taxonID	type	description	language	source
03EC5B01BB71FF90FF79FB3DFEA2EC2A.taxon	description	(Fig. 1)	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB71FF90FF79FB3DFEA2EC2A.taxon	materials_examined	Type material. HOLOTYPE ♂. France, Cervières (05027), Refuge Napoléon, 44.8226, 6.7279, 2300 m, 23. vii. 2025, D. Langlois, unimproved subalpine grassland MNHN, ED 13471. PARATYPES: FRANCE. 1 ♀, Abries-Ristolas (05001), Ristolas-Mont-Viso RNN, Ovin bas, 44.7046, 7.0387, 2160 m, 27. vi. – 18. vii. 2017, B. Tissot, TM 73, Hygrophilic thicket with Salix glaucosericea and Rhododendron ferrugineum, MNHN, ED 13472 & coll DL; 1 ♂, Cervières (05027), Refuge Napoléon, 44.8230, 6.7283, 2300 m, 23. vii. 2025, L. Fisler, unimproved subalpine grassland, coll DL; 1 ♂, 1 ♀, Puy-Saint-André (05107) Partias RNR, 44.8883, 6.5644, 2088 m, 10. – 27. vii. 2022, C. Lauriaut, TM 16, mesophilous subalpine grassland and Larix forest, coll. DL.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB71FF90FF79FB3DFEA2EC2A.taxon	etymology	Etymology. Named to highlight the very particular outgrowths on tergite 8 and sternite 7.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB71FF90FF79FB3DFEA2EC2A.taxon	diagnosis	Diagnosis. Medium-sized greyish species (5.5 – 6 mm), with swellings on each side of tergite 8; cerci enclosed within epandrium, rather thin phallus S-shaped at tip; anal vein complete, dusted thorax and abdomen, pale halter, recurrent outgrowth under posterior part of sternite 7 topped with a tuft of black setae, wing hyaline in male, infuscated in female; legs of female with pennate scales.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB71FF90FF79FB3DFEA2EC2A.taxon	distribution	Distribution. France, Southern Alps (05) (Fig. 4)	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB71FF90FF79FB3DFEA2EC2A.taxon	description	Description. Male (Fig. 1 A, B, D, E). Head. Occiput dull black to dusted grey, covered with black setae; row of black postoculars curved towards front; postgena with pale hairs; dusted ocellar triangle. Eyes holoptic with upper ommatidia enlarged. Frons reduced to small sub-shiny triangle above antennae. Face and clypeus dusted, dull brown. Antenna dull brownish; scape slightly longer and thinner than rounded pedicel; postpedicel 2.5 times longer than width at base, 2.5 times scape length. Stylus almost as long as postpedicel. Proboscis brownish, labrum about twice head height; labellum long and slender; palpus brown with a few black bristly hairs. Thorax dull black with grey pruinosity. Antepronotum with black setae above and white hairs on side. Proepisternum with tuft of white hairs. Postpronotum with 2 long black setae and slightly shorter white hairs. Acrostichals biserial, rather short, absent in prescutellar depression; dorsocentrals biserial, slightly longer than acrostichals, ending in 3 long setae in prescutellar depression. Laterotergite with pale fan. Scutellum with 2 long apical black setae, 2 finer and shorter subapicals. Other strong, long black setae as follows: 3 presutural supra-alars, 3 notopleurals, 4 postsutural supra-alars including smaller one, 1 postalar. Legs dark brown. Coxae with pale hairs and setae. Fore femur with short brown dorsal setae, ventrals less numerous; fore tibia with posterodorsal row of setae as long as tibia depth, ventrals more numerous and shorter, with 4 apical setae; tarsomeres with dorsal and apical setae, with shorter and paler ventral hairs. Mid femur with dorsal brown setae shorter than femur depth, 2 anteroventral and posteroventral rows of strong setae longer than femur depth, except apically; mid tibia with 2 anteroventral and posteroventral rows of rather strong setae, as long as tibia depth, 4 strong posterodorsal setae 4 times longer than tibia depth; tarsomeres with ventral and apical setae, thinner setae and hairs dorsally. Hind femur with many brown dorsal setae shorter than femur depth, a few longer than femur depth ventrally; hind tibia with many ventral setae, shorter or as long as tibia depth, dorsals less numerous but mixed with a few longer; hind tarsomere 1 somewhat swollen with setae all around, remaining tarsomeres with rather short setae stronger ventrally. Wing (length = 6.1 – 7.2 mm) hyaline, distinct dark pterostigma. Anal vein complete. Halter pale yellow with brown stem; pale calypter with white haired fringe. Abdomen dull brown; tergites mainly white haired, longer on side, mixed with black hairs dorsally and black setae posteriorly; 2 swellings on each side of tergite 8 (Fig. 1 B, E), partly shiny; sternites with white hairs ventrally as long as lateral tergal setae but less numerous; tuft of black setae, paler apically, on posterior part of sternites 3 – 6 (Fig. 1 B); posterior part of sternite 7 with recurrent outgrowth topped with tuft of black setae, paler apically (Fig. 1 B, E); sternite 8 with yellowish long hairs ventrally and row of long yellowish setae posteriorly. Hypopygium (Fig. 1 B, D, E) dull brown, mainly with white hairs and setae; cercus enclosed within epandrium, partly shiny with dorsal fan of brownish setae; elongated and subtriangular epandrial lamella with long pale setae on posterior margin; hypandrium bare, forming subtriangular ventral plate, narrowed along base of phallus; phallus yellowish, enlarged along hypandrium, narrowed and enclosed within epandrium, S-shaped at tip (Fig. 1 D). Female (Fig. 1 C). Similar to male except for sexual differences and following characters: eyes dichoptic, all ommatidia of equal size; frons as wide as face; wing infuscated and enlarged; sternites without outgrowths and tufts of setae; tergite 8 without swellings. Some of leg parts with pennate scales (absent on fore femur and mid coxa): fore tibia with dorsal pennate scales longer than tibia depth, ventral pennate scales shorter; mid femur with dorsal and ventral pennate scales longer than femur depth; mid tibia with dorsal pennate scales longer than tibia depth, except at tip, short ventral pennate scales at base; mid tarsomere 1 with short dorsal and ventral pennate scales; hind femur with dorsal pennate scales as long as femur depth, ventral pennate scales longer; hind tibia with dorsal pennate scales longer than tibia depth and ventral pennate scales at basal half; hind tarsomere 1 less swollen, with dorsal pennate scales.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB71FF90FF79FB3DFEA2EC2A.taxon	discussion	Remarks. Empis inaequalis sp. nov. is difficult to classify among the nine Empis - groups proposed by Chvála & Pont (2015). The male is very characteristic with the hairy outgrowth under sternite 7, the two swellings on tergite 8 and the shape of the phallus. Many species belonging to Empis (s. str.) nigripes - group (Syrovátka 1991 a; Kustov & Shamshev 2014 a) or Empis (s. str.) simulium - group (Chvála & Pont 2016) and occurring in Europe have swellings on tergite 8 or on its sides. But these swellings look like tubercles or are finger-like, those of E. inaequalis sp. nov. have a single projection, are more voluminous or less sharp. And none of E. nigripes - group or E. simulium - group species have such a recurrent outgrowth under sternite 7. Empis malleola Becker, 1887 (Syrovátka 1991 a, fig. 7) have a characteristic outgrowth under posterior sternites, but that of E. inaequalis sp. nov. is recurrent; secondly the phallus of E. malleola is short, not S-shaped at tip. The shape of the apex of the phallus of E. inaequalis sp. nov. is close to that of E. nuntia Meigen, 1838 (Collin 1961, fig. 195) and E. pseudonuntia Syrovátka, 1991 (Syrovátka 1991 b, fig. 4). The phallus of these two species of Empis end in a complete loop while that of E. inaequalis sp. nov. is just S-shaped. In E. nuntia, the phallus is stouter at the level of the loop and sternite 7 has no outgrowth. Empis pseudonuntia has a very small outgrowth under sternite 7 (Syrovátka did not mention this in his description of the species but it is visible in figure 4) but lacks a swelling on the side of tergite 8 and the female fore femora has a few pennate scales.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB73FF96FF79FA71FE24ECCE.taxon	description	(Fig. 2)	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB73FF96FF79FA71FE24ECCE.taxon	materials_examined	Type material. HOLOTYPE ♂. France, Frotey-lès-Vesoul (70261), Sabot de Frotey RNN, 47.6354, 6.1821, 304 m, 20. vi. – 3. vii. 2023, B. Tissot, TM 180, calcareous grassland, MNHN, ED 13473. PARATYPES: FRANCE. 1 ♂, Frotey-lès-Vesoul (70261), Sabot de Frotey RNN, 47.6300, 6.1860, 327 m, 02. – 20. vi. 2023, B. Tissot, TM 181, calcareous grassland, coll. DL; 1 ♀, Frotey-lès-Vesoul (70261), Sabot de Frotey RNN, 47.6354, 6.1821, 304 m, 20. vi. – 3. vii. 2023, B. Tissot, TM 180, calcareous grassland, MNHN, ED 13474; 1 ♂, 1 ♀, Solutré-Pouilly (71526), Roche de Solutré, 46.3033, 4.7226, 403 m, 09. vi. – 05. vii. 2023, R. Decoin, TM 168, marly grassland, MNHN ED 13475 & ED 13476; 1 ♂, Pouilly-sur-Loire (58215), Val de Loire RNN, Les loges, 47.2921, 2.9381, 152 m, 22. v. – 09. vi. 2023, R. Decoin, TM 187, alluvial forest and grassland, coll DL; 1 ♂, Mailly-le-château (89238), Bois du parc RNN, 47.5861, 3.6428, 152 m, 02. – 19. vi. 2020, R. Decoin, TM 137, Quercus forest and calcareous grassland, coll DL.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB73FF96FF79FA71FE24ECCE.taxon	etymology	Etymology. In honor of Hugues Pinston, who has managed the Sabot de Frotey RNN for 30 years, where this species was collected, recognizing his strong contribution to the knowledge of nature in Franche-Comté as well as in Burgundy.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB73FF96FF79FA71FE24ECCE.taxon	diagnosis	Diagnosis. Rather small greyish species (4.3 – 4.8 mm), with brown halter, black laterotergite fan; anal vein partly indistinct, wing clear in both sexes; cercus longer than epandrial lamella, phallus thin and S-shaped at tip; dusted thorax; white abdominal pubescence mixed with long dark setae; pennate scales of female legs confined to mid femur, and mid and hind tibiae.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB73FF96FF79FA71FE24ECCE.taxon	distribution	Distribution. France: Bourgogne-Franche-Comté (Fig. 4)	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB73FF96FF79FA71FE24ECCE.taxon	description	Description. Male (Fig. 2 A, B, C, E). Head. Occiput dull black, 2 rows of black setae; postoculars curved towards front; postgena with paler hairs. Eyes holoptic with upper ommatidia enlarged. Frons reduced to small triangle above antennae; equilateral ocellar triangle with black setae, 2 longer at centre, posterior ocelli close to eye margin. Face and clypeus dusted black. Antenna dull, blackish, scape as long as but thinner than rounded pedicel; postpedicel 3 times longer than width at base, 3.5 times scape length and 2 times stylus length. Proboscis blackish, labrum about twice head height; labellum long and slender; palpus black with a few black bristly-hairs, one longer at tip. Thorax dull black with grey pruinosity. Antepronotum with row of short black setae. Proepisternum with small tuft of black hairs. Postpronotum with one long black seta and shorter ones. Acrostichals biserial, rather long, absent in prescutellar depression; dorsocentrals irregularly biserial, slightly longer than acrostichals, ending in 2 strong, long setae in prescutellar depression. Laterotergite with fan of black setae. Scutellum with 2 long intersecting apical setae. Other strong, long black setae as follows: 2 presutural supra-alars, 3 notopleurals, 3 postsutural supra-alars including 2 small ones close to suture, 2 postalars including small one. Legs dark brown, slightly dull, with brown setae and bristles. Fore and mid coxae dull with brownish hairs and setae; hind coxa dull with a few longer setae slightly longer than coxa depth. Fore femur with very short dorsal setae, thinner and longer apically and ventrally; fore tibia with anterodorsal and posterodorsal rows of distinct setae as long as tibia depth, longer apically, ventrals much shorter; tarsomeres with dorsal setae, longer apically, shorter ventrally. Mid femur with white hairs mixed with row of anterodorsal setae and 2 rows of anteroventral and posteroventral setae, all rows 2 – 3 times longer than femur depth, becoming shorter apically; mid tibia with small setae mixed with 3 strong anterodorsal setae (2 near base and 1 apically), 4 times longer than tibia depth and 2 rows of anteroventral and posteroventral setae twice longer than tibia depth; mid tarsomeres with setae, longer apically. Hind femur with short setae mixed with row of anterodorsal setae as long as femur depth in middle, slightly longer at base and apically, with 2 rows of anteroventral and posteroventral setae as long as femur depth; hind tibia basally slender, stouter towards tip, with 2 rows of anterodorsal and posterodorsal setae, shorter than tibia depth basally, longer apically, ventrally densely short bristled; hind tarsomere 1 somewhat swollen but more slender than tip of hind tibia, with distinct apical setae and one longer bristle dorsally in middle, remaining tarsomeres with rather short setae, longer apically. Wing (length = 3.4 – 3.5 mm) hyaline, distinct brown pterostigma. Anal vein partly indistinct between middle and wing margin. Halter brown although pale in Figure 2 A due to reverberation of lamp on specimen in alcohol (with dried specimen, halter clearly brown); calypter dark with black haired fringe. Abdomen brown, slightly dull. White abdominal pubescence short, mixed with long dark setae, especially on posterior part of each tergite and at centre of posterior part of each sternite, except for sternite 1. Tergite 1 with many long dark setae posteriorly. Hypopygium brown (Fig. 2 B, C, E), slightly dull, with black hairs and setae; cercus longer than epandrial lamella, thin, curved and pointed forwards, with dorsal fan of fine setae, denser at base (Fig. 2 B); epandrial lamella triangular, about 2.5 times higher than wide, with long setae on posterior margin (Fig. 2 B); hypandrium forming broad short haired ventral plate; phallus yellowish, thin, S-shaped and thinner at tip (Fig. 2 C, E), just reaching dorsal part of cerci (Fig. 2 B). Female (Fig. 2 D). Similar to male except for usual sexual differences and following characters: setae shorter; eyes dichoptic, all ommatidia of equal size; frons as wide as face with row of setae along margin of eyes. Some leg parts with pennate scales: mid femur with dorsal pennate scales as long as femur depth, shorter apically, ventral pennate scales shorter, absent basally; hind femur with dorsal pennate scales as long as depth of femur, absent apically, ventral pennate scales as long as femur depth, absent basally; hind tibia with dorsal pennate scales as long as depth of tibia; hind tarsomere 1 less swollen.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB73FF96FF79FA71FE24ECCE.taxon	discussion	Remarks. Again, this species is difficult to classify among the nine Empis - groups proposed by Chvála & Pont (2015). Empis pinstoni sp. nov. genitalia are close to those of E. intercepta Melander, 1928 (Syrovátka 1991 a, fig 5). The form of the hypandrium, epandrial lamella and phallus are very similar but the cercus of E. pinstoni sp. nov. is much longer than the epandrial lamella, thinner, curved and pointed, the apex of the phallus is erected forward while that of E. intercepta is erected backward. In addition, E. intercepta has pale halteres, mid femora with 4 – 6 very long pale bristles posteroventrally, hind femora with a very long bristle anterodorsally in the middle and female legs are only short ciliated.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB75FF94FF79FA55FC20EDBA.taxon	description	(Fig. 3)	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB75FF94FF79FA55FC20EDBA.taxon	materials_examined	Type material. HOLOTYPE ♂. France, Monéteau (89263), Bois de grand Pien, 47.8678, 3.5953, 110 m, 13. viii. – 14. x. 2024, R. Decoin, TM 197, wooded fen, MNHN, ED 13477. PARATYPE: 1 ♂ with same data as holotype, coll. DL.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB75FF94FF79FA55FC20EDBA.taxon	etymology	Etymology. In honor of Bruno Tissot, recognizing his strong involvement in the development of knowledge of Diptera in the network of French nature reserves and beyond.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB75FF94FF79FA55FC20EDBA.taxon	diagnosis	Diagnosis. Small greyish species (4 mm) belonging to the subgenus Coptophlebia, with vein M 1 complete, epandrium with a long dorsal projection, dusted thorax and shiny abdomen, white abdominal pubescence mixed with black setae in the centre of the tergites, brownish halter with brown stem, and black laterotergite fan; anal vein indistinct at middle, wing clear, less visible veins in the lower part; strongly curved and recurrent phallus; female unknown.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB75FF94FF79FA55FC20EDBA.taxon	distribution	Distribution. France: Burgundy (89) (Fig. 4).	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB75FF94FF79FA55FC20EDBA.taxon	description	Description. Male (Fig. 3 A, B). Head. Occiput dull black, black setae, postoculars curved towards front; pale postgena hairs and setae. Eyes holoptic with upper ommatidia enlarged. Frons reduced to small triangle above antennae; ocellar triangle shiny. Face and clypeus shiny brown. Antenna dull, brownish; scape as long as but thinner than rounded pedicel; postpedicel 3.5 times longer than width at base, 4 times length of scape, 1.5 times length of stylus. Proboscis pale, labrum dark and shiny, 1.5 – 2 times head height; long and slender labellum, dark and shiny; palpus dusted, dark with apical black bristly-hair. Thorax dull black with grey pruinosity. Antepronotum with row of short black setae. Proepisternum with a few short dark setae. Postpronotum with one long black seta and shorter and thinner white ones. Acrostichals irregularly biserial, absent in prescutellar depression; dorsocentrals irregularly biserial, longer than acrostichals, ending in 3 strong, long setae in prescutellar depression. Laterotergite with fan of black setae. Scutellum with 2 long apical setae. Other strong, long black setae as follows: 2 presutural supra-alars, 3 notopleurals, 1 postsutural supra-alar, 2 postalars including small one. Legs dark brown, shiny, very slightly dusted. Fore and mid coxae dull with white hairs and setae; hind coxa dull with brown setae. Fore femur with short, dorsal brown setae, less numerous, thinner and longer ventrally but shorter than femur depth; fore tibia with anterodorsal and posterodorsal rows of distinct setae longer than tibia depth, longer apically, with ventrals much shorter; tarsomeres with dorsal setae, longer apically, shorter ventrally. Mid femur with short setae mixed with 1 row of anterodorsal setae 2 times longer than femur depth, 2 rows of anteroventral and posteroventrals 3 times longer than femur depth, becoming shorter apically; mid tibia with short hairs dorsally mixed with 3 strong dark setae, 3 times longer than tibia depth and 2 rows of anteroventral and posteroventral setae as long as tibia depth; mid tarsomeres with short setae, longer apically. Hind femur with short brown setae dorsally mixed with 2 rows of anterodorsal and posterodorsal setae as long as femur depth, 2 rows of thinner anteroventral and posteroventral setae as long as femur depth and less numerous; hind tibia basally slender, stouter towards tip, dorsally with short setae mixed with 5 – 6 dark setae slightly longer than tibia depth, one row of anteroventral setae slightly shorter than tibia depth; hind tarsomere 1 somewhat swollen but thinner than tibia at tip, ventrally with setae as long as depth of metatarsus, longer dorsally, remaining tarsomeres with rather short setae, longer apically. Wing (Length = 3.8 mm) very clear, veins less visible in basal area, inconspicuous small yellow pterostigma. Anal vein partly indistinct between middle and costa. Halter brownish although pale in Figure 3 A due to reverberation of lamp on specimen in alcohol (with dried specimen, halter clearly brownish), with darker stem; calypter pale with fringe of white and brown hairs. Abdomen shiny dark brown, 2 anterior sternites dusted. Abdominal hairs white ventrally and on sides of tergites, as long as postpedicel, white hairs 2 times shorter on tergites, mixed with shorter black setae in center of tergites, longer on tergite 1. Hypopygium shiny dark brown (Fig. 3 B); cercus with dorsal fan of short brownish hairs, slightly longer than epandrial lamella; epandrial lamella triangular and rounded posteriorly, as high as wide, with a few brown setae on posterior margin, hiding most of cercus, linked by broad dorsal sclerite, each lamella with long dorsal projection surmounted by dense fringe of dark setae; hypandrium forming broad ventral plate with narrow apex inserted inside widening of phallus; phallus dark brown, bumpy at exit of hypandrium, then thinner, paler and strongly bent forwards over cercus. Female. Unknown.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
03EC5B01BB75FF94FF79FA55FC20EDBA.taxon	discussion	Remarks. The subgenus Coptophlebia is very close to the subgenus Empis. The species were for a long time differentiated by the abbreviation of vein M 1 (Collin 1961; Chvála 1994). Daugeron et al. (2011) considered the abbreviation of vein M 1 should no longer be used to define taxonomic units within Empidinae. Empis tissoti sp. nov. is part of those Empis (Coptophlebia) species which have vein M 1 complete. It doesn’t belong to the Empis (C.) hyalipennis - group (Daugeron et al. 2011), because of the presence of acrostichals. Its strongly curved phallus is close to that of Empis (C.) abbreviata Loew, 1869 (Engel 1956, fig. 213) whose vein M 1 can be “ complete on both wings and reaching the wing-margin as in Empis s. str. ” (Chvála 1994). But E. abbreviata has the epandrial lamella without long dorsal projection. Empis (C.) hyalea - group (Daugeron & Grootaert 2005) is distributed almost everywhere in the world but only reaches the Palaearctic in Japan. Like E. tissoti sp. nov., species of this group have the epandrial lamellae linked by a broad dorsal sclerite, some species have also the epandrial lamellae with a long dorsal projection. Another autapomorphic character of this group is the presence of an unsclerotized zone in the middle of labella, E. tissoti sp. nov. doesn’t have such character. Its epandrium with such a long dorsal projection is presently unique among European Empis s. str. or Coptophlebia species.	en	Langlois, Dominique (2025): Three new Empis species of the subgenus Empis Linnaeus, 1758 and Coptophlebia Bezzi, 1909 (Diptera, Empididae, Empidinae) from France. Zootaxa 5737 (3): 414-426, DOI: 10.11646/zootaxa.5737.3.7, URL: https://doi.org/10.11646/zootaxa.5737.3.7
