taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03EA87C1FFF42D1A83D0FDB1FB9FFD09.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191022/files/figure.png	https://doi.org/10.5281/zenodo.19191022	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF42D1A83D0FDB1FB9FFD09.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191024/files/figure.png	https://doi.org/10.5281/zenodo.19191024	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF42D1A83D0FD0DFB1AFCC1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191022/files/figure.png	https://doi.org/10.5281/zenodo.19191022	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF42D1A83D0FC91FE08FC31.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191020/files/figure.png	https://doi.org/10.5281/zenodo.19191020	FIGURE 6. Malthinus filiformis Wittmer, 1997 (A: female paratype) and Phaneromalthinus malickyi (Wittmer, 1997) comb. nov. (B: holotype; C, G–I, N: female paratype; D–F, J, K–M: male paratype). A–C, habitus (B: with labels); D & G, head, dorsal view; E & H, head, pronotum, scutellum and elytral basal area, dorsal view; F & I, ditto, lateral view; J, elytra, dorsal view; K, aedeagus, caudal view; L, ditto, lateral view; M, ditto, separated basal piece, ventral view; N, female genitalia, ventral view. Abbreviations: bs, basal shaft; c, coxite; lp, lapel of coxite; p, proctiger; pp, paraproct; v, valvifer.	FIGURE 6. Malthinus filiformis Wittmer, 1997 (A: female paratype) and Phaneromalthinus malickyi (Wittmer, 1997) comb. nov. (B: holotype; C, G–I, N: female paratype; D–F, J, K–M: male paratype). A–C, habitus (B: with labels); D & G, head, dorsal view; E & H, head, pronotum, scutellum and elytral basal area, dorsal view; F & I, ditto, lateral view; J, elytra, dorsal view; K, aedeagus, caudal view; L, ditto, lateral view; M, ditto, separated basal piece, ventral view; N, female genitalia, ventral view. Abbreviations: bs, basal shaft; c, coxite; lp, lapel of coxite; p, proctiger; pp, paraproct; v, valvifer.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF42D1A83D0FC91FE08FC31.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191022/files/figure.png	https://doi.org/10.5281/zenodo.19191022	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF42D1A83D0FC91FE08FC31.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191024/files/figure.png	https://doi.org/10.5281/zenodo.19191024	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF42D1A83D0FC25FD27FBE9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191022/files/figure.png	https://doi.org/10.5281/zenodo.19191022	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF42D1A83D0FBEDFA4EFBA1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191022/files/figure.png	https://doi.org/10.5281/zenodo.19191022	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF42D1A83D0FB95FD86FB59.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191005/files/figure.png	https://doi.org/10.5281/zenodo.19191005	FIGURE 3. Habitus of Malthinellus and similar Malthinus species. A, Malthine. crenulatomimus (Wittmer, 1984), holotype, male, with its labels; B, Malthine. crenulatus (Wittmer, 1979), holotype, male, with its labels; C, ditto, paratype, male; D, Malthinu. palingenesis Wittmer, 1993, holotype, male; E, Malthinu. sikkimensis Pic, 1924, non-type, male; F, ditto, non-type, female. Scale bars for habitus photographs are 1.0 mm each.	FIGURE 3. Habitus of Malthinellus and similar Malthinus species. A, Malthine. crenulatomimus (Wittmer, 1984), holotype, male, with its labels; B, Malthine. crenulatus (Wittmer, 1979), holotype, male, with its labels; C, ditto, paratype, male; D, Malthinu. palingenesis Wittmer, 1993, holotype, male; E, Malthinu. sikkimensis Pic, 1924, non-type, male; F, ditto, non-type, female. Scale bars for habitus photographs are 1.0 mm each.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF42D1A83D0FB5DFDB0FAC9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191005/files/figure.png	https://doi.org/10.5281/zenodo.19191005	FIGURE 3. Habitus of Malthinellus and similar Malthinus species. A, Malthine. crenulatomimus (Wittmer, 1984), holotype, male, with its labels; B, Malthine. crenulatus (Wittmer, 1979), holotype, male, with its labels; C, ditto, paratype, male; D, Malthinu. palingenesis Wittmer, 1993, holotype, male; E, Malthinu. sikkimensis Pic, 1924, non-type, male; F, ditto, non-type, female. Scale bars for habitus photographs are 1.0 mm each.	FIGURE 3. Habitus of Malthinellus and similar Malthinus species. A, Malthine. crenulatomimus (Wittmer, 1984), holotype, male, with its labels; B, Malthine. crenulatus (Wittmer, 1979), holotype, male, with its labels; C, ditto, paratype, male; D, Malthinu. palingenesis Wittmer, 1993, holotype, male; E, Malthinu. sikkimensis Pic, 1924, non-type, male; F, ditto, non-type, female. Scale bars for habitus photographs are 1.0 mm each.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF42D1A83D0FB5DFDB0FAC9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191022/files/figure.png	https://doi.org/10.5281/zenodo.19191022	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF42D1A83D0FACDFBCFFA65.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191007/files/figure.png	https://doi.org/10.5281/zenodo.19191007	FIGURE 4. Habitus and labels of Malthinellus and similar Malthinus species. A, Malthine. multispinosus (Wittmer, 1995) comb. nov., holotype, male; B, Malthine. hoabinensis (Wittmer, 1995) comb. nov., holotype, male; C, ditto, paratype, female; D, Malthinu. tonkineus Pic, 1923, lectotype, male. Scale bars for habitus photographs are 1.0 mm each.	FIGURE 4. Habitus and labels of Malthinellus and similar Malthinus species. A, Malthine. multispinosus (Wittmer, 1995) comb. nov., holotype, male; B, Malthine. hoabinensis (Wittmer, 1995) comb. nov., holotype, male; C, ditto, paratype, female; D, Malthinu. tonkineus Pic, 1923, lectotype, male. Scale bars for habitus photographs are 1.0 mm each.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF42D1A83D0FACDFBCFFA65.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191024/files/figure.png	https://doi.org/10.5281/zenodo.19191024	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF22D1E8380FA83FDCDFD7D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19190997/files/figure.png	https://doi.org/10.5281/zenodo.19190997	FIGURE 1. Malthinellus chinensis Wittmer, 1956, allotype, male. A, Habitus (before dissection), dorsal view; B, ditto, lateral view; C, head, dorsal view; D, head and pronotum, dorsal view; E, labels; F, aedeagus, caudal view; G, ditto, ventral view; H, ditto, dorsal view, I, ditto, dorso-lateral view; J, ditto, lateral view. Abbreviations: al, anterior lobe; at, anterior terminal of inner basal piece; bp, basal piece; ibp, inner basal piece; ol, outer lateral lobe; pe, penis.	FIGURE 1. Malthinellus chinensis Wittmer, 1956, allotype, male. A, Habitus (before dissection), dorsal view; B, ditto, lateral view; C, head, dorsal view; D, head and pronotum, dorsal view; E, labels; F, aedeagus, caudal view; G, ditto, ventral view; H, ditto, dorsal view, I, ditto, dorso-lateral view; J, ditto, lateral view. Abbreviations: al, anterior lobe; at, anterior terminal of inner basal piece; bp, basal piece; ibp, inner basal piece; ol, outer lateral lobe; pe, penis.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF22D1E8380FA83FDCDFD7D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191022/files/figure.png	https://doi.org/10.5281/zenodo.19191022	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF22D1E8380FA83FDCDFD7D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191024/files/figure.png	https://doi.org/10.5281/zenodo.19191024	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF22D1E8380FA83FDCDFD7D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191033/files/figure.png	https://doi.org/10.5281/zenodo.19191033	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF02D108380F929FCD0F89E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191005/files/figure.png	https://doi.org/10.5281/zenodo.19191005	FIGURE 3. Habitus of Malthinellus and similar Malthinus species. A, Malthine. crenulatomimus (Wittmer, 1984), holotype, male, with its labels; B, Malthine. crenulatus (Wittmer, 1979), holotype, male, with its labels; C, ditto, paratype, male; D, Malthinu. palingenesis Wittmer, 1993, holotype, male; E, Malthinu. sikkimensis Pic, 1924, non-type, male; F, ditto, non-type, female. Scale bars for habitus photographs are 1.0 mm each.	FIGURE 3. Habitus of Malthinellus and similar Malthinus species. A, Malthine. crenulatomimus (Wittmer, 1984), holotype, male, with its labels; B, Malthine. crenulatus (Wittmer, 1979), holotype, male, with its labels; C, ditto, paratype, male; D, Malthinu. palingenesis Wittmer, 1993, holotype, male; E, Malthinu. sikkimensis Pic, 1924, non-type, male; F, ditto, non-type, female. Scale bars for habitus photographs are 1.0 mm each.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF02D108380F929FCD0F89E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191011/files/figure.png	https://doi.org/10.5281/zenodo.19191011	FIGURE 5. Aedeagi of Malthinellus species. A, Malthine. multispinosus (Wittmer, 1995) comb. nov., dorso-lateral view; B, ditto, dorsal view; C, Malthine. hoabinensis (Wittmer, 1995) comb. nov., dorsal view; D, ditto, dorso-lateral view; E, Malthine. crenulatus (Wittmer, 1979), lateral view; F, ditto, dorsal view; G, Malthine. crenulatomimus (Wittmer, 1984), dorso-lateral view; H, ditto, dorsal view. Not to scale. A–D: modified from Wittmer (1995), E–F: modified from Wittmer (1979), G–H: modified from Wittmer (1984). Abbreviations: at, anterior terminal of inner basal piece; bp, basal piece; ibp, inner basal piece; il, inner lateral lobe; ol, outer lateral lobe; pe, penis.	FIGURE 5. Aedeagi of Malthinellus species. A, Malthine. multispinosus (Wittmer, 1995) comb. nov., dorso-lateral view; B, ditto, dorsal view; C, Malthine. hoabinensis (Wittmer, 1995) comb. nov., dorsal view; D, ditto, dorso-lateral view; E, Malthine. crenulatus (Wittmer, 1979), lateral view; F, ditto, dorsal view; G, Malthine. crenulatomimus (Wittmer, 1984), dorso-lateral view; H, ditto, dorsal view. Not to scale. A–D: modified from Wittmer (1995), E–F: modified from Wittmer (1979), G–H: modified from Wittmer (1984). Abbreviations: at, anterior terminal of inner basal piece; bp, basal piece; ibp, inner basal piece; il, inner lateral lobe; ol, outer lateral lobe; pe, penis.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF02D108380F929FCD0F89E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191024/files/figure.png	https://doi.org/10.5281/zenodo.19191024	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF02D108380F929FCD0F89E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191033/files/figure.png	https://doi.org/10.5281/zenodo.19191033	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFFF2D148380FAA4FD58FCA5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191005/files/figure.png	https://doi.org/10.5281/zenodo.19191005	FIGURE 3. Habitus of Malthinellus and similar Malthinus species. A, Malthine. crenulatomimus (Wittmer, 1984), holotype, male, with its labels; B, Malthine. crenulatus (Wittmer, 1979), holotype, male, with its labels; C, ditto, paratype, male; D, Malthinu. palingenesis Wittmer, 1993, holotype, male; E, Malthinu. sikkimensis Pic, 1924, non-type, male; F, ditto, non-type, female. Scale bars for habitus photographs are 1.0 mm each.	FIGURE 3. Habitus of Malthinellus and similar Malthinus species. A, Malthine. crenulatomimus (Wittmer, 1984), holotype, male, with its labels; B, Malthine. crenulatus (Wittmer, 1979), holotype, male, with its labels; C, ditto, paratype, male; D, Malthinu. palingenesis Wittmer, 1993, holotype, male; E, Malthinu. sikkimensis Pic, 1924, non-type, male; F, ditto, non-type, female. Scale bars for habitus photographs are 1.0 mm each.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFFF2D148380FAA4FD58FCA5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191011/files/figure.png	https://doi.org/10.5281/zenodo.19191011	FIGURE 5. Aedeagi of Malthinellus species. A, Malthine. multispinosus (Wittmer, 1995) comb. nov., dorso-lateral view; B, ditto, dorsal view; C, Malthine. hoabinensis (Wittmer, 1995) comb. nov., dorsal view; D, ditto, dorso-lateral view; E, Malthine. crenulatus (Wittmer, 1979), lateral view; F, ditto, dorsal view; G, Malthine. crenulatomimus (Wittmer, 1984), dorso-lateral view; H, ditto, dorsal view. Not to scale. A–D: modified from Wittmer (1995), E–F: modified from Wittmer (1979), G–H: modified from Wittmer (1984). Abbreviations: at, anterior terminal of inner basal piece; bp, basal piece; ibp, inner basal piece; il, inner lateral lobe; ol, outer lateral lobe; pe, penis.	FIGURE 5. Aedeagi of Malthinellus species. A, Malthine. multispinosus (Wittmer, 1995) comb. nov., dorso-lateral view; B, ditto, dorsal view; C, Malthine. hoabinensis (Wittmer, 1995) comb. nov., dorsal view; D, ditto, dorso-lateral view; E, Malthine. crenulatus (Wittmer, 1979), lateral view; F, ditto, dorsal view; G, Malthine. crenulatomimus (Wittmer, 1984), dorso-lateral view; H, ditto, dorsal view. Not to scale. A–D: modified from Wittmer (1995), E–F: modified from Wittmer (1979), G–H: modified from Wittmer (1984). Abbreviations: at, anterior terminal of inner basal piece; bp, basal piece; ibp, inner basal piece; il, inner lateral lobe; ol, outer lateral lobe; pe, penis.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFFF2D148380FAA4FD58FCA5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191024/files/figure.png	https://doi.org/10.5281/zenodo.19191024	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFFF2D148380FAA4FD58FCA5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191033/files/figure.png	https://doi.org/10.5281/zenodo.19191033	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFFA2D158380FC54FDF0FA81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191003/files/figure.png	https://doi.org/10.5281/zenodo.19191003	FIGURE 2. Malthinellus mushanus sp. nov., males. A, Habitus, holotype; B, ditto, paratype from Meifeng; C, head and pronotum, holotype; D, ditto, paratype from Meifeng; E, aedeagus, caudal view; F, ditto, ventral view; G, ditto, dorsal view, H, ditto, lateral view; I, dorso-lateral view.Abbreviations: at, anterior terminal of inner basal piece; bp, basal piece; ibp, inner basal piece; il, inner lateral lobe; ol1, outer lateral lobe 1; ol2, outer lateral lobe 2; pe, penis.	FIGURE 2. Malthinellus mushanus sp. nov., males. A, Habitus, holotype; B, ditto, paratype from Meifeng; C, head and pronotum, holotype; D, ditto, paratype from Meifeng; E, aedeagus, caudal view; F, ditto, ventral view; G, ditto, dorsal view, H, ditto, lateral view; I, dorso-lateral view.Abbreviations: at, anterior terminal of inner basal piece; bp, basal piece; ibp, inner basal piece; il, inner lateral lobe; ol1, outer lateral lobe 1; ol2, outer lateral lobe 2; pe, penis.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFFA2D158380FC54FDF0FA81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191022/files/figure.png	https://doi.org/10.5281/zenodo.19191022	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFFA2D158380FC54FDF0FA81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191024/files/figure.png	https://doi.org/10.5281/zenodo.19191024	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFFA2D158380FC54FDF0FA81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191033/files/figure.png	https://doi.org/10.5281/zenodo.19191033	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF92D178380FF60FD9DFB8A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191007/files/figure.png	https://doi.org/10.5281/zenodo.19191007	FIGURE 4. Habitus and labels of Malthinellus and similar Malthinus species. A, Malthine. multispinosus (Wittmer, 1995) comb. nov., holotype, male; B, Malthine. hoabinensis (Wittmer, 1995) comb. nov., holotype, male; C, ditto, paratype, female; D, Malthinu. tonkineus Pic, 1923, lectotype, male. Scale bars for habitus photographs are 1.0 mm each.	FIGURE 4. Habitus and labels of Malthinellus and similar Malthinus species. A, Malthine. multispinosus (Wittmer, 1995) comb. nov., holotype, male; B, Malthine. hoabinensis (Wittmer, 1995) comb. nov., holotype, male; C, ditto, paratype, female; D, Malthinu. tonkineus Pic, 1923, lectotype, male. Scale bars for habitus photographs are 1.0 mm each.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF92D178380FF60FD9DFB8A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191011/files/figure.png	https://doi.org/10.5281/zenodo.19191011	FIGURE 5. Aedeagi of Malthinellus species. A, Malthine. multispinosus (Wittmer, 1995) comb. nov., dorso-lateral view; B, ditto, dorsal view; C, Malthine. hoabinensis (Wittmer, 1995) comb. nov., dorsal view; D, ditto, dorso-lateral view; E, Malthine. crenulatus (Wittmer, 1979), lateral view; F, ditto, dorsal view; G, Malthine. crenulatomimus (Wittmer, 1984), dorso-lateral view; H, ditto, dorsal view. Not to scale. A–D: modified from Wittmer (1995), E–F: modified from Wittmer (1979), G–H: modified from Wittmer (1984). Abbreviations: at, anterior terminal of inner basal piece; bp, basal piece; ibp, inner basal piece; il, inner lateral lobe; ol, outer lateral lobe; pe, penis.	FIGURE 5. Aedeagi of Malthinellus species. A, Malthine. multispinosus (Wittmer, 1995) comb. nov., dorso-lateral view; B, ditto, dorsal view; C, Malthine. hoabinensis (Wittmer, 1995) comb. nov., dorsal view; D, ditto, dorso-lateral view; E, Malthine. crenulatus (Wittmer, 1979), lateral view; F, ditto, dorsal view; G, Malthine. crenulatomimus (Wittmer, 1984), dorso-lateral view; H, ditto, dorsal view. Not to scale. A–D: modified from Wittmer (1995), E–F: modified from Wittmer (1979), G–H: modified from Wittmer (1984). Abbreviations: at, anterior terminal of inner basal piece; bp, basal piece; ibp, inner basal piece; il, inner lateral lobe; ol, outer lateral lobe; pe, penis.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF92D178380FF60FD9DFB8A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191024/files/figure.png	https://doi.org/10.5281/zenodo.19191024	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF92D178380FF60FD9DFB8A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191033/files/figure.png	https://doi.org/10.5281/zenodo.19191033	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF92D088380FB33FD8EFEB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191007/files/figure.png	https://doi.org/10.5281/zenodo.19191007	FIGURE 4. Habitus and labels of Malthinellus and similar Malthinus species. A, Malthine. multispinosus (Wittmer, 1995) comb. nov., holotype, male; B, Malthine. hoabinensis (Wittmer, 1995) comb. nov., holotype, male; C, ditto, paratype, female; D, Malthinu. tonkineus Pic, 1923, lectotype, male. Scale bars for habitus photographs are 1.0 mm each.	FIGURE 4. Habitus and labels of Malthinellus and similar Malthinus species. A, Malthine. multispinosus (Wittmer, 1995) comb. nov., holotype, male; B, Malthine. hoabinensis (Wittmer, 1995) comb. nov., holotype, male; C, ditto, paratype, female; D, Malthinu. tonkineus Pic, 1923, lectotype, male. Scale bars for habitus photographs are 1.0 mm each.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF92D088380FB33FD8EFEB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191011/files/figure.png	https://doi.org/10.5281/zenodo.19191011	FIGURE 5. Aedeagi of Malthinellus species. A, Malthine. multispinosus (Wittmer, 1995) comb. nov., dorso-lateral view; B, ditto, dorsal view; C, Malthine. hoabinensis (Wittmer, 1995) comb. nov., dorsal view; D, ditto, dorso-lateral view; E, Malthine. crenulatus (Wittmer, 1979), lateral view; F, ditto, dorsal view; G, Malthine. crenulatomimus (Wittmer, 1984), dorso-lateral view; H, ditto, dorsal view. Not to scale. A–D: modified from Wittmer (1995), E–F: modified from Wittmer (1979), G–H: modified from Wittmer (1984). Abbreviations: at, anterior terminal of inner basal piece; bp, basal piece; ibp, inner basal piece; il, inner lateral lobe; ol, outer lateral lobe; pe, penis.	FIGURE 5. Aedeagi of Malthinellus species. A, Malthine. multispinosus (Wittmer, 1995) comb. nov., dorso-lateral view; B, ditto, dorsal view; C, Malthine. hoabinensis (Wittmer, 1995) comb. nov., dorsal view; D, ditto, dorso-lateral view; E, Malthine. crenulatus (Wittmer, 1979), lateral view; F, ditto, dorsal view; G, Malthine. crenulatomimus (Wittmer, 1984), dorso-lateral view; H, ditto, dorsal view. Not to scale. A–D: modified from Wittmer (1995), E–F: modified from Wittmer (1979), G–H: modified from Wittmer (1984). Abbreviations: at, anterior terminal of inner basal piece; bp, basal piece; ibp, inner basal piece; il, inner lateral lobe; ol, outer lateral lobe; pe, penis.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFF92D088380FB33FD8EFEB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191033/files/figure.png	https://doi.org/10.5281/zenodo.19191033	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFE62D098380F91DFE0FFA1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191020/files/figure.png	https://doi.org/10.5281/zenodo.19191020	FIGURE 6. Malthinus filiformis Wittmer, 1997 (A: female paratype) and Phaneromalthinus malickyi (Wittmer, 1997) comb. nov. (B: holotype; C, G–I, N: female paratype; D–F, J, K–M: male paratype). A–C, habitus (B: with labels); D & G, head, dorsal view; E & H, head, pronotum, scutellum and elytral basal area, dorsal view; F & I, ditto, lateral view; J, elytra, dorsal view; K, aedeagus, caudal view; L, ditto, lateral view; M, ditto, separated basal piece, ventral view; N, female genitalia, ventral view. Abbreviations: bs, basal shaft; c, coxite; lp, lapel of coxite; p, proctiger; pp, paraproct; v, valvifer.	FIGURE 6. Malthinus filiformis Wittmer, 1997 (A: female paratype) and Phaneromalthinus malickyi (Wittmer, 1997) comb. nov. (B: holotype; C, G–I, N: female paratype; D–F, J, K–M: male paratype). A–C, habitus (B: with labels); D & G, head, dorsal view; E & H, head, pronotum, scutellum and elytral basal area, dorsal view; F & I, ditto, lateral view; J, elytra, dorsal view; K, aedeagus, caudal view; L, ditto, lateral view; M, ditto, separated basal piece, ventral view; N, female genitalia, ventral view. Abbreviations: bs, basal shaft; c, coxite; lp, lapel of coxite; p, proctiger; pp, paraproct; v, valvifer.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFE62D098380F91DFE0FFA1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191022/files/figure.png	https://doi.org/10.5281/zenodo.19191022	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	FIGURE 7. Left protibiae (A–F) and heads (G–P) of Malthinellus (A–B, G–I), Malthinus (C–E, J–M) and Phaneromalthinus gen. nov. (F, N–P) species. A & G, Malthine. bicolor Kiesenwetter, 1874 (male); B & I, Malthine. mushanus sp. nov. (male, B: holotype, I: paratype); C & J, Malthinu. flaveolus (Herbst, 1786) (C: male, J: female); D & K, Malthinu. sikkimensis Pic, 1924 (female); E & M, Malthinu. humeralis Kiesenwetter, 1874 (female); F, N, O & P, P. malickyi (Wittmer, 1997) comb. nov. (F: female, paratype, N–P: male, paratype); H, Malthine. iheyanus Takahashi & Nakamura, 2022 (female, paratype); L, Malthinu. filiformis Wittmer, 1997 (female, paratype). A–F, O: dorsal view, G–N, P: ventral view. White arrows indicate the position of the gula. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFE62D098380F91DFE0FFA1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191024/files/figure.png	https://doi.org/10.5281/zenodo.19191024	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	FIGURE 8. Male terminal abdominal ventrites (A–D), sternite VIII (O), sternite IX (E–M) and tergite IX (N) of Malthinellus (A–J), Malthinus (K–L) and Phaneromalthinus gen. nov. (M–O) species. A, E & F, Malthine. bicolor Kiesenwetter, 1874; B, Malthine. chinensis Wittmer, 1956 (allotype); C & H, Malthine. mushanus sp. nov. (paratype); D & I, Malthine. crenulatomimus (Wittmer, 1984); G, Malthine. multispinosus (Wittmer, 1995) comb. nov. (holotype); J, Malthine. crenulatus (Wittmer, 1979) (paratype); K, Malthinu. tonkineus Pic, 1923 (lectotype); L, Malthinu. filiformis Wittmer, 1997 (holotype); M–O, P. malickyi (Wittmer, 1997) comb. nov. (paratype). A–E, G–M, O: ventral view, F: lateral view, N: dorsal view. Not to scale.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFE62D098380F91DFE0FFA1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191033/files/figure.png	https://doi.org/10.5281/zenodo.19191033	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFE72D0A8380F912FD8EFEE5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191030/files/figure.png	https://doi.org/10.5281/zenodo.19191030	FIGURE 9. Mimomalthinus niger Pic, 1931, lectotype, male (dissected and mounted by the late Michel Brancucci).A, Habitus; B, labels; C, abdomen.	FIGURE 9. Mimomalthinus niger Pic, 1931, lectotype, male (dissected and mounted by the late Michel Brancucci).A, Habitus; B, labels; C, abdomen.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
03EA87C1FFE72D0A8380F912FD8EFEE5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19191033/files/figure.png	https://doi.org/10.5281/zenodo.19191033	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	FIGURE 10. Distribution map of Malthinellus (circle symbols), Mimomalthinus (diamond symbols) and Phaneromalthinus gen. nov. (star symbol) species. The distribution range of Malthine. bicolor is only outlined, not plotted.	2026-03-02	Nakamura, Ryo		Zenodo	biologists	Nakamura, Ryo			
