taxonID	type	description	language	source
03EB87BCFFF2FFC8FF446E0BDCD73BE0.taxon	description	Figs 1 – 14	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFF2FFC8FF446E0BDCD73BE0.taxon	type_taxon	Type species Strigoderma costulipennis Bates, l 888: 260, by present designation.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFF2FFC8FF446E0BDCD73BE0.taxon	diagnosis	Diagnosis Elongated body (Fig. 1). Frons and clypeus roughly-punctate (Fig. 2 A – H). Pronotum finely punctate (Fig. 2 I – P). Elytra with 5 – 10 longitudinal ridges notably raised; intercostal area finely tomentose (Figs 3 – 4). Humeral callus extending laterally to the outer margin, interrupting the epipleura (Figs 6 B to 12 B). Inner protarsal claw with the upper ramus very long and pointed (Figs 6 E to 12 E). Parameres with the apex dorsoventrally depressed, rounded apex (Figs 6 G – I, 12 G – I).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFF2FFC8FF446E0BDCD73BE0.taxon	etymology	Etymology From the Latin ‘ costa ’ (‘ ridge’) and ‘ tergum ’ (‘ back’ or ‘ spine’), in reference to the prominent elytral ridges, diagnostic for this subgenus.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFF2FFC8FF446E0BDCD73BE0.taxon	description	Variation Slightly evident mainly in the puncture density on head, pronotum and pygidium. Noticeable in the coloration, where species with yellow tones on the elytra have varying degrees of dark spots, which may be absent (completely yellow elytra), in addition to melanic specimens. Sexual dimorphism Females exhibit marked dimorphism in antennae, elytra, and front legs. Antennae with antennal club approximately 1 / 4 to 1 / 5 shorter. Elytra with external ridge and epipleura very widened. Protibia with distal denticle longer and curved; protarsomeres thinner, with proximal tarsomere longer than the three subsequent ones; internal protarsal claw with upper and lower ramus of similar thickness, almost equal length. Females generally with slightly less deep and dense punctation on head and pronotum.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFF2FFC8FF446E0BDCD73BE0.taxon	distribution	Distribution The species of this subgenus have a distribution mainly on the Pacific slope, between Chiapas and Sinaloa, with penetration towards the centre of the country in Guerrero and Morelos, in plant communities of deciduous tropical forest and corn plantations, located from sea level to 1300 m a. s. l., with records for the states of Chiapas, Colima, Guerrero, Oaxaca, Morelos and Sinaloa. This distribution fits well with the Typical Neotropical Distribution Pattern with high level of penetration, as described by Halffter (1964, 1974, 1976) and reviewed by Halffter & Morrone (2017), whose species have occupied Mexico through the tropical lowlands from South American lineages. Included species	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFF2FFC8FF446E0BDCD73BE0.taxon	discussion	Comments Bader (1992) considered S. tomentosa and S. costulipennis (Fig. 1 A) within the same group “ Costulipennis ”, justifying this by the granular surface of the elytra. However, the body shape, the unique condition of the elytra very different in both species (completely granular surface in S. tomentosa vs surface with notable ridges in S. costulipennis), and different genital configuration do not support this grouping, in addition to the fact that their geographic distributions present different biogeographic patterns. Additionally, Bader (1992) considered S. costulipennis as a species with a very wide and disjunct geographical distribution, with records that could not be corroborated. Explanatory notes Strigoderma tomentosa has traditionally been grouped together with S. costulipennis, since the former has shallow ridges, apart from this characteristic, they do not share much. S. tomentosa represents a particular taxon within the genus, so far recorded only from Juquila (Oaxaca, Mexico), and its morphology is unmistakable. It differs from S. costulipennis and from all the species in the subgenus S. (Costatergus) subgen. nov. (Fig. 1) by presenting as autapomorphy a ‘ coat’ of short and soft hairs on the dorsal surface (frons and pronotum) (finely tomentose surface on elytra in S. (Costatergus )), it has a uniform green coloration (black coloration, sometimes with spots in S. (Costatergus )), in addition, the protarsal claw have both the upper and lower ramus long (claw with a very short lower ramus in S. (Costatergus )); the mesometasternal process is broadly rounded, but slightly protruding (narrow and protruding in S. (Costatergus )), and the parameres are elongated, thin towards the base, separated and with slightly concave tips (protruding in S. (Costatergus )).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFBFFCCFE246D99DB263883.taxon	description	Figs 1 A, 2 A, I, 3 A, 4 A, 5 A – B, 6, 14 Strigoderma costulipennis Bates, l 888: 260.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFBFFCCFE246D99DB263883.taxon	diagnosis	Diagnosis Elytra with nine elevated ridges; ridge 3, 5 and 7 only marked at base; ridge 8 and 9 convergent in distal third. 5 th protarsomere with developed preapical denticle. Inner protarsal claw with rhomboidal lower ramus, slightly longer than half of upper ramus. Parameres elongated, thin, with curved outer margin.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFBFFCCFE246D99DB263883.taxon	materials_examined	Type material Lectotype (here designated) MEXICO – Guerrero • ♀; “ Chilpancingo, Guerrero. Höge; // Sp. figured; // Type; // Strigoderma costulipennis Bates (handwritten); // B. C. A., Coll., II (2). Strigoderma costulipennis; // Strigoderma (Costatergus) costulipennis Bates LECTOTYPE ♀ Andalco-Cid & Ramírez-Ponce, des. ” [yellow label printed]; BMNH. Paralectotypes MEXICO – Guerrero • 1 ♀; “ Chilpancingo, Guerrero, 4600 ft. June. H. H. Smith; // B. C. A., Coll., II (2). Strigoderma costulipennis; // Strigoderma (Costatergus) costulipennis Bates PARALECTOTYPE ♀ Andalco-Cid et al. 2024 des. ” [yellow label, printed]; BMNH. – Oaxaca • 1 ♀; “ Juquila, México. Hoege. // B. C. A., Coll., II (2). Strigoderma costulipennis; // Strigoderma costulipennis Bates // Strigoderma (Costatergus) costulipennis Bates PARALECTOTYPE ♀ Andalco-Cid et al. 2024 des. ” [yellow label, printed]; BMNH ” Two specimens of the original type series housed at the BMNH belonging to the collection of the Biologia Centrali-Americana (Bates 1888: 260) are regarded here as representatives of different species; one female from “ Juquila, México ” of Höge (Fig. 5 C) is only identified at this stage to subgenus level, and one male from “ Mexico ” (Fig. 5 D) is now a paratype of S. (C.) grossipenis sp. nov. Other material examined MEXICO – Guerrero • 1 ♂; “ Mexcala, Gro. Mex. VI- 29 - 51; // P. D. Hurd Collector; // UC Berkeley EMEC 1324375 “ QR Code ”; // Strigoderma costulipennis Bates, Det. A. Bader 1989; // Strigoderma costulipennis Bates, Ramírez-Ponce & Andalco det. ”; EMEC • 1 ♂; “ MEX: 2 Km S Xalitla, Gro. VII- 25 - 83 J. Chemsak, A & M Michelbacher; // UC Berkeley EMEC 1324374 “ QR Code ”; // Strigoderma costulipennis Bates, Det. A. Bader 1989; // Strigoderma costulipennis Bates, Ramírez-Ponce & Andalco det. ”; EMEC.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFBFFCCFE246D99DB263883.taxon	description	Description Male (UC Berkeley, EMEC 1324375) BODY SIZE. Total length: 6.81 mm. Maximum elytral width: 2.88 mm (Figs 1 A, 6 A). BODY COLOUR. Head, pronotum, and scutellum black with coppery reflections. Elytra ochre-yellow with wide dark coppery bands preceding the outer edge and contiguous to the suture. Abdomen dark reddish black (Fig. 6 A – C). CLYPEUS. Subrectangular, broad. Anterior angles broadly rounded; anterior edge slightly sinuate; outer margin moderately raised, interrupted in the centre. Surface with latero-apical excavation on each side. Punctation large, rough, deep and dense, with transverse pattern (Fig. 2 A). FRONS. Convex. Rough-punctate sculpture, wider and deeper than in clypeus. ANTENNA. Antennomeres 1 – 3 of similar length. Club as long as the pedicel and funiculus together (Fig. 6 D). PRONOTUM. Subtrapezoidal shape, clearly wider than long; noticeably widened on anterior half. Anterior angles short. Surface with longitudinal median groove reaching the middle. Punctures small, very dense, heterogeneous, almost rough (Fig. 2 I). Lateral silhouette slight and uniformly convex (lateral view) (Fig. 6 B). SCUTELLUM. Triangular shape, wide and angular apex. Sculpture with punctation similar in width and depth to the pronotum. ELYTRA. Subparallel to the distal two thirds. Surface with nine elytral ridges with rounded edges; the 3 rd, 5 th, and 7 th incomplete (only evident at the elytral base) (Figs 1 A, 3 A); ridges 8 and 9 join together as one on the distal third (Fig. 4 A); intercostal space rough. Elytral suture elevated up to the 2 nd distal third. Humeral callus discrete, developed up to the lateral margins of the elytra, interrupts the epipleura. Epipleura flat, thin and elevated (Fig. 6 B). PYGIDIUM. Frontal shape wider than long, with a broadly rounded apex. Lateral shape noticeably convex in the median part. Surface uniform: sculpture with dense, deep punctures, smaller than propygidium and some overlapping. Vestiture with thin, long setae in the apical area. METASTERNUM. Metasternal process narrow, slightly projecting; metasternal process uniform and deeply punctate (Fig. 6 C). ABDOMEN. Sternites 1, 2 and 3 almost equal in length. Sternite 4 as thick as the third sternite plus 1 / 4 of any of the first three. Surface with uniform punctation, separated by one puncture diameter (more dispersed on anal plate), sub-oval transverse, shallow (Fig. 6 C). LEGS. Protibia thick; distal denticle long, slightly curved, with rounded apex (Fig. 6 F). Protarsomeres 1 – 4 short and moderately broad, successively slightly shorter (Fig. 6 E). Arrangement of ventral setae: 3 spines on each protarsomere, short to moderate, successively thicker. Protarsomere 5 robust, broad, slightly longer than combined length of protarsomeres 1 – 4. Ventral lobe inserted subapically, poorly developed. Inner protarsal claw broad and deeply notched: lower ramus rhomboidal, reaching slightly more than half of upper ramus (Fig. 6 E). Mesotibia gradually thickened towards the apex; apex with 9 – 10 short spines. Surface with three transverse setiferous carinae; the proximal one with 3 – 4 thick and moderately long spines; the middle one with 4 – 5 thick and long spines; the distal one with 5 – 6 long spines. Surface with moderately deep and regular to irregular punctation. Apical spurs straight: the inner one with rounded apex, as long as the first two tarsomeres plus part of the third; the outer one with rounded apex, of similar length to the first one plus almost the second mesotarsomere. Mesotarsomeres 1 – 4 of similar length and thickness. Dorsal setae pattern: first tarsomere with 2 long, thick spines; second and third with 2 moderately thick, moderately long setae; fourth with two long, thin setae. Ventral setae pattern: first and second tarsomere with three long spines and one seta; third and fourth with four long, thick spines. Metatibia clearly widened in the middle and apical part; apex with 12 – 13 short spines. Surface with three transverse setiferous carinae: the proximal one with 2 – 3 thick spines; the middle one with 6 – 7 moderate spines, the distal one with 8 – 9 long spines. Apical spurs straight: the inner one very thick with rounded apex, as long as the first tarsomere; the outer one with rounded apex, of a length similar to half of the first mesotarsomere. Metatarsomeres similar in length, thickening upwards. Dorsal setae pattern: tarsomere 1 with 2 moderately long and moderately thin spines; protarsomere 2 and 3 with 2 spines successively thinner; fourth not noticeable. Ventral setae pattern: protarsomere 1 with 4 long and thick spines; protarsomeres 2, 3 and 4 with 3 long and successively thinner spines and 1 long seta successively thinner. GENITALIA. Parameres long, with curved outer margin, narrower before the apex; rounded apices (Fig. 6 H – I). Apical figure (lateral view) slightly curved ventrally (Fig. 6 G). VARIATION. Males may have a coppery or greenish reflection on pronotum and head. Female Similar to the male, except for the following differences. Head with punctation slightly more superficial and slightly less dense on clypeus and frons; antennal club shorter. Pronotum with slightly smaller punctation, noticeably less dense on the posterior central area. Elytra with very thickened outer ridge, clearly sinuate in its middle portion. Pygidium subtly concave (in lateral view).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFBFFCCFE246D99DB263883.taxon	distribution	Distribution This species is distributed in the centre of the state of Guerrero (southern Mexico), in localities located between 500 and 1400 m a. s. l., characterized by scrubland and low deciduous forest (Fig. 14).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFBFFCCFE246D99DB263883.taxon	biology_ecology	Natural history It has been collected on flowers of Acacia Mill. Its life cycle is unknown. Adult specimens were collected in June and July.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFBFFCCFE246D99DB263883.taxon	discussion	Taxonomic comments The type specimens housed at the BMHN (three females and one male; Fig. 5), represented a mixture of three different species, which, upon review and comparison with the other species, were able to be associated based on morphology and biogeographical correspondence. In the case of the specimen cited from Juquila (Fig. 5 C) (one single female), it is an unpublished species different from those described here, while the male specimen cited as “ Mexico ” (Fig. 5 D) corresponds to Strigoderma (C.) grossipenis sp. nov. We reject the record from Etla for this species cited by Machatschke (1957, 1972), because the species of the subgenus have restricted distributions, as demonstrated in the specific definitions present.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFFFFD3FE876EBEDFD73A25.taxon	description	Figs 1 B, 2 B, J, 3 B, 4 B, 7, 14	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFFFFD3FE876EBEDFD73A25.taxon	diagnosis	Diagnosis Pronotum dense and finely punctate, with projecting anterior angles. Elytra with seven ridges; the sixth and seventh not converging to form a single ridge in the distal third. Third ridge only developed at the base of the elytra. Internal protarsal claw with inferior ramus less than half the length of the upper ramus. Parameres elongate, subtriangular, with almost straight external margins.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFFFFD3FE876EBEDFD73A25.taxon	etymology	Etymology From the Latin ‘ tenebrōsus, tenebra ’ (‘ darkness’), alluding to its intense dark colour.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFFFFD3FE876EBEDFD73A25.taxon	materials_examined	Type material Holotype (revised) MEXICO – Chiapas • ♂; “ MEXICO: Chiapas, Villaflores, Finca Santa Elena, 12 - VI- 2001, C. J. Morales col. ”; IEXA. Paratype (revised) MEXICO – Chiapas • ♂; same data as for holotype; EMAC. Other material examined MEXICO – Chiapas • 2 ♂♂; “ Villa Flores, Chiapas, México, 10 - 06 - 1984, Aguilar. A // Strigoderma tenebrosa Delgado & Mora-Aguilar, 2012. Ramírez Ponce, 2021 det. ”; ARPC.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFFFFD3FE876EBEDFD73A25.taxon	description	Redescription Holotype male (IEXA) BODY SIZE. Total length: 7.7 mm. Maximum elytral width: 4.1 mm (Fig. 1 B). BODY COLOUR. Completely black, with greenish reflections mainly on the protarsus (Fig. 7 A – C). CLYPEUS. Sub-trapezoidal, elongate. Anterior angles rounded; anterior edge slightly sinuate; external margin weakly raised. Surface with latero-basal excavation above the fronto-clypeal suture. Surface with broad, rough, deep and dense punctation (Fig. 2 B). FRONS. Convex. Sculpture roughly-punctate, very dense, wider and deeper than in clypeus on disc (Fig. 2 B). ANTENNA. Antennomere 3 longer than the preceding ones. Club longer than the pedicel and funiculus together (Fig. 7 D). PRONOTUM. Subrectangular, longer than wide; moderately widened in anterior half. Anterior angles projecting. Surface without longitudinal median groove. Punctation small, very dense, homogeneous, fine (Fig. 2 J). Lateral view moderately convex (Fig. 7 B). SCUTELLUM. Broad sub-triangular, with rounded apex. Surface punctation similar in width to pronotum, deeper. ELYTRA. Subparallel to the 2 nd third. Surface with seven elevated ridges with acute edges; the 3 rd, incomplete (only evident at the elytral base) (Fig. 3 B); ridges six and seven not joint together as one on the distal third (Fig. 4 B); intercostal space wide, shallow and smooth. Elytral suture elevated up to the 2 nd distal third. Humeral callus discrete, developed up to the lateral margins of the elytra, interrupts the epipleura. Epipleura flat, thin and elevated. PYGIDIUM. Wider than long, with a broadly rounded apex. Punctation small, deep, dense and well-defined. METASTERNUM. Metaventral process slightly broad and slightly projecting; metasternum uniform and somewhat finely punctate (Fig. 7 C). ABDOMEN. Sternites 1, 2 and 3 almost equal in length. Sternite fourth as thick as the third sternites plus ½ of any of the first two (Fig. 7 C). Surface with uniform punctation, separated by two puncture diameters (more dispersed on anal plate), almost rounded, shallow. LEGS. Protibia thick; distal denticle long, slightly curved, with rounded apex (Fig. 7 F). Protarsomeres 1 – 4 conspicuously short and broad, successively slightly longer (Fig. 7 E). Ventral setae with three short to moderately long spines, very thick to moderately thin, successively decreasing in thickness. 5 th protarsomere robust, broad, slightly longer than tarsomeres 1 – 4 together. Ventral lobe subapical, developed. Inner protarsal claw broad and deeply notched: lower ramus lanceolate, not reaching half the length of the upper one (Fig. 7 E). Mesotibia thickened in the middle; apex with 12 – 13 short spines. Surface with two transverse setiferous carinae; the upper one with 4 to 5 thick and medium spines; the lower one with 12 – 13 medium spines. Surface with very large, irregular and superficial puncture-like impressions. Apical spurs straight: the inner one with a rounded apex, as long as the first two tarsomeres; the outer one with a rounded apex, of similar length to the first mesotarsomere. Mesotarsomeres 1 – 4 of similar in length and thickness. Dorsal setae pattern: mesotarsomeres 1 – 3 with 2 long, moderately thin spines; mesotarsomere 4 with 2 long, thin setae. Ventral setae pattern: mesotarsomeres 1 – 3 with 4 short to long, thick spines; mesotarsomere 4 with 3 short to long spines and 1 moderately thick, long seta. Metatibia clearly widened in the middle and apex; apex with 11 – 12 short spines. Surface with three transverse setiferous carinae: the proximal one with 2 – 3 thick spines; the middle one with 5 – 6 moderate spines, the distal one with 8 – 9 long spines. Apical spurs straight: the inner one with rounded apex, as long as the first one plus a small part of the second tarsomere; the outer one with sharp apex, of similar length to the first mesotarsomere. Metatarsomeres thickened and elongated in descending order. Dorsal setae pattern: metatarsomere 1 with 2 thick spines; metatarsomeres 2 and 3 with 2 long, moderately thick spines; metatarsomere 4 with 2 long, thin setae. Ventral setae pattern: metatarsomeres 1 and 2 with 4 long, thick spines; metatarsomere 3 with 3 long, thick spines and 1 long, thin setae; metatarsomere 4 with 2 long, moderately thick spines and 2 long setae. GENITALIA. Parameres long, subtriangular, with outer margin almost straight, gradually narrowing; apices slightly acute (Fig. 7 H – I). Figure (lateral view) slightly curved ventrally (Fig. 7 G). VARIATION. Males may have more intense greenish reflections on the head and pronotum. The surface between the ridges may vary from longitudinally concave to slightly convex. Female Unknown.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFFFFD3FE876EBEDFD73A25.taxon	distribution	Distribution This species has only been recorded from the central depression of Chiapas (Fig. 14).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFFFFD3FE876EBEDFD73A25.taxon	biology_ecology	Natural history Collected from corn plants and tropical deciduous forest areas at 600 m a. s. l. (Delgado & Mora-Aguilar 2012).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFFFFFD3FE876EBEDFD73A25.taxon	discussion	Taxonomic comments This species presents the greatest similarity with S. (C.) juliani sp. nov. and S. (C.) borealis sp. nov., characterized by the absence of three ridges (in relation to the species that all have nine or 10), making the spaces between the ridge very wide and lacking punctation. However, S. (C.) tenebrosa is distinguished by the presence of the complete 2 nd ridge (blurred in the central part of the elytra in S. (C.) juliani), and the lower ramus shorter than half the length of the upper one (longer than half the length of the upper one in S. (C.) borealis).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE0FFD6FF0A6CDBDD823A9F.taxon	description	urn: lsid: zoobank. org: act: 47 F 19470 - 2 C 21 - 4864 - A 41 B-DB 48 FBD 8 AB 4 B Figs 1 C, 2 C, K, 3 C, 4 C, 8, 14	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE0FFD6FF0A6CDBDD823A9F.taxon	diagnosis	Diagnosis Pronotum with heterogeneous (small to moderate) and sparse punctation (punctures not touching) over the entire surface. Elytra with 5 complete, rounded elevated ridges; interestrial area with wide, very shallow punctation; 5 th and 6 th subparallel ridges throughout. 5 th protarsomere with barely apparent basal denticle. Parameres with almost straight lateral margins.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE0FFD6FF0A6CDBDD823A9F.taxon	etymology	Etymology From the Latin ‘ pauci ’ (‘ few’), and the Latin ‘ punctum ’ (‘ prick, sharp point’), in reference to the scattered punctation of the pronotum; ‘ paucipunctata ’, ‘ with few punctures’.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE0FFD6FF0A6CDBDD823A9F.taxon	materials_examined	Type material Holotype MEXICO – Chiapas • ♂; “ Sepuleura [Sepultura] Canyon, Chiapas, México VI- 27 - 87; // 16 Km north, Arriaga, Chiapas, Mexico VI- 27 - 87; // Coll. by W. F. Chamberlain; // TAMU ENTO X 0043337 “ barcode ”; // Strigoderma costulipennis Bates, Det. 1997 by Paul K. Lago; // Strigoderma (Costatergus) paucipunctata Andalco-Cid & Ramírez-Ponce, 2025 HOLOTYPE ” [red label, printed]; TAMU.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE0FFD6FF0A6CDBDD823A9F.taxon	description	Description BODY SIZE. Total length: 6.77 mm. Maximum elytral width: 2.89 mm (Fig. 1 C). BODY COLOUR. Completely black with intense greenish reflections, except elytra without reflections (Fig. 8 A – C). CLYPEUS. Subrectangular, elongate. Anterior angles broadly rounded; anterior edge slightly sinuate; outer margin weakly raised. Surface with an anterolateral excavation on each side. Punctation broad, rough, deep and dense (Fig. 2 C). FRONS. Flat. Rough-punctate sculpture, denser than on clypeus (Fig. 2 C). ANTENNA. Antennomere 3 longer than preceding ones. Club as long as pedicel and funiculus together (Fig. 8 D). PRONOTUM. Subtrapezoidal, distinctly broader than long; widened in anterior half. Anterior angles short. Surface without longitudinal median groove. Mixed punctation; small to moderate, dense in small areas, mainly on sides (Fig. 2 K). Lateral view markedly convex (side view) (Fig. 8 B). SCUTELLUM. Triangular in shape, wide, with rounded apex. Sculpture with abundant punctation, very small, defined, thick and deep. ELYTRA. Subparallel to the 2 nd third. Surface with six elevated ridges with rounded edges (Fig. 3 C); ridges five and six not joining together to form only one in the distal third (Fig. 4 C); intercostal space slightly rough and punctate, with shallow and very wide punctures. Elytral suture elevated up to the 2 nd distal third. Humeral callus prominent, developed to the lateral margins of the elytra, interrupting the epipleura. Epipleura thick at the base, with a noticeable fold above the episternum, the rest flat and elevated. PYGIDIUM. Wider than long, with a broadly rounded apex. Shape (in lateral view) noticeably convex in the middle part. Surface regular and uniform, with lateral depressions in the anterior portion. Surface punctation moderate in size, deep and dense. METASTERNUM. Metaventral process narrow, not projecting; metasternum uniform and moderately punctate, punctation somewhat small and scattered (Fig. 8 C). ABDOMEN. Sternites 1, 2 and 3 almost equal in length. Sternite 4 th as thick as any of the first three sternites plus the half of another. Surface with small uniform punctation, separated by three puncture diameters (more dispersed on anal plate), sub-oval transverse, shallow (Fig. 8 C). LEGS. Protibia thick; distal denticle long, slightly curved, with rounded apex (Fig. 8 F). Protarsomeres 1 – 4 conspicuously short and broad, of similar size and thickness (Fig. 8 E). Ventral setae on tarsomere 1 and 2 with three spines, very short to long, moderately thin; third and fourth protarsomeres with three thick spines, very short to long. Protarsomere 5 robust, broad, slightly longer than tarsomeres 1 – 4 together. Ventral lobe poorly developed, medially-internally positioned, widely rounded (Fig. 8 E). Mesotibia gradually thickened towards the apex; apex with 9 – 10 medium spines. Surface with two transverse setiferous carinae; the upper one with 5 – 6 thick and moderately long spines; the lower one with 7 – 8 long spines. Sculpture with very deep and irregular punctation. Apical spurs straight, the inner one with acute apex, as long as the first two tarsomeres; the outer one with acute apex, of similar length to the first mesotarsomere. Mesotarsomeres 1 – 4 of similar length and thickness. Dorsal setae pattern: first mesotarsomere with 2 thick, moderately long spines; second and third mesotarsomeres with 2 moderately long, moderately thick spines; fourth mesotarsomere with 2 thin setae. Ventral setae pattern: first, second, and third mesotarsomeres with 4 thick, moderately long spines; fourth mesotarsomere with 4 moderately long, moderately thin spines. Metatibia clearly widened in the middle and apical part; apex with 11 – 12 medium spines. Surface with two transverse setiferous carinae: the proximal one with 4 – 5 thick spines; the distal one with 7 – 8 long spines. Sculpture with very deep and irregular punctation. Apical spurs straight: the inner one with a rounded apex, as long as the first two tarsomeres; the outer one short, very thick with a rounded apex, slightly shorter than the first mesotarsomere. Metatarsomeres dorsal setae pattern: fourth metatarsomere with 2 moderately long and stout spines; third metatarsomere with two moderately long and slender setae; second metatarsomere with 2 long and slender setae. Ventral setae pattern: first metatarsomere with 4 long and stout spines; second metatarsomere with 3 long and stout spines and one setae; third metatarsomere with 2 spines and two setae. GENITALIA. Parameres short, subtriangular, with outer margin almost straight, gradually narrowing; apices rounded (Fig. 8 H – I). Apical figure (lateral view) slightly curved ventrally (Fig. 8 G). Female Unknown.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE0FFD6FF0A6CDBDD823A9F.taxon	distribution	Distribution This species has only been recorded from the type locality (16 km north of Arriaga, Chiapas), close to the La Sepultura Biosphere Reserve and within the “ Selva Zoque-La Sepultura ” Terrestrial Priority Region for Conservation (RTP- 132) (Arriaga et al. 2000), characterized by an intermixture of extensive forest masses of tropical forests, mesophilous forests and pine forests, high rates of endemism and a Pleistocene refuge for diverse groups (Fig. 14).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE0FFD6FF0A6CDBDD823A9F.taxon	biology_ecology	Natural history Feeding habits and life cycle are unknown. The only specimen was collected in June.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE0FFD6FF0A6CDBDD823A9F.taxon	discussion	Taxonomic comments This species is easy to distinguish as it has the widest, most heterogeneous and least dense pronotal punctation (Figs 2 K, 8 A). It shares with S. (C.) grossipenis sp. nov. and S. (C.) uniungula sp. nov. the intercostal area with wide punctation; however, S. (C.) grossipenis has convergent ridges 5 and 6 in the apical callus (subparallel in S. (C.) paucipunctata sp. nov.), and in S. (C.) uniungula the intercostal punctation is very marked (shallow in S. (C.) paucipunctata), in addition to being the only species with a simple inner protarsal claw (split in S. (C.) paucipunctata).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE5FFD5FED86CB2DCDB3CFA.taxon	description	urn: lsid: zoobank. org: act: ED 68 ADF 7 - C 51 A- 46 E 9 - AFB 2 - 3 AD 25 A 273 C 5 F Figs 1 D, 2 D, L, 3 D, 4 D, 9, 14	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE5FFD5FED86CB2DCDB3CFA.taxon	diagnosis	Diagnosis Pronotum with uniform, very dense and fine punctation. Elytra with 5 complete, sharp elevated ridges, the 2 nd only marked at the ends; inter-costal area without punctation; ridges 5 and 6 subparallel throughout. Protarsomere 5 th with basal denticle very poorly developed. Parameres with slightly curved lateral margins.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE5FFD5FED86CB2DCDB3CFA.taxon	etymology	Etymology We dedicate this species to the memory of our colleague and entomologist Julián Hernández Cruz, who worked with the scarab beetle fauna of Oaxaca, mainly in the Sierra Sur where this species comes from.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE5FFD5FED86CB2DCDB3CFA.taxon	materials_examined	Type material Holotype MEXICO – Oaxaca • ♂; “ MEXICO: Oaxaca 12 Km W. El Camarón 22 June 1974 W. E. Steiner; // Strigoderma (Costatergus) juliani Andalco-Cid & Ramírez-Ponce, 2025 HOLOTYPE ” [red label, printed]; UNSM. Paratype MEXICO – Oaxaca • 1 ♀; “ Oax. nr Totolapan Aug. 5, 1965 Flint & Ortíz // STRIGODERMA COSTULIPENNIS BATES, Det. M. E. Jameson 1997; // Strigoderma (Costatergus) juliani Andalco-Cid & Ramírez-Ponce, 2025 PARATYPE ” [yellow label, printed]; UNSM.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE5FFD5FED86CB2DCDB3CFA.taxon	description	Description BODY SIZE. Total length: 7.50 mm. Maximum elytral width: 3.23 mm (Fig. 1 D). BODY COLOUR. Head, pronotum, scutellum and legs shiny black, with greenish reflections on clypeus and legs. Elytra ochre-yellow, with wide black bands longitudinally covering the outer third and a wide area adjacent to the suture (Fig. 9 A – B). Abdomen dark reddish black (Fig. 9 C). CLYPEUS. Sub-quadrangular, elongate. Anterior angles rounded; anterior edge broadly sinuate; outer margin weakly raised. Surface punctation broad, rough, deep and dense; points joined forming irregular transverse groove patterns (Fig. 2 D). FRONS. Sculpture rough-punctate, dense; defined points slightly smaller than on clypeus (Fig. 2 D). ANTENNA. Antennomere 3 longer than the preceding ones. Club slightly shorter than the pedicel and funiculus together (Fig. 9 D). PRONOTUM. Subrectangular, longer than wide; moderately widened in anterior half. Anterior angles projected. Surface without longitudinal median groove. Punctation small, very dense, homogeneous, fine (Fig. 2 D). Lateral figure markedly convex (lateral view) (Fig. 9 B). Scutellum triangular, wide, with rounded apex. Sculpture with punctation very small, defined, thick, deep and dense. ELYTRA. Subparallel to the 2 nd third. Surface with six elevated ridges with sharp edges; the 2 nd incomplete (diffuse in the middle portion) (Fig. 3 D); ridges five and six not converging to form one in its distal third (Fig. 4 D); intercostal space wide and uniform. Elytral suture elevated up to the 2 nd distal third. Humeral callus prominent, developed to the lateral margins of the elytra, interrupting the epipleura. Epipleura thick at the base, with a noticeable fold above the episternum, remainder of the structure flat and elevated. PYGIDIUM. Wider than long, with broadly rounded apex. Shape (in lateral view) noticeably convex in the middle part. Surface regular and uniform, with lateral depressions on the anterior portion. Sculpture with abundant, small, thick and deep punctation. METASTERNUM. Mesometasternal process somewhat broad, slightly projected; metasternum uniform and deeply punctate (Fig. 9 C). ABDOMEN. Sternites 1, 2 and 3 are almost the same size. Sternite 4 th as thick as any of the first three sternites plus one half as much as another. Surface with uniform punctation, separated by two puncture diameters (more dispersed on anal plate), almost rounded, bit deep (Fig. 9 C). LEGS. Protibia thick; distal denticle long, slightly curved, with rounded apex (Fig. 9 F). Protarsomeres 1 – 4 conspicuously short and broad, of similar size and thickness (Fig. 9 E). Arrangement of ventral setae: protarsomere 1 st and 2 nd with three very short to long, moderately thin to thin spines; protarsomere 3 rd and 4 th with three thick and very short to long spines. Protarsomere 5 slender, slightly longer than tarsomeres 1 – 4 together. The ventral lobe poorly developed, in medial-internal position (Fig. 9 E). Mesotibia g radually thickened towards the apex; apex with 9 – 10 medium spines. Surface with two transverse setiferous carinae; the upper one with 5 – 6 thick and moderately long spines; the lower one with 7 – 8 long spines. Sculpture with very deep and irregular punctation. Apical spurs straight: the inner one with acute apex, as long as the first two tarsomeres; the outer one with acute apex, of similar length to the first mesotarsomere. Mesotarsomeres 1 – 4 of similar length and thickness. Dorsal setae pattern: first mesotarsomere with 2 thick, moderately long spines; second and third mesotarsomere with 2 moderately long, moderately thick spines; fourth mesotarsomere with 2 thin setae. Ventral setae pattern: first, second, and third mesotarsomere with 4 thick, moderately long spines; fourth mesotarsomere with 4 moderately long, moderately thin spines. Metatibia clearly widened in the middle and apical part; apex with 11 – 12 medium spines. Surface with two transverse setiferous carinae: the proximal one with 4 – 5 thick spines; the distal one with 7 – 8 long spines. Sculpture with very deep and irregular punctation. Apical spurs straight: the inner one with a rounded apex, as long as the first two tarsomeres; the outer one short, very thick with a rounded apex, slightly shorter than the first mesotarsomere. Metatarsomeres dorsal setae pattern: fourth metatarsomere with 2 moderately long and thick spines; third metatarsomere with two moderately thin long setae; second metatarsomere with 2 long and thin setae. Ventral setae pattern: first metatarsomere with 4 long and thick spines; second metatarsomere with 3 long and thick spines and one seta; third metatarsomere with 2 spines and two setae. GENITALIA. Parameters long, with curved outer margins, narrower before the apex; rounded apices (Fig. 9 H – I). Apical figure (lateral view) slightly curved ventrally (Fig. 9 G). Female Similar to holotype except by: pronotum with denser and finer punctation; elytra entirely deep orange, base of fifth ridge extremely thickened; mesometasternal space considerably broader and metasternum longer; abdomen extremely inflated; protibia with longer and curved distal denticle; protarsomeres 1 – 4 thinner and inner protarsal claw like both rami similar in thickness and length.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE5FFD5FED86CB2DCDB3CFA.taxon	distribution	Distribution This species has been recorded in the Sierra Sur of the Oaxaca State, between 700 and 930 m a. s. l. (El Camarón and Totolapan). This is an area belonging to the Terrestrial Priority Region for Conservation “ Sierra Sur and Coast of Oaxaca ” (RTP- 129), characterized mainly by coniferous forests and medium altitude rainforest (Arriaga et al. 2000) (Fig. 14).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE5FFD5FED86CB2DCDB3CFA.taxon	biology_ecology	Natural history The feeding habits and life cycle of this species are unknown. The specimens were collected in June and August.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE5FFD5FED86CB2DCDB3CFA.taxon	discussion	Taxonomic comments This species is easy to distinguish as it is the only one with the 2 nd elytral ridges only marked at its ends, since the most closely related species, S. (C.) borealis sp. nov. (from Nayarit), has the 2 nd ridge complete along its entire length, although very thin. Other distinguishing characteristics are: internal protarsal claw with lower ramus less than half the length of the upper ramus (half the length in S. (C.) borealis), pronotum with broader punctation (denser and smaller in S. (C.) borealis), elytra with the yellowish area covering the entire length of the elytra (only the first anterior third in S. (C.) borealis), and long and thin parameres (short and wide in S. (C.) borealis).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE6FFD8FEFF6A96DA9F3E37.taxon	description	urn: lsid: zoobank. org: act: 23 C 176 C 5 - 52 F 5 - 4 EF 4 - B 71 E-D 49 F 99 C 9 EB 3 A Figs 1 E, 2 E, M, 3 E, 4 E, 5 D, 10, 14	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE6FFD8FEFF6A96DA9F3E37.taxon	diagnosis	Diagnosis Antennal club longer than scape, pedicel and funiculus together. Elytra with 10 elevated ridges; ridges 3, 5, 6 and 8 only marked at elytral base; complete ridges 5 th and 6 th (last external) convergent in distal third. 5 th protarsomere with developed internal preapical denticle. Internal protarsal claw with rhomboidal lower ramus, slightly longer than half the length of the upper ramus. Short parameres, with slightly curved lateral margins and widely rounded apex.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE6FFD8FEFF6A96DA9F3E37.taxon	etymology	Etymology From late Latin ‘ grossus ’ (‘ thick’), and Latin ‘ penis ’ (‘ tail, penis’), alluding to the notable thickness of the parameres in this species.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE6FFD8FEFF6A96DA9F3E37.taxon	materials_examined	Type material Holotype MEXICO – Guerrero • ♂; “ México, Guerrero, Techan de Galeana, Piedra de Tlacoyunque km. 50 Acapulco - Zihuatanejo 17 º 15 ’ 34 ’’ N, 100 º 59 ’ 55 ’’ O, 006 msnm 19 - XI- 2006, G. Ortega, L. Cervantes, C.; // 45234 “ QR code ”; // IBUNAM: CNIN: CO 45234 “ BAR CODE ”; // Colección del Instituto de Biología, UNAM. México, D. F.; // Strigoderma (Costatergus) grossipenis Andalco-Cid & Ramírez-Ponce, 2025 HOLOTYPE ” [red label, printed]; CNIN. Paratype MEXICO – 1 ♂; “ Genus N. Mexico; // Strigoderma costulipennis Bates [illegible]; // 500; // 6745; // Strigoderma (Costatergus) grossipenis Andalco-Cid & Ramírez-Ponce, 2025 PARATYPE ” [yellow label, printed]; BMNH.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE6FFD8FEFF6A96DA9F3E37.taxon	description	Description BODY SIZE. Total length: 6.5 mm. Maximum elytral width: 2.96 mm (Fig. 1 E). BODY COLOUR. Head and legs shiny coppery black; frons and clypeus with greenish reflections; rest of body completely black with greenish reflections (Fig. 10 A – C). CLYPEUS. Subrectangular, broad. Anterior angles broadly rounded; anterior edge almost straight; external margin well elevated. Surface with a latero-apical excavation on each side. Punctation large, rough, deep and dense (Fig. 2 E). FRONS. Convex. Sculpture roughly-punctate, similar to that of the clypeus (Fig. 2 E). ANTENNA. Pedicel slightly longer than the 1 st and 2 nd flagellomere. Club as long as the scape, pedicel and funiculus together. Pedicel slightly longer than the 1 st and 2 nd flagellomere (Fig. 10 D). PRONOTUM. Subtrapezoidal, clearly broader than long; noticeably widened in anterior half. Short anterior angles. Surface without longitudinal median groove. Punctation small, very dense, homogeneous and fine (Fig. 2 M). Lateral figure markedly convex (in lateral view) (Fig. 10 B). SCUTELLUM. Rounded, with thickened edge. Sculpture with punctation slightly less wide, denser and deeper than that of the pronotum. ELYTRA. Subparallel to the 2 nd third. Surface with 10 elevated ridges with rounded edges; the 3 rd, 5 th, 6 th and 8 th incomplete (only evident at the elytral base) (Fig. 3 E); ridges nine and ten join together as one on the distal third (Fig. 4 E); intercostal space slightly rough. Elytral suture elevated up to the 2 nd distal third. Humeral callus prominent, developed to the anterior and lateral margins of the elytra, interrupting the epipleura. Epipleura interrupted at the base by the humeral callus but continuous flat and elevated. METASTERNUM. Metaventral process broad, slightly projecting; metasternum uniform and deeply punctate (Fig. 10 C). ABDOMEN. Sternites 1, 2 and 3 almost equal in length. Surface with uniform, dense and deep punctation, separated by one puncture diameter (more dispersed on anal plate), sub-oval transverse (Fig. 10 C). LEGS. Protibia thick; distal denticle long, slightly curved, with rounded apex. Protarsomeres 1 – 4 noticeably short and broad, successively slightly broader (Fig. 10 E). Arrangement of ventral setae: 3 spines on each protarsomere, from very short to long, and a thin and long outer seta. Protarsomere 5 th robust, broad, almost equal to tarsomeres 1 – 4 together. Ventral lobe well developed, in subapical-internal position. The inner protarsal claw broad and deeply notched: lower ramus rhomboidal, reaching half the length of upper ramus (Fig. 10 E). Mesotibia thickened towards the middle; apex with 10 – 11 short spines. Surface with two transverse setiferous carinae; the upper one with 4 to 5 thick and long spines; the lower one with 6 – 7 long spines. Sculpture with moderate, deep and irregular punctation. Apical spurs straight: the inner one with acute apex, as long as the first two tarsomeres; the outer one with acute apex, of similar length to the first mesotarsomere. Mesotarsomeres 1 – 4 short and broad, slightly thickened towards the apex. Dorsal setae pattern: first three tarsomeres with 3 long, moderately thick spines and the fourth mesotarsomere with two very thin and very long setae. Ventral setae pattern: all tarsomeres with 4 spines, on each tarsomere the arrangement of spines is moderately long to long and thick. Metatibia clearly widened towards the apical part; apex with 12 – 13 moderately short spines. Surface with three transverse setiferous carinae: the proximal one with 2 – 3 thick spines; the middle one with 5 – 6 moderate spines, the distal one with 8 – 9 moderately long spines. Sculpture with large and irregular punctation, like an impression. Apical spurs straight: the inner one with a rounded apex, as long as the first one plus part of the second tarsomere; the outer one with a rounded apex, slightly shorter than the first mesotarsomere. Metatarsomeres thickness increases gradually towards apex; length is variable, but second metatarsomere is prominent. Dorsal setae pattern: first and second metatarsomere with two moderately long spines, the inner one slightly thinner than the outer one; third metatarsomere with two moderately long and moderately thick spines; fourth metatarsomere with one long, thin setae. Ventral setae pattern: four long spines on each tarsomere, but their thickness decreases successively on each tarsomere. GENITALIA. Parameres short, with the outer margin slightly curved, slightly narrower before the apex; apices broadly rounded (Fig. 10 H – I). Apical form (lateral view) subtly curved ventrally (Fig. 10 G). VARIATION. The reflections on head, pronotum and forelegs are less intense. Other features are not apparent due to the age and condition of the specimen. Female Unknown.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE6FFD8FEFF6A96DA9F3E37.taxon	distribution	Distribution This species has only been recorded from the type locality, in the Sierra Sur of the state of Guerrero, on the Acapulco-Zihuatanejo highway (Municipality Tecpan de Galeana). This area belongs to the “ Sierra Sur de Guerrero ” Terrestrial Priority Conservation Region (RTP- 117, Arriaga et al. 2000), characterized by being an isolated area of high endemism and species richness. The locality comprises medium and low altitude forests, although pine forests, oak forests and cloud forests are also found in the region (Fig. 14).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE6FFD8FEFF6A96DA9F3E37.taxon	biology_ecology	Natural history Feeding habits and life cycle are unknown. The holotype was collected in November.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFE6FFD8FEFF6A96DA9F3E37.taxon	discussion	Taxonomic comments This species is most similar to the nominal species, S. (C.) costulipennis, both from the state of Guerrero. They share the maximum number of elytral ridges and are the only ones with the two external ridges convergent in the apical third (besides S. (C.) vulcanica sp. nov.). However, they differ in the presence of ridge 6 in the basal area of the elytra (absent in S. (C.) costulipennis), fine and very dense punctation (wider and thicker in S. (C.) costulipennis), the wide intercoxal space of the mesosternum (narrow in S. (C.) costulipennis), in addition to the short, wide parameres, with almost straight external margins (thin and long parameres, widely curved on the external margins in S. (C.) costulipennis).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEAFFDFFEF36E43DD583E93.taxon	description	urn: lsid: zoobank. org: act: BB 76569 E- 0 C 28 - 4081 - B 960 - 7 E 530 B 779529 Figs 1 F, 2 F, N, 3 F, 4 F, 11, 14	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEAFFDFFEF36E43DD583E93.taxon	diagnosis	Diagnosis Simple inner protarsal claw. Elytra with inter-costal space markedly punctate.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEAFFDFFEF36E43DD583E93.taxon	etymology	Etymology From the Latin numerical prefix ‘ uni ’ (‘ one’), and from the Latin ‘ ungula ’ (‘ claw, spur or hoof’), in reference to the fact that this species is the only one with a simple internal protarsal claw.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEAFFDFFEF36E43DD583E93.taxon	materials_examined	Type material Holotype MEXICO – Guerrero • ♂; “ 8 miles sw. Iguala, gro., Mexico. VII- 10 - 66. P. M. & P. K. Wagner; // T A & M; // TAMU – ENTO X 0043336 “ barcode ”; // Strigoderma costulipennis Bates., Det. A. Bader 1989; // Strigoderma (Costatergus) uniungula Andalco-Cid & Ramírez-Ponce, 2025 HOLOTYPE ” [red label, printed]; TAMU.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEAFFDFFEF36E43DD583E93.taxon	description	Description BODY SIZE. Total length: 7.17 mm. Maximum elytral width: 3.30 mm (Fig. 1 F). BODY COLOUR. Antennal club, pronotum, scutellum and abdomen black with coppery-greenish reflections, apex of protibia and tarsi slightly reddish (Fig. 11 A – C). CLYPEUS. Markedly trapezoidal, elongate. Anterior angles rounded; anterior edge sinuate; external margin slightly elevated. Surface with shallow anterolateral excavation on each side. Punctation very broad, rough, deep and dense (Fig. 2 F). FRONS. Slightly convex, with a lateral depression at the base of the suture. Sculpture roughly-punctate, similar to the clypeus (Fig. 2 F). ANTENNA. 1 st to 3 rd flagellomeres similar in length and thickness; club slightly longer than the pedicel and funiculus together (Fig. 11 D). PRONOTUM. Shape sub-quadrangular, nearly as wide as long; broadened on anterior half. Anterior angles slightly projecting. Surface without longitudinal median groove. A cluster of 3 – 5 setae on each side of anterior half. Mixed punctation; larger and sparser toward anterior margin and smaller and denser toward posterior margin; heterogeneous (Fig. 2 N). Lateral figure markedly convex (side view) (Fig. 11 B). SCUTELLUM. Rounded with thickened edge. Sculpture with punctation less wide than that of the pronotum, denser and deeper. ELYTRA. Subparallel to the 2 nd third. Surface with seven elevated ridges; all complete, except the third one developed to the elytral half (Fig. 3 F). Ridges six and seven not convergent in the distal third to form a single one (Fig. 4 F). Space between ridges notable and irregularly punctate, wide and poorly defined points. Elytral suture elevated to the 2 nd distal third. Humeral callus prominent, developed to the anterior and lateral margins of the elytra, interrupting the epipleura. Epipleura interrupted at the base by the humeral callus but continuous, flat and elevated. METASTERNUM. Metaventral process narrow, slightly projecting; metasternal process uniform and deeply punctate (Fig. 11 C). ABDOMEN. Sternites 1, 2 and 3 sub-equal in length. Surface with uniform punctation, separated by one puncture diameter (more dispersed on anal plate), sub-rounded, shallow (Fig. 11 C). LEGS. Protibia very thick; distal denticle long, almost straight, with acute apex (Fig. 11 F). Protarsomeres 1 – 4 noticeably short and broad, successively slightly broader (Fig. 11 E). Arrangement of ventral setae: 3 spines on each protarsomere with very short to long arrangement and a long thin outer seta. Protarsomere 5 somewhat thin, longer than tarsomeres 1 – 4 together. Ventral lobe very poorly developed, in sub-basalinternal position. Inner protarsal claw simple, not split (Fig. 2 C). Mesotibia thickened towards the middle; apex with 10 – 11 short spines. Surface with two transverse setiferous carinae; the upper one with 4 – 5 thick and long spines; the lower one with 6 – 7 long spines. Sculpture with moderate, deep and irregular punctation. Apical spurs straight: the inner one with acute apex, as long as the first two tarsomeres; the outer one with acute apex, of similar length to the first mesotarsomere. Mesotarsomeres 1 – 4 short and wide, slightly thickened towards the apex. Dorsal setae pattern: first three tarsomeres with 3 long, moderately thick spines; fourth mesotarsomere with two very thin and very long setae. Ventral setae pattern: all tarsomeres with 4 long spines, progressively longer and thicker. Metatibia clearly widened towards the apical part; apex with 12 – 13 moderately short spines. Surface with three transverse setiferous carinae: the proximal one with 2 – 3 thick spines; the middle one with 5 – 6 moderate spines, the distal one with 8 – 9 moderately long spines. Sculpture with large and irregular punctation, like an impression. Apical spurs straight: the inner one with a rounded apex, as long as the first one plus part of the second tarsomere; the outer one with a rounded apex, slightly shorter than the first mesotarsomere. Metatarsomeres slightly thicker towards the apex; length is variable, but the second one stands out. Dorsal setae pattern: first and second metatarsomere with two moderately long spines, the inner one slightly thinner than the outer one; third metatarsomere with two moderately long and moderately thick spines; the fourth metatarsomere with a long, thin setae. Ventral setae pattern: each metatarsomere with four long spines, the thickness of which decreases progressively. GENITALIA. Parameres long and slender, with the outer margin slightly curved, slightly narrower before the apex; rounded apices (Fig. 11 H – I). Apical figure (lateral view) slightly curved ventrally (Fig. 11 G). Female Unknown.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEAFFDFFEF36E43DD583E93.taxon	distribution	Distribution This species has only been recorded from the type locality, Iguala, Guerrero (650 m a. s. l.), characterized by low deciduous forest vegetation. This area belongs to the “ Sierras de Taxco-Huautla ” Terrestrial Priority Region for Conservation (RTP- 120, Arriaga et al. 2000), characterized as a biological corridor with high ecological integrity and high biological richness (Fig. 14).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEAFFDFFEF36E43DD583E93.taxon	biology_ecology	Natural history Feeding habits and life cycle are unknown. The holotype was collected in July.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEAFFDFFEF36E43DD583E93.taxon	discussion	Taxonomic comments This species is easy to distinguish as it is the only one with a simple inner protarsal claw. It shares with S. (C.) paucipunctata sp. nov. the same intercostal area with wide punctation, however, they are differentiated by the more marked intercostal punctation (shallow in S. (C.) paucipunctata), as well as the pronotum with the densest punctation and with coppery reflections (pronotum with less dense punctation and intense greenish reflections in S. (C.) paucipunctata).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEFFFE3FEF66E43DCDC3CC2.taxon	description	urn: lsid: zoobank. org: act: 5 FBF 7775 - AB 91 - 4 D 47 - 80 F 2 - 595 DB 0 D 5995 A Figs 1 G, 2 G, Ñ, 3 G, 4 G, 12, 14	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEFFFE3FEF66E43DCDC3CC2.taxon	diagnosis	Diagnosis Elytra with eight elevated ridges; ridges 3 rd, and 5 th only marked at base; ridges seven and eight convergent in distal third. 5 th protarsomere with barely perceptible preapical denticle. Inner protarsal claw with lanceolate lower ramus, longer than half the length of upper ramus. Short parameres, with almost straight outer margin, and broad, widely rounded apices.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEFFFE3FEF66E43DCDC3CC2.taxon	etymology	Etymology The specific epithet refers to the locality site where the type series was collected, “ Volcán de Colima ”. From the Latin ‘ Vulcānus ’ (Vulcan: Roman god of fire; ‘ volcano’), and the noun suffix ‘ ica ’ (to denote the geographic region where the species is found); vulcanica, inhabitant of the volcano.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEFFFE3FEF66E43DCDC3CC2.taxon	materials_examined	Type material Holotype MEXICO – Colima • ♂; “ C. C. Hoffman, Cólima, VII- 1916, 023; // Strigoderma costulipennis Bates 1888, Det. A. Bader, Coll. Change; // Field Museum // Strigoderma (Costatergus) vulcanica Andalco Cid & Ramírez Ponce, 2021 HOLOTYPE ♂ ” [red label, printed]; CNHM. Paratypes (6 ♂♂, 6 ♀♀) MEXICO – Colima • 3 ♂♂; “ México, Coll. Lange (handwritten); // Specimen orig. without labels in colln. but associated with specimen bearing attached data; // Strigoderma costulipennis Bates, Det. A. Bader 1989; // CNHM 1955, Karl Brancsik Colln. ex Eduard Knirsch; // Strigoderma (Costatergus) vulcanica Andalco Cid & Ramírez Ponce, 2024 PARATYPE ♂ ” [yellow label, printed]; CNHM • 1 ♂; “ Colima, Colima. MEX. Mex., VIII-I- 41; // Col. & pres by. C. H. Seevers; // Strigoderma costulipennis Bates 1888, Det. A. Bader 1989; // Strigoderma (Costatergus) vulcanica Andalco Cid & Ramírez Ponce, 2024 PARATYPE ♂ ” [yellow label, printed]; CNHM • 1 ♀; “ México, Coll. Lange // CNHM 1955, Karl Brancsik Colln. ex Eduard Knirsch // Strigoderma (Costatergus) vulcanica Andalco Cid & Ramírez Ponce, 2024 PARATYPE ♀ ” [yellow label, printed]; CNHM • 2 ♀♀; “ Colima Vulcano Mex / L. Conrad // USNM // Ohaus determ. Strigoderma costulipennis ♀ W. Bts. // Strigoderma costulipennis Bates, Det. A. Bader 1989 // Strigoderma (Costatergus) vulcanica Andalco Cid & Ramírez Ponce, 2024 PARATYPE ♀ ” [yellow label, printed]; USNM • 1 ♀; “ Colima, Col. Mex. Conradt // USNM // Ohaus determ. Strigoderma costulipennis ♀ W. Bts. // Strigoderma costulipennis Bates, Det. A. Bader 1989 // Strigoderma (Costatergus) vulcanica Andalco Cid & Ramírez Ponce, 2024 PARATYPE ♀ ” [yellow label, printed]; USNM • 1 ♂; “ C. C. Hoffman, Colima, VII, 1916 // C. C. Hoffman, det. 023 // IBUNAM: CNIN: CO 45138 (barcode) // 45138 (QR code) // Strigoderma (Costatergus) vulcanica Andalco Cid & Ramírez Ponce, 2024 PARATPE ♂ ” [yellow label printed]; CNIN • 1 ♂; “ C. C. Hoffman Colima VII- 1916 // IBUNAM: CNIN: CO 45127 (barcode) // 45127 (QR code) // Strigoderma (Costatergus) vulcanica Andalco Cid & Ramírez Ponce, 2024 PARATYPE ♂ ” [yellow label, printed]; CNIN • 1 ♀; “ C. C. Hoffman Colima VII- 1916 // IBUNAM: CNIN: CO 45129 (barcode) // 45129 (QR code) // Strigoderma (Costatergus) vulcanica Andalco Cid & Ramírez Ponce, 2024 PARATYPE ♀ [yellow label, printed]; CNIN • 1 ♀; “ C. C. Hoffman Colima VII- 1916 // IBUNAM: CNIN: CO 45130 (barcode) // 45130 (QR code) // Strigoderma (Costatergus) vulcanica Andalco Cid & Ramírez Ponce, 2024 PARATYPE ♀ ” [yellow label printed]; CNIN.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEFFFE3FEF66E43DCDC3CC2.taxon	description	Description BODY SIZE. Total length: 6.31 mm. Maximum elytral width: 2.64 mm (Fig. 1 G). BODY COLOUR. Head, pronotum, scutellum black with coppery reflections. Elytra, legs, scape and pedicel intense yellow (Fig. 12 A – B). Abdomen dark reddish black (Fig. 12 C). CLYPEUS. Subtrapezoidal, broad; anterior angles broadly rounded; anterior edge sinuates; margin slightly raised. Surface with a noticeable latero-apical excavation on each side. Punctation large, rough, deep and dense (Fig. 2 O). FRONS. Convex. Sculpture with punctation slightly less dense than that of the clypeal (Fig. 2 O). ANTENNA. 1 st to 3 rd flagellomeres similar in length and thickness. Club slightly longer than pedicel and funiculus together (Fig. 12 D). PRONOTUM. Subtrapezoidal, distinctly broader than long; markedly widened in anterior half. Anterior angles short. Surface without longitudinal median groove. Punctation small, dense, somewhat heterogeneous, almost rough (Fig. 2 O). Lateral figure moderately convex (side view) (Fig. 12 B). SCUTELLUM. Wide. Sculpture with wide and deep punctation similar to the pronotum. ELYTRA. Subparallel to 2 nd third. Surface with eight elevated ridges; 3 rd, 4 th and 6 th incomplete (only marked at base of elytra) (Fig. 3 G); ridges 8 and 9 convergent, forming only one in distal third (Fig. 4 G). Elytral suture elevated to distal 2 nd third. Humeral callus discrete, developed to lateral margins of elytra, interrupting epipleura. Epipleura thin and elevated. PYGIDIUM. Broader than long, with a broadly rounded apex. Form (side view) noticeably convex in the median part. Sculpture punctation dense, oblique, small and deep. Vestiture with sparse short, thin, transparent setae over the apical area. METASTERNUM. Metaventral process broad, slightly projecting; metasternal dense and deeply punctate (Fig. 12 C). ABDOMEN. Sternites 1, 2, and 3 almost the same size. Sternite 4 as thick as the third sternites plus a part of any of the first three. Surface with uniform and small punctation, separated by two puncture diameters (more dispersed on anal plate), sub-rounded and shallow (Fig. 12 C). LEGS. Protibia little thick; distal denticle long, slightly curved, with rounded apex (Fig. 12 F). Protarsomeres 1 – 4 noticeably short and somewhat widened, progressively slightly longer (Fig. 12 E). Arrangement of ventral setae: protarsomeres 1 and 2 with 3 moderately thin spines and one seta; protarsomeres 3 and 4 with three short to long, stout spines. Protarsomere 5 slender, longer than tarsomeres 1 – 4 together. Ventral lobe barely visible, in inner sub-apical position. Inner protarsal claw deep and narrowly cleft: lower ramus lanceolate, half the length of the upper ramus (Fig. 12 E). Mesotibia gradually thickened in the middle; apex with 12 – 13 short spines. Surface with two transverse setiferous carinae; the upper one with 3 – 4 thick and short spines; the lower one with 6 – 7 long spines. Surface with large and irregular punctate print-like markings. Apical spurs straight: the inner one with acute apex, as long as the first one plus part of the second tarsomere; the outer one with acute apex, of similar length to the first mesotarsomere. Mesotarsomeres 1 – 4 similar in length and thickness. Dorsal setae pattern: mesotarsomeres 1 with 2 thin spines, one short and one very long; mesotarsomeres 2, 3 and 4 with 2 thin and long setae. Ventral setae pattern: mesotarsomeres 1, 2 and 3 with 3 spines that are very thick to moderately thick, short to long; mesotarsomere 4 with 2 moderately thin and moderately long spines and 1 seta. Metatibia clearly widened in the middle and apical part; apex with 14 – 15 short spines. Surface with three transverse setiferous carinae: the proximal one with 2 – 3 moderately thick and moderately long spines; the middle one with 4 – 5 moderate spines; the distal one with 8 – 9 moderately thick and moderately long spines. Apical spurs straight: the inner one with rounded apex, as long as the first two tarsomeres; the outer one with rounded apex, slightly longer than the first mesotarsomere. Metatarsomeres with similar thickness and successively decreasing length. Dorsal setae pattern: metatarsomeres 1 to 3 with 1 moderately thick spine and 1 moderately thin seta; metatarsomere 4 with two long, thin setae. Ventral setae pattern: metatarsomeres 1 and 2 with 4 long, thick spines; metatarsomere 3 with 3 spines and two long, moderately thick setae; metatarsomere 4 with 2 moderately long, thick spines and 2 setae. GENITALIA. Parameres short, with the outer margin slightly curved, slightly narrower before the apex; apices broadly rounded (Fig. 12 H – I). Apical figure (lateral view) slightly curved ventrally (Fig. 12 G). VARIATION. Pronotum with slightly more or less dense punctation and coppery or greenish reflections. Elytra completely bright yellow or with a complete longitudinal stripe covering from the penultimate ridge and another central longitudinal spot. Legs bright-yellow, ochre-yellow or black. Female Similar to holotype, except by: antennal club slightly shorter. Pronotum with finer punctation, less dense and deeper posteriorly; external elytral ridge noticeably wider, mainly at base; penultimate elytral ridge poorly developed. Abdomen noticeably inflated. Protibia with more elongated and curved distal denticle; thinner protarsi and internal protarsal claw with lower and upper ramus similar in thickness and length.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEFFFE3FEF66E43DCDC3CC2.taxon	distribution	Distribution This species is only known from Colima (‘ volcan’). It was probably collected on the periphery of the geological body where dry tropical vegetation exists (Fig. 14).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEFFFE3FEF66E43DCDC3CC2.taxon	biology_ecology	Natural history Feeding habits and life cycle are unknown. Specimens were collected in July and August.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFEFFFE3FEF66E43DCDC3CC2.taxon	discussion	Taxonomic comments This species presents the greatest similarity with S. (C.) costulipennis and S. (C.) grossipenis sp. nov., since they share the convergent arrangement in the distal third of the external elytral ridges, however, they differ in the presence of eight elytral ridges (10 in S. (C.) grossipenis); internal protarsal claw with both rami very close (clearly separated in S. (C.) costulipennis and S. (C.) grossipenis); and short and thick parameres (thin in S. (C.) costulipennis).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFD0FFE7FEE06AFFDDA73E6C.taxon	description	urn: lsid: zoobank. org: act: 51 F 8 CD 71 - CF 51 - 4 AE 4 - BA 94 - 3 B 173 C 7323 FA Figs 1 H, 2 H, O, 3 H, 4 H, 13 – 14	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFD0FFE7FEE06AFFDDA73E6C.taxon	diagnosis	Diagnosis Pronotum finely and densely punctate. Elytral ridges 3, 5 and 7 absent, 2 nd complete but subtly defined. Inner protarsal claw with lower ramus reaching more than half the length of upper ramus. Parameres very short and broad.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFD0FFE7FEE06AFFDDA73E6C.taxon	etymology	Etymology From the Latin adjective ‘ borealis ’ (‘ relating to the north or boreas’), as this species has the most northerly distribution of any species described so far.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFD0FFE7FEE06AFFDDA73E6C.taxon	materials_examined	Type material Holotype MEXICO – Nayarit • ♂; “ Jesús María Nayarit, MEX. VII- 6 - 1955 // B. Malkin Collector // SCARABAEIDAE: ANOMALINAE Strigoderma costulipennis Bates ♂, Det: R W L Potts, 1977 // Strigoderma (Costatergus) borealis Andalco-Cid & Ramírez-Ponce, 2025 HOLOTYPE ♂ ” [red label, printed]; EMEC. Paratypes (4 ♂♂, 2 ♀♀) MEXICO – Nayarit • 1 ♂, 1 ♀; “ Jesús María Nayarit, MEX. VII- 6 - 1955 // B. Malkin Collector // UCB // UC Berkeley EMEC 1324373 (female specimen with 1324376) “ QR code ” // Strigoderma costulipennis Bates, Det. A Bader 1989 // Strigoderma (Costatergus) borealis Andalco-Cid & Ramírez-Ponce, 2025 PARATYPE ” [yellow label, printed]; EMEC • 1 ♂; “ Arroyo Canaveral nr. Jesus Maria Nay. MEX. VII- 15 - 55 // B. Malkin Collector // UC Berkeley EMAC 1324369 “ QR code ” // Strigoderma (Costatergus) borealis Andalco-Cid & Ramírez-Ponce, 2025 PARATYPE ♂ ” [yellow label, printed]; EMEC • 1 ♂; “ Arroyo Canaveral nr. Jesus Maria Nay. MEX. VII- 15 - 55 // B. Malkin Collector // UC Berkeley EMAC 1324370 “ QR code ” // Strigoderma costulipennis Bates, Det. H. F. HOWDEN // Strigoderma (Costatergus) borealis Andalco-Cid & Ramírez-Ponce, 2025 PARATYPE ♂ ” [yellow label, printed]; EMEC • 1 ♀; “ MEX. Nay. Jesús Maria VII- 27 - 55 / B. Malkin Collector // UC Berkeley EMEC 1324372 “ QR code ” // UCB // Strigoderma costulipennis Bates, Det. A Bader 1989 // Strigoderma (Costatergus) borealis Andalco-Cid & Ramírez-Ponce, 2025 PARATYPE ♀ ” [yellow label, printed]; EMEC • 1 ♂; “ Arroyo Santiago nr. Jesús Maria Nay. MEX. VII- 5 - 55 // B. Malkin Collector // UCB // UC Berkeley EMEC 1324371 “ QR code ” // Strigoderma costulipennis Bates, Det. A Bader 1989 // Strigoderma (Costatergus) borealis Andalco-Cid & Ramírez-Ponce, 2025 PARATYPE ♂ ” [yellow label, printed]; EMEC.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFD0FFE7FEE06AFFDDA73E6C.taxon	description	Description Male holotype BODY SIZE. Total length: 6.56 mm. Maximum elytral width: 2.95 mm (Fig. 1 H). BODY COLOUR. Body completely black, except for a wide orange spot on the first third of each elytra; tibiae apically reddish and tarsi and claws with deeper reddish tone (Fig. 13 A – C). CLYPEUS. Subrectangular, broad. Anterior angles broadly rounded; anterior edge slightly sinuate; outer margin weakly raised. Surface with a shallow anterolateral excavation on each side. Punctation very wide, rough, deep and dense (Fig. 2 H). FRONS. Almost flat. Rough-punctate sculpture, similar to that of the clypeal in the central area (Fig. 2 H). ANTENNA. 3 rd flagellomere longer than the previous ones. Club almost as long as the pedicel and funiculus together (Fig. 13 D). PRONOTUM. Subtrapezoidal, broader than long; markedly widened in anterior half. Anterior angles short, and lateral margins broadly rounded. Surface without longitudinal median groove. Punctation small, very dense, homogeneous, fine (Fig. 2 P). Lateral figure markedly convex (side view) (Fig. 13 B). SCUTELLUM. Sub-rounded with thickened edge. Sculpture with a width slightly less than that of the pronotum, denser and deeper. ELYTRA. Subparallel to 2 nd third. Surface with seven elevated ridges; 3 rd ridge only marked at base of elytra (Figs 3 H, 4 H). Inter-costal space very wide, subtly irregular, without defined punctation. Elytral suture elevated to distal 2 nd third. Humeral callus prominent, developed to anterior and lateral margins of elytra, interrupting epipleura. Epipleura interrupted at base by humeral callus but continuous, flat and elevated. METASTERNUM. Metaventral process broad, slightly projecting; metasternum scattered and shallowly punctate (Fig. 13 C). ABDOMEN. Sternites 1, 2 and 3 almost equal in length. Sternite 4 as thick as the third sternites plus ½ of any of the first three. Surface with uniform punctation, separated by one puncture diameter (more dispersed on anal plate), oval transverse, broad and shallow (Fig. 13 C). LEGS. Protibia very thick; distal denticle long, almost straight, with rounded apex (Fig. 13 F). Protarsomeres 1 – 4 noticeably short and broadened, successively slightly broader (Fig. 13 E). Arrangement of ventral setae: protarsomeres with 3 spines, very short to long and a long thin outer seta. 5 th protarsomere slightly thin, longer than tarsomeres 1 – 4 together. The ventral lobe developed, in internal sub-apical position. Inner protarsal claw narrow and shortly cleft: lower ramus lanceolate, reaching more than half the length of upper ramus (Fig. 13 E). Mesotibia thickened towards the middle; apex with 10 – 11 short spines. Surface with two transverse setiferous carinae; the upper one with 4 – 5 thick and long spines; the lower one with 6 – 7 long spines. Sculpture with moderate, deep and irregular punctation. Apical spurs straight: the inner one with acute apex, as long as the first two tarsomeres; the outer one with acute apex, of similar length to the first mesotarsomere. Mesotarsomeres 1 – 4 short and broad, slightly thickened towards the apex. Dorsal setae pattern: first three tarsomeres with 3 long, moderately thick spines and the fourth tarsomere with two very thin and very long setae. Ventral setae pattern: tarsomeres with 4 spines, on each tarsomere the arrangement of spines is moderately long to long and thick. Metatibia clearly widened towards the apical part; apex with 12 – 13 moderately short spines. Surface with three transverse setiferous carinae: the proximal one with 2 – 3 thick spines; the middle one with 5 – 6 moderate spines, the distal one with 8 – 9 moderately long spines. Sculpture with large and irregular punctation, like an impression. Apical spurs straight: the inner one with a rounded apex, as long as the first one plus part of the second tarsomere; the outer one with a rounded apex, slightly shorter than the first mesotarsomere. Metatarsomeres thickness increases gradually towards apex; length is variable, but the second is prominent. Dorsal setae pattern: first and second metatarsomeres with two moderately long spines, the inner one slightly thinner than the outer one; third metatarsomere with two moderately long and moderately thick spines; fourth metatarsomere with one long, thin seta. Ventral setae pattern: four long spines on each tarsomere, but the thickness of these decreases successively with the number of tarsomeres. GENITALIA. Parameres very short, with the outer margin slightly curved, slightly narrower before the apex; apices broadly rounded (Fig. 13 H – I). Apical figure (lateral view) curved ventrally (Fig. 13 G). VARIATION. There is little variation in the size of the pronotal punctures. The colour of the pronotum may have intense greenish reflections, and the orange spots on the anterior part of the elytra may be absent, or only faintly marked on the elytral margin. Female Similar to holotype except for the following differences: head with antennal club shorter. Pronotum punctuation much smaller and less dense, mainly towards the posterior portion, where there are smooth areas close to the margin. Elytra with outer ridge strongly thickened across its length. Abdomen noticeably inflated. Anterior protibia with longer and curved distal denticle. Protarsomeres 1 – 4 thinner; inner protarsal claw with the upper ramus slightly longer, but both of similar thickness.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFD0FFE7FEE06AFFDDA73E6C.taxon	distribution	Distribution This species has only been collected at the type locality, Jesús María Nayarit (approximately 500 m a. s. l.), in the north of the state (Fig. 14).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFD0FFE7FEE06AFFDDA73E6C.taxon	biology_ecology	Natural history Its feeding habits and life cycle are unknown. Adults were collected in June and July.	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
03EB87BCFFD0FFE7FEE06AFFDDA73E6C.taxon	discussion	Taxonomic comments This species has the greatest morphological similarity to S. (C.) tenebrosa and S. (C.) juliani sp. nov., since they are characterized by having ridges 3, 5, and 7 (also ridge 2 in the case of S. (C.) juliani) absent or only marked at the base of the elytra, such that the defined ridges appear very separated, in addition to the unpunctuated inter-costal space. However, S. (C.) borealis sp. nov. is distinguished by having the lower ramus of the inner protarsal claw clearly longer than half the length of the upper ramus (shorter than half the upper ramus in S. (C.) tenebrosa and S. (C.) juliani), and very short and wide parameres at the apex (long and thin in S. (C.) tenebrosa and S. (C.) juliani).	en	Andalco-Cid, Karla Selene, Zaragoza-Caballero, Santiago, Estrada Torres, Arturo, Rodríguez-Mirón, Giovanni, Ramírez-Ponce, Andrés (2026): Description of a new Mexican endemic subgenus of Strigoderma Burmeister with six new species (Coleoptera, Melolonthidae, Anomalini). European Journal of Taxonomy 1052: 21-61, DOI: 10.5852/ejt.2026.1052.3265, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3265/14433
