ (PCHN). I–K. ♀ (PCHN). A. Abdominal colour variation, dorsal view. B. Thorax with humeral pit, lateral view. C. Wing. D. Hypopygium, dorsal view. E. Anal point. F. Inferior volsella, showing variation. G–H. Gonostylus, showing variation. I. Genitalis, ventral view. J. Genitalis, dorsal view. K. Tergite IX, showing variation.	Fig. 3. Rheocricotopus (P.) chalybeatus (Edwards, 1929). A. ♂ (JNU 2402). B–H. ♂ (PCHN). I–K. ♀ (PCHN). A. Abdominal colour variation, dorsal view. B. Thorax with humeral pit, lateral view. C. Wing. D. Hypopygium, dorsal view. E. Anal point. F. Inferior volsella, showing variation. G–H. Gonostylus, showing variation. I. Genitalis, ventral view. J. Genitalis, dorsal view. K. Tergite IX, showing variation.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507268/files/figure.png	https://doi.org/10.5281/zenodo.18507268	Fig. 23. Rheocricotopus (P.) protuberans sp. nov., holotype, ♂ (JNU 160206012). A. Thorax with humeral pit, lateral view. B. Wing. C. Hypopygium, dorsal view.	Fig. 23. Rheocricotopus (P.) protuberans sp. nov., holotype, ♂ (JNU 160206012). A. Thorax with humeral pit, lateral view. B. Wing. C. Hypopygium, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507205/files/figure.png	https://doi.org/10.5281/zenodo.18507205	Fig. 1. Rheocricotopus (P.) amamipubescius (Sasa, 1990). A–C. ♂ (PCHN). D. Holotype, ♂ (NSMT- I-Dip 4690). E. ♂ (JNU 21081601). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D–E. Gonostylus, showing variation.	Fig. 1. Rheocricotopus (P.) amamipubescius (Sasa, 1990). A–C. ♂ (PCHN). D. Holotype, ♂ (NSMT- I-Dip 4690). E. ♂ (JNU 21081601). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D–E. Gonostylus, showing variation.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507243/files/figure.png	https://doi.org/10.5281/zenodo.18507243	Fig. 15. Rheocricotopus (P.) beibengensis sp. nov., holotype, ♂ (JNU 25121501). A. Wing. B. Humeral pit. C. Hypopygium, dorsal view.	Fig. 15. Rheocricotopus (P.) beibengensis sp. nov., holotype, ♂ (JNU 25121501). A. Wing. B. Humeral pit. C. Hypopygium, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507221/files/figure.png	https://doi.org/10.5281/zenodo.18507221	Fig. 7. Rheocricotopus (P.) serratus Liu, Lin & Wang, 2014, ♂ (JNU 181210050). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D. Gonostylus.	Fig. 7. Rheocricotopus (P.) serratus Liu, Lin & Wang, 2014, ♂ (JNU 181210050). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507227/files/figure.png	https://doi.org/10.5281/zenodo.18507227	Fig. 10. Rheocricotopus (P.) tokarakeleus Sasa & Suzuki, 1995, holotype, ♂ (NSMT I-Dip-5003). A. Humeral pit. B. Wing. C. Setal arrangement on tergite IV. D. Hypopygium, dorsal view. E. Gonostylus.	Fig. 10. Rheocricotopus (P.) tokarakeleus Sasa & Suzuki, 1995, holotype, ♂ (NSMT I-Dip-5003). A. Humeral pit. B. Wing. C. Setal arrangement on tergite IV. D. Hypopygium, dorsal view. E. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507217/files/figure.png	https://doi.org/10.5281/zenodo.18507217	Fig. 5. Rheocricotopus (P.) hidakadeeus Sasa & Suzuki, 2001, holotype, ♂ (NSMT-I-Dip 5367). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view.	Fig. 5. Rheocricotopus (P.) hidakadeeus Sasa & Suzuki, 2001, holotype, ♂ (NSMT-I-Dip 5367). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507260/files/figure.png	https://doi.org/10.5281/zenodo.18507260	Fig. 20. Rheocricotopus (P.) masarui sp. nov. A–B, E–H. Holotype, ♂ (JNU 211013H03B). C–D, I–J. Paratype, ♂ (JNU 22082001). K–N. Pupa (JNU 22082001). A–C. Abdominal colour pattern, showing variation. D. Humeral pit. E. Wing. F. Setal arrangement on tergite II. G. Setal arrangement on tergite IV. H. Hypopygium, dorsal view. I. Anal point. J. Gonostylus. K. Frontal apotome. L. Thoracic horn. M. Abdomen, dorsal view. N. Abdomen, ventral view.	Fig. 20. Rheocricotopus (P.) masarui sp. nov. A–B, E–H. Holotype, ♂ (JNU 211013H03B). C–D, I–J. Paratype, ♂ (JNU 22082001). K–N. Pupa (JNU 22082001). A–C. Abdominal colour pattern, showing variation. D. Humeral pit. E. Wing. F. Setal arrangement on tergite II. G. Setal arrangement on tergite IV. H. Hypopygium, dorsal view. I. Anal point. J. Gonostylus. K. Frontal apotome. L. Thoracic horn. M. Abdomen, dorsal view. N. Abdomen, ventral view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507284/files/figure.png	https://doi.org/10.5281/zenodo.18507284	Fig. 27. Rheocricotopus (s. str.) tatequintus Sasa, 1996. A–C. Holotype, ♂ (NSMT I-Dip-4998). D–F. Paratype, ♂ (NSMT No. 284: 59). G–H. Paratype, ♀ (NSMT No. 284: 61). A, G. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D. Superior volsella. E. Inferior volsella. F. Gonostylus. H. Genitalia, ventral view.	Fig. 27. Rheocricotopus (s. str.) tatequintus Sasa, 1996. A–C. Holotype, ♂ (NSMT I-Dip-4998). D–F. Paratype, ♂ (NSMT No. 284: 59). G–H. Paratype, ♀ (NSMT No. 284: 61). A, G. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D. Superior volsella. E. Inferior volsella. F. Gonostylus. H. Genitalia, ventral view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507286/files/figure.png	https://doi.org/10.5281/zenodo.18507286	Fig. 28. Rheocricotopus (s. str.) togapeniculus Sasa & Okazawa, 1992. A–E. ♂ (PCHN). F–G. ♀ (PCHN). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D. Superior volsella, showing variation. E. Gonostylus, showing variation. F. Genitalia, ventral view. G. Genitalia, dorsal view.	Fig. 28. Rheocricotopus (s. str.) togapeniculus Sasa & Okazawa, 1992. A–E. ♂ (PCHN). F–G. ♀ (PCHN). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D. Superior volsella, showing variation. E. Gonostylus, showing variation. F. Genitalia, ventral view. G. Genitalia, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE2FF89FDF2FC8DCB4ACC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507276/files/figure.png	https://doi.org/10.5281/zenodo.18507276	Fig. 25. Rheocricotopus (s. str.) tamahumeralis Sasa, 1981. A–N. ♂ (PCHN). O–P. ♀ (PCHN). A. Thorax with humeral pit, lateral view. B–D. Humeral pit, showing variation. E. Wing. F. Hypopygium, dorsal view. G–H. Superior volsella, showing variation. I–J. Inferior volsella, showing variation. K–N. Gonostylus, showing variation. O. Genitalia, ventral view. P. Genitalia, dorsal view.	Fig. 25. Rheocricotopus (s. str.) tamahumeralis Sasa, 1981. A–N. ♂ (PCHN). O–P. ♀ (PCHN). A. Thorax with humeral pit, lateral view. B–D. Humeral pit, showing variation. E. Wing. F. Hypopygium, dorsal view. G–H. Superior volsella, showing variation. I–J. Inferior volsella, showing variation. K–N. Gonostylus, showing variation. O. Genitalia, ventral view. P. Genitalia, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFE9FF85FE25FCF2CF12CC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507205/files/figure.png	https://doi.org/10.5281/zenodo.18507205	Fig. 1. Rheocricotopus (P.) amamipubescius (Sasa, 1990). A–C. ♂ (PCHN). D. Holotype, ♂ (NSMT- I-Dip 4690). E. ♂ (JNU 21081601). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D–E. Gonostylus, showing variation.	Fig. 1. Rheocricotopus (P.) amamipubescius (Sasa, 1990). A–C. ♂ (PCHN). D. Holotype, ♂ (NSMT- I-Dip 4690). E. ♂ (JNU 21081601). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D–E. Gonostylus, showing variation.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFEAFF80FE14FD22CA16CC72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507209/files/figure.png	https://doi.org/10.5281/zenodo.18507209	Fig. 2. Rheocricotopus (P.) bifasciatus Wang & Zheng, 1991. A, F, H. Holotype, ♂(NKU). B–E, G, I. ♂ (JNU). A–B. Abdominal colour pattern, showing variation. C. Colour pattern of foreleg. D. Colour pattern of mid leg. E. Humeral pit. F. Wing. G. Setal arrangement on tergites II–IV. H. Hypopygium, dorsal view. I. Gonostylus.	Fig. 2. Rheocricotopus (P.) bifasciatus Wang & Zheng, 1991. A, F, H. Holotype, ♂(NKU). B–E, G, I. ♂ (JNU). A–B. Abdominal colour pattern, showing variation. C. Colour pattern of foreleg. D. Colour pattern of mid leg. E. Humeral pit. F. Wing. G. Setal arrangement on tergites II–IV. H. Hypopygium, dorsal view. I. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFEAFF80FE14FD22CA16CC72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507205/files/figure.png	https://doi.org/10.5281/zenodo.18507205	Fig. 1. Rheocricotopus (P.) amamipubescius (Sasa, 1990). A–C. ♂ (PCHN). D. Holotype, ♂ (NSMT- I-Dip 4690). E. ♂ (JNU 21081601). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D–E. Gonostylus, showing variation.	Fig. 1. Rheocricotopus (P.) amamipubescius (Sasa, 1990). A–C. ♂ (PCHN). D. Holotype, ♂ (NSMT- I-Dip 4690). E. ♂ (JNU 21081601). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D–E. Gonostylus, showing variation.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFEFFF98FE3AFD06CA4DCD78.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507211/files/figure.png	https://doi.org/10.5281/zenodo.18507211	Fig. 3. Rheocricotopus (P.) chalybeatus (Edwards, 1929). A. ♂ (JNU 2402). B–H. ♂ (PCHN). I–K. ♀ (PCHN). A. Abdominal colour variation, dorsal view. B. Thorax with humeral pit, lateral view. C. Wing. D. Hypopygium, dorsal view. E. Anal point. F. Inferior volsella, showing variation. G–H. Gonostylus, showing variation. I. Genitalis, ventral view. J. Genitalis, dorsal view. K. Tergite IX, showing variation.	Fig. 3. Rheocricotopus (P.) chalybeatus (Edwards, 1929). A. ♂ (JNU 2402). B–H. ♂ (PCHN). I–K. ♀ (PCHN). A. Abdominal colour variation, dorsal view. B. Thorax with humeral pit, lateral view. C. Wing. D. Hypopygium, dorsal view. E. Anal point. F. Inferior volsella, showing variation. G–H. Gonostylus, showing variation. I. Genitalis, ventral view. J. Genitalis, dorsal view. K. Tergite IX, showing variation.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFEFFF98FE3AFD06CA4DCD78.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507213/files/figure.png	https://doi.org/10.5281/zenodo.18507213	Fig. 4. Rheocricotopus (P.) chalybeatus (Edwards, 1929). A–C. Pupa (PCHN). D–J. Larva (PCHN). A. Frontal apotome. B. Thoracic horn. C. Abdomen, dorsal view. D. Labro-epipharyngeal region. E. Antenna. F. Mandible. G. Maxilla. H. Mentum. I. Posterior abdomen, lateral view. J. Posterior parapod claws.	Fig. 4. Rheocricotopus (P.) chalybeatus (Edwards, 1929). A–C. Pupa (PCHN). D–J. Larva (PCHN). A. Frontal apotome. B. Thoracic horn. C. Abdomen, dorsal view. D. Labro-epipharyngeal region. E. Antenna. F. Mandible. G. Maxilla. H. Mentum. I. Posterior abdomen, lateral view. J. Posterior parapod claws.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFF7FF96FE1DFC08CCC7CAD6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507217/files/figure.png	https://doi.org/10.5281/zenodo.18507217	Fig. 5. Rheocricotopus (P.) hidakadeeus Sasa & Suzuki, 2001, holotype, ♂ (NSMT-I-Dip 5367). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view.	Fig. 5. Rheocricotopus (P.) hidakadeeus Sasa & Suzuki, 2001, holotype, ♂ (NSMT-I-Dip 5367). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFF8FF92FDD1F96BCDF3CB91.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507219/files/figure.png	https://doi.org/10.5281/zenodo.18507219	Fig. 6. Rheocricotopus (P.) oiraprimus Sasa, 1991. A–E. ♂ (PCHN). F. Holotype, ♂ (NSMT-I-Dip 5989). A–B. Abdominal colour pattern, showing variation. C. Thorax with humeral pit, lateral view. D. Wing. E. Hypopygium, dorsal view. F. Gonostylus.	Fig. 6. Rheocricotopus (P.) oiraprimus Sasa, 1991. A–E. ♂ (PCHN). F. Holotype, ♂ (NSMT-I-Dip 5989). A–B. Abdominal colour pattern, showing variation. C. Thorax with humeral pit, lateral view. D. Wing. E. Hypopygium, dorsal view. F. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFFCFFAEFE2AFC72CA4FCC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507221/files/figure.png	https://doi.org/10.5281/zenodo.18507221	Fig. 7. Rheocricotopus (P.) serratus Liu, Lin & Wang, 2014, ♂ (JNU 181210050). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D. Gonostylus.	Fig. 7. Rheocricotopus (P.) serratus Liu, Lin & Wang, 2014, ♂ (JNU 181210050). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFC1FFAFFE3CFD38CD47CF84.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507231/files/figure.png	https://doi.org/10.5281/zenodo.18507231	Fig. 11. Rheocricotopus (P.) valgus Chaudhuri & Sinharay, 1983. A–H. ♂ (JNU 170208001). I. Pupa (JNU 170308002). A. Abdominal colour pattern. B. Colour pattern of foreleg. C. Colour pattern of mid leg. D. Wing. E. Hypopygium, dorsal view. F–H. Gonostylus, showing variation. I. Abdomen, dorsal view.	Fig. 11. Rheocricotopus (P.) valgus Chaudhuri & Sinharay, 1983. A–H. ♂ (JNU 170208001). I. Pupa (JNU 170308002). A. Abdominal colour pattern. B. Colour pattern of foreleg. C. Colour pattern of mid leg. D. Wing. E. Hypopygium, dorsal view. F–H. Gonostylus, showing variation. I. Abdomen, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFC0FFABFE09FE7FCF48C983.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507223/files/figure.png	https://doi.org/10.5281/zenodo.18507223	Fig. 8. Rheocricotopus (P.) togakuroasi (Sasa & Okazawa, 1992). A–B, D–F. ♂ (PCHN). C. Holotype, ♂ (NSMT-I-Dip 4757). G. ♀ (PCHN). A. Thorax with humeral pit, lateral view. B. Wing. C. Tergite IV, showing setal arrangement. D. Hypopygium, dorsal view. E. Superior volsella. F. Gonostylus, showing variation. G. Genitalia, dorsal view, showing tergite IX.	Fig. 8. Rheocricotopus (P.) togakuroasi (Sasa & Okazawa, 1992). A–B, D–F. ♂ (PCHN). C. Holotype, ♂ (NSMT-I-Dip 4757). G. ♀ (PCHN). A. Thorax with humeral pit, lateral view. B. Wing. C. Tergite IV, showing setal arrangement. D. Hypopygium, dorsal view. E. Superior volsella. F. Gonostylus, showing variation. G. Genitalia, dorsal view, showing tergite IX.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFC0FFABFE09FE7FCF48C983.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507225/files/figure.png	https://doi.org/10.5281/zenodo.18507225	Fig. 9. Rheocricotopus (P.) togakuroasi (Sasa & Okazawa, 1992). A–B. Pupa (PCHN). C–H. Larva (PCHN). A. Frontal apotome. B. Abdomen, dorsal view. C. Labro-epipharyngeal region. D. Antenna. E. Mandible. F. Maxilla. G. Postmentum. H. Procercus.	Fig. 9. Rheocricotopus (P.) togakuroasi (Sasa & Okazawa, 1992). A–B. Pupa (PCHN). C–H. Larva (PCHN). A. Frontal apotome. B. Abdomen, dorsal view. C. Labro-epipharyngeal region. D. Antenna. E. Mandible. F. Maxilla. G. Postmentum. H. Procercus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFC0FFABFE09FE7FCF48C983.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507233/files/figure.png	https://doi.org/10.5281/zenodo.18507233	Fig. 12. A. Rheocricotopus (P.) kongi Lin & Wang, 2020, ♂ (NKU, XL-1853). B–J. R. (P.) orientalis Wang, 1995. B–E. ♂(JNU161009005). F–G. ♀ (JNU 24040903). H–J. Pupa (JNU 24041002). A. Hypopygium, dorsal view (two red arrows showing ventral and inner projections). B. Abdominal colour pattern. C. Thorax, lateral view (arrow showing humeral pit). D. Wing. E. Hypopygium, dorsal view. F. Genitalia, ventral view. G. Genitalia, dorsal view. H. Frontal apotome. I. Thoracic horn. J. Abdomen, dorsal view.	Fig. 12. A. Rheocricotopus (P.) kongi Lin & Wang, 2020, ♂ (NKU, XL-1853). B–J. R. (P.) orientalis Wang, 1995. B–E. ♂(JNU161009005). F–G. ♀ (JNU 24040903). H–J. Pupa (JNU 24041002). A. Hypopygium, dorsal view (two red arrows showing ventral and inner projections). B. Abdominal colour pattern. C. Thorax, lateral view (arrow showing humeral pit). D. Wing. E. Hypopygium, dorsal view. F. Genitalia, ventral view. G. Genitalia, dorsal view. H. Frontal apotome. I. Thoracic horn. J. Abdomen, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFC7FFA6FE1BFEB2CD89CFFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507227/files/figure.png	https://doi.org/10.5281/zenodo.18507227	Fig. 10. Rheocricotopus (P.) tokarakeleus Sasa & Suzuki, 1995, holotype, ♂ (NSMT I-Dip-5003). A. Humeral pit. B. Wing. C. Setal arrangement on tergite IV. D. Hypopygium, dorsal view. E. Gonostylus.	Fig. 10. Rheocricotopus (P.) tokarakeleus Sasa & Suzuki, 1995, holotype, ♂ (NSMT I-Dip-5003). A. Humeral pit. B. Wing. C. Setal arrangement on tergite IV. D. Hypopygium, dorsal view. E. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFC9FFA5FE04FD95CD05CDBA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507231/files/figure.png	https://doi.org/10.5281/zenodo.18507231	Fig. 11. Rheocricotopus (P.) valgus Chaudhuri & Sinharay, 1983. A–H. ♂ (JNU 170208001). I. Pupa (JNU 170308002). A. Abdominal colour pattern. B. Colour pattern of foreleg. C. Colour pattern of mid leg. D. Wing. E. Hypopygium, dorsal view. F–H. Gonostylus, showing variation. I. Abdomen, dorsal view.	Fig. 11. Rheocricotopus (P.) valgus Chaudhuri & Sinharay, 1983. A–H. ♂ (JNU 170208001). I. Pupa (JNU 170308002). A. Abdominal colour pattern. B. Colour pattern of foreleg. C. Colour pattern of mid leg. D. Wing. E. Hypopygium, dorsal view. F–H. Gonostylus, showing variation. I. Abdomen, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFCDFFA2FDC4FC18CD30C97E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507233/files/figure.png	https://doi.org/10.5281/zenodo.18507233	Fig. 12. A. Rheocricotopus (P.) kongi Lin & Wang, 2020, ♂ (NKU, XL-1853). B–J. R. (P.) orientalis Wang, 1995. B–E. ♂(JNU161009005). F–G. ♀ (JNU 24040903). H–J. Pupa (JNU 24041002). A. Hypopygium, dorsal view (two red arrows showing ventral and inner projections). B. Abdominal colour pattern. C. Thorax, lateral view (arrow showing humeral pit). D. Wing. E. Hypopygium, dorsal view. F. Genitalia, ventral view. G. Genitalia, dorsal view. H. Frontal apotome. I. Thoracic horn. J. Abdomen, dorsal view.	Fig. 12. A. Rheocricotopus (P.) kongi Lin & Wang, 2020, ♂ (NKU, XL-1853). B–J. R. (P.) orientalis Wang, 1995. B–E. ♂(JNU161009005). F–G. ♀ (JNU 24040903). H–J. Pupa (JNU 24041002). A. Hypopygium, dorsal view (two red arrows showing ventral and inner projections). B. Abdominal colour pattern. C. Thorax, lateral view (arrow showing humeral pit). D. Wing. E. Hypopygium, dorsal view. F. Genitalia, ventral view. G. Genitalia, dorsal view. H. Frontal apotome. I. Thoracic horn. J. Abdomen, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFCCFFBEFDD4FEB2CAB1C8AD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507233/files/figure.png	https://doi.org/10.5281/zenodo.18507233	Fig. 12. A. Rheocricotopus (P.) kongi Lin & Wang, 2020, ♂ (NKU, XL-1853). B–J. R. (P.) orientalis Wang, 1995. B–E. ♂(JNU161009005). F–G. ♀ (JNU 24040903). H–J. Pupa (JNU 24041002). A. Hypopygium, dorsal view (two red arrows showing ventral and inner projections). B. Abdominal colour pattern. C. Thorax, lateral view (arrow showing humeral pit). D. Wing. E. Hypopygium, dorsal view. F. Genitalia, ventral view. G. Genitalia, dorsal view. H. Frontal apotome. I. Thoracic horn. J. Abdomen, dorsal view.	Fig. 12. A. Rheocricotopus (P.) kongi Lin & Wang, 2020, ♂ (NKU, XL-1853). B–J. R. (P.) orientalis Wang, 1995. B–E. ♂(JNU161009005). F–G. ♀ (JNU 24040903). H–J. Pupa (JNU 24041002). A. Hypopygium, dorsal view (two red arrows showing ventral and inner projections). B. Abdominal colour pattern. C. Thorax, lateral view (arrow showing humeral pit). D. Wing. E. Hypopygium, dorsal view. F. Genitalia, ventral view. G. Genitalia, dorsal view. H. Frontal apotome. I. Thoracic horn. J. Abdomen, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFD0FFBAFDE2FC70CF07CA6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507237/files/figure.png	https://doi.org/10.5281/zenodo.18507237	Fig. 13. Rheocricotopus (P.) bannaensis sp. nov. A–D. Holotype, ♂ (JNU 170429003). E–G. Paratype, ♀ (JNU). A. Abdominal colour pattern. B. Thorax with humeral pit, lateral view. C. Wing. D. Hypopygium, dorsal view. E. Thorax, lateral view (arrow showing humeral pit). F. Genitalia, ventral view. G. Genitalia, dorsal view, showing tergite IX.	Fig. 13. Rheocricotopus (P.) bannaensis sp. nov. A–D. Holotype, ♂ (JNU 170429003). E–G. Paratype, ♀ (JNU). A. Abdominal colour pattern. B. Thorax with humeral pit, lateral view. C. Wing. D. Hypopygium, dorsal view. E. Thorax, lateral view (arrow showing humeral pit). F. Genitalia, ventral view. G. Genitalia, dorsal view, showing tergite IX.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFD5FFB9FD8AFB19CDAFCA47.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507241/files/figure.png	https://doi.org/10.5281/zenodo.18507241	Fig. 14. Rheocricotopus (P.) albulus sp. nov. A. ♂ (JNU). B–G. Holotype, ♂ (JNU 170512011). H–I. Paratype, pupa (JNU). A–B. Abdominal colour pattern, showing variation. C. Foreleg colour pattern. D. Humeral pit. E. Wing. F. Setal arrangement on tergites II–IV. G. Hypopygium, dorsal view. H. Frontal apotome. I. Abdomen, dorsal view.	Fig. 14. Rheocricotopus (P.) albulus sp. nov. A. ♂ (JNU). B–G. Holotype, ♂ (JNU 170512011). H–I. Paratype, pupa (JNU). A–B. Abdominal colour pattern, showing variation. C. Foreleg colour pattern. D. Humeral pit. E. Wing. F. Setal arrangement on tergites II–IV. G. Hypopygium, dorsal view. H. Frontal apotome. I. Abdomen, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFD6FFB7FDE9FB33CA4BCBA7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507243/files/figure.png	https://doi.org/10.5281/zenodo.18507243	Fig. 15. Rheocricotopus (P.) beibengensis sp. nov., holotype, ♂ (JNU 25121501). A. Wing. B. Humeral pit. C. Hypopygium, dorsal view.	Fig. 15. Rheocricotopus (P.) beibengensis sp. nov., holotype, ♂ (JNU 25121501). A. Wing. B. Humeral pit. C. Hypopygium, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFD8FFB2FDFEFA52CA4CCCA0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507247/files/figure.png	https://doi.org/10.5281/zenodo.18507247	Fig. 16. Rheocricotopus (P.) binotatus sp. nov. A, C, G. ♂ (JNU 23020507). B, D–F. Holotype, ♂ (JNU 170723009). A–B. Abdominal colour pattern, showing variation. C. Humeral pit. D. Wing. E. Setal arrangement on tergites II–IV. F. Hypopygium, dorsal view. G. Gonostylus.	Fig. 16. Rheocricotopus (P.) binotatus sp. nov. A, C, G. ♂ (JNU 23020507). B, D–F. Holotype, ♂ (JNU 170723009). A–B. Abdominal colour pattern, showing variation. C. Humeral pit. D. Wing. E. Setal arrangement on tergites II–IV. F. Hypopygium, dorsal view. G. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFDDFFB0FDC2FD50CD0BC8AC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507249/files/figure.png	https://doi.org/10.5281/zenodo.18507249	Fig. 17. Rheocricotopus (P.) chinogodavarius sp. nov. A, D–H. Holotype, ♂ (JNU 170724006). B–C, I–J. Paratype, ♂ (JNU). A–C. Abdominal colour pattern, showing variation. D. Colour pattern of foreleg. E. Colour pattern of mid leg. F. Humeral pit. G. Wing. H. Hypopygium, dorsal view. I. Inferior volsella, showing variation. J. Gonostylus, showing variation.	Fig. 17. Rheocricotopus (P.) chinogodavarius sp. nov. A, D–H. Holotype, ♂ (JNU 170724006). B–C, I–J. Paratype, ♂ (JNU). A–C. Abdominal colour pattern, showing variation. D. Colour pattern of foreleg. E. Colour pattern of mid leg. F. Humeral pit. G. Wing. H. Hypopygium, dorsal view. I. Inferior volsella, showing variation. J. Gonostylus, showing variation.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFDEFFCFFDE5FEB2CB0CCD62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507251/files/figure.png	https://doi.org/10.5281/zenodo.18507251	Fig. 18. Rheocricotopus (P.) fasciolatus sp. nov., holotype, ♂ (JNU 170509002). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Setal arrangement on tergites II–IV. E. Hypopygium, dorsal view.	Fig. 18. Rheocricotopus (P.) fasciolatus sp. nov., holotype, ♂ (JNU 170509002). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Setal arrangement on tergites II–IV. E. Hypopygium, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFA0FFCDFDE1FC11CD4DCA81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507257/files/figure.png	https://doi.org/10.5281/zenodo.18507257	Fig. 19. Rheocricotopus (P.) hanmiensis sp. nov., holotype, ♂ (JNU 251215A1). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Hypopygium, dorsal view. E. Gonostylus.	Fig. 19. Rheocricotopus (P.) hanmiensis sp. nov., holotype, ♂ (JNU 251215A1). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Hypopygium, dorsal view. E. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFA2FFC9FDF5FB71CF42CF4B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507260/files/figure.png	https://doi.org/10.5281/zenodo.18507260	Fig. 20. Rheocricotopus (P.) masarui sp. nov. A–B, E–H. Holotype, ♂ (JNU 211013H03B). C–D, I–J. Paratype, ♂ (JNU 22082001). K–N. Pupa (JNU 22082001). A–C. Abdominal colour pattern, showing variation. D. Humeral pit. E. Wing. F. Setal arrangement on tergite II. G. Setal arrangement on tergite IV. H. Hypopygium, dorsal view. I. Anal point. J. Gonostylus. K. Frontal apotome. L. Thoracic horn. M. Abdomen, dorsal view. N. Abdomen, ventral view.	Fig. 20. Rheocricotopus (P.) masarui sp. nov. A–B, E–H. Holotype, ♂ (JNU 211013H03B). C–D, I–J. Paratype, ♂ (JNU 22082001). K–N. Pupa (JNU 22082001). A–C. Abdominal colour pattern, showing variation. D. Humeral pit. E. Wing. F. Setal arrangement on tergite II. G. Setal arrangement on tergite IV. H. Hypopygium, dorsal view. I. Anal point. J. Gonostylus. K. Frontal apotome. L. Thoracic horn. M. Abdomen, dorsal view. N. Abdomen, ventral view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFA6FFC7FDE8FE3ECAB1CB7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507262/files/figure.png	https://doi.org/10.5281/zenodo.18507262	Fig. 21. Rheocricotopus (P.) monovittatus sp. nov. A–D. Holotype, ♂ (JNU 23032701). E. Paratype, ♂ (JNU 24033120). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Hypopygium, dorsal view. E. Gonostylus.	Fig. 21. Rheocricotopus (P.) monovittatus sp. nov. A–D. Holotype, ♂ (JNU 23032701). E. Paratype, ♂ (JNU 24033120). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Hypopygium, dorsal view. E. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFA8FFC2FDC3FA0ACB7EC8DA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507266/files/figure.png	https://doi.org/10.5281/zenodo.18507266	Fig. 22. Rheocricotopus (P.) nippobifasciatus sp. nov. A–D. Holotype, ♂ (NSMT-I-Dip 36256). E. Paratype, ♂ (NSMT-I-Dip 36262). F–G. Paratype, ♀ (NSMT-I-Dip 36269). A. Abdominal colour pattern. B. Thorax with humeral pit, lateral view. C. Wing. D. Hypopygium, dorsal view. E. Gonostylus. F. Genitalia, ventral view. G. Genitalia, dorsal view, showing tergite IX.	Fig. 22. Rheocricotopus (P.) nippobifasciatus sp. nov. A–D. Holotype, ♂ (NSMT-I-Dip 36256). E. Paratype, ♂ (NSMT-I-Dip 36262). F–G. Paratype, ♀ (NSMT-I-Dip 36269). A. Abdominal colour pattern. B. Thorax with humeral pit, lateral view. C. Wing. D. Hypopygium, dorsal view. E. Gonostylus. F. Genitalia, ventral view. G. Genitalia, dorsal view, showing tergite IX.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFACFFC1FDEDFEB2CF1DCCEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507268/files/figure.png	https://doi.org/10.5281/zenodo.18507268	Fig. 23. Rheocricotopus (P.) protuberans sp. nov., holotype, ♂ (JNU 160206012). A. Thorax with humeral pit, lateral view. B. Wing. C. Hypopygium, dorsal view.	Fig. 23. Rheocricotopus (P.) protuberans sp. nov., holotype, ♂ (JNU 160206012). A. Thorax with humeral pit, lateral view. B. Wing. C. Hypopygium, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFAEFFDFFDEAFCA4CAFDC927.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507274/files/figure.png	https://doi.org/10.5281/zenodo.18507274	Fig. 24. Rheocricotopus (P.) yunnanensis sp. nov. A–C, E. Holotype, ♂ (JNU 170723007). D, F. Paratype, ♂ (JNU 170614009). G. Paratype, ♂ (JNU 17061406). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Setal arrangement on tergites II–IV. E. Hypopygium, dorsal view. F. Superior volsella. G. Gonostylus.	Fig. 24. Rheocricotopus (P.) yunnanensis sp. nov. A–C, E. Holotype, ♂ (JNU 170723007). D, F. Paratype, ♂ (JNU 170614009). G. Paratype, ♂ (JNU 17061406). A. Abdominal colour pattern. B. Humeral pit. C. Wing. D. Setal arrangement on tergites II–IV. E. Hypopygium, dorsal view. F. Superior volsella. G. Gonostylus.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFB4FFD5FE23FCF4CFA8CAE2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507276/files/figure.png	https://doi.org/10.5281/zenodo.18507276	Fig. 25. Rheocricotopus (s. str.) tamahumeralis Sasa, 1981. A–N. ♂ (PCHN). O–P. ♀ (PCHN). A. Thorax with humeral pit, lateral view. B–D. Humeral pit, showing variation. E. Wing. F. Hypopygium, dorsal view. G–H. Superior volsella, showing variation. I–J. Inferior volsella, showing variation. K–N. Gonostylus, showing variation. O. Genitalia, ventral view. P. Genitalia, dorsal view.	Fig. 25. Rheocricotopus (s. str.) tamahumeralis Sasa, 1981. A–N. ♂ (PCHN). O–P. ♀ (PCHN). A. Thorax with humeral pit, lateral view. B–D. Humeral pit, showing variation. E. Wing. F. Hypopygium, dorsal view. G–H. Superior volsella, showing variation. I–J. Inferior volsella, showing variation. K–N. Gonostylus, showing variation. O. Genitalia, ventral view. P. Genitalia, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFB4FFD5FE23FCF4CFA8CAE2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507280/files/figure.png	https://doi.org/10.5281/zenodo.18507280	Fig. 26. Rheocricotopus (s. str.) tamahumeralis Sasa, 1981. A–E. Pupa (PCHN). F–L. Larva (PCHN). A. Frontal apotome. B. Thoracic horn. C. Abdomen, dorsal view. D. Segment III, dorsal view, showing variation. E. Segment VII, dorsal view. F. Labro-epipharyngeal region. G. Antenna. H. Mandible. I. Maxilla. J. Mentum. K. Procercus. L. Posterior parapod claws.	Fig. 26. Rheocricotopus (s. str.) tamahumeralis Sasa, 1981. A–E. Pupa (PCHN). F–L. Larva (PCHN). A. Frontal apotome. B. Thoracic horn. C. Abdomen, dorsal view. D. Segment III, dorsal view, showing variation. E. Segment VII, dorsal view. F. Labro-epipharyngeal region. G. Antenna. H. Mandible. I. Maxilla. J. Mentum. K. Procercus. L. Posterior parapod claws.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFBAFFD0FDCBFA91CB4ACD76.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507284/files/figure.png	https://doi.org/10.5281/zenodo.18507284	Fig. 27. Rheocricotopus (s. str.) tatequintus Sasa, 1996. A–C. Holotype, ♂ (NSMT I-Dip-4998). D–F. Paratype, ♂ (NSMT No. 284: 59). G–H. Paratype, ♀ (NSMT No. 284: 61). A, G. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D. Superior volsella. E. Inferior volsella. F. Gonostylus. H. Genitalia, ventral view.	Fig. 27. Rheocricotopus (s. str.) tatequintus Sasa, 1996. A–C. Holotype, ♂ (NSMT I-Dip-4998). D–F. Paratype, ♂ (NSMT No. 284: 59). G–H. Paratype, ♀ (NSMT No. 284: 61). A, G. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D. Superior volsella. E. Inferior volsella. F. Gonostylus. H. Genitalia, ventral view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFBFFFEAFE60FC02CD3FCB38.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507286/files/figure.png	https://doi.org/10.5281/zenodo.18507286	Fig. 28. Rheocricotopus (s. str.) togapeniculus Sasa & Okazawa, 1992. A–E. ♂ (PCHN). F–G. ♀ (PCHN). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D. Superior volsella, showing variation. E. Gonostylus, showing variation. F. Genitalia, ventral view. G. Genitalia, dorsal view.	Fig. 28. Rheocricotopus (s. str.) togapeniculus Sasa & Okazawa, 1992. A–E. ♂ (PCHN). F–G. ♀ (PCHN). A. Humeral pit. B. Wing. C. Hypopygium, dorsal view. D. Superior volsella, showing variation. E. Gonostylus, showing variation. F. Genitalia, ventral view. G. Genitalia, dorsal view.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
03F55F24FFBFFFEAFE60FC02CD3FCB38.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18507288/files/figure.png	https://doi.org/10.5281/zenodo.18507288	Fig. 29. Rheocricotopus (s. str.) togapeniculus Sasa & Okazawa, 1992. A–C. Pupa (PCHN). D–J. Larva (PCHN). A. Frontal apotome. B. Abdomen, dorsal view. C. Segment III, dorsal view, showing variation. D. Labro-epipharyngeal region. E. Antenna. F. Mandible. G. Maxilla. I. Procercus. J. Posterior parapod claws.	Fig. 29. Rheocricotopus (s. str.) togapeniculus Sasa & Okazawa, 1992. A–C. Pupa (PCHN). D–J. Larva (PCHN). A. Frontal apotome. B. Abdomen, dorsal view. C. Segment III, dorsal view, showing variation. D. Labro-epipharyngeal region. E. Antenna. F. Mandible. G. Maxilla. I. Procercus. J. Posterior parapod claws.	2026-01-30	Niitsuma, Hiromi;Tang, Hongqu		Zenodo	biologists	Niitsuma, Hiromi;Tang, Hongqu			
