identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03EA87C1FFF42D1A83D0FDB1FB9FFD09.text	03EA87C1FFF42D1A83D0FDB1FB9FFD09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinellus bicolor Kiesenwetter 1874	<div><p>Malthinellus bicolor Kiesenwetter, 1874:</p><p>holotype (by monotypy), male (NHMUK), “Type / H. T.”, “ Japan / G. Lewis / 1910-320”, “ Malthinellus / bicolor / Ksw.”, “SYN- / TYPE”; non-type (all RNC): 1 male, Japan, Kumamoto Pref., Yabe-machi, Komagaeri-pass, 30. VII. 1997, S. Imasaka leg.; 1 female, Japan, Fukuoka Pref., Kitakyûshû-shi, Tashiro, 30. VII. 1996, S. Imasaka leg.; 3 males, Japan, Chiba Pref., Narita-shi, Ôtake, Sakatagaikesôgô-kôen (alt. ca. 30 m), 15. VI. 2016, Ryo Nakamura leg. (Figs. 7A, G, 8A, E–F).</p></div>	https://treatment.plazi.org/id/03EA87C1FFF42D1A83D0FDB1FB9FFD09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFF42D1A83D0FD0DFB1AFCC1.text	03EA87C1FFF42D1A83D0FD0DFB1AFCC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinellus iheyanus Takahashi & Nakamura 2022	<div><p>Malthinellus iheyanus Takahashi &amp; Nakamura, 2022:</p><p>paratype, 1 female (RNC), “[JAPAN] Iheya-jima Is. / Mt. Koshidake, alt. 100–160m / 21. III. 2022 / Ryo Nakamura leg. / 伊ṃDzẽ DZē ” (Fig. 7H).</p></div>	https://treatment.plazi.org/id/03EA87C1FFF42D1A83D0FD0DFB1AFCC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFF42D1A83D0FC91FE08FC31.text	03EA87C1FFF42D1A83D0FC91FE08FC31.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinus filiformis Wittmer 1997	<div><p>Malthinus filiformis Wittmer, 1997:</p><p>holotype, male (MNMB), “ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.49&amp;materialsCitation.latitude=25.22" title="Search Plazi for locations around (long 98.49/lat 25.22)">YUNNAN</a> 1500–2500m / 25.22N 98.49E 17–24.5 / GAOLIGONG mts. / Vít Kubáň leg. 1995”; paratypes, 2 females (NHMB), same collecting data as the holotype (Figs. 6A, 7L, 8L).</p></div>	https://treatment.plazi.org/id/03EA87C1FFF42D1A83D0FC91FE08FC31	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFF42D1A83D0FC25FD27FBE9.text	03EA87C1FFF42D1A83D0FC25FD27FBE9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinus flaveolus (Herbst 1786)	<div><p>Malthinus flaveolus (Herbst, 1786):</p><p>non-type, 1 male, 1 female (RNC), Czech Republic, Bohemia centr, Jabkenice, 25. VI. 1992, Ing. Brokes leg. (Figs. 7C, J).</p></div>	https://treatment.plazi.org/id/03EA87C1FFF42D1A83D0FC25FD27FBE9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFF42D1A83D0FBEDFA4EFBA1.text	03EA87C1FFF42D1A83D0FBEDFA4EFBA1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinus humeralis Kiesenwetter 1874	<div><p>Malthinus humeralis Kiesenwetter, 1874:</p><p>non-type, 1 male, 1 female (RNC), Japan, Nara Pref., Tenkawamura, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=135.876&amp;materialsCitation.latitude=34.241" title="Search Plazi for locations around (long 135.876/lat 34.241)">Mt. Kannonmine-yama</a> (alt. ca. 940 m), 34.241°N, 135.876°E, 17. VII. 2021, Ryudai Ito leg. (Figs. 7E, M) .</p></div>	https://treatment.plazi.org/id/03EA87C1FFF42D1A83D0FBEDFA4EFBA1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFF42D1A83D0FB95FD86FB59.text	03EA87C1FFF42D1A83D0FB95FD86FB59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinus palingenesis Wittmer 1993	<div><p>Malthinus palingenesis Wittmer, 1993:</p><p>holotype, male (NHMB), “ Nr. Paling / Taoyuan Pref. / TAIWAN ”, “ 6- IV-1982 / T. Shimomura leg.” (Fig. 3D).</p></div>	https://treatment.plazi.org/id/03EA87C1FFF42D1A83D0FB95FD86FB59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFF42D1A83D0FB5DFDB0FAC9.text	03EA87C1FFF42D1A83D0FB5DFDB0FAC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinus sikkimensis Pic 1924	<div><p>Malthinus sikkimensis Pic, 1924:</p><p>non-type, 1 male (NHMUK), “ NEPAL: / Ghanpokhara / 5,500–7,000 ft. / 2.v.1954. / J. Quinlan.”; ditto, 1 male (NHMB), “ Nepal / Kathmandu V. ”, “ Godawari 1500m / 17.v.1983 ” (Fig. 3E); ditto, 1 female (NHMB), “ Neentale / 2160 m 30. V.”, “O. Nepal 1979 / Bhakta B. Ch. ”(Figs. 3F, 7D, K). All specimens were identified by W. Wittmer.</p></div>	https://treatment.plazi.org/id/03EA87C1FFF42D1A83D0FB5DFDB0FAC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFF42D1A83D0FACDFBCFFA65.text	03EA87C1FFF42D1A83D0FACDFBCFFA65.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinus tonkineus Pic 1923	<div><p>Malthinus tonkineus Pic, 1923:</p><p>lectotype (designated here; bearing a lectotype label by W. Wittmer, although not formally designated previously), male (MNHN), “ Hoa Binh ”, “ Malthinus / tonkineus / n sp”, “LECTOTYPE”, “REM / 94 / 1.4”, “ Malthinus / tonkineus / Pic / det. W. Wittmer ”(Fig. 4D, 8K).</p></div>	https://treatment.plazi.org/id/03EA87C1FFF42D1A83D0FACDFBCFFA65	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFF52D1C8380FB39FBEFFEB9.text	03EA87C1FFF52D1C8380FB39FBEFFEB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinellus Kiesenwetter 1874	<div><p>Genus Malthinellus Kiesenwetter, 1874</p><p>Malthinellus Kiesenwetter, 1874: 280 .</p><p>Type species: Malthinellus bicolor Kiesenwetter, 1874, by monotypy.</p><p>Diagnosis. Within the tribe Malthinini, this genus is recognized by the combination of the following characteristics: body small (BL 2.5–6.3 mm, usually less than 5.0 mm); a combination of blackish brown and yellowish to reddish orange body, without a yellowish-white marking at the apex of each elytron as present in many Malthinus species; head moderately rounded, gradually narrowed posteriorly, not parallel-sided at neck, with the dorsum densely punctate; antennae filiform; gular sutures relatively indistinct; gula somewhat wide and not deeply concave, reaching the posterior margin of head; pronotum densely punctate, with both anterior and posterior margins bordered, lateral margins arcuate in dorsal view, and epipleura moderately developed without any fissures; legs not modified as in some Malthinus species; pro- and mesocoxae elongate; metacoxae not elongate, length and basal width nearly equal; tibiae each with one spur at apex; elytra longitudinally punctate and sometimes with some longitudinal costae, slightly abbreviated, usually with apical one or two abdominal segments and apices of hind wings a little protruding beyond elytral apices; sternite VIII widely emarginate in a V-shape on posterior margin; male sternite IX weakly sclerotized, oval, lamellate, gently convex ventrally; aedeagus rounded or subquadrate shape in dorsal and ventral outline; basal piece wide, widely covering the ventral area of aedeagus, lacking basal shaft; inner basal piece with anterior terminal projecting dorsally in various shape; centrophyse bearing one or two pairs of dorsally directed lateral lobes, with penis and complex processes situated medially; coxites divided, apical portion flattened and oval to triangular; proctiger indistinct; paraprocts separated laterally; valvifers indistinct.</p><p>The moderately wide and neither grooved nor concave gula and the presence of dorsally projected lateral lobes on the centrophyse of the aedeagus would be autapomorphic characteristics of Malthinellus, which distinguish it from the morphologically similar genera Malthinus and Phaneromalthinus gen. nov. Although the monophyly of two known Mimomalthinus species remain unknown (see Discussion), Malthinellus can be distinguished from the type species Mi. niger Pic, 1931 by the gula structure. From the other species Mi. shokhrini Kazantsev, 2006, Malthinellus differs in the shapes of head and male sternite IX. See also Table 1.</p><p>Distribution. Japan, Taiwan, China (Guandong, Hubei), Vietnam (Hòa Bình).</p></div>	https://treatment.plazi.org/id/03EA87C1FFF52D1C8380FB39FBEFFEB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFF22D1E8380FA83FDCDFD7D.text	03EA87C1FFF22D1E8380FA83FDCDFD7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinellus chinensis Wittmer 1956	<div><p>Malthinellus chinensis Wittmer, 1956</p><p>(Figs. 1A–J, 7I, 8B, 10)</p><p>Malthinellus chinensis Wittmer, 1956: 307 (original description); Kazantsev &amp; Brancucci, 2007: 276 (catalogue); N. Takahashi, 2018: 395 (checklist).</p><p>Type material examined. Allotype, male (NHMB), “ Suisapa, 1000 M. / Lichuan Distr. / W. Hupoh, China / VII-25-48”, “♂”, “Allotypus”, “ Malthinellus / chinensis / Wittm / det. W.Wittmer ”, “Naturhistorisches / Museum Basel / Coll. W. Wittmer ”, “ CANTHARIDAE [QR code] / CANTH00016078 ”.</p><p>Supplementary description. Male. Sternite VIII (Fig. 8B) with posterior margin broadly emarginate in a Vshape to more than half of its length. Sternite IX narrowed behind faint angulations on sides in posterior portion, weakly produced posteriorly and bilobate at apex.</p><p>Aedeagus (Figs. 1F–J) stout, subquadrate with rounded corners in dorsal and ventral outline; basal piece broadly rounded triangular in ventral outline, almost entirely covering the inner basal piece, with apical margin slightly emarginate, ventral surface with a medio-longitudinal furrow, gently convex medio-ventrally and weakly curved dorsally in profile; inner basal piece with broad anterior terminal strongly curved dorsally, apex divided into two lateral subrectangular parts in contact with each other; outer lateral lobe broad, rounded in profile, apically twisted, sharply tapering anteromedially in dorsal view; inner lateral lobe absent; anterior lobe existing at basal 1/4 of aedeagus, simply projecting dorsally in cultriform in profile; penis robust, divided into two lateral parts, each projecting dorsally in a bluntly rectangular shape in profile; central part of centrophyse with a pair of small laterally directed spine-like processes medially, and a pair of anteriorly directed, twisted process situated at basal 1/4, digitiform in dorsal view and triangular in profile.</p><p>Differential diagnosis. Malthinellus chinensis Wittmer is one of the smallest species in the genus, and only this species and small individuals of Malthine. exiguus N. Takahashi have a body length less than 3.0 mm. This species is unique and differs from all other congeners in possessing a pair of anterior lobes on the aedeagus. The absence of the inner lateral lobe is reminiscent of the two Japanese species from the Okinawa Islands, Malthine. masatakai Takahashi &amp; N. Takahashi and Malthine. iheyanus Takahashi &amp; Nakamura, but the shape of the outer lateral lobes and the body coloration differ considerably. As stated in the original description, this species resembles Malthine. bicolor Kiesenwetter in general appearance but can be distinguished by the predominantly yellowishbrown coloration of head (in Malthine. bicolor, only clypeus is yellowish brown). The body coloration is more similar to that of Malthine. tsuchidai Takahashi, Malthine. multispinosus (Wittmer) comb. nov. and Malthine. hoabinensis (Wittmer) comb. nov., but the differences in the morphology of the aedeagus, particularly the lateral lobes, are conspicuous.</p><p>Remarks. This species was originally described based on the holotype and allotype collected at the same locality on the same day (Wittmer, 1956). Although Wittmer’s original description is almost entirely based on the male, it is apparent that the allotype is actually a male. Therefore, the holotype, which is reportedly housed at the California Academy of Sciences (USA), is presumed to be a female, though it was not examined in this study. Another possibility is that the “ allotype ” examined here is actually the holotype, as in the case of Peltariosilis scutulata (Wittmer, 1952) whose actual holotype was labelled as “ allotype ” (cf. Biffi and Geiser, 2020). According to the original description, the collectors were “Gressitt und Djou”, but this information was not present on the labels.</p><p>The allotype was in rather bad condition. Before dissection, it was mounted at the tip of a triangular card, with both mesothoracic legs and the left metathoracic leg almost detached, barely attached between the card and the body. Other legs and most of the antennae were missing, and a few detached antennal segments were found stuck between the card and the body (Fig. 1A–B). These legs, antennomeres and the head detached when the specimen was softened in water, but all parts, including dissected aedeagus, were subsequently remounted together with the remaining body on the same white square card.</p><p>The aedeagus of this species is illustrated and described for the first time. In the allotype, the right one of the pair of small, laterally directed spine-like processes on the central part of the centrophyse is broken (Figs. 1H, I).</p><p>Distribution. China (Hubei).</p></div>	https://treatment.plazi.org/id/03EA87C1FFF22D1E8380FA83FDCDFD7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFF02D108380F929FCD0F89E.text	03EA87C1FFF02D108380F929FCD0F89E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinellus crenulatomimus (Wittmer 1984)	<div><p>Malthinellus crenulatomimus (Wittmer, 1984)</p><p>(Figs. 3A, 5G–H, 8D, I, 10)</p><p>Malthinus crenulatomimus Wittmer, 1984: 154 (original description), figs. 93–94 (aedeagus).</p><p>Malthinus (Malthinus) crenulatomimus: Kazantsev &amp; Brancucci, 2007: 278 (catalogue).</p><p>Malthinellus crenulatomimus: K. Takahashi &amp; N. Takahashi, 2007: 147 (new combination, additional record); Satô et al., 2014: 59 (checklist); N. Takahashi, 2018: 395 (checklist).</p><p>Type material examined. Holotype, male (NHMB), “ Mt. Nanfengshan / 1000m Nr. Liukuei / Kaohsiung Pref. ”, “ TAIWAN / 21-III-1981 / T. Shimomura leg.”, “154”, “HOLOTYPUS”, “ Malthinus / crenulato- / mimus / Wittm. / det. W.Wittmer ”, “Naturhistorisches / Museum Basel / Coll. W. Wittmer”, “ CANTHARIDAE [QR code] / CANTH00020137 ” . Paratypes, 2 males (NHMB), same collecting data as the holotype .</p><p>Additional material examined. 1 male (NMNS ENT5237-5735), “ Taiwan Nantou / Chunyang / III/12– IV/9/2002 / C. S. Lin &amp; W. T. Yang / Malaise trap (KCN)” ; 1 male (NMNS ENT5237-1840), same collecting data as the preceding but “ IV/9– V /7/2002 ”; 1 male (YHC), “ 4.IV.2013 / Malunshan / Yun Hsiao leg.”; 1 male (YHC), “ 25.IV.2015 / Xitou, Nantou, Taiwan / J.-F. Hsieh leg.”; 1 male (RNC), Taiwan, Chiayi County, Alishan Township, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.717&amp;materialsCitation.latitude=23.477" title="Search Plazi for locations around (long 120.717/lat 23.477)">Leye Village</a>, 23.477N, 120.717E, alt. 1320– 45 m, 23. IV.2025, Ryo Nakamura leg.</p><p>Supplementary description. Male. Sternite VIII (Fig. 8D) with apical margin deeply emarginate in a V-shape, the depth of the incision reaching approximately half the length of the sternite, bottom of the incision rounded. Sternite IX (Fig. 8I) elongate, oval, with apex narrowing gently and slightly bilobate.</p><p>Aedeagus (Figs. 5G–H) 0.65–0.72 mm in length (n = 4); stout, nearly circular in dorsal and ventral outline; basal piece rounded in ventral outline, almost entirely covering the inner basal piece, with apical margin protruding in quadrate-shape at the central 1/3, ventral surface with a medio-longitudinal furrow, gently convex medio-ventrally and weakly curved dorsally in profile; inner basal piece with anterior terminal divided into two subrectangular parts, each gently projecting dorsally, with apices slightly recurved ventrad; outer lateral lobe flattened, spoon-shaped, with a small, claw-like, sharp projection at the posterodistal end; inner lateral lobe flattened, elongate spoon-shaped, twisted dorso-posteriorly, with the apex twisted and recurved ventrally; penis robust, strongly projecting dorsally as if fused from both sides; central part of centrophyse without any distinct processes.</p><p>Differential diagnosis. This species shares the same coloration with three species from the Ryukyu Islands, Japan, Malthinellus exiguus N. Takahashi, Malthine. masatakai Takahashi &amp; N. Takahashi, and Malthine. iheyanus Takahashi &amp; Nakamura, but it clearly differs from all of them in the structure of the aedeagus. This species also resembles the other two Taiwanese species in the aedeagus. For distinguishing characters from other Taiwanese species, see the corresponding section under Malthine. mushanus sp. nov.</p><p>Remarks. This species was originally described from southern Taiwan, and it is also recorded from central Taiwan in the present study. At Chunyang, both Malthinellus crenulatus and Malthine. mushanus sp. nov. were collected using the same malaise trap, and the latter was obtained during the same period.</p><p>Distribution. Taiwan (Chiayi, Kaohsiung, Nantou).</p></div>	https://treatment.plazi.org/id/03EA87C1FFF02D108380F929FCD0F89E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFFF2D148380FAA4FD58FCA5.text	03EA87C1FFFF2D148380FAA4FD58FCA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinellus crenulatus (Wittmer 1979)	<div><p>Malthinellus crenulatus (Wittmer, 1979)</p><p>(Figs. 3B–C, 5E–F, 8J, 10)</p><p>Malthinus crenulatus Wittmer, 1979: 335 (original description), figs. 6–7 (aedeagus); Wittmer, 1984: 155 (part; additional records).</p><p>Malthinus (Malthinus) crenulatus: Kazantsev &amp; Brancucci, 2007: 278 (catalogue).</p><p>Malthinellus crenulatus: K. Takahashi &amp; N. Takahashi, 2007: 147 (new combination, additional record); Satô et al., 2014: 59 (checklist); N. Takahashi, 2018: 395 (checklist).</p><p>Type material examined. Holotype, male (NHMB), “ Taiwan 1977.23.4 / J. &amp; S. Klapperich ”, “ Fenchihu / 1400m ”, “441”, “HOLOTYPUS”, “ Malthinus / crenulatus / Wittm. / det. W.Wittmer ”, “Naturhistorisches / Museum Basel / Coll. W. Wittmer”, “ CANTHARIDAE [QR code] / CANTH00017356 ” . Paratypes (all NHMB): 1 male, same collecting data as the holotype; 1 male, same collecting data as the holotype but without date .</p><p>Additional material examined. 1 male (NMNS ENT5249-3074), “ Taiwan Nantou / Chunyang / V/11– VII/13/2004 / C. S. Lin &amp; W. T. Yang / Malaise trap (KCN)” .</p><p>Supplementary description. Male. Infuscate maculae on posterior parts of head and center of pronotum somewhat variable in size (Figs. 3B–C).</p><p>Sternite VIII with apical margin deeply emarginate in a V-shape, the depth of the incision reaching approximately half the length of the sternite, bottom of the incision rounded. Sternite IX (Fig. 8J) elongate, oval, with apex narrowing gently and slightly bilobate.</p><p>Aedeagus (Figs. 5E–F) 0.73–0.89 mm in length (n = 2); stout, subquadrate with rounded corners in dorsal and ventral outline; basal piece broadly rounded triangular in ventral outline, almost entirely covering the inner basal piece, with apical margin slightly emarginate, ventral surface with a medio-longitudinal furrow, gently convex medio-ventrally and weakly curved dorsally in profile; inner basal piece with broad anterior terminal, the left and right parts being extremely close to each other, weakly elevated dorsally as a whole, appearing wedge-shaped in profile and slightly triangularly raised in caudal view; outer lateral lobe thin, whip-like, projecting dorsally while slightly curved medially, with the apex sharply pointed and directed anteriorly; inner lateral lobe flattened, elongate spoon-shaped, slightly twisted, with the apex sharply pointed triangularly in profile, directed posteriorly; penis rod-like, divided into two lateral parts, each projecting dorsally, with apices bluntly triangular in profile and directed posteriorly; central part of centrophyse with a pair of rod-like processes medially, slightly expanded towards the apex.</p><p>Differential diagnosis. This species resembles the other two Taiwanese species. For distinguishing characters, see the corresponding section under Malthinellus mushanus sp. nov.</p><p>Remarks. This species is currently known only from central Taiwan. At Chunyang, both Malthinellus crenulatomimus (Wittmer) and Malthine. mushanus sp. nov. were collected using the same malaise trap.</p><p>After original description, Wittmer (1984) recorded at least six specimens of this species (the exact number was not specified), but two of them located in NHMB were misidentified and actually belong to Malthine. mushanus sp. nov., to which they are designated as paratypes below. Two and three specimens were also found in EUMJ and OMNH respectively, but all were females or broken and could not be reliably identified. Other specimens recorded by Wittmer (1984) could not be examined and confirmed its identification.</p><p>As stated in the original description, this species is somewhat similar to Malthinus sikkimensis Pic, 1924 (Figs. 3E–F), a Himalayan species of a different genus. However, Malthinu. sikkimensis can be distinguished from Malthine. crenulatus by the different antennal length and stick-like and curved sclerotized sternite IX of male. Wittmer (1993) also noted Malthinu. palingenesis Wittmer, 1993 (Fig. 3D) from Taiwan externally resembles this species and it is indeed difficult to distinguish these two species based solely on dorsal appearance.</p><p>Distribution. Taiwan (Chiayi, Nantou).</p></div>	https://treatment.plazi.org/id/03EA87C1FFFF2D148380FAA4FD58FCA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFFA2D158380FC54FDF0FA81.text	03EA87C1FFFA2D158380FC54FDF0FA81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinellus mushanus Nakamura 2026	<div><p>Malthinellus mushanus sp. nov.</p><p>(Figs. 2A–I, 7B, I, 8C, H, 10)</p><p>Malthinus crenulatus: Wittmer, 1984: 155 (part; misidentification).</p><p>Type material. Holotype, male (NMNS), “ Taiwan Nantou / Chunyang / IV/6–V/1/2004 / C. S. Lin &amp; W. T. Yang / Malaise trap (KCN)”, “ NMNS ENT / 5249-2303 ” . Paratypes (8 males): 3 males (NMNS ENT5237-1817, 1839, 3223), same collecting data as the holotype but “ IV/9–V/7/2002 ”; 1 male (TARI), “ Taiwan: Nantou / Chientai trail (尖台ūã) / 12.III.2019. leg. W.-C. Liao ” ; 2 males (EUMJ), “ 台=南üñ仁Ēae力n'āDzãw / 24.055°N, 121.155°E / [TAIWAN] Nantou Hsien, / Ren’ai Township, / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=121.155&amp;materialsCitation.latitude=24.055" title="Search Plazi for locations around (long 121.155/lat 24.055)">Lixing Industry Road</a>, / 7 May 2016, Yu HISASUE leg.” ; 1 male (NHMB), “[TAIWAN] / Meifeng / Nantou Hsien ”, “ 26. VI. 1971 / Y. Miyatake ” ; 1 male (NHMB), “ Nanzan-kei / Formosa / 29. III. 1967 / S. Fukuda ” .</p><p>Etymology. Named after the locality name “Musha (mữ)”. This is a traditional name by indigenous people and corresponds to Ren'ai Township, Nantou County, Taiwan. This new species has been collected only from this area.</p><p>Description. Male. BL 3.88–4.13 (4.13 in the holotype = HT) mm. Body dark blackish-brown; head, pronotum, scutellum, trochanters, and femora yellowish orange to orange (but the posterior part of head, median area of pronotum, and scutellum infuscate in one paratype from Meifeng); palpomeres and sternite IX dark brown; mandibles yellowish orange in basal halves, dark orange in apical halves; antennomere I yellowish orange, slightly darkened apically and antennomeres II–XI dark brown; tibiae and tarsi yellowish orange to brown, usually darkened towards the apices. Body densely covered with yellowish-white short pubescence.</p><p>Head rounded, HW 0.81–0.87 (0.86 in HT) mm, HW/PW 1.08–1.19 (1.19 in HT); eyes globular and moderately protruding, interocular distance 3.00–4.23 (3.38 in HT) times as wide as the diameter of an eye; antennae filiform, reaching basal 3/5 of elytra, comparative lengths of antennomeres as follows: 1.89: 1: 1.27: 1.46: 1.53: 1.52: 1.51: 1.39: 1.34: 1.35: 1.55.</p><p>Pronotum trapezoidal, sides slightly widened posteriorly, anterior angles rounded, lateral margins slightly sinuate near slightly produced or angulate hind angles, disc coarsely punctate, moderately convex but flattened in the median area; PL 0.54–0.61 (0.57 in HT) mm, PW 0.71–0.80 (0.73 in HT) mm, PW/PL 1.27–1.31 (1.28 in HT). Scutellum trapezoidal with rounded apex. Elytra moderately long, sides parallel, slightly narrowed and rounded at apex; EL 2.67–2.95 (2.88 in HT) mm, EW 1.03–1.08 (1.05 in HT) mm, EW/PW 1.34–1.51 (1.45 in HT), EL/EW 2.60–2.74 (2.73 in HT). Legs slender; femora mostly straight; tibiae feebly arcuate.</p><p>Sternite VIII (Fig. 8C) with apical margin deeply and broadly emarginate in a V-shape, the depth of the incision reaching approximately 3/4 of the length of the sternite, bottom of the incision smoothly rounded. Sternite IX (Fig. 8H) broadly oval, abruptly narrowed and curved dorsally at apical 1/6, apex slightly angulate or bilobate.</p><p>Aedeagus (Figs. 2E–I), 0.93–1.02 (1.02 in HT) mm in length; stout, subquadrate with rounded corners in dorsal and ventral outline; basal piece broadly rounded triangular in ventral outline, mostly covering the inner basal piece, with apical margin slightly emarginate, ventral surface with a medio-longitudinal furrow, gently convex medio-ventrally and weakly curved dorsally in profile; inner basal piece with broad anterior terminal, the left and right parts nearly fused, weakly elevated dorsally as a whole, appearing wedge-shaped in profile and projecting as quadrate plates in caudal view; outer lateral lobe divided into two parts: outer lateral lobe 1 robust, extending dorsally, then abruptly narrowed and strongly curved medially, with its apical part further elongated antero-posteriorly, apex rounded in profile and forming an elongate oval in dorsal view; outer lateral lobe 2 arising from the base of lobe 1, strongly digitiform, weakly curved posteriorly, apex acute in profile; inner lateral lobe flattened, elongate spoon-shaped, extending postero-medially with a gentle curve, apex broadly rounded; penis robust, divided into two lateral parts, each quadrate and projecting dorsally in profile; central part of centrophyse with a pair of rod-like processes medially, as long as penis, slightly undulate and slightly curved posteriorly.</p><p>Differential diagnosis. The three species occurring in Taiwan are externally very similar and share an elongate, spoon-shaped inner lateral lobe, but they are different in the following points. Dorsal coloration: posterior part of head, median area of pronotum, and scutellum are infuscate in Malthine. crenulatus, but entirely orange in Malthine. crenulatomimus, and both forms are present in Malthine. mushanus sp. nov.; sternite VIII: the V-shaped incision reaching approximately half the length of the sternite in Malthine. crenulatus and Malthine. crenulatomimus, while reaching approximately 3/4 of the length of the sternite in Malthine. mushanus sp. nov.; size of aedeagus: larger (more than 0.93 mm) in Malthine. mushanus sp. nov., smaller (less than 0.72 mm) in Malthine. crenulatomimus, and intermediate size in Malthine. crenulatus; basal piece of aedeagus: apical margin is protruding in quadrate-shape at the central 1/ 3 in Malthine. crenulatomimus, while not protruding and slightly emarginate in Malthine. crenulatus and Malthine. mushanus sp. nov.; outer lateral lobe of aedeagus: spoon-shaped, with a small sharp projection at the posterodistal end in Malthine. crenulatomimus, very thin and whip-like in Malthine. crenulatus, and divided into two parts, one of which is long and complex in shape, while the other is shorter and digitiform in Malthine. mushanus sp. nov.</p><p>Remarks. The two paratypes in NHMB were misidentified and recorded as Malthinus crenulatus by Wittmer (1984). Two specimens bearing the same label as the paratype from Meifeng were found in OMNH, but both were damaged and lacking the abdomen, making specific identification impossible.</p><p>This species is currently known only from Ren'ai Township, Nantou County, central Taiwan. At Chunyang, both Malthinellus crenulatomimus (Wittmer) and Malthine. crenulatus (Wittmer) were collected using the same malaise trap, and the former was obtained during the same period.</p><p>Distribution. Taiwan (Nantou).</p></div>	https://treatment.plazi.org/id/03EA87C1FFFA2D158380FC54FDF0FA81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFF92D178380FF60FD9DFB8A.text	03EA87C1FFF92D178380FF60FD9DFB8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinellus multispinosus (Wittmer 1995) Nakamura 2026	<div><p>Malthinellus multispinosus (Wittmer, 1995) comb. nov.</p><p>(Figs. 4A, 5A–B, 8G, 10)</p><p>Malthinus multispinosus Wittmer, 1995: 155 (original description), figs. 61–62 (aedeagus).</p><p>Malthinus (Malthinus) multispinosus: Kazantsev &amp; Brancucci, 2007: 281 (catalogue).</p><p>Type material examined. Holotype, male (NHMB), “ Kwangtung, S. China. / Loh Fau Shan, / Poh-lo District / April 6–8, 1934 / R. C. Yeung ”, “REM / 94 / 3.3”, “HOLOTYPUS”, “M. / multispinosus / Wittm. / det. W.Wittmer ”, “Naturhistorisches / Museum Basel / Coll. W. Wittmer”, “ CANTHARIDAE [QR code] / CANTH00017395 ”.</p><p>Supplementary description. Male. Pronotum trapezoidal, sides slightly widened posteriorly, lateral margines distinctly bordered, disc with a medio-longitudinal furrow not reaching the anterior or posterior margins.</p><p>Sternite IX (Fig. 8G) somewhat elongate, oval, with apex slightly bilobate.</p><p>Aedeagus (Figs. 5A–B) stout, broadly rounded in dorsal and ventral outline; basal piece rounded in ventral outline, almost entirely covering the inner basal piece, with apical margin deeply emarginate in a V-shape; inner basal piece with anterior terminal not strongly expanded, curved dorsally, apex divided into two rod-like parts; outer lateral lobe straight and rod-shaped, apically sharply pointed, bearing two sharp spines along the outer margin; inner lateral lobe simple and spine-like, slightly shorter than outer lateral lobe, and weakly curved anteriad in profile; penis robust, divided into two lateral parts, each rod-shaped, densely set with small spines apically; central part of centrophyse with a pair of long, sharply pointed processes curved posteriorly, opposing penis, and a small V-shaped projection situated medially at the posterior area.</p><p>Differential diagnosis. Malthinellus multispinosus (Wittmer) comb. nov. is similar to Malthine. chinensis Wittmer in general appearance but differs from it in somewhat larger body (BL 3.5 mm) and different shape of the aedeagus. The deeply V-shaped incision of the apical margin of the basal piece is likely unique to this species within the genus.</p><p>Remarks. Although the original description stated that the aedeagus of this species is peculiar and cannot be compared to any other Malthinus species described from China, there is no doubt that this species belongs to the genus Malthinellus based on the aedeagal structure, and a new combination is herein proposed.</p><p>Distribution. China (Guandong).</p></div>	https://treatment.plazi.org/id/03EA87C1FFF92D178380FF60FD9DFB8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFF92D088380FB33FD8EFEB9.text	03EA87C1FFF92D088380FB33FD8EFEB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthinellus hoabinensis (Wittmer 1995) Nakamura 2026	<div><p>Malthinellus hoabinensis (Wittmer, 1995) comb. nov.</p><p>(Figs. 4B–C, 5C–D, 10)</p><p>Malthinus hoabinensis Wittmer, 1995: 148 (original description), figs. 52–53 (aedeagus).</p><p>Type material examined. Holotype, male (MNHN), “ Hoo-Binh / Tonkin ”, “REM / 94 / 1.3”, “HOLOTYPUS”, “ Malthinus / hoabinensis / Wittm. / det. W.Wittmer ” . Paratype, 1 female (MNHN), “ Hoa / Binh ”, “PARATYPUS”, “ Malthinus / hoabinensis / Wittm. / det. W.Wittmer ” .</p><p>Supplementary description. Pronotum nearly rectangular, as wide at the anterior margin as at the posterior, lateral margines distinctly bordered.</p><p>Male. Aedeagus (Figs. 5C–D) stout, subquadrate with rounded corners in dorsal and ventral outline; inner basal piece with anterior terminal curved dorsally, apex deeply emarginate in U-shape with both lateral ends slightly projecting; outer lateral lobe flattened, dorsally projecting, weakly curved medially with a slight undulation, and apically sharply pointed; inner lateral lobe whip-like, extending posteriorly and then abruptly curved medially, with apex concealed behind the penis in dorsal view; penis robust, divided into two lateral parts, each projecting dorsally in a bluntly rectangular shape in profile; central part of centrophyse bearing a pair of thin plate-like processes, each with a sharp anteriorly directed spine at the base, the distal portion extending toward the penis and terminating in a triangular projection on the ventral surface.</p><p>Differential diagnosis. Malthinellus hoabinensis (Wittmer) comb. nov. is somewhat similar to M. chinensis Wittmer and Malthine. tsuchidai Takahashi in general appearance but differs from them in the shape of the aedeagus, particularly the peculiar-shaped inner lateral lobes. The slender and whip-like inner lateral lobe is somewhat reminiscent of the two Japanese species from the Ryukyu Islands, Malthine. chujoi (Wittmer) and Malthine. exiguus N. Takahashi, but the body coloration and other characteristics of the aedeagus are quite different.</p><p>Remarks. This species is also transferred to the genus Malthinellus based on the aedeagal structure.</p><p>As stated in the original description, this species externally resembles Malthinus tonkineus Pic, 1923, a species of a different genus occurring in the same locality. They are certainly similar and difficult to distinguish by dorsal appearance, though Malthinu. tonkineus has thicker antennae and larger eyes.</p><p>Distribution. Vietnam (Hòa Bình).</p></div>	https://treatment.plazi.org/id/03EA87C1FFF92D088380FB33FD8EFEB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFE62D088380FE65FA92F961.text	03EA87C1FFE62D088380FE65FA92F961.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaneromalthinus Nakamura 2026	<div><p>Genus Phaneromalthinus gen. nov.</p><p>Type species. Malthinellus malickyi Wittmer, 1997, hereby designated and monotypy.</p><p>Etymology. Derived from the Greek word phaneros, meaning “evident” or “conspicuous”, and the genus name Malthinus, referring to its close affinity. The name highlights the distinctive external appearance of the type species. Gender: masculine.</p><p>Diagnosis. Within the tribe Malthinini, this genus is recognized by the combination of the following characteristics: body relatively large (BL ca. 4.0– 5.3 mm); head moderately rounded, gradually narrowed posteriorly with rounded margines, not parallel-sided at neck, with the dorsum densely punctate; antennae slightly serrate; gular sutures separated from each other; gula broad and very deeply concave, not reaching the posterior margin of head, with an apodeme near the middle of the concavity; pronotum densely punctate, with both anterior and posterior margins bordered, lateral margins arcuate in dorsal view, hind angles protruding laterally, and epipleura moderately developed without any fissures; legs not modified as in some Malthinus species; pro- and mesocoxae elongate; metacoxae somewhat elongate, length approximately 1.3 times the basal width; tibiae each with one spur at apex; elytra longitudinally punctate and with three longitudinal costae, becoming indistinct posteriorly, long enough to completely cover hind wings and abdomen; sternite VIII widely emarginate in U-shape on posterior margin; male sternite IX weakly sclerotized, oval, lamellate, gently convex ventrally; general structure of aedeagus as in Malthinus; basal piece with basal shaft; inner basal piece with anterior terminal projecting dorsally and with a pair of slender arcuate processes directed posterad at the apical part; centrophyse with dorsally projected penis medially; female genitalia flattened, not three-dimensionally structured as in some Malthinus; proctiger distinct and wide but scarcely sclerotized; paraprocts slightly sclerotized, symmetrically divided, each nearly oval in ventral view, and each with a warped basal fragment connected to the lapel of coxite; each coxite slenderly ovate in ventral view, with a elongate lapel at the base; valvifers distinct and twisted, with apices somewhat membranous.</p><p>In Phaneromalthinus gen. nov., the gular sutures are separated from each other, differing from the condition in Malthinus species and Mimomalthinus niger Pic, but the deeply concave gula and the presence of the apodeme are also inconsistent with the diagnosis of Malthinellus . In addition, unlike Malthinus (excluding the subgenus Indomalthinus), Malthinellus and Mi. niger Pic, the elytra completely cover the hind wings and abdomen. The pronotum with prominent hind angles resembles that of Mi. shokhrini Kazantsev, but this species differs from it in the shape of the head and male abdominal sternite IX. See also Table 1.</p><p>Remarks. The structure of the aedeagus of Phaneromalthinus gen. nov. is mostly consistent with that of Malthinus, while the dorsal surface structure of the head, pronotum, and elytra resembles that of Malthinellus, suggesting a close relationship between the new genus and these genera. The entirely orange dorsal coloration and the presence of a cruciform sclerotized structure on the dorsal surface of the basal piece of the aedeagus may also represent autapomorphic characters of Phaneromalthinus gen. nov. However, the former is known from only one type species, and the latter cannot be confirmed due to the insufficient descriptions of the basal piece in many Malthinus species. Therefore, these characters are not included in the generic diagnosis in the present study.</p></div>	https://treatment.plazi.org/id/03EA87C1FFE62D088380FE65FA92F961	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFE62D098380F91DFE0FFA1D.text	03EA87C1FFE62D098380F91DFE0FFA1D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaneromalthinus malickyi (Nakamura 1997) Nakamura 2026	<div><p>Phaneromalthinus malickyi (Wittmer, 1997) comb. nov.</p><p>(Figs. 6B–N, 7F, N–P, 8M–O, 10)</p><p>Malthinellus malickyi Wittmer, 1997: 351 (original description), figs. 244–245 (aedeagus); N. Takahashi, 2018: 395 (checklist).</p><p>Type material examined. Holotype, male (NHMB), “ Thailand 25.XII.–2.I.90 / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.95&amp;materialsCitation.latitude=18.816668" title="Search Plazi for locations around (long 98.95/lat 18.816668)">Chiangmai Zoo Lichtfalle</a> / 400m, 98°57’E, 18°49’N / Chantaramongkol &amp; Malicky ”, “REM / 95 / 10.6”, “HOLOTYPUS”, “Maltinellus / malickyi / Wittm. / det. W.Wittmer ” . Paratypes, 3 males, 4 females (NHMB), same collecting data as the holotype .</p><p>Supplementary description. Pronotum (Figs. 6E–F, H–I) trapezoidal, sides slightly widened posteriorly, anterior angles angulate, lateral margins bordered and slightly sinuate near slightly produced hind angles, disc moderately convex, coarsely and densely punctate throughout, with a medio-longitudinal furrow not reaching the anterior or posterior margins, of which the length somewhat variable.</p><p>Aedeagus (Figs. 6K–L; figs. 244–245 in Wittmer, 1997) elongate, angular subelliptical in dorsal and ventral outline; basal piece rhomboidal, posterior margin shallowly emarginate in a V-shape, with a slender basal shaft in the anterior part, and with a cruciform sclerotized structure on dorsal surface; inner basal piece with anterior terminal curved dorsally, deeply divided into two parts, each apically expanded into a spatulate form, and with a pair of slender arcuate processes directed posterad at the apical part; penis robust, undivided, projecting dorsally, with a membranous sac at apex; centrophyse with an additional sclerotized process projecting dorso-anteriorly, similarly shaped to penis.</p><p>Female genitalia (Fig. 6N) flat; proctiger distinct and wide but scarcely sclerotized; paraprocts slightly sclerotized, symmetrically divided, each nearly oval in ventral outline, somewhat longer than the apical part of coxite, each with a warped basal fragment connected to the lapel of coxite; each coxite slenderly ovate in ventral outline, thicker than paraprocts, with a elongate lapel at the base; valvifers distinct and twisted, approximately as long as the paraprocts, with apices rather membranous.</p><p>Differential diagnosis. Without referring to the generic diagnosis, this species can be immediately distinguished from all other members of Malthinini by its highly distinctive coloration.</p><p>Remarks. Although the original description states that the spurs on the protibiae are short and almost invisible (Wittmer, 1997), the present observation shows that they are short but clearly visible (Fig. 7F). The original description gives the body length as “ 4–5 mm ”, but the largest female paratype measured approximately 5.3 mm.</p><p>The rationale behind Wittmer's placement of this species in Malthinellus is unclear.The structure of the aedeagus is mostly consistent with that of Malthinus, and I initially considered transferring the species to that genus. However, the peculiar external appearance and the head morphology do not fall within the range of variation observed in Malthinus species, and the establishment of a new genus was necessary to accept this species.</p><p>When Fanti and Castiglione (2017) described a new genus and species † Mantimalthinus balticus from Eocene Baltic amber, they placed it in the tribe Malthinini and stated that Malthinellus malickyi Wittmer (= Phaneromalthinus malickyi comb. nov.) was the only known species resembling it. However, the placement of † Man. balticus within Malthinini was uncertain, as its mandibles could not be observed. The two species are similar mainly in having long elytra covering the entire abdomen, but the former was different from the latter in the pronotum structure and the absence of tibial spurs and elytral costae (Fanti and Castiglione, 2017). In addition to these differences, † Man. balticus appears to have an elongate and sclerotized male sternite IX, which is clearly different from that of P. malickyi comb. nov. Subsequently, Fanti and Damgaard (2019) and Fanti (2022) added two more species to † Mantimalthinus, and the latter study transferred the genus to the tribe Malchinini .</p><p>Distribution. Thailand.</p></div>	https://treatment.plazi.org/id/03EA87C1FFE62D098380F91DFE0FFA1D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFE72D098380F9C1FBE5F96C.text	03EA87C1FFE72D098380F9C1FBE5F96C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mimomalthinus Pic 1931	<div><p>Genus Mimomalthinus Pic, 1931</p><p>Mimomalthinus Pic, 1931: 98 .</p><p>Type species: Mimomalthinus niger Pic, 1931, by monotypy.</p></div>	https://treatment.plazi.org/id/03EA87C1FFE72D098380F9C1FBE5F96C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
03EA87C1FFE72D0A8380F912FD8EFEE5.text	03EA87C1FFE72D0A8380F912FD8EFEE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mimomalthinus niger Pic 1931	<div><p>Mimomalthinus niger Pic, 1931</p><p>(Figs. 9A–C, 10)</p><p>Mimomalthinus niger Pic, 1931: 98 (original description); Brancucci, 1980: 305 (redescription, notes), figs. 17–18 (head), 78 (mandible), 126 (pronotum).</p><p>Type material examined. Lectotype (designated here), male (MNHN), “ Hoa / Binh ”, “Mimomalthinus / niger Pic”, “HOLOTYPUS”, “ Mimomalthinus / niger Pic / det. M. Brancucci ”.</p><p>Remarks. The total number of type specimens was not specified in the original description. Brancucci (1980) regarded the single specimen he examined as the holotype, but for the sake of nomenclatural stability, this specimen is designated as the lectotype here. No additional specimens of this species have been recorded or found.</p><p>Distribution. Vietnam (Hòa Bình).</p></div>	https://treatment.plazi.org/id/03EA87C1FFE72D0A8380F912FD8EFEE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nakamura, Ryo	Nakamura, Ryo (2026): A taxonomic study of Asian soldier beetles of the tribe Malthinini (Coleoptera, Cantharidae, Malthininae), with description of a new genus and a new species of Malthinellus, and notes on the genera Malthinus and Mimomalthinus. Zootaxa 5763 (1): 69-92, DOI: 10.11646/zootaxa.5763.1.3, URL: https://doi.org/10.11646/zootaxa.5763.1.3
