taxonID	type	description	language	source
03EA8F4FFFF1F91BFF0FFC41FAD4FA6D.taxon	description	(Figs. 1 - 6; Pl. I, Figs. I and II)	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
03EA8F4FFFF1F91BFF0FFC41FAD4FA6D.taxon	description	Description: Male (Fig. 1; Pl. I, Figs. I and II). Body length: 7.52, width: 2.31. Body (Fig. 1; Pl. I, Fig. I) black, surface shining. Setae bicoloured, glossy white and brown-black, thick, curved. Head (Fig. 1) black, punctation simple. Frons broader than long, concave. Frontal carina faintly impressed. Vertex slightly concave. Eye round with distinct facets. Labrum black, raised, broader than long, anterior margin convex. Mandible black, dentate, apically subpointed. Antennae brown, extending to the middle of the pronotum. First segment robust, longest. Third smallest. Fourth onward serrate and subequal in length. The last segment constricted near the apex. Pronotum (Fig. 1) black, wider than long, disc strongly convex, punctation (Fig. 2) simple. Submarginal line present. Anterior margin nearly straight, with a round anterior angle. Lateral margin entire, slightly convex, sinuate before hind angle. Hind angle subpointed. Scutellum (Fig. 1) pentagonal, black, flat; basally arcuate, depressed in middle. Elytra (Fig. 1) 2.09 × longer than pronotum; black, striae punctate, interstriae flat, punctation (Fig. 3) simple; basally emarginate; blunt apically. Prosternum black, convex, with distinct chin piece; punctation simple. Pronotosternal suture concave, broadly open to little beyond the middle of the prosternum. Hypomeron strongly grooved for accommodating protarsi. Prosternal spine long, emarginated, apically round. Mesepisternum and mesepimeron do not form part of the margin of the mesocoxal cavity. Metasternum strongly grooved for receiving mesotarsi. Metacoxal plate slightly round at middle. Abdomen black, punctation simple. First four ventrites with patches of setae. Legs (Fig. 4) brown; tarsal claw (Fig. 5) simple, setae present at base. Genitalia (Fig. 6; Pl. I, Fig. II): Basal piece wide, arms acute at apex, anterior margin ‘ V’ shape, posterior margin nearly straight, lateral margin arcuate, outer margin strongly sclerotised, thin and line like, rest feebly sclerotised; median lobe slightly longer than parameres, uniformly sclerotised throughout, arms of median lobe long, acute, round at base, not exceeding ventral posterior margin of parameres, gradually narrowing from base to apex, apex rounded, furcae not reaching the anterior margin of parameres; outer lateral margin of parameres from basal to distal and inner lateral margin throughout strongly sclerotised, outer lateral margin sinuate, basally broad, medially concave, distally highly concave, forming posteriorly directed long, pointed distal teeth and apex rounded.	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
03EA8F4FFFF1F91BFF0FFE85FE25FC6E.taxon	distribution	Distribution: All continents except South America and Antarctica. (Von Hayek, 1973).	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
03EA8F4FFFF3F919FF0FFE7EFC45FB33.taxon	description	Description: Male (Fig. 7; Pl. I, Figs. III, IV). Body length: 5.8, width: 1.9. Body (Fig. 7; Pl. I, Fig. III) dark brown, integument shining, setae white, these at regions forming a mosaic pattern, unevenly distributed. Head (Fig. 7) red brown, punctation simple, coarse, frons broader than long, concave. Frontal carina faintly impressed. Eye pale brown, round, moderately large with distinct facets. Labrum red brown, raised, broader than long, anterior margin convex. Mandible medially red brown, distally black, moderate, dentate, apically narrow, acute. Antennae pale brown, extending just beyond the middle of the pronotum. First segment robust, Third smallest, Fourth onward distinctly serrate and subequal in length to each other. Entirely longitudinally carinate. Pronotum (Fig. 7) dark brown, broader than long, disc strongly convex. Medially with a longitudinal shallow groove, flanked by a pair of tubercles near the posterior slope. Punctation (Fig. 8) simple. Setae white, denser toward the lateral margin. Submarginal line present. Anterior margin concave with a round anterior angle. Lateral margin entire, convex, sinuate before hind angle. Hind angle truncate; carina present, short. Scutellum (Fig. 7) dark brown, pentagonal, flat, setae dense, basally emarginated. Elytra (Fig. 7) 2.12 × longer than pronotum, red-brown. Striae punctate. Interstriae flat, punctures (Fig. 9) simple, with white setae. Basally emarginated. Apically round. Prosternum red brown, longer than broad, convex, with distinct chin piece. Punctation simple, coarse, with white pubescence. Prosternopleural suture slightly concave, broadly open up to the middle of the prosternum. Hypomeron with shallow depression for accommodating protarsi. Prosternal spine long, emarginated, medially with a longitudinal narrow keel, apically round. Mesepisternum and mesepimeron do not form part of the mesocoxal cavity. Metasternum with a shallow groove for receiving mesotarsi. Metacoxal plate incurved at middle. Abdomen red-brown. Punctuation doubled. Pubescence evenly distributed. Leg (Fig. 10) brown, claw (Fig. 11) simple, setae present at base of claw. Genitalia (Fig. 12; Pl. I, Fig. IV): Basal piece wide, arms blunt at apex, anterior margin broadly ‘ V’ shape, posterior margin weakly concave, lateral margin feebly arcuate, outer margin moderately sclerotized, thin and line like, rest feebly sclerotized; median lobe longer than parameres, arms strongly sclerotized compare to rest, moderate, arcuate, pointed at base, not exceeding ventral posterior margin of parameres, median lobe gradually narrowing from base to apex, apex rounded, furcae not reaching the anterior margin of parameres; inner lateral margin of parameres throughout and base of outer lateral margin strongly sclerotised, outer lateral margin sinuate, gradually broadening from basal to proximal, proximal to distal parallel sided, distal teeth small, pointed, apex rounded, with more than three hairs.	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
03EA8F4FFFF3F919FF0FFE7EFC45FB33.taxon	discussion	Remarks: None of the Indian congeners examined shows any resemblance to the present species.	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
03EA8F4FFFF3F919FF0FFE7EFC45FB33.taxon	etymology	Etymology: The species name is derived from the collection location.	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
03EA8F4FFFF3F919FF0FFE7EFC45FB33.taxon	materials_examined	Material examined: Holotype: Male: India, West Bengal, Jalpaiguri, BTR, Nimati, Coll. D. Raychaudhuri, 24 - vi- 2003.	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
03EA8F4FFFF3F91CFCF2FAF2FD92F898.taxon	description	(Figs. 13 - 18; Pl. I, Figs. V, VI)	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
03EA8F4FFFF3F91CFCF2FAF2FD92F898.taxon	description	Description: Male (Fig. 13; Pl. I, Figs. V, VI). Body length: 9.13, width: 2.43. Body (Fig. 13; Pl. I, Fig. V) red brown, surface shining. Setae pale brown, curved, few erect, evenly distributed. Head (Fig. 13) red brown, marginally blackish. Setae pale brown. Punctation simple. Frons broader than long. Frontal carina faintly impressed. Vertex weakly concave. Eye dark brown, round, with distinct facets. Labrum brown, black, flat, broader than long, anterior margin convex. Mandible medially red brown, margin black, dentate, apically round, blunt. Antennae brown, extending beyond the middle of the pronotum. First segment robust. Third smallest. Fourth onward serrate and subequal. The last segment constricted near the apex. Pronotum (Fig. 13) red brown, with anterior margin blackish, as long as wide, disc convex, medially with a longitudinal shallow groove near slope, punctation (Fig. 14) simple. Submarginal line present. Anterior margin straight, with round anterior angle. Lateral margin crenulate, more or less straight, sinuate before hind angle. Hind angle truncated. Scutellum (Fig. 13) brown, pentagonal, flat, setae dense, basally emarginate, convex. Elytra (Fig. 13) 2.33 x longer than pronotum, red brown, striae punctate, interstriae flat, punctation (Fig. 15) simple, basally emarginate, apically round, weakly spinose along suture. Prosternum red brown, longer than wide, convex, with distinct brown chin piece, punctation simple. Pronotosternal suture concave, broadly open up to the middle of the prosternum. Hypomeron strongly grooved for accommodating protarsi. Prosternal spine long, emarginated, apically round. Mesepisternum and mesepimeron do not form part of the margin of the mesocoxal cavity. Metasternum with groove for receiving mesotarsi. Metacoxal plate round at middle. Abdomen red brown, punctation simple, setae evenly distributed. Leg (Fig. 16) brown, tarsal claw (Fig. 17) simple, setae present at base. Genitalia (Fig. 18; Pl. I, Fig. VI): Basal piece wide, arms rounded at apex, anterior margin broadly ‘ U’ shape, posterior margin curved, corners of posterior and lateral margin projected, lateral margin straight, uniformly sclerotized; median lobe longer than parameres, uniformly sclerotised, arms of median lobe long, curved, pointed at base, not exceeding ventral posterior marginof parameres, median lobe gradually narrowing from base to apex, apex rounded, furcae not reaching the anterior margin of parameres; inner lateral margin of parameres strongly sclerotised, outer lateral margin sinuate, beyond broadend, distally deeply concave, forming posteriorly directed distal teeth, apex rounded, more than seven hairs on apex.	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
03EA8F4FFFF6F912FCECFC91FF1DF881.taxon	description	(Figs. 19 - 24; Pl. II, Figs. vii and viii)	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
03EA8F4FFFF6F912FCECFC91FF1DF881.taxon	description	Description: Male (Fig. 19; Pl. II, Figs. VII and VIII). Body length: 8.77, width: 2.85. Body (Fig. 19; Pl. II, Fig. VII) brown, marbled with yellow setae, rest of the area with golden setae. Head (Fig. 19) brown, punctation simple. Frons broader than long, concave. Frontal carina faintly impressed. Vertex weakly concave. Eye pale brown, round, with distinct facets. Labrum brown, raised, broader than long, anterior margin convex. Mandible medially red brown, distally black. Antennae pale brown, extending to the middle of the pronotum. First segment robust, third smallest and fourth onward serrate and subequal in length. Pronotum (Fig. 19) brown, wider than long, disc convex, medially with a longitudinal shallow groove extending almost up to the anterior margin. Either side of which with a gibbosity on the posterior slope. Punctation (Fig. 20) simple. Submarginal line present. Anterior margin red brown, straight with round anterior angle. Lateral margin crenulate, sinuate before hind angle. Hind angle truncated. Scutellum (Fig. 19) brown, pentagonal, flat, basally emarginated, convex, setae dense. Elytra (Fig. 19) 2.01 x longer than pronotum, brown, striae punctate, interstriae flat, punctation (Fig. 21) simple, basally emarginate, apically round and weakly spinose along suture. Prosternum brown, longer than wide, convex, with distinct chin piece, punctation simple. Pronotosternal suture slightly concave, broadly open to middle of prosternum. Hypomeron with depression for accommodating anterior tarsi. Prosternal spine long, emarginated, apically round. Mesepisternum and mesepimeron do not form part of the margin of the mesocoxal cavity. Metasternum without any groove for receiving mesotarsi. Metacoxal plate slightly round at middle. Abdomen brown, punctation double. Setae evenly distributed. Legs (Fig. 22) brown, claw (Fig. 23) simple, setae present at base. Genitalia (Fig. 24; Pl. II, Fig. VIII): Basal piece wide, arms blunt at apex, anterior margin broadly ‘ V’ shape, posterior margin weakly concave, lateral margin feebly arcuate, outer margin moderately sclerotised, thin and line like, rest feebly sclerotized; median lobe longer than parameres, arms strongly sclerotised compare to rest, long, arcuate, pointed at base, not exceeding ventral posterior margin of parameres, median lobe gradually narrowing from base to apex, apex rounded, furcae not reaching the anterior margin of parameres; base of outer lateral margin of paramere and inner lateral margin throughout strongly sclerotised, outer lateral margin sinuate, gradually broadening from basal to proximal, proximal to distal nearly straight, apex strongly narrow, rounded, distal teeth small, pointed and more than nine hairs on apex.	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
03EA8F4FFFF6F912FCECFC91FF1DF881.taxon	discussion	Remarks: No Indian congeners examined show any resemblance to the present species.	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
03EA8F4FFFF6F912FCECFC91FF1DF881.taxon	etymology	Etymology: The species name is derived from the body pattern.	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
03EA8F4FFFF6F912FCECFC91FF1DF881.taxon	materials_examined	Material examined: Holotype: Male: India, West Bengal, Jalpaiguri, BTR, Nimati, Coll. D. Raychaudhuri, 05 - vii- 2003.	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
03EA8F4FFFF8F911FF2AF8A0FD14FA66.taxon	description	(Figs. 25 - 30; Pl. II, Figs. ix and x) Description: Male (Figs. 25; Pl. II, Figs. IX and X). Body length: 7.88, width: 2.51. Body (Fig. 25; Pl. V, Fig. IX) black, surface shining. Setae pale brown, dense, curved. Head (Fig. 25) black, Punctation simple. Frons broader than long, slightly concave. Frontal carina faintly impressed. Eye pale brown, round. Labrum dark brown, raised, broader than long, anterior margin convex. Mandible dentate, apically black and round. Antennae brown, extending to the middle of the pronotum. First segment robust, longest. Second and third subequal. Fourth onward serrate and subequal in length. The last segment constricted near the apex. Pronotum (Fig. 25) black, disc convex, wider than long, punctation (Fig. 26) simple, Submarginal line present. Anterior margin straight, with round anterior angle. Lateral margin crenulate, convex, sinuate before hind angle. Carina reaches ¾ th of the pronotum. Hind angle truncated. Scutellum (Fig. 79) black, pentagonal, basally emarginated, concave. Elytra (Fig. 25) 2.09 x longer than pronotum, black, striae punctate. Interstriae slightly convex, punctation (Fig. 27) simple, basally emarginate, round. Apically round, and weak dentate. Prosternum black, strongly convex, with a distinct chin piece, punctation simple. Pronotosternal suture straight, broadly open to middle of prosternum. Hypomeron without any depression for accommodating anterior tarsi. Prosternal spine long, emarginated, apically round. Mesepisternum and mesepimeron do not form part of the margin of the mesocoxal cavity. Metasternum without any groove for receiving mesotarsi. Abdomen black, punctation simple, setae evenly distributed. Leg (Fig. 28) brown, tarsal claw (Fig. 29) simple, setae present at base. Genitalia (Fig. 30; Pl. II, Fig. X): Basal piece wide, arms rounded at apex, both anterior and posterior margin concave, lateral margin weakly arcuate, uniformly sclerotised throughout; median lobe longer than parameres, uniformly sclerotised throughout, Plate I, II. Agrypnus buxaensis sp. nov.; III, IV Agrypnus nimatiensis sp. nov.; V, VI. Agrypnus indicus sp. nov. Five new click beetles of AgrypNUS Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India Plate VII, VIII. Agrypnus variegatus sp. nov.; IX, X. Agrypnus crenulatus sp. nov. apex more so, arms of median lobe long, feebly arcuate, pointed at base, not exceeding ventral posterior margin of parameres, median lobe medially concave, proximal to distal gradually broadened, apex sharply narrow, acute, furcae not reaching the anterior margin of parameres; parameres uniformly sclerotised throughout, outer lateral margin broadened basally, medially strongly concave, proximal to distal straight, distal teeth small, pointed, apex rounded, more than five hairs on parameres.	en	Sarkar, Sutirtha, Saha, Sumana, Raychaudhuri, Dinendra (2025): Five new click beetles of Agrypnus Eschscholtz, 1829 (Elateridae: Agrypninae) from Buxa Tiger Reserve, West Bengal, India. Records of the Zoological Survey of India 125 (2): 97-109, DOI: 10.26515/rzsi/v125/i2/2025/172475, URL: https://doi.org/10.26515/rzsi/v125/i2/2025/172475
