taxonID	type	description	language	source
03EC87C3FFF7FFCBFDFCFDE1FE8CFB1B.taxon	type_taxon	Type species: Panurgus lustrans Cockerell, 1897 a, by original designation.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFF7FFCBFDFCFDE1FE8CFB1B.taxon	diagnosis	Subgeneric diagnosis Lasioglossum (Hemihalictus) in western North America can only be diagnosed by a combination of characters. Lasioglossum (Hemihalictus) belongs to the weak-veined Lasioglossum, also known as the Hemihalictus series. In this series, vein 1 rs-m is notably weaker compared to the more proximal wing veins, resembling the weakened 2 rs-m and 2 m-cu, or 1 rs-m is absent entirely. Both sexes have the body entirely black, without metallic reflections (brightly metallic in Antillean species formerly considered in subgenus Habralictellus; dull metallic green in some Asian species formerly in Sudila). The propodeum of female L. (Hemihalictus) is characterized by either a weak lateral carina that incompletely separates the lateral and posterior propodeal surfaces, or in cases where the lateral carina seems robust, the entire mesosoma is coarsely sculptured. Males of L. (Hemihalictus) have metasomal sterna with moderately elongate pubescence (1 – 2.5 OD), retrorse lobe present (small or absent in some extralimital species), male antennae usually relatively short, female-like (except long in the L. nitidiusculum species group), and male heads usually unmodified, with rounded clypeus, but some males are macrocephalic, with a flat clypeus, and long mandibles. Bees in the Lasioglossum series can be differentiated by the presence of a strong vein 1 rs-m (less evident in males). Male L. (Lasioglossum) are usually larger, typically with a flat clypeus, and the metasoma is usually consistently densely punctate to the apex. Lasioglossum (Sphecodogastra) Ashmead, 1899 females are identifiable by their carinate propodeum, coupled with a more finely sculptured mesosoma. In cases where the carina is either absent or reduced, which is common in Onagraceae-specialist L. (Sphecodogastra), females exhibit a modified metafemoral scopa, forming a distinctive rake-like row of setae. Lasioglossum (Sphecodogastra) males have very sparse, short pubescence (<1 OD) on the sterna, with the exception of the Onagraceae specialists. These specialists can be identified by a raised longitudinal area on S 6 and enlarged ocelli. Lasioglossum (Dialictus) and L. (Eickwortia s. lat.) are distinctly metallic, at least on the head and mesosoma. Lasioglossum (Eickwortia s. str.) occur at high elevations from Mexico to Costa Rica and are black but have a strongly bidentate mandible in the female. Males of Lasioglossum (Eickwortia s. str.) have modified hind legs: these are long and slender in L. nycteris (Vachal, 1904) and L. hienae Gibbs & Dumesh, 2013, whereas L. alexanderi (McGinley, 1999) has a broadened metabasitarsus (Hinojosa-Díaz et al. 2025) .. Lasioglossum (Evylaeus) females can be distinguished by combination of the wide head, inner metatibial spur denticulate-serrate, teeth not exceeding width of rachis, mesepisternum rugulose to rugose; propodeum lateral carina not extending to dorsal surface (except in L. cinctipes (Provancher, 1888 )); and T 1 without appressed hairs. Lasioglossum (Evylaeus) male genitalia are distinguished from those of L. (Hemihalictus) males by the absence of the retrorse lobe.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFF6FFCCFE19FAF4FC3DFD84.taxon	diagnosis	Diagnosis Females in the L. arizonense group are diagnosable using a combination of the following characters: typically orange metasoma (except in the dark morph of L. vanduzeei; light morphs of L. aspilurus (Cockerell, 1925) and L. ovaliceps (Cockerell, 1898) have orange metasoma); body covered by conspicuous tomentum on the head and mesosoma (no conspicuous tomentum on the body in most L. (Hemihalictus), but L. swenki (Crawford, 1906 b), L. subobscurum and L. robertsoni might have some tomentum, which is not as conspicuous as in these bees); small bees, length 5.50 – 6.50 mm (L. aspilurus, L. ovaliceps and L. swenki are large bees with orange abdomens, 6.5 – 8 mm); head mostly wide, length / width ratio 0.83 – 0.92 (longer in L. ovaliceps, L. swenki, length / width> 1); polished mesoscutum (duller sculpturing in L. swenki, L. subobscurum); and polished mesepisternum (rough in L. nitidiusculum group, L. subobscurum). Males in the group share similar traits. They can be diagnosed using a combination of the following characters; small size, length 4 – 5 mm (larger species, such as L. pathiranae sp. nov., L. lustrans, L. pectinatum, L. sopinci (Crawford, 1932), L. fedorense (Crawford, 1906 a), and L. swenki, range from 5 to 8.5 mm); elongated mandibles, nearly reaching opposing mandibular base (mandible shorter in L. nitidiusculum group, L. glabriventre (Crawford, 1907), L. ovaliceps, L. aspilurus, L. buccale, L. angusticulum sp. nov., L. pectoraloides, L. sequoiae (Michener, 1936), L. subobscurum, L. villosulum, L. engleri sp. nov., and L. macoupinense (Robertson, 1895 )); wide head, length / width ratio 0.84 – 0.88 (longer in L. ovaliceps, and L. pathiranae sp. nov., length / width> 1); abundant tomentum on the head and mesosoma; usually orange metasoma (though L. ovaliceps, L. aspilurus, and L. swenki also have orange metasomas, while L. subobscurum has a brown metasoma with pale marginal areas); polished mesoscutum (duller sculpturing in L. swenki and L. subobscurum); and polished mesepisternum (rough in L. nitidiusculum group and L. subobscurum).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFF1FFD6FE71FD67FDBDFBD1.taxon	description	Figs 6 – 8, 9 B, 10 B, 11 – 12, 16, 95 B, 96 B, 115 A, 120 A	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFF1FFD6FE71FD67FDBDFBD1.taxon	diagnosis	Diagnosis Female L. absurdiceps has the frons rugulose-punctate to polished, with dense punctures (IS <1 PD). In contrast, L. tomentosum sp. nov. also exhibits a somewhat rugulose-punctate frons from certain angles, but it is doubly punctate with small punctures throughout. It is further distinguished by its extremely dense, pale-yellow tomentum, which often obscures the underlying sculpturing and puncture patterns unless removed. The frons of L. vanduzeei and L. arizonense are polished with distinct double punctation, featuring minute, shallow punctures throughout. These ‘ minute’ punctures are smaller than the small punctures found on the frons of L. tomentosum. These punctures mark the points where hairs arise; however, without removing the hairs, these characters may be difficult to observe clearly. The frons punctures of L. arizonense are sparser (IS = 4 – 6 PD) than L. vanduzeei (IS = 2 – 4 PD) (check Fig. 9). The mesoscutal sculpture of L. vanduzeei and L. absurdiceps is comparatively less polished, with imbricate or tessellate sculpturing anteromedially. In contrast, L. arizonense and L. tomentosum sp. nov. exhibit a highly polished mesoscutum. Lasioglossum arizonense has comparatively very shallow and sparser (IS = 2 – 7 PD) punctures, with a distinctly doubly punctate pattern featuring minute punctures throughout. Lasioglossum tomentosum also possesses a doubly punctate mesoscutum, but with deeper minute punctures and denser density (IS = 1 – 3 PD, laterad of parapsidal line). In contrast, both L. absurdiceps and L. vanduzeei lack double punctation on the mesoscutum, and their punctures are deeper and denser comparatively (IS = 1 – 2 PD) laterad of parapsidal line (check Fig. 10). Females of Lasioglossum arizonense have a smooth and polished mesepisternum, which is doubly punctate with minute punctures throughout. Lasioglossum vanduzeei also has sparse punctation (IS = 1 – 3 PD) on the mesepisternum, which becomes progressively sparser ventrally, and it is doubly punctate, though not as conspicuously as in L. arizonense. The punctures are sparse (IS = 1 – 3 PD). In contrast, the other two species (L. absurdiceps and L. tomentosum sp. nov.) lack a smooth mesepisternum, as they exhibit an indistinctly reticulate-punctate surface structure with large, dense punctures (IS <1 PD) dorsally, though the surface remains polished. Unlike L. absurdiceps and L. tomentosum sp. nov., where T 3 is mostly fully covered by tomentum, females of L. vanduzeei display basal tomentum on T 3, and the overall body tomentum is not dense. Due to wear of tomentum, it is difficult to comment on T 3 tomentum in L. arizonense. Females of Lasioglossum vanduzeei have the protibial malus extending beyond the velum, nearly ¼ of length of velum, but the mali of other three species do not reach beyond the velum. Another reliable diagnostic character is the shape of the malus on the female’s protibial spur. In L. vanduzeei, the malus is longer, measuring nearly ⅓ the length of the velum (Fig. 17 F). In contrast, the other three species do not have a malus that extends significantly beyond the velum (as in Figs 6 F, 13 F, 15 F). Notably, in L. absurdiceps, there is no developed malus (Fig. 6 F). Female L. aspilurus are similarly coloured but lack distinct tomentum on the mesoscutum and mesepisternum. Lasioglossum absurdiceps have pale wings with light setae, but setae are darker in L. aspilurus. Lasioglossum swenki occur further east and are much larger, with a longer face (length / width 1.01), duller sculpturing on the mesoscutum, and less dense tomentum. Lasioglossum ovaliceps have a much longer face, length / width 1.10 (± 0.02 SD) and lack distinct tomentum. Only L. absurdiceps and L. vanduzeei males are known. Males of L. vanduzeei have no dense tomentum and have no distinct punctures on the mesoscutum as males of L. absurdiceps. Males of L. absurdiceps have orange metasoma and in some macrocephalic males (including the holotype), the gena is tuberculate at midlength with a short process. Males of L. vanduzeei have brown to dark brown metasoma. Mesoscutum punctures of L. vanduzeei are indistinct and sparser (IS = 2 – 7 PD), but L. absurdiceps have distinct punctures which are moderately sparse (IS = 1 – 2 PD), becoming more sparsely punctate (IS = 1 – 4 PD) submedially. Males of L. swenki and L. subobscurum can resemble the males of L. arizonense and L. vanduzeei but are distinguished by their duller mesoscutum, which results from microsculpturing. Males of Lasioglossum subobscurum have shorter mandibles, nearly reaching the opposing clypeal angle, whereas males of L. arizonense and L. vanduzeei possess elongated mandibles that nearly reach the opposing mandibular base. Additionally, males of L. swenki are larger (length 5 – 7 mm), while L. arizonense and L. vanduzeei are smaller (length 4 – 5 mm).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFF1FFD6FE71FD67FDBDFBD1.taxon	etymology	Etymology Timberlake (1960) named this species for its ‘ cephalic pecularities’.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFF1FFD6FE71FD67FDBDFBD1.taxon	materials_examined	Type material Holotype of Halictus (Evylaeus) absurdiceps (by original designation; examined) UNITED STATES – California • ♂; Merced Co., 8 mi. SW of Merced; [37.22 ° N, 120.59 ° W]; 5 Sep. 1956; R. R. Snelling leg.; CAS (Figs 7, 11). Specimen labels “ 8 mi. SW / Merced, Cal. / Sept. 5, 1956 // Snelling / Coll. // On Flowers of / Solidago // Holotype / H. absurdiceps ”. Other material examined UNITED STATES – California • 1 ♀; Fresno Co.; [36.76 ° N, 119.65 ° W]; 23 May 1949; E. G. Linsley, J. W. MacSwain and R. F. Smith leg.; EMEC • 2 ♀♀; Inyo Co., 2 mi. E of Big Pine; [37.164 ° N, 118.253 ° W]; 7 Jun. 1975; J. D. Pinto leg.; UCRC • 1 ♂; Inyo Co., 3 mi. N of Olancha; [36.324 ° N, 118.006 ° W]; 11 Sep. 1962; J. T. Doyen leg.; UCRC • 2 ♂♂; Inyo Co., 5.5 mi. S of Shoshone; [35.892 ° N, 116.272 ° W]; 29 Aug. 1979; J. C. Hall and N. Evenhuis leg.; UCRC (Fig. 8) • 7 ♀♀; Inyo Co., Keeler; [36.483 ° N, 117.873 ° W]; 28 Aug. 1977; E. G. Linsley and J. M. Linsley leg.; ex Tamarix aralensis; EMEC • 1 ♀; Inyo Co., 4 mi. E of Independence; [36.802 ° N, 118.128 ° W]; 30 May 1977; E. G. Linsley and J. M. Linsley leg.; ex Tamarix ramosissima; EMEC • 1 ♀; Inyo Co., 5 mi. N of Ballarat; [36.119 ° N, 117.227 ° W]; 20 Sep. 1965; LACM • 1 ♀; Inyo Co., Olancha; [36.282 ° N, 118.007 ° W]; 23 Aug. 1952; LACM • 1 ♀; Inyo Co., 12 mi. S of Olancha; [36.107 ° N, 118.006 ° W]; 8 Jul. 1982; J. C. Hall and E. M. Hall leg.; USNM • 1 ♂; Inyo Co., 3.5 mi. W of Lone Pine; [36.604 ° N, 118.126 ° W]; 8 Jul. 1961; H. V. Daly leg.; UCRC • 12 ♀♀; Inyo Co., Saratoga Springs, Death Valley National Park; [35.682 ° N, 116.424 ° W]; 17 Apr. 1974; M. Wasbauer and F. Andrews leg.; ex Tamarix gallica; CSCA (Fig. 9 B) • 1 ♂; Inyo Co., 3 mi. N of Olancha; [36.324 ° N, 118.006 ° W]; 11 Sep. 1962; J. T. Doyen leg.; EMEC • ♂; Inyo Co., Furnace Creek Death Valley; [36.425 ° N, 116.886 ° W]; 1 Apr. 1951; J. W. MacSwain leg.; ex Heliotropium; EMEC • 3 ♀♀; Inyo Co., Stovepipe Wells Death Valley; [36.606 ° N, 117.147 ° W]; 30 Mar. 1953; J. W. MacSwain leg.; ex Larrea; EMEC (Fig. 6) • 1 ♀; Merced Co., Los Banos; [37.059 ° N, 120.851 ° W]; 23 May 1918; E. P. Van Duzee leg.; CAS • 1 ♀; San Bernardino Co., 1 km S of Zzyzx, Soda Dry Lake; 35.133333 ° N, 116.1 ° W; elev. 290 m; 29 May 1999; D. Yanega leg.; UCRC • 2 ♀♀; San Bernardino Co., 9 air mi. S of Baker, Zzyzx Springs; [35.136 ° N, 116.1036 ° W]; 20 Apr. 1977; ex Larrea; EMEC. – Nevada • 1 ♂; Pershing Co., Black Rock Desert T 33 N R 26 E; [40.912 ° N, 119.056 ° W]; elev. 1219 m; 10 Sep. 1972; Wm. H. and M. H. Clark leg.; UCRC.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFF1FFD6FE71FD67FDBDFBD1.taxon	description	Redescription Female BODY MEASUREMENTS. Length 6.10 mm (± 0.20 SD). Head length 1.47 mm (± 0.02 SD). Face length 1.23 mm (± 0.02 SD). Head width 1.71 mm (± 0.02 SD). Wing length 4.31 mm (± 0.13 SD). n = 5. COLOURATION. Head and mesosoma dark brown to black. Clypeus apex dark brown to brown. Labrum brown. Mandible orange basally, reddish brown apically. Flagellum orange. Pronotal lobe orange. Tegula translucent pale amber. Wing membrane pale, veins with subcosta amber, otherwise pale amber. Legs orange, but coxae, trochanters, femora mostly brown. Metasoma orange, occasionally featuring irregular dark spots. PUBESCENCE. Body hair colour white. Tomentum on entire body except clypeus, postgena, pronotum anterior face, propodeum dorsal face and lateral face medially, T 1 (except laterally), T 2 medially and on rim and sterna; dense on supraclypeal area, paraocular area, frons, vertex, gena, pronotal angle, lobe and collar, space between pronotal lobe and tegula, mesoscutum marginally, metanotum, preëpisternum, metepisternum; moderately dense on mesoscutum (except marginal area), mesoscutellum, T 1 laterally, T 2 basolaterally and T 3 – T 4; sparse on propodeum lateral face near marginally and posterior face. Other than the tomentum, mesosoma covered by erect, weakly plumose, short hairs (<1 OD) on mesoscutum and erect, plumose, long hairs (<3 OD) on other areas. Metasomal sterna with dense, erect, plumose, long hairs (<5 OD). Metafemoral scopa with dense plumose hairs. Wing hairs light, very short and sparse. SURFACE SCULPTURING. Clypeus polished throughout; punctures large, irregularly spaced (IS <3 PD) on apical half, becoming small and moderately dense (IS = 1 – 2 PD) basally. Supraclypeal area polished, weakly imbricate medially; punctures dense (IS <1 PD) medially, becoming moderately sparse (IS = 1 – 2 PD) apically and laterally. Frons ruguloso-punctate, becoming polished dorsally; punctures dense (IS <1 PD). Ocellocular area polished; punctures sparse (IS = 1 – 4 PD), doubly punctate with small punctures. Gena polished, becoming weakly rugulose posteriorly; punctures moderately dense (IS = 1 – 2 PD). Postgena polished, but merely imbricate adjacent to hypostomal carina. Tegula impunctate. Mesoscutum polished, becoming imbricate anteromedially and merely imbricate mesad of parapsidal line anteriorly; punctures moderately dense (IS = 1 – 2 PD), becoming more sparsely punctate on mesad of parapsidal line (IS = 1 – 4 PD), diversopunctate, but not that conspicuous. Mesoscutellum polished, punctures moderately dense (IS = 1 – 2 PD) on median line and near lateral and posterior margins, becoming sparse (IS = 1 – 5 PD) submedially, diversopunctate. Metanotum imbricate to rugulose, punctures obscure. Propodeal triangle with rugae moderately strong, nearly reaching posterior margin, strongly anastomosing; interspace sculpture shiny, but weakly imbricate among rugae and laterally, becoming narrowly polished posteriorly. Preëpisternum reticulate-punctate. Hypoepimeron interspace sculpture shiny; punctures sparse (IS = 1 – 3 PD). Mesepisternum polished, but indistinctly reticulate-punctate; punctures dense (IS <1 PD). Propodeum lateral face imbricate, becoming shiny anteriorly; large punctures sparsely (IS = 3 – 6 PD) scattered on imbricate area and minute punctures sparse (IS = 1 – 3 PD) on shiny edge; posterior face weakly imbricate, punctures sparsely (IS = 2 – 6 PD) scattered. T 1 anterior face polished, punctures present laterally, sparse (IS = 3 – 5 PD); dorsal surface polished, with punctures sparse (IS = 1 – 3 PD), becoming more sparsely punctate (IS = 2 – 7 PD) in small subapicolateral boss and punctures fine, minute and sparse (IS = 1 – 3 PD) on rim, becoming impunctate on apical half and medially on rim. T 2 disc polished, punctures moderately dense (IS = 1 – 2 PD) basally, becoming sparse (IS = 1 – 3 PD) apically; rim weakly to distinctly coriarious, punctures minute and sparse (IS = 1 – 3 PD), becoming narrowly impunctate apically. STRUCTURE. Head wide, length / width ratio 0.85 (± 0.02 SD). Face length / width ratio 0.72 (± 0.01 SD). Clypeus projecting ~ 53 % below suborbital tangent; clypeus length / width ratio 0.97 (± 0.09 SD). Mandible short, nearly reaching beyond opposing clypeal angle. Eyes convergent below [UOD / LOD ratio 1.13 (± 0.03 SD)]. Ocelli normal. Gena / eye width ratio 0.75 (± 0.05 SD). Pronotal angle obtuse. Intertegular span 1.12 mm (± 0.04 SD). Mesoscutum length / width ratio 1.22 (± 0.02 SD); mesoscutum / scutellum length ratio 2.81 (± 0.13 SD); scutellum / metanotum length ratio 1.48 (± 0.06 SD); metanotum / propodeal triangle length ratio 0.87 (± 0.06 SD). Protibial spur with apical serrations longer in length to width of malus; malus not reaching beyond velum. Inner metatibial spur pectinate, teeth 3, basal teeth somewhat longer than width of rachis. Propodeal triangle posterior margin smoothly rounded. Propodeal lateral carina not reaching dorsal margin; oblique carina absent. VARIATION. Some specimens have an orange clypeus as well as legs that are entirely orange (3 specimens out of all). Male BODY MEASUREMENTS. Length 4.47 mm (± 0.49 SD). Head length 1.30 mm (± 0.11 SD). Face length 1.04 mm (± 0.08 SD). Head width 1.49 mm (± 0.12 SD). Wing length 3.38 mm (± 0.17 SD). n = 3. MACROCEPHALIC HOLOTYPE. Length 5 mm. Head length 1.45 mm. Face length 1.28 mm. Head width 1.72 mm. Wing length 3.9 mm. COLOURATION. Head and mesosoma dark brown to black. Clypeus apical half pale yellow. Labrum pale yellow. Mandible yellow with reddish-brown apical third. Flagellum orange. Pronotal lobe pale yellow. Tegula translucent pale amber. Wing membrane pale, veins with subcosta amber, otherwise faintly amber. Legs orange with brown coxae, trochanters, and femora. Metasoma mostly orange, occasionally with irregular dark spots. PUBESCENCE. Body hair colour white. Tomentum dense on head except vertex area, pronotal angle, lobe and collar, space between pronotal lobe and tegula, metanotum, preëpisternum, and mesepisternum; moderately dense on mesoscutum, mesoscutellum, propodeum lateral face and posterior face dorsally; sparse on T 1 laterally, T 2 basolaterally, T 3 basolaterally or nearly completely (due to wear difficult to say) and T 4. Mesosoma covered by erect, weakly plumose, short hairs (<1 OD) on mesoscutum and erect, plumose, long hairs (<3 OD) on other areas. Metasomal sterna with moderately dense, erect, moderately plumose, short hairs (0.5 – 2 OD). Wing setae pale, short and sparse. SURFACE SCULPTURE. Clypeus polished, punctures minute, moderately dense (IS = 1 – 2 PD) basally, becoming moderately sparse (IS = 1 – 2 PD) apically. Supraclypeal area polished, punctures minute, moderately dense (IS = 1 – 2 PD) to dense (IS = 1 – 2 PD). Frons polished; punctures moderately dense (IS = 1 – 2 PD). Ocellocular area polished; punctures sparse (IS = 2 – 4 PD). Gena polished; punctures moderately dense (IS = 1 – 2 PD). Postgena polished. Tegula impunctate. Mesoscutum well-polished; punctures moderately sparse (IS = 1 – 2 PD), becoming more sparsely punctate (IS = 1 – 4 PD) submedially, diversopunctate, but not that conspicuous. Mesoscutellum well-polished; punctures moderately sparse (IS = 1 – 2 PD) marginally and on median line, sparse submedially (IS = 3 – 6 PD). Metanotum polished, but weakly rugulose, punctures sparse (IS = 1 – 4 PD). Propodeal triangle with rugae weak or nearly absent, not reaching posterior margin; mostly polished, but merely imbricate laterally and basally. Preëpisternum rugulose. Hypoepimeron polished; punctures sparse (IS = 1 – 3 PD) and sometime obscure. Mesepisternum smooth and polished; punctures minute, sparse (IS = 1 – 5 PD). Propodeum lateral face imbricate, becoming polished anteriorly; posterior face polished to weakly imbricate. T 1 anterior face polished, few punctures present laterally, sparse (IS = 1 – 4 PD); dorsal surface polished, with punctures sparse (IS = 1 – 4 PD), becoming more sparsely punctate (IS = 2 – 5 PD) in small subapicolateral boss and towards premarginal line medially, impunctate on rim. T 2 disc polished; punctures sparse (IS = 1 – 4 PD); rim weakly coriarious, impunctate. STRUCTURE. Head wide, length / width ratio 0.88 (± 0.02 SD). Face length / width ratio 0.71 (± 0.01 SD). Clypeus projecting ~ 58 % below suborbital tangent. Mandible elongate and long, extending to opposing suborbital tangent, or go beyond, and sometime reaching opposing mandibular base. Flagellomeres short, F 1: pedicel length ratio 0.68 (± 0.18 SD); F 2: F 1 length ratio 2.03 (± 0.32 SD); F 2 length / width ratio 1.01 (± 0.06 SD). Gena / eye width ratio 0.77 (± 0.11 SD), sometime tuberculate at midlength. Pronotal angle obtuse. Intertegular span 0.93 mm (± 0.03 SD). Mesoscutum length / width ratio 1.23 (± 0.11 SD). Propodeal triangle posterior margin smoothly rounded. Propodeal lateral carina reaching dorsolateral slope; oblique carina absent. GENITALIA (Figs 11, 120 A). Gonostylus short, slightly longer than broad and bears short setae. Retrorse lobe somewhat attenuated apically, about 3 times as long as broad, covered in short setae. VARIATION. The male holotype exhibits an enlarged head with a prominent, elongated tubercular process on the gena, and has completely orange legs, making it a more unusual form compared to most other males. Additionally, the clypeal punctures are sparse (IS = 1 – 4 PD) basally, becoming sparser (IS = 3 – 5 PD) apically; the frons punctures moderately sparse (IS = 1 – 2 PD). In some specimens, the hypoepimeron has sparse (IS = 1 – 4 PD) punctures. STRUCTURE OF MACROCEPHALIC HOLOTYPE. Head wide, length / width ratio 0.84. Face length / width ratio 0.75. Clypeus projecting same as above. Mandible elongate and long, extending beyond opposing suborbital tangent. Flagellomeres short, F 1: pedicel length ratio 0.94; F 2: F 1 length ratio 1.60; F 2 length / width ratio 0.96. Gena width (considered up to end of tubercular process) / eye width ratio 1.21, elongated tubercular process on the gena (0.74 mm long from the ocular margin). Pronotal angle obtuse. Intertegular span 1.04 mm. Mesoscutum length / width ratio 1.24. Propodeal triangle posterior margin smoothly rounded. Propodeum lateral carinae reaching dorsolateral slope; oblique carina absent.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFF1FFD6FE71FD67FDBDFBD1.taxon	distribution	Range Central and northern California to northwestern Nevada (Figs 12, 16). USA (California, Nevada).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFF1FFD6FE71FD67FDBDFBD1.taxon	biology_ecology	Floral hosts ASTERACEAE: Solidago sp. • BORAGINACEAE: Heliotropium sp. • TAMARICACEAE: Tamarix aralensis Bunge, Tamarix gallica L., Tamarix ramosissima Ledeb. • EUPHORBIACEAE: Euphorbia sp. • ZYGOPHYLLACEAE: Larrea sp.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFEBFFDAFE6EFBBCFDEFFD4F.taxon	description	Figs 9 D, 10 D, 13 – 14, 16	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFEBFFDAFE6EFBBCFDEFFD4F.taxon	diagnosis	Diagnosis Females of L. arizonense can usually be distinguished from other species in the L. arizonense group by the following characteristics: orange clypeal apex; polished frons with distinct double punctation (large and minute punctures) and sparser puncture density (IS = 4 – 6 PD); highly polished mesoscutum with comparatively very shallow and sparser punctures (IS = 2 – 7 PD), exhibiting a distinctly doubly punctate pattern; and smooth and polished mesepisternum with sparse, conspicuous double punctation (IS = 1 – 3 PD). In some instances, a few females of L. absurdiceps have an orange clypeal apex. For additional minute characters, please refer to the diagnosis of L. absurdiceps, which diagnoses females of L. arizonense. Males of L. arizonense are unknown.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFEBFFDAFE6EFBBCFDEFFD4F.taxon	etymology	Etymology Crawford (1907 a) did not explain the epithet. The specific epithet is a presumably reference to the type locality ‘ Arizona’ with the adjectival suffix ‘ - ensis ’.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFEBFFDAFE6EFBBCFDEFFD4F.taxon	materials_examined	Type material Holotype of Halictus arizonensis (by monotypy; examined) UNITED STATES – Arizona • ♀; C. F. Baker leg.; USNM Type No. 10930 (Fig. 14). Specimen labels “ Ariz / 2341 // Collection / CF Baker // Type / No 10930 / U. S. M. M. // USNMENT / 00535045 ”. Other material examined UNITED STATES – Arizona • 1 ♀; Yuma Co., Yuma; [32.69 ° N, 114.63 ° W]; 1 – 31 May 1937; UCRC (Figs 9 D, 10 D, 13 A – B). – California • 1 ♀; Imperial Co., Winterhaven; [32.74 ° N, 114.64 ° W]; 29 Mar. 1954; T. Gallion leg.; CSCA • 1 ♀; Riverside Co., 4 mi. SW of Ripley; [33.484 ° N, 114.706 ° W]; 30 Apr. 1952; P. H. Timberlake leg.; ex Heliotropium [illegible]; UCRC (Fig. 13 C – D) • 1 ♀; Riverside Co., 6 mi. W of Indio; [33.718 ° N, 116.337 ° W]; 30 Apr. 1949; E. G. Linsley, J. W. MacSwain and R. F. Smith leg.; ex Melilotus; UCRC.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFEBFFDAFE6EFBBCFDEFFD4F.taxon	description	Description Female BODY MEASUREMENTS. Length 5.74 mm (± 0.19 SD). Head length 1.34 mm (± 0.04 SD). Face length 1.16 mm (± 0.02 SD). Head width 1.53 mm (± 0.06 SD). Wing length 3.99 mm (± 0.25 SD). n = 4. COLOURATION. Head and mesosoma brown. Clypeus orange, except basally. Labrum orange. Mandible brown to orange basally, reddish brown in apically. Flagellum orange. Pronotal lobe orange. Tegula translucent pale amber. Wing membrane pale, veins with subcosta amber, otherwise pale amber. Legs orange. Metasoma orange, sometimes with irregular dark spots. PUBESCENCE. Body hair colour off-white. Tomentum dense on paraocular area, gena, pronotal angle, lobe and collar, space between pronotal lobe and tegula, mesoscutum marginally and metanotum anteromedially, preëpisternum, metepisternum, and propodeum lateral face; moderately dense on vertex (behind the lateral ocelli), mesepisternum, and propodeum posterior face; moderately dense or sparse (possibly due to wear) on supraclypeal area, frons, mesoscutum medially, T 2 basolaterally and T 3 – T 4. Other than the tomentum, mesosoma with erect, weakly plumose, short hairs (<1 OD) on mesoscutum and erect, plumose, long hairs (<3 OD) on other areas. Metasomal sterna with moderately dense, erect, plumose, long hairs (<4 OD). Metafemoral scopa with dense plumose hairs. Wing setae light brown, very short and sparse. SURFACE SCULPTURING. Clypeus polished throughout; large punctures present, irregularly spaced (IS <3 PD) apically, nearly impunctate apicolaterally, and becoming smaller and denser basally (IS = 1 – 3 PD), diversopunctate. Supraclypeal area polished; punctures sparse (IS = 1 – 3 PD), diversopunctate. Frons highly polished; punctures sparse (IS = 4 – 6 PD), distinctly doubly punctate with minute, shallow punctures throughout (at the points where hairs arise). Ocellocular area highly polished; punctures sparse (IS = 4 – 6 PD), doubly punctate with minute, shallow punctures. Gena polished, becoming weakly rugulose posteriorly; punctures moderately dense (IS = 1 – 2 PD). Postgena polished. Tegula impunctate. Mesoscutum highly polished, becoming merely imbricate anteromedially; punctures sparse (IS = 2 – 7 PD), distinctly doubly punctate with minute, shallow punctures throughout. Mesoscutellum highly polished, punctures sparse (IS = 2 – 4 PD) marginally and on median line, sparse submedially (IS = 2 – 6 PD), diversopunctate. Metanotum weakly rugulose, punctures minute and sparse (IS = 1 – 4 PD). Propodeal triangle with rugae vary; moderately strong rugae, nearly reaching posterior margin medially, anastomosing medially and weakly so laterally or weakly developed rugae, not reaching halfway to posterior margin; interspace sculpture shiny, but weakly imbricate among rugae and laterally, becoming polished posteriorly. Preëpisternum weakly reticulate-punctate, but interspaces shiny. Hypoepimeron highly polished, with large and minute punctures intermixed, sparse (IS = 2 – 4 PD). Mesepisternum smooth and highly polished; punctures sparse (IS = 3 – 5 PD), distinctly doubly punctate with minute, shallow punctures throughout. Propodeum lateral face imbricate, becoming polished anteriorly; large punctures sparsely (IS = 3 – 5 PD) scattered medially and minute punctures sparse (IS = 1 – 3 PD) on shiny edge; posterior face polished to weakly imbricate, punctures obscure. T 1 anterior face polished, punctures present laterally, sparse (IS = 1 – 4 PD); dorsal surface polished, punctures sparse (IS = 1 – 4 PD), few present on subapicolateral boss and rim punctures minute and nearly impunctate (short hairs present, but these not arising from distinct punctures). T 2 disc polished; punctures sparse (IS = 1 – 4 PD); rim polished or merely coriarious; punctures minute and nearly impunctate (short hairs present, but these not arising from distinct punctures). STRUCTURE. Head wide, length / width ratio 0.87 (± 0.01 SD). Face length / width ratio 0.76 (± 0.02 SD). Clypeus projecting ~ 52 % below suborbital tangent; clypeus length / width ratio 0.92 (± 0.10 SD). Mandible short, reaching opposing clypeal angle. Eyes convergent below [UOD / LOD ratio 1.13 (± 0.02 SD)]. Ocelli normal. Gena / eye width ratio 0.66 (± 0.05 SD). Pronotal angle obtuse. Intertegular span 1.13 mm (± 0.07 SD). Mesoscutum length / width ratio 1.13 (± 0.07 SD); mesoscutum / scutellum length ratio 2.87 (± 0.17 SD); scutellum / metanotum length ratio 1.41 (± 0.07 SD); metanotum / propodeal triangle length ratio 0.94 (± 0.03 SD). Protibial spur with apical serrations longer in length to width of malus; malus not reaching much beyond velum. Inner metatibial spur pectinate, teeth 3 – 4, basal teeth somewhat longer or sometime subequal to width of rachis. Propodeal triangle posterior margin smoothly rounded. Propodeal lateral carina not reaching dorsal margin; oblique carina absent. n = 4. Male Unknown	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFEBFFDAFE6EFBBCFDEFFD4F.taxon	distribution	Range Southern California (specifically the few examined specimens were collected from Colorado River lower basin and the Coachella Valley) to Arizona (Fig. 16). USA (Arizona, California).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFEBFFDAFE6EFBBCFDEFFD4F.taxon	biology_ecology	Floral hosts FABACEAE: Melilotus sp • BORAGINACEAE: Heliotropium sp.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFEBFFDAFE6EFBBCFDEFFD4F.taxon	discussion	Remarks See remarks under L. vanduzeei.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE7FFDDFE3FFD38FDE9FAC0.taxon	description	urn: lsid: zoobank. org: act: 6065 B 011 - 57 A 8 - 4 B 65 - B 342 - EA 0 DFBD 4 D 96 E Figs 9 C, 10 C, 15 – 16, 95 A, 96 A	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE7FFDDFE3FFD38FDE9FAC0.taxon	diagnosis	Diagnosis Females of L. tomentosum sp. nov. can be easily recognised from any closely related bees by extremely dense, yellowish tomentum, on the body. Even the posterior face of the propodeum is covered by tomentum. The dense tomentum poses challenges in observing the sculpturing and puncture pattern on the body, but it makes this species easily stand out from others. For additional minute characters, please refer to the diagnosis of L. absurdiceps. Males of this species are unknown.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE7FFDDFE3FFD38FDE9FAC0.taxon	etymology	Etymology The specific epithet ‘ tomentosum ’ is a Latin adjective, which refers to the bees’ dense layer of woolly hairs. The prominent layer of pale-yellow tomentum covering the body gives it a distinct appearance.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE7FFDDFE3FFD38FDE9FAC0.taxon	materials_examined	Type material Holotype UNITED STATES – Utah • ♀; Washington Co., St George; [37.104 ° N, 113.584 ° W]; 14 Apr. 1962; G. F. Knowlton leg.; ex Tamarix sp.; SEMC (Fig. 15). Specimen labels “ UTAH Washington Co. / St George, 14 Apr. / 1962 (G. F. Knowlton) // on flowers of / Tamarix // Lasioglossum / GIBBS – 13406 // HOLOTYPE / Lasioglossum (Hemihalictus) tomentosum Hettiarachchi and Gibbs ”. Paratypes UNITED STATES – Utah • 4 ♀♀; Washington Co., St George; [37.104 ° N, 113.584 ° W]; 14 Apr. 1962; G. F. Knowlton leg.; ex Tamarix sp.; SEMC (Fig. 10 C) • 2 ♀♀; Washington Co., Santa Clara; [37.133 ° N, 113.654 ° W]; 15 Apr. 1962; G. F. Knowlton leg.; SEMC (Fig. 9 C) • 3 ♀♀; Washington Co., Washington; [37.131 ° N, 113.508 ° W]; 15 Apr. 1962; G. F. Knowlton leg.; ex Tamarix; SEMC.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE7FFDDFE3FFD38FDE9FAC0.taxon	description	Description Female BODY MEASUREMENTS. Length 5.24 mm (± 0.45 SD). Head length 1.34 mm (± 0.03 SD). Face length 1.15 mm (± 0.03 SD). Head width 1.48 mm (± 0.03 SD). Wing length 3.79 mm (± 0.11 SD). n = 5. COLOURATION. Head and mesosoma brown. Clypeus orange, except brown basal half. Labrum orange. Mandible orange basally, reddish brown apically. Flagellum mostly orange, sometime brown dorsally. Pronotal lobe orange. Tegula translucent pale amber. Wing membrane hyaline, veins with subcosta light brown, otherwise pale amber. Legs orange. Metasoma orange, sometimes with irregular dark spots. PUBESCENCE. Body hair colour pale yellow. Tomentum dense on entire body except clypeus, postgena, pronotum anterior face, propodeum dorsal face, T 1, T 2 medially and on rim and sterna. Other than the tomentum, mesosoma covered by erect, weakly plumose, short hairs (<1 OD) on mesoscutum and erect, plumose, long hairs (<3 OD) on other areas. Metasomal sterna with moderately dense, erect, plumose, long hairs (<4 OD). Metafemoral scopa with dense plumose hairs. Wing hairs light, very short and sparse. SURFACE SCULPTURING. Clypeus polished throughout; few large punctures present with irregularly spaced (IS <3 PD) apically, impunctate apicolaterally, and becoming smaller and denser basally (IS = 1 – 3 PD), diversopunctate. Supraclypeal area polished; punctures moderately sparse (IS = 1 – 2 PD), diversopunctate. Frons weakly ruguloso-punctate (or more rugulose from some angles) to highly polished; punctures sparse (IS = 1 – 3 PD), doubly punctate with small punctures throughout (at the points where hairs arise). Ocellocular area highly polished; punctures sparse (IS = 2 – 5 PD), doubly punctate with minute punctures (at the points where hairs arise). Gena polished, becoming weakly rugulose posteriorly; punctures moderately dense (IS = 1 – 2 PD). Postgena polished. Mesoscutum highly polished; punctures sparse (IS = 1 – 3 PD), becoming more sparser on mesad of parapsidal line (IS = 2 – 5 PD) and denser posteriorly (IS = 1 – 2 PD), doubly punctate. Mesoscutellum highly polished, punctures moderately sparse (IS = 1 – 2 PD) posteriorly and on median line, sparser submedially (IS = 2 – 6 PD), diversopunctate. Metanotum weakly rugulose, punctures minute and sparse (IS = 1 – 4 PD). Propodeal triangle with rugae weak, nearly reaching posterior margin medially; anastomosing medially and weakly so laterally; sculpture shiny, becoming weakly imbricate laterally. Preëpisternum reticulate-rugulose. Hypoepimeron weakly reticulate-rugulose, polished ventrally. Mesepisternum polished, but weakly reticulate-rugulose dorsally and becoming indistinct ventrally, minute punctures sparse (IS = 1 – 3 PD). Propodeum lateral face imbricate, becoming weakly imbricate anteriorly; large punctures sparsely (IS = 3 – 6 PD) scattered medially; posterior face imbricate to weakly imbricate, few punctures sparsely (IS = 2 – 6 PD) scattered. T 1 anterior face polished, punctures present posteriorly, sparse (IS = 1 – 4 PD), diversopunctate; dorsal surface polished, becoming weakly coriarious on rim, punctures sparse (IS = 1 – 4 PD), becoming more sparsely punctate (IS = 4 – 7 PD) in small subapicolateral boss and punctures sparse (IS = 1 – 3 PD) on rim, becoming impunctate on apical half of rim. T 2 disc polished; punctures sparse (IS = 1 – 3 PD); rim coriarious; punctures minute and sparse (IS = 1 – 3 PD), becoming impunctate towards edge. STRUCTURE. Head wide to round, length / width ratio 0.91 (± 0.01 SD). Face length / width ratio 0.77 (± 0.01 SD). Clypeus projecting ~ 53 % below suborbital tangent; clypeus length / width ratio 0.97 (± 0.09 SD). Mandible short, reaching opposing clypeal angle. Eyes convergent below [UOD / LOD ratio 1.13 (± 0.03 SD)]. Ocelli normal. Gena / eye width ratio 0.75 (± 0.05 SD). Pronotal angle obtuse. Intertegular span 1.12 mm (± 0.04 SD). Mesoscutum length / width ratio 1.22 (± 0.02 SD); mesoscutum / scutellum length ratio 2.81 (± 0.13 SD); scutellum / metanotum length ratio 1.48 (± 0.06 SD); metanotum / propodeal triangle length ratio 0.87 (± 0.06 SD). Protibial spur with apical serrations longer in length to width of malus; malus not reaching much beyond velum. Inner metatibial spur pectinate, teeth 3, basal teeth shorter or subequal to width of rachis. Propodeal triangle posterior margin smoothly rounded. Propodeal lateral carina not reaching dorsal margin; oblique carina absent. n = 5. Male Unknown.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE7FFDDFE3FFD38FDE9FAC0.taxon	distribution	Range Washington County, Utah (Fig. 16). USA (Utah).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE7FFDDFE3FFD38FDE9FAC0.taxon	biology_ecology	Floral host TAMARICACEAE: Tamarix sp.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE0FFE5FEE3FAA3FE06FEA4.taxon	description	Figs 9 A, 10 A, 17 – 21, 96 C, 100 C, 107 A, 120 B	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE0FFE5FEE3FAA3FE06FEA4.taxon	diagnosis	Diagnosis Females of L. vanduzeei can usually be distinguished from other species in the L. arizonense group by the following characteristics: polished frons with distinct double punctation consisting of large and minute punctures, punctures relatively sparse (IS = 2 – 4 PD) (not sparser as in L. arizonense); mesoscutum less polished (well-polished in L. arizonense and L. tomentosum sp. nov.) with deeper and denser punctures (shallow in L. arizonense); T 3 with a basal tomentum (T 3 fully covered by tomentum in L. absurdiceps and L. tomentosum); and the malus of the protibial spur is longer, measuring nearly ¼ the length of the velum (unlike in other species in the L. arizonense group, which lack an extended malus). For additional minute characters, please refer to the diagnosis of L. arizonense, which also diagnoses females of L. vanduzeei. Females of L. vanduzeei occur in two morphs: a light morph, which has a mostly orange metasoma, and a dark morph, which has a mostly brown metasoma. The dark morph can be easily confused with L. subobscurum, as the latter species also exhibits some tomentum on the body. However, L. subobscurum can be distinguished by its strongly imbricate frons with dense punctures (IS <1 PD), imbricate-tessellate mesoscutum with dense punctures (IS ≤ 1 PD), and a duller mesepisternum that lacks punctures. For the diagnosis of males of L. vanduzeei, refer to L. absurdiceps.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE0FFE5FEE3FAA3FE06FEA4.taxon	etymology	Etymology Sandhouse & Cockerell (1924) did not state the derivation of the epithet, but the specific epithet is presumably named for Mr Edward P. Van Duzee (1861 – 1940), former curator of entomology at the California Academy of Sciences, who collected the type series.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE0FFE5FEE3FAA3FE06FEA4.taxon	materials_examined	Type material Holotype of Halictus vanduzeei (by original designation; examined by JG ca 2013 and re-examined from photographs) MEXICO – Baja California Sur • ♀; La Paz; [24.14 ° N, 110.31 ° W]; 5 Jun. 1921; E. P. Van Duzee leg.; CAS 1545. Specimen labels “ La Paz / Baja Calf. / June 5 1921 // EPVanDuzee / Collector // Halictus / vanduzeei / Cockerell + Sandhouse / Type // California Academy of Sciences / Type No. 1545 ”. Paratypes of Halictus vanduzeei (by original designation; examined) MEXICO – Baja California • 1 ♀; Las Animas Bay; [28.39 ° N, 113.18 ° W]; 8 May 1921; E. P. Van Duzee leg.; USNM 39979 • 1 ♀; Angeles Bay; [28.91 ° N, 113.56 ° W]; 26 Jun. 1921; E. P. Van Duzee leg.; USNM 39979. Specimen labels “ Las Animas / Bay Gulf Cal. / May 8 1921 // EPVan Duzee / Collector // ParatypeNo. / 39979. / U. S. N. M. // Halictus / vanduzeei / Sandhouse & Ckll. / Paratype. // EVYLAEUS / ARIZONENSIS / (CRAWFORD) ”. “ Angeles Bay / Gulf Calif. / Jun. 26 1921 // EPVan Duzee / Collector // ParatypeNo. / 39979. / U. S. N. M. // Halictus / vanduzeei / Sandhouse & Ckll. / Paratype. ”. Holotype of Halictus chionocephalus (by original designation; examined by JG ca 2013 and re-examined from photographs) MEXICO – Baja California Sur • ♂; Loreto; [26.01, 111.34 ° W]; 20 May 1921; E. P. Van Duzee leg.; CAS 1546 • 1 ♂; Las Animas Bay; [28.39 ° N, 113.18 ° W]; 8 May 1921; E. P. Van Duzee leg.; EMEC. Specimen labels “ Loreto / Baja Cal. / May 20 1921 // EPVanDuzee / Collector // Halictus / chionocephalus / Cockerell + Sandhouse / Type // California Academy of Sciences / Type No. 1546 ”. Paratype of Halictus chionocephalus (by original designation; examined) MEXICO – Baja California • 1 ♂; Las Animas Bay; [28.39 ° N, 113.18 ° W]; 8 May 1921; E. P. Van Duzee leg.; EMEC (Figs 19 – 20). Specimen labels “ Las Animas / Bay Gulf Cal. / May 8 1921 // EPVan Duzee / Collector // Halictus / chionocephalus / Cockerell + Sandhouse / Cotype ”. Othe material examined MEXICO – Baja California Sur • 1 ♀; Bahía San Nicolás, San Sebastián; [26.617 ° N, 111.568 ° W]; 7 Mar. 1975; P. DeBach leg.; ex Malaise; WRME 0512988 • 1 ♀; same data as for preceding; 19 May 1975; P. DeBach leg.; WRME 0512989 • 1 ♀; Las Barracas; [23.475 ° N, 109.454 ° W]; 7 May 1984; P. DeBach leg.; WRME 0512991 • 1 ♀; 2 mi. S of La Paz; [24.1107 ° N, 110.3131 ° W]; 2 Aug. 1966; E. Linsley, J. Linsley, J. Chemsak and P. D. Hurd leg.; ex at light; EMEC (Fig. 17 B) • 6 ♀♀; same data as for preceding; 31 Jul. 1966; E. G. Linsley and J. M. Linsley leg.; EMEC (Figs 9 A, 10 A, 17 A, 17 C – D, 18 A) • 6 ♀♀; 9 mi. SW of La Paz; [24.048 ° N, 110.414 ° W]; 14 Aug. 1966; J. Powell leg.; EMEC (Fig. 18 B) • 1 ♀; 2 mi. S of La Paz; [24.111 ° N, 110.313 ° W]; 7 Aug. 1966; J. A. Chemsak leg.; ex Cleomella refracta (verbatim: Wislizenia refracta); EMEC • 1 ♀; 7 mi. SW of La Paz; [24.068 ° N, 110.392 ° W]; 4 Aug. 1966; J. A. Chemsak leg.; ex Cleomella refracta (verbatim: Wislizenia refracta); EMEC • 1 ♀; El Pescadero, Playa Los Cerritos; [23.330 ° N, 110.177 ° W]; 16 – 17 Apr. 1979; M. Wasbauer leg.; CSCA.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE0FFE5FEE3FAA3FE06FEA4.taxon	description	Description Female BODY MEASUREMENTS. Length 5.77 mm (± 0.56 SD). Head length 1.45 mm (± 0.07 SD). Face length 1.23 mm (± 0.06 SD). Head width 1.66 mm (± 0.08 SD). Wing length 3.86 mm (± 0.24 SD). n = 6. COLOURATION. Head and mesosoma dark brown to black. Clypeus apex dark brown to brown. Labrum brown. Mandible brown basally, light brown apically with reddish brown apex. Flagellum orange to brown. Pronotal lobe orange. Tegula translucent pale amber. Wing membrane pale, veins with subcosta light brown, otherwise amber. Legs orange with more brownish coxae, trochanters, and femora. Metasoma brown, with rims of terga and sterna broadly light brown. PUBESCENCE. Body hair colour off-white. Tomentum dense on paraocular area, gena, pronotal angle, lobe and collar, space between pronotal lobe and tegula, and metanotum anteromedially; moderately dense on supraclypeal area, frons, vertex, mesoscutum marginally, preëpisternum, metepisternum, propodeum lateral face near marginally, T 2 basolaterally, and T 3 basally; sparse on mesoscutum (except marginal area), mesepisternum, and T 4. Other than the tomentum, mesosoma covered by erect, weakly plumose, short hairs (<1 OD) on mesoscutum and erect, plumose, long hairs (<3 OD) on other areas. Metasomal sterna with moderately dense, erect, plumose, long hairs (<5 OD). Metafemoral scopa with dense plumose hairs. Wing setae light brown, very short and sparse. SURFACE SCULPTURING. Clypeus polished throughout; punctures large, irregularly spaced apically (IS <3 PD), becoming small basally and sometime minute punctures scattered basally. Supraclypeal area weakly imbricate; minute punctures scattered. Frons polished; punctures sparse (IS = 2 – 4 PD), distinctly doubly punctate with minute punctures throughout (at the points where hairs arise). Ocellocular area polished; punctures sparse (IS = 2 – 4 PD), doubly punctate with minute punctures. Gena polished, becoming weakly rugulose posteriorly; punctures moderately dense (IS = 1 – 2 PD). Postgena polished, becoming imbricate adjacent to hypostomal carina. Tegula impunctate. Mesoscutum polished, becoming imbricate to tessellate anteromedially; punctures moderately sparse (IS = 1 – 2 PD), becoming sparser on mesad of parapsidal line (IS = 1 – 5 PD), diversopunctate. Mesoscutellum highly polished, punctures moderately dense (IS = 1 – 2 PD) near lateral and posterior margins, sparse (IS = 1 – 4 PD) on median line and sparser or nearly impunctate submedially, diversopunctate. Metanotum weakly rugulose, punctures sparse (IS = 1 – 4 PD). Propodeal triangle with rugae moderately strong, not reaching posterior margin, anastomosing; interspace sculpture weakly imbricate becoming polished posteriorly. Preëpisternum rugulose. Hypoepimeron polished; punctures irregularly spaced (IS <3 PD) and sometime obscure. Mesepisternum polished; punctures sparse (IS = 1 – 3 PD) dorsally and anteriorly, doubly punctate with minute punctures (not conspicuous from some angles), becoming indistinct, smaller and more sparse ventro-posteriorly (IS = 3 – 6 PD). Propodeum lateral face weakly tessellate to imbricate, becoming weakly so towards anterior margin; few large punctures sparsely (IS = 2 – 5 PD) scattered on posteriorly and minute punctures sparse (IS = 1 – 3 PD) anteriorly; posterior face weakly imbricate, punctures sparsely (IS = 2 – 6 PD) scattered. T 1 anterior face polished, punctures present laterally, sparse (IS = 1 – 4 PD); dorsal surface polished, becoming weakly coriarious on rim, punctures sparse (IS = 1 – 3 PD), becoming more sparsely punctate (IS = 2 – 7 PD) in small subapicolateral boss and punctures sparse (IS = 1 – 3 PD) on rim, becoming impunctate on apical half of rim. T 2 disc polished, punctures moderately dense (IS = 1 – 2 PD) basally, becoming sparse (IS = 1 – 3 PD) apically; rim coriarious sometimes, punctures minute and sparse (IS = 1 – 3 PD), becoming impunctate towards edge. STRUCTURE. Head wide, length / width ratio 0.87 (± 0.01 SD). Face length / width ratio 0.74 (± 0.02 SD). Clypeus projecting ~ 54 % below suborbital tangent; clypeus length / width ratio 1.00 (± 0.11 SD). Mandible short, just extending beyond opposing clypeal angle. Eyes convergent below [UOD / LOD ratio 1.15 (± 0.06 SD)]. Ocelli normal. Gena / eye width ratio 0.74 (± 0.02 SD). Pronotal angle obtuse. Intertegular span 1.23 mm (± 0.08 SD). Mesoscutum length / width ratio 1.14 (± 0.05 SD); mesoscutum / scutellum length ratio 2.99 (± 0.23 SD); scutellum / metanotum length ratio 1.54 (± 0.15 SD); metanotum / propodeal triangle length ratio 0.91 (± 0.11 SD). Protibial spur with apical serrations longer in length to width of malus; malus extending beyond the velum, nearly ¼ of length of velum. Inner metatibial spur pectinate, teeth 4 – 5, basal teeth somewhat longer or sometime subequal to width of rachis. Propodeal triangle posterior margin smoothly rounded. Propodeal lateral carina nearly reaching dorsolateral slope; oblique carina absent. n = 6. VARIATION. The mesoscutum is occasionally tessellate anteriorly and medially, while other are regions weakly tessellate; the mesoscutellum can also be weakly tessellate to tessellate, particularly in posterior areas; in some instances, the rugae on the propodeal triangle can appear strong, and the sculpture may exhibit weak tessellation posteriorly; The rim area of T 1 also can occasionally appear coriaceous. The colour of the metasoma varies between orange and brown. Some dark morphs exhibit a reddish-brown coloration, while others have an entirely brown metasoma with light-brown rims on the terga and sterna. Although all type specimens represent the dark morph with a brown metasoma, the light morph, characterized by an orange metasoma, is noted here as a variation. Male BODY MEASUREMENTS. Length 4.9 mm. Head length 1.38 mm. Face length 1.2 mm. Head width 1.57 mm. Wing length 3.5 mm. n = 1. COLOURATION. Head and mesosoma dark brown to black. Clypeus light brown apically. Labrum light brown. Mandible orange with reddish brown apical ⅓. Flagellum orange. Pronotal lobe orange. Tegula translucent pale amber. Wing membrane pale, veins with subcosta light brown, otherwise amber. Legs light brown. Metasoma brown to black with rims of T 4 – 5 light brown. PUBESCENCE. Body hair colour white. Tomentum dense on head except vertex area, pronotal angle, lobe and collar, space between pronotal lobe and tegula, and metanotum; moderately dense on mesoscutum, scutellum, and T 2 – T 3 basolaterally. Due to the wear on the specimen, assessing the tomentum on other parts of the body is challenging. Mesosoma covered by erect, weakly plumose, short hairs (<1 OD) on mesoscutum and erect, plumose, long hairs (<3 OD) on other areas. Metasomal sterna with moderately dense, erect, moderately plumose, short hairs (<2 OD). Wing hairs light, short and sparse. SURFACE SCULPTURE. Clypeus polished, punctures, moderately sparse (IS = 1 – 2 PD) basally, becoming sparse (IS = 1 – 4 PD) apically, diversopunctate. Supraclypeal area polished, punctures dense (IS ≤ 1 PD). Frons polished; punctures minute, moderately dense (IS = 1 – 2 PD). Ocellocular area polished; punctures minute, sparse (IS = 3 – 7 PD). Gena polished, becoming ruguloso-lineate posteriorly; punctures moderately dense (IS = 1 – 2 PD). Postgena imbricate. Tegula impunctate. Mesoscutum well-polished; punctures sparse (IS = 2 – 4 PD), becoming more indistinct and sparse mesad of parapsidal line (IS = 3 – 7 PD). Mesoscutellum well-polished; punctures sparse (IS = 1 – 3 PD) marginally and on median line, nearly impunctate submedially. Metanotum weakly rugulose, punctures sparse (IS = 1 – 4 PD). Propodeal triangle with rugae absent; sculpture tessellate, becoming polished posteriorly. Preëpisternum weakly rugulose. Hypoepimeron polished; punctures sparse (IS = 1 – 4 PD), diversopunctate. Mesepisternum polished; punctures sparse (IS = 1 – 3 PD), becoming smaller and more sparsely punctate (IS = 2 – 6 PD) ventrally, diversopunctate. Propodeum lateral face imbricate, becoming ruguloso-lineate dorsoposteriorly; posterior face polished to weakly imbricate. T 1 anterior face polished, few punctures present laterally; dorsal surface polished, punctures sparse (IS = 1 – 2 PD), becoming more sparsely punctate (IS = 2 – 6 PD) in small subapicolateral boss, and punctures sparse (IS = 1 – 3 PD) on rim, becoming impunctate on apical half of rim. T 2 disc polished; punctures sparse (IS = 1 – 6 PD) basally, becoming closer medially (IS = 1 – 3 PD); rim polished, punctures sparse (IS = 1 – 3 PD) becoming impunctate apically. STRUCTURE. Head wide, length / width ratio 0.88. Face length / width ratio 0.77. Clypeus projecting ~ 54 % below suborbital tangent. Mandible long and elongate, nearly reaching opposing mandibular base. Flagellomeres short, F 1: pedicel length ratio 0.7; F 2: F 1 length ratio 1.93; F 2 length / width ratio 1.13. Gena / eye width ratio 0.67. Forewing with 3 submarginal cells. Pronotal angle obtuse. Intertegular span 0.99 mm. Mesoscutum length / width ratio 1.18. Propodeal triangle posterior margin smoothly rounded. Propodeal lateral carina reaching dorsolateral slope; oblique carina absent. n = 1. GENITALIA (Figs 20, 120 B). Gonostylus approximately twice as long as broad, with short setae. Retrorse lobe rounded apically, about 3 times as long as broad, covered in short setae.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE0FFE5FEE3FAA3FE06FEA4.taxon	distribution	Range Baja California Peninsula (Figs 16, 21).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE0FFE5FEE3FAA3FE06FEA4.taxon	biology_ecology	Floral host CAPPARACEAE: Cleomella refracta (Engelm.) J. C. Hall & Roalson • BORAGINACEAE: Heliotropium curassavicum var. oculatum (A. Heller) Thorne • FABACEAE: Melilotus sp. • TAMARICACEAE: Tamarix aralensis Bunge, Tamarix ramosissima Ledeb. • ZYGOPHYLLACEAE: Larrea tridentata (DC.) Coville.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFE0FFE5FEE3FAA3FE06FEA4.taxon	discussion	Remarks Halictus vanduzeei and H. chionocephalus were described in the same paper. Although their similarity was noted, differences in the propodeal sculpturing led the authors to keep them separate. They were later both treated as synonyms of L. arizonense in a catalogue of halictid bees (Moure & Hurd 1987), which was attributed to George Eickwort. Our study of the L. arizonense group has led to the resurrection of Lasioglossum vanduzeei as a valid species, distinct from L. arizonense, with H. chionocephalus remaining its synonym.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD8FFECFE17FE86FC15FADB.taxon	description	Figs 22 – 26, 95 E, 98 B, 104 K, 112 E, 119 B, 120 C	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD8FFECFE17FE86FC15FADB.taxon	diagnosis	Diagnosis Both sexes are readily identifiable by their short head (length / width ratio <1) and reddish orange metasoma, which is a unique combination for L. (Hemihalictus) west of the Sierra Nevada Mountains. Lasioglossum ovaliceps is similarly coloured but has a long head (length / width ratio> 1.1) (see Diagnosis for L. ovaliceps). Both sexes of L. aspilurus have been recorded with dark morphs, characterized by brown abdomens, though such individuals are exceedingly rare (Fig. 24). Lasioglossum aspilurus lack the tomentum present in the L. arizonense group. Lasioglossum aspilurus may be mistaken for Sphecodes Latreille, 1804, but the generic characters for Lasioglossum will distinguish them and the yellow maculation of the male is not present in Sphecodes.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD8FFECFE17FE86FC15FADB.taxon	etymology	Etymology The specific epithet is presumably a reference to the absence of tomentum on the metasoma. It is apparently derived from the Greek ‘ σπίλος’ for ‘ spot’ and ‘ οὐρά’ for ‘ tail’, and the prefix ‘ a- ’ indicating their absence. The ending ‘ - urus’ is ambiguous and could be a Greek noun ending (- ος) or Latin adjectival ending (- us). Following article 31.2.2 and 34.2.1 (ICZN 1999), we treat it as a noun in apposition and retain the original spelling.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD8FFECFE17FE86FC15FADB.taxon	materials_examined	Type material Holotype of Halictus aspilurus (by original designation; examined by JG ca 2013 and re-examined from photographs) UNITED STATES – California • ♀; Monterey Co., Pleyto; [35.86 ° N, 120.99 ° W]; 22 May 1920; E. P. Van Duzee leg.; CAS 1651. Specimen labels “ Pleyto Cal. / MontereyCo. / May 22 1920 // EPVan Duzee / Collector // Halictus / aspilurus / Ckll TYPE // California Academy / of Sciences / Type No. 1651 ”. Holotype of Halictus (Evylaeus) humboldtensis (by original designation; examined by JG ca 2013 and re-examined from photographs) UNITED STATES – California • ♀; Humboldt Co., Garberville; [40.10 ° N, 123.80 ° W]; 19 Apr. 1935; E. P. Van Duzee leg.; CAS 4200. Specimen labels “ Garberville Cal. / Humboldt Co. / IV- 19 - 35 // EPVan Duzee / Collector // Ceanothus // Halictus / humboldtensis / Mich. / TYPE // California Academy / of Sciences / Type No. 4200 ”. Material examined CANADA – British Columbia • 1 ♀; Okanagan Falls, Vaseux Lake Provincial Park; 49.29913 ° N, 119.53065 ° W; 7 Jul. – 16 Aug. 2021; H. Singh and T. McGratton (J. H) leg.; RSKM. UNITED STATES – California • 1 ♂; Alameda Co., Arroyo Mocho; 20 mi. S of Livermore; [37.558 ° N, 121.581 ° W]; 22 Jul. 1955; ex Asclepias; EMEC (Figs 23, 24 D) • 4 ♀♀; Butte Co., Chico; [39.729 ° N, 121.8371 ° W]; 26 Apr. 1967; R. G. Malin leg.; UCRC (Figs 22, 24 A) • 1 ♀; Contra Costa Co., Pine Canyon, Mount Diablo; [38.456 ° N, 121.978 ° W]; 23 May 1937; R. L. Usinger leg.; EMEC • 1 ♀; El Dorado Co., Snowline Camp; [38.747 ° N, 120.624 ° W]; 20 Jun. 1948; P. D. Hurd leg.; ex Chamaebatia foliolosa; EMEC • 2 ♀♀; Humboldt Co., Garberville; [40.10 ° N, 123.80 ° W]; 19 Apr. 1935; ex Cercocarpus betuloides; UCRC (Fig. 24 B) • 1 ♂; Humboldt Co., Redwood Creek at Stover Road, Redwood Valley N. of Hwy 299; [37.505 ° N, 122.213 ° W]; elev. 198.12 m; 10 Aug. 1966; H. B. Leech leg.; CAS (Fig. 25) • 2 ♀♀; Humboldt Co.; [40.75 ° N, 123.87 ° W]; 23 May 1911; F. W. Nunenmacher leg.; USNM • 1 ♀; Inyo Co., Haiwee Reservoir; [36.155 ° N, 117.953 ° W]; 23 Apr. 1944; R. D. Flrich leg.; UCRC • 1 ♀; Los Angeles Co., 5 mi. S of Pearblossom; [34.431 ° N, 117.910 ° W]; 23 Apr. 1956; E. G. Linsley leg.; ex Tamarix gallica; EMEC • 2 ♀♀; Los Angeles Co., Acton; [34.470 ° N, 118.197 ° W]; 19 Mar. 1956; E. G. Linsley leg.; EMEC • 3 ♀♀; same data as for preceding; 19 Mar. 1936; E. G. Linsley leg.; ex Salix; EMEC • 1 ♀; same data as for preceding; 19 Apr. 1936; E. G. Linsley leg.; ex Salix; EMEC • 1 ♀; Los Angeles Co., San Gabriel Can. near Dam no. 1; [34.2064 ° N, 117.8603 ° W]; 17 Apr. 1955; Hogue leg.; LACM • 1 ♀; Los Angeles Co., Santa Monica Mountains; [34.120 ° N, 118.932 ° W]; 18 Apr. 1953; A. Fukushima leg.; LACM • 1 ♀; Madera Co.; [37.25 ° N, 119.7 ° W]; 19 May 1942; A. J. Walz leg.; ex Lupinus; EMEC • 1 ♀; Mariposa Co., Yosemite; [37.833 ° N, 119.500 ° W]; 9 May 1941; Cockerell leg.; ex Nemophila; UCRC • 1 ♀; Mariposa Co., Yosemite National Park; [37.865 ° N, 119.538 ° W]; 1 – 31 May 1910; LACM • 1 ♀; Mariposa Co., Yosemite Valley; [37.746 ° N, 119.594 ° W]; 19 Jun. 1921; E. O. VanDyke leg.; CAS • 1 ♀; same data as for preceding; 17 Jun. 1921; E. P. Van Duzee leg.; CAS • 1 ♀; Mariposa Co.; [37.51 ° N, 119.88 ° W]; 2 Jun. 1914; F. W. Nunenmacher leg.; EMEC • 1 ♀; Mendocino Co., Big Rock Creek; [34.408 ° N, 117.828 ° W]; 25 Apr. 1926; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; Mendocino Co., Hopland; [38.979 ° N, 123.108 ° W]; 9 May 1925; E. P. Van Duzee leg.; CAS • 1 ♀; Mendocino Co., UC Hopland Field Station, near Headquarters; [39.001 ° N, 123.083 ° W]; elev. 268.22 m; 17 May 1968; W. J. Turner leg.; EMEC • 2 ♀♀; Monterey Co., Arroyo Seco Campground; [36.236 ° N, 1231.482 ° W]; 24 Apr. 1976; M. Wasbauer leg.; CSCA • 1 ♀; Nevada Co., 3 mi. NW of Newton; [38.735 ° N, 120.719 ° W]; 14 Apr. 1934; P. H. Timberlake leg.; ex Cryptantha intermedia; UCRC • 2 ♀♀; Plumas Co., Belden; [40.006 ° N, 121.249 ° W]; 12 Apr. 1949; J. W. MacSwain leg.; ex Rhus; EMEC • 1 ♀; Riverside Co., Riverside; [33.956 ° N, 117.372 ° W]; elev. 290 m; 19 Feb. 1928; P. H. Timberlake leg.; ex Salix vestita (verbatim: Salix leiolepis); UCRC • 1 ♀; same data as for preceding; 2 May 1939; P. H. Timberlake leg.; UCRC • 1 ♀; same data as for preceding; 6 Apr. 1940; P. H. Timberlake leg.; ex Salix gooddingii; UCRC • 2 ♀♀; same data as for preceding; 13 May 1951; P. H. Timberlake leg.; ex boysenberry; UCRC • 1 ♀; same data as for preceding; 4 May 1950; P. H. Timberlake leg.; ex Sisymbrium irio; UCRC • 1 ♀; same data as for preceding; 4 May 1950; P. H. Timberlake leg.; ex Salix vestita (verbatim: Salix leiolepis); UCRC • 1 ♀; same data as for preceding; 22 Feb. 1935; P. H. Timberlake leg.; ex Sedum; UCRC • 1 ♀; same data as for preceding; 24 May 1925; P. H. Timberlake leg.; ex? Margarita; UCRC • 1 ♀; same data as for preceding; 18 May 1947; P. H. Timberlake leg.; ex Veronica; UCRC • 1 ♂; Riverside Co., Riverside; Santa Ana River; [33.988 ° N, 117.396 ° W]; 26 Jun. 1965; G. R. Ballmer leg.; UCRC • 1 ♀; San Benito Co., Paicines; [36.7305 ° N, 121.2791 ° W]; 24 Apr. 1968; J. A. Chemsak leg.; ex Baccharis; EMEC • 2 ♀♀; San Bernardino Co., Tetley Park; [34.246 ° N, 117.309 ° W]; 08 May 1940; Philip H. Timberlake leg.; ex Erodium cicutarium; UCRC • 1 ♀; same data as for preceding; 15 May 1937; P. H. Timberlake leg.; ex Salix laevigata; UCRC • 1 ♀; San Bernardino Co., 15 km SSE of Hesperia, Hwy 173; [34.301 ° N, 117.238 ° W]; elev. 1021.1 m; 21 Apr. 1981; J. A. Bethke leg.; UCRC • 1 ♀; San Bernardino Co., Lytle Creek Ranger Sta., 10 mi. N of Fontana; [34.234 ° N, 117.480 ° W]; 14 Apr. 1965; C. Slobodchikoff leg.; EMEC • 1 ♀; San Bernardino Co., Lytle Creek; 8 mi. of N of Fontana; [34.259 ° N, 117.500 ° W]; 14 Apr. 1965; J. Powell leg.; ex Ceanothus; EMEC • 1 ♀; San Bernardino Co., Mill Creek; [40.360 ° N, 121.493 ° W]; elev. 1341 m; 1 – 31 May 1930; ex Cryptantha intermedia; UCRC • 1 ♀; San Bernardino Co., Miller Canyon, San Bernardino Mountains; [34.13 ° N, 116.88 ° W]; 19 May 1941; E. O. VanDyke leg.; CAS • 1 ♀; San Bernardino Co., Wildwood Canyon; 5 mi. E of Calimesa; [34.020 ° N, 116.999 ° W]; 14 May 1969; M. E. Irwin leg.; UCRC • 1 ♀; San Diego Co., Cuyumaca; [32.946 ° N, 116.576 ° W]; 19 Apr. 1950; E. G. Linsley leg.; ex Baeria; EMEC • 1 ♀; San Diego Co., Descanso; [32.853 ° N, 116.616 ° W]; 30 Mar. 1961; P. D. Hurd leg.; ex Salix; EMEC • 1 ♀; San Diego Co., Rincon; [33.324 ° N, 116.969 ° W]; 16 Jul. 1928; E. O. VanDyke leg.; CAS • 1 ♀; San Diego Co., Warner Springs; [33.282 ° N, 116.634 ° W]; 19 Apr. 1950; E. G. Linsley leg.; ex Cryptantha; EMEC • 2 ♀♀; San Luis Obispo Co., 5 mi. NW of Nipomo; [35.093 ° N, 120.539 ° W]; 19 Mar. 1969; P. A. Opler leg.; ex Ceanothus; EMEC • 1 ♀; San Luis Obispo Co., Nacimiento Dam; [35.855 ° N, 120.926 ° W]; 14 Apr. 1967; P. A. Rude leg.; EMEC • 1 ♀; San Luis Obispo Co., Paso Robles; [36.293 ° N, 120.703 ° W]; 22 Apr. 1919; E. P. Van Duzee leg.; CAS • 1 ♀; San Luis Obispo Co., Pozo; [35.305 ° N, 120.375 ° W]; 28 Apr. 1962; C. A. Toschi leg.; EMEC • 1 ♀; same data as for preceding; 27 Apr. 1962; J. Powell leg.; EMEC • 1 ♀; same data as for preceding; 20 Apr. 1962; C. A. Toschi leg.; EMEC • 1 ♂; San Mateo Co., Corte de Madera Creek, Portola; [37.397 ° N, 122.316 ° W]; 27 Jun. 1959; P. H. Arnaud Jr leg.; CAS • 1 ♀; Santa Barbara Co., Los Prietos; [34.542 ° N, 119.802 ° W]; 25 Jun. 1965; E. M. Omi leg.; EMEC • 1 ♀; same data as for preceding; 23 Jun. 1965; E. M. Omi leg.; EMEC • 1 ♀; same data as for preceding; 23 Jun. 1965; J. Powell leg.; EMEC • 1 ♂; same data as for preceding; 23 Jun. 1965; R. W. Gear leg.; EMEC (Fig. 24 E) • 2 ♀♀; Santa Clara Co., Stanford University; [37.43 ° N, 122.17 ° W]; 4 Apr. 1956; P. H. Arnaud Jr leg.; CAS • 1 ♀; same data as for preceding; 10 Apr. 1956; P. H. Arnaud Jr leg.; ex Stellaria media; CAS • 1 ♀; Santa Clara Co., Stevens Creek Reservoir; [37.297 ° N, 122.077 ° W]; 13 Apr. 1953; D. Burdick leg.; EMEC • 1 ♂; Santa Cruz Co., Mount Hermon; [37.051 ° N, 122.058 ° W]; 7 – 30 Jul. 1932; F. E. Blaisdell leg.; CAS • 1 ♀; Shasta Co., 7 mi. NE of Burney; [40.953 ° N, 121.566 ° W]; 30 May 1952; J. G. Rozen leg.; EMEC • 1 ♀; Shasta Co., Castle Rock State Park; [37.230 ° N, 122.125 ° W]; elev. 2133.6 m; 3 May 1953; J. D. Lattin leg.; EMEC • 1 ♀; Shasta Co., Old Station; [40.675 ° N, 121.431 ° W]; 22 Jun. 1955; E. E. Lindquist leg.; EMEC • 1 ♀; Shasta Co., Redding; [40.579 ° N, 122.394 ° W]; 7 Jul. 1979; T. R. Haig leg.; CSCA • 1 ♀; Shasta Co.; [40.79 ° N, 121.85 ° W]; 2 May 1913; F. W. Nunenmacher leg.; EMEC • 1 ♀; Stanislaus Co., La Grange; [37.664 ° N, 120.464 ° W]; 15 Apr. 1961; R. R. Snelling and M. D. Snelling leg.; ex Salix; LACM • 1 ♀; Trinity Co., 6 mi. NE of Hayfork; [40.611 ° N, 123.101 ° W]; 20 May 1973; J. Chemsak leg.; ex Rhamnus; EMEC • 1 ♀; Trinity Co., Hayfork; [40.555 ° N, 123.184 ° W]; 23 May 1973; J. Chemsak leg.; EMEC • 1 ♀; Trinity Co., Hayfork Ranger Station; [40.553 ° N, 123.165 ° W]; elev. 701 m; 18 May 1973; EMEC • 1 ♀; Tulare Co., 2 mi. E of Johnsondale; [35.975 ° N, 118.504 ° W]; 27 Apr. 1964; C. A. Toschi leg.; EMEC • 1 ♀; same data as for preceding; 27 Apr. 1964; C. A. Toschi leg.; ex Prunus subcordata; EMEC • 1 ♀; same data as for preceding; 27 Apr. 1964; J. Powell leg.; ex Prunus subcordata; EMEC • 2 ♀♀; same data as for preceding; 27 Apr. 1964; P. Rude leg.; ex Salix sp.; EMEC • 1 ♀; Tulare Co., Fairview; [37.679 ° N, 122.051 ° W]; 27 Apr. 1964; C. A. Toschi leg.; EMEC • 1 ♀; Tulare Co., Visalia; [36.330 ° N, 119.292 ° W]; Culbertson leg.; USNM • 1 ♀; Tuolumne Co., Basin Creek campground; [38.166 ° N, 120.184 ° W]; 1 Jul. 1963; P. H. Arnaud Jr leg.; CAS • 1 ♀; Ventura Co., Chuchupate Ranger Station, base Frazier Mtn; [34.816 ° N, 119.011 ° W]; 5 May 1959; G. I. Stage leg.; ex Descurainia sophia; EMEC • 1 ♀; Ventura Co., Grade Valley; 9 mi. SW of Stauffer P. O.; [34.655 ° N, 119.109 ° W]; 7 May 1959; C. W. O’Brien leg.; EMEC • 10 ♀♀; Ventura Co., Lockwood Creek near Stauffer P. O.; [34.730 ° N, 119.033 ° W]; 5 May 1959; P. D. Hurd leg.; ex Descurainia sophia; EMEC • 1 ♀; Ventura Co., Lockwood Creek, near Stauffer P. O.; [34.730 ° N, 119.033 ° W]; 5 May 1959; J. R. Powers leg.; ex Descurainia sophia; EMEC • 2 ♀♀; same data as for preceding; 7 May 1959; P. D. Hurd leg.; ex Descurainia sophia; EMEC • 1 ♀; Ventura Co., Lockwood Creek, near Stauffer P. O.; [34.729 ° N, 119.033 ° W]; 7 May 1959; J. R. Powers leg.; EMEC • 1 ♀; same data as for preceding; 7 May 1959; P. D. Hurd leg.; EMEC • 1 ♀; same data as for preceding; 5 May 1959; C. W. O’Brien leg.; EMEC • 1 ♀; same data as for preceding; 5 May 1959; G. I. Stage leg.; EMEC • 1 ♀; same data as for preceding; 5 May 1959; G. I. Stage leg.; ex Salix; EMEC • 1 ♀; Ventura Co., Lockwood Valley; [34.733 ° N, 119.042 ° W]; 10 May 1954; P. H. Timberlake leg.; ex Phacelia ciliata; UCRC • 1 ♀; same data as for preceding; 5 May 1959; P. H. Timberlake leg.; ex Eleutherine bulbosa (verbatim: Sisyrinchium bulbosum); UCRC • 1 ♀; Ventura Co., Lockwood Creek; near Stauffer P. O.; [34.730 ° N, 119.033 ° W]; 7 May 1959; J. Powell leg.; ex Euphorbia albomarginata; EMEC • 1 ♀; same data as for preceding; 7 May 1959; J. Powell leg.; ex Hesperochiron californicus; EMEC • 1 ♀; same data as for preceding; 5 May 1959; C. W. O’Brien leg.; ex Descurainia sophia; EMEC • 1 ♀; Ventura Co., Lockwood Valley; [34.733 ° N, 119.042 ° W]; 10 May 1954; P. H. Timberlake leg.; ex Phacelia ciliata; UCRC • 1 ♀; same data as for preceding; 5 May 1959; P. H. Timberlake leg.; ex Eleutherine bulbosa (verbatim: Sisyrinchium bulbosum); UCRC • 1 ♀; Ventura Co., S. Gorge; [34.579 ° N, 119.258 ° W]; 15 May 1955; Menke and Stange leg.; LACM • 1 ♀; Cuyama Valley; [35.02 ° N, 119.87 ° W]; 26 Apr. 1905; P. M. Daumirs leg.; UCRC. – Washington • 1 ♀; Benton Co., Prosser, Irr. Exp. Sta. Roza Unit; [46.267 ° N, 119.741 ° W]; 17 Apr. 1951; E. Klostermeyer leg.; WSU • 1 ♀; Whitman Co., Almota; [46.703 ° N, 117.469 ° W]; WSU • 1 ♀; Whitman Co., Pullman; [46.730 ° N, 117.182 ° W]; 14 May 1904; WSU • 1 ♀; same data as for preceding; 2 May 1949; H. F. Menke leg.; WSU • 1 ♀; Whitman Co., Wawawai; [46.637 ° N, 117.375 ° W]; 14 May 1910; WSU • 1 ♀; same data as for preceding; R. W. Doane leg.; WSU • 1 ♀; same data as for preceding; WSU • 1 ♀; Yakima Co., Selah; [46.654 ° N, 120.530 ° W]; 14 May 1919; RBCM.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD8FFECFE17FE86FC15FADB.taxon	description	Description GENITALIA (Figs 25, 120 C). Gonostylus short, approximately as long as broad, with short setae. Retrorse lobe attenuated apically, about 3.5 times as long as broad, covered in short sparse setae.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD8FFECFE17FE86FC15FADB.taxon	distribution	Range Canada (British Columbia) and the USA (California, Oregon, and Washington) (Fig. 26).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD8FFECFE17FE86FC15FADB.taxon	biology_ecology	Floral hosts ANACARDIACEAE: Rhus sp. • APOCYNACEAE: Asclepias sp. • ASTERACEAE: Argyranthemum frutescens (L.) Sch. Bip., Lasthenia sp. • BORAGINACEAE: Cryptantha intermedia (A. Gray) Greene • BRASSICACEAE: Descurainia sophia (L.) Webb ex Prantl, Sisymbrium irio L., Thysanocarpus curvipes Hook. • CRASSULACEAE: Sedum sp. • GERANIACEAE: Erodium cicutarium (L.) L’Hér. • HYDROPHYLLACEAE: Nemophila sp., Phacelia ciliata Benth., Phacelia distans Benth. • PAEONIACEAE: Paeonia brownii Douglas ex Hook. • GROSSULARIACEAE: Ribes roezlii Regel • RANUNCULACEAE: Ranunculus californicus Benth. • RHAMNACEAE: Ceanothus leucodermis Greene, Ceanothus sp., Rhamnus ilicifolia Kellogg, Rhamnus sp. • ROSACEAE: Cercocarpus betuloides Nutt., Prunus subcordata Benth., Rubus sp. • SALICACEAE: Salix gooddingii C. R. Ball, Salix laevigata Bebb, Salix vestita Pursh, Salix sp. • TAMARICACEAE: Tamarix gallica L.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD8FFECFE17FE86FC15FADB.taxon	discussion	Remarks Both Cockerell (1925) and Michener (1936) only refer to a single specimen in their original descriptions. The type localities are separated by over 500 km along the California coast. Michener (1936) distinguished L. humboldtense from L. aspilurus based on the black apex of the metasoma and the sparser mesoscutal punctures. Two specimens collected with the L. humboldtense holotype have reddish brown T 5 and small impunctate areas mesad of the parapsidal line. Individuals of L. aspilurus show variation in the density of mesoscutal punctures in this area. Some are rather close, but there is too much variation even within a single location to discriminate species on this basis.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD1FFF1FE35FAB5FD92F956.taxon	description	Figs 27 – 30, 95 I, 97 E, 112 C, 116 B, 119 C, 120 D	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD1FFF1FE35FAB5FD92F956.taxon	diagnosis	Diagnosis Both sexes of L. buccale are entirely black, and easily diagnosed from North American Lasioglossum by their size (females 9 – 10 mm, males 7.5 – 8.5 mm); long heads (females length / width 1.25 (± 0.04), males length / width 1.28 (± 0.02 SD )); and coarse mesepisternal punctures. The female inner metatibial spur has 10 – 14 teeth, shorter than the width of the primary rachis (Fig. 27 E). Lasioglossum ovaliceps is the most similar but is easily separated by the orange-red metasoma in both sexes, and the female inner metatibial pectinate with 3 – 5 long branches.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD1FFF1FE35FAB5FD92F956.taxon	etymology	Etymology Pérez (1903) did not explain the epithet. The specific epithet likely has its roots in the Latin ‘ bucca ’ meaning ‘ jaw’ or ‘ mouth’, with an adjectival suffix, ‘ - alis ’; however, the author’s intent is unknown.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD1FFF1FE35FAB5FD92F956.taxon	materials_examined	Material examined AUSTRIA – Burgenland • 1 ♀; Illmitz, Seedamm 400 m N of Biol. Station; 47.7731 ° N, 16.7663 ° E; 11 – 12 Aug. 2023; L. Bertsch and K. Kraus leg.; Leander Bertsch’s personal collection • 1 ♀; same data as for preceding; 12 Aug. 2023; L. Bertsch leg.; Leander Bertsch’s personal collection • 1 ♀; same data as for preceding; 11 Aug. 2023; L. Bertsch leg.; WRME. CANADA – British Columbia • 2 ♀♀; Oliver, Inkaneep Provincial Park; 49.22381 ° N, 119.53921 ° W; 14 Jun. – 20 Jul. 2017; K. G. Marks leg.; RSKM • 2 ♀♀; Inkaneep Provincial Park; 49.22381 ° N, 119.53921 ° W; 14 Jun. – 20 Jul. 2017; D. Marks and K. Grant (J. Heron) leg.; RSKM. GERMANY – Baden-Württemberg • 1 ♀; Achkarren am Kaiserstuhl, Schneckenberg; 48.0695 ° N, 7.6253 ° E; 8 May 2022; Handfang leg.; Leander Bertsch’s personal collection. UNITED STATES – Oregon • 1 ♀; Marion Co., Independence; 44.839 ° N, 123.166 ° W; 15 May 2017; G. Hoffman leg.; WRME • 1 ♀; Multnomah Co., Oxbow Meadow; 45.4844 ° N, 122.3015 ° W; 30 Sep. 2020; J. D. Engler leg.; OSAC • 1 ♀; Multnomah Co., Oxbow Meadow; 45.485 ° N, 122.302 ° W; 27 Jul. 2021; J. D. Engler leg.; OSAC • 1 ♀; same data as for preceding; 28 Jul. 2021; J. D. Engler leg.; OSAC • 1 ♂; Multnomah Co., Oxbow Pkway; 45.494 ° N, 122.313 ° W; 3 Aug. 2021; J. D. Engler leg.; OSAC • 1 ♂; Multnomah Co., Oxbow Meadow; 45.485 ° N, 122.302 ° W; 31 Jul. 2021; J. D. Engler leg.; OSAC (Fig. 29) • 1 ♂; same data as for preceding; 2 Sep. 2021; J. D. Engler leg.; OSAC • 1 ♀; same data as for preceding; 7 Jun. 2022; J. D. Engler leg.; OSAC • 1 ♀; same data as for preceding; 15 Jun. 2022; J. D. Engler leg.; OSAC. – Washington • 1 ♀; Chelan Co., Ardenvoir, Cougar Creek Fire area; 47.772 ° N, 120.422 ° W; 29 – 30 May 2023; A. Maust leg.; WSU • 1 ♀; Chelan Co., Chelan; [47.83 ° N, 120.02 ° W]; 5 Jul. – 1 Aug. 2022; A. Maust leg.; WSU • 2 ♀♀; same data as for preceding; 22 Jun. – 6 Jul. 2021; A. Maust leg.; WSU • 2 ♀♀; same data as for preceding; 22 Jun. – 3 Aug. 2021; A. Maust leg.; WSU • 2 ♀♀; same data as for preceding; 28 Apr. – 3 Aug. 2021; A. Maust leg.; WSU • 1 ♀; same data as for preceding; 8 Jun. – 6 Jul. 2021; A. Maust leg.; WSU • 1 ♀; Chelan Co., Chelan; 47.939 ° N, 120.055 ° W; 12 – 13 Jun. 2022; A. Maust leg.; WSU • 2 ♀♀; Chelan Co., Chelan; 47.939 ° N, 120.055 ° W; 10 – 11 Jul. 2023; A. Maust leg.; WSU • 1 ♀; Chelan Co., Chelan; 47.940 ° N, 120.042 ° W; 12 – 13 Jun. 2023; A. Maust leg.; WSU • 1 ♀; Chelan Co., Chelan; 47.940 ° N, 120.042 ° W; 29 – 30 May 2023; A. Maust leg.; WSU • 1 ♀; Chelan Co., Chelan; 47.964 ° N, 120.047 ° W; 12 – 13 Jun. 2023; A. Maust leg.; WSU • 1 ♀; Chelan Co., Chelan; 47.953 ° N, 120.036 ° W; 26 – 27 Jun. 2023; A. Maust leg.; WSU • 1 ♀; Chelan Co., Chelan; 47.940 ° N, 120.042 ° W; 12 – 13 Jun. 2023; A. Maust leg.; WSU • 2 ♂♂; Jefferson Co., near Crowberry Bog; 47.805 ° N, 124.201 ° W; elev. 144 m; 6 Aug. 2023; I. Carmean leg.; ex Hypochaeris radicata; WSU • 1 ♂; Jefferson Co., Jefferson, near Forks; 47.805 ° N, 124.201 ° W; elev. 144 m; 6 Aug. 2023; A. Wilson leg.; ex Chamaenerion angustifolium; WSU • 2 ♀♀; same data as for preceding; 6 Aug. 2023; A. Wilson leg.; ex Digitalis purpurea; WSU • 4 ♀♀; King Co., Beacon Food Forest; 47.5686 ° N, 122.3126 ° W; 16 Jul. 2014; E. Bloom leg.; WSU • 1 ♀; King Co., Interbay; 47.6410 ° N, 122.3762 ° W; 18 Jul. 2014; E. Bloom leg.; WSU • 1 ♀; King Co., Magnuson Park; 47.6826 ° N, 122.2585 ° W; 18 Jul. 2014; E. Bloom leg.; WSU • 1 ♀; King Co., Marra Farm; 47.5215 ° N, 122.3289 ° W; 15 Jul. 2014; E. Bloom leg.; WSU • 1 ♀; King Co., Rainier Beach Urban Farm and Wetland; 47.5259 ° N, 122.2628 ° W; 4 Aug. 2016; E. Bloom leg.; WSU • 1 ♀; King Co., Seattle; 47.4274 ° N, 122.3081 ° W; 5 Jul. 2020; Peterman W. leg.; WRME (Fig. 27) • 1 ♀; King Co., Seattle; 47.6062 ° N, 122.3321 ° W; 6 Sep. 1964; OSAC • 1 ♀; King Co., Seattle; 47.5059 ° N, 122.2672 ° W; 10 Jun. 2016; R. Redmond and E. A. Sugden leg.; WRME • 1 ♀; King Co., Seattle; 47.5039 ° N, 122.2757 ° W; 19 Apr. 2015; R. Redmond and E. A. Sugden leg.; WRME • 1 ♀; King Co., Seattle; 47.4285 ° N, 122.3036 ° W; 6 Aug. 2018; E. A. Sugden and W. Peterman leg.; WRME • 1 ♀; King Co., Seattle; 47.4274 ° N, 122.3081 ° W; 5 Jul. 2020; W. Peterman leg.; WRME • 1 ♀; King Co., Steel Wheel Farm; 47.5817 ° N, 121.9175 ° W; 21 Jul. 2014; E. Bloom leg.; WSU • 1 ♀; Lincoln Co., Lamona; 47.360732 ° N, 118.476279 ° W; 18 May 2016; T. D. Hatten leg.; WRME • 1 ♀; Lincoln Co., Lamona, Sage Downs Rd N, S of Coal Coule; 47.377576 ° N, 118.391616 ° W; 23 May 2016; T. D. Hatten leg.; WRME • 1 ♀; Lincoln Co., Lamona; 47.360732 ° N, 118.476279 ° W; 20 May 2016; T. D. Hatten leg.; WRME • 1 ♀; Lincoln Co., Lamona; 47.360732 ° N, 118.476279 ° W; 2 Aug. 2017; Sprague R. leg.; WRME • 1 ♀; Lincoln Co., Lamona; 47.360732 ° N, 118.476279 ° W; 27 Jul. 2017; T. D. Hatten leg.; WRME • 1 ♀; same data as for preceding; 2 Aug. 2017; T. D. Hatten leg.; WRME • 1 ♂; Spokane Co.; 47.62 ° N, 117.4 ° W; 15 Oct. 1967; Robin D. Gray leg.; WSU • 1 ♀; Spokane Co., Fairchild Airforce Base, 10 mi. W of Spokane; 47.60393 ° N, 117.62817 ° W; elev. 735 m; 7 Jun. 2024; R. S. Zack leg.; WSU • 1 ♀; Spokane Co., Turnbulls; 47.42553 ° N, 117.565653 ° W; 25 Jul. 2017; Dibble M. leg.; WRME • 1 ♂; Snohomish County Co., Everett; 47.93 ° N, 122.2852 ° W; 30 Jul. 2018; E. A. Sugden and W. Peterman leg.; WSU • 1 ♀; Thurston Co.; 46.953 ° N, 123.025 ° W; elev. 57 m; 26 Jul. 2009; C. Fimbel leg.; PCYU • 1 ♀; Thurston Co., Wendell Berry Garden; 47.0521 ° N, 122.8732 ° W; 13 Jul. 2014; R. Bietz leg.; WSU • 1 ♀; Yakima Co., White Swan; 46.4693 ° N, 120.7016 ° W; 20 Jul. 2013; A. A. Jordan leg.; WSU.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD1FFF1FE35FAB5FD92F956.taxon	description	Description GENITALIA (Figs 29, 120 D). Gonostylus short and broad approximately as long as broad, with setae short. Retrorse lobe broad and curved, about 3 times as long as broad, rounded at apex, covered in circular bumps.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD1FFF1FE35FAB5FD92F956.taxon	distribution	Range Oregon, Washington, and west of British Columbia (Fig 30).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD1FFF1FE35FAB5FD92F956.taxon	biology_ecology	Floral hosts ASTERACEAE: Hypochaeris radicata L. • CARYOPHYLLACEAE: Silene spaldingii S. Watson. • ONAGRACEAE: Chamaenerion angustifolium (L.) Schur • SCROPHULARIACEAE: Digitalis purpurea L. In its native range, L. buccale has been associated with Linaria vulgaris Mill. and related taxa in Plantaginaceae Juss. like Digitalis L. and Antirrhinum L. (Burger 2020; Ebmer 2003). Linaria vulgaris is a widespread, introduced species in North America.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFD1FFF1FE35FAB5FD92F956.taxon	discussion	Remarks Ebmer (2003) reports seeing a female entering a nest in a stone wall “ fing ich beim Einflug zum Nest in einer Trockenmauer ”. The collection records from Seattle, Washington, in September 1964, and Spokane, Washington, in October 1967, indicate that this species has been present in the region for several decades, suggesting it is not a recent introduction. However, it is interesting that this species began to appear again in collections data after 2013 (Hatten et al. 2024), raising questions about its recent presence or increased documentation in the region. DNA barcodes DNA barcodes from Washington, Austria, and France were compared and found to form a discrete cluster from other Lasioglossum species.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFCCFFF9FDDFF936FC51FEA4.taxon	description	urn: lsid: zoobank. org: act: 1995 AA 5 A- 9 DB 0 - 4422 - 9 A 74 - B 991 C 5237149 Figs 31 – 34, 102 A, 117 D, 119 E, 120 E	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFCCFFF9FDDFF936FC51FEA4.taxon	diagnosis	Diagnosis Females of L. engleri sp. nov. can be recognised by the following combination of characters: head long, length / width 1.05 (± 0.03 SD); supraclypeal punctation sparse (IS = 1 – 3 PD) medially and anteriorly, becoming dense (IS ≤ 1 PD) laterally and posteriorly; mesepisternum distinctly punctate; and T 2 – T 3 basolaterally tomentum. Males of L. engleri sp. nov. can be recognised by the following combination of characters: head long, length / width ratio 1.06 (± 0.02 SD); supraclypeal punctation dense (IS ≤ 1 PD); and mesepisternum distinctly punctate. Eastern North American L. macoupinense (Robertson, 1895) and L. birkmanni (Crawford, 1906 a) are very closely related to L. engleri sp. nov. However, both sexes of L. birkmanni can be distinguished by their shorter heads (length / width ratio 0.95 – 1.03). Females of L. engleri can be distinguished from L. macoupinense by the presence of short setae overhanging the margin of T 2 – 3, which are particularly evident laterally (Fig. 102). While some L. macoupinense females may also exhibit some short setae overhanging the margin of T 2 – 3, L. engleri is characterized by significantly denser and longer hairs on the metasoma, especially on T 2 – 3. Additionally, females of L. macoupinense have a coriarious microsculpture on the T 2 apical impressed area, while females of L. engleri lack or exhibit only very weak coriarious microsculpture in this region (Fig. 102). Males of L. macoupinense can be distinguished from L. engleri sp. nov. by their supraclypeal area: L. engleri has a polished supraclypeal area with deep punctures, while L. macoupinense males exhibit shallow punctures on an imbricate supraclypeal area.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFCCFFF9FDDFF936FC51FEA4.taxon	etymology	Etymology The specific epithet ‘ engleri ’ is a dedication to Joseph D. Engler, U. S. Fish and Wildlife Service, Regional Refuge Biologist and Pacific Northwest Pollinator Conservation Coordinator (retired) for sharing specimens, DNA barcode information, and valuable context regarding this species.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFCCFFF9FDDFF936FC51FEA4.taxon	materials_examined	Type material examined Holotype UNITED STATES – Oregon • ♀; Sherman Co., Fulton Canyon; 45.6345 ° N, 120.86424 ° W; 10 May 2011; J. D. Engler leg.; OSAC (Fig. 31). Specimen labels “ USA: OR: Sherman Co. / Fulton Canyon / N 45.63450, W 120.86424 / 10 May 2011; J. D. Engler // Lasioglossum (Hemihalictus) macoupinense / Det: JD Engler 2021 // Lasioglossum / GIBBS- 14928 // HOLOTYPE / Lasioglossum (Hemihalictus) engleri Hettiarachchi and Gibbs ”. Paratypes CANADA – Alberta • 5 ♀♀; Cypress Co., Medicine Hat near Gas City Campgr., wet coulee bottom; 50.037222 ° N, 110.729722 ° W; 4 Jan. – 31 May 2016; M. Buck leg.; PMAE • 1 ♀; Cypress Co., Redcliff, floodplain, river sand; 50.054722 ° N, 110.786667 ° W; 26 – 28 Jan. 2017; M. Buck leg.; PMAE • 3 ♀♀; Newell Co., Dinosaur Provincial Park, Sandhill Creek Valley; 50.755833 ° N, 111.519167 ° W; 14 – 15 Jul. 2010; Buck and Heule leg.; PMAE • 1 ♀; Warner No. 5, Writing-on-Stone Provincial Park; [49.08 ° N, 111.62 ° W]; 21 – 28 Jul. 1990; D. McCorquodale leg.; PMAE • 2 ♀♀; Edmonton, Kinnaird Park, vertical river bank; 53.556944 ° N, 113.454167 ° W; 10 Aug. 2018; M. Buck leg.; PMAE. UNITED STATES – California • 1 ♀; Cruz Co., Bates Creek, 2 mi. NE of Soquel; [37.008 ° N, 121.932 ° W]; 2 May 1971; J. Doyen leg.; EMEC • 3 ♀♀; El Dorado Co., Pino Grande, SW of Lake Edson; [37.631 ° N, 119.524 ° W]; 29 Jun. 1967; W. Turner leg.; EMEC • 1 ♀; El Dorado Co.; [38.76 ° N, 120.57 ° W]; 30 Jun. 1912; F. W. Nunenmacher leg.; EMEC • 1 ♀; El Dorado Co., Pollock Pines; [38.761 ° N, 120.587 ° W]; 27 May 1970; D. P. Levin leg.; LACM • 1 ♀; Humboldt Co., 3 mi. N of Gargerville; [40.143 ° N, 123.795 ° W]; 23 May 1976; R. Wharton and R. Coville leg.; EMEC • 1 ♀; Mendo Co., 5 mi. N of Branscomb; [39.725 ° N, 123.626 ° W]; 24 – 25 May 1976; R. Wharton leg.; EMEC • 1 ♀; Mendocino Co., Russian River, Hopland; [38.973 ° N, 123.111 ° W]; 3 Apr. 1968; J. Powell leg.; EMEC • 1 ♂; Modoc Co., Alturas Insp. Sta; [41.523 ° N, 120.476 ° W]; 1 Oct. 1975; T. R. Haig leg.; CSCA (Fig. 32) • 1 ♂; Mono Co., 7 mi. S of Mill Creek Canyon, Junction Highway; [33.986 ° N, 117.039 ° W]; 25 – 26 Aug. 1979; M. Wasbauer and P. Adams leg.; CSCA (Fig. 33) • 1 ♀; Sacramento Co., Walnut Grove; [38.242 ° N, 121.511 ° W]; 6 Apr. 1931; ex Pear tree; CSCA • 1 ♀; Santa Clara Co., Palo Alto; [37.429 ° N, 122.138 ° W]; 25 Mar. 1995; EMEC • 1 ♀; Shasta Co., Old Station; [40.675 ° N, 121.431 ° W]; J. W. MacSwain leg.; EMEC • 1 ♀; Shasta Co., 7 mi. SW of Viola; [40.446 ° N, 121.772 ° W]; elev. 1219.2 m; 20 May 1966; J. Powell leg.; EMEC • 1 ♀; Trinity Co., Hayfork Ranger Station; [40.548 ° N, 123.164 ° W]; elev. 701 m; 18 May 1973; J. Powell leg.; EMEC. – New Mexico • 1 ♀; Rio Arriba Co. Co., Ojo Caliente; [36.1094 ° N, 106.143 ° W]; 13 Jun. 1999; L. Packer leg.; PCYU. – Oregon • 1 ♀; Baker Co., Big Creek, 26 mi. SE of Union; [44.977 ° N, 117.428 ° W]; elev. 1280.2 m; 1 – 3 Jul. 1976; E. J. Davis leg.; WSU • 1 ♂; Baker Co., Cougar Creek, 37 mi. SE of Union; [44.879 ° N, 117.252 ° W]; elev. 1243.6 m; 22 – 25 Aug. 1976; E. J. Davis leg.; WSU • 1 ♀; Benton Co., 4 mi. SW of Corvallis; [44.523 ° N, 123.320 ° W]; 1 May 1956; J. D. Lattin leg.; ex Ranunculus; EMEC • 1 ♀; Benton Co., Corvallis; [44.567 ° N, 123.283 ° W]; 23 May 1945; Grace Bennett leg.; OSAC • 1 ♀; Crook Co., Pineville; 44.171 ° N, 120.545 ° W; elev. 1024 m; 9 May 2019; A. Bessette leg.; OSAC • 1 ♀; Douglas Co., Roseburg; [43.223 ° N, 123.352 ° W]; 6 Apr. 1930; J. Wilcox leg.; OSAC • 1 ♀; Josephine Co., Gallice; 42.562 ° N, 123.584 ° W; elev. 215 m; 19 Mar. 2020; J. Maxwell leg.; OSAC • 1 ♀; Josephine Co., Wolf Creek; 42.651 ° N, 123.59 ° W; elev. 223 m; 6 May 2019; E. Watrous leg.; OSAC • 1 ♀; Lake Co., Booth State Park; 14 mi. W of Lakeview; [42.197 ° N, 120.591 ° W]; 18 Jun. 1952; V. Roth leg.; OSAC • 1 ♀; Lane Co., Oakridge; 43.881 ° N, 122.225 ° W; elev. 701 m; 23 Aug. 2019; L. Humphreys leg.; WRME • 1 ♀; Linn Co.; [44.63 ° N, 123.09 ° W]; 12 Jul. 1929; J. E. Davis leg.; OSAC • 1 ♀; Harney Co., Malheur NWR, Tower; 43.2365 ° N, 118.8421 ° W; 4 – 17 Jun. 2011; T. Coe leg.; OSAC • 1 ♀; Sherman Co., Fulton Canyon; 45.6345 ° N, 120.86424 ° W; 10 May 2011; J. D. Engler leg.; OSAC • 1 ♀; Umatilla Co., Cayuse; [45.676 ° N, 118.574 ° W]; 13 May 1938; J. Schuh leg.; OSAC • 1 ♀; Union Co., La Grange, NF- 5176; 45.1451 ° N, 118.3709 ° W; 22 Jun. 2018; G. Pederson leg.; WRME • 1 ♀; Union Co., La Grange, HWY 244; 45.2737 ° N, 118.3437 ° W; 22 Jun. 2018; G. Pederson leg.; OSAC • 1 ♀; Union Co., Union; [45.21 ° N, 117.8647 ° W]; elev. 827.8 m; 6 Sep. 1926; H. A. Scullen leg.; OSAC • 1 ♀; Wallowa Co., Wallowa Canyon Fount; 5 mi. E of Minam; [45.617 ° N, 117.552 ° W]; elev. 823 m; 21 Jul. 1929; H. A. Scullen leg.; OSAC • 1 ♀; Washington Co., Beaverton; 45.450 ° N, 122.874 ° W; elev. 205 m; 28 Apr. 2021; S. Albright leg.; WRME • 1 ♀; Yamhill Co., McMinnville; 45.211 ° N, 123.174 ° W; elev. 33 m; 12 Apr. 2019; M. O’Loughlin leg.; WRME. – Washington • 1 ♂; Asotin Co., Fields Spring State Park, 4 mi. of S of Anatone; [46.0814 ° N, 117.171 ° W]; elev. 1097.3 m; 5 Jul. 1984; W. Turner leg.; WSU • 1 ♂; Clark Co., Vancouver; [45.63 ° N, 122.67 ° W]; 10 Jul. 1970; WSU • 1 ♂; Clark Co., Vancouver; [45.631 ° N, 122.675 ° W]; 10 Jul. 1970; WSU • 1 ♀; Island Co., Whidbey Island, Fort Casey; [48.164 ° N, 122.679 ° W]; 18 Jun. 1980; D. F. Frechin leg.; WSU • 1 ♀; King Co., Seattle; [47.61 ° N, 122.33 ° W]; UCMC • 1 ♀; same data as for preceding; OSAC • 1 ♂; Spokane Co., Dartford; [47.78 ° N, 117.42 ° W]; 29 – 31 Jul. 1969; R. D. Gray leg.; WSU • 1 ♀; Thurston Co., Olympia; [47.0451 ° N, 122.895 ° W]; UCMC • 1 ♀; Whitman Co., Pullman; [46.73 ° N, 117.17 ° W]; 7 May 1969; R. W. Dawson leg.; WSU • 2 ♂♂; same data as for preceding; 13 Jul. 1901; WSU • 2 ♀♀; same data as for preceding; 27 May 1965; R. D. Akre leg.; WSU • 1 ♀; same data as for preceding; 26 May 1965; R. D. Akre leg.; WSU • 1 ♀; same data as for preceding; 11 May 1965; R. D. Akre leg.; WSU • 1 ♂; same data as for preceding; 16 Aug. 1965; R. D. Akre leg.; WSU • 1 ♀; Whitman Co., Almota; [46.703 ° N, 117.467 ° W]; 6 May 1972; W. J. Turner leg.; WSU • 1 ♀; same data as for preceding; 27 Apr. 1972; W. J. Turner leg.; WSU • 1 ♀; Whitman Co., Almota; [46.703 ° N, 117.467 ° W]; 28 Jun. 1975; J. Powell leg.; EMEC • 1 ♀; Whitman Co., 10 mi. SW of Colton, Steptoe Canyon; [46.485 ° N, 117.301 ° W]; 3 Aug. 1974; W. J. Turner leg.; WSU. – Wyoming • 1 ♀; Teton Co., Jackson Hole Research Station, Grand Teton National Park; [43.833 ° N, 110.701 ° W]; elev. 2042.2 m; 11 Jul. 1967; H. E. Evans and M. A. Evans leg.; UCRC.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFCCFFF9FDDFF936FC51FEA4.taxon	description	Description Female BODY MEASUREMENTS. Length 6.73 mm (± 0.63 SD). Head long 1.72 mm (± 0.06 SD). Face length 1.41 mm (± 0.05 SD). Head width 1.64 mm (± 0.05 SD). Wing length 4.97 mm (± 0.30 SD). n = 6. COLOURATION. Head and mesosoma black. Clypeus apex black to dark brown. Labrum brown. Mandible dark brown basally, becoming light brown medially and reddish brown apically. Flagellum dark brown to brown dorsally, light brown ventrally. Pronotal lobe brown. Tegula light brown, with brown base. Wing membrane faintly brown, veins brown. Legs brown to light brown. Metasoma black to dark brown with rims of terga and sterna broadly light brown. PUBESCENCE. Body hair colour off-white, but more yellowish. Tomentum dense on pronotal angle, and lobe, space between pronotal lobe and tegula, metanotum anteromedially, and T 2 – T 3 basolaterally. Mesosoma covered by sub-appressed to erect, simple to weakly plumose, short hairs (0.5 – 1 OD) on mesoscutum and erect, plumose, long hairs (<1.5 OD) on other areas. Metafemoral scopa with dense plumose hairs. Wing hairs brown, very short and sparse. SURFACE SCULPTURING. Clypeus polished, becoming weakly imbricate in the basal third; punctures irregularly sparse (IS ≤ 2 PD) apically, becoming smaller and denser (IS ≤ 1 PD) basally. Supraclypeal area imbricate, becoming weakly so or weakly polished medially; punctures sparse (IS = 1 – 3 PD) anteromedially, becoming dense (IS ≤ 1 PD) posterolaterally. Frons reticulate-punctate; punctures crowded (IS = 0 PD). Ocellocular area polished; punctures moderately sparse (IS = 1 – 2 PD) anterolaterally, diversopunctate. Gena polished, becoming lineolate posteriorly; punctures dense (IS ≤ 1 PD), becoming obscure posteriorly. Postgena lineolate. Tegula broadly impunctate, minute punctures present anteriorly and basally, moderately sparse (IS = 1 – 2 PD). Mesoscutum polished, becoming imbricate anteriorly and medially halfway on median line; punctures dense (IS ≤ 1 PD), becoming moderately dense (IS = 1 – 2 PD) mesad of parapsidal line and anteromedially. Mesoscutellum polished; punctures dense (IS ≤ 1 PD) marginally and on median line, sparse submedially (IS = 1 – 4 PD). Metanotum rugulose. Propodeal triangle with rugae moderately strong, reaching ¾ towards posterior margin, weakly anastomosing medially, lateral rugae extending to dorsolateral slope of propodeum lateral face; interspace sculpture imbricate to tessellate. Preëpisternum weakly rugulose, punctures crowded (IS = 0 PD). Hypoepimeron polished, becoming rugulose dorsally; punctures dense (IS ≤ 1 PD). Mesepisternum polished, but weakly rugulose from some angles; punctures sparse (IS = 1 – 3 PD), doubly punctate, but not that conspicuous. Propodeum lateral face strongly imbricate, punctures obscure; posterior face imbricate-rugulose, but weakly strigate, punctures obscure. T 1 anterior face polished, punctures present laterally, sparse (IS = 1 – 6 PD); dorsal surface polished, punctures present laterally, sparse (IS = 1 – 3 PD), becoming sparser (IS = 1 – 6 PD) and fine towards premarginal line and nearly impunctate medially and in small subapicolateral boss, punctures fine, sparse (IS = 3 – 6 PD) on rim. T 2 disc polished, punctures moderately sparse (IS = 1 – 2 PD) basally becoming sparse (IS = 1 – 4 PD) apically; rim weakly coriarious, punctures fine, sparse (IS = 1 – 4 PD). STRUCTURE. Head long, length / width ratio 1.05 (± 0.03 SD). Face length / width ratio 0.86 (± 0.01 SD). Clypeus projecting ~ 70 % below suborbital tangent. Mandible short, reaching opposing clypeal angle. Eyes convergent below [UOD / LOD ratio 1.20 (± 0.03 SD)]. Ocelli normal. Gena / eye width ratio 0.76 (± 0.05 SD). Pronotal angle obtuse. Intertegular span 1.20 mm (± 0.05 SD). Mesoscutum length / width ratio 0.93 (± 0.06 SD); mesoscutum / scutellum length ratio 2.69 (± 0.23 SD); scutellum / metanotum length ratio 1.85 (± 0.16 SD); metanotum / propodeal triangle length ratio 0.76 (± 0.09 SD). Protibial spur with apical serrations shorter than width of malus; malus extending beyond the velum, nearly ¾ the length of velum. Inner metatibial spur pectinate, teeth 3 – 5, basal teeth somewhat longer than width of rachis. Propodeal triangle posterior margin rounded onto posterior propodeal surface. Propodeal lateral carina nearly reaching dorsolateral slope, oblique carina obscure. Male BODY MEASUREMENTS. Length 5.13 mm (± 0.46 SD). Head length 1.56 mm (± 0.03 SD). Face length 1.26 mm (± 0.03 SD). Head width 1.46 mm (± 0.04 SD). Wing length 4.01 mm (± 0.13 SD). n = 4. COLOURATION. Head and mesosoma black. Clypeus apical half light brown to yellow. Labrum light brown. Mandible dark brown basally, becoming yellow medially and reddish brown apically. Flagellum light brown to brown. Pronotal lobe brown. Tegula light brown, with brown base. Wing membrane faintly brown, veins brown. Legs brown, except tarsi, and protibia light brown. Metasoma dark brown with rims of terga and sterna narrowly light brown. PUBESCENCE. Body hair colour off-white, but more yellowish. Tomentum dense on pronotal angle, and lobe, space between pronotal lobe and tegula; moderately dense on supraclypeal area laterally, and paraocular area; very sparse on mesepisternum ventrally, and T 1 basolaterally. Mesosoma covered by erect, mostly simple, short hairs (0.25 – 1 OD) on mesoscutum and erect, plumose, long hairs (<2 OD) on other areas. Metasomal sterna with dense, erect and sub-appressed, plumose, short (<1.5 OD) hairs. Wing hairs brown, very short and sparse. SURFACE SCULPTURING. Clypeus polished apical; punctures irregularly sparse (IS ≤ 2 PD) apically, becoming smaller and denser (IS ≤ 1 PD) basally. Supraclypeal area polished; punctures dense (IS ≤ 1 PD). Frons reticulate-rugose, becoming polished dorsally; punctures crowded (IS <0 PD). Ocellocular area polished; punctures dense (IS ≤ 1 PD). Gena polished; punctures moderately dense (IS = 1 – 2 PD). Postgena lineolate. Tegula broadly impunctate, punctures present anteriorly and basally, moderately sparse (IS = 1 – 2 PD). Mesoscutum mostly polished, imbricate near anteriorly; punctures dense (IS ≤ 1 PD), becoming moderately dense (IS = 1 – 2 PD) mesad of parapsidal line. Mesoscutellum polished; punctures dense (IS ≤ 1 PD) marginally and on median line, sparse submedially (IS = 1 – 4 PD). Metanotum rugulose. Propodeal triangle with rugae moderately strong, extending about halfway toward posterior margin, weakly anastomosing medially, lateral rugae extending to dorsolateral slope of propodeum lateral face; interspace sculpture imbricate to tessellate. Preëpisternum weakly rugulose. Hypoepimeron polished, becoming rugulose dorsally; punctures dense (IS ≤ 1 PD). Mesepisternum polished; punctures dense (IS ≤ 1 PD) dorsally, becoming sparse (IS = 1 – 3 PD) ventrally, diversly punctate ventrally with minute punctures. Propodeum lateral face strongly imbricate, punctures obscure; posterior face imbricate-rugulose, but weakly strigate, punctures obscure. T 1 anterior face polished, few minute punctures present laterally; dorsal surface polished, few minute punctures present laterally, sparse (IS = 3 – 6 PD), becoming nearly impunctate medially, in subapicolateral boss and on rim. T 2 disc polished, punctures sparse (IS = 1 – 5 PD); rim polished, impunctate. STRUCTURE. Head long, length / width ratio 1.06 (± 0.02 SD). Face length / width ratio 0.86 (± 0.03 SD). Clypeus projecting ~ 63 % below suborbital tangent. Mandible short, reaching opposing clypeal angle. Flagellomeres short, F 1 length / pedicel length ratio 0.96 (± 0.08 SD); F 2 length / F 1 length ratio 1.22 (± 0.14 SD); F 2 length / F 2 width ratio 1.00 (± 0.03 SD). Gena / eye width ratio 0.69 (± 0.08 SD). Forewing with 3 submarginal cells. Pronotal angle obtuse. Intertegular span 0.91 mm (± 0.02 SD). Mesoscutum length / width ratio 0.91 (± 0.06 SD). Propodeal lateral carina nearly reaching dorsolateral slope; oblique carina absent. n = 4. GENITALIA (Figs 33, 120 E). Gonostylus elongate, about 2 times as long as broad, and slightly concave on outer margin, with short setae. Retrorse lobe attenuated apically, about 3 times as long as broad, covered in short setae.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFCCFFF9FDDFF936FC51FEA4.taxon	distribution	Range Cascadia bioregion, California, Oregon and Washington to Alberta, New Mexico, Wyoming (Fig. 34).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFCCFFF9FDDFF936FC51FEA4.taxon	biology_ecology	Floral hosts ROSACEAE: Pyrus sp. • RANUNCULACEAE: Ranunculus sp.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFCCFFF9FDDFF936FC51FEA4.taxon	discussion	DNA barcodes DNA barcodes provide additional support for recognizing L. engleri sp. nov. as distinct from L. macoupinense. Minimum interspecific sequence divergence between the two species exceeds 1.5 %, and mean interspecific sequence divergence exceeds 3.8 %. The level of barcode divergence is comparable to other closely related species of Lasioglossum.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFC4FF83FE66FE87FD21F929.taxon	description	Figs 35 – 38, 93 F, 101 A – B, 109 F, 110 B, 120 F	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFC4FF83FE66FE87FD21F929.taxon	diagnosis	Diagnosis Both sexes of L. glabriventre can be recognized by the following combination of characters: tegula enlarged (mostly broadly punctate) and mesepisternum smooth, distinctly punctate. Lasioglossum kincaidii has an enlarged tegula, but the mesepisternum is coarsely rugose.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFC4FF83FE66FE87FD21F929.taxon	etymology	Etymology Crawford (1907 b) did not explain the epithet. The specific epithet is interpreted as a compound Latin adjective based on the roots ‘ glaber ’ meaning ‘ bald, smooth’ and ‘ venter ’ referring to ‘ the belly’.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFC4FF83FE66FE87FD21F929.taxon	materials_examined	Type material Holotype of Halictus Vachali (by monotypy; examined by photos) UNITED STATES – Oregon • ♀; Corvallis; [44.57 ° N, 123.28 ° W]; 6 Jun. 1898; USNM 29424. Specimen labels “ Corvallis / 6 - 6 - 98. Or // 26 // Type No. / 29424 / U. S. N. M. // HOLOTYPE // USNMENT / 01735684 ”. Type remarks The original description states it was based on a single female, so a holotype is designated by monotypy. Other material examined CANADA – Alberta • 1 ♀; Newell Co., Dinosaur Provincial Park, Sandhill Creek Valley; 50.755833 ° N, 111.519167 ° W; 14 Jul. 2010; M. Buck leg.; PMAE • 1 ♀; Oldman River near Kipp, Sandy floodplain; 49.740556 ° N, 112.990278 ° W; 4 Jul. 2020; M. Buck leg.; PMAE • 1 ♀; Forty Mile Co., Pinhorn Provincial Grazing Reserve, Milk River 3 km E of HQ; 49.127222 ° N, 110.854167 ° W; 10 Aug. 2017; M. Buck leg.; PMAE • 1 ♀; Redcliff; 50.054722 ° N, 110.786667 ° W; 28 Jun. 2017; M. Buck leg.; PMAE • 1 ♀; Writing-On-Stone Provincial Park; [49.09 ° N, 111.62 ° W]; 9 – 13 Jul. 1990; M. Klassen leg.; PMAE • 1 ♀; same data as for preceding; 6 – 12 Jun. 1990; D. McCorquodale leg.; PMAE • 1 ♀; same data as for preceding; 26 May – 6 Jun. 1990; D. McCorquodale leg.; PMAE • 1 ♀; Writing-On-Stone Provincial Park, 6 km E at bridge; [49.089 ° N, 111.536 ° W]; 13 Jul. 1982; D. E. McCorquodale leg.; PMAE • 1 ♀; Townshi Rd 102, Cranford; 49.81496 ° N, 112.2562 ° W; elev. 767 m; 1 – 15 Jul. 2016; S. Robinson leg.; PMAE. – British Columbia • 1 ♀; Blind Bay; [50.886 ° N, 119.398 ° W]; 1 – 30 Jun. 1987; C. A. Elsey leg.; PMAE • 1 ♀; Kalamalka Lake Provincial Park; 50.20773 ° N, 119.2342 ° W; 27 – 29 May 2015; D. Marks leg.; RSKM • 1 ♀; Okan, Similk; 49.17648 ° N, 119.76225 ° W; 25 Sep. – 17 Oct. 2015; C. Dawson leg.; RSKM • 2 ♂♂; Okanagan Mountain Provincial Park; 49.76453 ° N, 119.59564 ° W; 28 Jul. 2016; M. Sanelli leg.; RSKM • 5 ♀♀; Okanagan-Similkameen Reg. Dist., Oliver, Ripley Lake; 49.2435 ° N, 119.6324 ° W; elev. 925 m; 13 Jul. 2008; L. R. Best leg.; PCYU • 1 ♀; Okanagan-Similkameen Reg. Dist., Oliver, Ripley Lake; 49.2429 ° N, 119.6338 ° W; elev. 925 m; 11 Jul. 2008; L. R. Best leg.; PCYU • 1 ♀; Okanagan-Similkameen Reg. Dist., Oliver, Ripley Lake; 49.2432 ° N, 119.6329 ° W; elev. 925 m; 11 Jul. 2008; L. R. Best leg.; PCYU • 3 ♀♀; Okanagan-Similkameen Reg. Dist., Oliver, Ripley Lake; 49.2432 ° N, 119.6332 ° W; elev. 925 m; 16 Jul. 2008; L. R. Best leg.; PCYU • 9 ♀♀; Okanagan-Similkameen Reg. Dist., Oliver, Ripley Lake; 49.2432 ° N, 119.6329 ° W; elev. 925 m; 12 Jul. 2008; L. R. Best leg.; PCYU • 1 ♀; Okanagan-Similkameen Reg. Dist., Oliver, Sawmill Lake; 49.2072 ° N, 119.614 ° W; elev. 740 m; 29 Jul. 2008; L. R. Best leg.; PCYU • 1 ♀; Okanagan-Similkameen Reg. Dist., Mt Oliver, Ripley Lake; 49.2429 ° N, 119.6326 ° W; elev. 935 m; 31 Jul. – 3 Aug. 2009; L. R. Best leg.; PCYU • 1 ♀; North Okanagan Reg. Dist. Lumby, Twin Creeks; 50.2499 ° N, 118.9593 ° W; elev. 500 m; 26 May 2008; L. R. Best leg.; PCYU • 1 ♀; Osoyoos; [49.0251 ° N, 119.4376 ° W]; 12 Jul. – 10 Aug. 2007; Scudder leg.; RSKM • 1 ♀; Osoyoos, Osoyoos West Bench; 49.03033 ° N, 119.50034 ° W; 17 Jul. – 9 Aug. 2017; D. Marks and K. Grant (J. Heron) leg.; RSKM • 1 ♀; Osoyoos, Strawberry Creek Road; 49.038471 ° N, 119.508261 ° W; 28 Jun. 2020; T. Haapalainen and K. Needham leg.; UBCZ • 1 ♀; Penticton; [49.500 ° N, 119.593 ° W]; 5 Jun. 2019; E. R. Buckell leg.; UBCZ • 2 ♂♂; Shuswap Lake, Blind Bay; [50.886 ° N, 119.398 ° W]; 31 Aug. 1986; C. A. Elsey leg.; PMAE • 1 ♀; Vernon Military Camp-DND; 50.2302 ° N, 119.3004 ° W; 6 – 22 May 2018; T. Ehlers and I. Zwart leg.; RSKM • 17 ♀♀; Vernon, CFB Vernon; 50.226 ° N, 119.302 ° W; 1 – 30 Jun. 2018; T. Ehlers leg.; RSKM • 5 ♀♀; Vernon, Kalamalka Lk PP; 50.20961 ° N, 119.2329 ° W; 29 Jun. – 2 Jul. 2017; D. Marks, K. Grant and J. Heron leg.; RSKM • 5 ♀♀; Vernon, Vernon Military Camp-DND; 50.2255 ° N, 119.306 ° W; 6 – 22 May 2018; T. Ehlers and I. Zwart leg.; RSKM • 2 ♀♀; Vernon, Vernon Military Camp – DND; 50.2252 ° N, 119.3085 ° W; 6 – 22 May 2018; T. Ehlers and I. Zwart leg.; RSKM • 1 ♀; Vernon, Vernon Military CampDND; 50.2256 ° N, 119.3084 ° W; 6 – 22 May 2018; T. Ehlers and I. Zwart leg.; RSKM. – Saskatchewan • 1 ♀; Leader; 50.68337 ° N, 109.2231 ° W; S. Stecyk and S. MacDonald leg.; RSKM • 1 ♀; Maple Creek Golf Course; 49.9021 ° N, 109.4654 ° W; 9 Aug. 2018; RSKM. UNITED STATES – California • 1 ♀; El Dorado Co., Fallen Leaf; [38.883 ° N, 120.073 ° W]; 14 Aug. 1931; O. H. Swezey leg.; UCRC • 2 ♀♀; Butte Co., 6.2 mi. NW of Jarbo Pass; [39.801 ° N, 121.580 ° W]; 1 Jul. 1963; C. A. Toschi leg.; ex Clarkia garcilis albicaulis; EMEC • 1 ♀; Contra Costa Co., Bethel Harbor; [38.031 ° N, 121.641 ° W]; 12 Sep. 1956; J. Powell leg.; EMEC • 1 ♀; El Dorado Co., 3 mi. S of Camino; [38.694 ° N, 120.674 ° W]; 23 Jun. 1948; K. W. Tucker leg.; ex Compositae; EMEC • 1 ♀; El Dorado Co., Pollock Hill; [38.76 ° N, 120.59 ° W]; 21 Jun. 1953; G. D. Butler leg.; ex Chamaebatiaria millefolium; UCRC • 2 ♀♀; El Dorado Co., Riverton; [38.771 ° N, 120.449 ° W]; 18 Jun. 1968; S. Paul Welles leg.; ex Clarkia rhomboidea; EMEC • 1 ♀; same data as for preceding; 18 Jun. 1968; S. Paul Welles leg.; ex Gayophytum; EMEC • 4 ♀♀; same data as for preceding; 11 Jun. 1968; S. Paul Welles leg.; ex Clarkia rhomboidea; EMEC • 1 ♀; same data as for preceding; 11 Jun. 1968; S. Paul Welles leg.; ex Clarkia rhomboidea; UCRC • 1 ♀; El Dorado Co., Snowline Camp; [38.747 ° N, 120.624 ° W]; 20 Jul. 1948; P. D. Hurd leg.; EMEC • 1 ♀; Fresno Co., 2 mi. W of Pine Ridge; [37.063 ° N, 119.397 ° W]; elev. 1219.2 m; 19 Jul. 1960; R. R. Snelling leg.; ex Eriogonum; LACM • 1 ♀; Fresno Co., Dinkey Creek; [37.086 ° N, 119.157 ° W]; 8 Aug. 1956; R. O. Schuster leg.; EMEC • 1 ♀; Fresno Co., Pinehurst; [36.695 ° N, 119.016 ° W]; elev. 1249.7 m; 24 Jun. 1960; R. R. Snelling leg.; LACM • 2 ♀♀; Fresno Co., Prather; [37.038 ° N, 119.514 ° W]; 29 Jun. 1956; R. O. Schuster leg.; EMEC • 1 ♂; same data as for preceding; 29 Jun. 1956; R. O. Schuster leg.; EMEC • 1 ♀; Fresno Co., Shaver Lake; [37.089 ° N, 119.318 ° W]; 27 Jul. 1968; J. B. Hoy leg.; EMEC • 1 ♀; same data as for preceding; 13 Jul. 1968; J. B. Hoy leg.; EMEC • 2 ♂♂; same data as for preceding; 6 Jul. 1968; J. B. Hoy leg.; EMEC • 3 ♀♀; same data as for preceding; 8 Aug. 1956; R. O. Schuster leg.; EMEC • 1 ♂; Fresno Co., Watts Valley; [36.934 ° N, 119.358 ° W]; 22 Jun. 1956; R. O. Schuster leg.; EMEC • 1 ♀; Humboldt Co., Redwood Creek at Stover Rd, Redwood Valley, N of Hwy 299; [40.97 ° N, 123.85 ° W]; elev. 198.1 m; 10 Aug. 1966; H. B. Leech leg.; CAS • 1 ♀; Inyo Co., Antelope Springs; [37.331 ° N, 118.087 ° W]; 15 Jun. 1961; C. A. Toschi leg.; ex Cleome lutea; EMEC • 1 ♀; Inyo Co., Lone Pine; [36.61 ° N, 118.06 ° W]; 13 May 1969; P. Welles leg.; ex Aster; EMEC • 3 ♀♀; Lassen Co., Hallelujah Junction; [39.775 ° N, 120.039 ° W]; 22 Jun. 1964; M. E. Irwin leg.; UCRC • 1 ♀; Lassen Co.; [40.67 ° N, 120.59 ° W]; 3 Jun. 1913; F. W. Nunenmacher leg.; EMEC • 1 ♀; same data as for preceding; 4 Jun. 2013; EMEC • 1 ♀; Los Angeles Co., Arroyo Seco, Pasadena; [34.118 ° N, 118.169 ° W]; 9 Aug. 2017; R. May leg.; EMEC • 1 ♂; Los Angeles Co., Claremont; [34.097 ° N, 117.719 ° W]; Baker leg.; USNM • 1 ♀; Los Angeles Co., El Segundo Dunes; [33.932 ° N, 118.434 ° W]; 30 Apr. 1939; J. A. Comatook leg.; ex Eremalche exilis (verbatim: Malvastrum exile); LACM • 1 ♀; Los Angeles Co., Green Valley, 4 mi. S of Cuyamaca; [32.906 ° N, 116.582 ° W]; 8 Aug. 1951; P. H. Timberlake leg.; ex Eriogonum davidsonii; UCRC • 1 ♀; Los Angeles Co., Los Angeles; [34.054 ° N, 118.243 ° W]; 21 Apr. 1918; R. May leg.; EMEC • 1 ♀; Los Angeles Co., Mts near Claremont; [34.161 ° N, 117.716 ° W]; Baker leg.; USNM • 1 ♀; Los Angeles Co., Newhall; [34.595 ° N, 118.662 ° W]; 20 Apr. 1940; R. M. Bohart leg.; LACM • 1 ♀; Los Angeles Co., Tanbark Flat; [34.204 ° N, 117.760 ° W]; 22 Jun. 1950; P. D. Hurd leg.; EMEC • 1 ♀; Madera Co., 2 mi. N of Nipinnawasee; [37.433 ° N, 119.733 ° W]; 4 Jul. 1960; G. I. Stage and R. R. Snelling leg.; ex Clarkia williamsonii; EMEC • 2 ♂♂; Madera Co., Chiquito Creek, 0.25 mi. below Upper Chiquito Campground; [37.498 ° N, 119.409 ° W]; elev. 2078.7 m; 11 Aug. 1971; H. B. Leech leg.; CAS • 1 ♀; Madera Co., Oakhurst; [37.328 ° N, 119.649 ° W]; 1 Jun. 1942; EMEC • 1 ♀; Madera Co., Oakhurst cemetery; [37.333 ° N, 119.653 ° W]; 16 May 1963; J. W. MacSwain leg.; ex Oenothera campestris; EMEC • 2 ♀♀; Mariposa Co., 0.8 mi. S of Mormon Bar; [37.4503 ° N, 119.9479 ° W]; 16 Jun. 1959; R. W. Thorp leg.; ex Clarkia williamsonii; EMEC • 3 ♀♀; same data as for preceding; 16 Jun. 1959; G. I. Stage leg.; ex Clarkia williamsonii; EMEC • 1 ♀; same data as for preceding; 16 Jun. 1959; G. I. Stage leg.; ex Clarkia dudleyana; EMEC • 1 ♀; Mariposa Co., 10 mi. E of Mormon Bar; [37.462 ° N, 119.766 ° W]; 4 Jul. 1960; G. I. Stage and R. R. Snelling leg.; ex Clarkia williamsonii; EMEC • 1 ♀; Mariposa Co., 5.5 mi. N of Coulterville; [37.788 ° N, 120.197 ° W]; 18 Jul. 1963; R. W. Thorp leg.; ex Clarkia dudleyana; EMEC • 1 ♀; Mariposa Co., Yosemite; [37.833 ° N, 119.500 ° W]; 20 May 1934; O. Bryant leg.; CAS • 1 ♀; same data as for preceding; 25 May 1938; N. F. Hardman leg.; EMEC • 1 ♀; Mariposa Co.; [37.57 ° N, 119.90 ° W]; 23 May 1938; N. F. Hardman leg.; EMEC • 1 ♀; Modera Co.; [36.99 ° N, 120.01 ° W]; 21 May 1938; N. F. Hardman leg.; EMEC • 2 ♀♀; Modoc Co., 5 mi. NE of Alturas; [41.538 ° N, 120.474 ° W]; 29 May 1976; T. R. Haig leg.; CSCA • 1 ♀; Modoc Co., 9 mi. N of Adin; [41.325 ° N, 120.944 ° W]; elev. 1493.5 m; 5 Aug. 1966; UCRC • 2 ♀♀; Modoc Co., Davis Creek; [41.733 ° N, 120.372 ° W]; 14 Jul. 1922; C. L. Fox leg.; CAS • 2 ♀♀; Modoc Co., Lake City; [41.643 ° N, 120.217 ° W]; 30 Jul. 1922; C. L. Fox leg.; CAS • 1 ♀; Modoc Co., Adin Summit, 10 mi. S of Canby; [41.299 ° N, 120.871 ° W]; 13 Jun. 1964; C. W. Baker and D. R. Smith leg.; EMEC • 1 ♀; Mono Co., 7 mi. E of Bodie; [38.212 ° N, 118.885 ° W]; 2 Jul. 1964; P. Rude leg.; EMEC • 1 ♂; Mono Co., Leavitt Meadow; [38.316 ° N, 119.550 ° W]; elev. 2194.6 m; 13 Aug. 1963; H. B. Leech leg.; CAS • 1 ♀; same data as for preceding; 13 Aug. 1963; H. B. Leech leg.; CAS • 1 ♂; same data as for preceding; 11 Aug. 1960; C. A. Toschi leg.; EMEC • 1 ♀; Monterey Co., Indians Guard Staion, 17 air mi. NW of Jolon; [36.144 ° N, 121.392 ° W]; elev. 640 m; 9 May 1975; R. Wharton leg.; EMEC • 1 ♀; Monterey Co., Tassajara Hot Springs; [36.234 ° N, 121.550 ° W]; elev. 457.2 m; 26 May 1954; O. Bryant leg.; CAS • 1 ♀; Napa Co., Chiles Valley; [38.625 ° N, 122.371 ° W]; 7 May 1939; E. C. Van Dyke leg.; CAS • 1 ♂; Nevada Co., 3.5 mi. N of Truckee; [39.378 ° N, 120.183 ° W]; 8 Aug. 1963; J. W. MacSwain leg.; ex Gayophytum heterozygum; EMEC • 1 ♀; Nevada Co., 4 mi. N of Truckee; [39.386 ° N, 120.183 ° W]; 5 Sep. 1992; E. G. Linsley leg.; ex Chrysothamnus viscidiflorus var. typicus; EMEC • 1 ♀; Nevada Co., 7 mi. SE of Truckee; [39.254 ° N, 120.091 ° W]; 24 Jun. 1954; R. H. Goodwin leg.; EMEC • 1 ♀; Nevada Co., Boca; [39.387 ° N, 120.090 ° W]; 3 Jul. 1954; P. D. Hurd leg.; ex Chaenactis glabriuscula var. lanosa; EMEC • 1 ♀; Nevada Co., near Hobart Mills; [39.400 ° N, 120.185 ° W]; 17 Jun. 1964; M. E. Irwin leg.; UCRC • 1 ♀; Nevada Co., Sagehen near Hobart Mills; [39.43 ° N, 120.20 ° W]; 21 Jun. 1954; P. D. Hurd leg.; ex Phacelia humilis; EMEC • 1 ♀; same data as for preceding; 25 Jun. 1954; J. A. Powell leg.; EMEC • 1 ♀; same data as for preceding; 9 Jul. 1954; R. H. Goodwin leg.; EMEC • 1 ♀; same data as for preceding; 4 Jul. 1954; P. D. Hurd leg.; EMEC • 2 ♀♀; Placer Co., Summit; [39.313 ° N, 120.359 ° W]; elev. 2133.6 m; W. M. Giflard leg.; CAS • 2 ♀♀; Placer Co., Lake Tahoe, Carnelian Bay; [39.227 ° N, 120.082 ° W]; 24 Jun. 1954; J. A. Powell leg.; EMEC • 3 ♀♀; Plumas Co., 23 mi. SW of Quincy; [39.700 ° N, 121.254 ° W]; 2 Jul. 1963; J. W. MacSwain leg.; ex Clarkia purpurea; EMEC • 1 ♀; Plumas Co., 5.5 mi S of Vinton; [39.722 ° N, 120.175 ° W]; 6 Aug. 1963; J. W. MacSwain leg.; ex Taraxia tanacetifolia (verbatim: Oenothera tanacetifolia); EMEC • 1 ♀; Plumas Co., Bucks Lake; [39.871 ° N, 121.184 ° W]; 1 Jul. 1949; J. W. MacSwain leg.; EMEC • 1 ♂; same data as for preceding; 13 Aug. 1959; J. W. MacSwain leg.; ex Gayophytum humile (verbatim: Gayophytum nuttallii); EMEC • 1 ♀; same data as for preceding; 13 Aug. 1959; J. W. MacSwain leg.; ex Gayophytum humile (verbatim: Gayophytum nuttallii); EMEC • 1 ♀; Plumas Co., Portola; [39.811 ° N, 120.469 ° W]; 24 Aug. 1976; T. R. Haig leg.; CSCA • 1 ♀; Riverside Co., 1 mi. S of Temecula; [33.477 ° N, 117.149 ° W]; 2 Jul. 1963; J. Powell leg.; EMEC • 1 ♀; same data as for preceding; 15 Jul. 1963; J. Powell leg.; EMEC • 1 ♀; Riverside Co., Bautista Canyon, T 65, R 2 E, S 18; [33.70 ° N, 116.85 ° W]; 24 Apr. 1984; F. Corcel leg.; UCRC • 1 ♀; Riverside Co., Bautista Canyon, San Bernardino National Forest (S. Valle Vista); [33.70 ° N, 116.85 ° W]; 23 May 1968; S. Frommer and R. Somerby leg.; UCRC • 1 ♀; Riverside Co., Hemet; [33.748 ° N, 116.972 ° W]; 15 Jul. 1945; Ray F. Smith leg.; EMEC • 2 ♂♂; same data as for preceding; 11 – 19 Aug. 1946; C. Grainger leg.; ex Goldenrod; EMEC • 1 ♀; Riverside Co., Idyllwild; [33.748 ° N, 116.715 ° W]; 25 Jun. 1928; E. C. Van Dyke leg.; CAS • 1 ♀; Riverside Co., Idyllwild, San Jacinto Mountains; [33.748 ° N, 116.715 ° W]; 12 Jun. 1939; J. G. Shanafelt leg.; LACM • 1 ♀; same data as for preceding; 20 May 1939; E. G. Linsley leg.; EMEC • 4 ♀♀; Riverside Co., Keen Camp, San Jacinto Mountains; [33.687 ° N, 116.704 ° W]; 10 Jun. 1939; E. S. Ross leg.; EMEC • 1 ♀; same data as for preceding; 10 Jun. 1939; E. G. Linsley leg.; EMEC • 1 ♀; Riverside Co., Pine Flat, San Jacinto Mountains; [33.585 ° N, 116.456 ° W]; 15 Jun. 1946; C. D. Michener leg.; ex Cryptantha micrantha; EMEC • 1 ♀; Riverside Co., Riverside; [33.956 ° N, 117.372 ° W]; elev. 290 m; 12 May 1926; P. H. Timberlake leg.; ex (?); UCRC • 1 ♀; same data as for preceding; 17 Apr. 1925; P. H. Timberlake leg.; ex Eschscholzia californica Cham; UCRC • 1 ♀; same data as for preceding; 10 Apr. 1928; P. H. Timberlake leg.; ex Gilia achilleifolia Benth. subsp. multicaulis (verbatim: Gilia multicaulis); UCRC • 1 ♀; same data as for preceding; 10 Apr. 1925; P. H. Timberlake leg.; ex Heterotheca grandiflora; UCRC • 1 ♀; same data as for preceding; 17 Mar. 1925; P. H. Timberlake leg.; ex Eschscholzia; UCRC • 1 ♀; same data as for preceding; 5 Mar. 1926; P. H. Timberlake leg.; ex Cryptantha intermedia; UCRC • 1 ♀; same data as for preceding; 15 Feb. 1935; P. H. Timberlake leg.; ex Clarkia (verbatim: Godetia); UCRC • 1 ♀; same data as for preceding; 24 Apr. 1930; P. H. Timberlake leg.; ex Clarkia purpurea subsp. viminea (verbatim: Godetia viminea); UCRC • 1 ♀; same data as for preceding; 6 Oct. 1935; P. H. Timberlake leg.; ex Polygonum lapathifolium; UCRC • 1 ♀; same data as for preceding; 6 May 1929; P. H. Timberlake leg.; ex Gilia achilleifolia subsp. multicaulis (verbatim: Gilia multicaulis); UCRC • 1 ♀; same data as for preceding; 7 Jul. 1929; P. H. Timberlake leg.; ex Portulaca; UCRC (Fig. 35) • 1 ♀; same data as for preceding; 10 May 1929; P. H. Timberlake leg.; ex Gilia achilleifolia subsp. multicaulis (verbatim: Gilia multicaulis); UCRC • 1 ♀; same data as for preceding; 7 Apr. 1936; P. H. Timberlake leg.; ex Gilia achilleifolia subsp. multicaulis (verbatim: Gilia multicaulis); UCRC • 1 ♀; same data as for preceding; 25 Apr. 1928; P. H. Timberlake leg.; ex Cryptantha intermedia; UCRC • 1 ♀; same data as for preceding; 28 Apr. 1938; P. H. Timberlake leg.; ex Chaenactis glabriuscula; UCRC • 1 ♂; same data as for preceding; 4 Oct. 1937; P. H. Timberlake leg.; ex Phyla nodiflora (verbatim: Lippia filiformis); UCRC • 1 ♂; same data as for preceding; 14 Sep. 1941; P. H. Timberlake leg.; ex Conyza bonariensis (verbatim: Erigeron linifolius); UCRC • 1 ♀; same data as for preceding; 27 Aug. 1953; P. H. Timberlake leg.; ex Picris echioides; EMEC • 1 ♀; Riverside Co., Tahquitz valley; [33.770 ° N, 116.658 ° W]; 23 Jul. 1928; P. H. Timberlake leg.; ex Geranium; UCRC • 1 ♀; San Benito Co., 6 mi. SE of Idria; [36.355 ° N, 120.598 ° W]; 28 Jun. 1954; J. D. Lattin leg.; EMEC • 1 ♀; San Benito Co., 6 mi. SE of Idria; [36.355 ° N, 120.598 ° W]; 29 Jun. 1954; J. D. Lattin leg.; EMEC • 1 ♀; San Benito Co., Idria; [36.417 ° N, 120.674 ° W]; 29 Jun. 1955; M. Wasbauer leg.; ex Mimulus fremonti; EMEC • 7 ♀♀; San Benito Co., Idria (gem mine); [36.336 ° N, 120.606 ° W]; 5 Jul. 1954; R. F. Smith and E. G. Linsley leg.; ex Achillea millefolium; EMEC • 1 ♀; same data as for preceding; 29 Jun. 1954; C. D. MacNeill leg.; EMEC • 1 ♀; same data as for preceding; 29 Jun. 1955; P. D. Hurd leg.; ex Phacelia; EMEC • 1 ♀; San Benito Co., Idria (Gem Mine); [36.336 ° N, 120.606 ° W]; 15 Jun. 1955; D. J. Burdick leg.; EMEC • 2 ♂♂; San Benito Co., Idria (Gem Mine); [36.336 ° N, 120.606 ° W]; 5 Jul. 1954; R. F. Smith and E. G. Linsley leg.; ex Achillea millefolium; EMEC • 1 ♀; San Benito Co.; [36.62 ° N, 121.12 ° W]; 24 Jun. 1968; D. Veirs leg.; EMEC • 1 ♀; San Bernardino Co., Lone Pine Canyon; [34.323 ° N, 117.563 ° W]; elev. 1371.6 m; 16 Jun. 1928; P. H. Timberlake leg.; ex Eriastrum virgatum (verbatim: Hugelia virgata); UCRC • 1 ♀; San Bernardino Co., Lytle Creek Park; [34.095 ° N, 117.309 ° W]; 23 May 1936; P. H. Timberlake leg.; ex Iris hartwegii; UCRC • 1 ♀; same data as for preceding; 23 May 1936; P. H. Timberlake leg.; ex Fragaria vesca subsp. californica (verbatim: Fragaria californica); UCRC • • 1 ♂; San Bernardino Co., Mill Creek; [34.083 ° N, 116.894 ° W]; elev. 1890 m; 7 Aug. 1946; P. H. Timberlake leg.; ex Eriogonum; UCRC (Figs 36, 37 A – C) • 1 ♀; San Bernardino Co., 1 mi. E of Highland; [34.110 ° N, 117.153 ° W]; 6 May 1965; M. E. Irwin leg.; UCRC • 2 ♀♀; San Bernardino Co., Lake Arrowhead, San Bernardino Mountains; [34.248 ° N, 117.189 ° W]; 9 Jul. 1956; P. D. Hurd leg.; ex Aster; EMEC • 2 ♂♂; same data as for preceding; 9 Jul. 1956; P. D. Hurd leg.; ex Aster; EMEC • 1 ♀; San Bernardino Co., Seven Oaks, San Bernardino Mountain; [34.186 ° N, 116.910 ° W]; 11 Jul. 1964; E. I. Schlinger leg.; UCRC • 1 ♀; San Bernardino Co., Sugarloaf; [34.243 ° N, 116.828 ° W]; 15 Aug. 1953; H. T. Spieth leg.; USNM • 1 ♂; same data as for preceding; 6 Jul. 1954; A. L. Melander leg.; USNM • 1 ♀; San Bernardino Co., up Santa Ana River; [34.05 ° N, 117.34 ° W]; 15 Sep. 1954; A. L. Melander leg.; EMEC • 4 ♀♀; same data as for preceding; 8 Jun. 1955; A. L. Melander leg.; USNM • 1 ♀; same data as for preceding; 29 Aug. 1956; A. L. Melander leg.; USNM • 1 ♀; San Bernardino Co., Wildwood Canyon, 5 mi. E of Calimesa; [34.002 ° N, 116.975 ° W]; 11 Jun. 1968; M. E. Irwin leg.; UCRC • 2 ♀♀; San Diego Co., Mount Laguna; [32.87 ° N, 116.42 ° W]; 21 Jun. 1963; P. D. Hurd leg.; EMEC • 1 ♀; same data as for preceding; 5 Jul. 1963; H. L. Griffin leg.; EMEC • 2 ♀♀; San Diego Co., Mt Palomar; [33.363 ° N, 116.836 ° W]; 28 Jun. 1963; P. D. Hurd leg.; ex Cryptantha intermedia; EMEC • 2 ♀♀; San Diego Co., Warner Springs, Agua Caliente Creek; [33.279 ° N, 116.720 ° W]; elev. 944.9 m; 23 – 25 Aug. 1980; M. Wasbauer and P. Adams leg.; CSCA • 1 ♀; San Diego Co., William Heise County Park; [33.037 ° N, 116.592 ° W]; 28 Jul. 1982; J. C. Hall leg.; USNM • 2 ♀♀; San Diego Co.; [33.03 ° N, 116.77 ° W]; 8 Jun. 2021; SDMC • 1 ♀; San Diego Co., Warner Springs; [33.282 ° N, 116.634 ° W]; 6 Aug. 2021; EMEC • 1 ♀; San Joaquin Co., Tracy; [37.738 ° N, 121.434 ° W]; 21 Jun. 1949; Ray F. Smith leg.; EMEC • 2 ♀♀; same data as for preceding; 20 Jun. 1949; Ray F. Smith leg.; ex Convolvulus; EMEC • 2 ♂♂; same data as for preceding; 21 Jun. 1949; Ray F. Smith leg.; ex Melilotus; EMEC • 3 ♀♀; same data as for preceding; 21 Jun. 1949; Ray F. Smith leg.; ex Melilotus; EMEC • 4 ♀♀; same data as for preceding; 5 May 1949; ex Brassica; EMEC • 1 ♂; same data as for preceding; 13 Jun. 1949; J. W. MacSwain leg.; EMEC • 1 ♀; San Luis Obispo Co., 5 mi. NE of Santa Margarita; [35.441 ° N, 120.547 ° W]; 25 Jun. 1964; J. W. MacSwain leg.; ex Oenothera; EMEC • 1 ♀; same data as for preceding; 25 Jun. 1964; J. W. MacSwain leg.; ex Clarkia unguiculata; EMEC • 1 ♂; Santa Barbara Co.; 34.0225 ° N, 119.8728 ° W; 10 Oct. 2012; S. Teel leg.; BIML • 3 ♂♂; same data as for preceding; 18 Sep. 2012; S. Teel leg.; BIML • 1 ♀; Los Angeles Co., Big Pine Camp; [34.396 ° N, 117.734 ° W]; elev. 1850 m; 16 Jul. 1927; P. H. Timberlake leg.; ex Erigeron; UCRC • 3 ♀♀; same data as for preceding; 12 Jul. 1927; P. H. Timberlake leg.; ex Gilia; UCRC • 1 ♀; same data as for preceding; 16 Jul. 1927; P. H. Timberlake leg.; UCRC • 1 ♀; same data as for preceding; 13 Jul. 1927; P. H. Timberlake leg.; ex Eriogonum; UCRC • 1 ♀; same data as for preceding; 15 Jul. 1927; P. H. Timberlake leg.; UCRC • 1 ♀; Santa Clara Co., Mountain Home Canyon; [7.121 ° N, 121.796 ° W]; 19 Sep. 1949; A. L. Melander leg.; UCRC • 1 ♀; Santa Clara Co., San Antonio Valley; [37.344 ° N, 121.453 ° W]; 22 Jul. 1955; M. Wasbauer leg.; EMEC • 2 ♂♂; Santa Cruz Co., Laurel; [37.117 ° N, 121.965 ° W]; 2 Jul. 1956; D. J. Burdick leg.; EMEC • 1 ♀; Shasta Co., 2 mi. W of Viola; [40.518 ° N, 121.716 ° W]; 20 May 1941; C. D. Michener leg.; EMEC • 1 ♀; Shasta Co., Burney Falls; [41.012 ° N, 121.652 ° W]; E. C. Van Dyke leg.; ex Grindelia; CAS • 1 ♀; Shasta Co., Cassel; [40.919 ° N, 121.548 ° W]; 5 Jul. 1955; E. E. Lindquist leg.; EMEC • 1 ♀; Shasta Co., Old Station; [40.675 ° N, 121.431 ° W]; 24 Jun. 1955; E. E. Lindquist leg.; EMEC • 1 ♀; Shasta Co., Old Station; [40.673 ° N, 121.423 ° W]; 22 Jun. 1955; J. W. MacSwain leg.; EMEC • 1 ♀; Shasta Co., Redding; [40.583 ° N, 122.367 ° W]; 4 Jul. 1975; T. R. Haig leg.; CSCA • 1 ♀; same data as for preceding; 7 Aug. 1975; T. R. Haig leg.; CSCA • 1 ♀; Shasta Co., Shingletown; [40.496 ° N, 121.86 ° W]; 22 May 1941; EMEC • 1 ♀; Sierra Co., 5 mi. NW of Calpine; [39.717 ° N, 120.5062 ° W]; 31 May 1950; J. G. Rozen leg.; ex Ranunculus; EMEC • 1 ♀; Sierra Co.; [39.58 ° N, 120.53 ° W]; 19 Jun. 1913; F. W. Nunenmacher leg.; EMEC • 4 ♀♀; Siskiyou Co., 1 mi. SE of Bartle; [41.247 ° N, 121.808 ° W]; 8 – 10 Jun. 1974; R. Coville leg.; EMEC • 1 ♀; Siskiyou Co., 9 mi. N of Bartle; [41.388 ° N, 121.821 ° W]; 12 – 15 Jun. 1974; R. Coville leg.; EMEC • 1 ♀; Siskiyou Co., Ash Creek Ranger Station; 9 mi. E of McCloud; [41.253 ° N, 121.965 ° W]; elev. 1066.8 m; 10 – 12 Jun. 1974; R. Coville leg.; EMEC • 1 ♀; same data as for preceding; 13 – 16 Jun. 1974; R. Coville leg.; EMEC • 1 ♀; Siskiyou Co., Brown’s Lake, SW of Mt Shasta City; [41.299 ° N, 122.330 ° W]; 13 – 14 Jun. 1974; J. Powell leg.; EMEC • 1 ♀; Siskiyou Co., Dunsmuir; [41.22 ° N, 122.27 ° W]; Wickham leg.; USNM • 1 ♀; Siskiyou Co., Mt Shasta City; [41.328 ° N, 122.325 ° W]; 20 Jun. 1958; J. Powell leg.; EMEC • 1 ♂; Siskiyou Co., Salmon Trinity Alps Wilderness Area, Big Flat Campground; [41.017 ° N, 123.089 ° W]; elev. 1524 m; 2 Aug. 1968; H. B. Leech leg.; CAS • 1 ♀; Siskiyou Co., Shasta Springs; [41.2468 ° N, 122.2611 ° W]; 8 Jun. 1920; C. L. Fox leg.; CAS • 2 ♀♀; Sonoma Co., Pitkin Ranch, 6 mi. N of Sebastopol; [38.487 ° N, 122.824 ° W]; 25 Jul. 1965; J. W. MacSwain leg.; ex Clarkia imbricata; EMEC • 1 ♀; same data as for preceding; 22 Jul. 1965; J. W. MacSwain leg.; ex Clarkia imbricata; EMEC • 1 ♂; same data as for preceding; 25 Jul. 1965; J. W. MacSwain leg.; ex Clarkia imbricata; EMEC • 1 ♂; Stanislaus Co., Ceres; [37.601 ° N, 120.957 ° W]; 28 Aug. 1953; W. W. Middlekauff leg.; EMEC • 1 ♀; Trinity Co., Hayfork Ranger Station; [40.553 ° N, 123.165 ° W]; elev. 701 m; 18 May 1973; J. A. Chemsak leg.; EMEC • 1 ♀; Trinity Co., Junction City; [40.74 ° N, 123.06 ° W]; 30 Jun. 1975; T. R. Haig leg.; CSCA • 1 ♀; same data as for preceding; 21 Apr. 1977; T. R. Haig leg.; CSCA • 1 ♂; same data as for preceding; 13 Jun. 1975; T. R. Haig leg.; CSCA • 1 ♀; Tulare Co., 3 mi. W of Three Rivers; [36.437 ° N, 118.958 ° W]; 10 May 1936; P. H. Timberlake leg.; ex Camissonia contorta (verbatim: Oenothera dentata); UCRC • 1 ♀; Tulare Co., California Hot Springs; [35.880 ° N, 118.677 ° W]; 2 Jun. 1939; E. C. Van Dyke leg.; CAS • 1 ♀; Tulare Co., Badger; [36.631 ° N, 119.013 ° W]; 20 Jun. 1924; P. H. Timberlake leg.; ex Brodiaea coronaria; UCRC • 1 ♂; same data as for preceding; 26 Jun. 1929; P. H. Timberlake leg.; ex Frangula californica (verbatim: Rhamnus californica); UCRC • 2 ♀♀; Tuolumne Co., Berkeley Tuolumne Camp; [37.811 ° N, 119.944 ° W]; 12 Jun. 1968; S. Paul Welles leg.; ex Clarkia rhomboidea; EMEC • 1 ♀; same data as for preceding; 12 Jun. 1968; S. Paul Welles leg.; EMEC • 4 ♀♀; same data as for preceding; 4 Jun. 1968; S. Paul Welles leg.; ex Clarkia rhomboidea; EMEC • 1 ♀; Tuolumne Co., Camp Bob MacBride, Pinecrest; [37.942 ° N, 120.262 ° W]; elev. 1645.9 m; 1 Aug. 1952; R. Snelling leg.; LACM • 1 ♀; Tuolumne Co., Pinecrest; [38.189 ° N, 119.992 ° W]; 12 Jul. 1953; J. G. Rozen leg.; ex Phacelia; EMEC • 1 ♀; Tuolumne Co., Strawberry; [38.198 ° N, 120.009 ° W]; 24 Jun. 1953; J. G. Rozen leg.; EMEC • 1 ♀; same data as for preceding; 18 May 1953; J. G. Rozen leg.; EMEC • 2 ♀♀; same data as for preceding; 27 Aug. 1960; P. D. Hurd leg.; ex Ericameria bloomeri (verbatim: Haplopappus bloomeri subsp. angustatus); EMEC • 1 ♀; same data as for preceding; 29 Jul. 1960; R. W. Thorp leg.; ex Erigeron; EMEC • 1 ♀; same data as for preceding; 21 Jul. 1960; P. D. Hurd leg.; ex Erigeron nudum; EMEC • 1 ♀; same data as for preceding; 2 Aug. 1960; E. Jessen leg.; EMEC • 2 ♀♀; Tuolumne Co., Strawberry; [38.20 ° N, 120.01 ° W]; 1 Aug. 1960; C. A. Toschi leg.; EMEC • 1 ♀; Tuolumne Co., Tuolumne City; [37.961 ° N, 120.237 ° W]; 22 Jun. 1953; J. G. Rozen leg.; EMEC • 1 ♂; Tuolumne Co., Strawberry; [38.20 ° N, 120.01 ° W]; 21 Aug. 1960; P. D. Hurd leg.; ex Eriogonum nudum; EMEC • 1 ♀; Ventura Co., Lockwood Creek, near Stauffer P. O.; [34.730 ° N, 119.033 ° W]; 7 May 1959; P. D. Hurd leg.; ex Phacelia fremontii; EMEC • 3 ♀♀; Ventura Co., Lockwood Valley; [34.7325 ° N, 119.042 ° W]; 5 May 1959; P. H. Timberlake leg.; ex Sisymbrium; UCRC • 3 ♀♀; Ventura Co., Lockwood Creek, near Stauffer P. O.; [34.730 ° N, 119.033 ° W]; 5 May 1959; J. R. Powers leg.; ex Descurainia sophia; EMEC • 1 ♀; same data as for preceding; 7 May 1959; P. D. Hurd leg.; ex Leptosiphon aureus; EMEC • 2 ♀♀; same data as for preceding; 7 May 1959; P. D. Hurd leg.; ex Descurainia sophia; EMEC. – Colorado • 10 ♀♀; La Plata Co., Los Pinos; [37.09 ° N, 106.49 ° W]; 22 May 1999; Carl F. Baker leg.; ex Erigeron; USNM • 1 ♀; Larimer Co., Hewlett Gulch; [40.706 ° N, 105.314 ° W]; 1 Jul. 1983; H. E. Evans leg.; UCMC • 2 ♀♀; Montrose Co., East Portal; [39.92 ° N, 105.66 ° W]; elev. 1981.2 m; 27 Jul. 1962; U. N. Lanham leg.; UCMC • 1 ♀; Rio Grande Co., Monte Vista; 37.583333 ° N, 106.15 ° W; elev. 2331.7 m; 16 Jun. 2019; UCRC. – Idaho • 1 ♀; Benewah Co., Saint Maries; [47.32 ° N, 116.57 ° W]; 24 Jun. 2019; H. A. Scullen leg.; OSAC • 1 ♀; Blaine Co., Magic Reservoir; 43.3277 ° N, 114.3999 ° W; 7 Jul. 2009; F. Morrison leg.; WRME • 1 ♀; Cassia or Minidoka Co., Burley; [42.54 ° N, 113.79 ° W]; 14 Jun. 1929; ex Solidago altissima; USNM • 1 ♀; Twin Falls Co., Hollister; [42.35 ° N, 114.58 ° W]; 8 Jul. 1930; A. Rosea and S. Pestifer leg.; EMEC • 1 ♂; same data as for preceding; 24 Jul. 1930; EMEC. – Montana • 1 ♀; Missoula Co., Florence MPG Ranch; 46.6905 ° N, 114.0265 ° W; elev. 1136 m; 8 – 10 Jul. 2019; M. Seidensticker leg.; CBG • 2 ♀♀; Missoula Co., Florence MPG Ranch; 46.6776 ° N, 114.0392 ° W; elev. 986 m; 16 – 18 Jul. 2019; M. Seidensticker leg.; CBG • 1 ♀; same data as for preceding; 9 – 11 Jul. 2019; M. Seidensticker leg.; CBG • 1 ♀; same data as for preceding; 23 – 25 Jul. 2019; M. Seidensticker leg.; CBG • 1 ♀; Missoula Co., Florence MPG Ranch; 46.6819 ° N, 114.0288 ° W; elev. 1018 m; 2 – 4 Jul. 2019; M. Seidensticker leg.; CBG. – Nevada • 1 ♀; Elko Co., Wells; [41.11 ° N, 114.97 ° W]; 20 Jun. 1927; J. M. Aldrich leg.; EMEC • 1 ♀; Mineral Co., 4.5 mi. S of Schurz; [38.8812 ° N, 118.8117 ° W]; 20 Jul. 1982; W. J. Pulawski leg.; CAS • 8 ♀♀; Ormsby Co.; [39.16 ° N, 119.77 ° W]; Jul.; Baker leg.; USNM • 1 ♀; Washoe Co., 10 mi. S of Reno on hyw. 395; [39.625 ° N, 119.913 ° W]; 19 Jun. 1959; R. W. Thorp leg.; ex Chylismia claviformis subsp. integrior (verbatim: Oenothera claviformis var. cruciformis); EMEC • 1 ♀; Washoe Co., Verdi; [39.518 ° N, 119.989 ° W]; 25 Jun. 1954; J. A. Powell leg.; EMEC. – Oregon • 1 ♂; Benton Co., Corvallis; [44.567 ° N, 123.283 ° W]; 28 Aug. 1931; USNM • 1 ♀; same data as for preceding; 17 Sep. 1935; G. Ferguson leg.; EMEC • 2 ♀♀; same data as for preceding; 10 May 1936; N. P. Laraon leg.; OSAC • 1 ♀; same data as for preceding; 14 Jul. 1935; OSAC • 1 ♀; same data as for preceding; 4 Jun. 1948; OSAC • 1 ♀; Crook Co., 15 mi. SE of Prineville; [44.145 ° N, 120.620 ° W]; elev. 1066.8 m; 22 May 1971; Oman leg.; OSAC • 1 ♂; Deschutes Co., Sisters; [44.2909 ° N, 121.5492 ° W]; 17 Aug. 1941; ex Chrysothamnus; OSAC • 1 ♀; Douglas Co., Yoncalla; [43.599 ° N, 123.283 ° W]; 26 May 1937; R. M. and G. E. Bohart leg.; CAS • 1 ♀; Grant Co., 19 mi. S of Dale; [44.713 ° N, 118.947 ° W]; 17 Jul. 1965; J. Wolf leg.; ex Clarkia pulchella; EMEC • 5 ♀♀; Grant Co., 9 mi. S of Dale; [44.858 ° N, 118.947 ° W]; 17 Jul. 1965; R. L. Langston leg.; ex Clarkia pulchella; EMEC • 2 ♀♀; Harney Co., Malheur National Wildlife Refuge, HQT; 43.2652 ° N, 118.8433 ° W; 8 – 9 Jul. 2010; Linda Beck leg.; OSAC • 1 ♂; Klamath Co., Algoma, Klamath Falls; [42.225 ° N, 121.782 ° W]; 23 Aug. 1958; Joe Schuh leg.; OSAC • 1 ♀; Klamath Co., Klamath Falls, Geary Ranch; [42.277 ° N, 121.884 ° W]; 10 Jun. 1958; J. Schuh leg.; OSAC • 1 ♀; Klamath Co., Klamath Lake, Alkali Pond; [42.39 ° N, 121.88 ° W]; 19 Aug. 1958; J. Schuh leg.; OSAC • 1 ♀; Lane Co., Cottage Grove; [43.798 ° N, 123.059 ° W]; 1 – 31 Jul. 1939; G. Moxiey leg.; LACM • 1 ♀; Lane Co., Eugene; [43.95 ° N, 122.88 ° W]; 29 May 1933; O. H. Swezey leg.; CAS • 1 ♀; Lane Co., Springfield; [41.666 ° N, 110.934 ° W]; W. Holleman leg.; EMEC • 1 ♀; Linn Co., High Cascade Mountains; [46.85 ° N, 121.76 ° W]; 20 Jul. 2009; J. C. Bridwell leg.; EMEC • 1 ♀; Milton Co.; [45.94 ° N, 118.39 ° W]; 22 Jun. 1938; K. Gray and J. Schuh leg.; OSAC • 5 ♀♀; Multnomah Co., Oxbow Meadow; 45.4844 ° N, 122.3015 ° W; 2 Jun. 2021; J. D. Engler leg.; OSAC • 1 ♀; Multnomah Co., Oxbow Meadow; 45.485 ° N, 122.3015 ° W; 23 Jul. 2021; J. D. Engler leg.; OSAC • 1 ♂; same data as for preceding; 23 Jul. 2021; J. D. Engler leg.; ex Tarax / Hypochaer; OSAC • 1 ♂; same data as for preceding; 23 Jul. 2021; J. D. Engler leg.; ex Erigeron; OSAC • 1 ♂; Polk Co., Grand Island; [45.127 ° N, 123.038 ° W]; 6 Aug. 1941; R. E. Rieder leg.; OSAC • 1 ♀; Union Co., Union; [45.210 ° N, 117.865 ° W]; elev. 823 m; 11 Jun. 1926; H. A. Scullen leg.; OSAC • 2 ♀♀; Wallowa Co., 5 mi. E of Minam, Wallowa Canyon Fount; [45.621 ° N, 117.626 ° W]; elev. 823 m; 21 Jul. 1929; H. A. Scullen leg.; OSAC. – Utah • 3 ♀♀; Duchesne Co., 5.5 mi. W of Roosevelt; [40.299 ° N, 110.093 ° W]; 20 May 1962; J. W. MacSwain leg.; EMEC • 3 ♂♂; Uinta Co.; [41.26 ° N, 110.57 ° W]; CMNH • 1 ♂; same data as for preceding; 8 Sep. 2011; CMNH. – Washington • 1 ♀; Klickitat Co., Conboy Lake National Wildlife Refuge, ARN; 45.9812 ° N, 121.2616 ° W; 14 Jul. 2011; L. Wilson leg.; OSAC • 2 ♂♂; Klickitat Co., Conboy Lake National Wildlife Refuge, LRL; 45.9283 ° N, 121.3783 ° W; 11 Sep. 2012; Diedre Shirley leg.; OSAC • 1 ♀; Snohomish Co., 13 mi. E of Snohomish Pass; [47.391 ° N, 121.123 ° W]; 12 Aug. 1943; S. C. Jones leg.; ex Solidago; OSAC • 2 ♀♀; Stevens Co., Little Pend Oreille National Wildlife Refuge, LPO; 48.4654 ° N, 117.7358 ° W; 11 – 12 Jul. 2012; J. Koelsch leg.; OSAC • 4 ♀♀; same data as for preceding; 20 – 21 Jul. 2012; J. Koelsch leg.; OSAC • 1 ♀; Stevens Co., Little Pend Oreille National Wildlife Refuge, LPO; 48.4604 ° N, 117.7331 ° W; 16 – 17 Jul. 2012; J. Koelsch leg.; OSAC • 1 ♀; Stevens Co., Little Pend Oreille National Wildlife Refuge, LPO; 48.4654 ° N, 117.7358 ° W; 23 Apr. 2012; Mike Munts leg.; OSAC • 1 ♀; Spokane Co., Turnbull National Wildlife Refuge, SSA; 47.39 ° N, 117.51 ° W; 7 – 20 Jul. 2011; J. Bastow leg.; OSAC • 2 ♀♀; Spokane Co., Turnbull National Wildlife Refuge, SSA; 47.39 ° N, 117.51 ° W; 8 – 15 Jul. 2011; Mike Rule leg.; OSAC • 1 ♀; same data as for preceding; 22 – 28 Sep. 2011; Mike Rule leg.; OSAC • 1 ♀; Spokane Co., Turnbull National Wildlife Refuge, SSA; 47.4116 ° N, 117.5852 ° W; 17 – 18 May 2016; D. Cousins leg.; OSAC • 1 ♀; Spokane Co., Turnbull National Wildlife Refuge, SSA; 47.39845 ° N, 117.605648 ° W; 26 – 27 May 2016; M. Ploegman leg.; OSAC • 1 ♂; Whitman Co., Pullman; [46.7333 ° N, 117.1667 ° W]; 13 Sep. 2008; EMEC • 1 ♀; Whitman Co., Wanawai; [37.5369 ° N, 119.6564 ° W]; 12 Jul. 1925; C. L. Fox leg.; CAS. – Wyoming • 2 ♀♀; Lincoln Co., Bridger Basin; [41.66606 ° N, 110.9335 ° W]; S. Garman leg.; EMEC • 1 ♀; Natrona Co., South Fork Powder River at Hwy 20, 1.8 mi. W of Town of Powder River; [43.0316 ° N, 107.0227 ° W]; 21 Aug. 1965; H. B. Leech leg.; CAS. • 1 ♀; Park Co.; 44.9701 ° N, 110.6999 ° W; 20 – 21 Jun. 2012; A. Rodman leg.; BIML.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFC4FF83FE66FE87FD21F929.taxon	description	Description GENITALIA (Figs 37, 120 F). Gonostylus elongate, approximately twice as long as broad, and bears short setae. Retrorse lobe broad, about 2 times as long as broad, covered in fine setae.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFC4FF83FE66FE87FD21F929.taxon	distribution	Range Canada (Alberta, British Columbia, Saskatchewan), USA (California, Colorado, Idaho, Nevada, Oregon, Utah, Washington) (Fig. 38).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFC4FF83FE66FE87FD21F929.taxon	biology_ecology	Floral hosts APIACEAE: Perideridia gairdneri (Hook. & Arn.) Mathias • ASTERACEAE: Achillea millefolium L., Aster sp., Chaenactis glabriuscula DC., Chaenactis glabriuscula DC. var. lanosa (DC.) H. M. Hall, Chrysothamnus sp., Chrysothamnus viscidiflorus (Hook.) Nutt. var. typicus (H. M. Hall & Clem.) Cronquist, Conyza bonariensis (L.) Cronquist, Ericameria bloomeri (A. Gray) J. F. Macbr., Erigeron divergens Torr. & A. Gray, Erigeron sp., Grindelia sp., Heterotheca grandiflora Nutt., Picris echioides L., Solidago altissima L., Solidago sp., Taraxacum sp. or Hypochaeris sp. • BORAGINACEAE: Cryptantha intermedia (A. Gray) Greene, Cryptantha micrantha (Torr.) I. M. Johnst. • BRASSICACEAE: Brassica sp., Descurainia sophia (L.) Webb ex Prantl, Sisymbrium irio L., Sisymbrium sp. • CAPPARACEAE: Cleome lutea Hook. • CONVOLVULACEAE: Convolvulus sp. • FABACEAE: Melilotus sp. • GERANIACEAE: Geranium sp. • HYDROPHYLLACEAE: Phacelia fremontii Torr., Phacelia humilis Torr. & A. Gray, Phacelia sp. • IRIDACEAE: Iris hartwegii Baker • LILIACEAE: Brodiaea coronaria (Salisb.) Engl. • MALVACEAE: Eremalche exilis (A. Gray) Greene • ONAGRACEAE: Camissonia claviformis (Torr. & Frém.) P. H. Raven subsp. cruciformis (Kellogg) P. H. Raven, Camissonia campestris (Greene) P. H. Raven subsp. campestris, Camissonia sierrae P. H. Raven, Taraxia tanacetifolia (Torr. & A. Gray) Piper, Clarkia dudleyana (Abrams) J. F. Macbr., Clarkia gracilis subsp. albicaulis (Jeps.) F. H. Lewis & M. E. Lewis, Clarkia heterandra (Torr.) F. H. Lewis & P. H. Raven, Clarkia imbricata F. H. Lewis & M. E. Lewis, Clarkia pulchella Pursh, Clarkia purpurea subsp. viminea (Douglas) F. H. Lewis & M. E. Lewis, Clarkia rhomboidea Douglas, Clarkia sp., Clarkia rubicunda (Lindl.) F. H. Lewis & M. E. Lewis, Clarkia unguiculata Lindl., Clarkia williamsonii (Durand & Hilg.) F. H. Lewis & M. E. Lewis, Gayophytum heterozygum F. H. Lewis & Szweyk., Gayophytum humile Juss., Gayophytum sp., Camissonia contorta (Douglas ex Lehm.) Kearney, Oenothera sp. • PAPAVERACEAE: Eschscholzia californica Cham., Eschscholzia sp. • POLEMONIACEAE: Eriastrum virgatum (Benth.) H. Mason, Gilia achilleifolia subsp. multicaulis (Benth.) A. D. Grant & V. E. Grant, Gilia sp., Leptosiphon aureus (Nutt.) J. M. Porter & L. A. Johnson subsp. aureus • POLYGONACEAE: Eriogonum davidsonii Greene, Eriogonum gracile Benth., Eriogonum nudum Douglas ex Benth., Eriogonum sp., Polygonum lapathifolium L. • PORTULACACEAE: Portulaca sp. • RANUNCULACEAE: Ranunculus sp. • RHAMNACEAE: Frangula californica (Eschsch.) A. Gray • ROSACEAE: Chamaebatiaria millefolium (Torr.) Maxim., Fragaria vesca L. subsp. californica (Cham. & Schltdl.) Staudt • SCROPHULARIACEAE: Mimulus fremontii (Benth.) A. Gray, Scrophularia californica Cham. & Schltdl. • VERBENACEAE: Phyla nodiflora (L.) Greene	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFC4FF83FE66FE87FD21F929.taxon	discussion	Biological remarks Lasioglossum glabriventre is a ground-nesting species. Hicks (1936) reports excavating a nest of this species in August. The nest was three inches deep and made in fine sand. DNA barcodes Six full DNA barcodes and two partial barcodes are available, all from British Columbia. They form a discrete cluster with little intraspecific variation.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFBEFF84FECAF905FC92FE91.taxon	diagnosis	Diagnosis This group is distinguished by a combination of the following characters: the male antenna relatively long, reaching to the metasoma, much shorter in other species of L. (Hemihalictus); the propodeal triangle long, nearly as long as mesoscutellum in both sexes; and the gonostylus of the male genitalia is slender and weakly clavate (Ebmer & Sakagami 1985).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFB9FF99FDF8FE72FD7BFB52.taxon	description	Figs 39, 40 A – C, 41 A – C, 42 – 45, 116 A, 117 C, 118 A, D, G, J, 120 G	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFB9FF99FDF8FE72FD7BFB52.taxon	diagnosis	Diagnosis All species in the L. nitidiusculum species group are very similar in appearance, with the exception of L. synthyridis (formerly known as L. inconditum (Cockerell, 1916 )). Both sexes of L. synthyridis can be easily distinguished from other species in the group by the rugae of the propodeal triangle which reach the sharply edged posterior margin of the dorsal surface of the propodeum. In contrast, both sexes of all other species in the group have propodeal triangle with rugae that do not reach the smoothly rounded posterior margin. Females of L. foxii are very similar to females of L. ruficorne. However, females of L. ruficorne have the base of T 2 – T 3 densely punctate (IS ≤ 1 PD), becoming moderately dense or moderately sparse (IS = 1 – 2 PD) near or on the premarginal line. In contrast, females of L. foxii have the base of T 2 – T 3 densely punctate (IS ≤ 1 PD), but the punctures become noticeably sparser (IS = 2 – 6 PD) near the premarginal line, and punctures are comparatively smaller. Females of L. foxii typically have a more uniformly imbricate mesoscutum with pronounced microsculpture, whereas females of L. ruficorne usually have a mostly imbricate mesoscutum with a subtle weakening or, at times, a nearly polished appearance submedially. However, as a variation, L. ruficorne can occasionally exhibit pronounced microsculpturing on the mesoscutum as well. In some instances, particularly where the ranges of eastern North American L. foxii and western North American L. ruficorne overlap, distinguishing the two species becomes very difficult, as individuals may exhibit intermixed characters. Females of L. diatretum have mostly polished mesoscutum, becoming imbricate anteriorly and medially halfway on median line, punctures moderately dense to moderately sparse (IS = 1 – 2 PD), becoming broadly sparse (IS = 1 – 3 PD) submedially. But females of L. ruficorne and L. foxii have imbricate mesoscutum with or without subtle polished area submedially and punctures dense throughout (IS ≤ 1 PD) or sparser (IS = 1 – 1.5 PD) submedially. Female L. ruficorne have more translucent yellowish tegula whereas in most of the cases, L. diatretum have comparatively more brownish tegula. The mesoscutum punctures in females of L. diatretum are slightly larger than those of L. ruficorne and L. foxii. Additionally, L. diatretum females possess a larger head, with a mean length of 1.83 mm (± 0.06 SD) and a mean width of 1.76 mm (± 0.04 SD). Both sexes of L. diatretum are generally larger than those of L. ruficorne. In contrast, females of L. ruficorne have a smaller head, with a mean length of 1.65 mm (± 0.06 SD) and a mean width of 1.60 mm (± 0.05 SD). Due to the many variations and overlapping ranges of L. diatretum and L. ruficorne, it is recommended to use a combination of characteristics to separate them. Lasioglossum diatretum females are easily identified when they exhibit sparser punctures on a polished mesoscutum and have brownish tegulae. However, in some cases, they may have a less polished mesoscutum with dense punctures throughout (IS ≤ 1 PD), except for moderately sparse or moderately dense punctures (IS = 1 – 2 PD) mesad of the parapsidal line, making them closely resemble L. ruficorne. Similarly, L. ruficorne females are easily identified when they have denser punctures on a more imbricate mesoscutum and more translucent yellowish tegulae. However, in some instances, their mesoscutum may become more polished submedially, with punctures becoming sparser submedially (IS = 1 – 3 PD), making them closely resemble L. diatretum. Males of L. synthyridis can be easily distinguished from other males in the L. nitidiusculum species group by the propodeal triangle with rugae, which reach the sharply edged posterior margin of the dorsal surface. However, males of L. synthyridis also possess genitalia that are very similar to those of L. diatretum. Both species feature a broad retrorse lobe, slightly folded in laterally, and a very long gonostylus (4 – 5 times as long as broad). The key difference lies in the shape of the retrorse lobe: males of L. diatretum have a rounded retrorse lobe at the apex, while L. synthyridis males exhibit a somewhat angled apex. If the genitalia are exposed, L. diatretum can be easily distinguished from males of L. foxii and L. ruficorne. Males of L. foxii and L. ruficorne have very slender retrorse lobes that are attenuated at the apex, along with a comparatively short gonostylus (2 – 3 times as long as broad) that is triangular-shaped and broad at the apex. In comparison, males of L. foxii and L. ruficorne have very similar-looking genitalia. Males of L. foxii can be separated from L. diatretum and L. ruficorne by the punctation pattern on T 2 – T 3. In L. foxii, the punctures become sparser (IS = 1 – 3 PD) near the premarginal line and are entirely impunctate on the apical rim. In contrast, L. diatretum and L. ruficorne have T 2 – T 3 densely punctate (IS ≤ 1 PD) to moderately dense (IS = 1 – 2 PD) up to the premarginal line, becoming nearly impunctate on the apical rim. When comparing males of L. diatretum and L. ruficorne, several distinguishing features can be observed. Males of L. diatretum have largely brown flagellomeres, dark brown to brown legs, and a comparatively broad abdomen, with T 3 – 4 apical rim slightly depressed or nearly flat. In contrast, males of L. ruficorne have more yellowish-brown flagellomeres, legs that are brown except for the tibiae and tarsi which are largely yellowish brown, and a slender abdomen with the T 3 – 4 apical rim distinctly depressed relative to the disc.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFB9FF99FDF8FE72FD7BFB52.taxon	etymology	Etymology Vachal (1904) did not explain the derivation of the epithet. The specific epithet is treated as a Latin adjective meaning ‘ pierced with holes, carved, embossed’. The same root is used for a category of luxury, patterned vessels used by ancient Romans.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFB9FF99FDF8FE72FD7BFB52.taxon	materials_examined	Type material Lectotype of Halictus diatretus (designated by Moure & Hurd 1987; examined by JG ca 2013) UNITED STATES – Colorado • ♀; NHMW. Syntype of Halictus supranitens (designated herein; examined) UNITED STATES – Colorado • 1 ♀; Larimer Co., Long Peak trail, above timberline; [40.254 ° N, 105.614 ° W]; 20 Jul. 1906; Cockerell leg.; USNM Type No. 27999 (Fig. 45). Specimen labels “ Long Peak / trail, above / timber line / July 20. (Ckll) // Halictus supranitens Ckll. TYPE // Type No. 27999 U. S. N. M. ”. Type remarks Cockerell (1919) did not specify the composition of the type series. Following recommendation 73 F (ICZN 1999), we assume the existence of syntypes, although it may have been described from a single female. Other material examined CANADA – Alberta • 1 ♀; Pincher Creek, West Castle River; 49.3333 ° N, 114.4167 ° W; 28 Jun. 1984; Spanton and Adamski leg.; PMAE • 1 ♀; 15 km S of Three Hills, Hwy 21; 51.49484 ° N, 113.23214 ° W; elev. 702 m; 15 Jun. 2007; J. Gibbs and C. Sheffield leg.; PCYU • 1 ♀; Cypress Co., Medicine Hat, W of Gas City Campground; 50.0358 ° N, 110.7319 ° W; 1 Jun. 2016; M. Buck leg.; PMAE. – British Columbia • 1 ♀; Osoyoos West Bench, Osoyoos; 49.001392 ° N, 119.49994 ° W; 17 May 2017; Dawn Marks (J. Heron) leg.; RSKM • 1 ♀; Osoyoos West Bench, Osoyoos; 49.00142 ° N, 119.49765 ° W; 19 – 20 Jul. 2017; D. Marks and K. Grant (J. Heron) leg.; RSKM • 1 ♀; Osoyoos West Bench (Crown Land), Osoyoos; 49.01235 ° N, 119.4957 ° W; elev. 479 m; 6 Jun. 2017; J. Heron, D. Marks and K. Grant leg.; RSKM • 1 ♀; Osoyoos West Bench, Osoyoos; 49.0038 ° N, 119.49455 ° W; 17 – 19 Jun. 2017; D. Marks and K. Grant (J. Heron) leg.; RSKM • 1 ♂; Osoyoos Mt Kobau; 49.11363 ° N, 119.66677 ° W; 14 Aug. 2014; RSKM • 1 ♀; Okanagan-Similkameen Reg. Dist., Oliver, Ripley Lake; 49.2432 ° N, 119.6329 ° W; elev. 925 m; 12 Jul. 2008; L. R. Best leg.; PCYU • 1 ♀; Okanagan-Similkameen Reg. Dist., Oliver, Mt Baldy, McKinney Camp Rd; 49.1297 ° N, 119.3704 ° W; elev. 1140 m; 24 – 31 May 2010; L. R. Best leg.; PMAE • 1 ♀; Okanagan-Similkameen Reg. Dist., Mt Oliver, Ripley Lake; 49.2431 ° N, 119.6332 ° W; elev. 940 m; 5 – 9 Jun. 2010; L. R. Best leg.; PMAE • 3 ♀♀; same data as for preceding; 18 – 19 May 2010; L. R. Best leg.; PMAE • 3 ♀♀; same data as for preceding; 20 – 21 May 2010; L. R. Best leg.; PMAE • 1 ♀; Robinson; [49.334 ° N, 117.691 ° W]; 24 Sep. 1964; R. R. Foxlee leg.; EMEC • 1 ♀; Vernon, Rattlesnake Point; [50.201 ° N, 119.294 ° W]; 9 May 1948; E. S. Ross leg.; CAS. MEXICO – Baja California • 1 ♂; 8 mi. E of Tecate, HYW 2; [32.571 ° N, 116.506 ° W]; 29 May 1980; Brown Faulkner leg.; SDMC. UNITED STATES – Arizona • 1 ♂; Navajo Co., 5 mi. W of Fort Apache; [33.790 ° N, 110.077 ° W]; 21 Jun. 1957; F. Werner and G. Butler leg.; ASUHIC. – California • 2 ♀♀; Alameda Co., Alameda Foothills; [37.70 ° N, 122.1 ° W]; 15 Apr. or Apr. 1915; W. M. Giffard leg.; CAS • 6 ♂♂; Alpine Co., 1 mi. S of Monitor Pass; [38.660 ° N, 119.62 ° W]; elev. 2499.4 m; 22 – 23 Aug. 1979; M. Wasbauer and P. Adams leg.; CSCA • 1 ♀; same data as for preceding; CSCA • 1 ♀; Alpine Co., Hope Valley; [38.776 ° N, 119.932 ° W]; 29 Jun. 1957; J. Powell leg.; ex Penstemon procerus; EMEC • 1 ♂; same data as for preceding; 20 Aug. 1949; C. D. MacNeill leg.; EMEC • 1 ♀; Alpine Co., Kinney Reservoir; [38.555 ° N, 119.811 ° W]; 8 Jun. 1960; G. I. Stage leg.; EMEC • 1 ♀; Alpine Co., Lake Alpine; [38.477 ° N, 120.003 ° W]; 8 Jun. 1960; R. W. Thorp leg.; ex Salix; EMEC • 3 ♀♀; same data as for preceding; 8 Jun. 1960; G. I. Stage leg.; ex Salix; EMEC • 1 ♀; Alpine Co., Markleville; [38.692 ° N, 119.781 ° W]; 29 May 1957; R. P. Allen leg.; EMEC • 1 ♀; Butte Co., Pentz; [39.655 ° N, 121.584 ° W]; 5 Apr. 1928; H. H. Keifer leg.; CAS • 3 ♀♀; Butte Co., Yankee Hill; [39.739 ° N, 121.47 ° W]; 20 Mar. 1928; H. H. Keifer leg.; CAS • 3 ♂♂; Calaveras Co., Murphys; [38.137 ° N, 120.461 ° W]; 25 May 1936; F. E. Blaisdell leg.; CAS • 1 ♀; Chaffee Co., Maysville; [38.534 ° N, 106.171 ° W]; 18 Jun. or Jun. 1918; T. D. A. Cockerell leg.; CAS • 1 ♀; El Dorado Co., Blodgett Forest, 13 mi. E of Georgetown; [38.911 ° N, 120.661 ° W]; 28 May 1977; J. Powell leg.; ex Arctostaphylos; EMEC • 1 ♀; El Dorado Co., Blodgett Forest, 13 mi. E of Georgetown; [38.911 ° N, 120.596 ° W]; 13 Jul. 1967; J. Powell leg.; EMEC • 2 ♂♂; El Dorado Co., Blodgett Forest, 13 mi. E of Georgetown; [38.911 ° N, 120.596 ° W]; 13 Jul. 1967; J. Powell leg.; EMEC • 1 ♀; El Dorado Co., Echo Lake; [38.834 ° N, 120.042 ° W]; elev. 2270 m; 6 Apr. 1934; E. O. Essig leg.; EMEC • 1 ♀; same data as for preceding; 29 Jun. 1953; W. W. Middlekauff leg.; EMEC • 1 ♀; same data as for preceding; 3 Jul. 1953; W. W. Middlekauff leg.; EMEC • 1 ♀; El Dorado Co., Fallen Leaf; [38.883 ° N, 120.073 ° W]; 1 Jul. 1931; UCRC • 1 ♂; same data as for preceding; 19 Aug. 1931; UCRC • 1 ♂; same data as for preceding; 7 Aug. 1931; UCRC (Figs 41 B, 42 – 43) • 1 ♀; El Dorado Co., N. end of Folsom Lake; [38.770 ° N, 121.107 ° W]; 29 Apr. 1967; A. Keuter leg.; CAS • 1 ♂; El Dorado Co., Pilot Creek; [38.929 ° N, 120.648 ° W]; 15 Jul. 1967; J. Powell leg.; EMEC • 2 ♂♂; El Dorado Co., Snowline Camp; [38.746 ° N, 120.627 ° W]; 20 Jul. 1948; P. D. Hurd leg.; EMEC • 1 ♂; same data as for preceding; 7 Jul. 1948; K. W. Tucker leg.; ex Compositae; EMEC • 2 ♀♀; Fresno Co., General Grant National Park; [36.747 ° N, 118.975 ° W]; 12 Jun. 1925; P. H. Timberlake leg.; ex Ribes; UCRC • 1 ♀; Fresno Co., Huntington Lake; [37.232 ° N, 119.238 ° W]; 5 Jul. 1917; I. McCracken leg.; ex Compositae; EMEC • 1 ♀; same data as for preceding; 7 Jul. 1919; E. P. Van Duzee leg.; CAS • 1 ♀; Fresno Co., Pine Springs Ranch, Rivers; [33.7083 ° N, 116.6508 ° W]; elev. 1595 m; 16 Mar. 1940; P. H. Timberlake leg.; ex Arctostaphylos glauca; UCRC • 4 ♂♂; Fresno Co., Pinehurst; [36.696 ° N, 119.016 ° W]; elev. 1250 m; 24 Jun. 1960; R. R. Snelling leg.; EMEC • 3 ♀♀; Humboldt Co., 4 mi. E of Shelter Cove; [40.033 ° N, 123.999 ° W]; 21 May 1976; J. Powell leg.; ex Phacelia; EMEC • 1 ♀; same data as for preceding; 21 May 1976; J. Chemsak leg.; ex Phacelia; EMEC • 4 ♀♀; Humboldt Co., 7 mi. SE of Shelter Cove; [39.962 ° N, 123.978 ° W]; 22 May 1976; J. Powell leg.; ex Ceanothus incanus; EMEC • 1 ♂; Inyo Co., Kearsarge Pass Road, 8 mi. W of Independence; [36.771 ° N, 118.349 ° W]; 29 Aug. 1979; M. Wasbauer and P. Adams leg.; EMEC • 1 ♂; Inyo Co., White Mountains, Campito Mountain; [37.499 ° N, 118.201 ° W]; 1 Aug. 1981; Brown Faulkner leg.; SDMC • 1 ♀; Kern Co., Double Mountain; [35.035 ° N, 118.445 ° W]; 9 Jul. 1955; B. Hudson leg.; EMEC • 2 ♀♀; Kern Co., Mt Pinos; [34.816 ° N, 119.145 ° W]; elev. 2438 m; 1 Jul. 1965; J. Powell leg.; EMEC • 1 ♂; same data as for preceding; 1 Jul. 1965; J. Powell leg.; EMEC • 3 ♀♀; Lassen Co., Black’s Mountain; [36.149 ° N, 116.659 ° W]; 13 Jun. 1941; C. D. Michener leg.; EMEC • 1 ♀; Lassen Co., Cinder Cone; [40.493 ° N, 121.889 ° W]; 18 Jun. 1941; P. D. Hurd leg.; ex Ribes; EMEC • 1 ♀; Lassen Co., Cinder Cone, Lassen National Park; [40.548 ° N, 121.32 ° W]; 19 Jun. 1941; EMEC • 1 ♀; Lassen Co., Susanville; [40.416 ° N, 120.653 ° W]; 27 Jun. 1977; T. R. Haig leg.; CSCA • 2 ♂♂; Los Angeles Co., Big Pine Camp; [34.3956 ° N, 117.7339 ° W]; elev. 1850 m; 2 Aug. 1944; P. H. Timberlake leg.; UCRC (Fig. 41 C) • 3 ♂♂; same data as for preceding; 2 Aug. 1944; P. H. Timberlake leg.; ex Eriodictyon trichocalyx; UCRC • 1 ♂; same data as for preceding; 17 Jul. 1939; P. H. Timberlake leg.; UCRC • 1 ♀; Madera Co., Bass Lake; [37.338 ° N, 119.582 ° W]; 30 May 1942; E. G. Linsley leg.; ex Potentilla; UCRC • 1 ♀; Madera Co., Oakhurst; [40.493 ° N, 121.889 ° W]; 1 Jun. 1942; EMEC • 1 ♂; Madera Co., Oakhurst; [37.328 ° N, 119.649 ° W]; 26 May 1942; A. J. Walz leg.; ex Eriodictyon; UCRC • 2 ♀♀; Madera Co., Soquel Basin; [37.053 ° N, 121.923 ° W]; 11 Jun. 1942; A. J. Walz leg.; ex Salix; EMEC • 1 ♀; Marin Co., Inverness; [38.1008 ° N, 122.857 ° W]; 25 Mar. 1962; C. A. Toschi leg.; EMEC • 1 ♂; Marin Co., Mill Valley; [37.9042 ° N, 122.5421 ° W]; 8 – 11 Oct. 1965; P. H. Arnaud Jr leg.; CAS • 1 ♀; Mariposa Co., Fish Camp; [37.476 ° N, 119.635 ° W]; 21 May 1942; EMEC • 1 ♀; Mariposa Co., Miami Ranger Station; [37.419 ° N, 119.745 ° W]; 18 May 1942; E. G. Linsley leg.; UCRC • 2 ♀♀; same data as for preceding; 4 Jun. 1942; A. J. Walz leg.; UCRC • 1 ♀; same data as for preceding; 16 May 1942; E. G. Linsley leg.; ex Arctotaphylos; UCRC • 2 ♀♀; same data as for preceding; 4 Jun. 1942; E. G. Linsley leg.; UCRC • 1 ♀; same data as for preceding; 22 May 1942; A. J. Walz leg.; UCRC • 1 ♀; same data as for preceding; 29 May 1942; C. Kennett leg.; EMEC • 1 ♂; Mariposa Co., Wawona; [37.547 ° N, 119.644 ° W]; 6 Jun. 1942; A. J. Walz leg.; ex Eriogonum; EMEC • 1 ♀; same data as for preceding; 6 Jun. 1942; A. J. Walz leg.; ex Eriogonum; EMEC • 1 ♂; Mariposa Co., Yosemite; [37.863 ° N, 119.539 ° W]; elev. 1180 – 1220 m; 12 Jun. 1928; Walter Ebeling leg.; UCRC • 1 ♀; same data as for preceding; 28 May 1921; E. C. VanDyke leg.; CAS • 1 ♀; same data as for preceding; 16 May 1934; Bryant Lot leg.; CAS • 1 ♀; same data as for preceding; 1 Jun. 1921; E. C. VanDyke leg.; CAS • 2 ♂♂; Mendocino Co., Hopland Field Station; [39.001 ° N, 123.08 ° W]; 10 Jun. 1958; W. W. Middlekauff leg.; EMEC • 1 ♀; same data as for preceding; 10 Jun. 1958; W. W. Middlekauff leg.; EMEC • 1 ♀; Mendocino Co., Ryan Creek; [39.476 ° N, 123.381 ° W]; 13 Apr. 1941; EMEC • 1 ♀; Modoc Co., 1 mi. E of Pine Creek; [41.573 ° N, 120.156 ° W]; 13 Jun. 1970; J. A. Chemsak leg.; EMEC • 2 ♀♀; Modoc Co., 5 mi. N of Davis Creek; [41.832 ° N, 120.371 ° W]; 3 Jun. 1971; ex Lithospermum ruderale; EMEC • 1 ♀; Modoc Co., 5.2 mi. NW of Cedarville; [41.584 ° N, 120.099 ° W]; elev. 1981 m; 13 Jun. 1969; R. R. and R. A. Snelling leg.; ex Phacelia; EMEC • 1 ♀; Modoc Co., Buck Creek Ranger Station; [41.870 ° N, 120.293 ° W]; elev. 1570 m; 5 – 7 Jun. 1970; P. Opler leg.; ex Isatis tinctoria; EMEC • 2 ♀♀; same data as for preceding; 5 – 7 Jun. 1970; J. A. Chemsak leg.; EMEC • 2 ♀♀; Modoc Co., Cedar Pass; [41.562 ° N, 120.269 ° W]; 29 Jun. 1955; J. W. MacSwain leg.; EMEC • 2 ♀♀; same data as for preceding; 29 Jun. 1955; C. L. Hogue leg.; EMEC • 1 ♀; Modoc Co., Fandango Pass; [41.802 ° N, 120.207 ° W]; elev. 1860 m; 7 Jun. 1970; J. A. Chemsak leg.; EMEC • 1 ♀; Modoc Co., Lily Lake; [41.976 ° N, 120.21 ° W]; 11 Jul. 1957; J. Powell leg.; EMEC • 2 ♀♀; Mono Co., Blanco’s Corral, White Mountains; [37.594 ° N, 118.228 ° W]; elev. 3048 m; 5 Jul. 1953; H. Nakakihara leg.; UCRC • 1 ♀; same data as for preceding; 7 Jul. 1953; J. W. MacSwain leg.; EMEC • 1 ♂; same data as for preceding; 25 Aug. 1960; L. R. Nault leg.; EMEC • 1 ♂; Mono Co., Blanco’s Corral, White Mountains; [37.510 ° N, 118.19 ° W]; 25 Aug. 1960; P. D. Hurd leg.; ex Senecio; EMEC • 3 ♂♂; Mono Co., Bodie; [38.212 ° N, 119.011 ° W]; 21 Sep. 1958; Monke, Stange leg.; EMEC • 1 ♀; Mono Co., Fales Hot Springs; [38.351 ° N, 119.399 ° W]; elev. 2164 m; 16 Jun. 1969; R. R. and R. A. Snelling leg.; EMEC • 1 ♂; Mono Co., Leavitt Meadow; [38.336 ° N, 119.611 ° W]; 13 Aug. 1963; H. B. Leech leg.; CAS • 1 ♀; same data as for preceding; 14 Aug. 1963; H. B. Leech leg.; CAS • 1 ♀; same data as for preceding; 16 Jun. 1961; C. A. Toschi leg.; ex Ribes viscosissimum; EMEC • 1 ♀; Mono Co., Lee Vining; [37.961 ° N, 119.121 ° W]; 7 Mar. 1933; R. M. and G. E. Bohart leg.; CAS • 1 ♀; Mono Co., Tioga Pass-Hall Area site; [37.932 ° N, 119.263 ° W]; 18 Jul. 1969; A. R. Moldenke leg.; ex Phacelia hastata var. compacta (verbatim: Phacelia frigida); CAS • 1 ♂; Mono Co., 1 mi. W of Tom’s Place; [37.561 ° N, 118.699 ° W]; D. D. Linsdale leg.; EMEC • 1 ♀; Mono Co., White Mountains, 3 mi. N of Inyo Co., near Naval Research Station, N. Fork of Crooked Creek; [37.659 ° N, 118.16 ° W]; elev. 3094 m; 20 Aug. 1968; H. B. Leech leg.; CAS • 1 ♂; Napa Co., Mount Saint Helena; [38.667 ° N, 122.638 ° W]; 9 Jun. 1918; E. P. Van Duzee leg.; CAS • 2 ♀♀; Nevada Co., Donner Pass; [39.314 ° N, 120.327 ° W]; 4 Jul. 1962; J. Powell leg.; EMEC • 1 ♀; Nevada Co., Russell Valley; [39.438 ° N, 120.133 ° W]; 26 Jun. 1964; M. E. Irwin leg.; UCRC • 2 ♀♀; Nevada Co., Sagehen; 39.43413 ° N, 120.25929 ° W; 22 Jun. 2019; A. Zell leg.; ex Camassia quamash; UCDC • 1 ♀; Nevada Co., Sagehen Creek, near Hobart Mills; [39.432 ° N, 120.201 ° W]; 7 Jul. 1964; M. E. Irwin leg.; UCRC • 1 ♀; Nevada Co., Sagehen Creek; [39.432 ° N, 120.201 ° W]; 19 Jun. 1962; R. L. Wescott leg.; EMEC • 1 ♀; Nevada Co., Truckee; [39.357 ° N, 120.185 ° W]; Ximena McGlashan leg.; EMEC • 2 ♀♀; same data as for preceding; 19 Jun. 1927; H. Van Duzee leg.; CAS • 4 ♀♀; same data as for preceding; 14 Jun. 1927; E. P. Van Duzee leg.; CAS • 1 ♀; same data as for preceding; 12 Jun. 1927; E. P. Van Duzee leg.; CAS • 1 ♀; same data as for preceding; 13 Jun. 1927; E. P. Van Duzee leg.; CAS • 1 ♀; same data as for preceding; 21 Jun. 1927; E. P. Van Duzee leg.; CAS • 3 ♀♀; Nevada Co., White Cloude Campground; [39.317 ° N, 120.934 ° W]; 30 May 1964; P. H. Arnaud Jr leg.; CAS • 4 ♀♀; same data as for preceding; 31 May 1964; P. H. Arnaud Jr leg.; CAS • 1 ♂; Placer Co., 3 mi. NE of Auburn; [38.957 ° N, 121.041 ° W]; 5 May 1958; Menke and Stange leg.; EMEC • 1 ♀; Placer Co., East end of Bear Valley; [39.236 ° N, 120.597 ° W]; 1 Jun. 1964; P. H. Arnaud Jr leg.; CAS • 1 ♀; Placer Co., Pyramid Ranger Station; [39.284 ° N, 120.654 ° W]; 2 Jun. 1963; J. W. MacSwain leg.; EMEC • 1 ♀; Placer Co., Tahoe NF, 34 km NE of Foresthill; 39.1733 ° N, 120.5075 ° W; elev. 1955 m; 8 Jun. 2009; M. Buck leg.; PMAE • 3 ♀♀; Placer Co., Summit; [39.318 ° N, 120.327 ° W]; W. M. Giffard leg.; CAS • 1 ♂; Plumas Co., Little Long Valley Creek, 6 mi. E of Spring Garden; [39.909 ° N, 120.533 ° W]; 4 – 10 Aug. 1977; M. Wasbauer leg.; CAS • 4 ♀♀; Riverside Co., 12 mi. NW of Oak Grove on Hwy 79; [33.470 ° N, 116.949 ° W]; 5 Apr. 1969; Saul Frommer, Suzy Frommer, Kate Reed and Marshall Reed leg.; UCRC • 1 ♂; Riverside Co., Dark Canyon, San Jacinto Mountains; [33.775 ° N, 116.777 ° W]; 6 Jun. 1939; E. S. Ross leg.; EMEC • 2 ♀♀; Riverside Co., Gavilan Hills; [33.7944 ° N, 117.3558 ° W]; elev. 620 m; 21 Mar. 1941; P. H. Timberlake leg.; ex Rhus trilobata; UCRC • 3 ♀♀; same data as for preceding; 19 Mar. 1936; P. H. Timberlake leg.; ex Rhus trilobata; UCRC • 3 ♀♀; same data as for preceding; 7 Apr. 1939; P. H. Timberlake leg.; ex Cryptantha intermedia; UCRC • 1 ♀; same data as for preceding; 2 Mar. 1937; P. H. Timberlake leg.; ex Ribes indecorum; UCRC • 1 ♀; same data as for preceding; 2 Mar. 1937; P. H. Timberlake leg.; ex Salix vestita (verbatim: Salix leiolepis); UCRC • 1 ♂; Riverside Co., Idyllwild; [33.744 ° N, 116.718 ° W]; Jul. 1936; E. S. Ross leg.; CAS • 1 ♂; Riverside Co., Idyllwild, San Jacinto Mountains; [33.744 ° N, 116.718 ° W]; 15 Jun. 1941; E. C. VanDyke leg.; CAS • 3 ♀♀; Riverside Co., Keen Camp, San Jacinto Mountains; [33.7011 ° N, 116.715 ° W]; 10 Jun. 1939; J. G. Shanafelt leg.; EMEC • 1 ♀; Riverside Co., Millard Canyon; [34.205 ° N, 118.166 ° W]; elev. 762 – 853 m; 26 Apr. 1969; B. Hunt leg.; ex Salix; UCRC • 5 ♀♀; Riverside Co., Pine Cove; [33.7586 ° N, 116.7389 ° W]; 3 Jun. 1939; P. H. Timberlake leg.; ex Arctostaphylos pringlei spp. drupacea (verbatim: Arctostaphylos drupacea); UCRC • 1 ♂; Riverside Co., San Jacinto Mt Trail, San Jacinto Mts; [33.812 ° N, 116.679 ° W]; 1 Jul. 1952; J. F. Powers leg.; EMEC • 3 ♀♀; San Benito Co., Pinnacles; [36.529 ° N, 121.145 ° W]; 25 Mar. 1940; E. G. Linsley leg.; ex Cryptantha; EMEC • 1 ♀; San Bernardino Co., 15 km SSE of Hesperia, Hwy 173; [34.337 ° N, 117.259 ° W]; elev. 1022 m; 21 Apr. 1981; J. A. Bethke leg.; UCRC • 1 ♀; San Bernardino Co., 8 mi. N of Fontana; [34.192 ° N, 117.431 ° W]; 14 Apr. 1965; C. A. Toschi leg.; EMEC • 1 ♀; San Bernardino Co., Above Lake Arrowhead; [34.270 ° N, 117.17 ° W]; 16 Jul. 1933; UCRC • 1 ♂; San Bernardino Co., Big Bear Valley; [34.2650 ° N, 116.8483 ° W]; elev. 2055 m; 9 Aug. 1933; P. H. Timberlake leg.; UCRC • 1 ♂; same data as for preceding; 8 Aug. 1933; P. H. Timberlake leg.; ex Eriogonum umbellatum; UCRC • 1 ♂; San Bernardino Co., Grass Valley, San Bernardino Mountains; [34.255 ° N, 117.217 ° W]; 30 Jul. 1961; D. P. Gregory leg.; ex Oenothera elata subsp. hookeri (verbatim: Oenothera hookeri); UCRC • 4 ♀♀; San Bernardino Co., Mill Creek; [34.0825 ° N, 116.8936 ° W]; elev. 1830 m; 16 May 1948; P. H. Timberlake leg.; ex Arctostaphylos patula; UCRC • 1 ♀; same data as for preceding; 28 Apr. 1946; P. H. Timberlake leg.; ex Arctostaphylos patula; UCRC • 2 ♀♀; same data as for preceding; 2 Apr. 1961; P. H. Timberlake leg.; ex Arctostaphylos patula; UCRC • 4 ♀♀; same data as for preceding; 8 May 1949; P. H. Timberlake leg.; ex Arctostaphylos patula; UCRC • 1 ♀; same data as for preceding; 23 May 1948; P. H. Timberlake leg.; ex Arctostaphylos patula; UCRC • 4 ♀♀; same data as for preceding; 2 Apr. 1961; P. H. Timberlake leg.; ex Arctostaphylos patula; UCRC • 1 ♀; same data as for preceding; 3 Apr. 1961; P. H. Timberlake leg.; ex Arctostaphylos patula; UCRC • 1 ♀; same data as for preceding; 16 May 1948; P. H. Timberlake leg.; ex Arctostaphylos patula; UCRC • 2 ♀♀; San Bernardino Co., Mill Creek; [34.0806 ° N, 116.8875 ° W]; elev. 1890 m; 22 May 1940; P. H. Timberlake leg.; ex Arctostaphylos pringlei subsp. drupacea (verbatim: Arctostaphylos drupacea); UCRC • 3 ♀♀; same data as for preceding; 29 May 1938; P. H. Timberlake leg.; ex Arctostaphylos pringlei subsp. drupacea (verbatim: Arctostaphylos drupacea); UCRC • 2 ♀♀; same data as for preceding; 29 May 1938; P. H. Timberlake leg.; ex Arctostaphylos pringlei subsp. drupacea (verbatim: Arctostaphylos drupacea); UCRC • 2 ♀♀; same data as for preceding; 30 May 1940; P. H. Timberlake leg.; ex Arctostaphylos pringlei subsp. drupacea (verbatim: Arctostaphylos drupacea); UCRC • 1 ♀; San Bernardino Co., Mill Creek; [34.0825 ° N, 116.8936 ° W]; elev. 1830 m; 4 Jun. 1951; P. H. Timberlake leg.; ex Ceanothus cordulatus; UCRC • 1 ♀; same data as for preceding; 16 May 1938; P. H. Timberlake leg.; ex Ceanothus cordulatus; UCRC • 1 ♀; same data as for preceding; 27 May 1951; P. H. Timberlake leg.; ex Eremocarya lepida (verbatim: Cryptantha lepida); UCRC • 2 ♂♂; same data as for preceding; 10 Sep. 1945; P. H. Timberlake leg.; ex Eriogonum wrightii var. subscaposum (verbatim: Eriogonum subscaposum); UCRC • 1 ♀; San Bernardino Co., Mill Creek; [34.0781 ° N, 116.8786 ° W]; elev. 1980 m; 20 Jun. 1956; P. H. Timberlake leg.; ex Penstemon grinnelli; UCRC • 1 ♀; San Bernardino Co., Mill Creek; [34.0825 ° N, 116.8936 ° W]; elev. 1830 m; 22 Oct. 1950; P. H. Timberlake leg.; ex Penstemon grinnelli; UCRC • 1 ♀; same data as for preceding; 6 Jun. 1940; P. H. Timberlake leg.; ex Frangula californica (verbatim: Rhamnus californica); UCRC • 2 ♀♀; same data as for preceding; 16 May 1948; P. H. Timberlake leg.; ex Ribes roezlii; UCRC • 1 ♀; same data as for preceding; 6 Sep. 1955; P. H. Timberlake leg.; UCRC • 1 ♀; San Bernardino Co., near Crestline; [34.2403 ° N, 117.2894 ° W]; 30 Jul. 1935; P. H. Timberlake leg.; ex Heracleum maximum (verbatim: Heracleum lanatum); UCRC • 1 ♂; same data as for preceding; 30 Jun. 1935; P. H. Timberlake leg.; ex Eriophyllum lanatum; UCRC • 1 ♀; San Bernardino Co., Rim of the World; [34.227 ° N, 117.245 ° W]; 14 Sep. 1934; ex Ericameria nauseosa var. bernardina (verbatim: Crysothamnus bernardinus); UCRC • 4 ♀♀; San Bernardino Co., San Bernardino Mountains, Van Dusen Canyon, near Holcomb; [34.302 ° N, 116.855 ° W]; 1 Jun. 1969; E. I. Schlinger leg.; ex Ribes; EMEC • 1 ♀; San Bernardino Co., Tetley’s Mountain Camp, Crestline; [34.2464 ° N, 117.3094 ° W]; elev. 1380 m; 15 May 1937; P. H. Timberlake leg.; ex Malus; UCRC • 2 ♂♂; San Bernardino Co., Valley of the Falls; [34.0833 ° N, 116.9125 ° W]; elev. 1745 m; 28 Aug. 1935; P. H. Timberlake leg.; UCRC • 1 ♀; San Bernardino Co., Van Dusen Canyon; [34.294 ° N, 116.905 ° W]; elev. 2150 – 2180 m; 1 Jun. 1969; W. A. Hunt leg.; ex Erysimum; UCRC • 1 ♀; San Bernardino Co., Van Dusen Creek, Holcomb Valley; [34.302 ° N, 116.893 ° W]; elev. 1700 m; 1 Jun. 1969; G. Goddard leg.; UCRC • 1 ♀; San Diego Co., 5 mi. N of Santa Ysabel; [33.175 ° N, 116.675 ° W]; 7 Feb. 1952; J. D. Rozen leg.; ex Salix; EMEC • 1 ♀; San Diego Co., Coastal Sage site; [32.716 ° N, 117.256 ° W]; A. R. Moldenke leg.; CAS • 1 ♀; San Diego Co., Cuyamaca; [32.928 ° N, 116.568 ° W]; 19 Apr. 1950; J. W. MacSwain leg.; EMEC • 1 ♀; same data as for preceding; 19 Apr. 1950; E. G. Linsley leg.; ex Lasthenia (verbatim: Baeria); EMEC • 1 ♀; San Diego Co., Cuyamaca Rancho State Park; [32.933 ° N, 116.567 ° W]; 10 Jun. 1964; L. D. Anderson leg.; UCRC • 2 ♀♀; San Diego Co., Laguna Mountains; [32.840 ° N, 116.438 ° W]; 3 May 1946; John S. Perry leg.; UCRC • 1 ♀; San Diego Co., Mt Laguna; [32.867 ° N, 116.42 ° W]; 21 Jun. 1963; J. Powell leg.; EMEC • 1 ♀; same data as for preceding; 21 Jun. 1963; P. D. Hurd leg.; EMEC • 1 ♂; same data as for preceding; 5 Jul. 1963; R. L. Langston leg.; EMEC • 1 ♀; San Diego Co., Mt Laguna site; [32.867 ° N, 116.42 ° W]; A. R. Moldenke leg.; CAS • 1 ♀; San Diego Co., Mt Palomar; [33.363 ° N, 116.836 ° W]; 28 Jun. 1963; P. D. Hurd leg.; ex Cryptantha intermedia; EMEC • 1 ♀; San Diego Co., Ramona; [40.493 ° N, 121.889 ° W]; 19 Apr. 1950; E. G. Linsley leg.; ex Cryptantha; EMEC • 1 ♀; Santa Clara Co., 1 mi. E of Mt Hamilton; [37.341 ° N, 121.625 ° W]; 6 May 1957; Don Burdick leg.; EMEC • 1 ♀; Santa Clara Co., Isabel Creek. Mt Hamilton; [37.3573 ° N, 121.6549 ° W]; 1 May 1956; D. J. Burdick leg.; EMEC • 1 ♀; Santa Clara Co., San Antonio R. S.; [37.47 ° N, 121.81 ° W]; 11 Mar. 1948; P. D. Hurd leg.; EMEC • 1 ♀; Shasta Co., 4 mi. S of Hat Lake; [40.77 ° N, 121.516 ° W]; 5 Jun. 1941; P. D. Hurd leg.; ex Wyethia; EMEC • 6 ♀♀; Shasta Co., Castle Rock State Park; [37.231 ° N, 122.096 ° W]; 3 May 1953; J. D. Lattin leg.; EMEC • 1 ♀; Shasta Co., Hat Lake; [40.509 ° N, 121.466 ° W]; 20 Jun. 1941; P. D. Hurd leg.; ex Ribes; EMEC • 3 ♀♀; same data as for preceding; 3 Jun. 1944; P. D. Hurd leg.; ex Salix; EMEC • 1 ♀; Shasta Co., Hat Lake; [40.493 ° N, 121.889 ° W]; 14 Jun. 1941; P. D. Hurd leg.; ex Salix; EMEC • 2 ♀♀; Shasta Co., Hat Lake; [40.509 ° N, 121.466 ° W]; 3 Jun. 1944; C. D. Michener leg.; ex Arctostaphylos; EMEC • 6 ♀♀; same data as for preceding; 14 Jun. 1941; C. D. Michener leg.; ex Salix; EMEC • 4 ♀♀; same data as for preceding; 3 Jun. 1941; C. D. Michener leg.; ex Salix; EMEC • 2 ♀♀; same data as for preceding; 3 Jun. 1941; P. D. Hurd leg.; ex Salix; EMEC • 1 ♀; same data as for preceding; 4 Jun. 1941; P. D. Hurd leg.; EMEC • 1 ♀; same data as for preceding; 14 Jun. 1941; C. W. Anderson leg.; ex Ribes; EMEC • 2 ♀♀; same data as for preceding; 14 Jun. 1941; C. W. Anderson leg.; ex Ribes cereum; EMEC • 1 ♀; Shasta Co., Lassen National Park, Hat Lake; [40.509 ° N, 121.466 ° W]; 14 Jun. 1941; C. W. Anderson leg.; ex Ribes cereum; EMEC • 1 ♀; Shasta Co., Mountains west of Nieubieber; [41.117 ° N, 121.35 ° W]; 4 Jun. 1938; E. C. VanDyke leg.; CAS • 1 ♀; Shasta Co., Shingletown; [40.493 ° N, 121.889 ° W]; 7 Jun. 1941; P. D. Hurd leg.; EMEC • 1 ♂; same data as for preceding; 2 Jun. 1941; C. D. Michener leg.; EMEC • 1 ♀; same data as for preceding; 21 Jun. 1941; EMEC • 1 ♀; same data as for preceding; 28 May 1941; E. G. Linsley leg.; EMEC • 2 ♀♀; Shasta Co., Viola; [40.518 ° N, 121.678 ° W]; 18 May 1941; C. D. Michener leg.; EMEC • 3 ♀♀; same data as for preceding; 19 May 1941; C. D. Michener leg.; ex Salix; EMEC • 2 ♀♀; same data as for preceding; 19 May 1941; P. D. Hurd leg.; EMEC • 2 ♀♀; same data as for preceding; 18 May 1941; C. D. Michener leg.; ex Taraxacum officinale (verbatim: Taraxacum vulgare); EMEC • 2 ♀♀; same data as for preceding; 18 May 1941; C. D. Michener leg.; EMEC • 3 ♀♀; Shasta Co.; [40.75 ° N, 122.12 ° W]; 2 May 1913; F. W. Nunenmacher leg.; EMEC • 1 ♀; Sierra Co., Carmen; 39.69693 ° N, 120.45803 ° W; 7 Jun. 2019; M. Page leg.; ex Camassia quamash; UCDC • 1 ♀; Sierra Co., Gold Lake; [39.672 ° N, 120.66 ° W]; 13 Jul. 1921; C. L. Fox leg.; CAS • 1 ♂; same data as for preceding; 23 Jul. 1921; C. L. Fox leg.; CAS • 1 ♀; same data as for preceding; 14 Jul. 1921; C. L. Fox leg.; CAS • 1 ♀; Sierra Co., Harvey South; 39.69379 ° N, 120.43872 ° W; 3 Jun. 2019; M. Page leg.; ex Camassia quamash; UCDC • 1 ♀; same data as for preceding; 7 Jun. 2019; D. Deterding leg.; ex Camassia quamash; UCDC • 1 ♀; Sierra Co., Holcomb; 39.48892 ° N, 120.26687 ° W; 15 Jun. 2019; A. Zell leg.; ex Camassia quamash; UCDC • 1 ♀; Sierra Co., Mitchell; 39.47896 ° N, 120.28642 ° W; 1 Jul. 2019; M. Page leg.; ex Penstemon rydbergii; UCDC • 1 ♀; Siskiyou Co., 1 mi. SE of Bartle; [41.249 ° N, 121.805 ° W]; 7 – 9 Jun. 1974; R. Coville leg.; EMEC • 1 ♀; same data as for preceding; 7 – 9 Jun. 1974; L. Green leg.; EMEC • 1 ♀; Siskiyou Co., Ash Creek Ranger Station; [41.270 ° N, 121.944 ° W]; 7 – 9 Jun. 1974; J. Sorensen leg.; EMEC • 1 ♀; Siskiyou Co., Ash Creek Ranger Station; [41.27 ° N, 121.944 ° W]; elev. 1067 m; 7 – 9 Jun. 1974; D. Green leg.; ex Trifolium; EMEC • 1 ♀; Siskiyou Co., Castle Lake; [41.228 ° N, 122.384 ° W]; 22 Jul. 1958; J. Powell leg.; EMEC • 4 ♂♂; Siskiyou Co., Headwaters E. Fork of S. Fork Salmon River, Cecilville-Callahan Road; [41.222 ° N, 123.05 ° W]; elev. 1829 m; 31 Jul. 1968; H. B. Leech leg.; CAS • 2 ♂♂; same data as for preceding; 30 Jul. 1968; H. B. Leech leg.; CAS • 3 ♀♀; same data as for preceding; 30 Jul. 1968; H. B. Leech leg.; CAS • 1 ♂; Siskiyou Co., McBride Springs; [41.353 ° N, 122.282 ° W]; elev. 1585 m; 10 – 14 Jun. 1974; J. Chemsak and J. Doyen leg.; EMEC • 1 ♀; same data as for preceding; 10 – 14 Jun. 1974; R. Coville leg.; EMEC • 5 ♀♀; same data as for preceding; 10 – 14 Jun. 1974; L. Green leg.; EMEC • 5 ♂♂; Siskiyou Co., Road to Taylor Lake S of Sawyers, Bar-Etna Road; [41.256 ° N, 123.086 ° W]; elev. 1737 m; 28 Jul. 1968; H. B. Leech leg.; CAS • 5 ♀♀; same data as for preceding; 28 Jul. 1968; H. B. Leech leg.; CAS • 1 ♂; Siskiyou Co., Shasta Springs; [41.247 ° N, 122.261 ° W]; 17 Jun. 1920; C. L. Fox leg.; CAS • 1 ♀; Sonoma Co., Trinity P. O.; [38.293 ° N, 122.452 ° W]; 3 Apr. 1937; R. M. and G. E. Bohart leg.; CAS • 1 ♀; Stanislaus Co., Del Puerto Canyon; [37.477 ° N, 121.23 ° W]; 2 Apr. 1961; R. R. and M. D. Snelling leg.; OSAC • 2 ♂♂; Trinity Co., Carrville; [40.859 ° N, 122.705 ° W]; elev. 732 – 762 m; 9 Jun. 1984; E. C. VanDyke leg.; CAS • 2 ♀♀; Tulare Co., 2 mi. E of Johnsondale; [35.970 ° N, 118.498 ° W]; 27 Apr. 1964; C. A. Toschi leg.; EMEC • 1 ♀; same data as for preceding; 27 Apr. 1964; P. Rude leg.; ex Salix; EMEC • 4 ♀♀; same data as for preceding; 27 Apr. 1964; C. A. Toschi leg.; ex Prunus subcordata; EMEC • 8 ♀♀; same data as for preceding; 27 Apr. 1964; J. Powell leg.; ex Prunus subcordata; EMEC • 1 ♀; Tulare Co., Giant Forest; [36.5625 ° N, 118.75139 ° W]; elev. 2130 m; 12 Jun. 1939; P. H. Timberlake leg.; ex Arctostaphylos patula; UCRC • 1 ♀; Tulare Co., Portuguese Pass; [35.800 ° N, 118.582 ° W]; elev. 2234 m; 14 Jun. 1961; G. I. Stage and R. R. Snelling leg.; EMEC • 1 ♂; Tulare Co., Redwood Meadows; [36.541 ° N, 118.635 ° W]; 3 Aug. 1923; C. L. Fox leg.; CAS • 2 ♀♀; Tulare or Fresno Co., General Grant National Park; [36.747 ° N, 118.975 ° W]; 26 Jun. 1929; P. H. Timberlake leg.; ex Senecio lugens; UCRC • 1 ♀; Tuolumne Co., 2 mi. W of Sonora Pass; [38.323 ° N, 119.682 ° W]; 16 Aug. 1959; G. I. Stage and R. R. Snelling leg.; ex Angelica capitellata (verbatim: Sphenosciadium capitellatum); EMEC • 2 ♂♂; Tuolumne Co., 4 mi. W of Sonora Pass; [38.327 ° N, 119.71 ° W]; 8 Aug. 1959; G. I. Stage and R. R. Snelling leg.; EMEC • 1 ♀; Tuolumne Co., 5 mi. NW of Columbia; [38.085 ° N, 120.467 ° W]; 6 May 1953; R. R. Snelling leg.; ex Eriodictyon californicum; EMEC • 3 ♀♀; Tuolumne Co., Buck Meadows – Mather site; [37.820 ° N, 120.067 ° W]; A. R. Moldenke leg.; CAS • 6 ♀♀; Tuolumne Co., Chinese Camp; [37.87 ° N, 120.433 ° W]; 2 Apr. 1960; R. R. Snelling leg.; EMEC • 1 ♀; Tuolumne Co., Chipmunk Flat; [38.324 ° N, 119.693 ° W]; 13 Jun. 1962; C. A. Toschi leg.; EMEC • 1 ♀; same data as for preceding; 9 Aug. 1966; J. W. MacSwain leg.; ex Ranunculus; EMEC • 1 ♀; Tuolumne Co., Columbia; [38.036 ° N, 120.401 ° W]; 2 Apr. 1960; R. R. and M. D. Snelling leg.; ex Ceanothus; EMEC • 1 ♀; Tuolumne Co., Lyons Reservoir; [38.046 ° N, 120.153 ° W]; 3 Jun. 1963; P. H. Arnaud Jr leg.; CAS • 1 ♀; Tuolumne Co., Pinecrest; [38.192 ° N, 119.994 ° W]; 18 Jul. 1953; J. G. Rozen leg.; EMEC • 7 ♂♂; Tuolumne Co., Pinecrest; [38.192 ° N, 119.995 ° W]; 20 Jul. 1929; P. H. Timberlake leg.; ex Solidago velutina subsp. californica (verbatim: Solidago californica); UCRC • 1 ♀; Tuolumne Co., Sonora Pass; [38.328 ° N, 119.637 ° W]; 9 Aug. 2000; John M. Burns leg.; EMEC • 1 ♀; Tuolumne Co., Strawberry; [38.200 ° N, 120.005 ° W]; 18 Jun. 1957; J. W. MacSwain leg.; EMEC • 1 ♀; same data as for preceding; 15 Jun. 1957; John M. Burns leg.; EMEC • 1 ♀; same data as for preceding; 18 Jun. 1951; R. W. Morgan leg.; EMEC • 3 ♀♀; Tuolumne Co., Tuolumne City; [37.962 ° N, 120.242 ° W]; 23 Mar. 1966; J. Powell leg.; EMEC • 1 ♀; Ventura Co., Chuchupate River Station; [34.8078 ° N, 119.0117 ° W]; elev. 1610 m; 8 May 1959; P. H. Timberlake leg.; ex Sisymbrium; UCRC • 1 ♀; Ventura Co., Mt Pinos; [34.816 ° N, 119.145 ° W]; elev. 2286 m; 8 May 1959; J. R. Powers leg.; EMEC. – Colorado • 2 ♀♀; Archuleta Co., 14 mi. WSW of Pagosa Springs; [37.191 ° N, 107.245 ° W]; 29 May 1977; U. Lanham and P. Byron leg.; ex Prunus virginiana; UCMC • 1 ♀; same data as for preceding; 27 May 1977; U. Lanham and P. Byron leg.; ex Amelanchier; UCMC • 2 ♀♀; same data as for preceding; 27 May 1977; U. Lanham and P. Byron leg.; ex Prunus virginiana; UCMC • 1 ♀; Boulder Co., 1 mi. NW of Boulder; [40.027 ° N, 105.286 ° W]; 17 Apr. 1963; U. N. and C. C. Lanham leg.; ex Salix; UCMC • 1 ♀; Boulder Co., 1 mi. SW of Boulder; [40.003 ° N, 105.285 ° W]; 17 Apr. 1963; U. N. and C. C. Lanham leg.; ex Salix; UCMC • 1 ♀; Boulder Co., 1 mi. W of Gold Hill; [40.063 ° N, 105.429 ° W]; 4 Jun. 1963; U. N. Lanham and B. Vogel leg.; ex Euphorbia esula (verbatim: Esula); UCMC • 8 ♀♀; Boulder Co., 2 mi. E of Ward; [40.072 ° N, 105.471 ° W]; 31 May 1963; U. N. Lanham and B. Vogel leg.; ex Barbarea; UCMC • 1 ♀; Boulder Co., 2 mi. SW of Boulder; [39.994 ° N, 105.297 ° W]; elev. 1981 m; 18 May 1963; U. N. Lanham leg.; UCMC • 1 ♀; Boulder Co., 2 mi. W of Boulder; [40.015 ° N, 105.308 ° W]; elev. 1676 m; 22 May 1963; U. N. Lanham and B. Nielsen leg.; UCMC • 1 ♀; same data as for preceding; 18 May 1963; U. N. Lanham et al. leg.; UCMC • 2 ♀♀; Boulder Co., 4 km N of Lyons; 40.24847 ° N, 105.29742 ° W; elev. 1705 m; 19 May 2007; C. Sheffield and J. Gibbs leg.; RSKM • 3 ♀♀; Boulder Co., 4 mi. N of Boulder; [40.073 ° N, 105.271 ° W]; elev. 1676 m; 21 May 1980; U. Lanham and P. Robinson leg.; ex Physaria belli; UCMC • 1 ♀; same data as for preceding; 23 May 1980; U. N. Lanham leg.; ex Physaria belli; UCMC • 4 ♀♀; same data as for preceding; 25 May 1980; K. Noff leg.; ex Physaria belli; UCMC • 1 ♀; same data as for preceding; 25 May 1980; P. Robinson leg.; ex yellow crucifer; UCMC • 2 ♀♀; same data as for preceding; 10 May 1980; U. N. Lanham leg.; ex Physaria belli; UCMC • 7 ♀♀; Boulder Co., 4 mi. SW of Boulder; [39.974 ° N, 105.324 ° W]; elev. 2042 m; 11 May 1963; B. Vogel leg.; ex Ribes; UCMC • 6 ♀♀; Boulder Co., 4 mi. W of Boulder; [40.015 ° N, 105.347 ° W]; elev. 1676 m; 25 May 1980; K. Noff leg.; ex Physaria belli; UCMC • 1 ♀; Boulder Co., 5 mi. N of Boulder; [40.089 ° N, 105.27 ° W]; 9 May 1939; Rodeck and Lanham leg.; UCMC • 2 ♀♀; Boulder Co., 5 mi. NW of Boulder; [40.066 ° N, 105.337 ° W]; 31 May 1963; U. N. Lanham and B. Vogel leg.; ex Barbarea; UCMC • 4 ♀♀; same data as for preceding; 31 May 1963; U. N. Lanham and B. Vogel leg.; ex Prunus virginiana; UCMC • 1 ♀; same data as for preceding; 31 May 1963; U. N. Lanham and B. Vogel leg.; ex Crataegus; UCMC • 1 ♀; Boulder Co., 6 mi. N of Boulder; [40.103 ° N, 105.27 ° W]; elev. 2103 m; 21 Apr. 1981; U. N. Lanham leg.; ex Salix; UCMC • 1 ♀; Boulder Co., 6 mi. NW of Boulder; [40.077 ° N, 105.351 ° W]; elev. 1829 m; 22 May 1981; U. Lanham leg.; ex Physaria vitulifera; UCMC • 1 ♂; Boulder Co., 6 mi. S of Ward; [40.0119 ° N, 105.5056 ° W]; elev. 2710 m; 1 Jul. 1939; P. H. Timberlake leg.; ex Potentilla glandulosa; UCRC • 3 ♀♀; Boulder Co., Allenspark; [40.194 ° N, 105.526 ° W]; 19 Jun. 1963; U. N. Lanham leg.; ex Prunus virginiana; UCMC • 1 ♀; same data as for preceding; 19 Jun. 1963; U. N. Lanham leg.; ex Barbarea; UCMC • 1 ♂; Boulder Co., Beaver Reservoir; [40.1206 ° N, 105.5228 ° W]; elev. 2800 m; 1 Jul. 1939; P. H. Timberlake leg.; ex Drymocallis glandulosa (verbatim: Potentilla glandulosa); UCRC • 1 ♀; Boulder Co., Boulder; [40.02 ° N, 105.27 ° W]; 14 May 1939; R. Bauer leg.; UCMC • 1 ♀; same data as for preceding; 13 May 1913; M. M. Ellis leg.; UCRC • 1 ♀; same data as for preceding; 22 May 1913; M. M. Ellis leg.; UCRC • 1 ♀; same data as for preceding; 28 Apr. 1913; M. D. Ellis leg.; ex Prunus; UCRC • 1 ♀; same data as for preceding; 6 May 1913; M. D. Ellis leg.; ex Malus; UCRC • 2 ♀♀; same data as for preceding; 24 May 1913; M. D. Ellis leg.; UCRC • 1 ♀; same data as for preceding; 23 May 1913; M. D. Ellis leg.; UCRC • 1 ♀; same data as for preceding; 28 Apr. 1913; M. D. Ellis leg.; ex Ribes; UCRC • 1 ♀; same data as for preceding; 21 May 1913; M. D. Ellis leg.; UCRC • 1 ♀; same data as for preceding; 4 May 1906; S. G. Robins (?) leg.; UCRC • 1 ♀; same data as for preceding; 20 May 1933; H. W. Campbell leg.; ex Taraxacum; UCRC • 1 ♀; same data as for preceding; 17 May 1933; H. W. Campbell leg.; ex Taraxacum; UCRC • 1 ♀; same data as for preceding; 2 Apr. 1933; H. W. Campbell leg.; UCRC • 1 ♀; same data as for preceding; May 1920 (?); Backer (?) leg.; UCRC • 1 ♀; same data as for preceding; 12 May 1922; M. Pope leg.; UCRC • 1 ♀; same data as for preceding; 20 May 1922; M. Pope leg.; UCRC • 1 ♀; same data as for preceding; 12 May 1922; D. W. MaC (?) leg.; UCRC • 1 ♀; same data as for preceding; 21 May 1922; E. M. Foster leg.; UCRC • 1 ♀; same data as for preceding; 31 May 1980; P. Robinson leg.; ex Senecio; UCMC • 1 ♀; same data as for preceding; 23 May 1978; U. Lanham leg.; ex Prunus virginiana; UCMC • 1 ♀; same data as for preceding; 16 May 1965; U. N. Lanham leg.; ex Ribes; UCMC • 1 ♀; same data as for preceding; 16 May 1965; U. N. Lanham leg.; ex Malus; UCMC • 1 ♀; same data as for preceding; 3 Jun. 1938; U. Lanham leg.; UCMC • 1 ♀; same data as for preceding; 17 May 1963; U. N. Lanham leg.; ex Prunus virginiana; UCMC • 1 ♀; same data as for preceding; 24 Apr. 1974; U. N. Lanham leg.; UCMC • 3 ♀♀; same data as for preceding; 30 Apr. 1925; C. H. Hicks leg.; UCMC • 1 ♀; same data as for preceding; 8 May 1925; C. H. Hicks leg.; UCMC • 1 ♀; same data as for preceding; 12 May 1925; C. H. Hicks leg.; UCMC • 3 ♀♀; same data as for preceding; 4 May 1925; C. H. Hicks leg.; UCMC • 1 ♀; same data as for preceding; 17 May 1925; C. H. Hicks leg.; UCMC • 1 ♀; same data as for preceding; 14 Apr. 1907; S. A. Rohwer leg.; ex Rubus; EMEC • 2 ♀♀; same data as for preceding; 14 May 1907; S. A. Rohwer leg.; ex Ribes; MCZ • 1 ♀; same data as for preceding; 16 May 1907; S. A. Rohwer leg.; ex Ribes (cultivated); MCZ • 1 ♀; same data as for preceding; 12 Apr; Beanutt leg.; ex Taraxacum; MCZ • 1 ♀; same data as for preceding; 21 May; T. D. A. Cockerell leg.; ex Rubus deliciosus; CAS • 1 ♀; same data as for preceding; 21 May; T. D. A. Cockerell leg.; ex Rubus deliciosus; CMNH • 1 ♀; same data as for preceding; 20 Apr; W. P. Cockerell leg.; ex Claytonia perfoliata; CMNH • 1 ♀; same data as for preceding; 20 Apr.; T. D. A. Cockerell leg.; ex Clematis hirsutissima (verbatim: Pulsatilla hirsutissima); CMNH • 1 ♀; Boulder Co., near Boulder; [40.01 ° N, 105.27 ° W]; 2 May 1974; U. N. Lanham leg.; UCMC • 1 ♀; Boulder Co., White Rocks; [40.062 ° N, 105.132 ° W]; 4 May 1925; C. H. Hicks leg.; UCMC • 3 ♀♀; Chaffee Co., Maysville; [38.534 ° N, 106.171 ° W]; 18 Jun.; M. Stach leg.; CAS • 1 ♀; Chaffee Co., Poncha Pass; [38.534 ° N, 106.171 ° W]; 20 Jun. 1933; M. Stach leg.; CAS • 1 ♀; same data as for preceding; 20 Jun. 1933; W. Steele leg.; CAS • 1 ♀; same data as for preceding; 20 Jun. 1933; C. A. Wagner leg.; CAS • 7 ♀♀; Chaffee Co., Poncha Springs; [38.512 ° N, 106.077 ° W]; 24 May 1977; U. Lanham and P. Byron leg.; ex Taraxacum; UCMC • 1 ♀; Conejos Co., 9 mi. W of Capulin; [37.283 ° N, 106.276 ° W]; 25 May 1977; U. Lanham and P. Byron leg.; ex Ribes; UCMC • 4 ♀♀; same data as for preceding; 12 May 1974; U. N. Lanham leg.; ex Salix; UCMC • 4 ♀♀; same data as for preceding; 12 May 1974; U. N. Lanham leg.; ex Ribes; UCMC • 1 ♂; Custer Co., 6 mi. NNW of San Isabel; [38.159 ° N, 105.153 ° W]; 3 Jul. 1975; U. N. Lanham leg.; ex Prunus virginiana var. demissa (verbatim: Prunus melanocarpus); UCMC • 1 ♀; Delta Co., Grand Mesa; [39.040 ° N, 107.95 ° W]; 8 Jul. 1938; R. Bauer leg.; UCMC • 1 ♂; Delta Co., Lands End, Grand Mesa; [39.026 ° N, 108.223 ° W]; 28 Jul. 1962; U. N. Lanham et al. leg.; UCMC • 1 ♀; Dolores Co., 6 mi. E of Cahone; [37.690 ° N, 108.771 ° W]; elev. 2500 m; 28 May 1977; U. Lanham and P. Byron leg.; ex Amelanchier; UCMC • 1 ♀; Douglas Co., 10 mi. NE of Deckers; [39.360 ° N, 105.095 ° W]; elev. 2134 m; 6 May 1964; U. N. Lanham leg.; ex Salix; UCMC • 4 ♀♀; Douglas Co., 10 mi. SSE of Franktown; [39.257 ° N, 104.679 ° W]; 26 Apr. 1963; U. N. Lanham leg.; ex Salix; UCMC (Fig. 39) • 1 ♀; same data as for preceding; 5 May 1963; U. N. Lanham leg.; ex Salix; UCMC • 17 ♀♀; Douglas Co., 5 mi. SSW of Franktown; [39.324 ° N, 104.789 ° W]; 12 Apr. 1963; U. N. Lanham et al. leg.; ex Salix; UCMC (Fig. 40 A) • 5 ♀♀; same data as for preceding; 26 Apr. 1963; U. N. Lanham leg.; ex Salix; UCMC • 1 ♀; same data as for preceding; 7 May 1964; U. N. Lanham leg.; ex Salix; UCMC • 1 ♀; Elbert Co., Running Creek Field Station; [39.502 ° N, 104.561 ° W]; Susan Condie leg.; WRME • 1 ♀; Grand Co., 2 mi. E of Hot Sulfur Springs; [40.074 ° N, 106.065 ° W]; 20 Jun. 1963; B. Vogel leg.; ex Prunus virginiana; UCMC • 4 ♀♀; Grand Co., 2 mi. S of Kremmling; [40.030 ° N, 106.391 ° W]; elev. 2286 m; 20 May 1964; U. N. Lanham leg.; ex Salix; UCMC • 3 ♀♀; Huerfano Co., 20 mi. W of Walsenberg; [37.623 ° N, 105.144 ° W]; 24 Apr. 1963; B. Vogel and S. M. Sutton leg.; ex Salix; UCMC • 2 ♀♀; Huerfano Co., 3 mi. W of Walsenburg; [37.618 ° N, 104.836 ° W]; 30 Apr. 1981; U. N. Lanham leg.; ex Ribes; UCMC • 7 ♀♀; same data as for preceding; 30 Apr. 1981; U. N. Lanham leg.; ex Salix; UCMC • 1 ♀; Jefferson Co., 10 mi. SE of Bailey; [39.299 ° N, 105.345 ° W]; 12 Jun. 1963; U. N. Lanham leg.; ex Prunus virginiana; UCMC • 1 ♀; Jefferson Co., Morrison; [39.653 ° N, 105.191 ° W]; 6 Jun. 1952; R. S. Beal leg.; EMEC • 1 ♀; Jefferson Co., Plainview; [39.8958 ° N, 105.2788 ° W]; 6 Apr. 1963; B. Vogel and S. M. Sutton leg.; ex Pulsatilla nuttalliana (verbatim: Anemone patens); UCMC • 1 ♀; Jefferson Co., Plainview, 13 mi. N of Golden; [39.944 ° N, 105.221 ° W]; 23 May 1962; J. W. MacSwain leg.; ex Crepis runcinata; EMEC • 1 ♂; La Plata Co., 16.2 mi. W of Durango; [37.3256 ° N, 108.1328 ° W]; elev. 2415 m; 27 Aug. 1967; P. H. Timberlake leg.; ex yellow Composite; UCRC • 4 ♀♀; La Plata Co., Durango; [37.275 ° N, 107.881 ° W]; 21 May 1953; R. S. Beal leg.; EMEC • 1 ♀; Larimer Co., 10 mi. SW of Loveland; [40.339 ° N, 105.249 ° W]; 26 May 1980; U. N. Lanham leg.; ex Physaria belli; UCMC • 1 ♀; same data as for preceding; 26 May 1980; F. Robinson leg.; ex Physaria belli; UCMC • 1 ♀; same data as for preceding; 26 May 1980; F. Robinson leg.; UCMC • 1 ♀; Larimer Co., 10 mi. WSW of Loveland; [40.339 ° N, 105.249 ° W]; elev. 1981 m; 15 May 1963; U. N. Lanham leg.; ex Ribes; UCMC • 3 ♀♀; same data as for preceding; 15 May 1963; U. N. Lanham leg.; ex Salix; UCMC • 2 ♀♀; Larimer Co., 13 mi. WSW of Loveland; [40.321 ° N, 105.302 ° W]; elev. 1981 m; 30 Apr. 1963; U. N. Lanham leg.; ex Salix; UCMC • 12 ♀♀; Larimer Co., 14 mi. W of Fort Collins; [40.585 ° N, 105.351 ° W]; elev. 2012 m; 6 May 1963; U. N. Lanham leg.; ex Salix; UCMC • 1 ♀; Larimer Co., 16 mi. N of Fort Collins; [40.817 ° N, 105.084 ° W]; 10 May 1963; U. Lanham leg.; ex Salix; UCMC • 3 ♀♀; Larimer Co., 16 mi. WNW of Fort Collins; [40.671 ° N, 105.367 ° W]; elev. 2134 m; 9 May 1963; U. N. Lanham leg.; UCMC • 1 ♀; same data as for preceding; 9 May 1963; U. N. Lanham leg.; ex Salix; UCMC • 1 ♀; Larimer Co., 7 mi. W of Berthoud; [40.304 ° N, 105.214 ° W]; elev. 1676 m; 15 May 1963; U. Lanham leg.; ex Salix; UCMC • 2 ♀♀; same data as for preceding; 15 May 1963; U. Lanham leg.; ex Rhus trilobata; UCMC • 4 ♀♀; Larimer Co., 8 mi. WSW of Loveland; [40.352 ° N, 105.209 ° W]; elev. 1676 m; 30 Apr. 1963; U. N. Lanham leg.; ex Prunus; UCMC • 1 ♂; Larimer Co., Glacier View Meadows; [40.737 ° N, 105.422 ° W]; 2 Sep. 1984; H. E. and M. A. Evans leg.; UCMC • 1 ♀; Larimer Co., Hewlett Gulch; [40.655 ° N, 105.345 ° W]; 29 Jun. 1979; H. E. Evans leg.; UCMC • 1 ♀; same data as for preceding; 28 Jun. 1984; H. E. Evans leg.; UCMC • 5 ♀♀; Larimer Co., RMNP Moraine Park; [40.353 ° N, 105.598 ° W]; elev. 2286 m; 12 Jun. 1963; S. M. Sutton leg.; UCMC • 1 ♀; Larimer Co., Virginia Dale; [40.951 ° N, 105.346 ° W]; 10 May 1963; U. N. Lanham leg.; ex Salix; UCMC • 1 ♀; Larimer Co.; 40.4302 ° N, 105.5076 ° W; 5 Aug. 2010; C. Whitten leg.; BIML • 2 ♀♀; Larimer Co.; 40.4012 ° N, 105.6278 ° W; 5 Aug. 2010; C. Flynn leg.; BIML (Fig. 40 B) • 1 ♀; Mesa Co., 17 mi. SW of Whitewater; [38.754 ° N, 108.693 ° W]; 18 May 1963; C. J. McCoy and B. Vogel leg.; ex Taraxacum; UCMC • 1 ♀; Mesa Co., Colorado National Monument; [39.051 ° N, 108.692 ° W]; elev. 1463 m; 2 May 1963; B. Vogel and C. J. McCoy leg.; UCMC • 1 ♀; same data as for preceding; 16 May 1985; U. Lanham and M. Weissmann leg.; ex Lepidium montanum; UCMC • 1 ♀; Mesa Co., Mud Springs, Pinyon Mesa; [38.904 ° N, 108.739 ° W]; 22 Jun. 1963; C. J. McCoy and B. Vogel leg.; UCMC • 1 ♀; Mesa Co., Vega Res; [39.222 ° N, 107.786 ° W]; 16 Jun. 1977; E. Byron leg.; ex Prunus virginiana; UCMC • 1 ♀; Moffat Co., 12 mi. N of Dinosaur; [40.414 ° N, 109.008 ° W]; elev. 2438 m; 24 – 25 May 1974; U. N. Lanham leg.; ex Cercocarpus ledifolius; UCMC • 1 ♂; Moffat Co., 15 mi. NE of Craig; [40.669 ° N, 107.345 ° W]; 3 Aug. 1979; U. N. Lanham leg.; ex Solidago; UCMC • 1 ♀; Moffat Co., 23 mi. SW of Maybell; [40.285 ° N, 108.404 ° W]; elev. 1829 m; 26 May 1974; U. N. Lanham leg.; UCMC • 1 ♀; Montezuma Co., 10 mi. NW of Dolores; [37.577 ° N, 108.626 ° W]; 28 May 1977; U. Lanham and P. Byron leg.; ex Amelanchier; UCMC • 1 ♀; Montezuma Co., 10 mi. NW of Dolores; [37.577 ° N, 108.626 ° W]; 28 May 1977; U. Lanham and P. Byron leg.; ex Amelanchier; UCMC • 1 ♀; Montezuma Co., 11 mi. NW of Dolores; [37.587 ° N, 108.639 ° W]; 28 May 1977; U. Lanham and P. Byron leg.; ex Prunus virginiana; UCMC • 3 ♀♀; Montrose Co., 1 mi. S of Buckeye Reservoir; [38.360 ° N, 108.931 ° W]; 19 May 1963; B. Vogel and C. J. McCoy leg.; ex Amelanchier; UCMC • 2 ♀♀; Montrose Co., 17 mi. SSW of Naturita; [38.101 ° N, 108.673 ° W]; elev. 1981 m; 29 May 1977; U. Lanham and P. Byron leg.; ex Cardaria; UCMC • 1 ♀; Montrose Co., Buckeye Reservoir; [38.438 ° N, 109.041 ° W]; 19 May 1966; B. Vogel and C. J. McCoy leg.; ex Taraxacum; UCMC • 1 ♀; Montrose Co., Gunnison River Portal; [38.524 ° N, 107.65 ° W]; 26 Jun. 1963; C. J. and P. K. McCoy leg.; UCMC • 1 ♀; Pueblo Co., Colorado City; [37.945 ° N, 104.836 ° W]; elev. 1829 m; 20 May 1975; U. Lanham leg.; ex Rhus trilobata; UCMC • 1 ♀; San Miguel Co., 9 mi. NE of Placerville; [38.108 ° N, 107.936 ° W]; 25 Jun. 1963; B. Vogel, C. J. and P. K. McCoy leg.; ex Prunus virginiana; UCMC • 1 ♂; Teller Co., 7 mi. W of Woodland Park, South Meadows Camp; [39.064 ° N, 105.093 ° W]; elev. 2438 m; 21 – 28 Jul. 1977; S. and J. Peck leg.; ex Salix; EMEC • 1 ♀; Teller Co., Florissant; [38.946 ° N, 105.29 ° W]; 11 Jun. 1907; A. Hoburn leg.; ex Ribes?; UCRC. – Idaho • 1 ♂; Bannock Co., Pocatello; [42.86 ° N, 112.45 ° W]; 18 Jun. 1958; UCRC • 1 ♀; Benewah Co., St Maries; [47.314 ° N, 116.563 ° W]; 24 Jun. 1919; H. A. Scullen leg.; OSAC • 1 ♀; Bingham Co., Blackfoot; [43.191 ° N, 112.345 ° W]; 21 Jun. 1932; Louise Ireland leg.; CAS • 1 ♀; Latah Co., Moscow Mountains; [46.803 ° N, 116.869 ° W]; elev. 975 m; 3 Jun. 1970; Paul Oman leg.; OSAC • 1 ♀; same data as for preceding; 27 Jul. 1925; C. L. Fox leg.; OSAC • 1 ♂; same data as for preceding; 27 Jul. 1925; C. L. Fox leg.; CAS • 1 ♀; same data as for preceding; 27 Jul. 1925; C. L. Fox leg.; CAS • 2 ♂♂; same data as for preceding; 27 Jul. 1925; C. L. Fox leg.; CAS • 1 ♀; Valley Co., near Smiths Ferry, Hwy 55; 44.29777 ° N, 116.08869 ° W; 28 May 2007; J. Gibbs and C. Sheffield leg.; PCYU. – Montana • 1 ♀; Carbon Co., 3 mi. SSE of Roscoe; [45.321 ° N, 109.26 ° W]; elev. 1646 m; 21 Jun. 1981; U. N. Lanham leg.; UCMC • 2 ♀♀; Carbon Co., E. of Rosebud Lake; [45.197 ° N, 109.636 ° W]; 21 Jun. 1962; B. Vogel leg.; ex Lupinus argenteus; UCMC • 1 ♀; Fergus Co., Big Snowy Mountains; [46.640 ° N, 109.18 ° W]; elev. 2560 m; 5 Aug. 1961; B. Vogel leg.; UCMC • 2 ♀♀; Gallatin Co., Hwy 286; 44.80887 ° N, 111.24994 ° W; elev. 2014 m; 23 Jun. 2007; J. Gibbs, C. Sheffield leg.; PCYU • 1 ♀; Mineral Co., 3 mi. SE of DeBorgia; [47.366 ° N, 115.303 ° W]; 8 Jun. 1963; D. R. Smith leg.; OSAC • 1 ♀; Missoula Co., Florence MPG Ranch; 46.7093 ° N, 114.0038 ° W; elev. 1359 m; 20 – 22 May 2019; M. Seidensticker leg.; CBG • 1 ♀; Missoula Co., Florence MPG Ranch; 46.6901 ° N, 113.9847 ° W; elev. 1380 m; 16 – 18 Jul. 2019; M. Seidensticker leg.; CBG. – Nevada • 1 ♀; Clark Co., Kyle Canyon Campground, Spring Mountains; [36.270 ° N, 115.653 ° W]; elev. 2133 m; 15 May 1971; P. H. and M. M. Arnaud leg.; CAS • 2 ♀♀; Elko Co., Secret Pass, Ruby Mountains; [40.815 ° N, 115.198 ° W]; elev. 1981 m; 19 Jun. 1955; C. D. MacNeill leg.; EMEC • 1 ♀; Eureka Co., 3 mi. W of Eureka; [39.512 ° N, 116.016 ° W]; 24 Jun. 1968; J. T. Doyen and B. Manion leg.; EMEC • 1 ♀; Washoe Co., Reno; [39.530 ° N, 119.814 ° W]; 27 Jun. 1927; E. P. Van Duzee leg.; CAS • 1 ♀; White Pine Co., Lehman Creek, 6 mi. E of Baker; [39.012 ° N, 114.012 ° W]; 24 Jun. 1968; J. T. Doyen and B. Manion leg.; EMEC • 1 ♂; White Pine Co., 22 mi. W of Ely; [39.246 ° N, 115.299 ° W]; 20 Jul. 1958; E. G. Linsley and J. W. MacSwain leg.; EMEC (Fig. 41 A) • 1 ♀; Kings Canyon Ranch, Carson City; [39.161 ° N, 119.799 ° W]; elev. 1524 m; 23 Jun. 1964; D. Bolinger leg.; OSAC. – New Mexico • 2 ♂♂; Bernalillo Co., Juan Tabo, Sandia Mountains; [35.205 ° N, 106.499 ° W]; elev. 2134 m; 8 Jul. 1945; W. O. Griesel leg.; EMEC • 1 ♀; Bernalillo or Sandoval Co., Sandia Mountain; [35.208 ° N, 106.407 ° W]; (?); UCRC • 1 ♀; Santa Fe Co., 20 mi. W of Santa Fe; [35.686 ° N, 106.293 ° W]; 28 Apr. 1981; U. N. Lanham leg.; UCMC • 1 ♀; Socorro Co., 10 mi. W of Socorro; [34.044 ° N, 107.068 ° W]; 28 Mar. 1963; U. N. Lanham leg.; UCMC. – Oregon • 7 ♂♂; Baker Co., Cornucopia; [45.008 ° N, 117.192 ° W]; elev. 2210 m; 24 Jul. 1936; E. A. Scullen leg.; OSAC • 1 ♀; same data as for preceding; 24 Jul. 1936; R. E. Rieder leg.; OSAC • 1 ♀; same data as for preceding; 24 Jul. 1936; H. A. Scullen leg.; OSAC • 3 ♀♀; Benton Co., Corvallis; [44.564 ° N, 123.262 ° W]; 8 May 1936; G. Ferguson leg.; OSAC • 1 ♀; same data as for preceding; 28 Apr. 1936; G. Ferguson leg.; OSAC • 1 ♀; same data as for preceding; 11 May 1925; OSAC • 1 ♀; same data as for preceding; 6 Jul. 1931; OSAC • 1 ♀; same data as for preceding; 9 Jun. 1932; OSAC • 1 ♀; same data as for preceding; 10 Apr. 1929; H. A. Scullen leg.; OSAC • 1 ♀; same data as for preceding; 24 Mar. 1926; H. A. Scullen leg.; OSAC • 1 ♀; same data as for preceding; 14 Mar. 1926; H. A. Scullen leg.; OSAC • 1 ♀; same data as for preceding; 1 Jul. 1931; Joe Schuh leg.; OSAC • 1 ♀; Benton Co., Marys Peak Summit; [44.504 ° N, 123.553 ° W]; 4 Jul. 1941; G. R. Ferguson leg.; OSAC • 1 ♀; Benton Co., Monroe; [44.314 ° N, 123.297 ° W]; just May 24; J. E. Davis leg.; OSAC • 2 ♀♀; Columbia Co., Scappoose; [45.757 ° N, 122.878 ° W]; 23 Apr. 1938; K. Gray and J. Schuh leg.; OSAC • 2 ♀♀; same data as for preceding; 1 May 1937; OSAC • 2 ♀♀; same data as for preceding; 5 – 6 May 1937; OSAC • 1 ♀; same data as for preceding; 6 May 1937; OSAC • 2 ♀♀; Columbia Co., St Helens – Scappoose; [45.808 ° N, 122.847 ° W]; 6 May 1938; OSAC • 1 ♀; Columbia Co., St Helens to Scappoose; [45.808 ° N, 122.847 ° W]; 28 Apr. 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♀; Crook Co., 5 mi. E of Ochoco Ranger Station; [44.396 ° N, 120.37 ° W]; 3 May 1939; K. Gray and J. Schuh leg.; OSAC • 1 ♂; Crook Co., Milton; [44.349 ° N, 120.906 ° W]; 23 Jun. 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♀; Deschutes Co., Camp Abbot; [43.868 ° N, 121.442 ° W]; 21 May 1944; Paul H. Arnaud leg.; CAS • 1 ♀; Douglas Co., Yoncalla; [43.596 ° N, 123.283 ° W]; 26 May 1937; R. M. and G. E. Bohart leg.; CAS • 1 ♀; Grant Co., Strawberry Lake to Straw Lake; [44.380 ° N, 118.678 ° W]; elev. 2073 – 2134 m; 16 Jul. 1936; H. A. Scullen leg.; OSAC • 1 ♀; Harney Co., 17 mi. N of French-Glen; [43.052 ° N, 118.918 ° W]; 19 May 1956; J. D. Lattin leg.; OSAC • 1 ♀; Harney Co., 3 mi. S of Fish Lake, Steens Mountain; [42.614 ° N, 118.596 ° W]; 3 Jul. 1947; P. Ellerton leg.; PMAE • 1 ♀; Harney Co., Buena Vista Ranch, Blitzen Valley; [43.272 ° N, 118.853 ° W]; 19 Apr. 1936; S. G. Jewett leg.; OSAC • 1 ♀; Harney Co., Fish Lake, Steens Mountains; [42.729 ° N, 118.653 ° W]; elev. 2195 m; 8 Jul. 1947; J. Ellertson leg.; OSAC (Fig. 40 C) • 1 ♀; same data as for preceding; 4 Jul. 1947; F. Ellerton leg.; OSAC • 6 ♀♀; same data as for preceding; 11 Jul. 1927; H. A. Scullen leg.; OSAC • 1 ♀; same data as for preceding; 14 Jul. 1953; J. E. Davis leg.; OSAC • 1 ♀; Harney Co., Malheur National Wildlife Refuge, SBV; 42.8188 ° N, 118.8688 ° W; 17 Apr. 2012; J. D. Engler leg.; OSAC • 1 ♀; Hood River Co., Homestead Inn, Mt Hood; [45.413 ° N, 121.592 ° W]; 30 Jun. 1927; E. C. VanDyke leg.; CAS • 1 ♀; Hood River Co., Hood River; [45.705 ° N, 121.522 ° W]; 12 Jun. 1924; E. C. VanDyke leg.; CAS • 2 ♂♂; Hood River Co., Mt Hood Ghostridge; [45.411 ° N, 121.652 ° W]; elev. 1829 m; 30 Jul. 1921; A. L. Melander leg.; UCRC • 1 ♀; Jackson Co., 1.5 mi. NE of Mt Ashland Ski Bowl; [42.097 ° N, 122.684 ° W]; 4 Jul. 1970; P. Rude leg.; EMEC • 1 ♀; Jackson Co., Buckhorn Mineral Springs, 11 mi. ESE of Ashland. Emmigrant Creek; [42.133 ° N, 122.511 ° W]; elev. 853 m; 19 May 1960; J. D. Lattin leg.; OSAC • 1 ♀; Jackson Co., Little Squaw Lake, 7 mi. E of Copper; [42.098 ° N, 123.109 ° W]; elev. 975 m; 22 May 1964; J. D. Lattin leg.; OSAC • 2 ♀♀; Jefferson Co., Suttle Lake; [44.423 ° N, 121.747 ° W]; 2 May 1939; Schuh and Gray leg.; OSAC • 1 ♀; Jefferson Co., Suttle Lake; [44.426 ° N, 121.728 ° W]; 2 May 1939; K. Gray and J. Schuh leg.; OSAC • 2 ♀♀; Klamath Co., Eagle Ridge, Klamath Lake; [42.249 ° N, 121.968 ° W]; 19 May 1924; C. L. Fox leg.; CAS • 1 ♀; same data as for preceding; 23 May 1924; C. L. Fox leg.; CAS • 1 ♀; same data as for preceding; 15 May 1924; C. L. Fox leg.; CAS • 1 ♀; Klamath Co., Klamath Falls; [42.225 ° N, 121.782 ° W]; 9 May 1924; C. L. Fox leg.; CAS • 2 ♀♀; same data as for preceding; 7 May 1924; C. L. Fox leg.; CAS • 4 ♀♀; Klamath Co.; [42.68 ° N, 121.65 ° W]; 18 May 1913; F. W. Nunenmacher leg.; EMEC • 1 ♀; Lake Co., Hart Mountain Antelope Refuge; [42.549 ° N, 119.642 ° W]; 24 Apr. 1954; Otto C. Nelson leg.; ex Currant; OSAC • 1 ♀; Lake Co., Lakeview; [42.188 ° N, 120.346 ° W]; 6 May 1954; Otto C. Nelson leg.; ex Prunus; OSAC • 1 ♀; Lake Co., Picture Rock Pass; [43.048 ° N, 120.799 ° W]; 19 Apr. 1950; Black and Davis leg.; OSAC • 1 ♀; Lane Co., Alder Springs, McKenzie Pass; [44.179 ° N, 121.913 ° W]; elev. 922 m; 30 Jun. 1927; H. A. Scullen leg.; OSAC • 3 ♀♀; Lane Co., White Brunch Meadow, Three Sisters, Frog Camp; [44.215 ° N, 121.858 ° W]; elev. 1676 m; 19 Jul. 1927; H. A. Scullen leg.; OSAC • 2 ♀♀; Lincoln Co., Alsea Mountain, 15 mi. W of Corvallis; [44.564 ° N, 123.566 ° W]; 16 Mar. 1936; K. Gray and J. Schuh leg.; OSAC • 4 ♀♀; same data as for preceding; 17 Mar. 1936; K. Gray and J. Schuh leg.; ex Salix; OSAC • 1 ♂; Multnomah Co., Laurelhurst St Prk; [45.521 ° N, 122.627 ° W]; 20 Jun. 1962; C. A. Toschi leg.; EMEC • 1 ♂; Umatilla Co., 8 mi. W of Meacham; [45.507 ° N, 118.586 ° W]; elev. 1646 m; 19 Jul. 1929; H. A. Scullen leg.; ex Holodiscus discolor; OSAC • 2 ♀♀; Umatilla Co., Cabbage Hill, E. of Pendleton; [45.555 ° N, 118.611 ° W]; 8 Jun. 1955; Paul O. Ritcher leg.; OSAC • 1 ♀; Umatilla Co., Meacham; [45.506 ° N, 118.421 ° W]; elev. 1122 m; 8 May 1927; H. A. Scullen leg.; OSAC • 1 ♀; Umatilla Co., Milton; [45.934 ° N, 118.391 ° W]; 22 Jun. 1938; K. Gray and J. Schuh leg.; OSAC • 2 ♂♂; same data as for preceding; 22 Jun. 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♀; Umatilla Co., Weston; [45.813 ° N, 118.425 ° W]; 28 Apr. 1939; K. Gray and J. Schuh leg.; OSAC • 1 ♀; same data as for preceding; 29 Apr. 1939; K. Gray leg.; ex Brassica; OSAC • 6 ♀♀; Umatilla Co., Wild Horses Mountain, Athena; [45.802 ° N, 118.481 ° W]; 14 May 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♀; Union Co., 20 mi. SW of LaGrande; [45.119 ° N, 118.377 ° W]; elev. 1219 – 1524 m; 11 May 1930; OSAC • 2 ♂♂; Union Co., La Grande; [45.323 ° N, 118.088 ° W]; 20 Jul. 1929; H. A. Scullen leg.; ex Convolvulus; OSAC • 1 ♀; Wasco Co., The Dalles; [45.601 ° N, 121.185 ° W]; 2 May 1927; H. A. Scullen leg.; OSAC • 1 ♀; same data as for preceding; 24 May 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♂; Wheeler Co., Twickenham; [44.747 ° N, 120.176 ° W]; 24 Jun. 1948; J. E. Davis leg.; OSAC. – Texas • 1 ♂; Patricio Co., Welder Preserve, 7 mi. NE of Sinton; [28.105 ° N, 97.438 ° W]; 25 Apr. 1977; M. Wasbauer and T. Eichlis leg.; EMEC. – Utah • 1 ♂; Cache Co., Logan Canyon; [41.78 ° N, 111.644 ° W]; 17 Jul. 1958; E. G. Linsley leg.; ex Asclepias; EMEC • 1 ♀; Cache Co., Millville; [41.685 ° N, 111.822 ° W]; 23 May 1964; G. F. Knowlton leg.; SEMC • 1 ♀; Cache Co., Providence; [41.706 ° N, 111.818 ° W]; 24 May 1964; G. F. Knowlton leg.; ex Daucus; SEMC • 1 ♀; same data as for preceding; 4 May 1965; G. F. Knowlton leg.; ex Umbelliferae; SEMC • 1 ♀; Cache Co., Smithfield; [41.836 ° N, 111.838 ° W]; 12 May 1963; G. F. Knowlton leg.; SEMC • 1 ♀; Iron Co., Parowan Canyon; [37.835 ° N, 112.819 ° W]; 4 Mar. 1963; ex Salix; UCRC • 1 ♀; Salt Lake Co., Salt Lake; [40.761 ° N, 111.891 ° W]; 4 May 1913; P. H. Timberlake leg.; UCRC • 4 ♀♀; same data as for preceding; 4 May 1913; P. H. Timberlake leg.; ex Salix; UCRC • 1 ♀; same data as for preceding; 17 May 1913; P. H. Timberlake leg.; UCRC • 1 ♀; Salt Lake Co., Salt Lake City; [40.761 ° N, 111.891 ° W]; 31 May 1928; W. W. Jones leg.; UCRC • 1 ♀; same data as for preceding; 12 Jun. 1941; CAS • 1 ♀; Uintah Co., 6 mi. NW of Vernal; [40.517 ° N, 109.609 ° W]; 1 Jun. 1977; U. Lanham and P. Byron leg.; ex Amelanchier; UCMC • 6 ♀♀; Uintah Co., 8 mi. NW of Vernal; [40.537 ° N, 109.636 ° W]; 1 Jun. 1977; U. Lanham and P. Byron leg.; ex Taraxacum; UCMC • 1 ♂; Uintah Co.; [40.455 ° N, 109.528 ° W]; CMNH • 1 ♂; Utah Co., 4 mi. N of Teat Mountain; [40.081 ° N, 111.366 ° W]; E. G. Linsley leg.; EMEC • 1 ♂; Utah Co., Aspen Grove; [40.404 ° N, 111.605 ° W]; 1 Aug. 1938; H. P. Chandler leg.; CAS • 1 ♀; Utah Co., Provo; [40.234 ° N, 111.659 ° W]; H. Chandler leg.; CAS • 2 ♀♀; Utah Co., Provo Canyon; [40.355 ° N, 111.575 ° W]; 22 May 1960; S. C. Bromley leg.; EMEC. – Washington • 1 ♂; Asotin Co., Fields Spring State Park, S of Antone; [46.087 ° N, 117.172 ° W]; 28 Jul. 1985; Finnamore and Thormin leg.; PMAE • 1 ♂; Chelan or King Co., White Rock Spring, Steven Pass, Cascade Mountains; [47.744 ° N, 121.089 ° W]; 30 Jul. 1930; E. C. VanDyke leg.; CAS • 1 ♀; Klickitat Co., Conboy Lake National Wildlife Refuge, LRL; 45.9283 ° N, 121.3783 ° W; 25 – 26 May 2012; Diedre Shirley leg.; OSAC • 1 ♀; same data as for preceding; 5 – 6 May 2011; L. Wilson leg.; OSAC • 1 ♀; Stevens Co., Little Pend Oreille National Wildlife Refuge, LPO; 48.4654 ° N, 117.7358 ° W; 20 – 21 Jul. 2012; J. Koelsch leg.; OSAC. – Wyoming • 3 ♀♀; Albany Co., 3 mi. S of Woods Landing; [41.085 ° N, 105.997 ° W]; 15 May 1978; U. Lanham leg.; ex Salix; UCMC • 5 ♀♀; same data as for preceding; 16 May 1978; U. Lanham leg.; ex Salix; UCMC • 1 ♀; Albany Co., National Forest Boundary W. of Woods Landing; [41.185 ° N, 106.072 ° W]; 13 Jun. 1978; U. Lanham leg.; ex Prunus virginiana; UCMC • 1 ♀; Carbon Co., 18 air mi. SE of Saratoga; [41.270 ° N, 106.562 ° W]; 30 Jun. 1982; W. J. Pulawski leg.; CAS • 1 ♂; Lincoln Co., Bridger Basin; [41.666 ° N, 110.934 ° W]; S. Garman leg.; EMEC • 1 ♀; Lincoln Co.; 41.8650 ° N, 110.7653 ° W; 4 Jun. 2013; BIML • 1 ♀; same data as for preceding; 17 – 18 Jul. 2012; P. Knecht and C. Bossi leg.; BIML • 1 ♀; Lincoln Co.; 41.8704 ° N, 108.7772 ° W; 13 May 2013; BIML • 1 ♀; same data as for preceding; 20 – 21 Sep. 2011; BIML • 1 ♂; Sweetwater Co., 25 mi. S of Bitter Creek; [41.185 ° N, 108.555 ° W]; EMEC • 1 ♀; Teton Co., Grand Teton National Park; [43.790 ° N, 110.681 ° W]; 23 Jun. 1938; E. C. VanDyke leg.; CAS • 1 ♀; Teton Co., Jenny Lake, Grand Teton; [43.754 ° N, 110.72 ° W]; 25 Jun. 1938; E. C. VanDyke leg.; CAS	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFB9FF99FDF8FE72FD7BFB52.taxon	description	Redescription Female BODY MEASUREMENTS. Length 7.58 mm (± 0.30 SD). Head length 1.86 mm (± 0.06 SD). Face length 1.54 mm (± 0.03 SD). Head width 1.76 mm (± 0.04 SD). Wing length 5.62 mm (± 0.31 SD). n = 5. COLOURATION. Head and mesosoma black. Clypeus apex black. Labrum black to dark brown. Mandible black to dark brown basally, becoming light brown medially and reddish brown apically. Flagellum dark brown to brown dorsally, light brown ventrally. Pronotal lobe dark brown. Tegula light brown, with brown base. Wing membrane faintly brown, veins light brown. Legs brown. Metasoma black to dark brown with rims of terga and sterna broadly light brown. PUBESCENCE. Body hair colour off-white. Tomentum dense on pronotal angle, and lobe, space between pronotal lobe and tegula; very sparse or sometimes absent on T 2 – T 3 basolaterally. Mesosoma covered by erect plumose hairs (1 – 2.5 OD) and sub-appressed simple short hairs (<0.5 OD). Metafemoral scopa with dense plumose hairs. Wing hairs brown, very short and sparse. SURFACE SCULPTURING. Clypeus polished, becoming weakly imbricate in the basal third; punctures irregularly sparse (IS ≤ 2 PD) apically, becoming smaller and denser (IS ≤ 1 PD) basally. Supraclypeal area polished, becoming imbricate posterolaterally; punctures sparse (IS = 1 – 3 PD) anteromedially, becoming dense (IS ≤ 1 PD) posterolaterally. Frons reticulate-punctate; punctures crowded (IS = 0 PD). Ocellocular area polished; punctures dense (IS <1 PD). Gena polished and weakly lineolate posteriorly; punctures sparse (IS = 1 – 3 PD), becoming obscure posteriorly. Postgena polished, but weakly lineolate adjacent to hypostomal carina and posteriorly. Tegula broadly impunctate, minute punctures present anteriorly. Mesoscutum mostly polished, transitioning to imbricate anteriorly and medially halfway on median line; punctures moderately dense to moderately sparse (IS = 1 – 2 PD), becoming broadly sparse (IS = 1 – 3 PD) submedially, diversopunctate. Mesoscutellum polished; punctures dense (IS <1 PD) marginally and on median line, sparse submedially (IS = 1 – 6 PD). Metanotum rugulose laterally, ruguloso-punctate medially with punctures crowded (IS = 0 PD). Propodeal triangle with rugae moderately strong, reaching ¾ towards posterior margin, anastomosing medially and weakly so laterally, lateral rugae not extending to dorsolateral slope of propodeum lateral face; interspace sculpture imbricate to tessellate. Preëpisternum weakly rugulose. Hypoepimeron imbricate to weakly rugulose; impunctate. Mesepisternum finely rugulose, imbricate ventrally; impunctate. Propodeum lateral face strongly imbricate, punctures obscure; posterior face polished to weakly imbricate, with punctures moderately dense anteriorly (IS = 1 – 2 PD), becoming irregularly sparser posteriorly (IS <3 PD). T 1 anterior face polished, punctures present laterally, sparse (IS = 1 – 6 PD); dorsal surface polished, punctures present laterally, sparse (IS = 1 – 3 PD), becoming sparser (IS = 1 – 6 PD) and fine towards premarginal line and nearly impunctate medially; rim weakly coriarious, punctures sparse (IS = 1 – 3 PD). T 2 disc coriarious to polished, with punctures densely spaced basally (IS ≤ 1 PD), becoming sparser apically (IS = 1 – 4 PD); rim coriarious, with minute and sparse punctures (IS = 1 – 3 PD), becoming impunctate narrowly after laterally premarginal line. STRUCTURE. Head long, length / width ratio 1.06 (± 0.03 SD). Face length / width ratio 0.88 (± 0.01 SD). Clypeus projecting ~ 80 % below suborbital tangent. Mandible short, extending just past the opposing clypeal angle. Eyes convergent below, UOD / LOD ratio 1.05 (± 0.03 SD). Ocelli normal. Gena / eye width ratio 0.72 (± 0.08 SD). Pronotal angle obtuse. Intertegular span 1.00 mm (± 0.05 SD). Mesoscutum length / width ratio 0.90 (± 0.03 SD); mesoscutum / scutellum length ratio 2.78 (± 0.14 SD); scutellum / metanotum length ratio 1.84 (± 0.11 SD); metanotum / propodeal triangle length ratio 0.66 (± 0.06 SD). Protibial spur with apical serrations shorter than width of malus; malus extending beyond the velum, nearly ¾ the length of velum. Inner metatibial spur pectinate, teeth 3 – 5, basal teeth longer than width of rachis. Propodeal triangle posterior margin rounded onto posterior propodeal surface. Propodeal lateral carina not reaching dorsal margin, oblique carina absent. VARIATION. The tegula colour varies from light brown to brown. Occasionally, the rims of the metasomal terga become darker, lacking the typical pronounced broad light-brown rims. The ocellocular area is sometimes weakly rugulose, with crowded punctures (IS = 0 PD). The mesoscutum exhibits variation in puncture density and the extent of the polished area. In some individuals, the mesoscutum is polished only submedially, with dense punctures throughout (IS ≤ 1 PD), except for moderately sparse or moderately dense punctures (IS = 1 – 2 PD) mesad of the parapsidal line. Occasionally, the punctures can even become more broadly sparse submedially (IS = 3 – 6 PD). In some specimens, the hypoepimeron appears to have a few shallow, obscure punctures. Sometimes, the punctures on the posterior face of the propodeum are very obscure and more crowded. In some instances, these punctures may extend onto the dorsolateral slope, but they can only be observed from the posterior view of the bee. In some specimens, the mandible reaches the opposing clypeal angle but does not extend beyond it. Male BOSY MEASUREMENTS. Length 6.78 mm (± 0.38 SD). Head length 1.71 mm (± 0.10 SD). Face length 1.43 mm (± 0.08 SD). Head width 1.59 mm (± 0.08 SD). Wing length 4.83 mm (± 0.39 SD). n = 5. COLOURATION. Head and mesosoma black. Clypeus apical half yellow. Labrum brown. Mandible dark brown basally, becoming yellow medially and reddish brown apically. Flagellum dark brown to brown dorsally, light brown ventrally. Pronotal lobe brown. Tegula light brown, with brown base. Wing membrane faintly brown, veins brown. Legs dark brown to brown, except slightly light brown tarsi. Metasoma dark brown with rims of terga and sterna narrowly lighter. PUBESCENCE. Body hair colour off-white. Tomentum dense on paraocular area, clypeus basally, supraclypeal area laterally, pronotal angle, and lobe, space between pronotal lobe and tegula. Mesosoma covered by erect plumose hairs (1 – 2.5 OD) and sub-appressed simple short hairs (<0.5 OD). Metasomal sterna with dense, erect and sub-appressed, plumose, short (<1.5 OD) hairs. Wing hairs brown, very short and sparse. SURFACE SCULPTURING. Clypeus polished; punctures irregularly sparse (IS ≤ 2 PD) apically, becoming smaller and denser (IS ≤ 1 PD) basally. Supraclypeal area polished; punctures dense (IS ≤ 1 PD). Frons reticulate-punctate; punctures crowded (IS = 0 PD). Ocellocular area weakly rugulose, with crowded punctures (IS = 0 PD). Gena polished and weakly lineolate posteriorly; punctures moderately sparse (IS = 1 – 2 PD), becoming obscure posteriorly. Postgena lineolate. Tegula broadly impunctate. Mesoscutum mostly polished, slightly imbricate anteriorly; punctures dense (IS ≤ 1 PD), becoming sparse (IS = 1 – 4 PD) mesad of parapsidal line, diversopunctate. Mesoscutellum polished; punctures dense (IS ≤ 1 PD) marginally and on median line, sparse submedially (IS = 1 – 4 PD). Metanotum rugulose. Propodeal triangle with rugae moderately strong, reaching ¾ towards posterior margin, anastomosing medially and weakly so laterally, lateral rugae not extending to dorsolateral slope of propodeum lateral face; interspace sculpture imbricate to tessellate and becoming polished posteriorly on posterior margin. Preëpisternum weakly rugulose. Hypoepimeron imbricate to weakly rugulose; punctures crowded (IS = 0 PD). Mesepisternum reticulate-punctate with crowded punctures, becoming imbricate ventrally where the punctures become obscure or impunctate. Propodeum lateral face strongly imbricate, and becoming rugulose dorsolateral slope, punctures obscure; posterior face weakly rugulose-imbricate, with dense punctures anteriorly (IS <1 PD). T 1 anterior face polished, few scattered punctures present laterally; dorsal surface polished, punctures present laterally, sparse (IS = 1 – 3 PD), becoming sparser (IS = 1 – 5 PD) medially and on subapicolateral boss; rim polished, punctures sparse (IS = 1 – 3 PD) and becoming impunctate laterally and apically. T 2 disc weakly coriarious basally, becoming polished apically, with moderately sparse punctures (IS = 1 – 2 PD); rim polished, with minute, sparse punctures (IS = 1 – 4 PD), becoming impunctate apically. STRUCTURE. Head long, length / width ratio 1.08 (± 0.03 SD). Face length / width ratio 0.90 (± 0.02 SD). Clypeus projecting ~ 64 % below suborbital tangent. Mandible short, reaching opposing clypeal angle. Flagellomeres long, F 1 length / pedicel length ratio 1.20 (± 0.15 SD); F 2 length / F 1 length ratio 2.65 (± 0.14 SD); F 2 length / F 2 width ratio 1.61 (± 0.05 SD). Gena / eye width ratio 0.58 (± 0.03 SD). Forewing with 3 submarginal cells. Pronotal angle obtuse. Intertegular span 0.78 mm (± 0.11 SD). Mesoscutum length / width ratio 1.02 (± 0.13 SD). Propodeal lateral carina not reaching dorsal margin, oblique carina absent. n = 5. VARIATION. Typically, there is a slight brown-to-light-brown gradient between the tibia and basitarsus. However, this gradient can rarely become more pronounced, as seen in the legs of male L. ruficorne. In some individuals, the mesoscutum punctures become more broadly sparse submedially (IS = 2 – 5 PD) submedially. The punctures on the hypoepimeron are obscure in some specimens. Sometimes, the mesepisternum exhibits reticulate-rugulose or rugulose sculpturing dorsally, with punctures that are difficult to observe. GENITALIA (Figs 43, 120 G). Gonostylus elongate and truncated, about 4 times as long as broad, with short setae. Retrorse lobe broad, about 3 times as long as broad, covered in short setae.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFB9FF99FDF8FE72FD7BFB52.taxon	distribution	Range Canada (Alberta, British Columbia), Mexico (Baja California), USA (Arizona, California, Colorado, Idaho, Montana, Nevada, New Mexico, Oregon, Texas, Utah, Washington, Wyoming) (Fig. 44).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFB9FF99FDF8FE72FD7BFB52.taxon	biology_ecology	Floral hosts ANACARDIACEAE: Rhus trilobata Nutt. • APIACEAE: Angelica capitellata (A. Gray) Spalik, Reduron & S. R. Downie, Daucus sp., Heracleum maximum W. Bartram • ASCLEPIADACEAE: Asclepias sp. • ASTERACEAE: Crepis runcinata (E. James) Torr. & A. Gray, Ericameria nauseosa var. bernardina (H. M. Hall) G. L. Nesom & G. I. Baird, Eriophyllum lanatum (Pursh) J. Forbes, Lasthenia sp., Senecio lugens Richardson, Senecio sp., Solidago velutina subsp. californica (Nutt.) Semple, Solidago sp., Taraxacum officinale F. H. Wigg., Taraxacum sp., Wyethia sp. • BORAGINACEAE: Cryptantha intermedia (A. Gray) Greene, Cryptantha sp., Eremocarya lepida (A. Gray) Greene, Lithospermum ruderale Douglas ex Lehm. • BRASSICACEAE: Barbarea sp., Brassica sp., Cardaria sp., Erysimum sp., Isatis tinctoria L., Lepidium montanum Nutt., Physaria bellii G. A. Mulligan, Physaria vitulifera Rydb., Sisymbrium sp. • CONVOLVULACEAE: Convolvulus sp. • ERICACEAE: Arctostaphylos glauca Lindl., Arctostaphylos patula Greene, Arctostaphylos pringlei subsp. drupacea (Parry) P. V. Wells, Arctostaphylos sp. • EUPHORBIACEAE: Euphorbia esula L. • FABACEAE: Lupinus argenteus Pursh, Trifolium sp. • GROSSULARIACEAE: Ribes cereum Douglas, Ribes indecorum Eastw., Ribes roezlii Regel, Ribes sp., Ribes viscosissimum Pursh • HYDROPHYLLACEAE: Eriodictyon californicum (Hook. & Arn.) Torr., Eriodictyon sp., Eriodictyon trichocalyx A. Heller, Phacelia hastata var. compacta (Brand) Cronquist (verbatim: Phacelia frigida), Phacelia sp. • LILIACEAE: Camassia quamash (Pursh) Greene • ONAGRACEAE: Oenothera elata subsp. hookeri (Torr. & A. Gray) W. Dietr. & W. L. Wagner • POLYGONACEAE: Eriogonum sp., Eriogonum umbellatum Torr., Eriogonum wrightii var. subscaposum S. Watson • PORTULACACEAE: Claytonia perfoliata Donn ex Willd. • RANUNCULACEAE: Clematis hirsutissima Pursh, Pulsatilla nuttalliana (DC.) Bercht. & J. Presl • RHAMNACEAE: Ceanothus cordulatus Kellogg, Ceanothus incanus Torr. & A. Gray, Ceanothus sp., Frangula californica (Eschsch.) A. Gray, Ranunculus sp. • ROSACEAE: Amelanchier sp., Cercocarpus ledifolius Nutt., Crataegus sp., Holodiscus discolor (Pursh) Maxim., Malus sp., Drymocallis glandulosa (Lindl.) Rydb., Potentilla sp., Prunus sp., Prunus subcordata Benth., Prunus virginiana var. demissa (Nutt.) Torr., Prunus virginiana L., Rubus deliciosus Torr., Rubus sp. • SALICACEAE: Salix sp., Salix vestita Pursh • SCROPHULARIACEAE: Penstemon grinnellii Eastw., Penstemon procerus Douglas ex Graham, Penstemon rydbergii A. Nelson	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFB9FF99FDF8FE72FD7BFB52.taxon	discussion	DNA barcodes DNA barcodes of L. diatretum form a single cluster based on sequences from Alberta, British Columbia, Saskatchewan, Colorado, Idaho, Montana.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFA4FFAFFDF6FB32FB0BFDF7.taxon	description	Figs 40 D – F, 41 D – F, 46 – 51, 108 A, 109 E, 118 B, E, H, K, 120 H	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFA4FFAFFDF6FB32FB0BFDF7.taxon	etymology	Etymology Crawford (1907 c) did not explain the epithet. The specific epithet is treated as a compound Latin adjective, likely derived from ‘ rufus’ and ‘ cornis ’, meaning ‘ red horned’.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFA4FFAFFDF6FB32FB0BFDF7.taxon	diagnosis	Diagnosis Refer to L. diatretum diagnosis.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFA4FFAFFDF6FB32FB0BFDF7.taxon	materials_examined	Type material Lectotype of Halictus ruficornis (designated herein; examined) UNITED STATES – Nevada • ♂; Carson City; [39.16 ° N, 119.78 ° W]; Jul.; Baker leg.; USNM. Specimen labels “ OrmsbyCo. Nev. / July. Baker // 4688 // Type / No. 12049 / U. S. N. M. // Halictus / ♂ ruficornis / Cwfd. Type // LECTOTYPE // USNMENT / 01735683 ”. Paralectotype of Halictus ruficornis (designated herein; examined) UNITED STATES – Nevada • 1 ♂; Carson City; [39.16 ° N, 119.78 ° W]; Jul.; Baker leg.; USNM. Specimen labels “ Ormsby Co. Nev. / July. Baker // 5029 // Paratype / No. 12049 / U. S. N. M. // Halictus / ♂ ruficornis / Cwfd. Paratype ”. Syntype of Halictus vaporellus (designated herein; examined) UNITED STATES – Colorado • 1 ♀; Routt Coson Co., Steamboat Springs; [40.50 ° N, 106.83 ° W]; Jul.; Cockerell leg.; USNM (Type No. 23324) (Fig. 51). Specimen labels “ Steamboat Spgs. / Colo. May 27. (Ckll) // TypeNo. / 23324 / U. S. N. M. // Halictus / vaporellus / Ckll. TYPE. // USNMENT 01735685 ”. Syntype of Halictus pulveris (examined by photos) UNITED STATES – Colorado • 1 ♀; Montezuma Co., Mesa Verde; [37.183 ° N, 108.500 ° W]; 3 – 7 Jul. 1919; elev. 2011.7 m; Cockerell leg.; AMNH IZC 00323614 (Fig. 50). Specimen labels “ Mesa Verde, Colo. / July 3 – 7, ᾽ 19 / F 4371. // About 37 ° 11 ᾽ N. / 108 ° 30 ᾽ W. / About 6600 ft. // Halictus / pulveris / Ckll. TYPE // AMNH _ IZC 00323614 ”. Material examined CANADA – Alberta • 1 ♀; Cypress Co., Medicine Hat; [50.041 ° N, 110.677 ° W]; 24 Apr. 1926; Owen Bryant leg.; CAS • 1 ♀; Cypress Co., Medicine Hat, near Gas City Campground; 50.039 ° N, 110.726 ° W; 29 May 2014; M. Buck leg.; PMAE • 1 ♀; same data as for preceding; 31 May – 4 Jun. 2016; M. Buck leg.; PMAE • 4 ♀♀; same data as for preceding; 31 May – 4 Jun. 2016; M. Buck leg.; PMAE • 1 ♀; same data as for preceding; 1 Jun. 2016; M. Buck leg.; PMAE • 1 ♂; Forty Mile Co., Manyberries; [49.402 ° N, 110.693 ° W]; 7 Aug. 1981; D. B. McCorquodale leg.; PMAE • 2 ♂♂; Newell Co., Dinosaur Provincial Park, Sandhill Creek Valley; 50.7558 ° N, 111.5192 ° W; 14 – 15 Jul. 2010; Buck and Huele leg.; PMAE • 3 ♂♂; Warner Co., Writing-On-Stone Provincial Park; [49.0819 ° N, 111.6169 ° W]; 1 Aug. 1990; M. Klassen leg.; PMAE • 1 ♂; Warner Co., Writing-On-Stone Provincial Park; [49.0819 ° N, 111.6169 ° W]; 8 Aug. 1990; M. Klassen leg.; PMAE • 6 ♂♂; same data as for preceding; 18 – 25 Jul. 1990; M. Klassen leg.; PMAE • 1 ♀; same data as for preceding; 26 May – 6 Jun. 1990; D. McCorquodale leg.; PMAE • 1 ♀; same data as for preceding; 5 – 13 Jul. 1990; M. Klassen leg.; PMAE • 5 ♂♂; same data as for preceding; 5 – 13 Jul. 1990; M. Klassen leg.; PMAE • 1 ♂; Warner Co., Writing-On-Stone Provincial Park; [49.0819 ° N, 111.6169 ° W]; 30 Aug. – 10 Sep. 1990; M. Klassen leg.; PMAE • 1 ♂; same data as for preceding; 16 – 25 Jul. 1990; M. Klassen leg.; PMAE • 2 ♂♂; Warner Co., Writing-On-Stone Provincial Park; [49.0819 ° N, 111.6169 ° W]; 16 Jul. 1990; M. Klassen leg.; PMAE • 1 ♀; Drumheller; 51.465 ° N, 112.71 ° W; 8 Jun. 2013; R. Poulin leg.; RSKM • 2 ♀♀; Lethbridge, Gold Canyon; 49.6547 ° N, 112.8197 ° W; 3 Jul. 2020; M. Buck leg.; PMAE. – British Columbia • 1 ♀; Central Interior, Prince George; 53.9294 ° N, 122.75 ° W; elev. 572 m; 12 Jun. 2022; A. White leg.; CBG • 1 ♂; East Kootenay B, Kikomun Creek Provincial Park; 49.25366 ° N, 115.23547 ° W; 12 Jul. – 19 Aug. 2021; S. Achal and H. Bates (J. Heron) leg.; RSKM • 1 ♂; East Kootenay B, Kikomun Creek Provincial Park; 49.25364 ° N, 115.23608 ° W; 14 Jun. – 12 Jul. 2021; S. Achal and H. Bates (J. Heron) leg.; RSKM • 1 ♀; Kamloops; [50.675 ° N, 120.327 ° W]; 17 Apr. 1943; E. R. Buckell leg.; EMEC • 1 ♀; same data as for preceding; 13 Apr. 1943; G. J. Spencer leg.; EMEC • 6 ♀♀; Kamloops; [50.675 ° N, 120.327 ° W]; 1 May 1932; Owen Bryant leg.; CAS • 1 ♀; Okanagan-Similkameen Reg. Dist., Camp McKinney Road, Sandy Cut; 49.1135 ° N, 119.3406 ° W; elev. 1190 m; 26 Apr. – 3 May 2009; L. R. Best leg.; PCYU • 1 ♀; Okanagan-Similkameen Reg. Dist., Mt Kruger, Strawberry Creek Road; 49.0359 ° N, 119.5007 ° W; elev. 390 m; 21 Apr. 2009; L. R. Best leg.; PCYU • 5 ♀♀; Okanagan-Similkameen; 49.16569 ° N, 119.73643 ° W; 28 Aug. – 9 Sep. 2015; D. Marks leg.; RSKM (Fig. 40 D) • 1 ♀; Okanagan-Similkameen, SOGPA, Nighthawk Road; 49.01614 ° N, 119.67998 ° W; 14 Jun. – 16 Jul. 2021; H. Singh and T. McGratton (JH) leg.; RSKM • 1 ♀; Osoyoos, Haynes Ecol. Res; [49.083 ° N, 119.515 ° W]; 20 May 1987; S. G. Cannings leg.; EMEC • 1 ♀; Osoyoos, Osoyoos West Bench; 49.00142 ° N, 119.49765 ° W; 19 – 20 Jul. 2017; D. Marks and K. Grant (J. Heron) leg.; RSKM • 1 ♀; Osoyoos, Osoyoos West Bank; 49.028954 ° N, 119.496864 ° W; 19 May – 8 Jul. 2016; C. Dawson and M. Senelli leg.; RSKM • 1 ♀; Osoyoos, Osoyoos West Bench; 49.02871 ° N, 119.49376 ° W; 14 Jun. – 17 Jul. 2017; D. Marks and K. Grant (J. Heron) leg.; RSKM • 1 ♀; Osoyoos, Osoyoos West Bench; 49.00233 ° N, 119.49658 ° W; 19 Jun. – 20 Jul. 2017; D. Marks and K. Grant (J. Heron) leg.; RSKM • 1 ♀; Osoyoos, Osoyoos West Bench; 49.01193 ° N, 119.4951 ° W; 12 Jul. – 8 Aug. 2017; D. Marks and K. Grant (J. Heron) leg.; RSKM • 1 ♂; Osoyoos, Osoyoos West Bench; 49.03033 ° N, 119.50034 ° W; 17 Jul. – 9 Aug. 2017; D. Marks and K. Grant (J. Heron) leg.; RSKM • 2 ♀♀; Osoyoos, Strawberry Creek Road; 49.038471 ° N, 119.508261 ° W; 28 Jun. 2020; T. Haapalainen and K. Needham leg.; EMEC • 1 ♀; Oliver; [49.183 ° N, 119.551 ° W]; 26 May 1945; D. Black leg.; EMEC • 1 ♀; Penticton; [49.499 ° N, 119.594 ° W]; 20 May 1919; E. R. Buckell leg.; CUIC • 1 ♀; Petitot River, Liard Hwy; [59.99 ° N, 123.1 ° W]; 26 Jun. 1985; B. A. Macdonald leg.; EMEC • 1 ♀; Robson; [49.334 ° N, 117.691 ° W]; 3 May 1957; R. R. Foxlee leg.; EMEC • 17 ♂♂; Shore, Shuswap Lake, near Salmon Arm; [50.700 ° N, 119.283 ° W]; 13 Jul. 1949; H. B. Leech leg.; CAS • 1 ♂; Columbia Shuswap, Shuswap Lake Shore, Salmon Arm; [50.7049 ° N, 119.2874 ° W]; 13 Jul. 1949; H. B. Leech leg.; EMEC • 1 ♀; Vernon; [50.267 ° N, 119.272 ° W]; 3 May 1928; P. N. Vroom leg.; EMEC • 1 ♀; Vernon, CFB Vernon; 50.226 ° N, 119.302 ° W; Jun. 2018; T. Ehlers leg.; RSKM. – Saskatchewan • 1 ♀; RM of Val Marie, Grasslands National Park; 49.1361 ° N, 107.6306 ° W; 8 Jul. 2010; N. de Silva leg.; PMAE. MEXICO – Baja California • 1 ♂; Ensenada Co., Sierra San Pedro Martir, La Encantada; [30.98 ° N, 115.392 ° W]; elev. 2134 m; 31 May 1958; J. Powell leg.; EMEC • 1 ♂; Sierra San Pedro Mártir, slope NE of Vallecitos; [31.05 ° N, 115.45 ° W]; 14 Jul. 1980; Brown Faulkner leg.; SDMC. – Sonora • 1 ♀; Alamos Municipality Co., San Bernardo, Rio Mayo; [27.4138 ° N, 108.8621 ° W]; 17 Aug. 1935; EMEC. UNITED STATES – Arizona • 2 ♂♂; Apache Co., Subalpine meadow, White Mountains; [33.850 ° N, 109.57 ° W]; A. R. Moldenke leg.; CAS • 1 ♂; Apache Co., White Mountains; [33.850 ° N, 109.4 ° W]; 7 Jul. 1933; Parker leg.; CAS • 1 ♀; Cochise Co., 3 mi. SW of Portal, along Cave Creek; 31.88687 ° N, 109.17358 ° W; elev. 1550 m; 25 – 30 Jun. 1987; SEMC • 1 ♀; Cochise Co., Chiricahua Mountains; [31.883 ° N, 109.208 ° W]; 9 Mar. 1917; V. W. Owen leg.; CAS • 1 ♂; Cochise Co., Mountain meadows, Chiricahua Mountains; [31.883 ° N, 109.208 ° W]; J. L. Neff and A. R. Moldenke leg.; CAS • 1 ♂; same data as for preceding; J. L. Neff leg.; CAS • 1 ♀; Cochise Co., Ramsey Canyon, 12 km S of Sierra Vista; [31.446 ° N, 110.303 ° W]; 22 Jun. 1987; B. Brown leg.; PMAE • 1 ♀; Cochise Co., S of Sierra Vista, Ramsey Canyon, Mile High Preserve; [31.488 ° N, 110.408 ° W]; elev. 1700 m; 1 – 8 Oct. 1986; B. Brown leg.; PMAE • 1 ♂; Coconino or Mohave Co., Grand Canyon; [36.300 ° N, 112.75 ° W]; elev. 2134 m; 22 Jun. 1925; A. A. Nichol leg.; ASUHIC • 2 ♀♀; Gila Co., Parker Creek; [33.747 ° N, 110.983 ° W]; 19 Apr. 1947; SEMC • 1 ♀; same data as for preceding; 2 May 1947; SEMC • 1 ♀; Graham Co., Marijilda Canyon, Graham Mountains; [32.701 ° N, 109.785 ° W]; 7 Apr. 1961; J. C. Bequaert leg.; ex Phacelia distans; ASUHIC • 1 ♀; Maricopa Co., 15 mi. E of Gila Bend; [32.945 ° N, 112.459 ° W]; 16 Feb. 1953; E. I. Schlinger leg.; EMEC • 1 ♂; Maricopa Co., 15 mi. W of Chandler; [33.306 ° N, 112.101 ° W]; 9 Jun. 1961; G. D. Butler leg.; ex Heliotropium; ASUHIC • 1 ♂; Maricopa Co., Gila Bend; [32.945 ° N, 112.717 ° W]; 11 May 1978; E. G. and J. M. Linsley leg.; ex Parkinsonia microphylla (verbatim: Cercidium microphyllum); EMEC • 1 ♂; same data as for preceding; 10 May 1978; E. G. and J. M. Linsley leg.; ex Parkinsonia microphylla (verbatim: Cercidium microphyllum); CUIC • 1 ♀; Maricopa Co., Granite Reef Dam; [33.516 ° N, 111.691 ° W]; 30 Mar. 1959; R. S. Beal leg.; ex Salix (verbatim: Salix catkins); ASUHIC • 1 ♂; Maricopa Co., Phoenix; [33.450 ° N, 112.07 ° W]; 20 Apr. 1933; R. H. Crandall leg.; ASUHIC • 1 ♀; Maricopa Co., Tortilla Flat; [33.530 ° N, 111.388 ° W]; 11 Apr. 1966; F. F. Hasbrouck leg.; ex Baileya multiradiata; ASUHIC • 4 ♀♀; Mohave Co., Peach Springs, N of Colorado River; 35.76667 ° N, 113.3717 ° W; 18 Mar. 2000; D. Brzoska leg.; ex Chrysothamnus; SEMC • 1 ♂; Mojave Co., Hualapai Mountains, S of Kingman; [35.078 ° N, 113.974 ° W]; elev. 1900 m; 30 Jul. 1965; M. E. Irwin leg.; UCRC • 3 ♂♂; same data as for preceding; 30 Jul. 1965; E. I. Schlinger leg.; UCRC • 2 ♂♂; Navajo Co., 23 mi. W of Winslow; [35.024 ° N, 111.103 ° W]; 3 Jul. 1953; E. S. Ross leg.; CAS • 1 ♂; Navajo Co., 5 mi. W of Fort Apache; [33.790 ° N, 110.077 ° W]; 21 Jun. 1957; F. Werner and G. Butler leg.; ASUHIC • 1 ♂; Navajo Co., Jadito Trading Post; [35.767 ° N, 110.133 ° W]; 28 Jun. 1966; J. H. Davidson, J. M. Davidson and M. A. Cazier leg.; ex Poliomintha incana; ASUHIC • 1 ♂; Navajo Co., Show Low; [34.253 ° N, 110.03 ° W]; 22 Aug. 1973; G. and K. Eickwort leg.; CUIC • 1 ♀; Pima Co., 37 mi. SW of Tucson; [31.842 ° N, 111.42 ° W]; 19 – 20 Mar. 1961; R. H. Painter leg.; ASUHIC • 1 ♀; Pima Co., Catalina Mountains; [32.430 ° N, 110.705 ° W]; 24 Mar. 1956; A. D. Telford leg.; ASUHIC • 1 ♂; Pima Co., Catalina Mountains, 19 mi. along Hitchhiker Highway; [32.488 ° N, 110.447 ° W]; 9 Jul. 1957; F. Werner leg.; ex Ceanothus; ASUHIC • 4 ♂♂; Pima Co., Catalina Mountains, 24 mi. along Hitchhiker Highway; [32.539 ° N, 110.387 ° W]; 21 Jul. 1957; F. Werner leg.; ASUHIC • 1 ♂; Pima Co., Catalina Mountains, 26 mi. along Hitchhiker Highway; [32.559 ° N, 110.362 ° W]; 9 Jun. 1956; F. G. Werner and G. D. Butler leg.; ASUHIC • 1 ♀; Pima Co., Catalina Mountains, 5 miles along Hitchcock Highway; [32.344 ° N, 110.617 ° W]; 17 Apr. 1955; G. D. Butler, F. G. Werner leg.; ex Fendlera; ASUHIC • 2 ♀♀; same data as for preceding; 10 Apr. 1955; G. D. Butler leg.; ex Salix; ASUHIC • 2 ♀♀; same data as for preceding; 9 Apr. 1955; G. D. Butler and GF. G. Werner leg.; ex Fendlera; ASUHIC • 3 ♀♀; same data as for preceding; 10 Apr. 1955; G. D. Butler leg.; ex Salix; ASUHIC • 1 ♀; same data as for preceding; 24 Mar. 1956; W. Nutting leg.; ASUHIC • 1 ♂; Pima Co., Desert Scrub Site, Santa Catalina Mountains; [32.350 ° N, 110.7 ° W]; J. L. Neff leg.; CAS • 1 ♂; Pima Co., Mt Lemmon, Santa Catalina Mountains; [32.442 ° N, 110.788 ° W]; 28 Jul. 1924; E. P. Van Duzee leg.; CAS • 1 ♂; Pima Co., Mt Lemmon, Santa Catalina Mountains; [32.442 ° N, 110.788 ° W]; elev. 2790 m; 1 Aug. 1992; L. Packer leg.; PCYU • 33 ♂♂; Pima Co., Quercus / Pinus Forest, Santa Catalina Mountains; [32.416 ° N, 110.7 ° W]; J. L. Neff leg.; CAS • 1 ♂; Pima Co., S slope of Kitt Peak, Quinlan Mountains; [31.961 ° N, 111.6 ° W]; 22 Apr. 1961; M. L. Noller leg.; ex Compositae; ASUHIC • 2 ♂♂; Pima Co., Saguaro Park; [32.183 ° N, 110.613 ° W]; 9 May 1942; A. L. Melander leg.; UCRC • 3 ♂♂; Pima Co., Santa Catalina Mountains; [32.443 ° N, 110.788 ° W]; 10 Jun. 1938; Bryant Lot leg.; CAS • 1 ♀; same data as for preceding; 10 May 1938; Bryant Lot leg.; CAS • 1 ♂; Pima Co., Tucson; [32.222 ° N, 110.926 ° W]; 4 May 1942; A. L. Melander leg.; UCRC • 1 ♂; same data as for preceding; 14 May 1961; R. H. and E. M. Painter leg.; ASUHIC • 2 ♂♂; same data as for preceding; 9 May 1953; G. D. Butler leg.; ex Lobularia maritima; ASUHIC • 1 ♂; same data as for preceding; 30 Apr. 1953; G. D. Butler leg.; ASUHIC • 1 ♀; Pima Co., Tucson (A-Mt); [32.213 ° N, 110.995 ° W]; 27 Mar. 1954; R. H. Painter leg.; ex Larrea tridentata; ASUHIC • 2 ♂♂; Pima Co., Tucson Mountains; [32.224 ° N, 111.117 ° W]; 27 Mar. 1954; G. D. Butler leg.; ASUHIC • 1 ♀; same data as for preceding; 27 Mar. 1954; G. D. Butler leg.; ASUHIC • 1 ♂; Pinal Co., B. Thompson Arboretum, Superior; [33.279 ° N, 111.159 ° W]; 3 May 1953; G. D. Butler leg.; ex Opuntia; ASUHIC • 3 ♀♀; Pinal Co., Coolidge; [32.977 ° N, 111.517 ° W]; 28 Feb. 1956; G. D. Butler leg.; ex Sphaeralcea; ASUHIC • 1 ♂; Pinal Co., Mammoth; [32.722 ° N, 110.64 ° W]; 3 May 1956; F. Werner and G. Butler leg.; ASUHIC • 1 ♀; Pinal Co., Peppersauce Canyon, northern foothills of the Santa Catalina Mountains; [32.520 ° N, 110.7 ° W]; 11 Apr. 1961; J. C. Bequaert leg.; ASUHIC • 1 ♀; Santa Cruz Co., 4 mi. SW of Patagonia; [31.500 ° N, 110.801 ° W]; 19 Feb. 1963; M. L. Noller leg.; ex Barkleyanthus salicifolius (verbatim: Senecio salignus); ASUHIC • 3 ♀♀; Santa Cruz Co., Patagonia; 31.8833 ° N, 111.2833 ° W; 24 Mar. 1994; B. V. Brown and E. Wilk leg.; RSKM • 1 ♀; same data as for preceding; 20 Mar. 1994; E. Wilk, B. Brown leg.; RSKM • 2 ♀♀; Santa Cruz Co., Patagonia; [31.883 ° N, 111.283 ° W]; 26 Mar. 1955; G. D. Butler and F. G. Werner leg.; ex Barkleyanthus salicifolius (verbatim: Senecio salignus); ASUHIC • 1 ♂; Santa Cruz Co., Sycamore Canyon; [31.365 ° N, 110.774 ° W]; J. L. Neff leg.; CAS • 1 ♀; Yavapai Co., 3 mi. S of Camp Verde; [34.520 ° N, 111.854 ° W]; 26 Mar. 1959; F. G. Werner leg.; ASUHIC • 2 ♂♂; Yavapai Co., Prescott; [34.539 ° N, 112.468 ° W]; elev. 1640 m; 27 Jul. 1932; P. H. Timberlake leg.; ex Asclepias tuberosa; UCRC (Fig. 41 F) • 1 ♂; same data as for preceding; 28 Jul. 1932; P. H. Timberlake leg.; ex Ceanothus; UCRC • 1 ♂; same data as for preceding; 27 Jul. 1932; P. H. Timberlake leg.; ex Frangula californica subsp. tomentella (verbatim: Rhamnus tomentella); UCRC • 1 ♂; Yuma Co., 2 mi. S of Stone Cabin; [33.231 ° N, 114.241 ° W]; 12 May 1978; E. G. Linsley and J. M. Linsley leg.; ex Parkinsonia microphylla (verbatim: Cercidium microphyllum); EMEC (Fig. 41 D). – California • 1 ♀; Alameda Co., Livermore; [37.6819 ° N, 121.7681 ° W]; 14 Mar. 1937; G. Ferguson leg.; OSAC • 1 ♀; Alameda Co., Tesla; [40.493 ° N, 121.889 ° W]; 21 Apr. 1953; W. F. Ehrhardt leg.; EMEC • 1 ♀; Alpine Co., 3 mi. W of Monitor Pass; [38.645 ° N, 119.695 ° W]; 22 Jun. 1962; C. D. MacNeill leg.; CAS • 1 ♀; Colusa Co., 9.5 mi. E of Hough Springs; [39.161 ° N, 122.434 ° W]; 20 May 1963; P. H. Raven leg.; ex Clarkia modesta; EMEC • 1 ♀; Contra Costa Co., Antioch; [38.002 ° N, 121.807 ° W]; 23 Oct. 1938; J. W. MacSwain leg.; EMEC • 1 ♀; Contra Costa Co., Walnut Creek; [37.908 ° N, 122.067 ° W]; 7 Mar. 1937; ex Salix vestita (verbatim: Salix leiolepis); UCRC • 1 ♂; El Dorado Co., Lake Tahoe; [39.227 ° N, 120.082 ° W]; 6 – 8 Mar. 1925; J. D. Gunder leg.; CAS • 1 ♀; Fresno Co., 13 mi. W of Coalinga; [36.139 ° N, 120.592 ° W]; 3 Nov. 1955; R. R. Snelling leg.; EMEC • 9 ♀♀; Fresno Co., Ciervo Hills, 18 air mi. SW of Mendota; [36.567 ° N, 120.611 ° W]; 16 Mar. 1975; J. Powell and P. Rude leg.; ex Phacelia tanacetifolia; EMEC • 19 ♀♀; same data as for preceding; 18 Mar. 1975; J. Powell and P. Rude leg.; ex Phacelia tanacetifolia; EMEC • 1 ♀; same data as for preceding; 18 Mar. 1975; J. T. Doyen leg.; EMEC • 6 ♀♀; same data as for preceding; 18 Mar. 1975; J. Powell and P. Rude leg.; EMEC • 2 ♂♂; Glenn Co., Black Butte; [39.726 ° N, 122.872 ° W]; elev. 1981 – 2255 m; 14 Jun. 1972; D. F. Veirs leg.; EMEC • 1 ♂; Glenn Co., Plaskett Meadows; [39.727 ° N, 122.843 ° W]; elev. 1920 m; 14 Jun. 1972; J. Doyen leg.; EMEC • 1 ♂; same data as for preceding; 14 Jun. 1972; J. Doyen leg.; ex Allium siskiyouense; EMEC • 3 ♂♂; same data as for preceding; 14 Jun. 1972; J. Doyen leg.; ex Calyptridium umbellatum; EMEC • 2 ♀♀; Imperial Co., 1.6 mi. S of Truckhaven; [33.2789 ° N, 115.9639 ° W]; elev. - 25 m; 12 Feb. 1952; P. H. Timberlake leg.; ex Astragalus crotalariae; UCRC • 2 ♀♀; Inyo Co., 2 mi. W of Westgard Pass; [37.300 ° N, 118.19 ° W]; 16 May 1969; J. T. Doyen leg.; ex Prunus andersonii; EMEC • 2 ♂♂; Inyo Co., Big Pines; [37.166 ° N, 118.289 ° W]; 2 Aug. 1944; A. L. Melander leg.; UCRC • 1 ♂; Inyo Co., Darwin Falls; [36.320 ° N, 117.525 ° W]; 14 May 1969; J. Powell leg.; EMEC • 1 ♂; Inyo Co., Darwin Falls; [36.32 ° N, 117.525 ° W]; 14 May 1969; A. Rude leg.; EMEC • 1 ♀; Inyo Co., Glacier Lodge, Big Pine; [37.124 ° N, 118.438 ° W]; 23 Jun. 1937; E. C. VanDyke leg.; CAS • 3 ♀♀; Inyo Co., Goodale Creek, N. of Lone Pine; [36.716 ° N, 118.124 ° W]; 3 Apr. 1953; H. B. Leech leg.; ex Salix exigua; CAS • 1 ♀; Inyo Co., Mazourka Canyon, 8 mi. NE of Independence; [36.874 ° N, 118.091 ° W]; 11 May 1969; J. A. Chemsak leg.; EMEC • 4 ♀♀; same data as for preceding; 11 May 1969; P. A. Opler leg.; EMEC • 1 ♀; same data as for preceding; 11 May 1969; P. A. Rude leg.; EMEC • 1 ♀; same data as for preceding; 11 May 1969; J. Powell leg.; EMEC • 1 ♀; Inyo Co., Panamint Mountains; [36.113 ° N, 117.151 ° W]; 29 May 1937; N. W. Frazier leg.; EMEC • 2 ♂♂; Inyo Co., Surprise Canyon; [36.113 ° N, 117.151 ° W]; 5 May 1961; P. M. Marsh leg.; UCDC • 1 ♀; Inyo Co., Surprise Canyon, Panamint Mountains; [36.1150 ° N, 117.1428 ° W]; 24 Apr. 1957; P. D. Hurd leg.; ex Cryptantha utahensis; EMEC • 4 ♀♀; same data as for preceding; 24 Apr. 1957; P. D. Hurd leg.; ex Encelia farinosa; EMEC • 2 ♂♂; same data as for preceding; 24 Apr. 1957; P. D. Hurd leg.; ex Encelia farinosa; EMEC • 1 ♀; same data as for preceding; 24 Apr. 1957; G. I. Stage leg.; ex Encelia farinosa; EMEC • 1 ♂; same data as for preceding; 24 Apr. 1957; P. D. Hurd leg.; ex Encelia farinosa; UCDC • 2 ♀♀; same data as for preceding; 24 Apr. 1957; P. H. Timberlake leg.; ex Encelia farinosa; UCRC • 1 ♂; same data as for preceding; 24 Apr. 1957; P. H. Timberlake leg.; ex Encelia farinosa; UCRC • 1 ♀; same data as for preceding; 24 Apr. 1957; P. D. Hurd leg.; ex Eriogonum inflatum; EMEC • 1 ♀; same data as for preceding; 24 Apr. 1957; G. I. Stage leg.; ex Eucnide urens; EMEC • 1 ♂; same data as for preceding; 24 Apr. 1957; P. D. Hurd leg.; ex Larrea; EMEC • 1 ♂; same data as for preceding; 24 Apr. 1957; P. D. Hurd leg.; ex Larrea tridentata (verbatim: Larrea glutinosa); EMEC • 3 ♂♂; same data as for preceding; 24 Apr. 1957; G. I. Stage leg.; ex Lepidium fremontii; EMEC • 2 ♂♂; same data as for preceding; 24 Apr. 1957; P. D. Hurd leg.; ex Lepidium fremontii; EMEC • 7 ♂♂; same data as for preceding; 24 Apr. 1957; G. I. Stage leg.; ex Lepidium fremontii; EMEC • 3 ♂♂; same data as for preceding; 24 Apr. 1957; P. H. Timberlake leg.; ex Lepidium fremontii; UCRC • 1 ♀; same data as for preceding; 24 Apr. 1957; J. Powell leg.; ex Lepidium fremontii; EMEC • 1 ♀; same data as for preceding; 24 Apr. 1957; P. D. Hurd leg.; ex Phacelia calthifolia; EMEC • 2 ♀♀; same data as for preceding; 24 Apr. 1957; G. I. Stage leg.; ex Salvia dorrii var. incana (verbatim: Salvia carnosa); EMEC • 1 ♂; same data as for preceding; 24 Apr. 1957; G. I. Stage leg.; ex Sphaeralcea ambigua; EMEC • 3 ♀♀; same data as for preceding; 24 Apr. 1957; J. Powell leg.; ex Stanleya pinnata; EMEC • 1 ♀; same data as for preceding; 25 Apr. 1957; P. H. Timberlake leg.; ex Stanleya pinnata; UCRC • 5 ♂♂; same data as for preceding; 24 Apr. 1957; G. I. Stage leg.; EMEC • 1 ♀; same data as for preceding; 29 Apr. 1945; J. C. Hall leg.; UCRC • 3 ♀♀; same data as for preceding; 24 Apr. 1957; J. Powell leg.; EMEC • 1 ♀; Inyo Co., Tuttle Creek, 2 mi. SW of Lone Pine; [36.58 ° N, 118.095 ° W]; 14 May 1969; P. A. Rude leg.; EMEC • 1 ♂; Inyo Co., Wildrose Canyon, Panamint Mts; [36.172 ° N, 117.26 ° W]; 27 May 1959; Truxal and Martin leg.; EMEC • 1 ♀; Inyo Co., Wyman Canyon (Vic. Deep Springs); [37.387 ° N, 117.979 ° W]; 9 Apr. 1970; Suzy Frommer leg.; ex Salix; UCRC • 1 ♀; Inyo Co., Wyman Canyon, White Mountains; [37.409 ° N, 117.982 ° W]; 10 Jul. 1967; Saul and Suzy Frommer leg.; EMEC • 2 ♀♀; Inyo or San Bernardino Co., Death Valley; [36.532 ° N, 116.933 ° W]; 28 Mar. 1940; CAS • 3 ♀♀; Kern Co., 16 mi. S of Weldon; [35.432 ° N, 118.277 ° W]; 26 Apr. 1964; P. Rude leg.; ex Salix; EMEC • 1 ♀; Kern Co., 3 mi. W of Wofford Heights; [35.711 ° N, 118.511 ° W]; 29 Apr. 1964; C. A. Toschi leg.; EMEC • 1 ♂; Kern Co., 4 mi. W of Frazier Park; [34.825 ° N, 119.015 ° W]; 20 Jul. 1963; J. Powell leg.; EMEC • 15 ♀♀; Kern Co., Caliente; [35.291 ° N, 118.629 ° W]; elev. 400 m; 24 Mar. 1932; P. H. Timberlake leg.; ex Cleome isomeris (verbatim: Isomeris arborea); UCRC • 2 ♀♀; Kern Co., Canebrake Creek, 3 mi. W of Walker Pass; [35.661 ° N, 118.079 ° W]; 30 Apr. 1964; J. Powell leg.; ex Plagiobothrys nothofulvus; EMEC • 1 ♀; Kern Co., Frazier Park; [34.815 ° N, 118.947 ° W]; 3 May 1959; G. I. Stage leg.; ex Oenothera; EMEC • 1 ♀; Kern Co., Frazier Park; [34.822 ° N, 118.945 ° W]; 4 May 1958; EMEC • 1 ♀; Kern Co., Indian Wells Can., W. off Hwy 6, 2 mi. W of Inyo / Kern turnoff; [36.048 ° N, 117.986 ° W]; 12 Apr. 1960; C. A. Toschi leg.; EMEC • 3 ♂♂; Kern Co., Mt Pinos; [34.816 ° N, 119.145 ° W]; elev. 2438 m; 1 Jul. 1965; J. Powell leg.; EMEC • 1 ♂; same data as for preceding; 1 Jul. 1965; E. M. Omi leg.; EMEC • 1 ♀; Kern Co., Short Canyon, 6 mi. W of Inyokern; [35.644 ° N, 117.921 ° W]; 10 Apr. 1960; J. W. MacSwain leg.; EMEC • 2 ♀♀; Kern Co., Walker Pass; [35.6628 ° N, 118.027 ° W]; 26 Apr. 1949; E. G. Linsley, J. W. MacSwain and R. F. Smith leg.; ex Salix; EMEC • 1 ♀; Lake Co., 10 mi. E of Clearlake Oaks; [39.027 ° N, 122.484 ° W]; 11 May 1961; J. Powell leg.; EMEC • 1 ♀; Lake Co., Lake Pillsbury; [39.430 ° N, 122.95 ° W]; 4 Apr. 1962; J. Powell leg.; EMEC • 2 ♂♂; Lassen Co., 10 mi. S of Ravendale; [40.652 ° N, 120.367 ° W]; 5 Feb. 1964; R. C. Dickson leg.; UCRC • 7 ♀♀; Lassen Co., 3.5 mi. S of Ravendale; [40.746 ° N, 120.365 ° W]; 20 Jun. 1959; G. I. Stage leg.; ex Taraxia tanacetifolia (verbatim: Oenothera tanecetifolia); EMEC • 26 ♀♀; Lassen Co., 6 mi. SE of Doyle; [39.965 ° N, 120.024 ° W]; 5 Jun. 1970; P. A. Rude leg.; ex Sisymbrium; EMEC • 1 ♂; Lassen Co., Hallelujah Junction; [39.775 ° N, 120.039 ° W]; 2 Jul. 1954; M. E. Irwin leg.; UCRC • 1 ♀; same data as for preceding; 22 Jun. 1964; M. E. Irwin leg.; UCRC • 1 ♂; same data as for preceding; 4 Jul. 1949; J. E. Gillaspy leg.; EMEC • 1 ♀; Lassen Co.; [40.67 ° N, 120.59 ° W]; 3 Jun. 1913; F. W. Nunenmacher leg.; EMEC • 2 ♀♀; Los Angeles Co., 2 mi. S of Pearblossom; [34.469 ° N, 117.911 ° W]; elev. 1067 m; 7 May 1965; R. R. Snelling leg.; EMEC (Fig. 40 F) • 1 ♀; Los Angeles Co., 5 mi. S of Pearblossom; [34.433 ° N, 117.909 ° W]; 23 Apr. 1956; E. G. Linsley leg.; ex Tamarix gallica; EMEC • 1 ♀; Los Angeles Co., 6 mi. SE of Pearblossom; [34.451 ° N, 117.831 ° W]; elev. 1067 m; 16 Mar. 1972; R. R. Snelling leg.; EMEC • 24 ♂♂; Los Angeles Co., Big Pine Camp; [34.3956 ° N, 117.7339 ° W]; elev. 1850 m; 2 Aug. 1944; P. H. Timberlake leg.; ex Eriodictyon trichocalyx; UCRC (Fig. 47) • 1 ♀; Los Angeles Co., Little Rock; [34.517 ° N, 117.993 ° W]; 12 Apr. 1936; R. M. Bohart and G. E. Bohart leg.; CAS • 1 ♀; Los Angeles Co., Westwood Hills; [34.056 ° N, 118.43 ° W]; 27 Jul. 1935; E. G. Linsley leg.; UCRC • 1 ♂; Madera Co., Buck Creek; [37.542 ° N, 119.487 ° W]; 20 Jul. 1922; C. L. Fox leg.; CAS • 1 ♀; Mariposa Co., Miami Ranger Station; [37.419 ° N, 119.745 ° W]; 20 May 1942; E. G. Linsley leg.; UCRC • 1 ♀; same data as for preceding; 4 Jun. 1942; EMEC • 1 ♀; Mendocino Co., Ryan Creek; [39.476 ° N, 123.381 ° W]; 13 Apr. 1941; EMEC • 1 ♀; Merced Co., Snelling; [37.519 ° N, 120.439 ° W]; 18 Apr. 1939; J. J. duBois leg.; ex Salix; EMEC • 1 ♀; Modoc Co., 15 mi. E of Cedarville; [41.529 ° N, 119.881 ° W]; 3 Jul. 1935; Joe Schuh leg.; OSAC • 1 ♀; Modoc Co., 5.2 mi. NW of Cedarville; [41.584 ° N, 120.099 ° W]; elev. 1981 m; 13 Jun. 1969; R. R. and R. A. Snelling leg.; ex Phacelia; EMEC • 1 ♀; Modoc Co., 6 mi. NW of Cedarville; [41.591 ° N, 120.09 ° W]; 22 May 1955; J. W. MacSwain leg.; EMEC • 1 ♀; Modoc Co., Buck Creek Ranger Station; [41.870 ° N, 120.293 ° W]; elev. 1570 m; 5 – 7 Jun. 1970; P. Opler leg.; ex Isatis tinctoria; EMEC • 1 ♀; same data as for preceding; 5 – 7 Jun. 1970; J. T. Doyen leg.; ex Cirsium cymosum; EMEC • 1 ♂; Mono Co., 1 mi. W of Tom’s Place; [37.561 ° N, 118.699 ° W]; D. D. Linsdale leg.; EMEC • 3 ♂♂; Mono Co., 4 mi. E of Monitor Pass; [38.677 ° N, 119.55 ° W]; 15 Jul. 1966; P. A. Rude leg.; EMEC • 1 ♂; Mono Co., 4 mi. SW of Tom’s Place; [37.520 ° N, 118.733 ° W]; 13 Jul. 1961; H. V. Daly leg.; EMEC • 2 ♂♂; Mono Co., 8 mi. NW of Bridgeport; [38.337 ° N, 119.336 ° W]; 9 Sep. 1957; J. A. Chemsak leg.; ex Chrysothamnus; EMEC • 1 ♀; Mono Co., Bodie; [38.216 ° N, 119.01 ° W]; 13 Jul. 1937; Dr John A. Comstock leg.; EMEC • 1 ♀; Monterey Co., Tassajara Hot Springs; [36.237 ° N, 121.558 ° W]; elev. 457 m; 26 May 1954; Owen Bryant leg.; CAS • 1 ♀; Nevada Co., Sagehen Creek, near Hobart Mills; [39.432 ° N, 120.201 ° W]; 16 Jul. 1964; M. E. Irwin leg.; UCRC • 1 ♀; Riverside Co., 2 mi. N of Palm Springs; [33.8742 ° N, 116.5056 ° W]; elev. 175 m; 7 Mar. 1936; P. H. Timberlake leg.; ex Larrea tridentata var. tridentata (verbatim: Larrea divaricata); EMEC • 4 ♀♀; same data as for preceding; 7 Mar. 1936; P. H. Timberlake leg.; ex Larrea tridentata var. tridentata (verbatim: Larrea divaricata); UCRC • 1 ♀; Riverside Co., 2 mi. S of Oasis; [33.4319 ° N, 116.0669 ° W]; elev. - 35 m; 8 Mar. 1936; P. H. Timberlake leg.; ex Larrea tridentata var. tridentata (verbatim: Larrea divaricata); EMEC (Fig. 40 E) • 1 ♀; same data as for preceding; 8 Mar. 1936; P. H. Timberlake leg.; ex Condea emoryi (verbatim: Hyptis emoryi); EMEC • 5 ♀♀; same data as for preceding; 8 Mar. 1936; P. H. Timberlake leg.; ex Larrea tridentata var. tridentata (verbatim: Larrea divaricata); UCRC • 1 ♂; Riverside Co., 3 mi. E of Ribbonwood; [33.570 ° N, 116.447 ° W]; 17 May 1963; E. I. Schlinger leg.; ex Yucca; UCRC • 1 ♀; Riverside Co., 3 mi. E of Ribbonwood; [33.57 ° N, 116.447 ° W]; 17 May 1963; E. I. Schlinger leg.; ex Yucca; EMEC • 1 ♀; Riverside Co., Andreas Canyon, near Palm Springs; [33.7614 ° N, 116.5522 ° W]; elev. 290 m; 23 Feb. 1929; P. H. Timberlake leg.; UCRC • 4 ♀♀; same data as for preceding; 5 Mar. 1933; P. H. Timberlake leg.; ex Prunus fremontii; UCRC • 1 ♀; same data as for preceding; 5 Mar. 1933; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 3 ♀♀; same data as for preceding; 22 Feb. 1939; P. H. Timberlake leg.; ex Cleome isomeris (verbatim: Isomeris arborea); UCRC • 1 ♀; same data as for preceding; 24 Mar. 1933; P. H. Timberlake leg.; ex Larrea tridentata; UCRC • 1 ♀; Riverside Co., Berdoo Canyon; [33.805 ° N, 116.176 ° W]; 5 Mar. 1968; J. C. Hall leg.; ex Poppy; UCRC • 4 ♀♀; Riverside Co., Colorado Desert, 5 mi. N of Palm Springs; [33.9175 ° N, 116.5017 ° W]; elev. 240 m; 7 Mar. 1934; P. H. Timberlake leg.; ex Cleome isomeris (verbatim: Isomeris arborea); UCRC • 16 ♀♀; Riverside Co., Colorado Desert, Varner Road, 5 mi. NW of Indio; [33.7667 ° N, 116.3208 ° W]; elev. 40 m; 18 Feb. 1928; P. H. Timberlake leg.; ex Sisymbrium; UCRC • 1 ♀; Riverside Co., Deep Canyon; [33.713 ° N, 116.365 ° W]; 24 Feb. 1964; M. E. Irwin leg.; ex Encelia; UCRC • 1 ♀; same data as for preceding; 1 Apr. 1964; E. I. Schlinger leg.; ex Hyptis; UCRC • 2 ♀♀; same data as for preceding; 27 Feb. 1963; E. I. Schlinger leg.; UCRC • 1 ♀; same data as for preceding; 24 Feb. 1964; M. E. Irwin leg.; UCRC • 2 ♀♀; same data as for preceding; 6 Mar. 1963; R. C. Dickson leg.; UCRC • 1 ♀; same data as for preceding; 15 Mar. 1963; E. I. Schlinger leg.; ex Hyptis; UCRC • 2 ♀♀; Riverside Co., Devil’s Garden; [34.0269 ° N, 116.6 ° W]; elev. 910 m; 24 Feb. 1967; P. H. Timberlake leg.; ex Cleome isomeris (verbatim: Isomeris arborea); UCRC • 1 ♂; Riverside Co., Gavilan Hills; [33.7944 ° N, 117.3558 ° W]; elev. 620 m; 14 Apr. 1965; G. R. Ballmer leg.; UCRC • 1 ♀; same data as for preceding; 6 Feb. 1940; P. H. Timberlake leg.; ex Salix vestita (verbatim: Salix leiolepis); UCRC • 1 ♀; same data as for preceding; 6 Feb. 1940; P. H. Timberlake leg.; ex Ribes indecorum; UCRC • 1 ♀; same data as for preceding; 16 Mar. 1938; P. H. Timberlake leg.; ex Ribes indecorum; UCRC • 1 ♀; Riverside Co., La Quinta; [33.6725 ° N, 116.3036 ° W]; elev. 15 m; 17 Feb. 1935; P. H. Timberlake leg.; ex Condea emoryi; UCRC • 2 ♀♀; Riverside Co., Lost Palms Canyon; [33.715 ° N, 115.748 ° W]; 9 Apr. 1935; R. C. Dick leg.; UCRC • 1 ♀; Riverside Co., near Palm Canyon; [33.7350 ° N, 116.5361 ° W]; elev. 250 m; 13 Mar. 1932; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; same data as for preceding; 15 Jun. 1941; P. H. Timberlake leg.; ex Cleome isomeris (verbatim: Isomeris arborea); UCRC • 1 ♀; Riverside Co., P. L. Boyd Desert Research Center. 3.5 mi. S of Palm Desert; [33.648 ° N, 116.377 ° W]; 4 – 6 May 1970; S. Frommer and R. Worley leg.; UCRC • 1 ♀; same data as for preceding; 25 – 27 Mar. 1978; UCRC • 1 ♀; Riverside Co., Palm Canyon; [33.747 ° N, 116.548 ° W]; 10 Feb. 1952; ASUHIC • 1 ♀; Riverside Co., Palm Springs; [33.829 ° N, 116.547 ° W]; 26 Feb. 1935; A. L. Melander leg.; UCRC • 1 ♀; Riverside Co., Railroad Canyon, 4 mi. E of Elsinore; [33.673 ° N, 117.202 ° W]; 17 Apr. 1965; C. Slobodchikoff leg.; EMEC • 1 ♀; Riverside Co., Riverside; [33.9556 ° N, 117.3722 ° W]; elev. 290 m; 4 Mar. 1926; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 4 ♀♀; same data as for preceding; 6 Mar. 1926; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; same data as for preceding; 11 Mar. 1926; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; same data as for preceding; 31 Mar. 1926; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; same data as for preceding; 1 Apr. 1929; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; same data as for preceding; 7 Feb. 1930; P. H. Timberlake leg.; ex Sinapis alba or Brassica alba (verbatim: Brassica alba); UCRC • 1 ♀; same data as for preceding; 12 Feb. 1930; P. H. Timberlake leg.; ex Sisymbrium; UCRC • 1 ♀; same data as for preceding; 3 Apr. 1930; P. H. Timberlake leg.; UCRC • 1 ♀; same data as for preceding; 4 Apr. 1930; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; same data as for preceding; 23 Mar. 1933; P. H. Timberlake leg.; ex Sisymbrium; UCRC • 1 ♀; same data as for preceding; 23 Feb. 1934; P. H. Timberlake leg.; ex Viburnum tinus; UCRC • 1 ♀; same data as for preceding; 22 Jun. 1935; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; same data as for preceding; 4 Apr. 1939; P. H. Timberlake leg.; ex Ellisia; UCRC • 1 ♀; same data as for preceding; 23 Feb. 1940; P. H. Timberlake leg.; ex Phacelia minor; UCRC • 2 ♀♀; same data as for preceding; 27 Feb. 1940; P. H. Timberlake leg.; ex Phacelia minor; UCRC • 1 ♀; same data as for preceding; 28 Feb. 1940; P. H. Timberlake leg.; ex Phacelia minor; UCRC • 1 ♀; same data as for preceding; 1 Mar. 1940; P. H. Timberlake leg.; ex Phacelia minor; UCRC • 1 ♀; same data as for preceding; 1 Mar. 1940; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 2 ♀♀; same data as for preceding; 1 Mar. 1940; P. H. Timberlake leg.; ex Phacelia minor; UCRC • 1 ♀; same data as for preceding; 1 Mar. 1940; P. H. Timberlake leg.; ex Cryptantha intermedia; UCRC • 1 ♀; same data as for preceding; 5 Mar. 1940; P. H. Timberlake leg.; ex Phacelia minor; UCRC • 1 ♀; same data as for preceding; 8 Mar. 1940; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 2 ♀♀; same data as for preceding; 17 Mar. 1941; P. H. Timberlake leg.; UCRC • 3 ♀♀; same data as for preceding; 5 Apr. 1948; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; Riverside Co., San Jacinto River – North Fork at Hwy 74; [33.7303 ° N, 116.8097 ° W]; elev. 640 m; 22 Feb. 1936; P. H. Timberlake leg.; ex Salix exigua var. hindsiana (verbatim: Salix hindsiana); UCRC • 1 ♂; Riverside Co., Santa Rosa Mountain, San Jacinto Mountains; [33.820 ° N, 116.675 ° W]; elev. 2286 m; 31 May 1940; C. D. Michener leg.; ex Eremocarya micrantha (verbatim: Cryptantha micrantha); EMEC • 4 ♂♂; Riverside Co., Santa Rosa Peak, Santa Rosa Mountains; [33.5408 ° N, 116.4656 ° W]; elev. 2286 m; 8 Jun. 1940; P. H. Timberlake leg.; ex Eremocarya micrantha (verbatim: Cryptantha micrantha); UCRC • 1 ♀; Riverside Co., Snow Creek, 5 mi. N of Palm Springs; [33.8989 ° N, 116.6431 ° W]; elev. 340 m; 24 Feb. 1954; P. H. Timberlake leg.; ex Cleomella arborea (verbatim: Isomeris arborea); UCRC • 1 ♀; Riverside Co., Soboba Hot Springs; [33.800 ° N, 116.929 ° W]; elev. 500 m; 22 Jul. 1936; P. H. Timberlake leg.; ex Baccharis salicifolia (verbatim: Baccharis viminea); EMEC • 1 ♀; Riverside Co., Thousand Palms; [33.8178 ° N, 116.3944 ° W]; elev. 70 m; 27 Mar. 1941; P. H. Timberlake leg.; ex Hyptis emoryi; UCRC • 1 ♀; Riverside Co., Whitewater Canyon; [33.9878 ° N, 116.6561 ° W]; elev. 670 m; 24 Feb. 1954; P. H. Timberlake leg.; ex Salix exigua; UCRC • 3 ♀♀; Routt Co., 3 mi. E of Phippsburg; [40.232 ° N, 106.886 ° W]; 8 May 1954; R. S. Beal leg.; EMEC • 1 ♀; San Benito Co., 1 mi. S of Hernandez; [36.445 ° N, 120.967 ° W]; 25 May 1956; P. D. Hurd leg.; ex Emmenanthe penduliflora; EMEC • 1 ♀; San Benito Co., 5 mi. S of Bitterwater; [36.308 ° N, 121.001 ° W]; 31 Mar. 1959; C. W. O’Brien leg.; EMEC • 1 ♀; San Benito Co., Panoche Hills; [36.617 ° N, 120.851 ° W]; 10 Apr. 1938; M. C. Van Duzee leg.; CAS • 1 ♀; same data as for preceding; 10 Apr. 1938; E. C. VanDyke leg.; CAS • 4 ♀♀; San Bernardino Co., 15 km SSE of Hesperia, Hwy 173; [34.337 ° N, 117.259 ° W]; elev. 1022 m; 21 Apr. 1981; James A. Bethke leg.; UCRC • 2 ♀♀; same data as for preceding; 21 Apr. 1981; Gordon Pratt leg.; UCRC • 6 ♀♀; San Bernardino Co., 15 mi. S of Needles; [34.628 ° N, 114.615 ° W]; 31 Mar. 1959; P. H. Raven leg.; ex Oenothera brevipes; EMEC • 1 ♀; San Bernardino Co., 25 mi. Sof Ivanpah; [34.967 ° N, 115.284 ° W]; 13 Oct. 1958; J. A. Chemsak leg.; ex Chrysothamnus; EMEC • 1 ♀; San Bernardino Co., 7 Oaks; [34.186 ° N, 116.914 ° W]; 20 Jun. 1945; A. L. Melander leg.; UCRC • 1 ♂; San Bernardino Co., Big Bear Valley; [34.263 ° N, 116.822 ° W]; 8 Aug. 1953; P. H. Timberlake leg.; ex Eriogonum umbellatum; UCRC • 1 ♀; San Bernardino Co., Black Canyon Mojave Desert; [35.1336 ° N, 117.2614 ° W]; elev. 770 m; 7 Oct. 1935; P. H. Timberlake leg.; UCRC • 1 ♀; San Bernardino Co., Cactus Flat; [34.324 ° N, 116.815 ° W]; 28 Apr. 1950; T. F. Leigh leg.; EMEC • 1 ♂; San Bernardino Co., Cajon Pass; [34.3258 ° N, 117.45 ° W]; elev. 1067 m; 26 May 1941; P. H. Timberlake leg.; ex Prunus ilicifolia; UCRC • 1 ♀; San Bernardino Co., Camp Angelus; [34.209 ° N, 118.141 ° W]; 26 May 1947; A. L. Melander leg.; UCRC • 1 ♂; San Bernardino Co., Clark Mountain; [35.5192 ° N, 115.5836 ° W]; elev. 1981 m; 8 Jul. 1936; P. H. Timberlake leg.; ex Agave utahensis var. nevadensis (verbitim: Agave nevadensis); UCRC • 1 ♂; San Bernardino Co., Clark Mountain; [35.5217 ° N, 115.6428 ° W]; elev. 1372 m; 8 Jul. 1936; P. H. Timberlake leg.; ex Eriogonum inflatum; UCRC • 1 ♂; San Bernardino Co., Granite Pass; [34.812 ° N, 115.61 ° W]; elev. 1160 – 1280 m; 26 Apr. 1968; J. Devol leg.; UCRC • 1 ♂; same data as for preceding; 27 Apr. 1968; R. Hobza leg.; ex Eriogonum fasciculatum; UCRC • 1 ♂; San Bernardino Co., Granite Pass, near Kelso; [34.812 ° N, 115.611 ° W]; 27 Apr. 1968; H. E. Evans leg.; MCZ • 1 ♂; San Bernardino Co., Little Morongo Canyon; [33.989 ° N, 116.525 ° W]; 1 May 1967; J. C. Hall leg.; UCRC • 1 ♀; San Bernardino Co., Lone Pine Canyon; [34.3217 ° N, 117.5581 ° W]; elev. 1450 m; 16 Jun. 1928; P. H. Timberlake leg.; UCRC • 1 ♂; San Bernardino Co., March’s Ranch Rd 2 mi. E of Baldwin Lake; [34.268 ° N, 116.831 ° W]; elev. 1525 – 1830 m; 25 May 1968; R. Hobza leg.; ex Haplopappus; UCRC • 1 ♀; San Bernardino Co., Morongo Valley; [34.046 ° N, 116.581 ° W]; 20 Apr. 1957; R. R. Snelling and M. D. Stage leg.; ex Gilia; EMEC • 1 ♀; same data as for preceding; 15 Oct. 1934; ex Gutierrezia microcephala (verbatim: Gutierrezia lucida); UCRC • 1 ♂; San Bernardino Co., mouth of Deep Creek; [34.3414 ° N, 117.235 ° W]; elev. 910 m; 5 May 1936; P. H. Timberlake leg.; ex Eriodictyon trichocalyx; EMEC • 1 ♀; same data as for preceding; 5 May 1936; P. H. Timberlake leg.; ex Eriodictyon trichocalyx; UCRC • 3 ♂♂; same data as for preceding; 5 May 1936; P. H. Timberlake leg.; ex Eriodictyon trichocalyx; UCRC • 2 ♂♂; San Bernardino Co., Phelan. Vic. Sheep Creek Rd; [34.429 ° N, 117.573 ° W]; 4 May 1966; Saul Frommer leg.; UCRC • 2 ♀♀; San Bernardino Co., Providence Mountains, Bonanza King Mine; [34.981 ° N, 115.506 ° W]; 16 Mar. 1967; D. C. Rentz leg.; EMEC • 1 ♀; San Bernardino Co., Tetley’s Mountain Camp, Crestline; [34.2464 ° N, 117.3094 ° W]; elev. 1380 m; 15 May 1937; P. H. Timberlake leg.; ex Malus; UCRC • 1 ♀; same data as for preceding; 15 May 1937; E. G. Linsley leg.; ex Malus; CAS • 1 ♂; San Bernardino Co., Valley of the Falls; [34.088 ° N, 116.929 ° W]; 28 Aug. 1935; P. H. Timberlake leg.; ex Eriogonum wrightii var. subscaposum (verbatim: Eriogonum subscaposum); UCRC • 1 ♀; San Diego Co., 15 mi. SE of Borrego; [33.052 ° N, 115.851 ° W]; 5 Feb. 1952; J. G. Rozen leg.; EMEC • 1 ♀; San Diego Co., 20 mi. SE of Borrego; [33.066 ° N, 116.058 ° W]; 4 Feb. 1952; J. G. Rozen leg.; EMEC • 2 ♀♀; San Diego Co., 6 mi. SW of Borrego; [33.175 ° N, 116.33 ° W]; 4 Feb. 1952; J. G. Rozen leg.; EMEC • 1 ♀; San Diego Co., Borrego; [33.223 ° N, 116.334 ° W]; 6 Feb. 1952; J. G. Rozen leg.; EMEC • 1 ♂; same data as for preceding; 31 Mar. 1953; P. D. Hurd leg.; ex Cryptantha barbigera; EMEC • 1 ♀; same data as for preceding; 31 Mar. 1952; P. D. Hurd leg.; ex Phacelia; EMEC • 1 ♀; San Diego Co., Borrego; [33.247 ° N, 116.372 ° W]; 27 Feb. 1950; G. H. Sperry and J. L. Sperry leg.; UCRC • 5 ♀♀; San Diego Co., Borrego Palm Canyon; [33.272 ° N, 116.407 ° W]; 31 Jan. 1965; G. R. Ballmer leg.; UCRC • 1 ♀; San Diego Co., Borrego Palm Canyon; [33.2881 ° N, 116.4472 ° W]; elev. 595 m; 29 Mar. 1936; P. H. Timberlake leg.; ex Larrea tridentata var. tridentata (verbatim: Larrea divaricata); UCRC • 1 ♀; San Diego Co., Borrego-Clark Lake, N End; [33.331 ° N, 116.273 ° W]; 23 Mar. 1978; Wasbauer, Slansky Adams leg.; CSCA • 1 ♂; San Diego Co., Clark Dry Lake, Borrego State Park; [33.331 ° N, 116.273 ° W]; 31 Mar. 1977; M. Wasbauer, J. Slansky leg.; CSCA • 1 ♀; San Diego Co., Coronado Beach; [32.680 ° N, 117.178 ° W]; 3 Apr. 1918; J. I. Carlson leg.; CAS • 1 ♀; San Diego Co., Cuyamaca; [32.928 ° N, 116.568 ° W]; 19 Apr. 1950; E. G. Linsley leg.; ex Lasthenia (verbatim: Baeria); EMEC • 1 ♀; San Diego Co., McCain Valley; [32.672 ° N, 116.237 ° W]; 3 May 1980; Brown Faulkner leg.; SDMC • 1 ♂; same data as for preceding; 3 May 1980; Brown Faulkner leg.; SDMC • 3 ♀♀; San Diego Co., Descanso; [32.859 ° N, 116.614 ° W]; 30 Mar. 1961; P. D. Hurd leg.; ex Salix; EMEC • 1 ♂; San Diego Co., Mt Laguna; [32.867 ° N, 116.42 ° W]; 21 Jun. 1963; P. D. Hurd leg.; EMEC • 2 ♂♂; same data as for preceding; 5 Jul. 1963; P. D. Hurd leg.; ex Heliomeris multiflora (verbatim: Viguiera multiflora); EMEC • 1 ♂; San Diego Co., Sentenac Canyon; [33.129 ° N, 116.432 ° W]; 22 Mar. 1953; R. Mackie leg.; SDMC • 1 ♂; San Luis Obispo Co., La Panza Camp; [35.354 ° N, 120.263 ° W]; 25 Apr. 1968; J. Doyen leg.; ex Rhamnus crocea; EMEC • 1 ♂; Santa Barbara Co., Cachuma Campground, Los Padres National Forest; [34.575 ° N, 119.957 ° W]; 24 – 25 May 1976; C. George and R. R. Snelling leg.; ex Frangula californica (verbatim: Rhamnus californica); EMEC • 1 ♀; Santa Barbara Co., Cuyama; [34.931 ° N, 119.615 ° W]; 18 Mar. 1958; R. P. Allen leg.; EMEC • 1 ♂; Santa Clara Co., Mt Hamilton; [37.342 ° N, 121.643 ° W]; 25 May 1950; P. D. Hurd leg.; ex Eriodictyon; EMEC • 1 ♀; Santa Clara Co., San Antonio Valley; [40.493 ° N, 121.889 ° W]; 11 Mar. 1948; P. D. Hurd leg.; EMEC • 1 ♀; Santa Clara Co., Stanford University; [37.428 ° N, 122.17 ° W]; 28 Mar. 1916; EMEC • 2 ♀♀; Shasta Co., Hat Lake; [40.509 ° N, 121.466 ° W]; 14 Jun. 1941; P. D. Hurd leg.; ex Salix; EMEC • 1 ♀; Shasta Co., Shasta Springs; [41.252 ° N, 122.275 ° W]; 6 Jun. 1920; C. L. Fox leg.; CAS • 2 ♀♀; Shasta Co., Shingletown; [40.493 ° N, 121.889 ° W]; 21 Jun. 1941; EMEC • 2 ♀♀; Shasta Co., Viola; [40.518 ° N, 121.678 ° W]; 19 May 1941; C. D. Michener leg.; ex Salix; EMEC • 2 ♀♀; Sierra Co., Gold Lake; [39.672 ° N, 120.66 ° W]; 14 Jul. 1921; C. L. Fox leg.; CAS • 1 ♀; same data as for preceding; 21 Jul. 1921; C. L. Fox leg.; CAS • 2 ♀♀; Stanislaus Co., Adobe Creek W; [37.396 ° N, 121.375 ° W]; 7 – 9 Mar. 1956; Don Burdick leg.; ex Salix; EMEC • 1 ♀; Stanislaus Co., Del Puerto Canyon; [37.477 ° N, 121.23 ° W]; 11 May 1948; Ray F. Smith leg.; ex Eriodictyon; EMEC • 1 ♀; same data as for preceding; 2 Apr. 1961; R. R. Snelling and M. D. Snelling leg.; EMEC • 1 ♀; Stanislaus Co., Del Puerto Canyon, 21 mi. W of Patterson; [37.470 ° N, 121.511 ° W]; 12 Mar. 1966; R. Langston leg.; ex Salix; EMEC • 1 ♂; Stanislaus Co., Del Puerto Canyon, W of Patterson; [37.488 ° N, 121.207 ° W]; 19 May 1950; T. F. Leigh leg.; EMEC • 3 ♀♀; Stanislaus Co., Westley; [37.548 ° N, 121.201 ° W]; 1 Apr. 1948; P. D. Hurd leg.; ex Brassica; EMEC • 1 ♀; Trinity Co., 5 mi. SE of Peanut; [40.307 ° N, 123.278 ° W]; 21 May 1973; J. Doyen leg.; EMEC • 3 ♀♀; Tulare Co., 2 mi. E of Johnsondale; [35.970 ° N, 118.498 ° W]; 27 Apr. 1964; C. A. Toschi leg.; ex Prunus subcordata; EMEC • 1 ♀; same data as for preceding; 27 Apr. 1964; J. Powell leg.; ex Arctostaphylos; EMEC • 1 ♀; Tuolumne Co., Chinese Camp; [37.870 ° N, 120.433 ° W]; 2 Apr. 1960; R. R. Snelling leg.; ex Ranunculus; EMEC • 2 ♀♀; Ventura Co., Chuchupate Ranger Station, base, Frazier Mountain; [34.808 ° N, 119.012 ° W]; 5 May 1959; P. D. Hurd leg.; ex Descurainia sophia; EMEC • 1 ♂; same data as for preceding; 5 May 1959; P. D. Hurd leg.; ex Descurainia sophia; EMEC • 1 ♂; same data as for preceding; 7 May 1959; J. Powell leg.; ex Descurainia sophia; EMEC • 1 ♀; Ventura Co., Lockwood Creek; [34.7331 ° N, 119.0958 ° W]; elev. 1555 m; 5 Mar. 1959; P. H. Timberlake leg.; ex Sisymbrium; UCRC • 1 ♀; Ventura Co., Lockwood Creek, near Stauffer P. O.; [34.730 ° N, 119.033 ° W]; 5 – 7 May 1959; P. D. Hurd leg.; ex Descurainia sophia; EMEC • 1 ♂; same data as for preceding; 5 – 7 May 1959; P. D. Hurd leg.; ex Descurainia sophia; EMEC • 1 ♂; Ventura Co., Lockwood Creek, near Stauffer P. O.; [34.730 ° N, 119.033 ° W]; 5 May 1959; G. I. Stage leg.; ex Descurainia sophia; EMEC • 2 ♂♂; same data as for preceding; 7 May 1959; P. D. Hurd leg.; ex Descurainia sophia; EMEC • 1 ♂; same data as for preceding; 5 May 1959; G. I. Stage leg.; ex Descurainia sophia; EMEC • 1 ♀; Ventura Co., Lockwood Valley, near Stauffer P. O.; [34.733 ° N, 119.04 ° W]; 3 May 1959; J. Powell leg.; ex Phacelia ciliata; EMEC • 1 ♀; Yolo Co., Putah Creek Canyon; [38.512 ° N, 122.048 ° W]; 19 Apr. 1970; D. P. Levin leg.; EMEC • 1 ♂; Yolo Co., Rumsey; [38.889 ° N, 122.239 ° W]; 1 May 1959; Menke and Stange leg.; EMEC. – Colorado • 1 ♀; Boulder Co., 2 mi. N of Boulder; [40.044 ° N, 105.27 ° W]; elev. 1676 m; 9 May 1980; U. N. Lanham leg.; ex Physaria belli; UCMC • 1 ♀; Boulder Co., 3 mi. E of Jamestown; [40.115 ° N, 105.331 ° W]; 31 May 1980; K. Neff and U. Lanham leg.; ex Physaria; UCMC • 2 ♀♀; Boulder Co., 4 km N of Lyons; 40.24847 ° N, 105.29742 ° W; elev. 1705 m; 19 May 2007; C. Sheffield and J. Gibbs leg.; RSKM • 2 ♀♀; Boulder Co., 4 mi. N of Boulder; [40.073 ° N, 105.271 ° W]; elev. 1676 m; 21 May 1980; U. N. Lanham and P. Robinson leg.; ex Physaria belli; UCMC • 1 ♀; Boulder Co., 5 mi. N of Boulder; [40.089 ° N, 105.27 ° W]; 9 May 1939; Rodeck and Lanham leg.; UCMC • 2 ♀♀; Boulder Co., 5 mi. NW of Boulder; [40.066 ° N, 105.337 ° W]; 31 May 1963; U. N. Lanham and B. Vogel leg.; ex Prunus virginiana; UCMC • 1 ♀; Boulder Co., 6 mi. NW of Boulder; [40.077 ° N, 105.351 ° W]; elev. 1829 m; 22 May 1981; U. N. Lanham leg.; ex Physaria vitulifera; UCMC • 2 ♀♀; Boulder Co., 9 mi. NW of Longmont Foothills; [40.259 ° N, 105.222 ° W]; 22 Jun. 1961; U. N. Lanham leg.; ex Amorpha fruticosa; UCMC • 1 ♀; Boulder Co., Boulder; [40.02 ° N, 105.27 ° W]; 21 May 1922; E. M. Foster leg.; UCRC • 1 ♀; same data as for preceding; 18 May 1913; M. D. Ellis leg.; UCRC • 1 ♀; same data as for preceding; 23 May 1913; M. D. Ellis leg.; ex Erysimum capitatum; UCRC • 1 ♀; same data as for preceding; 22 May 1913; M. M. Ellis leg.; ex Astragalus agrestis; UCRC • 1 ♀; same data as for preceding; 22 May 1913; M. M. Ellis leg.; UCRC • 1 ♀; same data as for preceding; 20 May 1922; C. Walton leg.; UCRC • 2 ♀♀; same data as for preceding; 14 May 1939; R. Bauer leg.; UCMC • 1 ♀; same data as for preceding; 14 May 1939; U. Lanham leg.; UCMC • 1 ♀; same data as for preceding; Spring – 1931; P. Lundy leg.; UCMC • 2 ♀♀; same data as for preceding; 14 Jul. 1961; Swenson and Lanham leg.; UCMC • 1 ♀; same data as for preceding; 20 Apr. 1965; U. Lanham leg.; ex Salix; UCMC • 1 ♀; same data as for preceding; 25 Apr. 1968; U. N. Lanham leg.; ex Salix; UCMC • 1 ♀; same data as for preceding; 15 May 1908; S. A. Rohwer leg.; ex Salix; MCZ • 1 ♀; same data as for preceding; 21 May; T. D. A. Cockerell leg.; ex Rubus deliciosus; MCZ • 1 ♀; same data as for preceding; 17 May 1922; D. W. Macomber leg.; CAS • 1 ♀; same data as for preceding; 29 Apr. 1906; S. A. Rohwer leg.; CAS • 1 ♀; Boulder Co., Boulder Canyon; [40.001 ° N, 105.386 ° W]; 22 May 1962; R. W. Thorp leg.; EMEC • 1 ♀; Boulder Co., near Altona; [40.148 ° N, 105.293 ° W]; 30 May 1975; U. Lanham leg.; ex Salix; UCMC • 1 ♀; Chaffee Co., Poncha Pass; [38.534 ° N, 106.171 ° W]; 20 Jun. 1933; C. A. Wagner leg.; CAS • 1 ♀; same data as for preceding; 20 Jun. 1933; W. Steele leg.; CAS • 1 ♀; Costilla Co., San Luis; [37.201 ° N, 105.424 ° W]; 26 Apr. 1955; H. G. Rodeck leg.; UCMC • 5 ♀♀; Delta Co., 4 mi. Escalante Canyon; [38.728 ° N, 108.14 ° W]; 21 May 1963; C. J. McCoy and B. Vogel leg.; UCMC • 2 ♀♀; Douglas Co., 10 mi. N of Castlerock; [39.517 ° N, 104.856 ° W]; 26 Apr. 1963; U. N. Lanham leg.; ex Salix; UCMC • 6 ♀♀; Douglas Co., 10 mi. SSE of Franktown; [39.319 ° N, 104.977 ° W]; 26 Apr. 1963; U. N. Lanham leg.; ex Salix; UCMC (Fig. 46) • 10 ♀♀; Douglas Co., 5 mi. SSW of Franktown; [39.324 ° N, 104.789 ° W]; 26 Apr. 1963; U. N. Lanham leg.; ex Salix; UCMC • 6 ♀♀; same data as for preceding; 12 Apr. 1963; U. N. Lanham et al. leg.; ex Salix; UCMC • 1 ♀; Eagle Co., 2 mi. N of Dotsero; [39.673 ° N, 107.067 ° W]; 27 Jun. 1963; B. Vogel leg.; ex Tamarix; UCMC • 1 ♀; El Paso Co., Solar Trails County Park; [38.7094 ° N, 104.6617 ° W]; elev. 1980 m; 26 May 1980; CUIC • 1 ♀; El Paso Co., Foster Ranch; [38.6391 ° N, 104.5515 ° W]; elev. 1740 m; 22 May 1977; F. M. Brown leg.; ex Cardaria; CUIC • 1 ♀; El Paso Co., Manitou; [38.857 ° N, 104.917 ° W]; 20 Jun. 1926; M. C. Van Duzee leg.; CAS • 2 ♀♀; Elbert Co., Elbert; [39.219 ° N, 104.537 ° W]; 21 May; W. P. Cockerell leg.; ex Salix; UCRC • 1 ♀; Elbert Co., Running Creek Field Station; [39.3319 ° N, 104.4852 ° W]; elev. 2120 m; 25 May 1977; Susan Condie leg.; ex Salix; CUIC • 1 ♀; Elbert Co., Running Creek Field Station; [39.3306 ° N, 104.4858 ° W]; elev. 2120 m; 27 May 1976; F. M. Brown leg.; ex Prunus; CUIC • 1 ♀; Garfield Co., 4 mi. N of Rifle; [39.592 ° N, 107.783 ° W]; 24 Apr. 1982; P. Robinson leg.; UCMC • 1 ♀; Huerfano Co., 3 mi. W of Walsenburg; [37.618 ° N, 104.836 ° W]; 30 Apr. 1981; U. N. Lanham leg.; ex Salix; UCMC • 1 ♀; Huerfano Co., Walsenburg; [37.622 ° N, 104.781 ° W]; 1 May 1981; U. N. Lanham leg.; ex Cardaria; UCMC • 1 ♀; Jefferson Co., 1 mi. S of Eldorado Springs; [39.917 ° N, 105.284 ° W]; 22 Apr. 1985; U. N. Lanham leg.; ex Salix; UCMC • 1 ♀; same data as for preceding; 29 Apr. 1965; U. N. Lanham leg.; ex Salix; UCMC • 1 ♀; Kit Carson Co., 12 mi. N of Burlington; [39.478 ° N, 102.27 ° W]; 9 May 1977; U. N. Lanham, P. Byron leg.; ex Salix; UCMC • 3 ♀♀; Larimer Co., 10 mi. WSW of Loveland; [40.339 ° N, 105.249 ° W]; elev. 1981 m; 15 May 1963; U. N. Lanham leg.; ex Salix; UCMC • 10 ♀♀; Larimer Co., 13 mi. WSW of Loveland; [40.321 ° N, 105.302 ° W]; elev. 1981 m; 30 Apr. 1963; U. N. Lanham leg.; ex Salix; UCMC • 17 ♀♀; Larimer Co., 14 mi. W of Fort Collins; [40.585 ° N, 105.351 ° W]; elev. 2012 m; 6 May 1963; U. N. Lanham leg.; ex Salix; UCMC • 1 ♀; Larimer Co., 16 mi. N of Fort Collins; [40.817 ° N, 105.084 ° W]; 10 May 1963; U. Lanham leg.; ex Salix; UCMC • 1 ♀; Larimer Co., 16 mi. WNW of Fort Collins; [40.671 ° N, 105.367 ° W]; elev. 2134 m; 9 May 1963; U. N. Lanham leg.; UCMC • 1 ♀; Larimer Co., 5 mi. N of Masonville; [40.560 ° N, 105.211 ° W]; 10 May 1963; U. Lanham leg.; ex Prunus virginiana; UCMC • 1 ♀; same data as for preceding; 10 May 1963; U. Lanham leg.; ex Rhus trilobata; UCMC • 6 ♀♀; Larimer Co., 7 mi. W of Berthoud; [40.304 ° N, 105.214 ° W]; elev. 1676 m; 15 May 1963; U. N. Lanham leg.; ex Salix; UCMC • 1 ♀; same data as for preceding; 15 May 1963; U. N. Lanham leg.; ex Rhus trilobata; UCMC • 1 ♀; Larimer Co., 8 mi. WSW of Loveland; [40.352 ° N, 105.209 ° W]; elev. 1676 m; 30 Apr. 1963; U. N. Lanham leg.; ex Prunus; UCMC • 1 ♂; Mesa Co., 4 mi. W of Mesa; [39.142 ° N, 108.194 ° W]; 18 Jun. 1938; R. Bauer leg.; UCMC • 6 ♀♀; Mesa Co., Colorado National Monument; [39.051 ° N, 108.692 ° W]; elev. 1753 m; 15 May 1963; C. J. McCoy leg.; ex Lepidium montanum; UCMC • 6 ♀♀; same data as for preceding; 8 May 1963; C. J. McCoy leg.; ex Lepidium montanum; UCMC • 2 ♀♀; same data as for preceding; 12 May 1963; C. J. McCoy leg.; ex Lepidium montanum; UCMC • 2 ♀♀; same data as for preceding; 16 May 1985; U. Lanham and M. Weissmann leg.; ex Lepidium montanum; UCMC • 1 ♀; Mesa Co., Gateway; [38.680 ° N, 108.977 ° W]; elev. 1676 m; 3 May 1978; U. Lanham leg.; ex Lepidium montanum; UCMC • 2 ♀♀; Mesa Co., Whitewater; [38.991 ° N, 108.453 ° W]; 18 May 1963; B. Vogel and C. J. McCoy leg.; ex Astragalus; UCMC • 1 ♀; Mesa Co.; 39.0295 ° N, 108.6276 ° W; 24 – 25 Aug. 2011; BIML • 1 ♀; Moffat Co., 23 mi. SW of Maybell; [40.285 ° N, 108.404 ° W]; elev. 1829 m; 26 May 1974; U. N. Lanham leg.; UCMC • 1 ♂; Montezuma Co., Mesa Verde; [37.183 ° N, 108.5 ° W]; 3 Jul. 1907; EMEC • 1 ♀; Montrose Co., 17 mi. SSW of Naturita; [38.101 ° N, 108.673 ° W]; elev. 1981 m; 29 May 1977; U. Lanham and P. Byron leg.; ex Carderia; UCMC • 1 ♀; Pueblo Co., Colorado City; [37.945 ° N, 104.836 ° W]; 2 May 1977; U. Lanham and P. Byron leg.; ex Prunus; UCMC • 1 ♀; same data as for preceding; 20 May 1975; U. Lanham leg.; ex Rhus trilobata; UCMC • 1 ♀; same data as for preceding; 20 May 1975; U. Lanham leg.; ex Prunus virginiana var. demissa (verbatim: Prunus melanocarpus); UCMC • 1 ♀; same data as for preceding; 20 May 1975; U. N. Lanham leg.; ex Rhus trilobata; UCMC • 1 ♀; same data as for preceding; 20 May 1975; U. N. Lanham leg.; ex Cercocarpus montanus; UCMC • 1 ♂; Routt Co., 4 mi. E of Steamboat Springs; [40.485 ° N, 106.75 ° W]; 28 Aug. 1979; U. N. Lanham leg.; UCMC • 1 ♂; Routt Co., 8 mi. NW of Steamboat Springs; [40.600 ° N, 106.95 ° W]; 28 Aug. 1979; U. N. Lanham leg.; ex Solidago; UCMC • 1 ♀; Routt Co., SE of Steamboat Springs; [40.485 ° N, 106.832 ° W]; 18 Jun. 1978; U. Lanham leg.; ex Ceanothus; UCMC • 2 ♂♂; Yuma Co., Beecher Island; [39.870 ° N, 102.185 ° W]; 21 Jun. 1979; U. N. Lanham leg.; ex Melilotus officinalis; UCMC. – Idaho • 1 ♀; Canyon Co., Deer Flat National Wildlife Refuge, DFS; 43.55281 ° N, 116.64137 ° W; 4 – 5 Oct. 2012; Addison Mohler leg.; OSAC • 2 ♀♀; Gooding Co., Hagerman Valley; [42.817 ° N, 114.898 ° W]; 16 May 1929; ex Solidago altissima; USNM • 1 ♀; Gooding Co., Wood River, 1 mi. NE of Goodling; [42.949 ° N, 114.698 ° W]; 6 – 7 Jul. 1980; M. S. Wasbauer and K. M. Wasbauer leg.; CSCA • 2 ♂♂; same data as for preceding; 6 – 7 Jul. 1980; M. S. Wasbauer and K. M. Wasbauer leg.; CSCA • 1 ♀; Latah Co., Moscow; [46.732 ° N, 117.001 ° W]; 4 Nov. 1966; UCRC • 4 ♂♂; Nez Perce Co., Top of Lewiston Grade; [46.462 ° N, 117.02 ° W]; 30 Jun. 1975; J. Powell leg.; EMEC • 1 ♂; Oneida Co., Rock Creek; [42.188 ° N, 112.655 ° W]; 21 Jul. 1971; G. F. Knowlton leg.; CUIC • 1 ♂; Oneida Co., Rock Creek; [42.188 ° N, 112.655 ° W]; 14 Jul. 1971; G. F. vKnowlton leg.; CUIC • 1 ♀; Oneida Co., 2 mi. N of Stone; [42.064 ° N, 112.895 ° W]; 16 May 1969; G. F. Knowlton and G. E. Bohart leg.; CUIC • 1 ♂; Owyhee Co., 6 mi. E of Silver City; [43.016 ° N, 116.614 ° W]; 20 Jul. 1965; R. L. Langston leg.; ex Clarkia pulchella; EMEC • 1 ♂; Owyhee Co., Jump Creek; [43.479 ° N, 116.926 ° W]; 23 Jun. 1958; P. H. Raven leg.; ex Chylismia claviformis subsp. integrior (verbatim: Oenothera claviformis subsp. purparescens); EMEC • 1 ♀; Twin Falls Co., Castleford; [42.521 ° N, 114.873 ° W]; 28 Jun. 1928; ex Solidago altissima; UCRC. – Montana • 2 ♀♀; Big Horn Co., Custer Battlefield; [45.57 ° N, 107.43 ° W]; 4 Jun. 1941; G. E. Ferguson leg.; OSAC • 4 ♂♂; Carbon Co., East Rosebud Canyon; [45.484 ° N, 109.457 ° W]; elev. 1800 m; 10 Jul. 1963; B. Vogel leg.; UCMC • 1 ♀; Richland Co., Elk Island, 3 _ Canopy; 47.46127 ° N, 104.31522 ° W; 29 Jun. – 1 Jul. 2021; A. R. Morphew leg.; WRME • 2 ♀♀; Missoula Co., Florence MPG Ranch; 46.6850 ° N, 113.9861 ° W; elev. 1291 m; 4 – 6 Jun. 2019; M. Seidensticker leg.; CBG • 1 ♀; Missoula Co., Florence MPG Ranch; 46.6888 ° N, 113.9922 ° W; elev. 1422 m; 11 – 13 Jun. 2019; M. Seidensticker leg.; CBG • 1 ♀; Missoula Co., Florence MPG Ranch; 46.6935 ° N, 114.0086 ° W; elev. 1198 m; 10 – 12 Jun. 2019; M. Seidensticker leg.; CBG • 1 ♀; Missoula Co., Florence MPG Ranch; 46.6901 ° N, 113.9847 ° W; elev. 1380 m; 4 – 6 Jun. 2019; M. Seidensticker leg.; CBG • 1 ♀; Missoula Co., Florence MPG Ranch; 46.7144 ° N, 113.995 ° W; elev. 1535 m; 20 – 22 May 2019; M. Seidensticker leg.; CBG • 1 ♀; Missoula Co., Florence MPG Ranch; 46.6901 ° N, 113.9847 ° W; elev. 1380 m; 11 – 18 Jun. 2019; M. Seidensticker leg.; CBG • 1 ♀; Custer Co., Ft. Keogh, RO Restoration Block 8; 46.40592 ° N, 105.87979 ° W; 18 – 20 May 2021; J. Muscha and A. Pearson leg.; WRME • 1 ♀; Richland Co., Seven Sisters, 2 _ Ground; 47.57646 ° N, 104.22487 ° W; 15 – 17 Jun. 2021; A. R. Morphew leg.; WRME. – Nevada • 1 ♀; Churchill Co., 25 mi. SE of Fallon; [39.300 ° N, 118.62 ° W]; 25 Apr. 1980; R. W. Rust leg.; ex Astragalus; CUIC • 1 ♀; Clark Co., Kyle Canyon, Mt Charleston; [36.2714 ° N, 115.5781 ° W]; elev. 1981 m; 3 Jun. 1941; P. H. Timberlake leg.; UCRC • 2 ♂♂; Carson City; [39.16 ° N, 119.78 ° W]; Jul.; Baker leg.; UCRC • 1 ♂; same data as for preceding; Jul.; Baker leg.; USNM • 1 ♂; same data as for preceding; 25 Jun. 1929; E. P. Van Duzee leg.; CAS • 1 ♀; Kings Canyon Ranch, Carson City; [39.161 ° N, 119.799 ° W]; elev. 1524 m; 23 Jun. 1964; D. Bolinger leg.; OSAC • 1 ♀; Clark Co., Kyle Canyon, Mt Charleston; [36.2725 ° N, 115.5275 ° W]; elev. 1707 m; 4 Jun. 1941; P. H. Timberlake leg.; UCRC • 1 ♀; Clark Co., Kyle Canyon, Mt Charleston; [36.2703 ° N, 115.5519 ° W]; elev. 1829 m; 5 Jun. 1941; P. H. Timberlake leg.; UCRC • 1 ♂; same data as for preceding; 5 Jun. 1941; P. H. Timberlake leg.; UCRC • 1 ♂; Clark Co., Lee Canyon, Charleston Mts; [36.311 ° N, 115.671 ° W]; elev. 1829 m; 3 Jun. 1941; P. H. Timberlake leg.; UCRC • 2 ♂♂; Douglas Co., Topaz Lake; [38.696 ° N, 119.519 ° W]; 11 Jul. 1967; W. J. Turner leg.; EMEC • 1 ♀; Elko Co., Elko; [40.832 ° N, 115.763 ° W]; 19 May 1926; M. C. Van Duzee leg.; CAS • 4 ♂♂; Lander Co., Kingston Canyon; [39.225 ° N, 117.139 ° W]; elev. 2134 m; 22 Jul. 1968; J. Powell leg.; EMEC • 1 ♀; Lincoln Co., 8 mi. E of Panaca; [37.787 ° N, 114.241 ° W]; 22 Jun. 1982; W. J. Pulawski leg.; CAS • 1 ♂; Storey Co., Ophir Grade, 2 mi. SW of Virginia City; [39.299 ° N, 119.66 ° W]; elev. 1950 m; 20 Jul. 1964; R. L. Langston leg.; EMEC • 3 ♂♂; Storey Co., Virginia City; [39.309 ° N, 119.65 ° W]; 5 Jul. 1964; C. N. Slobodchikoff leg.; CAS • 4 ♀♀; Washoe Co., 21 mi. SE of Eagleville, California; [41.093 ° N, 119.723 ° W]; 11 Jun. 1970; P. A. Rude leg.; ex Lepidium appelianum (verbatim: Cardaria pubescens); EMEC • 2 ♂♂; Washoe Co., Patrick; [39.547 ° N, 119.579 ° W]; 30 Jun. 1964; M. E. Irwin leg.; UCRC • 1 ♂; Washoe Co., Sparks; [39.535 ° N, 119.753 ° W]; 23 Jun. 1927; E. P. Van Duzee leg.; CAS. – New Mexico • 1 ♂; Bernalillo Co., 1 mi. W of Chilili; [34.871 ° N, 106.285 ° W]; 6 Jul. 1965; R. C. Dickson leg.; UCRC • 1 ♀; Grant Co., Cherry Creek Campground, 14 mi. N of Silver City; [32.914 ° N, 108.225 ° W]; elev. 2255 m; 15 – 16 Aug. 1993; J. E. O’Hara leg.; RSKM • 1 ♂; Grant Co., Cherry Creek Campground, 14 mi. N of Silver City; [32.914 ° N, 108.225 ° W]; elev. 2255 m; 15 – 16 Aug. 1993; J. E. O’Hara leg.; RSKM • 3 ♀♀; Socorro Co., 10 mi. W of Socorro; [34.044 ° N, 107.068 ° W]; 28 Mar. 1963; U. N. Lanham leg.; UCMC. – Oregon • 1 ♂; Baker Co., 4 mi. NW of Haines; [44.954 ° N, 117.998 ° W]; 8 May 1955; W. P. Stephen leg.; OSAC • 1 ♀; Baker Co., Oregon Trail Interpretive Center; 44.8118 ° N, 117.7265 ° W; 20 Jun. – 5 Jul. 2011; J. Rebholz leg.; OSAC • 1 ♂; Crook Co., 11 mi. SW of Prineville; [44.187 ° N, 120.99 ° W]; 16 Jul. 1965; R. L. Langston leg.; EMEC • 2 ♂♂; Crook Co., 25 mi. E of Prineville; [44.300 ° N, 120.666 ° W]; elev. 1097 m; 13 Jul. 1936; H. A. Scullen leg.; OSAC • 2 ♀♀; Deschutes Co., Redmond; [44.273 ° N, 121.174 ° W]; 11 May 1939; Schuh and Gray leg.; OSAC • 1 ♂; Grant Co., Fall Mountain Lookout Trail; [44.294 ° N, 119.043 ° W]; elev. 1585 – 1829 m; 14 Jul. 1936; H. A. Scullen leg.; OSAC • 1 ♀; Harney Co., 17 mi. N of French – Glen; [43.052 ° N, 118.918 ° W]; 19 May 1956; J. D. Lattin leg.; OSAC • 1 ♀; Harney Co., Blitzen Valley; [43.272 ° N, 118.853 ° W]; 15 May 1936; S. G. Jewett leg.; OSAC • 1 ♀; Harney Co., Blitzen Valley, Buena Vista Ranch; [43.272 ° N, 118.853 ° W]; 19 Apr. 1936; S. G. Jewett leg.; OSAC • 3 ♀♀; Harney Co., Buena Vista Ranch, Blitzen Valley; [43.272 ° N, 118.853 ° W]; 19 Apr. 1936; S. G. Jewett leg.; OSAC • 1 ♀; Harney Co., Malheur National Wildlife Refuge, BV; 43.06803 ° N, 118.8681 ° W; 18 – 19 May 2012; Marilyn Kircus leg.; OSAC • 1 ♀; Harney Co., Malheur National Wildlife Refuge, PSD; 42.80817 ° N, 118.87006 ° W; 17 – 18 Jun. 2012; Marilyn Kircus leg.; OSAC • 1 ♀; Harney Co., Malheur National Wildlife Refuge, SBV; 42.8188 ° N, 118.8688 ° W; 17 Apr. 2012; J. D. Engler leg.; OSAC • 1 ♀; Hood River Co., Hood River; [45.705 ° N, 121.522 ° W]; 8 Jul. 1927; E. C. VanDyke leg.; CAS • 1 ♀; Klamath Co., 10 mi. E of Bly; [42.397 ° N, 120.847 ° W]; 12 Jul. 1940; H. A. Scullen leg.; ex Eriogonum; OSAC • 2 ♀♀; Klamath Co., Klamath Falls; [42.225 ° N, 121.782 ° W]; 10 May 1924; C. L. Fox leg.; CAS • 1 ♀; same data as for preceding; 8 May 1924; C. L. Fox leg.; CAS • 2 ♀♀; same data as for preceding; 9 May 1924; C. L. Fox leg.; CAS • 1 ♀; Klamath Co., Merrill; [42.026 ° N, 121.599 ° W]; 10 May 1924; C. C. Wilson leg.; ex Mutarda arvensis (verbatim: Sinapis arvensis); UCRC • 1 ♀; Lake Co., Hart Mountain; [42.549 ° N, 119.642 ° W]; 17 Jun. 1938; K. Gray and J. Schuh leg.; OSAC • 3 ♀♀; Lake Co., Hart Mountain Antelope Refuge; [42.549 ° N, 119.642 ° W]; 24 Apr. 1954; Otto C. Nelson leg.; ex Ribes; OSAC • 1 ♀; same data as for preceding; 4 May 1954; Otto C. Nelson leg.; ex Taraxacum; OSAC • 1 ♀; Lake Co., Picture Rock Pass; [43.048 ° N, 120.799 ° W]; 19 Apr. 1950; Black and Davis leg.; OSAC • 1 ♀; Lake Co., Summer Lake; [42.973 ° N, 120.778 ° W]; 16 Jun. 1938; Gray and Schuh leg.; OSAC • 1 ♂; Malheur Co., Nyssa; [43.877 ° N, 116.995 ° W]; elev. 665 m; 16 Jun. 1926; R. W. Haegele leg.; UCRC • 1 ♂; Malheur Co., Ranaway Gulch; [42.4083 ° N, 117.8225 ° W]; elev. 1430 m; 14 Jul. 1984; William H. Clark and Donald R. Frohlich leg.; CUIC • 1 ♀; Malheur Co., Runaway Gulch; [42.4083 ° N, 117.8225 ° W]; elev. 1430 m; 23 Apr. 1983; Cynthia J. Clark, William H. Clark and Donald R. Frohlich leg.; CUIC • 1 ♂; Malheur Co., Sucker Creek Canyon; [42.550 ° N, 117.3 ° W]; 15 – 18 Jun. 1951; Borys Malkin leg.; CAS • 1 ♂; Malheur Co., Vale; [43.982 ° N, 117.239 ° W]; 29 Jun. 1902; Wicklund leg.; OSAC • 3 ♀♀; Umatilla Co., Cayuse; [45.676 ° N, 118.574 ° W]; 23 Apr. 1939; K. Gray and J. Schuh leg.; OSAC • 17 ♂♂; Umatilla Co., Milton; [45.934 ° N, 118.391 ° W]; 22 Jun. 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♀; same data as for preceding; 22 Jun. 1938; K. Gray leg.; EMEC • 1 ♀; Umatilla Co., Thorn Hollow; [45.689 ° N, 118.462 ° W]; 29 Apr. 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♂; Umatilla Co., Ukiah – Dale Forest Wayside, Camas Creek; [45.126 ° N, 118.971 ° W]; 24 Jul. 1974; Paul H. Arnaud Jr leg.; CAS • 21 ♀♀; Umatilla Co., Weston; [45.813 ° N, 118.425 ° W]; 28 Apr. 1939; K. Gray and J. Schuh leg.; OSAC • 22 ♀♀; same data as for preceding; 29 Apr. 1939; K. Gray leg.; ex Brassica; OSAC • 2 ♀♀; Umatilla Co., Wild Horses Mountain, Athena; [45.802 ° N, 118.481 ° W]; 14 May 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♀; Umatilla, Union, Grant, or Baker Co., Blue Mountain; [45.000 ° N, 118.25 ° W]; 25 Aug. 1948; J. E. Davis leg.; OSAC • 1 ♂; Union Co., La Grande; [45.323 ° N, 118.088 ° W]; 20 Jul. 1929; H. A. Scullen leg.; ex Convolvulus; OSAC • 1 ♀; same data as for preceding; 28 May 1943; G. R. Ferguson leg.; OSAC • 1 ♂; Wallowa Co., 5 mi. E of Minam, Wallowa Canyon Fount; [45.571 ° N, 117.631 ° W]; elev. 823 m; 21 Jul. 1929; H. A. Scullen leg.; OSAC • 1 ♂; Wallowa Co., Chief Joseph, Mt Joseph; [45.285 ° N, 117.262 ° W]; 12 Aug. 1950; G. H. Sperry and J. L. Sperry leg.; UCRC • 1 ♀; Wallowa Co., Joseph; [45.354 ° N, 117.23 ° W]; 29 Jul. 1941; R. E. Rieder leg.; ex Melilotus; OSAC • 2 ♂♂; Wasco Co., 10 mi. S of The Dalles; [45.459 ° N, 121.178 ° W]; 15 Jun. 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♀; same data as for preceding; 5 May 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♀; Wasco Co., 12 mi. S of Dufur; [45.316 ° N, 121.128 ° W]; 5 May 1938; K. Gray and J. Schuh leg.; OSAC • 3 ♀♀; Wasco Co., 5 mi. S of The Dalles; [45.519 ° N, 121.178 ° W]; 5 May 1938; K. Gray and J. Schuh leg.; OSAC • 9 ♀♀; Wasco Co., Mosier; [45.683 ° N, 121.398 ° W]; 24 May 1938; K. Gray and J. Schuh leg.; OSAC • 4 ♀♀; Wasco Co., The Dalles; [45.600 ° N, 121.187 ° W]; 24 May 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♀; Wheeler Co., 17 mi. E of Mitchell; [44.564 ° N, 119.807 ° W]; 30 Jun. 1967; S. Paul Welles leg.; ex Clarkia pulchella; EMEC. – Utah • 1 ♀; Box Elder Co., Curlew Junction; [41.987 ° N, 112.684 ° W]; 16 May 1969; G. F. Knowlton and G. E. Bohart leg.; CUIC • 1 ♀; Grand Co., 15 mi. NW of Cisco; [39.121 ° N, 109.529 ° W]; 22 May 1979; D. B. Wahi leg.; SEMC • 1 ♀; Grand Co., 8 mi. SE of Emery; [38.843 ° N, 111.142 ° W]; 22 May 1979; D. B. Wahi leg.; SEMC • 2 ♀♀; Salt Lake Co., Camp Douglas; [40.765 ° N, 111.841 ° W]; CMNH • 1 ♀; Tooele Co., Grantsville; [40.599 ° N, 112.465 ° W]; 19 May 1919; G. F. Knowlton leg.; CAS • 1 ♀; San Juan Co., Kigalia Ranger Station, Manti – La Sal National Forest; [37.675 ° N, 109.826 ° W]; 5 Jul. 1935; Brues leg.; MCZ • 1 ♀; Cache Co., Logan River; [41.733 ° N, 111.763 ° W]; 24 May 1954; G. E. Bohart leg.; ex Salix; SEMC • 3 ♀♀; Cache Co., Wellsville; [41.638 ° N, 111.933 ° W]; 13 May 1954; G. E. Bohart leg.; ex Lomatium dissectum; SEMC. – Washington • 1 ♀; Asotin Co., Asotin Creek; [46.332 ° N, 117.077 ° W]; 27 Apr. 1924; A. L. Melander leg.; UCRC • 8 ♀♀; Asotin Co., Clarkston; [46.416 ° N, 117.045 ° W]; CMNH • 45 ♂♂; Asotin Co., Fields Spring State Park, S of Antone; [46.087 ° N, 117.172 ° W]; 28 Jul. 1985; Finnamore and Thormin leg.; PMAE (Figs 41 E, 48) • 1 ♂; same data as for preceding; 28 Jul. 1985; Finnamore and Thormin leg.; EMEC • 1 ♀; Chelan Co., N of Dryden; [47.533 ° N, 120.602 ° W]; 23 Apr. 1925; A. L. Melander leg.; UCRC • 1 ♀; Douglas Co., Pine Canyon; [47.636 ° N, 120.166 ° W]; 2 May 1937; OSAC • 3 ♂♂; Pend Oreille Co., Metaline Falls; [48.851 ° N, 117.39 ° W]; 1 Aug. 1923; C. L. Fox leg.; CAS • 1 ♀; Walla Walla Co., Prescott; [46.300 ° N, 118.315 ° W]; L. R. Dice leg.; EMEC • 4 ♂♂; Walla Walla Co., Walla Walla; [46.063 ° N, 118.343 ° W]; 5 – 10 Jun. 1936; R. M. and G. E. Bohart leg.; EMEC • 5 ♀♀; same data as for preceding; 29 Mar. – 2 Jun. 1937; R. M. and G. E. Bohart leg.; CAS • 1 ♂; same data as for preceding; 5 – 10 Jun. 1936; R. M. and G. E. Bohart leg.; CAS • 1 ♀; Walla Walla Co., Walla Walla; [46.0625 ° N, 118.343 ° W]; 22 Apr. 1925; M. C. Lane leg.; OSAC • 2 ♂♂; Whitman Co., Almota; [46.703 ° N, 117.469 ° W]; Melander leg.; UCRC • 3 ♂♂; same data as for preceding; 24 Jun.; Melander leg.; MCZ • 1 ♀; Whitman Co., Colfax; [46.880 ° N, 117.364 ° W]; 9 Jul. 1925; C. L. Fox leg.; CAS • 2 ♀♀; same data as for preceding; 8 Jul. 1925; C. L. Fox leg.; CAS • 1 ♂; same data as for preceding; 8 Jul. 1925; C. L. Fox leg.; CAS • 1 ♂; same data as for preceding; 9 Jul. 1925; C. L. Fox leg.; CAS • 1 ♀; Whitman Co., Wawawai; [46.638 ° N, 117.379 ° W]; 24 Apr. 1909; W. M. Mann leg.; EMEC • 2 ♀♀; Yakima Co., Satus Creek; [46.084 ° N, 120.554 ° W]; 28 Apr. 1938; K. Gray leg.; ex Purshia tridentata; OSAC • 2 ♀♀; Yakima Co., Selah; [46.652 ° N, 120.535 ° W]; 14 May 1919; OSAC. – Wyoming • 1 ♀; Lincoln Co.; 41.8650 ° N, 110.7653 ° W; 20 – 21 Sep. 2011; BIML.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFA4FFAFFDF6FB32FB0BFDF7.taxon	description	Redescription Female BODY MEASUREMENTS. Length 6.46 mm (± 0.45). Head long 1.67 mm (± 0.05 SD). Face length 1.39 mm (± 0.05 SD). Head width 1.61 mm (± 0.05 SD). Wing length 4.82 mm (± 0.19 SD). n = 5. COLOURATION. Head and mesosoma black. Clypeus apex black. Labrum black to dark brown. Mandible black to dark brown basally, becoming light brown medially and reddish brown apically. Flagellum dark brown to brown dorsally, yellowish brown ventrally. Pronotal lobe dark brown. Tegula translucent yellowish brown, with brown base. Wing membrane faintly brown, veins light brown. Legs brown. Metasoma black to dark brown with rims of terga and sterna broadly light brown. PUBESCENCE. Body hair colour off-white. Tomentum dense on pronotal angle, and lobe, space between pronotal lobe and tegula; very sparse or sometimes absent on T 2 – T 3 basolaterally. Mesosoma covered by erect plumose hairs (1 – 2.5 OD) and sub-appressed simple short hairs (<0.5 OD). Metafemoral scopa with dense plumose hairs. Wing hairs brown, very short and sparse. SURFACE SCULPTURING. Clypeus polished, becoming weakly imbricate in the basal half; punctures irregularly sparse (IS ≤ 2 PD) apically, becoming smaller and denser (IS ≤ 1 PD) basally. Supraclypeal area polished, becoming imbricate posterolaterally; punctures sparse (IS = 1 – 3 PD) anteromedially, becoming dense (IS ≤ 1 PD) posterolaterally. Frons reticulate-punctate; punctures crowded (IS = 0 PD). Ocellocular area polished; punctures dense (IS <1 PD). Gena polished and weakly lineolate posteriorly; punctures sparse (IS = 1 – 2 PD), becoming obscure posteriorly. Postgena polished, weakly lineolate adjacent to hypostomal carina and posteriorly. Tegula broadly impunctate, minute punctures present anteriorly. Mesoscutum imbricate throughout with a subtle weakening submedially; punctures dense throughout (IS <1 PD), becoming moderately dense submedially (IS = 1 – 1.5 PD), diversopunctate. Mesoscutellum polished; punctures dense (IS <1 PD) marginally and on median line, sparse submedially (IS = 1 – 4 PD). Metanotum rugulose laterally, ruguloso-punctate medially with punctures crowded (IS = 0 PD). Propodeal triangle with rugae moderately strong, reaching half towards posterior margin, anastomosing medially and weakly so laterally, lateral rugae not extending to dorsolateral slope of propodeum lateral face; interspace sculpture imbricate to tessellate. Preepisternum weakly rugulose. Hypoepimeron imbricate to weakly rugulose; impunctate. Mesepisternum finely rugulose, imbricate ventrally; impunctate. Propodeum lateral face strongly imbricate, punctures obscure; posterior face polished to weakly imbricate, with punctures dense anteriorly (IS ≤ PD), becoming irregularly sparser posteriorly (IS <3 PD). T 1 anterior face polished, punctures present laterally, sparse (IS = 1 – 4 PD); dorsal surface polished, punctures present laterally, sparse (IS = 1 – 3 PD), becoming sparser (IS = 1 – 6 PD) towards premarginal line and nearly impunctate medially; rim coriarious, punctures sparse (IS = 1 – 3 PD). T 2 disc coriarious to polished, base densely punctate (IS ≤ 1 PD), becoming moderately dense to moderately sparse (IS = 1 – 2 PD) apically; rim coriarious, with minute and sparse punctures (IS = 1 – 4 PD), becoming impunctate narrowly after laterally premarginal line. STRUCTURE. Head long, length / width ratio 1.04 (± 0.01 SD). Face length / width ratio 0.86 (± 0.01 SD). Clypeus projecting ~ 74 % below suborbital tangent. Mandible short, reaching opposing clypeal angle. Eyes convergent below, UOD / LOD ratio 1.07 (± 0.02 SD). Ocelli normal. Gena / eye width ratio 0.68 (± 0.04 SD). Pronotal angle obtuse. Intertegular span 0.89 mm (± 0.05 SD). Mesoscutum length / width ratio 0.90 (± 0.03 SD); mesoscutum / scutellum length ratio 2.35 (± 0.25 SD); scutellum / metanotum length ratio 1.74 (± 0.17 SD); metanotum / propodeal triangle length ratio 0.64 (± 0.08 SD). Protibial spur with apical serrations shorter than width of malus; malus extending beyond the velum, nearly ¾ the length of velum. Inner metatibial spur pectinate, teeth 3 – 5, basal teeth longer than width of rachis. Propodeal triangle posterior margin rounded onto posterior propodeal surface. Propodeal lateral carina not reaching dorsal margin, oblique carina absent. VARIATION. Occasionally, the rims of the metasomal terga become conspicuously more yellow. Sometimes, the supraclypeal area is weakly imbricate medially. The ocellocular area is sometimes weakly rugulose, with crowded punctures (IS = 0 PD). The mesoscutum exhibits variation in puncture density and texture. In some individuals, it is more imbricate throughout, with a pronounced microsculpture. Others have a mostly imbricate mesoscutum that becomes nearly polished submedially. Occasionally, the punctures are dense throughout (IS <1 PD), but in some cases, they become sparser submedially (IS = 1 – 3 PD). Male BODY MEASUREMENTS. Length 6.65 mm (± 0.34 SD). Head length 1.57 mm (± 0.07 SD). Face length mm 1.36 (± 0.06 SD). Head width 1.45 mm (± 0.07 SD). Wing length 4.36 mm (± 0.10 SD). n = 5. COLOURATION. Head and mesosoma black. Clypeus apical half yellow. Labrum brown. Mandible brown basally, becoming yellow medially and reddish brown apically. Flagellum brown dorsally, yellow ventrally. Pronotal lobe brown. Tegula translucent yellowish brown. Wing membrane faintly brown, veins brown. Legs dark brown to brown, except yellow tarsi. Metasoma dark brown with rims of terga and sterna narrowly lighter. PUBESCENCE. Body hair colour off-white. Tomentum dense on paraocular area, clypeus basally, supraclypeal area laterally, pronotal angle, and lobe, space between pronotal lobe and tegula. Mesosoma covered by erect plumose hairs (1 – 2 OD) and sub-appressed simple short hairs (<0.5 OD). Metasomal sterna with dense, erect and sub-appressed, plumose, short (<1.5 OD) hairs. Wing hairs brown, very short and sparse. SURFACE SCULPTURING. Clypeus polished; punctures irregularly sparse (IS ≤ 2 PD) apically, becoming smaller and denser (IS ≤ 1 PD) basally. Supraclypeal area polished; punctures dense (IS <1 PD). Frons reticulate-punctate; punctures crowded (IS = 0 PD). Ocellocular area polished, punctures dense (IS ≤ 1 PD). Gena polished and weakly lineolate posteriorly; punctures moderately sparse (IS = 1 – 2 PD), becoming obscure posteriorly. Postgena polished anteriorly, become lineolate posteriorly. Tegula broadly impunctate. Mesoscutum mostly polished, slightly imbricate anteriorly; punctures dense (IS ≤ 1 PD), becoming sparse (IS = 1 – 2 PD) mesad of parapsidal line, diversopunctate. Mesoscutellum polished; punctures dense (IS ≤ 1 PD) marginally and on median line, sparse submedially (IS = 1 – 2 PD). Metanotum rugulose. Propodeal triangle with rugae moderately strong, reaching ¾ towards posterior margin, anastomosing medially and weakly so laterally, lateral rugae not extending to dorsolateral slope of propodeum lateral face; interspace sculpture imbricate to tessellate and becoming polished posteriorly on posterior margin. Preëpisternum weakly rugulose. Hypoepimeron imbricate to weakly rugulose; punctures obscure. Mesepisternum reticulate-punctate with crowded punctures, becoming imbricate ventrally where the punctures become obscure or impunctate. Propodeum lateral face strongly imbricate, and becoming rugulose dorsolateral slope, punctures obscure; posterior face weakly rugulose-imbricate, with dense punctures anteriorly (IS <1 PD). T 1 anterior face polished, few scattered punctures present laterally; dorsal surface polished, punctures present laterally, moderately sparse (IS = 1 – 2 PD), becoming sparser (IS = 1 – 4 PD) on subapicolateral boss; rim polished, punctures moderately sparse (IS = 1 – 2 PD) and becoming impunctate laterally and apically. T 2 disc coriarious basally, becoming polished apically, with dense (IS ≤ 1 PD); rim polished, with minute, sparse punctures (IS = 1 – 4 PD), becoming impunctate apically. STRUCTURE. Head long, length / width ratio 1.09 (± 0.00 SD). Face length / width ratio 0.93 (± 0.01 SD). Clypeus projecting ~ 67 % below suborbital tangent. Mandible short, reaching opposing clypeal angle. Flagellomeres long, F 1 length / pedicel length ratio 1.10 (± 0.11 SD); F 2 length / F 1 length ratio 2.46 (± 0.21 SD); F 2 length / F 2 width ratio 1.66 (± 0.15 SD). Gena / eye width ratio 0.60 (± 0.03 SD). Forewing with 3 submarginal cells. Pronotal angle obtuse. Intertegular span 0.73 mm (± 0.09 SD). Mesoscutum length / width ratio 0.99 (± 0.05 SD). Propodeal lateral carina not reaching dorsal margin, oblique carina absent. n = 5. GENITALIA (Figs 48, 120 H). Gonostylus elongate, about 3 times as long as broad, and concave on outer margin, with long setae. Retrorse lobe slender, attenuated apically, about 3 times as long as broad, covered in short setae. VARIATION. Sometimes, the tegula becomes more translucent yellow. Occasionally, the tarsi get more yellowish-brown. Typically, there is a strong colour gradient between the tibia and basitarsus. Rarely, some specimens exhibit a subtle colour gradient, similar to that observed in the legs of male L. ruficorne. In some instances, the punctures on the supraclypeal area can be nearly crowded in the middle. In some specimens, the mandible slightly extends beyond the opposing clypeal angle.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFA4FFAFFDF6FB32FB0BFDF7.taxon	distribution	Range Canada (Alberta, British Columbia, Saskatchewan), Mexico (Baja California, Sonora), USA (Arizona, California, Colorado, Idaho, Montana, Nevada, New Mexico, Oregon, Utah, Washington, Wyoming) (Fig. 49).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFA4FFAFFDF6FB32FB0BFDF7.taxon	biology_ecology	Floral hosts AGAVACEAE: Agave utahensis var. nevadensis Engelm. ex Greenm. & Roush, Yucca sp. • ANACARDIACEAE: Rhus trilobata Nutt. • APIACEAE: Lomatium dissectum (Nutt.) Mathias & Constance • ASCLEPIADACEAE: Asclepias tuberosa L. • ASTERACEAE: Baccharis salicifolia (Ruiz & Pav.) Pers., Lasthenia sp., Baileya multiradiata Harv. & A. Gray, Chrysothamnus sp., Cirsium cymosum (Greene) J. T. Howell, Encelia farinosa A. Gray ex Torr., Encelia sp., Gutierrezia microcephala (DC.) A. Gray, Haplopappus sp., Barkleyanthus salicifolius (Kunth) H. Rob. & Brettell, Solidago altissima L., Solidago sp., Taraxacum sp., Heliomeris multiflora Nutt. • BORAGINACEAE: Cryptantha barbigera (A. Gray) Greene, Cryptantha intermedia (A. Gray) Greene, Cryptantha utahensis (A. Gray) Greene, Eremocarya micrantha (Torr.) Greene, Heliotropium sp., Plagiobothrys nothofulvus (A. Gray) A. Gray • BRASSICACEAE: Brassica sp., Cardaria sp., Descurainia sophia (L.) Webb ex Prantl, Erysimum capitatum (Douglas ex Hook.) Greene, Isatis tinctoria L., Lepidium appelianum Al-Shehbaz, Lepidium fremontii S. Watson, Lepidium montanum Nutt., Lobularia maritima (L.) Desv., Mutarda arvensis (L.) D. A. German, Physaria bellii G. A. Mulligan, Physaria sp., Physaria vitulifera Rydb., Sinapis alba L. or Brassica olerace a L., Sisymbrium sp., Stanleya pinnata (Pursh) Britton • CACTACEAE: Opuntia sp. • CAPPARACEAE: Cleomella arborea (Nutt.) Roalson & J. C. Hall Greene • CAPRIFOLIACEAE: Viburnum tinus L. • CONVOLVULACEAE: Convolvulus sp. • ERICACEAE: Arctostaphylos sp. • FABACEAE: Amorpha fruticosa L., Astragalus agrestis Douglas ex G. Don, Astragalus crotalariae (Benth.) A. Gray, Astragalus sp., Parkinsonia microphylla Torr., Melilotus officinalis (L.) Lam., Melilotus sp. • GROSSULARIACEAE: Ribes indecorum Eastw., Ribes sp. • HYDRANGEACEAE: Fendlera sp. • HYDROPHYLLACEAE: Ellisia sp., Emmenanthe penduliflora Benth., Eriodictyon sp., Eriodictyon trichocalyx A. Heller, Phacelia calthifolia Brand, Phacelia ciliata Benth., Phacelia distans Benth., Phacelia minor (Harv.) Thell., Phacelia sp., Phacelia tanacetifolia Benth. • LAMIACEAE: Condea emoryi (Torr.) Harley & J. F. B. Pastore, Hyptis sp., Poliomintha incana (Torr.) A. Gray, Salvia dorrii var. incana (Benth.) Strachan • LILIACEAE: Allium siskiyouense Ownbey ex Traub • LOASACEAE: Eucnide urens Parry ex Coville • MALVACEAE: Sphaeralcea ambigua A. Gray, Sphaeralcea sp. • ONAGRACEAE: Chylismia claviformis subsp. integrior (P. H. Raven) W. L. Wagner & Hoch, Clarkia modesta Jeps., Clarkia pulchella Pursh, Oenothera brevipes A. Gray, Oenothera sp., Taraxia tanacetifolia (Torr. & A. Gray) Piper • PAPAVERACEAE: Papaver sp. • POLEMONIACEAE: Gilia sp. • POLYGONACEAE: Eriogonum fasciculatum Benth., Eriogonum inflatum Torr. & Frém., Eriogonum sp., Eriogonum umbellatum Torr., Eriogonum wrightii var. subscaposum S. Watson • PORTULACACEAE: Calyptridium umbellatum (Torr.) Greene • RANUNCULACEAE: Ranunculus sp. • RHAMNACEAE: Ceanothus sp., Frangula californica (Eschsch.) A. Gray, Rhamnus crocea Nutt., Frangula californica subsp. tomentella (Benth.) Kartesz & Gandhi • ROSACEAE: Cercocarpus montanus Raf., Malus sp., Prunus andersonii A. Gray, Prunus fremontii S. Watson, Prunus ilicifolia (Nutt. ex Hook. & Arn.) D. Dietr., Prunus sp., Prunus virginiana var. demissa (Nutt.) Torr., Prunus subcordata Benth., Prunus virginiana L., Purshia tridentata (Pursh) DC., Rubus deliciosus Torr. • SALICACEAE: Salix exigua Nutt., Salix vestita Pursh, Salix sp. • TAMARICACEAE: Tamarix gallica L., Tamarix sp. • ZYGOPHYLLACEAE: Larrea tridentata (DC.) Coville, Larrea sp.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FFA4FFAFFDF6FB32FB0BFDF7.taxon	discussion	DNA barcodes DNA barcodes of the L. ruficorne form a single cluster with sequences from Alberta, British Columbia, Northwest Territories, Saskatchewan, Colorado, Oregon, Utah, Nevada, and New Mexico.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF92FFB1FDCFFDD3FAF0F96E.taxon	description	Figs 52 – 53, 117 A, 120 I	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF92FFB1FDCFFDD3FAF0F96E.taxon	etymology	Etymology The specific epithet is a noun in the genitive case, in reference to the host plant of the holotype, Synthyris plantaginea.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF92FFB1FDCFFDD3FAF0F96E.taxon	diagnosis	Diagnosis Refer to L. diatretum diagnosis.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF92FFB1FDCFFDD3FAF0F96E.taxon	materials_examined	Type material Holotype of Halictus (Evylaeus) synthyridis (holotype not examined; current location unknown) UNITED STATES – Colorado • ♀; Florissant; 16 Jun. 1906; T. D. A. Cockerell leg.; ex Synthyris plantaginea; UCMC. Paratypes of Halictus (Evylaeus) synthyridis (designated herein; examined) UNITED STATES – Colorado • 1 ♀; Teller Co., Florissant; 9 Jul.; S. R. Rohwer leg.; USNM (Type No. 12048) (Figs 52 – 53) • 1 ♂; same data as for preceding; 18 Jul.; S. R. Rohwer leg.; Polemonium; UCRC. Specimen labels “ Florissant, / Colo. July 9. / S. R. Rohwer // {illegible} // H. synthyridis / Ckll Cotype // CoType / No 12048 / U. S. N. M. // PARATYPE ” • “ Florissant, / Colo. July 18. / S. R. Rohwer // Polemonium // H. synthyridis / Ckll Cotype // UCRC // PARATYPE ”. Type notes The holotype of L. synthyridis is cataloged under UCMC, but it was not located within UCMC collections, and its current whereabouts remain unknown. Among the examined material, one female paratype (USNM 12048) and one male paratype (UCRC) correspond to Lasioglossum inconditum (Cockerell, 1916). Nontype material identified by T. D. A. Cockerell closely resembles Lasioglossum diatretum (Vachal, 1904). Based on the original description, it is likely that additional paratypes of L. synthyridis exist, but these have not been located. Given the clear correspondence of its paratypes to L. inconditum, we formally synonymize L. synthyridis as a senior subjective synonym of L. inconditum (Cockerell, 1916).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF92FFB1FDCFFDD3FAF0F96E.taxon	description	Description GENITALIA (Figs 53, 120 I). Gonostylus elongate, about 4.5 times as long as broad, and with short setae. Retrorse lobe pointed apically, about 4 times as long as broad, covered in short setae.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF92FFB1FDCFFDD3FAF0F96E.taxon	discussion	Remarks For detailed description and distribution information (as L. inconditum), see Gibbs et al. (2013).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF8CFFB4FDC4F95EFBD0FB4D.taxon	description	urn: lsid: zoobank. org: act: C 03 DBFC 6 - 0 AD 1 - 49 BC- 8 EF 9 - FCAAF 9502 AA 9 Figs 54, 55, 93 B, 97 C, 104 H, 106 B	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF8CFFB4FDC4F95EFBD0FB4D.taxon	diagnosis	Diagnosis Females of L. opata sp. nov. are readily distinguished from other western North American species by their larger body size (length 8.18 mm ± 0.54 SD), enlarged tegula, round head (length / width ratio 0.96 ± 0.01 SD), the propodeal triangle with weak rugae and faint apical carina, T 1 with a fan bearing a narrow dorsal opening, and distinct mesepisternal sculpture featuring strigate-rugulose patterning anteroventrally. Male is unknown.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF8CFFB4FDC4F95EFBD0FB4D.taxon	etymology	Etymology The specific epithet ‘ opata’ is derived from the name of the Opata tribe, an Indigenous community historically residing in northern Mexico along the border with Arizona. This naming choice serves as a tribute to the Opata nation. It is treated as a noun in apposition.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF8CFFB4FDC4F95EFBD0FB4D.taxon	materials_examined	Type material Holotype MEXICO – Sonora • ♀; Nogales; [31.308 ° N, 110.942 ° W]; 4 Sep. 1962; A. E. Michelbacher leg.; EMEC (Fig. 54). Specimen labels “ MEX: Son. / Nogales, / IX – 4 – 65 // A. E. Michelbacher Collector // Lasioglossum GIBBS- 14503 // HOLOTYPE / Lasioglossum (Hemihalictus) opata Hettiarachchi and Gibbs ”. Paratypes MEXICO – Sonora • 1 ♀; Álamos; [27.023 ° N, 108.935 ° W]; 27 Feb. 1963; P. H. Arnaud Jr leg.; EMEC • 1 ♀; same data as for preceding; 24 Feb. 1963; P. H. Arnaud Jr leg.; EMEC • 24 ♀♀; Navojoa; [27.072 ° N, 109.444 ° W]; 19 Aug. 1962; A. E. Michelbacher and M. M. Michelbacher leg.; ex domestic squash; EMEC • 1 ♀; Nogales; [31.308 ° N, 110.942 ° W]; 4 Sep. 1962; A. E. Michelbacher leg.; EMEC. UNITED STATES – Arizona • 1 ♀; Santa Cruz Co., Patagonia; [31.54 ° N, 110.76 ° W]; 19 Apr. 1956; G. D. Butler leg.; ex alfalfa; ASUHIC.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF8CFFB4FDC4F95EFBD0FB4D.taxon	description	Description Female BODY MEASUREMENTS. Length 8.18 mm (± 0.54 SD). Head length 1.89 mm (± 0.04 SD). Face length 1.59 mm (± 0.03 SD). Head width 1.97 mm (± 0.05 SD). Wing length 5.60 mm (± 0.10 SD). n = 5. COLOURATION. Head and mesosoma black. Clypeus apex dark brown. Labrum dark brown. Mandible brown with dark brown base. Flagellum brown dorsally, light brown ventrally. Pronotal lobe brown. Tegula mostly dark brown, light brown medially. Wing membrane faintly brownish, veins with subcosta brown, otherwise light brown. Legs dark brown. Metasoma dark brown to brown with rims of terga, sterna and downcurved lateral areas of terga broadly light brown. PUBESCENCE. Body hair colour off-white. Tomentum dense on pronotal angle and lobe, space between pronotal lobe and tegula, metanotum, propodeum lateral surface anterodorsally, and T 2 – T 4 basolaterally; moderately dense on gena, anterolateral angle and marginally on mesoscutum; sparse on paraocular area, propodeum posterior surface mostly on dorsally. Mesosoma covered by appressed, simple, very short hairs (<0.25 OD) mixed with sparse, erect, weakly plumose, long hairs (<1 OD) on mesoscutum and erect, plumose, long hairs (<2 OD) on other areas. Metasomal sterna with moderately sparse, erect, weakly plumose, long (<3 OD) hairs. Wing hairs brown, very short and sparse. T 1 with a fan, narrow dorsal opening (0.5 OD), composed of overlapping sub-appressed simple hairs (<2 OD). SURFACE SCULPTURING. Clypeus tessellate, becoming polished apical half; punctures irregularly sparse (IS <4 PD) apically and medially, becoming smaller and denser (IS <1 PD) basolaterally. Supraclypeal area tessellate medially, imbricate posterolaterally; punctures fine, dense (IS ≤ 1 PD). Frons reticulate-punctate punctures crowded (IS = 0 PD). Ocellocular area polished; punctures moderately sparse (IS = 1 – 2 PD) and distinctly doubly punctate with minute punctures throughout. Gena polished, becoming lineolate posteriorly; punctures dense (IS ≤ 1 PD), becoming obscure posteriorly. Postgena polished, weakly lineolate adjacent to hypostomal carina. Tegula punctures dense (IS ≤ 1 PD) in the anterior half and basally, becoming impunctate posteriorly. Mesoscutum tessellate to imbricate throughout; punctures dense (IS ≤ 1 PD), becoming anteromedially moderately sparse (IS = 1 – 2 PD), doubly punctate. Mesoscutellum polished medially, becoming weakly rugulose to imbricate marginally; punctures dense (IS <1 PD) marginally and on median line, sparse submedially (IS = 1 – 3 PD), diversopunctate. Metanotum weakly rugulose, punctures obscure. Propodeal triangle with rugae weak, only visible laterally, weakly anastomosing, extending to dorsolateral slope; interspace sculpture strongly imbricate. Preëpisternum rugulose. Hypoepimeron strongly rugulose; punctures obscure. Mesepisternum rugulose, becoming strigate-rugulose anteroventrally. Propodeum lateral face imbricate; posterior face rugulose. T 1 anterior face dull, strongly coriarious; dorsal surface polished to coriarious, punctures sparse (IS = 1 – 3 PD), becoming more sparsely punctate (IS = 2 – 7 PD) in small subapicolateral boss and punctures minute, and sparse (IS = 1 – 3 PD) on rim, becoming impunctate apical half and medially on rim. T 2 disc weakly coriarious to polished, punctures moderately dense (IS = 1 – 2 PD) basally, becoming sparse (IS = 1 – 4 PD) apically; rim coriarious, punctures minute and sparse (IS = 1 – 3 PD). STRUCTURE. Head round, length / width ratio 0.96 (± 0.01 SD). Face length / width ratio 0.81 (± 0.01 SD). Clypeus projecting ~ 77 % below suborbital tangent; clypeus length / width ratio 1.07 (± 0.08 SD). Mandible short, just extending beyond opposing clypeal angle. Eyes convergent below, UOD / LOD ratio 1.13 (± 0.01 SD). Ocelli normal. Gena / eye width ratio 0.85 (± 0.06 SD). Pronotal angle obtuse. Intertegular span 1.47 mm (± 0.05 SD). Mesoscutum length / width ratio 0.93 (± 0.03 SD); mesoscutum / scutellum length ratio 2.90 (± 0.14 SD); scutellum / metanotum length ratio 1.53 (± 0.16 SD); metanotum / propodeal triangle length ratio 0.88 (± 0.10 SD). Protibial spur with apical serrations subequal in length to width of malus; malus extending beyond the velum, nearly as length as velum. Inner metatibial spur pectinate, teeth 4 – 5, basal teeth longer than width of rachis. Propodeal triangle posterior margin angled onto posterior propodeal surface, apical carina faint. Propodeal lateral carina reaching dorsolateral slope; oblique carina indistinct. n = 5. Male Unknown.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF8CFFB4FDC4F95EFBD0FB4D.taxon	distribution	Range Sonoran Deserts to southern Arizona (Fig. 55).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF8CFFB4FDC4F95EFBD0FB4D.taxon	biology_ecology	Floral hosts CUCURBITACEAE: Cucurbita sp. • FABACEAE: Medicago sativa L.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF88FF41FE17FEE2FAA3FD73.taxon	description	Figs 56 – 59, 95 F, 98 C, 104 L, 112 D, 119 A, 120 J	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF88FF41FE17FEE2FAA3FD73.taxon	etymology	Etymology The specific epithet is a reference to the head shape, which Cockerell describes as ‘ longitudinally oval’.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF88FF41FE17FEE2FAA3FD73.taxon	diagnosis	Diagnosis Both sexes of L. ovaliceps can be recognized by the combination of head long, female and male head length / width ratio 1.10 (± 0.02 SD) and 1.22 (± 0.02 SD), respectively; propodeal triangle with rugae partially extending to the posterior margin; and metasoma orange. Lasioglossum aspilurus also has an orange metasoma, but the later species has a much wider head, female and male head length / width ratio 0.91 (± 0.01 SD) and 0.96 (± 0.02 SD), respectively, and the rugae of the propodeal triangle almost reaching the posterior margin. There are dark morphs of male L. ovaliceps with brown metasoma. In this case, they could be confused with L. buccale males, but males of L. buccale are comparatively larger (7.5 – 8.5 mm); clypeus slightly yellow only apically; the rugae of the propodeal triangle reaching posterior margin. Lasioglossum ovaliceps are superficially similar in colour to Sphecodes, but the generic characters for Lasioglossum will distinguish them and the long head and yellow maculation of the male is not present in Sphecodes. Members of the L. (Dialictus) petrellus species group typically have long heads and reddish metasoma but have the head and mesosoma dull-metallic (Gibbs 2009).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF88FF41FE17FEE2FAA3FD73.taxon	materials_examined	Type material Syntype of Halictus ovaliceps (designated herewith; examined by photos) UNITED STATES – New Mexico • 1 ♀; Santa Fe Co., Santa Fe; [35.67 ° N, 105.96 ° W]; Aug; Boyle leg.; USNM Type No. 5832 .. Specimen labels “ Ckll. 4034. / Sta Fe. Aug. / Boyle. // Type / No. 5832 / U. S. N. M. // H. ovaliceps / Ckll. TYPE. // USNMENT / 01735680 ”. Type notes The species was described as part of a key and no information on the composition of the type series was available in the original description. Following recommendation 73 F of the Code, we assume the existence of a syntype series. A specimen at the USNM (above) is labelled as ‘ type’. The identity of the species is not in dispute, so no lectotype designation is required. Material examined CANADA – Alberta • 1 ♀; Cypress Co., Cypress Hills PP, Battle Creek Rd, Graburn Overlook; 49.6625 ° N, 110.042222 ° W; elev. 1220 – 1260 m; 6 – 7 Jun 2025; M. Buck leg.; PMAE • 1 ♀; Onefour; 49.156167 ° N, 110.273278 ° W; 29 Jul. 2010; N. de Silva leg.; RSKM • 1 ♀; Waterton ID, Waterton NP, Lower Waterton Lake, E side; 49.088333 ° N, 113.845833 ° W; elev. 1280 m; 19 Jul 2023; M. Buck leg.; PMAE. – British Columbia • 1 ♀; Ag. Can. Rea. Stn; [49.57 ° N, 119.64 ° W]; 27 Jun. 1990; S. G. Cannings leg.; UBCZ • 1 ♀; Nanaimo, Biological Station; [49.211 ° N, 123.955 ° W]; 23 Jun. 1920; E. P. Van Duzee leg.; CAS • 1 ♀; Okanagan-Similkameen Reg. Dist. Osoyoos, Mt Kruger, Strawberry Creek. Road., N-site; 49.0399 ° N, 119.5084 ° W; elev. 505 m; 21 Jun. 2008; L. R. Best leg.; WRME • 2 ♀♀; Okanagan-Similkameen Reg. Dist., Mt Oliver, Ripley Lake; 49.2429 ° N, 119.6326 ° W; elev. 935 m; 2 May 2009; L. R. Best leg.; PCYU • 1 ♀; Okanagan-Similkameen Reg. Dist., Ripley Lake; 49.243355 ° N, 119.634954 ° W; elev. 950 m; 25 Mar. 2009; L. R. Best leg.; PCYU • 1 ♀; Oliver, Burnell Lake; 49.207887 ° N, 119.612148 ° W; 27 Jun. 2020; UBCZ • 1 ♀; Penticton, West Bench; [49.495 ° N, 119.618 ° W]; 14 Jun. 1987; S. G. Cannings leg.; UBCZ • 1 ♀; same data as for preceding; 24 Jun. 1988; S. G. Cannings leg.; UBCZ • 1 ♀; Squamish, Keyhole Hotsprings; 50.668404 ° N, 123.460826 ° W; 16 Jun. 2020; J. Heron leg.; RSKM. UNITED STATES – California • 1 ♀; Alameda Co., Oakland; [37.804 ° N, 122.271 ° W]; 21 May 1937; E. S. Ross leg.; CAS • 1 ♀; same data as for preceding; 20 Apr. 1952; R. Schuster leg.; EMEC • 1 ♀; Alameda Co., Tesla; [37.62 ° N, 121.49 ° W]; 27 Apr. 1960; W. F. Ehrhardt leg.; EMEC • 1 ♂; Alpine Co., 1 mi. S of Monitor Pass; [38.661 ° N, 119.621 ° W]; elev. 2499.4 m; 22 – 23 Aug. 1979; M. Wasbauer and P. Adams leg.; CSCA • 1 ♀; Alpine Co., Hope Valley; [38.780 ° N, 119.930 ° W]; 18 Jul. 1948; EMEC • 1 ♀; Alpine Co., Woodfords; [38.79 ° N, 119.82 ° W]; 12 Jun. 1962; EMEC • 1 ♀; Calaveras Co., 1.5 mi. SE of Railroad Flat; [38.327 ° N, 120.492 ° W]; 1 – 2 Jun. 1970; EMEC • 1 ♀; same data as for preceding; 1 Jun. 1970; EMEC • 8 ♀♀; Calaveras Co., 4 mi. S of Railway Flat; [38.285 ° N, 120.513 ° W]; elev. 853.4 m; 18 May 1969; ex Chamaebatia foliolosa; EMEC (Fig. 56 D) • 4 ♀♀; same data as for preceding; 19 May 1969; EMEC • 1 ♀; Calaveras Co., Copperopolis; [37.981 ° N, 120.642 ° W]; 15 Apr. 1954; EMEC • 1 ♀; same data as for preceding; 25 Apr. 1954; EMEC • 1 ♀; Calaveras Co., Mokel Hill; [38.301 ° N, 120.706 ° W]; Apr; F. E. Blaisdell leg.; CAS • 1 ♀; Contra Costa Co., Mount Diablo; [37.882 ° N, 121.914 ° W]; 25 Jun. 1939; E. C. Van Dyke leg.; CAS • 1 ♀; Del Norte Co., Gasquet; [41.845 ° N, 123.969 ° W]; 6 Jul. 1960; T. R. Haig leg.; CSCA • 1 ♀; El Dorado Co., 3 mi. S of Camino; [38.694 ° N, 120.674 ° W]; 26 Jun. 1948; P. D. Hurd leg.; EMEC • 1 ♀; El Dorado Co., Angora Peak; [38.867 ° N, 120.073 ° W]; elev. 2628.9 m; 19 Jul. 1931; E. O. Essig leg.; EMEC • 1 ♀; El Dorado Co., Blodgett Forest, 13 mi. E of Georgetown; [38.892 ° N, 120.663 ° W]; 20 Jul. 1967; EMEC • 1 ♀; El Dorado Co., Camino; [38.738 ° N, 120.675 ° W]; 19 Jun. 1958; EMEC • 2 ♀♀; El Dorado Co., Echo Lake; [38.834 ° N, 120.042 ° W]; elev. 2255.5 m; 30 Jul. 1955; EMEC • 1 ♀; same data as for preceding; 17 Jul. 1961; EMEC • 4 ♀♀; same data as for preceding; 20 Jul. 1960; EMEC • 4 ♀♀; same data as for preceding; 8 Jul. 1961; EMEC • 6 ♀♀; same data as for preceding; 9 Jul. 1961; EMEC • 2 ♀♀; same data as for preceding; 13 Jul. 1956; EMEC • 1 ♀; same data as for preceding; 17 Jun. 1961; EMEC • 1 ♀; same data as for preceding; 1 Jul. 1954; W. W. Middlekauff leg.; EMEC • 1 ♀; same data as for preceding; 29 Jun. 1954; W. W. Middlekauff leg.; EMEC • 1 ♀; same data as for preceding; 17 Jul. 1956; W. W. Middlekauff leg.; EMEC • 1 ♀; same data as for preceding; 2 Jul. 1951; W. W. Middlekauff leg.; EMEC • 1 ♀; same data as for preceding; 6 Jul. 1951; W. W. Middlekauff leg.; EMEC • 1 ♀; same data as for preceding; 9 Jul. 1951; W. W. Middlekauff leg.; EMEC • 1 ♀; same data as for preceding; 21 Jul. 1948; P. D. Hurd leg.; EMEC • 1 ♀; Fresno Co., Sulphur Mdw., Shaver Lake; [37.1042 ° N, 119.3175 ° W]; 12 – 19 Jun. 1968; J. R. Hoy leg.; UCMC • 1 ♀; Humboldt Co., Orleans; [41.301 ° N, 123.54 ° W]; 12 Jun. 1961; T. W. Koerber leg.; USNM • 2 ♀♀; Inyo Co., Big Pine Creek Campground; [38.796 ° N, 116.85 ° W]; 14 Jul. 1927; P. H. Timberlake leg.; ex Castilleja applegatei subsp. pinetorum (verbatim: Castilleja pinetorum); UCRC • 1 ♂; same data as for preceding; 4 Jul. 1927; P. H. Timberlake leg.; ex Castilleja applegatei subsp. pinetorum (verbatim: Castilleja pinetorum); UCRC • 1 ♂; Inyo Co., Lone Pine; [36.61 ° N, 118.06 ° W]; 16 Jun. 1937; E. C. Van Dyke leg.; CAS • 2 ♀♀; Inyo Co., Panamint Mountains; [36.17 ° N, 117.089 ° W]; 25 Apr. 1953; P. H. Timberlake leg.; ex Datura inoxia (verbatim: Datura meteloides); UCRC • 1 ♀; Inyo Co., Surprise Canyon; [36.113 ° N, 117.16 ° W]; 28 Apr. 1953; C. A. Marsh leg.; ex Eucnide urens; EMEC • 1 ♀; Inyo Co., Surprise Canyon, Panamint Mountains; [36.113 ° N, 117.160 ° W]; 24 Apr. 1957; J. Powell leg.; EMEC • 1 ♀; Kern Co., 8 mi. NW of Mojave; [35.126 ° N, 118.271 ° W]; 13 Apr. 1957; EMEC • 1 ♀; Kern Co., Alder Creek Campground; [35.719 ° N, 118.612 ° W]; 19 Jul. 1958; EMEC • 1 ♀; Kern Co., Greenhorn MB; [35.729 ° N, 118.552 ° W]; 10 Jul. 1955; EMEC • 1 ♀; Kern or Tulare Co., Greenhorn Mountains; [35.85 ° N, 118.573 ° W]; 10 Jul. 1955; EMEC • 1 ♀; King Co.; 47.5039 ° N, 122.2757 ° W; 19 Apr. 2015; R. Redmond and E. A. Sugden leg.; WRME • 2 ♀♀; Lake Co., 0.1 mi. S of Hobergs; [38.842 ° N, 122.723 ° W]; 2 Jun. 1968; ex Clarkia rhomboidea; EMEC • 1 ♀; Lake Co., Bartlett Springs Road (8.2 mi. NE) junction Hwy 20; [38.136 ° N, 122.766 ° W]; elev. 1097.3 m; 28 Jul. 1980; ex Keckiella lemmonii; EMEC • 1 ♀; Lassen Co., Susanville; [40.42 ° N, 120.65 ° W]; 15 Jul. 2011; EMEC • 1 ♀; Los Angeles Co., Acton; [34.480 ° N, 118.190 ° W]; 14 Mar. 1936; E. G. Linsley leg.; ex Salix; EMEC • 1 ♀; same data as for preceding; 19 Mar. 1936; E. G. Linsley leg.; ex Salix; EMEC • 1 ♀; Los Angeles Co., Arcadia; [34.133 ° N, 118.036 ° W]; MCZ • 2 ♀♀; Los Angeles Co., Bell Canyon, San Dimas Experimental Forest; [34.206 ° N, 118.690 ° W]; elev. 218.3 m; 2 May 1965; EMEC • 2 ♀♀; Los Angeles Co., Benedict Canyon; [34.099 ° N, 118.431 ° W]; (?) Mar. 1971; EMEC • 1 ♀; Los Angeles Co., Catalina Island; [33.382 ° N, 118.430 ° W]; 6 Apr. 1938; W. P. Cockerell leg.; CAS • 1 ♀; Los Angeles Co., Claremont; [34.100 ° N, 117.720 ° W]; 15 May 1928; EMEC • 1 ♀; same data as for preceding; 4 May 1915; P. H. Timberlake leg.; ex Penstemon spectabilis; UCRC • 1 ♀; same data as for preceding; 27 Apr. 1927; T. Craig leg.; CAS • 1 ♀; same data as for preceding; 8 Mar. 1909; USNM • 7 ♀♀; same data as for preceding; Baker leg.; USNM • 2 ♀♀; Los Angeles Co., Devil’s Punchbowl; [34.420 ° N, 117.852 ° W]; 1 May 1968; EMEC • 1 ♀; Los Angeles Co., Fish Canon; [34.602 ° N, 118.665 ° W]; 25 Apr. 1928; EMEC • 1 ♀; Los Angeles Co., Griffith Park; [34.137 ° N, 118.294 ° W]; 5 Apr. 1936; E. G. Linsley leg.; ex Nemophila; EMEC • 1 ♀; Los Angeles Co., Los Angeles; [34.050 ° N, 118.240 ° W]; 21 Apr. 2017; EMEC • 1 ♀; Los Angeles Co., Monrovia; [34.150 ° N, 118.000 ° W]; 15 Aug. 1949; EMEC • 6 ♀♀; Los Angeles Co., Mts near Claremont; [34.161 ° N, 117.716 ° W]; Baker leg.; USNM • 1 ♀; same data as for preceding; Baker leg.; ex Salix; USNM • 1 ♀; Los Angeles Co., near Solromar; [34.052 ° N, 118.942 ° W]; 28 Jun. 1958; P. H. Arnaud leg.; ex Photinia; CSCA • 1 ♀; Los Angeles Co., Pasadena; [34.156 ° N, 118.131 ° W]; 2 Jun. 1944; MCZ • 1 ♀; Los Angeles Co., Puente Hills; [33.999 ° N, 117.917 ° W]; 8 May 1927; P. H. Timberlake leg.; ex Brassica nigra; UCRC • 1 ♀; Los Angeles Co., San Gabriel Canyon; [34.249 ° N, 117.866 ° W]; 7 Jul. 1946; EMEC • 1 ♀; Los Angeles Co., Tanbark Flat; [34.204 ° N, 117.760 ° W]; 10 Jul. 1950; EMEC • 1 ♀; same data as for preceding; 14 Jun. 1950; H. M. Graham leg.; EMEC • 1 ♀; Los Angeles Co., Tapia Park; [34.084 ° N, 118.707 ° W]; 9 May 1969; EMEC • 2 ♀♀; Los Angeles Co., Three Points; [34.736 ° N, 118.598 ° W]; 2 May 1968; EMEC • 1 ♀; Los Angeles Co., Westwood Hills; [38.302 ° N, 122.320 ° W]; 3 Jun. 1937; N. E. Hardman leg.; ex Brassica; EMEC • 1 ♀; Los Angeles Co., Whittier; [33.971 ° N, 118.031 ° W]; 13 Mar. 1926; P. H. Timberlake leg.; ex Mirabilis laevis var. laevis (verbatim: Mirabilis californica); UCRC • 1 ♀; same data as for preceding; 16 Apr. 1913; P. H. Timberlake leg.; ex Rubus vitifolius; USNM • 1 ♀; Los Angeles Co.; [34.33 ° N, 118.23 ° W]; 18 Jun. 1915; M. C. Van Duzee leg.; CAS • 1 ♀; same data as for preceding; 6321; EMEC • 2 ♀♀; same data as for preceding; USNM • 1 ♀; Madera Co., San Joaquin Experimental Range; [37.083 ° N, 119.716 ° W]; 18 Apr. 1953; P. D. Hurd leg.; ex Phacelia platyloba; EMEC • 1 ♀; Marin Co., Blue (?) Valley; 18 May 1919; C. L. Fox leg.; CAS • 1 ♀; Marin Co., Corte Madera C reek; [37.942 ° N, 122.502 ° W]; 1 – 30 Apr. 1915; EMEC • 1 ♀; Marin Co., Inverness; [38.101 ° N, 122.857 ° W]; 14 May 1961; C. A. Toschi leg.; EMEC • 1 ♀; Marin Co., Mill Valley; [37.906 ° N, 122.545 ° W]; 7 Nov. 1950; H. B. Leech leg.; CAS • 1 ♀; Mariposa Co., 1 mi. W of Mt Bullion; [37.507 ° N, 120.063 ° W]; 2 May 1960; ex Cryptantha; EMEC • 1 ♀; Mariposa Co., Exchequer Dam; [37.586 ° N, 120.269 ° W]; 10 Apr. 1960; ex Phacelia; EMEC • 1 ♀; Mendocino Co., UC Hopland Field Station near headquarters; [39.001 ° N, 123.083 ° W]; elev. 268.2 m; 2 Jun. 1968; EMEC • 2 ♀♀; Modoc Co., 1 mi. E of Pine Creek; [41.982 ° N, 120.256 ° W]; 13 Jun. 1970; EMEC • 4 ♀♀; Modoc Co., 5 mi. N of Davis Creek; [39.372 ° N, 119.827 ° W]; 3 Jun. 1971; ex Lithospermum ruderale; EMEC • 4 ♀♀; Modoc Co., Buck Creek Ranger Station; [48.269 ° N, 121.333 ° W]; elev. 1569.7 m; 5 – 7 Jun. 1970; EMEC • 4 ♀♀; same data as for preceding; 5 Jun. 1970; EMEC • 1 ♀; Modoc Co., Fandango Pass; [41.802 ° N, 120.207 ° W]; elev. 1859.3 m; 7 Jun. 1970; ex Wyethia; EMEC • 5 ♀♀; same data as for preceding; 7 Jun. 1970; EMEC • 2 ♀♀; Modoc Co., Rush Creek; 9 mi. NE of Adin; [41.263 ° N, 120.894 ° W]; 12 Jun. 1974; EMEC • 2 ♀♀; Mono Co., 7 mi. E of Bodie; [38.212 ° N, 118.885 ° W]; 2 Jul. 1964; EMEC • 1 ♀; Mono Co., Blanco’s Corral, White Mountains; [37.819 ° N, 118.476 ° W]; 25 Jul. 1960; EMEC • 1 ♀; same data as for preceding; 25 Aug. 1960; EMEC • 1 ♂; same data as for preceding; 25 Aug. 1960; P. D. Hurd leg.; EMEC • 1 ♂; same data as for preceding; 25 Aug. 1960; P. D. Hurd leg.; ex Senecio; EMEC • 1 ♀; Mono Co., McGee Creek; [37.573 ° N, 118.791 ° W]; 28 Jun. 1938; EMEC • 1 ♀; Mono Co., Paradise Camp; [38.288 ° N, 119.215 ° W]; 10 Aug. 1961; O. D. MacNeill leg.; CAS • 1 ♀; Mono Co., Silver Lake; [37.778 ° N, 119.123 ° W]; 16 Aug. 1948; W. S. Kelson leg.; EMEC • 1 ♂; Mono Co., 1 mi. W of Tom’s Place; [37.561 ° N, 118.700 ° W]; 13 Aug. 1957; EMEC • 1 ♀; Monterey Co., 4 mi. W of Soledad; [36.424 ° N, 121.398 ° W]; 17 Aug. 1964; G. R. Ballmer leg.; UCRC • 1 ♀; Monterey Co., 5 air mi. SW of Greenfield, Arroyo Seco River; [36.268 ° N, 121.308 ° W]; elev. 121.9 m; 5 May 1975; EMEC • 1 ♀; Monterey Co., Arroyo Seco River, 5 air mi. SW of Greenfield; [36.269 ° N, 121.307 ° W]; elev. 121.9 m; 5 May 1975; EMEC • 1 ♀; Monterey Co., Carmel; [36.555 ° N, 121.923 ° W]; 5 May 1930; L. S. Slevin leg.; CAS • 1 ♀; Monterey Co., Carmel Valley; [36.480 ° N, 121.735 ° W]; 6 Aug. 1967; T. Plichta and L. LaPre leg.; UCRC • 1 ♀; Monterey Co., Cone Peak, Santa Lucia Mountains; [36.031 ° N, 121.295 ° W]; elev. 1219.2 m; 25 Jul. 1980; ex Keckiella corymbosa; EMEC • 2 ♀♀; Monterey Co., Indians Rd, 3 air mi. S of Arroyo Seco Guard Station; [36.191 ° N, 121.493 ° W]; elev. 823 m; 7 May 1975; EMEC • 1 ♀; Monterey Co., Pebble Beach; [36.566 ° N, 121.947 ° W]; 1 – 30 April 1906; W. P. Cockerell leg.; ex Scrophularia villosa; MCZ • 4 ♀♀; Nevada Co., 6 mi. SW of Colfax; [39.039 ° N, 121.034 ° W]; 18 Apr. 1968; EMEC • 3 ♂♂; Nevada Co., Grass Valley; [39.219 ° N, 121.058 ° W]; elev. 762 m; 23 Jun. 1979; M. Wasbauer and F. Hillermann leg.; CSCA • 2 ♀♀; Nevada Co., Truckee; [39.325 ° N, 120.184 ° W]; 15 Jun. 1927; E. P. Van Duzee leg.; CAS • 2 ♀♀; same data as for preceding; 17 Jun. 1927; E. P. Van Duzee leg.; CAS • 1 ♀; same data as for preceding; 21 Jun. 1933; E. P. Van Duzee leg.; CAS • 1 ♀; same data as for preceding; 14 Jun. 1927; E. P. Van Duzee leg.; CAS • 3 ♀♀; Orange Co., Mount Santiago; [33.711 ° N, 117.534 ° W]; 19 Sep. 1926; P. H. Timberlake leg.; ex Ericameria parishii; UCRC • 1 ♀; same data as for preceding; 19 Sep. 1926; P. H. Timberlake leg.; ex Solidago velutina subsp. californica (verbatim: Solidago californica); UCRC • 1 ♀; Orange Co., Silverado; [33.746 ° N, 117.636 ° W]; 3 Nov. 1951; EMEC • 1 ♀; Orange Co., Starr Ranch Audubon Sanctuary, Santa Ana Mountains; [33.632 ° N, 117.555 ° W]; elev. 274.3 m; 13 Jul. 1980; ex Keckiella cordifolia; EMEC • 1 ♂; Placer Co., Ward Creek, 2 mi. S of Tahoe City; [39.129 ° N, 120.155 ° W]; elev. 1844 m; 31 Aug. 1974; EMEC • 1 ♀; Plumas Co., Meadow Valley; [39.937 ° N, 121.062 ° W]; 5 Jun. 1924; E. P. Van Duzee leg.; CAS • 2 ♀♀; Plumas Co., Spencer Lakes Road, 7 mi. SW of Johnsville; [39.6763 ° N, 120.7513 ° W]; 10 Sep. 1961; J. S. Buckett leg.; UCDC • 1 ♀; Riverside Co., 14 mi. SE of Upper Palm Canyon, Keen Camp Summit; [33.685 ° N, 116.704 ° W]; 8 Jul. 1964; E. I. Schlinger leg.; UCRC • 1 ♀; Riverside Co., Corona; [33.87 ° N, 117.57 ° W]; 9 Mar. 1972; ex Paeonia californica; EMEC • 2 ♀♀; same data as for preceding; 19 Mar. 1972; ex Paeonia californica; EMEC • 1 ♀; Riverside Co., Elsinore; [33.681 ° N, 117.345 ° W]; Baker leg.; USNM • 2 ♀♀; Riverside Co., Fern Valley; [33.759 ° N, 116.698 ° W]; elev. 1981.2 m; 25 Jul. 1965; E. I. Schlinger leg.; UCRC • 1 ♀; Riverside Co., Gavilan Hills; [33.7944 ° N, 117.3558 ° W]; elev. 620 m; 27 Apr. 1938; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 6 Apr. 1982; P. H. Timberlake leg.; ex Scrophularia; UCRC • 2 ♀♀; same data as for preceding; 7 Apr. 1939; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 6 Apr. 1956; P. H. Timberlake leg.; ex Salvia mellifera; UCRC • 1 ♀; same data as for preceding; 4 May 1938; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 2 ♀♀; same data as for preceding; 29 Mar. 1950; P. H. Timberlake leg.; ex Scrophularia heterophylla subsp. laciniata (verbatim: Scrophularia laciniata); UCRC • 1 ♀; Riverside Co., Idyllwild, San Jacinto Mountains; [33.748 ° N, 116.715 ° W]; 9 Jun. 1940; C. D. Michener leg.; ex Penstemon spectabilis; EMEC • 1 ♀; Riverside Co., Riverside; [33.9556 ° N, 117.3722 ° W]; elev. 290 m; 11 Jul. 1942; EMEC • 3 ♀♀; same data as for preceding; 3 Apr. 1928; P. H. Timberlake leg.; ex Scrophularia californica; MCZ • 1 ♀; same data as for preceding; 28 Apr. 1925; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 26 Mar. 1928; P. H. Timberlake leg.; ex Phacelia minor; UCRC • 1 ♀; same data as for preceding; 29 Mar. 1928; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; same data as for preceding; 21 Mar. 1926; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; same data as for preceding; 4 Mar. 1930; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; same data as for preceding; 20 Mar. 1939; P. H. Timberlake leg.; ex Phacelia minor; UCRC • 2 ♀♀; same data as for preceding; 20 Mar. 1934; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; same data as for preceding; 19 Mar. 1936; C. M. Dammers leg.; UCRC • 1 ♀; same data as for preceding; 27 Mar. 1929; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 22 Mar. 1938; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 30 Mar. 1938; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 2 ♀♀; same data as for preceding; 23 Mar. 1936; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 25 Mar. 1929; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 3 ♀♀; same data as for preceding; 16 Mar. 1928; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 30 Mar. 1926; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 21 Mar. 1926; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 2 ♀♀; same data as for preceding; 24 Mar. 1926; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 16 Mar. 1925; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 22 Mar. 1926; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 18 Mar. 1926; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 2 ♀♀; same data as for preceding; 2 Mar. 1925; P. H. Timberlake leg.; ex Scrophularia; UCRC • 1 ♀; same data as for preceding; 19 Mar. 1934; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 2 ♀♀; same data as for preceding; 10 Mar. 1936; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 2 ♀♀; same data as for preceding; 1 Mar. 1926; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 2 Mar. 1926; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 5 Mar. 1940; P. H. Timberlake leg.; ex Phacelia minor; UCRC • 1 ♀; same data as for preceding; 21 Apr. 1926; P. H. Timberlake leg.; ex Salvia polystachya; UCRC • 1 ♀; same data as for preceding; 17 Apr. 1939; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; same data as for preceding; 15 Apr. 1939; P. H. Timberlake leg.; ex Cryptantha intermedia; UCRC • 1 ♀; same data as for preceding; 29 Apr. 1937; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 3 ♀♀; same data as for preceding; 16 Apr. 1930; P. H. Timberlake leg.; ex Salvia mellifera; UCRC • 1 ♀; same data as for preceding; 29 Apr. 1929; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 26 Apr. 1935; P. H. Timberlake leg.; ex Salvia mellifera; UCRC • 1 ♀; same data as for preceding; 17 Apr. 1928; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 27 Apr. 1928; P. H. Timberlake leg.; ex Salvia apiana; UCRC • 1 ♀; same data as for preceding; 20 Apr. 1927; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 15 Apr. 1930; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 9 Apr. 1954; P. H. Timberlake leg.; ex Salvia; UCRC • 1 ♀; same data as for preceding; 10 Apr. 1926; P. H. Timberlake leg.; ex Salvia; UCRC • 1 ♀; same data as for preceding; 3 Apr. 1928; P. H. Timberlake leg.; ex Phacelia minor; UCRC • 1 ♀; same data as for preceding; 11 Apr. 1929; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 16 Apr. 1929; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 11 Apr. 1930; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 10 Apr. 1930; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 7 Apr. 1930; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 1 ♀; same data as for preceding; 8 Apr. 1936; P. H. Timberlake leg.; ex Acmispon glaber (verbatim: Lotus scoparius); UCRC • 1 ♀; same data as for preceding; 12 Apr. 1935; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 10 Apr. 1930; P. H. Timberlake leg.; ex Salvia columbariae (verbatim: Salvia columbarius); UCRC • 1 ♀; same data as for preceding; 9 Apr. 1934; P. H. Timberlake leg.; ex Salvia columbariae (verbatim: Salvia columbarius); UCRC • 1 ♀; same data as for preceding; 10 Apr. 1930; P. H. Timberlake leg.; ex Phacelia minor; UCRC • 1 ♀; same data as for preceding; 8 Apr. 1927; P. H. Timberlake leg.; ex Phacelia minor; UCRC • 1 ♀; same data as for preceding; 3 Apr. 1930; P. H. Timberlake leg.; ex Acmispon glaber (verbatim: Lotus scoparius); UCRC • 1 ♀; same data as for preceding; 2 Apr. 1930; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 2 Apr. 1936; P. H. Timberlake leg.; ex Acmispon glaber (verbatim: Lotus scoparius); UCRC • 1 ♀; same data as for preceding; 2 Apr. 1930; P. H. Timberlake leg.; UCRC • 1 ♀; same data as for preceding; 7 Apr. 1925; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 25 May 1929; P. H. Timberlake leg.; ex Calendula; UCRC • 1 ♀; same data as for preceding; 9 May 1926; P. H. Timberlake leg.; ex Melilotus indicus; UCRC • 1 ♀; same data as for preceding; 15 May 1926; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 15 May 1926; H. Compere leg.; UCRC • 1 ♀; same data as for preceding; 15 May 1928; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 19 May 1928; P. H. Timberlake leg.; ex Coreopsis; UCRC • 1 ♀; same data as for preceding; 6 May 1977; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 4 ♀♀; same data as for preceding; 22 Jul. 1927; P. H. Timberlake leg.; ex Canna; UCRC • 1 ♀; same data as for preceding; 19 Jun. 1939; P. H. Timberlake leg.; ex Scrophularia californica; UCRC • 1 ♀; same data as for preceding; 6 Apr. 1927; P. H. Timberlake leg.; ex Scrophularia californica; USNM • 1 ♀; same data as for preceding; 12 May 1934; P. H. Timberlake leg.; ex Calendula; EMEC • 1 ♀; same data as for preceding; 6 May 1934; P. H. Timberlake leg.; ex Scrophularia californica; EMEC • 1 ♀; Riverside Co., Tin Mine Canyon; [33.836 ° N, 117.610 ° W]; elev. 381 – 533.4 m; 6 Apr. 1968; E. I. Schlinger leg.; ex Salvia mellifera; UCRC • 5 ♀♀; Sacramento Co., 10 mi. NE of Folsom; [38.778 ° N, 121.044 ° W]; 10 Apr. 1960; M. Wasbauer leg.; ex Salvia sonomensis; CSCA • 1 ♀; Sacramento Co., Michigan Bar; [38.499 ° N, 121.045 ° W]; 19 Apr. 1961; M. E. Irwin leg.; UCDC • 2 ♀♀; same data as for preceding; 19 Apr. 1961; F. D. Parker leg.; UCDC • 1 ♀; Sacramento Co., Sacramento Valley; [39.000 ° N, 121.500 ° W]; 16 Jun. 1928; P. H. Timberlake leg.; ex Fremontodendron californicum (verbatim: Fremontia californica); UCRC • 1 ♀; San Benito Co., 5 mi. S of Bitterwater; [36.308 ° N, 121.001 ° W]; 2 Jul. 1954; C. W. O’Brien leg.; EMEC • 1 ♀; San Benito Co., Deep Creek; [34.271 ° N, 117.131 ° W]; 28 Apr. 1936; E. G. Linsley leg.; ex Lotus; EMEC • 1 ♀; San Benito Co., Idria (Gem Mine); [36.410 ° N, 120.670 ° W]; 29 Jun. 1955; P. D. Hurd leg.; ex Phacelia; EMEC • 2 ♀♀; San Benito Co., Lime Kiln Canyon, SW of Paicines; [34.273 ° N, 118.557 ° W]; 24 Apr. 1968; EMEC • 4 ♀♀; San Benito Co., Lime Kiln Road, 5 mi. SW of Paicines; [36.689 ° N, 121.344 ° W]; 24 Mar. 1966; EMEC • 1 ♀; San Bernardino Co., Camp Baldy; [34.169 ° N, 117.9261 ° W]; 12 Jun. 2016; EMEC • 1 ♀; San Bernardino Co., Lake Arrowhead; [34.248 ° N, 117.189 ° W]; 26 Jul. 1944; G. P. Mackenzie leg.; UCRC • 2 ♀♀; San Bernardino Co., Lytle Creek; 7 mi. N of Fontana; [34.195 ° N, 117.441 ° W]; elev. 1585 m; 17 Apr. 1965; EMEC • 1 ♀; San Bernardino Co., Mill Creek; [40.326 ° N, 121.523 ° W]; 16 May 1948; P. H. Timberlake leg.; ex Arctostaphylos patula; UCRC • 1 ♀; same data as for preceding; 4 May 1948; P. H. Timberlake leg.; ex Penstemon grinnellii; UCRC • 1 ♀; same data as for preceding; 1 Sep. 1947; P. H. Timberlake leg.; ex Erigeron divergens; UCRC • 1 ♂; same data as for preceding; 10 Sep. 1945; P. H. Timberlake leg.; ex Eriogonum; UCRC (Figs 57 – 58) • 1 ♀; San Bernardino Co., Redlands; [34.055 ° N, 117.183 ° W]; 1913; F. R. Cole leg.; USNM • 1 ♀; San Bernardino Co., San Bernardino Mountains; [34.130 ° N, 116.880 ° W]; elev. 1524 m; 19 Aug. 1934; P. H. Timberlake leg.; ex Canna; UCRC • 2 ♀♀; San Bernardino Co., San Bernardino Mountains Little Mojave Valley; [34.289 ° N, 116.654 ° W]; 6400; EMEC • 1 ♂; San Bernardino Co., San Bernardino Mountains, Camp O-Ongo near Running Springs; [34.208 ° N, 117.109 ° W]; 1 Sep. 1967; EMEC • 1 ♀; San Bernardino Co., Seven Oaks; [34.186 ° N, 116.910 ° W]; 28 Jul. 1953; A. L. Melander leg.; USNM • 1 ♀; San Bernardino Co., Victorville; [34.536 ° N, 117.291 ° W]; 5 May 1956; W. Wasbauer leg.; EMEC • 2 ♀♀; San Diego Co., Borrego; [33.247 ° N, 116.372 ° W]; 31 Mar. 1953; ex Cryptantha barbigera; EMEC • 2 ♀♀; San Diego Co., Harbison Canyon; [32.820 ° N, 116.830 ° W]; 9 May 1963; EMEC • 1 ♀; San Diego Co., La Mesa; [32.771 ° N, 117.023 ° W]; 2 Apr. 1953; F. X. Williams leg.; CAS • 1 ♀; San Diego Co., Ocean Beach; [32.750 ° N, 117.240 ° W]; 7 Apr. 2018; J. I. Carlson leg.; CAS • 1 ♂; San Diego Co., Palomar; 33.3333 ° N, 116.8934 ° W; 9 Jul. 2014; K. J. Hung leg.; ex Aquilegia formosa; SDMC • 1 ♀; San Diego Co., San Diego; [32.720 ° N, 117.160 ° W]; 27 Mar. 1933; F. X. Williams leg.; CAS • 1 ♀; San Diego Co., San Ysidro; [32.555 ° N, 117.044 ° W]; 29 Apr. 1945; P. G. Teach leg.; ex Rhus ovata; UCRC • 1 ♀; San Diego Co., Warner Springs, Agua Caliente Creek; [33.279 ° N, 116.720 ° W]; elev. 944.9 m; 23 – 25 Aug. 1980; M. Wasbauer and P. Adams leg.; CSCA • 1 ♀; same data as for preceding; 26 – 28 Aug. 1980; M. Wasbauer and P. Adams leg.; CSCA • 1 ♀; San Diego Co., San Felipe Creek; [35.940 ° N, 121.080 ° W]; 5 Jun. 1936; P. H. Timberlake leg.; ex Heliotropium curassavicum; UCRC • 1 ♀; San Luis Obispo Co., 10 mi. S of Simmler; [35.206 ° N, 119.986 ° W]; 3 May 1962; ex Eschscholtzia californica; EMEC • 1 ♀; San Luis Obispo Co., 10 mi. W of Simmler; [35.352 ° N, 120.164 ° W]; 3 May 1962; ex Eschscholtzia californica; EMEC • 1 ♂; San Luis Obispo Co., 5.1 mi. NE of Santa Margarita; [35.442 ° N, 120.545 ° W]; 8 Jun. 1962; P. D. Hurd leg.; EMEC • 1 ♀; San Mateo Co., Huddart Park on Kings Mountain Road (5 km W) Woodside; [37.440 ° N, 122.292 ° W]; 22 May 1973; P. H. Arnaud Jr leg.; CAS • 1 ♀; San Mateo Co., Jasper Ridge; [37.408 ° N, 122.227 ° W]; (?) May 1924; EMEC • 1 ♀; Santa Barbara Co., New Cuyama; [34.948 ° N, 119.689 ° W]; 28 Mar. 1957; EMEC • 1 ♀; Santa Barbara Co., Rancho San Marcos HQ, Santa Ynez Mountains; [34.600 ° N, 119.830 ° W]; 5 Jul. 1956; E. G. Linsley leg.; ex Phacelia grandiflora; EMEC • 1 ♀; Santa Barbara Co., Santa Barbara Botanic Garden; [34.457 ° N, 119.710 ° W]; 22 Aug. 1939; P. H. Timberlake leg.; UCRC • 1 ♀; Santa Barbara Co., Santa Cruz Island; [34.004 ° N, 119.726 ° W]; 18 May 1919; E. P. Van Duzee leg.; CAS • 2 ♀♀; Santa Clara Co., Alum Rock Park; [37.396 ° N, 121.802 ° W]; 16 May 1965; EMEC • 1 ♀; Santa Clara Co., Mountain Home Canyon; [37.121 ° N, 121.796 ° W]; 15 Oct. 1956; A. L. Melander leg.; USNM • 1 ♀; Santa Clara Co., Stanford University, W. o Quad; [37.436 ° N, 122.157 ° W]; 21 Mar. 1915; H. Morrison leg.; USNM • 2 ♀♀; Santa Clara Co., Del Puerto Canyon, 17.5 mi. W of Patterson; [37.466 ° N, 121.448 ° W]; 19 Jun. 1965; EMEC • 2 ♀♀; Santa Clara Co.; [37.29 ° N, 121.74 ° W]; EMEC • 1 ♀; Shasta Co., 4 mi. S of Hat Creek; [40.771 ° N, 121.514 ° W]; 4 Jun. 1941; EMEC • 1 ♀; Shasta Co., Hat Creek; [40.79 ° N, 121.474 ° W]; 5 Jun. 1941; C. D. Michener leg.; EMEC • 1 ♀; same data as for preceding; 4 Jun. 1941; C. D. Michener leg.; EMEC • 1 ♀; Sierra Co., Tamarack Lake; 41.545 ° N, 121.793 ° W; 19 Jul. 1931; E. O. Essig leg.; EMEC • 2 ♀♀; Sierra Co., Sierra; [39.573 ° N, 120.637 ° W]; 19 Jun. 2013; EMEC • 1 ♀; Siskiyou Co., 14 mi. WSW of Coffee Creek, Josephine Lake; [41.016 ° N, 122.946 ° W]; 15 Aug. 1977; S. Kuba leg.; CSCA • 1 ♀; Siskiyou Co., Ash Creek Ranger Station; 9 mi. E of McCloud, sandy area near stream; [41.2532 ° N, 121.965 ° W]; elev. 1066.8 m; 7 – 9 Jun. 1974; EMEC • 1 ♀; Siskiyou Co., Ash Creek Ranger Station; 9 mi. E of McCloud; [41.253 ° N, 121.965 ° W]; elev. 1066.8 m; 7 Jun. 1974; EMEC • 4 ♀♀; Siskiyou Co., Jct. Forest Service Roads 42 N 13 41 N 06, 14 air mi. NE of McCloud; [41.399 ° N, 121.947 ° W]; 10 Jun. 1974; EMEC • 1 ♀; Siskiyou Co., McBride Springs; [41.353 ° N, 122.282 ° W]; elev. 1585 m; 10 – 14 Jun. 1974; EMEC • 1 ♀; same data as for preceding; 10 Jun. 1974; EMEC • 1 ♀; Sonoma Co., Bear Flat; [38.598 ° N, 122.965 ° W]; elev. 1524 m; 23 Aug. 1920; P. H. Timberlake leg.; UCRC • 1 ♀; Stanislaus Co., Adobe Creek W; [37.396 ° N, 121.375 ° W]; 6 Apr. 1949; W. W. Middlekauff leg.; EMEC • 1 ♀; Stanislaus Co., Del Puerto Canyon; [37.488 ° N, 121.207 ° W]; 2 Apr. 1961; ex Phacelia; EMEC • 1 ♀; same data as for preceding; 20 Apr. 1949; P. D. Hurd leg.; EMEC • 1 ♀; Stanislaus Co., Knight’s Ferry; [37.820 ° N, 120.664 ° W]; 30 May 1961; EMEC • 1 ♀; Sutter Co., Wilson; [38.978 ° N, 121.626 ° W]; 5 Jun. 1915; H. Morrison leg.; USNM • 4 ♀♀; Tehama Co., Tehama; [40.024 ° N, 122.124 ° W]; 4862; EMEC • 1 ♀; Tulare Co., 10 mi. NW of California Hot Springs; [35.982 ° N, 118.803 ° W]; 14 May 1963; EMEC • 2 ♀♀; Tulare Co., 2 mi. E of Johnsondale; [35.975 ° N, 118.504 ° W]; 27 Apr. 1964; ex Arctostaphylos; EMEC • 1 ♀; Tulare Co., 3 mi. W of Three Rivers; [36.437 ° N, 118.958 ° W]; 1 Apr. 1939; P. H. Timberlake leg.; ex Phacelia distans; UCRC • 2 ♀♀; Tulare Co., 4 mi. N of Kaweah; [36.528 ° N, 118.918 ° W]; 13 May 1963; EMEC • 1 ♀; Tulare Co., Hot Springs; [35.880 ° N, 118.676 ° W]; 14 May 1963; EMEC • 1 ♀; Tulare Co., Mineral King Road; [36.444 ° N, 118.717 ° W]; 16 May 1970; ex Collinsia heterophylla; EMEC • 1 ♀; same data as for preceding; 21 May 1970; ex Collinsia heterophylla; EMEC • 1 ♀; same data as for preceding; 17 May 1970; EMEC • 1 ♀; same data as for preceding; 15 May 1970; ex Collinsia heterophylla; EMEC • 1 ♀; Tulare Co., Potwisha, Sequoia National Park; [36.515 ° N, 118.801 ° W]; elev. 609.6 – 1524 m; 13 Jun. 1929; E. C. Van Dyke leg.; CAS • 1 ♀; Tuolumne Co., 3 mi. NE of Tuolumne City; [37.931 ° N, 120.202 ° W]; 23 Mar. 1966; EMEC • 1 ♀; Tuolumne Co., 3 mi. SE of Tuolumne City; [37.931 ° N, 120.202 ° W]; 23 Mar. 1966; EMEC • 1 ♀; Tuolumne Co., Pinecrest; [38.189 ° N, 119.991 ° W]; 4 Aug. 1948; P. D. Hurd and J. W. MacSwain leg.; EMEC • 1 ♀; Tuolumne Co., Deadman Campground; [38.317 ° N, 119.750 ° W]; 6 Jun. 1960; W. A. Steffan leg.; EMEC • 1 ♀; Ventura Co., Calabasas Peak (1 mi. NW) Santa Monica Mountains; 15 Mar. 1971; EMEC • 2 ♀♀; Ventura Co., Hungry Valley; 5 mi. S of Gorman; [34.728 ° N, 118.866 ° W]; 6 May 1959; J. Powell leg.; ex Penstemon centranthifolius; EMEC • 2 ♀♀; Ventura Co., Hungry Valley; 5 mi. S of Gorman; [34.730 ° N, 118.866 ° W]; 6 May 1959; ex Ericameria cooperi (verbatim: Haplopappus cooperi); EMEC • 1 ♀; same data as for preceding; 4 May 1959; EMEC • 1 ♀; same data as for preceding; 4 May 1959; ex Ericameria cooperi (verbatim: Haplopappus cooperi); EMEC • 1 ♀; Ventura Co., Quatal Canyon; [34.837 ° N, 119.360 ° W]; 9 May 1959; EMEC • 1 ♀; same data as for preceding; 9 May 1959; ex Penstemon centranthifolius; EMEC • 1 ♀; Ventura Co., Quatal Canyon, NW corner Ventura; [34.831 ° N, 119.351 ° W]; 9 May 1959; ex Encelia actoni; EMEC • 1 ♀; same data as for preceding; 9 May 1959; ex Penstemon centranthifolius; EMEC • 1 ♀; Ventura Co., Santa Paula; [34.356 ° N, 119.068 ° W]; 6 Mar. 1959; W. E. Simonds leg.; CSCA • 2 ♀♀; Ventura Co., Wegis Ranch; 9 May 1963; EMEC • 1 ♀; Lassen National Park; [40.498 ° N, 121.428 ° W]; 3 Jun. 1941; C. D. Michener leg.; EMEC (Fig. 56 A – C) • 1 ♀; Sierra National Forest; [39.573 ° N, 120.637 ° W]; elev. 1920.2 m; 28 Jun. 1956; USNM. – Colorado • 1 ♀; Boulder Co., 1 mi. W of Gold Hill; [40.063 ° N, 105.429 ° W]; 4 Jun. 1963; U. N. Lanham and B. Vogel leg.; ex Esula; UCMC • 1 ♂; Boulder Co., Nederland, Ward; [40.070 ° N, 105.510 ° W]; 29 Aug. 1959; Ellen Ordway leg.; SEMC • 1 ♀; Jackson Co., Cowdrey; [40.858 ° N, 106.312 ° W]; elev. 2529.8 m; 22 Jun. 1961; G. L. Bush leg.; MCZ • 1 ♀; Mineral Co., Creede; [37.849 ° N, 106.925 ° W]; elev. 2743.2 m; 1 – 31 Jul. 1907; F. M. Carpenter leg.; MCZ • 1 ♂; Saguache Co., Valley View Springs, 7 mi. E of Mineral Hot Springs on W foot of Sangre de Cristo range; [38.160 ° N, 105.795 ° W]; elev. 2590.8 m; 13 Aug. 1965; H. B. Leech leg.; CAS • 1 ♀; Teller Co., Big Spring Ranch, Florissant; [38.940 ° N, 105.290 ° W]; 20 Jun. 1962; EMEC • 1 ♂; same data as for preceding; 30 Jul. 1966; EMEC • 1 ♂; Teller Co., South base Blue Mountain near Florissant; elev. 2568.2 m; 1 Aug. 1966; EMEC. – Idaho • 1 ♂; Latah Co., Moscow Mountains; [46.804 ° N, 116.869 ° W]; elev. 914.4 m; 27 Jul. 1925; C. L. Fox leg.; CAS • 1 ♀; Lemhi Co., 5 mi. N of Gibbonsville; [45.625 ° N, 113.927 ° W]; 21 Jul. 1983; J. D. Pinto and D. R. Pinto leg.; UCRC • 1 ♀; craters of the Moon National Monument and Preserve; [43.172 ° N, 113.482 ° W]; 16 Aug. 1964; Blaine, Butte, Lincoln, Minidoka and Power leg.; ex columbine (Aquilegia); EMEC. – Montana • 1 ♂; Gallatin Co., Madison River, Gallatin National Forest, below Hebgen Dam or reservoir; [45.513 ° N, 111.592 ° W]; 28 Aug. 1935; P. Wible leg.; CMNH. – Nevada • 2 ♀♀; Elko Co., Spruce Mountain; [40.552 ° N, 114.821 ° W]; elev. 2590.8 m; 24 Jun. 1956; EMEC • 2 ♀♀; Eureka Co., 3 mi. W of Eureka; [39.545 ° N, 116.016 ° W]; 24 Jun. 1968; EMEC • 1 ♀; Fresno Co., Huntington Lake; [37.236 ° N, 119.179 ° W]; elev. 2133.6 m; 7 Jul. 1919; E. P. Van Duzee leg.; CAS • 2 ♀♀; Gooding Co., Wood River, 1 mi. NE of Gooding; [42.948 ° N, 114.699 ° W]; 6 – 7 Jul. 1980; M. S. Wasbauer and K. M. Wasbauer leg.; CSCA • 1 ♀; Harney Co., Alder Springs Campground McKenzie Pass; [44.178 ° N, 121.911 ° W]; elev. 1104.9 m; 30 Jun. 1927; H. A. Scullen leg.; OSAC • 1 ♀; Huerfano Co., 12 mi. N of Gardner; [37.906 ° N, 105.156 ° W]; elev. 3048 m; 5 Jul. 1975; U. N. Lanham leg.; ex Castilleja miniata; UCMC • 1 ♀; Mesa Co., 19 mi. NE of Gateway; [38.862 ° N, 108.729 ° W]; 18 May 1963; C. J. McCoy and B. Vogel leg.; UCMC • 1 ♀; Ormsby Co.; [39.16 ° N, 119.77 ° W]; 6 Jul.; Baker leg.; USNM • 2 ♀♀; same data as for preceding; Jul.; Baker leg.; USNM • 1 ♂; White Pine Co., 22 mi. W of Ely; [39.251 ° N, 115.285 ° W]; 20 Jul. 1958; E. G. Linsley and J. W. MacSwain leg.; EMEC. – New Mexico • 2 ♀♀; Catron Co., 8 mi. SE of Lunc; elev. 2286 m; 9 – 14 Jul. 1979; S. Peck and J. Peck leg.; PMAE • 2 ♀♀; San Miguel Co., Pecos; [35.581 ° N, 105.678 ° W]; Jul. 07 ???; EMEC • 1 ♀; no place; 23 Jun. 1908; W. T. Cockerell leg.; ex Castilleja integra; USNM. – Oregon • 1 ♀; Columbia Co., 20 mi. SW of St Helens; [45.650 ° N, 123.117 ° W]; 21 May 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♀; Umatilla Co., Cayuse; [45.676 ° N, 118.574 ° W]; 13 May 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♀; Jackson Co., Chimney Gulch; [44.420 ° N, 118.993 ° W]; N. Banks leg.; MCZ • 1 ♀; Klamath Co., Crater Lake; [42.950 ° N, 122.100 ° W]; 16 Sep. 1934; A. L. Melander leg.; MCZ • 1 ♀; Klamath Co., Eagle Ridge County Park; [42.405 ° N, 121.950 ° W]; 20 May 1924; C. L. Fox leg.; CAS • 1 ♀; same data as for preceding; 24 Jun. 1924; C. L. Fox leg.; CAS • 1 ♀; Clackamas Co., Estacada; [45.290 ° N, 122.334 ° W]; 30 Jun. 1930; R. M. Scullen leg.; OSAC • 1 ♀; Wallowa Co., Joseph; [45.353 ° N, 117.23 ° W]; 6 Jun. 1970; OSAC • 1 ♀; Jackson Co., Little Squaw Lake (7 mi. E) Copper; [42.037 ° N, 123.025 ° W]; elev. 975.4 m; 22 May 1964; E. Copper leg.; OSAC • 1 ♀; Gilliam Co., Mayville; [45.080 ° N, 120.190 ° W]; 1 – 30 Jun. 1918; CAS • 4 ♀♀; Lake Co., Summer Lake; [42.830 ° N, 120.750 ° W]; 16 Jun. 1938; K. Gray, J. Schuh leg.; OSAC • 1 ♀; Umatilla Co., Weston; [45.816 ° N, 118.426 ° W]; 29 Apr. 1938; K. Gray leg.; ex mustard; OSAC. – South Dakota • 1 ♀; Lawrence Co., Spearfish Creek, T 3 N R 1 E, S 6; [44.41 ° N, 103.894 ° W]; 28 Jun. 1965; R. W. Hodges leg.; USNM. – Utah • 1 ♀; Beaver Co., Beaver Valley; [38.276 ° N, 112.639 ° W]; USNM • 1 ♀; Kane Co., Bryce Canyon; [37.617 ° N, 112.142 ° W]; 21 Jun. 1933; O. H. Swezey leg.; CAS • 1 ♀; Garfield Co., Calf Creek; [37.822 ° N, 111.418 ° W]; elev. 1645.9 – 1676.4 m; 28 – 29 Jul. 1982; A. Menke and K. Menke leg.; USNM • 2 ♀♀; Summit Co., Park City; [40.650 ° N, 111.501 ° W]; 8 Jul. 1922; E. P. Van Duzee leg.; CAS. – Washington • 1 ♀; Yakima Co., 7 km W of Naches, Hwy 12; 46.7367 ° N, 120.79103 ° W; 6 Aug. 2007; J. Gibbs and C. Sheffield leg.; PCYU • 1 ♀; King Co., Seattle; [47.610 ° N, 122.330 ° W]; 10420; OSAC • 1 ♀; same data as for preceding; 24 Apr. 1994; UCMC • 1 ♀; same data as for preceding; 5574; UCMC • 1 ♀; same data as for preceding; 20 Apr. 1998; ex Rubus ursinus; UCMC • 1 ♀; same data as for preceding; 25 Jun. 1997; UCMC • 1 ♀; same data as for preceding; 4540; UCMC • 1 ♀; same data as for preceding; 5254; UCMC • 1 ♀; same data as for preceding; 2733; UCMC • 1 ♀; same data as for preceding; 3765; UCMC • 1 ♀; same data as for preceding; 3775; UCMC • 1 ♀; same data as for preceding; 10 Aug. 1999; ex Anaphalis; UCMC • 1 ♀; same data as for preceding; 5956; UCMC • 1 ♀; same data as for preceding; 9 Apr. 1997; UCMC • 1 ♀; same data as for preceding; UCMC • 1 ♀; same data as for preceding; 10416; ex Crepis capillaris (verbatim: Crepis virens); UCMC • 1 ♀; same data as for preceding; 4898; UCMC • 1 ♀; same data as for preceding; 21 May 1999; ex Rubus ursinus; UCMC • 1 ♀; Skamania Co., Hanleys, Mount St Helens; [46.2 ° N, 122.186 ° W]; 3 May 1946; T. O. Thatcher leg.; EMEC • 1 ♀; Yakima Co., Yakima; [46.602 ° N, 120.508 ° W]; 18 Jun. 1955; H. P. Lanchester leg.; USNM. – Wyoming • 1 ♀; Teton Co., Grand Teton National Park Jenny Lake; [43.775 ° N, 110.72 ° W]; E. C. Van Dyke leg.; CAS • 2 ♀♀; Teton Co., Teton National Park, String Lake; [43.789 ° N, 110.732 ° W]; 18 Jun. 1963; C. A. Toschi leg.; EMEC • 6 ♀♀; Lincoln Co.; 41.865 ° N, 110.7653 ° W; 20 – 21 Sep. 2011; BIML • 1 ♀; Lincoln Co.; 41.8704 ° N, 110.7772 ° W; 13 May 2011; BIML • 1 ♂; Lincoln Co.; 41.865 ° N, 110.7653 ° W; 20 – 21 Sep. 2011; BIML.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF88FF41FE17FEE2FAA3FD73.taxon	description	Description GENITALIA. (Figs 58, 120 J). Gonostylus elongate, about 2 times as long as broad, with long setae. Retrorse lobe slender, about 5 times as long as broad. S 7 apical lobe broadly rounded.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF88FF41FE17FEE2FAA3FD73.taxon	distribution	Range Canada (Alberta, British Columbia) and USA (California, Colorado, Idaho, Montana, Nevada, New Mexico, Oregon, South Dakota, Utah, Washington) (Fig. 59).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF88FF41FE17FEE2FAA3FD73.taxon	biology_ecology	Floral hosts Cockerell (1903) recorded this species as a habitual visitor of Castilleja integra A. Gray (family Scrophulariaceae) in New Mexico. ANACARDIACEAE: Rhus ovata S. Watson • ASTERACEAE: Anaphalis sp., Baccharis sp. Calendula sp., Coreopsis sp., Crepis capillaris (L.) Wallr., Encelia actonii Elmer, Ericameria cooperi H. M. Hall, Ericameria parishii (Greene) H. M. Hall, Erigeron divergens Torr. & A. Gray, Eriogonum wrightii var. subscaposum S. Watson, Hazardia squarrosa (Hook. & Arn.) Greene, Senecio sp., Solidago velutina subsp. californica (Nutt.) Semple, Wyethia sp. • BORAGINACEAE: Cryptantha barbigera (A. Gray) Greene, Cryptantha intermedia (A. Gray) Greene, Cryptantha sp., Heliotropium curassavicum L., Lithospermum ruderale Douglas ex Lehm. • BRASSICACEAE: Brassica sp., Brassica nigra (L.) W. D. J. Koch • CANNACEAE: Canna sp. • ERICACEAE: Arctostaphylos sp., Arctostaphylos patula Greene • FABACEAE: Acmispon glaber (Vogel) Brouillet, Lotus sp., Melilotus indicus (L.) All. • HYDROPHYLLACEAE: Nemophila sp., Phacelia cicutaria var. hispida (A. Gray) J. T. Howell, Phacelia distans Benth., Phacelia grandiflora (Benth.) A. Gray, Phacelia imbricata Greene, Phacelia minor (Harv.) Thell., Phacelia platyloba A. Gray, Phacelia sp., Phacelia tanacetifolia Benth. • LAMIACEAE: Salvia apiana Jeps., Salvia columbariae Benth., Salvia mellifera Greene, Salvia sp., Salvia sonomensis Greene, Stachys bullata Benth. • LOASACEAE: Eucnide urens Parry ex Coville • NYCTAGINACEAE: Mirabilis laevis var. laevis • ONAGRACEAE: Clarkia rhomboidea Douglas • PAEONIACEAE: Paeonia brownii Douglas ex Hook., Paeonia californica Nutt. • PAPAVERACEAE: Eschscholzia californica Cham. • POLYGONACEAE: Eriogonum sp. • RANUNCULACEAE: Aquilegia formosa Fisch. ex DC., Aquilegia sp. • ROSACEAE: Chamaebatia foliolosa Benth., Photinia sp., Rubus ursinus Cham. & Schltdl., Rubus vitifolius Cham. & Schltdl. • SALICACEAE: Salix sp. • SCROPHULARIACEAE: Castilleja affinis Hook. & Arn., Castilleja applegatei var. pinetorum (Fernald) N. H. Holmgren, Castilleja integra A. Gray, Castilleja miniata Douglas ex Hook., Collinsia heterophylla Buist ex Graham, Cordylanthus pilosus A. Gray, Cordylanthus rigidus subsp. rigidus, Keckiella cordifolia (Benth.) Straw, Keckiella corymbosa (Benth. ex A. DC.) Straw, Keckiella lemmonii (A. Gray) Straw, Keckiella ternata (Torr. ex A. Gray) Straw, Penstemon centranthifolius (Benth.) Benth., Penstemon grinnellii Eastw., Penstemon heterodoxus A. Gray, Penstemon spectabilis Thurb. ex A. Gray, Scrophularia heterophylla subsp. laciniata (Waldst. & Kit.) Maire & Petitm., Scrophularia sp., Scrophularia villosa Pennell, Triphysaria versicolor subsp. faucibarbata (A. Gray) T. I. Chuang & Heckard • SOLANACEAE: Datura inoxia Mill. • STERCULIACEAE: Fremontodendron californicum (Torr.) Coville. Biological notes Lasioglossum ovaliceps is a ground-nesting species, based on observations of females digging in a clay bank is Pasadena, California in March (Hicks 1936).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF88FF41FE17FEE2FAA3FD73.taxon	discussion	DNA barcodes Ten full DNA barcodes and three partial barcodes were analyzed, from Alberta, British Columbia, Washington, Oregon, Utah, and California. They form a discrete cluster with little intraspecific variation.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF7CFF42FE0FFD5DFE38FD4E.taxon	diagnosis	Diagnosis The L. pectorale species group can be diagnosed by a combination of characters in both females and males. Females are consistently black bees (no orange metasoma); exhibit a completely rugose or rugulose propodeum, propodeal triangle with strong to moderately strong rugae reaching all the way to the posterior margin; possess an apical carina on the propodeal triangle posterior margin, which forms a sharp edge; and T 2 – T 3 with distinct basolateral tomentum. Males in this group do not exhibit any clypeal maculations (in contrast to L. fedorense species group, L. pectinatum species group, L. subobscurum, and most of other species in this subgenus); have short, female-like antennae (in contrast to L. nitidiusculum species group); usually “ normal ” (unmodified) heads, meaning that no broad clypeus, no genal tuberculation, and no extremely elongated mandibles, (in contrast to modifications present in males of L. lustrans, L. fedorense species group, L. pectinatum species group, or L. arizonense species group). Comparison with related species and exceptions Some females of this group show a strongly coriarious anterior face of T 1 with a well-developed fan of appressed hairs. But this character is absent in most species in this group, like L. kincaidii, L. nelumbonis, L. hewetti, L. pectorale, L. pectoraloides, and L. pseudopectorale. Lasioglossum lustrans is included in the L. pectorale species group despite lacking an apical carina and a completely rugose propodeum. An exception is L. lustrans, in which the male head is modified. Some species have one or more of the above traits but, except for L. lustrans, are not included in the pectorale group due to missing other key characters or having conflicting features. Notable comparisons and exceptions include: 1) L. opata sp. nov.: females of L. opata do exhibit an apical carina (faint), however, in L. opata the rugae of the propodeal triangle do not reach the posterior margin, and the propodeum is not completely rugose. 2) L. swenki: this species is somewhat problematic to classify. Females of L. swenki have an apical carina, and in some specimens the propodeal rugae reach the posterior margin. However, the males of L. swenki do not have the normal heads.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF7CFF42FE0FFD5DFE38FD4E.taxon	discussion	Taxonomic remarks The pectorale species group was previously treated in part as the L. robertsoni species group by Gibbs et al. (2013), but given the rarity and limited representation of L. robertsoni, we here propose using L. pectorale – a more commonly collected species – as the nominal representative. The pectorale species group is proposed as a practical working unit within this revision. Although it may not represent a strictly monophyletic lineage, the group provides a useful framework for comparing species that share a distinctive suite of morphological traits. Several included and excluded taxa remain taxonomically problematic, and the boundaries of the group are recognized as provisional. Nevertheless, defining this assemblage facilitates clearer discussion and future tracking of related species as additional morphological and molecular data become available.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF7FFF49FE36FD39FD64FDE2.taxon	description	urn: lsid: zoobank. org: act: 57 C 32036 - 1647 - 4 F 82 - 903 F-A 99297 DF 983 D Figs 60 – 63, 104 E, 105 A, 106 D, 109 D, 113 B, 114 B, 120 K	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF7FFF49FE36FD39FD64FDE2.taxon	diagnosis	Diagnosis Female L. angusticulum sp. nov. can be recognised by the combination of head long, length / width ratio 1.06 (± 0.03 SD) (remarkably long in L. pathiranae sp. nov.); mesoscutum mostly polished (rough in L. robertsoni), with distinct double punctation (no double punctation in L. angustius and L. robertsoni), sparse (IS = 1 – 3 PD) puncture density submedially (denser in L. angustius, L. pathiranae and L. robertsoni); rugae of the propodeal triangle reaching posterior margin (separating from species in which the rugae do not reach the posterior margin), propodeal triangle posterior margin sharply edged with a less prominent apical carina (this excludes all species with a smoothly rounded posterior margin; apical carina is prominently raised in L. pathiranae sp. nov. and L. robertsoni); T 1 anterior face mostly polished with broad dorsal opening in the fan (T 1 anterior face strongly coriarious with a narrower dorsal opening in the fan in L. robertsoni, L. pathiranae, L. angustius and L. opata sp. nov.); and T 2 – T 3 with distinct basolateral tomentum (separating it from species lacking basal patches or bands of tomentum on T 2 – T 3, such as L. nitidiusculum group). Males L. angusticulum sp. nov. can be recognised by the combination of head long, length / width ratio 1.08 (± 0.01 SD) (remarkably long in L. pathiranae sp. nov.); mesoscutum mostly polished and distinctly diversopunctate (indistinct in L. pathiranae), with moderately sparser (IS = 1 – 2 PD) interspaces submedially (denser in L. pathiranae); rugae of the propodeal triangle reaching posterior margin (separating from species in which the rugae do not reach the posterior margin); mesepisternum reticulate-rugose (finely rugulose in L. synthyridis); and propodeal triangle with apical carina indistinct or absent (prominently raised in L. pathiranae).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF7FFF49FE36FD39FD64FDE2.taxon	etymology	Etymology The specific epithet angusticulum is a diminutive form derived from the Latin ‘ angustius ’ (meaning ‘ narrow’), referencing Lasioglossum angustius, a species with which this new species was previously confused. The name acknowledges the historical taxonomic confusion between the two species.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF7FFF49FE36FD39FD64FDE2.taxon	materials_examined	Material examined Holotype UNITED STATES – Arizona • ♀; Pima Co., Catalina Mountains; [32.430 ° N, 110.705 ° W]; 21 Jul. 1942; E. C. Van Dyke leg.; CAS (Fig. 60). Specimen labels “ Catalina Mts / Ariz / VII- 21 - 42 // Presented by / EC VanDyk / Collector // Lasioglossum / GIBBS- 14439 // HOLOTYPE / Lasioglossum (Hemihalictus) angusticulum Hettiarachchi and Gibbs ”. Paratypes UNITED STATES – Arizona • 1 ♀; Cochise Co., Chiricahua Mountains, 3.5 mi. SW of Portal; [31.878 ° N, 109.184 ° W]; 13 Aug. 1952; H. B. Leech and J. W. Green leg.; CAS • 1 ♀; Coconino Co., 9 mi. N of Jacob Lake; [36.844 ° N, 112.217 ° W]; 11 Jun. 1966; J. M. Davidson and M. A. Cazier leg.; ex Sphaeralcea laxa; ASUHIC • 3 ♀♀; Coconino Co., Oak Creek Canyon; [34.913 ° N, 111.727 ° W]; F. H. Snow leg.; USNM • 1 ♀; Gila Co., Pioneer Pass; [33.270 ° N, 110.802 ° W]; 18 Aug. 1958; L. A. Stange leg.; LACM • 1 ♀; Gila Co., Young; [34.101 ° N, 110.964 ° W]; 20 Jul. 1996; N. F. Gunz leg.; RSKM • 2 ♀♀; Mojave Co., Hualapai Mountains, S of Kingman; [34.900 ° N, 113.884 ° W]; 30 Jul. 1965; M. E. Irwin leg.; UCRC • 1 ♀; same data as for preceding; 30 Jul. 1965; E. I. Schlinger leg.; UCRC • 1 ♀; Navajo Co., 5 mi. S of McNary; [34.000 ° N, 109.857 ° W]; 22 Jun. 1957; F. G. Werner and G. D. Butler leg.; ASUHIC • 1 ♀; Navajo Co., Kaventa; [36.722 ° N, 110.254 ° W]; 20 Jun. 1933; H. N. Hultgren leg.; CAS • 1 ♂; Navajo Co., 14 mi. S of Show Low; [34.050 ° N, 110.030 ° W]; 24 Jun. 1956; G. D. Butler, Gerhardt leg.; ASUHIC (Fig. 61) • 2 ♀♀; Navajo Co., 19 mi. SW of Kayenta; [36.531 ° N, 110.497 ° W]; 20 Jun. 1933; H. N. Hultgren leg.; CAS • 1 ♀; Pima Co., Catalina Mountains; [32.430 ° N, 110.705 ° W]; 31 May 1954; G. D. Butler leg.; ex Houstonia wrightii; ASUHIC • 1 ♂; Pima Co., Santa Catalina Mountains, Summerhaven; [32.445 ° N, 110.761 ° W]; 20 Jul. 1942; R. S. Beal leg.; UCRC • 1 ♀; Pima Co., Catalina Mountains; [32.430 ° N, 110.705 ° W]; 21 Jul. 1942; E. C. Van Dyke leg.; CAS • 1 ♂; same data as for preceding; CAS (Fig. 62) • 3 ♀♀; same data as for preceding; 20 Jul. 1942; E. C. Van Dyke leg.; CAS • 1 ♀; Pima Co., Catalina Mountains South; [32.375 ° N, 110.818 ° W]; 1 – 31 Jul. 1936; E. S. Ross leg.; CAS • 1 ♀; Pima Co., Bear Wallow, Station Catalina Mountains; [32.424 ° N, 110.740 ° W]; 8 Jun. 1961; R. H. Painter and E. M. Painter leg.; ASUHIC • 1 ♀; Pima Co., Catalina Mountains; [32.430 ° N, 110.705 ° W]; 30 Apr. 1953; G. D. Butler leg.; ex Thistle; ASUHIC • 1 ♀; same data as for preceding; 25 Jul. 1954; G. D. Butler leg.; ASUHIC • 1 ♀; Pima Co., Bear Wallow, Station Catalina Mountains; [32.424 ° N, 110.740 ° W]; 5 Jun. 1961; F. Werner leg.; ASUHIC • 1 ♀; Pima Co., Catalina Mountains, Hitchcock Hwy 22 mi; [32.446 ° N, 110.759 ° W]; 18 Jul. 1955; F. G. Werner and G. D. Butler leg.; ex yellow composite; ASUHIC • 1 ♀; Pima Co., Catalina Mountains, Hitchcock Hwy 23 mi; [32.444 ° N, 110.760 ° W]; 18 Jul. 1955; F. G. Werner and G. D. Butler leg.; ASUHIC • 1 ♂; Pima Co., Bear Wallow, Station Catalina Mountains; [32.424 ° N, 110.740 ° W]; 8 Jun. 1961; J. C. Bequaert leg.; ex Senecio; ASUHIC • 1 ♂; same data as for preceding; 5 Jun. 1961; F. G. Werner leg.; ex Senecio; ASUHIC • 1 ♂; Pima Co., Catalina Mountains; [32.430 ° N, 110.705 ° W]; 31 May 1954; G. D. Butler leg.; ASUHIC • 1 ♂; same data as for preceding; 31 May 1954; G. D. Butler leg.; ex Houstonia wrightii; ASUHIC • 1 ♂; same data as for preceding; 9 Jun. 1958; ASUHIC • 1 ♀; Pima and Pinal Co., Santa Catalina Mountains; [32.443 ° N, 110.788 ° W]; 20 Jul. 1942; H. A. Scullen leg.; SDMC • 3 ♀♀; Yavapai Co., 5.7 mi. S of Prescott; [34.460 ° N, 112.469 ° W]; 7 Jun. 1966; J. M. Davidson and M. A. Cazier leg.; ASUHIC. – Colorado • 1 ♀; Boulder Co., Boulder; [40.015 ° N, 105.2705 ° W]; 14 Aug. 1971; U. N. Lanham leg.; UCMC • 3 ♀♀; Boulder Co., 2 mi. SE of Eldorado Springs; [39.915 ° N, 105.252 ° W]; 18 Jun. 1963; B. Vogel and U. N. Lanham leg.; UCMC • 1 ♀; Boulder Co., Boulder; [40.015 ° N, 105.271 ° W]; elev. 1828.8 m; 31 May 1961; B. H. Poole leg.; CNC • 1 ♀; same data as for preceding; 16 Jun. 1961; B. H. Poole leg.; CNC • 1 ♀; Boulder Co., 4 mi. NW of Boulder; [40.053 ° N, 105.324 ° W]; elev. 2103.1 m; 8 Jun. 1961; B. H. Poole leg.; CNC • 1 ♀; Delta Co., Paonia; [38.868 ° N, 107.592 ° W]; 14 Jun. 1926; E. C. Van Dyke leg.; CAS • 1 ♀; Larimer Co.; 40.6833 ° N, 105.3975 ° W; 11 – 12 May 2004; S. W. Droege leg.; PCYU • 1 ♀; Mesa Co.; 39.0295 ° N, 108.6276 ° W; C. Stragar leg.; WRME. – New Mexico • 1 ♀; Bernalillo Co., Albuquerque; [35.084 ° N, 106.651 ° W]; 1 Jul. 1945; W. O. Griesel leg.; LACM. – Texas • 1 ♀; Culberson Co., Pine Springs; [31.908 ° N, 104.826 ° W]; 26 Apr. 1969; U. N. Lanham leg.; UCMC. – Utah • 1 ♀; Washington Co., 3 mi. E of Pine Valley, Juniper Campground; [37.391 ° N, 113.459 ° W]; 4 Jun. 1979; R. Hardy leg.; CSCA.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF7FFF49FE36FD39FD64FDE2.taxon	description	Description Female BODY MEASUREMENTS. Length 7.79 mm (± 0.73 SD). Head length 1.98 mm (± 0.08 SD). Face length 1.64 mm (± 0.08 SD). Head width 1.86 mm (± 0.10 SD). Wing length 5.41 mm (± 0.25 SD). n = 6. COLOURATION. Head and mesosoma black. Clypeus apex dark brown. Labrum dark brown. Mandible dark brown basally, becoming light brown medially and slightly reddish brown apically. Flagellum brown. Pronotal lobe dark brown. Tegula light brown. Wing membrane hyaline, veins with subcosta brown, otherwise light brown. Legs dark brown to brown. Metasoma dark brown or brown with rims of terga, sterna and downcurved lateral areas of terga narrowly ochreous. PUBESCENCE. Body hair colour off-white. Tomentum dense on pronotal angle, and lobe, space between pronotal lobe and tegula, metanotum anteromedially, propodeum lateral surface anterodorsally, and T 2 basolaterally, and T 3 basally; moderately dense on anterolateral angle and marginally of mesoscutum; very sparse on supraclypeal area dorsolaterally, paraocular area, gena near eye margin, propodeum lateral (except anterodorsal area) and posterior surface mostly dorsally. Mesosoma covered by appressed, simple, very short hairs (<0.25 OD) mixed with sparse, erect, weakly plumose, long hairs (<1.5 OD) on mesoscutum and erect, plumose, long hairs (<2 OD) on other areas. Metasomal sterna with moderately sparse, erect, simple to weakly plumose, long (<4 OD) hairs. Metafemoral scopa with dense plumose hairs. Wing hairs brown, very short and sparse. T 1 with fan, broad dorsal opening (3 – 4 OD), composed of overlapping sub-appressed simple hairs (<1.5 OD). SURFACE SCULPTURING. Clypeus imbricate to tessellate, becoming polished apical half; punctures irregularly sparse (IS <3 PD) apically and medially, becoming smaller and denser (IS <1 PD) basolaterally. Supraclypeal area imbricate or tessellate, becoming weakly so or polished medially; punctures dense (IS <1 PD), becoming moderately dense (IS = 1 – 2 PD) medially, diversopunctate. Frons reticulate-punctate; punctures crowded (IS = 0 PD). Ocellocular area polished; punctures sparse (IS = 1 – 3 PD) and diversopunctate posteriorly, becoming dense (IS <1 PD) anteriorly. Gena polished, becoming lineolate posteriorly; punctures dense (IS <1 PD), becoming obscure posteriorly. Postgena polished, weakly imbricate adjacent to hypostomal carina. Tegula broadly impunctate, minute punctures present anteriorly and basally, sparse (IS = 1 – 4 PD). Mesoscutum polished, becoming imbricate anteromedially and on median line; punctures moderately dense (IS = 1 – 2 PD), becoming sparse (IS = 1 – 3 PD) mesad of parapsidal line and anteromedially, distinctly doubly punctate. Mesoscutellum polished; punctures dense (IS ≤ 1 PD) marginally and on median line, sparse submedially (IS = 1 – 5 PD), distinctly diversopunctate. Metanotum rugulose, punctures obscure. Propodeal triangle with rugae strong, reaching posterior margin and even entering to propodeum posterior face, anastomosing medially and weakly so laterally; lateral rugae evidently extending to dorsolateral slope of propodeum lateral face; interspace sculpture shiny to weakly imbricate. Preëpisternum reticulate-rugose. Hypoepimeron rugulose; punctures crowded (IS = 0 PD), indistinct. Mesepisternum reticulate-rugose, becoming rugose ventrally. Propodeum lateral face strongly imbricate, becoming rugulose posteriorly; posterior face rugulose. T 1 anterior face mostly polished, but coriarious laterally, punctures present laterally, sparse (IS = 1 – 4 PD); dorsal surface polished, punctures sparse (IS = 1 – 3 PD), becoming sparser towards premarginal line and sometime on median line (IS = 1 – 5 PD) and in small subapicolateral boss (IS = 2 – 6 PD), diversopunctate, punctures moderately sparse (IS = 1 – 2 PD) on rim, becoming impunctate apical half and broadly impunctate medially on rim. T 2 disc polished, but somewhat weakly coriarious subapicolateral boss apically and on premarginal line, punctures moderately sparse (IS = 1 – 2 PD) basally and sometime moderately dense (IS = 1 – 2 PD) basally, becoming sparse (IS = 1 – 4 PD) apically; rim polished, punctures sparse (IS = 1 – 3 PD), becoming narrowly impunctate apically and sometimes medially. Punctures on T 2 also diversopunctate. STRUCTURE. Head long, length / width ratio 1.06 (± 0.03 SD). Face length / width ratio 0.88 (± 0.25 SD). Clypeus projecting ~ 81 % below suborbital tangent; clypeus length / width ratio 1.01 (± 0.08 SD). Mandible length mostly long, extending beyond opposing clypeal angle, and sometimes nearly reaching opposing mandibular base. Eyes convergent below, UOD / LOD ratio 1.20 (± 0.02 SD). Gena / eye width ratio 0.88 (± 0.07 SD). Ocelli normal. Pronotal angle obtuse. Intertegular span 1.44 mm (± 0.12 SD). Mesoscutum length / width ratio 0.91 (± 0.02 SD); mesoscutum / scutellum length ratio 2.87 (± 0.10 SD); scutellum / metanotum length ratio 1.59 (± 0.14 SD); metanotum / propodeal triangle length ratio 0.77 (± 0.11 SD). Protibial spur with apical serrations subequal in length to width of malus; malus extending beyond the velum, nearly ¾ the length of velum. Inner metatibial spur pectinate, teeth 4 – 5, basal teeth somewhat longer than width of rachis. Propodeal triangle posterior margin angled onto posterior propodeal surface; apical carina present, but a less prominent due to background sculpturing. Propodeal lateral carina reaching dorsolateral slope, oblique carina obscure. n = 6. Male BODY MEASUREMENTS. Length 6.86 mm (± 0.61 SD). Head length 1.75 mm (± 0.10 SD). Face length 1.44 mm (± 0.07 SD). Head width 1.61 mm (± 0.09 SD). Wing length 4.94 mm (± 0.17 SD). n = 5. COLOURATION. Head and mesosoma dark brown to black. Clypeus brown. Labrum dark brown. Mandible dark brown basally, becoming light brown medially and slightly reddish brown apically. Flagellum brown. Pronotal lobe brown. Tegula mostly brown, light brown medially. Wing membrane hyaline, veins with subcosta brown, otherwise light brown. Legs brown. Metasoma dark brown to brown with rims of terga, sterna and downcurved lateral areas of terga narrowly light brown. PUBESCENCE. Body hair colour off-white. Tomentum dense on pronotal angle, and lobe, space between pronotal lobe and tegula, and metanotum anteromedially; moderately dense on paraocular area, and T 2 – T 4 basolaterally. Mesosoma covered by appressed, simple, very short hairs (<0.25 OD) mixed with sparse, erect, weakly plumose, long hairs (<1.5 OD) on mesoscutum and erect, plumose, long hairs (<2 OD) on other areas. Metasomal sterna with dense, erect and sub-appressed, plumose, short (<1 OD) hairs. Wing hairs brown, very short and sparse. SURFACE SCULPTURE. Clypeus imbricate to tessellate, becoming polished apical ⅔; punctures irregularly sparse (IS <3 PD) apically and medially, becoming denser (IS <1 PD) basolaterally. Supraclypeal area imbricate, becoming weakly so or polished medially; punctures dense (IS <1 PD). Frons reticulate-punctate; punctures crowded (IS = 0 PD). Ocellocular area polished; punctures irregularly sparse (IS <3 PD), becoming dense (IS <1 PD) anteriorly. Gena polished, weakly lineolate, becoming lineolate posteriorly; punctures dense (IS <1 PD), becoming obscure posteriorly. Postgena weakly lineolate. Tegula broadly impunctate, minute punctures present anteriorly and basally, sparse (IS = 1 – 4 PD). Mesoscutum polished, becoming imbricate anteromedially; punctures dense (IS ≤ 1 PD), becoming moderately sparse (IS = 1 – 2 PD) mesad of parapsidal line and anteromedially, distinctly doubly punctate. Mesoscutellum polished; punctures moderately dense (IS = 1 – 2 PD) marginally and on median line, sparse submedially (IS = 1 – 5 PD), distinctly diversopunctate. Metanotum rugulose, punctures obscure. Propodeal triangle with rugae moderately strong, reaching posterior margin and even entering to propodeum posterior face, anastomosing medially and weakly so laterally; lateral rugae extending to dorsolateral slope; interspace sculpture shiny to weakly imbricate. Preëpisternum reticulate-rugose. Hypoepimeron rugulose; punctures obscure. Mesepisternum reticulate-rugose. Propodeum lateral face strongly imbricate, becoming rugulose posteriorly; posterior face rugulose to rugose, punctures obscure. T 1 anterior face polished, punctures present laterally, sparse (IS = 1 – 3 PD); dorsal surface polished, punctures moderately sparse (IS = 1 – 2 PD) or sparse (IS = 1 – 3 PD), becoming sparser towards premarginal line (IS = 1 – 4 PD) and in small subapicolateral boss (IS = 2 – 4 PD), diversopunctate, punctures sparse (IS = 1 – 3 PD) on rim, becoming impunctate apical half and impunctate medially on rim. T 2 disc polished, but somewhat weakly coriarious subapicolateral boss apically, punctures moderately sparse (IS = 1 – 2 PD) basally, becoming sparse (IS = 1 – 3 PD) apically; rim polished, punctures sparse (IS = 1 – 3 PD), becoming narrowly impunctate apically. STRUCTURE. Head long, length / width ratio 1.08 (± 0.01 SD). Face length / width ratio 0.90 (± 0.02 SD). Clypeus projecting ~ 68 % below suborbital tangent. Mandible long, reaching beyond opposing clypeal angle. Flagellomeres short, F 1 length / pedicel length ratio 0.90 (± 0.06 SD); F 2 length / F 1 length ratio 1.20 (± 0.08 SD); F 2 length / F 2 width ratio 0.95 (± 0.06 SD). Gena / eye width ratio 0.81 (± 0.05 SD). Forewing with 3 submarginal cells. Pronotal angle obtuse. Intertegular span 1.19 mm (± 0.12 SD). Mesoscutum length / width ratio 0.94 (± 0.05 SD). Propodeal triangle posterior margin angled onto posterior propodeal surface; apical carina present, but indistinct from background sculpturing. Propodeum lateral carinae reaching dorsolateral slope. n = 5. GENITALIA (Figs 62, 120 K). Gonostylus elongate, approximately twice as long as broad, and bears short setae. Retrorse lobe broad, about 3 times as long as broad, covered in short sparse setae.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF7FFF49FE36FD39FD64FDE2.taxon	distribution	Range Chihuahuan Desert of Arizona, New Mexico, Arizona, west Texas and to north of Colorado and south of Utah (Fig. 63).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF7FFF49FE36FD39FD64FDE2.taxon	biology_ecology	Floral hosts ASTERACEAE: Cirsium sp., Senecio sp. • MALVACEAE: Sphaeralcea laxa Wooton & Standl. • RUBIACEAE: Houstonia wrightii A. Gray	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF74FF4DFE6BFDCCFCE7FCE7.taxon	description	Figs 64, 65, 93 C, 97 D, 104 F, 105 B, 106 C	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF74FF4DFE6BFDCCFCE7FCE7.taxon	etymology	Etymology Cockerell (1897 c) did not explain the epithet. The specific epithet is a comparative form of the Latin adjective ‘ angustus ’, meaning ‘ narrow’ or ‘ constricted’; but the author’s intent is unknown.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF74FF4DFE6BFDCCFCE7FCE7.taxon	diagnosis	Diagnosis Female L. angustius can be recognised by combination of head long, length / width ratio 1.07 (± 0.03 SD) (remarkably long in L. pathiranae sp. nov.); mesoscutum imbricate (more polished in L. angusticulum sp. nov. and L. pathiranae; rough in L. robertsoni) with dense (IS ≤ 1 PD) punctures without double punctation (sparser and distinctive double punctation present in L. angustius and L. pathiranae); hypoepimeron partly polished (rugulose in L. angusticulum, L. pathiranae and L. robertsoni), punctures obscure (distinct in L. robertsoni and L. pathiranae); propodeal triangle with rugae reaching posterior margin (separating from species in which the rugae do not reach the posterior margin), propodeal triangle posterior margin sharply edged with a less prominent apical carina (this excludes all species with a smoothly rounded posterior margin; apical carina is prominently raised in L. pathiranae); T 1 strongly coriarious on anterior face with sub-appressed hairs but incomplete fan with narrow dorsal opening (T 1 anterior face mostly polished with broad dorsal opening in the fan in L. angusticulum); and T 2 – T 3 with distinct basolateral tomentum (removes all species lacking basal patches or bands of tomentum on T 2 – T 3, such as L. nitidiusculum group). Males of this species are unknown. Lasioglossum angustius are most similar to presumably undescribed species from various localities in Mexico, but these species exhibit punctate tegulae, coarser mesepisternum structures, and raised apical carinae.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF74FF4DFE6BFDCCFCE7FCE7.taxon	materials_examined	Type material Lectotype of Halictus angustior (designated herein; examined by photos) UNITED STATES – New Mexico • ♀; Las Cruces; [32.32 ° N, 106.76 ° W]; 25 Aug. 1895; Cockerell leg.; ex Flaveria (presumably); USNM Type No. 3388. Specimen labels “ Ckll. 4681. / Las Cruces. / on Flaveria? // H. angustior / TYPE // Type / No. 3388 / U. S. N. M. ”. Paralectotype of Halictus angustior (examined) UNITED STATES – New Mexico • 1 ♀; Las Cruces; [32.32 ° N, 106.76 ° W]; 25 Aug. 1895; Cockerell leg.; ex Flaveria (presumably); USNM Type No. 3388 (Fig. 64). Specimen labels “ Ckll. 4683. / Las Cruces. / on Flaveria? // Halictus / angustior / Ckll. // CoType / No. 3388 / U. S. N. M. ”. Type notes The original description was based on three females, two collected together in Las Cruces, New Mexico and another from Mesilla, New Mexico. The specimen above is designated as the lectotype to fix the name given the recognition of closely related species in this group (e. g., see below). Cockerell, 1897 c originally described a male specimen, which he tentatively identified as Lasioglossum angustius, collected from Ruidoso Creek, New Mexico, at an elevation of 6600 feet, on July 10, on flowers of Rhus (E. O. Wooton, no. 172). However, upon re-examination, the specimen was determined to represent a male of Lasioglossum diatretum. Material examined MEXICO – Durango • 1 ♀; Tlahualilo; [26.386 ° N, 103.618 ° W]; 3 Sep. 1904; A. W. Morrill leg.; ex Prunus persica (L.) Batsch (verbatim: peach blossoms); USNM. UNITED STATES – Arizona • 1 ♀; Cochise Co., Lewis Springs; [31.581 ° N, 110.143 ° W]; 18 Jun. 1915; Harold Morrison leg.; USNM. – New Mexico • 1 ♀; Doña Ana Co., Mesilla; [32.270 ° N, 106.801 ° W]; 13 Jul. 1897; Cockerell leg.; ex Datura inoxia (verbatim: Datura meteloides); USNM • 1 ♀; same data as for preceding; 1 – 31 May 1909; Cockerell leg.; ex yellow Composite like Crepis; USNM.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF74FF4DFE6BFDCCFCE7FCE7.taxon	description	Redescription Female BODY MEASUREMENTS. Length 7.58 mm (± 0.21 SD). Head length 1.87 mm (± 0.03 SD). Face length 1.50 mm (± 0.04 SD). Head width 1.74 mm (± 0.05 SD). Wing length 4.99 mm (± 0.09 SD). n = 4. COLOURATION. Head and mesosoma dark brown. Clypeus apex brown. Labrum brown. Mandible brown basally, slightly reddish brown apically. Flagellum brown dorsally, light brown ventrally. Pronotal lobe brown. Tegula mostly brown, light brown medially. Wing membrane faintly brownish, veins with subcosta brown, otherwise light brown. Legs brown. Metasoma brown with rims of terga, sterna and downcurved lateral areas of terga narrowly light brown. PUBESCENCE. Body hair colour off-white. Tomentum dense on pronotal angle and lobe, space between pronotal lobe and tegula, metanotum anteromedially, propodeum lateral surface anterodorsally, T 2 – T 3 basolaterally, and T 4 basally; moderately dense on anterolateral angle and marginally on mesoscutum; very sparse on supraclypeal area dorsolaterally, paraocular area, gena near eye margin, propodeum lateral (except anterodorsal area) and posterior surface mostly on dorsally. Mesosoma covered by appressed, simple, very short hairs (<0.25 OD) mixed with sparse, erect, weakly plumose, long hairs (<1 OD) on mesoscutum and erect, plumose, long hairs (<2 OD) on other areas. Metasomal sterna with sparse, erect, simple to weakly plumose, long (<3 OD) hairs. Metafemoral scopa with dense plumose hairs. Wing hairs brown, very short and sparse. T 1 with a fan, narrow dorsal opening (1.5 OD), composed of overlapping sub-appressed simple hairs (<1.5 OD). SURFACE SCULPTURING. Clypeus tessellate, becoming polished apical ⅓; punctures irregularly sparse (IS <3 PD) apically and medially, becoming slightly smaller and denser (IS <1 PD) on basolaterally. Supraclypeal area tessellate, becoming imbricate ventrally; punctures fine, sparse (IS = 1 – 3 PD). Frons reticulate-punctate; punctures crowded (IS = 0 PD). Ocellocular area polished; punctures moderately dense (IS = 1 – 2 PD) and diversopunctate posteriorly and, becoming crowded (IS = 0 PD) anteriorly. Gena rugulose to weakly lineolate, becoming lineolate posteriorly; punctures obscure. Postgena weakly imbricate to weakly lineolate. Tegula punctures sparse (IS = 1 – 3 PD) in the anterior half and basally, becoming impunctate posteriorly. Mesoscutum imbricate, but quite weakly so submedially; punctures dense (IS ≤ 1 PD), diversopunctate, but not that conspicuous. Mesoscutellum polished but imbricate marginally; punctures dense (IS ≤ 1 PD) marginally and on median line, sparse submedially (IS = 1 – 3 PD), diversopunctate. Metanotum rugulose; punctures obscure. Propodeal triangle with rugae moderately strong, reaching posterior margin, strongly anastomosing, lateral rugae extending to dorsolateral slope of propodeum lateral face; interspace sculpture shiny to weakly imbricate. Preëpisternum rugulose. Hypoepimeron polished, becoming finely rugulose dorsally; punctures dense (IS ≤ 1 PD), becoming crowded (IS = 0 PD) and obscure towards dorsally. Mesepisternum reticulate-rugulose, becoming rugulose or strongly imbricate posteroventrally; punctures obscure. Propodeum lateral face strongly imbricate; posterior face imbricate-rugulose. T 1 anterior face strongly coriarious, punctures present laterally; dorsal surface mostly polished, becoming weakly coriarious on small subapicolateral boss and rim, punctures moderately sparse (IS = 1 – 2 PD), becoming more sparsely punctate (IS = 2 – 6 PD) in small subapicolateral boss and punctures fine, minute, and sparse (IS = 1 – 4 PD) on rim, becoming impunctate apical half and medially on rim. T 2 disc polished to coriarious, punctures moderately dense (IS = 1 – 2 PD) basally, becoming sparse (IS = 1 – 3 PD) apically; rim coriarious, punctures minute and sparse (IS = 1 – 3 PD), becoming impunctate narrowly after premarginal line. STRUCTURE. Head long, length / width ratio 1.07 (± 0.03 SD). Face length / width ratio 0.86 (± 0.02 SD). Clypeus projecting ~ 86 % below suborbital tangent; clypeus length / width ratio 0.93 (± 0.05 SD). Mandible short, nearly reaching opposing clypeal angle. Eyes convergent below, UOD / LOD ratio 1.15 (± 0.01 SD). Ocelli normal. Gena / eye width ratio 0.75 (± 0.03 SD). Pronotal angle obtuse. Intertegular span 1.28 mm (± 0.04 SD). Mesocutum length / width ratio 0.90 (± 0.02 SD); mesoscutum / scutellum length ratio 2.68 (± 0.16 SD); scutellum / metanotum length ratio 1.52 (± 0.05 SD); metanotum / propodeal triangle length ratio 0.85 (± 0.03 SD). Protibial spur with apical serrations subequal in length to width of malus; malus extending beyond the velum, nearly ½ the length of velum. Inner metatibial spur pectinate, teeth 3 – 4, basal teeth somewhat longer than width of rachis. Propodeal triangle posterior margin angled onto posterior propodeal surface, apical carina indistinct from background sculpturing. Propodeal lateral carina reaching dorsolateral slope, oblique carina obscure. n = 4. VARIATION. Specimens from Arizona and Durango have hypoepimeron punctures that are mostly crowded. Male Unknown.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF74FF4DFE6BFDCCFCE7FCE7.taxon	distribution	Range Chihuahuan Desert from Mesilla, New Mexico and Tlahualilo, Durango, Mexico (Fig. 65).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF74FF4DFE6BFDCCFCE7FCE7.taxon	biology_ecology	Floral hosts ASTERACEAE: Flaveria sp. (presumably, like Crepis sp.) • SOLANACEAE: Datura inoxia Mill. • ROSACEAE: Prunus sp.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF74FF4DFE6BFDCCFCE7FCE7.taxon	discussion	Remarks Rare, with only four known specimens. The most recent record for this species dates to May 1909 in Mesilla, near Las Cruces, the location of the syntypes. Since then, there have been no additional records of this species based on observed materials. Previously, L. angusticulum sp. nov. was erroneously identified as Lasioglossum angustius. Superficially similar species have been observed in various localities in Mexico, some of which exhibit distinct characteristics such as punctate tegulae, coarser mesepisternum structures, and raised oblique carinae.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF70FF5CFE12FCC1FC7AFAD9.taxon	description	Figs 66 – 69, 93 A, 94 A, 105 D, 109 B, 110 A, 120 L	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF70FF5CFE12FCC1FC7AFAD9.taxon	diagnosis	Diagnosis Both sexes of Lasioglossum kincaidii can be recognized by the combination of an enlarged, punctate tegula and coarsely rugose mesepisternum. Lasioglossum kincaidii is unlikely to be confused with other species, yet it bears a resemblance to L. pectorale, particularly in the presence of coarsely rugose mesepisterna. However, distinguishing features include the broadly impunctate tegula and polished mesoscutum in L. pectorale, while the mesoscutum of L. kincaidii is comparatively duller. Lasioglossum glabriventre has an enlarged tegula but the mesepisternum is quite smooth.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF70FF5CFE12FCC1FC7AFAD9.taxon	etymology	Etymology Presumably named for Mr T. Kincaid, who collected specimens from Olympia, Washington that were subsequently described by Cockerell.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF70FF5CFE12FCC1FC7AFAD9.taxon	materials_examined	Type material Syntype of Halictus kincaidii (not examined) UNITED STATES – Washington • 1 ♀; Thurston Co., Olympia, [47.038 ° N, 122.901 ° W]; 13 Jun. 1895; T. Kincaid leg.; USNM Type No. 29421. Specimen labels “ Olympia, / Wash. // 6 Ju 95 // Type No. / 29421 / U. S. N. M. // H. kincaidii. / n. sp. // USNMENT / 01570341 ”. Material examined CANADA – British Columbia • 1 ♀; Kelowna, Summerhill Pyramid Winery; 49.80263 ° N, 119.50596 ° W; 31 Aug. – 14 Sep. 2016; C. Dawson leg.; RSKM • 1 ♀; Okan. – Similk; 49.17693 ° N, 119.76329 ° W; 27 Aug. – 17 Sep. 2015; C. Dawson leg.; RSKM • 2 ♀♀; Okan. – Similk; 49.187633 ° N, 119.765 ° W; 4 – 15 Sep. 2015; C. Dawson leg.; RSKM • 1 ♀; Okan. – Similk; 49.192341 ° N, 119.769038 ° W; 25 Aug. – 11 Sep. 2015; D. Marks leg.; RSKM • 1 ♀; Okan. – Similk; 49.17648 ° N, 119.76225 ° W; 25 Aug. – 12 Sep. 2015; C. Dawson leg.; RSKM • 1 ♀; Okan. – Similk; 49.188638 ° N, 119.766776 ° W; 21 Aug. – 15 Sep. 2015; C. Dawson leg.; RSKM • 2 ♀♀; Okanagan Falls, Vaseux Lake Provincial Park; 49.29901 ° N, 119.53063 ° W; 16 Aug. – 25 Sep. 2021; TM, HS and AB (J. Heron) leg.; RSKM • 2 ♀♀; same data as for preceding; 7 Jul. – 16 Aug. 2021; H. Singh and T. McGratton (J. H) leg.; RSKM • 1 ♀; Oliver, Phantom Creek Estates; 49.11641 ° N, 119.55425 ° W; 15 Jun. 2017; D. Marks, K. Grant and J. Heron leg.; RSKM • 1 ♀; Oliver, Phantom Creek Estates Winery; 49.1171 ° N, 119.5582 ° W; 27 Jul. – 18 Aug. 2017; D. Marks, K. Grant and J. Heron leg.; RSKM • 2 ♀♀; Osoyos, West Bench; 49.03687 ° N, 119.50581 ° W; 21 Jun. – 25 Jul. 2017; D. Marks and K. Grant (J. Heron) leg.; RSKM • 2 ♀♀; Osoyos, West Bench (Crown Land); 49.0304 ° N, 119.4997 ° W; elev. 438 m; 14 Jun. 2017; J. Heron, D. Marks and K. Grant leg.; RSKM. MEXICO – Baja California • 1 ♀; Ensenada; [31.858 ° N, 116.606 ° W]; 24 Jun. 1950; A. L. Melander leg.; UCRC • 1 ♂; San Vicente; [31.326 ° N, 116.247 ° W]; 8 Jul. 1963; J. Powell leg.; EMEC • 1 ♀; Sierra San Pedro Martir, La Encantada; [30.977 ° N, 115.384 ° W]; elev. 2133.6 m; 1 Jun. 1958; J. Powell leg.; EMEC • 4 ♀♀; Norte, Rio San Miguel, La Mision; [32.129 ° N, 116.8154 ° W]; 22 Aug. 1981; DKF leg.; SDMC. UNITED STATES – California • 2 ♀♀; Alameda Co., Alameda foothills; [37.789 ° N, 122.225 ° W]; W. M. Giffard leg.; CAS • 1 ♀; same data as for preceding; 1 – 31 Aug. 1915; W. M. Giffard leg.; CAS • 1 ♂; same data as for preceding; W. M. Giffard leg.; CAS • 1 ♀; Alameda Co., Altamont; [37.744 ° N, 121.663 ° W]; 13 May 1956; W. R. Powell leg.; EMEC • 1 ♀; Alameda Co., Berkeley; [37.872 ° N, 122.272 ° W]; 6 May 1952; J. D. Lattin leg.; EMEC • 1 ♂; same data as for preceding; 15 Jul. 1947; P. D. Hurd leg.; EMEC • 1 ♀; same data as for preceding; 24 Apr. 1949; P. D. Hurd leg.; EMEC • 1 ♀; same data as for preceding; 8 Mar. 1946; P. D. Hurd leg.; ex Narcissus (verbatim: daffodil); EMEC • 1 ♀; same data as for preceding; 8 Jun. 1948; P. D. Hurd leg.; EMEC • 1 ♀; same data as for preceding; EMEC • 1 ♀; same data as for preceding; 1 May 1949; P. D. Hurd leg.; EMEC • 8 ♂♂; same data as for preceding; 4 Sep. 1959; J. Powell leg.; ex Baccharis; EMEC • 1 ♀; same data as for preceding; 4 Sep. 1959; J. Powell leg.; ex Baccharis; EMEC • 2 ♂♂; same data as for preceding; 2 Oct. 1960; D. D. Linsdale leg.; EMEC • 17 ♀♀; same data as for preceding; 10 May 1912; J. C. Bridwell leg.; EMEC • 3 ♀♀; same data as for preceding; 27 Mar. 1912; J. C. Bridwell leg.; USNM • 1 ♀; same data as for preceding; 6 May 1912; J. C. Bridwell leg.; USNM • 1 ♀; same data as for preceding; 14 Apr. 1954; H. L. Hansen leg.; EMEC • 3 ♀♀; same data as for preceding; 1 Sep. 1948; EMEC • 1 ♀; same data as for preceding; 14 Mar. 1949; J. W. MacSwain leg.; EMEC • 1 ♀; same data as for preceding; 29 Oct. 1912; EMEC • 1 ♀; Alameda Co., Emeryville; [37.831 ° N, 122.285 ° W]; 5 Sep. 1936; G. E. Bohart leg.; CAS • 1 ♀; Alameda Co., Hillsback of Oakland; [37.804 ° N, 122.271 ° W]; 2 Aug. 1908; E. C. Van Dyke leg.; CAS • 1 ♀; Alameda Co., Niles; [37.584 ° N, 121.986 ° W]; 14 Apr. 1904; E. J. Brausigon leg.; UCRC • 1 ♀; Alameda Co., Oakland; [37.804 ° N, 122.271 ° W]; 1 – 31 Aug. 1936; E. S. Ross leg.; CAS • 1 ♀; same data as for preceding; 10 Apr. 1954; R. Schuster leg.; EMEC • 2 ♀♀; same data as for preceding; 6 Aug. 1920; P. H. Timberlake leg.; ex visiting Compositae; UCRC • 1 ♂; same data as for preceding; 5 Nov. 1955; B. J. Adelson leg.; EMEC • 1 ♀; Alameda Co., San Lorenzo; [37.681 ° N, 122.124 ° W]; 4 Sep. 1954; P. D. Hurd leg.; EMEC • 1 ♀; Alameda Co., U. C. Campus; [37.8 ° N, 122.27 ° W]; 2 Sep. 1910; J. C. Bridwell leg.; USNM • 1 ♂; Alameda Co., U. C. Campus, Berkeley; [37.873 ° N, 122.26 ° W]; 10 Aug. 1910; J. C. Bridwell leg.; USNM • 1 ♀; Alameda Co., University of California Campus, Barkeley; [37.87 ° N, 122.26 ° W]; 21 Jun. 1934; H. A. Scullen leg.; OSAC • 1 ♀; Alpine Co., Markleeville; [38.695 ° N, 119.78 ° W]; 2 Jul. 1963; R. P. Allen leg.; CSCA • 1 ♂; Butte Co., Chico; [39.740 ° N, 121.836 ° W]; 7 Jun. 1954; M. Wasbauer leg.; EMEC • 2 ♀♀; Colusa Co., Sycamore; [39.130 ° N, 121.940 ° W]; 15 Jul. 1916; EMEC • 3 ♀♀; Contra Costa Co., Antioch; [38.005 ° N, 121.806 ° W]; 8 May 1950; P. D. Hurd leg.; EMEC • 1 ♀; same data as for preceding; 21 May 1949; L. W. Quate leg.; EMEC • 1 ♀; Contra Costa Co., Danville; [37.820 ° N, 122 ° W]; 5 Oct. 1951; F. X. Williams leg.; CAS • 1 ♀; same data as for preceding; 19 Sep. 1951; F. X. Williams leg.; CAS • 1 ♀; same data as for preceding; 20 Jul. 1952; F. X. Williams leg.; CAS • 1 ♀; Contra Costa Co., Hampton Road, P. F. Yorks; [37.984 ° N, 122.113 ° W]; 25 Jul. 1975; ex Photinia arbutifolia (verbatim: Heteromeles arbutifolia); USNM • 1 ♀; Contra Costa Co., Mount Diablo; [37.882 ° N, 121.914 ° W]; 10 Jul. 1947; U. N. Lanham leg.; UCMC • 1 ♂; Contra Costa Co., Point Molette, Richmond; [37.948 ° N, 122.42 ° W]; 19 Jun. 1964; J. A. Powell leg.; EMEC • 1 ♂; Contra Costa Co., Richmond; [37.936 ° N, 122.347 ° W]; 23 Sep. 1950; EMEC • 1 ♂; Del Norte Co., Smith River; [41.94 ° N, 124.20 ° W]; 17 Jul. 1930; J. M. Aldrich leg.; EMEC • 1 ♀; Del Notre Co., Klamath; [41.5263 ° N, 124.0383 ° W]; 18 Jul. 1934; E. P. Van Duzee leg.; CAS • 1 ♂; El Dorado Co., Fallen Leaf Lake; [38.92 ° N, 120.06 ° W]; 24 Aug. 1953; Lattin and Lattin leg.; EMEC • 1 ♂; El Dorado Co., Chile Bar; [38.771 ° N, 120.813 ° W]; 5 Jul. 1948; J. W. Adams leg.; EMEC • 1 ♀; El Dorado Co., Fallen Leaf Lake; [38.92 ° N, 120.06 ° W]; 24 Jul. 1953; Lattin and Lattin leg.; EMEC • 1 ♀; El Dorado Co., North End Lake Folsom; [38.820 ° N, 121.092 ° W]; 23 May 1967; A. Keuter leg.; CAS • 1 ♀; Fresno Co., Laton; [36.433 ° N, 119.687 ° W]; 5 Oct. 1959; R. R. Snelling leg.; EMEC • 1 ♂; Fresno Co., Mono Hot Springs; [37.327 ° N, 119.018 ° W]; 8 Aug. 1956; R. O. Schuster leg.; EMEC • 1 ♀; Fresno Co., Shaver Lake; [37.089 ° N, 119.318 ° W]; 27 Jul. 1968; J. B. Hoy leg.; EMEC • 1 ♀; Glenn Co., Artois; [39.62 ° N, 122.19 ° W]; 2 Jun. 1953; J. W. MacSwain leg.; EMEC • 1 ♀; Glenn Co., Phelan Is. REM; 39.6785 ° N, 121.9639 ° W; 20 Jun. 2012; N. Williams leg.; WRME • 1 ♀; Humboldt Co., Samoa dunes; [40.763 ° N, 124.222 ° W]; 26 Jun. 1969; C. Slobodchikoff leg.; EMEC • 2 ♀♀; Humboldt Co., 1 mi. S of Blue Lake; [40.869 ° N, 123.984 ° W]; 12 Jun. 1959; T. R. Haig leg.; CSCA • 1 ♀; Humboldt Co., Arcata; [40.868 ° N, 124.091 ° W]; 5 Jun. 1969; R. R. Snelling and R. A. Snelling leg.; EMEC • 1 ♂; same data as for preceding; 4 Jun. 1969; J. Powell leg.; EMEC • 1 ♀; Humboldt Co., Dry Lagoon State Park; [41.225 ° N, 124.098 ° W]; 10 Aug. 1953; P. H. Arnaud Jr leg.; CAS • 3 ♀♀; Humboldt Co., Eureka; [40.802 ° N, 124.164 ° W]; 6 Jul. 1946; E. Mills leg.; ex on leaves of Lilium longiflorum; CSCA • 1 ♀; same data as for preceding; 6 Jul. 1946; E. Mills leg.; ex on leaves of Lilium longiflorum; UCRC • 1 ♀; Humboldt Co., Humboldt; [40.8 ° N, 123.8 ° W]; 3 Mar. 1911; J. C. Bridwell leg.; USNM • 2 ♀♀; Humboldt Co., Kneeland School; [40.726 ° N, 123.944 ° W]; 17 Aug. 1961; T. R. Haig leg.; CSCA • 1 ♂; Humboldt Co., Mad Riv. near Arcata; [40.914 ° N, 124.101 ° W]; 1 – 30 Jun. 1924; J. M. Aldrich leg.; USNM • 1 ♀; Humboldt Co., Maple Creek; [40.762 ° N, 123.869 ° W]; 15 Aug. 1968; R. P. Allen leg.; CSCA • 1 ♂; same data as for preceding; 15 Aug. 1968; R. P. Allen leg.; CSCA • 1 ♀; Inyo Co., Lone Pine; [36.61 ° N, 118.06 ° W]; 13 May 1969; P. Welles leg.; ex Salix; EMEC • 1 ♀; same data as for preceding; 13 May 1969; J. A. Chemsak leg.; EMEC • 1 ♀; Kern Co., 15 mi. E of Bakersfield; [35.368 ° N, 118.752 ° W]; 2 May 1953; C. D. MacNeill leg.; CAS • 1 ♀; Kern Co., Kern River Preserve, 1.5 mi. W of Weldon; [35.667 ° N, 118.317 ° W]; 31 Mar. 1983; J. D. Pinto and R. K. Velten leg.; UCRC • 1 ♀; Kern Co., Kernville; [35.7331 ° N, 118.4344 ° W]; 10 Jul. 1955; J. Powell leg.; EMEC • 1 ♂; Kern Co., Walker Pass; [35.663 ° N, 118.027 ° W]; elev. 1597.2 m; 11 Jul. 1964; R. R. Snelling leg.; EMEC • 1 ♀; King Co., 4 mi. NW of Straford; [36.230 ° N, 119.874 ° W]; 4 Nov. 1954; R. R. Snelling leg.; ex Grindelia; EMEC • 1 ♂; Los Angeles Co., Tanbark Flat; [34.204 ° N, 117.760 ° W]; 12 Jul. 1950; W. C. Bentinck leg.; EMEC • 1 ♀; Los Angeles Co., 16 mi. NE of Saugus; [34.570 ° N, 118.342 ° W]; 18 Jun. 1962; J. F. Lawrence leg.; EMEC • 1 ♂; Los Angeles Co., Alamitos Bay; [33.750 ° N, 118.118 ° W]; 6 May 1939; P. H. Timberlake leg.; ex Oenothera; UCRC • 1 ♀; Los Angeles Co., Ballona Wetlands near Playa del Rey; [33.97 ° N, 118.44 ° W]; 5 Apr. 1981; C. Nagano leg.; EMEC • 1 ♀; same data as for preceding; 26 Aug. 1981; C. Nagano leg.; EMEC • 3 ♀♀; same data as for preceding; 22 Apr. 1981; C. Nagano leg.; EMEC • 1 ♀; same data as for preceding; 6 Jun. 1981; C. Nagano leg.; EMEC • 2 ♀♀; same data as for preceding; 23 Jul. 1980; C. Nagano, J. Hogue and E. Birdsall leg.; EMEC • 1 ♀; same data as for preceding; 2 Jul. 1980; C. Nagano, J. Hogue and C. Chapelle leg.; EMEC • 1 ♀; same data as for preceding; 9 Jun. 1981; R. R. Snelling leg.; ex Heliotropium curassavicum; EMEC • 3 ♀♀; same data as for preceding; 11 Jun. 1981; R. R. Snelling leg.; ex Carpobrotus; EMEC • 1 ♀; same data as for preceding; 11 Jun. 1981; R. R. Snelling leg.; ex Heliotropium curassavicum; EMEC • 1 ♂; same data as for preceding; 25 Jul. 1981; R. R. Snelling leg.; ex Foeniculum vulgare; EMEC • 2 ♂♂; same data as for preceding; 9 May 1981; C. Nagano leg.; EMEC • 1 ♂; same data as for preceding; 11 Jun. 1981; R. R. Snelling leg.; EMEC • 1 ♂; same data as for preceding; 8 May 1981; R. R. Snelling leg.; EMEC • 1 ♂; same data as for preceding; 11 Jun. 1981; R. R. Snelling leg.; ex Heliotropium curassavicum; EMEC • 1 ♂; same data as for preceding; 6 Jun. 1981; C. Nagano leg.; EMEC • 5 ♀♀; Los Angeles Co., Claremont; [34.097 ° N, 117.719 ° W]; USNM • 1 ♀; Los Angeles Co., Covina; [34.092 ° N, 117.879 ° W]; 15 Apr. 1925; M. Kamal leg.; ex Brassica; UCRC • 4 ♀♀; Los Angeles Co., Huntington Park; [33.983 ° N, 118.217 ° W]; 28 Jul. 1940; A. Bauman leg.; EMEC • 1 ♀; Los Angeles Co., La Mirada; [33.902 ° N, 118.01 ° W]; 4 Apr. 1939; J. Roberds leg.; EMEC • 1 ♀; same data as for preceding; 21 Feb. 1939; J. Roberds leg.; EMEC • 1 ♀; same data as for preceding; 4 Apr. 1939; EMEC • 1 ♀; Los Angeles Co., Los Angeles; [34.053 ° N, 118.243 ° W]; 17 Apr. 1935; K. Hollan leg.; EMEC • 1 ♀; same data as for preceding; 31 Jul. 1938; EMEC • 1 ♀; same data as for preceding; 5 May 2017; R. May leg.; EMEC • 1 ♀; same data as for preceding; 10 Apr. 1916; EMEC • 1 ♂; same data as for preceding; 18 Aug. 2017; R. May leg.; EMEC • 1 ♀; same data as for preceding; 6 Apr. 2018; EMEC • 1 ♀; same data as for preceding; 1 Apr. 2016; EMEC • 2 ♀♀; same data as for preceding; 23 Jun. 1917; R. May leg.; EMEC • 1 ♀; same data as for preceding; 20 Apr. 1916; EMEC • 1 ♀; same data as for preceding; 14 Jun. 1947; H. Wilson leg.; EMEC • 1 ♀; Los Angeles Co., Pasadena; [34.148 ° N, 118.145 ° W]; 1 – 31 Aug. 1913; O. B. Hayes leg.; OSAC • 1 ♀; Los Angeles Co., Pomona; [34.061 ° N, 117.756 ° W]; 8 Jul. 1931; EMEC • 1 ♀; Los Angeles Co., Redondo; [33.84 ° N, 118.38 ° W]; 22 Jun. 1937; J. Roberds leg.; EMEC • 1 ♀; Los Angeles Co., Redondo Beach; [33.858 ° N, 118.379 ° W]; 21 Jun. 1937; J. Roberds leg.; EMEC • 1 ♀; Los Angeles Co., Rivera; [33.989 ° N, 118.089 ° W]; 7 May 1927; P. H. Timberlake leg.; UCRC • 1 ♀; Los Angeles Co., Santa Monica; [34.022 ° N, 118.481 ° W]; 30 Jun. 1935; E. G. Linsley leg.; EMEC • 1 ♀; Los Angeles Co., Valyermo; [34.446 ° N, 117.852 ° W]; 13 May 1944; A. L. Melander leg.; UCRC • 1 ♂; Los Angeles Co., West Hollywood Hills; [34.116 ° N, 118.370 ° W]; 15 Aug. 1949; R. G. Howell leg.; EMEC • 4 ♂♂; Los Angeles Co., Westwood Hills; [38.302 ° N, 122.32 ° W]; 15 Jul. 1930; J. Wilson leg.; EMEC • 1 ♀; same data as for preceding; 29 Feb. 1936; E. G. Linsley leg.; ex Helianthus; EMEC • 2 ♀♀; same data as for preceding; 3 Apr. 1937; N. F. Hardman leg.; ex willow; EMEC • 1 ♀; same data as for preceding; 18 Feb. 1935; E. G. Linsley leg.; EMEC • 3 ♀♀; Los Angeles Co., Whittier; [33.972 ° N, 118.031 ° W]; L. L. Muchmore leg.; EMEC • 1 ♂; same data as for preceding; L. L. Muchmore leg.; EMEC • 1 ♂; same data as for preceding; 10 Aug. 1920; P. H. Timberlake leg.; UCRC • 1 ♀; same data as for preceding; 30 Jul. 1926; P. H. Timberlake leg.; ex Calystegia occidentalis (verbatim: Convolvulus occidentalis); UCRC • 1 ♀; same data as for preceding; 30 Jul. 1934; P. H. Timberlake leg.; ex Symphyotrichum (verbatim: Michaelmas daisy); UCRC • 1 ♀; same data as for preceding; 30 Jul. 1926; P. H. Timberlake leg.; ex Convolvulus; UCRC • 1 ♂; same data as for preceding; 21 Aug. 1950; H. M. Graham leg.; EMEC • 2 ♀♀; Los Angeles Co.; [34.33 ° N, 118.23 ° W]; 16 Apr. 1935; K. Hollan leg.; EMEC • 1 ♀; Madera Co., 8 mi. SE of Madera; [37.042 ° N, 119.958 ° W]; 17 Jun. 1957; R. R. Snelling leg.; EMEC • 1 ♀; Madoc Co., Hackamore Reservoir; [41.558 ° N, 121.14 ° W]; elev. 1417.3 m; 5 Aug. 1965; T. W. Fisher and R. E. Orth leg.; UCRC • 1 ♂; Marin Co., Stinson Beach; [37.9 ° N, 122.64 ° W]; 16 Sep. 1949; CAS • 1 ♀; Mariposa Co., Wawona; [37.537 ° N, 119.656 ° W]; 27 Jun. 1948; C. H. Spitzer leg.; EMEC • 1 ♀; Mariposa Co., Yosemite; [37.833 ° N, 119.500 ° W]; elev. 1182.6 – 1219.2 m; 25 Feb. 1938; N. F. Hardman leg.; EMEC • 1 ♀; same data as for preceding; 25 Feb. 1938; E. D. Essig leg.; EMEC • 1 ♀; same data as for preceding; 25 Feb. 1938; K. D. Snyder leg.; EMEC • 4 ♀♀; Mariposa Co., Yosemite National Park, Mirror Lake; [37.748 ° N, 119.55 ° W]; 28 Jun. 1947; A. L. Melander leg.; UCRC • 1 ♀; Mariposa Co., Yosemite Valley; [37.7455 ° N, 119.5937 ° W]; 27 May 1934; L. S. Slevin leg.; CAS • 1 ♀; same data as for preceding; 24 Jun. 1936; P. H. Timberlake leg.; ex Helenium bigelovii; UCRC • 1 ♀; same data as for preceding; 24 Jun. 1936; P. H. Timberlake leg.; ex Asclepias speciosa; UCRC • 1 ♀; Mendocino Co., 5.7 mi. S of Leggett; [39.781 ° N, 123.714 ° W]; elev. 228.6 m; 4 Jun. 1969; R. R. Snelling and R. A. Snelling leg.; EMEC • 2 ♀♀; Mendocino Co., Caspar; [39.364 ° N, 123.816 ° W]; 20 May 1936; I. McCracken leg.; ex Phacelia californica; CAS • 3 ♀♀; Mendocino Co., Eel River Ranger Station; [39.8257 ° N, 123.0836 ° W]; elev. 457.2 m; 12 Jun. 1972; R. Coville leg.; ex Convolvulus; EMEC • 1 ♀; Mendocino Co., McKerricher Beach State Park, 3 mi. N of Fort Bragg; [39.489 ° N, 123.797 ° W]; 7 May 1976; J. A. Powell leg.; EMEC • 1 ♀; Merced Co., 2 mi. SW of Livington; [37.366 ° N, 120.749 ° W]; 5 Apr. 1959; G. I. Stage leg.; ex Camissonia campestris subsp. campestris (verbatim: Oenothera dentata var. campestris); EMEC • 1 ♀; Merced Co., 2.5 mi. S of Livington; [37.341 ° N, 120.723 ° W]; 20 Mar. 1960; R. R. Snelling leg.; ex Camissonia contorta (verbatim: Oenothera contorta); EMEC • 1 ♀; same data as for preceding; 26 Mar. 1960; R. R. Snelling leg.; ex Camissonia campestris subsp. campestris (verbatim: Oenothera dentata var. campestris); EMEC • 1 ♀; same data as for preceding; 26 Mar. 1960; R. R. Snelling leg.; ex Camissonia contorta (verbatim: Oenothera contorta); EMEC • 1 ♀; same data as for preceding; 2 Apr. 1960; R. R. Snelling leg.; ex Camissonia campestris subsp. campestris (verbatim: Oenothera dentata var. campestris); EMEC • 2 ♀♀; Merced Co., Los Banos; [37.059 ° N, 120.851 ° W]; 22 May 1918; E. P. Van Duzee leg.; CAS • 6 ♀♀; same data as for preceding; 23 May 1918; E. P. Van Duzee leg.; CAS • 2 ♂♂; same data as for preceding; 22 May 1918; E. P. Van Duzee leg.; CAS • 1 ♀; Modoc Co., Alturas; [41.49 ° N, 120.54 ° W]; 9 Jul. 1922; C. L. Fox leg.; CAS • 14 ♀♀; Modoc Co., Lake City; [41.643 ° N, 120.217 ° W]; 30 Jul. 1922; C. L. Fox leg.; CAS • 1 ♀; same data as for preceding; 1 Aug. 1922; C. L. Fox leg.; CAS • 1 ♂; same data as for preceding; 30 Jul. 1922; C. L. Fox leg.; CAS • 1 ♀; Mono Co., 3 mi. E of Coleville; [38.564 ° N, 119.451 ° W]; 24 Jun. 1962; J. Powell leg.; EMEC • 1 ♀; Mono Co., Benton Station; [37.819 ° N, 118.476 ° W]; 20 May 1963; R. P. Allen leg.; CSCA • 2 ♀♀; Monterey Co., Arroyo Seco River, 5 air mi. SW of Greenfield; [36.269 ° N, 121.307 ° W]; elev. 121.9 m; 7 – 8 May 1975; E. Rogers leg.; EMEC • 2 ♀♀; same data as for preceding; 5 May 1975; E. Rogers leg.; EMEC • 2 ♀♀; same data as for preceding; 5 May 1975; J. A. Chemsak leg.; EMEC • 1 ♂; Monterey Co., Asilomar; [36.62 ° N, 121.94 ° W]; 26 Sep. 1959; C. W. O’Brien leg.; EMEC • 1 ♂; Monterey Co., Carmel; [36.555 ° N, 121.923 ° W]; 19 Jun. 1921; L. S. Slevin leg.; CAS • 1 ♀; same data as for preceding; 29 Jul. 1933; L. S. Slevin leg.; CAS • 1 ♀; same data as for preceding; 20 May 1913; E. C. Van Dyke leg.; CAS • 1 ♀; Monterey Co., Lockwood; [35.94 ° N, 121.08 ° W]; 5 May 1959; P. H. Timberlake leg.; UCRC • 3 ♀♀; Monterey Co., Monterey; [36.603 ° N, 121.893 ° W]; 25 Sep. 1934; A. L. Melander leg.; MCZ • 3 ♀♀; Nevada Co., 1 mi. S of Hobart Mills; [39.386 ° N, 120.184 ° W]; 1 Sep. 1957; J. M. Linsley leg.; ex Chrysothamnus viscidiflorus var. typicus; EMEC • 1 ♀; Nevada Co., Grass Valley; [39.219 ° N, 121.058 ° W]; 5 Apr. 1959; ex Frangula californica (verbatim: Rhamnus californica); CSCA • 1 ♀; Orange Co., Anaheim; [33.835 ° N, 117.912 ° W]; 22 Jul. 1955; C. A. Toschi leg.; EMEC • 1 ♂; Orange Co., Buena Park; [33.856 ° N, 118.004 ° W]; 19 May 1944; A. L. Melander leg.; UCRC • 1 ♀; Orange Co., Costa Mesa; [33.665 ° N, 117.912 ° W]; 25 Jun. 1939; P. D. Hurd leg.; EMEC • 1 ♀; same data as for preceding; 24 Jun. 1939; P. D. Hurd leg.; EMEC • 1 ♀; same data as for preceding; 4 Jul. 1939; P. D. Hurd leg.; EMEC • 2 ♂♂; Orange Co., Laguna Beach; [33.53 ° N, 117.77 ° W]; 7 Aug. 1936; P. H. Timberlake leg.; ex Heliotropium curassavicum; UCRC • 1 ♀; same data as for preceding; Baker leg.; USNM • 1 ♂; Orange Co., Laguna Hills; [33.60 ° N, 117.7 ° W]; 18 Jul. 1926; P. Small leg.; CAS • 1 ♀; Orange Co., Newport; [33.62 ° N, 117.89 ° W]; 15 Jul. 1937; N. F. Hardman leg.; EMEC • 2 ♀♀; Orange Co., Newport (Upper) Bay; [33.65 ° N, 117.89 ° W]; 6 Apr. 1964; A. E. Michelbacher leg.; EMEC • 1 ♀; same data as for preceding; 7 Apr. 1964; A. E. Michelbacher leg.; EMEC • 1 ♀; Orange Co., Newport Beach; [33.62 ° N, 117.89 ° W]; 19 Aug. 1920; P. H. Timberlake leg.; UCRC • 2 ♀♀; same data as for preceding; 19 May 1964; A. E. Michelbacher leg.; EMEC • 1 ♀; Orange Co., Santa Ana; [33.74 ° N, 117.88 ° W]; 30 May 1936; C. E. Norland leg.; EMEC • 1 ♀; same data as for preceding; 12 Jul. 1942; P. D. Hurd leg.; EMEC • 1 ♀; Orange Co., Westminster; [33.751 ° N, 117.990 ° W]; 6 May 1939; EMEC • 1 ♀; Pacific Co., Nahcotta; [46.50 ° N, 124.03 ° W]; 25 Jul. 1954; OSAC • 2 ♀♀; Placer Co., 4 mi. S of Rocklin; [38.731 ° N, 121.234 ° W]; 27 May 1979; Wasbauer, Adams leg.; CSCA • 1 ♀; same data as for preceding; 26 May 1979; Wasbauer, Adams leg.; CSCA • 1 ♀; Placer Co., 5 mi. N of Auburn; [38.960 ° N, 121.079 ° W]; 25 Aug. 1948; P. D. Hurd leg.; EMEC • 1 ♀; Placer Co., Forest Hill; [39.02 ° N, 120.83 ° W]; 9 Jun. 1931; E. C. Van Dyke leg.; CAS • 1 ♀; Plumas Co., 4 mi. W of Quincy; [39.936 ° N, 121.022 ° W]; 20 Jun. 1949; C. I. Smith leg.; EMEC • 2 ♂♂; Plumas Co., 5.5. mi. S of Vinton; [39.722 ° N, 120.175 ° W]; 6 Aug. 1963; J. W. MacSwain leg.; ex Taraxia tanacetifolia (verbatim: Oenothera tanacetifolia); EMEC • 2 ♂♂; same data as for preceding; 6 Aug. 1963; R. W. Thorp leg.; ex Taraxia tanacetifolia (verbatim: Oenothera tanacetifolia); EMEC • 1 ♀; Plumas Co., Lake Almanor; [40.25 ° N, 121.16 ° W]; 8 Jul. 1949; P. D. Hurd leg.; EMEC • 1 ♂; Riverside Co., 5 mi. W of Sage; [33.583 ° N, 117.016 ° W]; 3 Jul. 1963; P. D. Hurd leg.; EMEC • 1 ♀; Riverside Co., Bautista Canyon; [33.698 ° N, 116.853 ° W]; 24 Apr. 1984; F. Corcel leg.; UCRC • 1 ♀; Riverside Co., Hemet; [33.748 ° N, 116.972 ° W]; 5 – 19 Jul. 1950; E. G. Linsley, J. W. MacSwain leg.; EMEC • 1 ♀; same data as for preceding; 26 Aug. 1946; J. W. MacSwain leg.; ex Tribulus; EMEC • 1 ♀; same data as for preceding; 24 Jul. 1946; J. W. NacSwain leg.; ex Helianthus; EMEC • 1 ♀; Riverside Co., Hurkey Creek, San Jacinto Mountains; [33.714 ° N, 116.677 ° W]; 10 Jun. 1950; EMEC • 1 ♀; Riverside Co., Riverside; [33.9556 ° N, 117.3722 ° W]; elev. 290 m; 14 Sep. 1939; C. M. Dammers leg.; EMEC • 1 ♀; same data as for preceding; 21 Feb. 1937; EMEC • 4 ♀♀; same data as for preceding; 12 Sep. 1953; P. H. Timberlake leg.; ex Picris echioides; EMEC • 1 ♂; same data as for preceding; 1 Jul. 1925; P. H. Timberlake leg.; ex Leucanthemum × superbum (verbatim: shasta daisy); UCRC • 1 ♂; same data as for preceding; 21 Jul. 1930; P. H. Timberlake leg.; ex Medicago sativa; UCRC • 1 ♂; same data as for preceding; 28 Jun. 1940; P. H. Timberlake leg.; ex Heliotropium curassavicum; UCRC • 1 ♀; same data as for preceding; 16 Feb. 1925; P. H. Timberlake leg.; ex Encelia farinosa; UCRC • 6 ♀♀; same data as for preceding; 24 May 1925; P. H. Timberlake leg.; ex marguerite daisy; UCRC • 1 ♀; same data as for preceding; 15 Jul. 1928; P. H. Timberlake leg.; ex Mentha spicata; UCRC • 11 ♀♀; same data as for preceding; 30 May 1925; P. H. Timberlake leg.; ex marguerite daisy; UCRC • 1 ♀; same data as for preceding; 29 Aug. 1926; P. H. Timberlake leg.; ex Polygonum lapathifolium; UCRC • 1 ♀; same data as for preceding; 23 Aug. 1925; P. H. Timberlake leg.; ex marguerite daisy; UCRC • 1 ♀; same data as for preceding; 15 Apr. 1928; P. H. Timberlake leg.; ex Calendula; UCRC • 1 ♀; same data as for preceding; 1 Jul. 1928; P. H. Timberlake leg.; ex Leucanthemum × superbum (verbatim: shasta daisy); UCRC • 1 ♀; same data as for preceding; 1 Jul. 1928; P. H. Timberlake leg.; ex marguerite daisy; UCRC • 1 ♀; same data as for preceding; 2 Sep. 1925; P. H. Timberlake leg.; ex Deinandra; UCRC • 1 ♀; same data as for preceding; 1 Apr. 1928; P. H. Timberlake leg.; ex Calendula; UCRC • 1 ♀; same data as for preceding; 4 Jul. 1936; P. H. Timberlake leg.; ex Cosmos; UCRC • 1 ♀; same data as for preceding; 25 Jul. 1926; P. H. Timberlake leg.; ex Ageratum; UCRC • 1 ♀; same data as for preceding; 22 Sep. 1934; P. H. Timberlake leg.; UCRC • 1 ♀; same data as for preceding; 26 Oct. 1939; P. H. Timberlake leg.; ex Polygonum lapathifolium; UCRC • 1 ♀; same data as for preceding; 3 Apr. 1932; P. H. Timberlake leg.; ex Erodium moschatum; UCRC • 1 ♀; same data as for preceding; 16 Nov. 1935; P. H. Timberlake leg.; UCRC • 1 ♀; same data as for preceding; 1 Sep. 1935; P. H. Timberlake leg.; ex Lobularia maritima (verbatim: Alyssum maritimum); UCRC • 1 ♀; same data as for preceding; 6 Oct. 1935; P. H. Timberlake leg.; ex Polygonum lapathifolium; UCRC • 2 ♀♀; same data as for preceding; 7 Jun. 1934; P. H. Timberlake leg.; ex Calendula; UCRC • 1 ♀; same data as for preceding; 19 Jun. 1932; P. H. Timberlake leg.; ex Coreopsis tinctoria; UCRC • 1 ♀; same data as for preceding; 21 Apr. 1924; P. H. Timberlake leg.; ex Salix nigra; UCRC • 1 ♀; same data as for preceding; 3 May 1932; P. H. Timberlake leg.; ex Erodium moschatum; UCRC • 1 ♀; same data as for preceding; 9 May 1939; P. H. Timberlake leg.; ex Sonchus oleraceus; UCRC • 1 ♀; same data as for preceding; 25 May 1940; P. H. Timberlake leg.; ex Salix gooddingii; UCRC • 1 ♀; same data as for preceding; 7 Oct. 1937; P. H. Timberlake leg.; ex Phyla nodiflora (verbatim: Lippia filiformis); UCRC • 1 ♀; same data as for preceding; 23 Sep. 1937; P. H. Timberlake leg.; ex Stephanomeria exigua; UCRC • 1 ♀; same data as for preceding; 7 Sep. 1940; P. H. Timberlake leg.; ex michaelmas daisy; UCRC • 1 ♀; same data as for preceding; 21 Jul. 1940; P. H. Timberlake leg.; ex Coreopsis lanceolata; UCRC • 1 ♀; same data as for preceding; 16 Aug. 1936; P. H. Timberlake leg.; ex Mentha spicata; UCRC • 1 ♀; same data as for preceding; 6 Apr. 1941; P. H. Timberlake leg.; ex Erodium moschatum; UCRC • 1 ♀; same data as for preceding; 12 Apr. 1942; P. H. Timberlake leg.; ex Sonchus oleraceus; UCRC • 1 ♀; same data as for preceding; 6 Oct. 1941; P. H. Timberlake leg.; ex Heterotheca grandiflora; UCRC • 1 ♀; same data as for preceding; 13 Aug. 1947; P. H. Timberlake leg.; ex Sisyrinchium; UCRC • 1 ♀; same data as for preceding; 18 May 1947; P. H. Timberlake leg.; ex Coreopsis lanceolata; UCRC • 1 ♀; same data as for preceding; 24 Aug. 1953; P. H. Timberlake leg.; ex Picris echioides; UCRC • 1 ♀; same data as for preceding; 29 Aug. 1953; P. H. Timberlake leg.; ex Picris echioides; UCRC • 1 ♀; same data as for preceding; 3 Jun. 1934; P. H. Timberlake leg.; ex ?? lanceolata; UCRC • 1 ♀; same data as for preceding; 15 Apr. 1950; P. H. Timberlake leg.; UCRC • 1 ♀; same data as for preceding; 14 Sep. 1952; P. H. Timberlake leg.; UCRC • 1 ♀; same data as for preceding; 29 Aug. 1963; P. H. Timberlake leg.; ex Picris echioides; UCRC • 1 ♀; same data as for preceding; 4 May 1929; P. H. Timberlake leg.; ex Gilia achilleifolia subsp. multicaulis (verbatim: Gilia multicaulis); UCRC • 1 ♂; same data as for preceding; 2 Aug. 1925; P. H. Timberlake leg.; ex mint; UCRC (Fig. 68) • 1 ♂; same data as for preceding; 15 Jul. 1925; P. H. Timberlake leg.; ex Mentha spicata; UCRC • 1 ♂; same data as for preceding; 7 Sep. 1940; P. H. Timberlake leg.; UCRC (Fig. 67) • 1 ♀; Riverside Co., Temecula; [33.503 ° N, 117.124 ° W]; 10 May 1945; A. Melander leg.; UCRC • 1 ♀; Riverside Co., Vail Lake; [33.491 ° N, 116.974 ° W]; 26 Aug. 1965; R. E. Orth leg.; UCRC • 1 ♀; Sacramento Co., Carmichael; [38.639 ° N, 121.321 ° W]; 23 Aug. 1965; R. F. Wilkey leg.; CSCA • 1 ♀; Sacramento Co., Sacramento; [38.58 ° N, 121.48 ° W]; 18 Mar. 1958; W. E. Simonds leg.; CSCA • 1 ♂; same data as for preceding; 2 Aug. 1968; R. A. Belmont leg.; CSCA • 1 ♀; Sacramento Co., Sacramento; [38.58 ° N, 121.48 ° W]; 7 Sep. 1959; I. Savage leg.; CSCA • 1 ♀; same data as for preceding; 11 May 1960; M. Washauer leg.; CSCA • 1 ♀; Sacramento Co., Sherman Island; [38.059 ° N, 121.733 ° W]; 16 Oct. 1925; E. C. Van Dyke leg.; CSCA • 1 ♀; San Benito Co., 6 mi. SE of Idria; [36.355 ° N, 120.598 ° W]; 28 Jun. 1954; J. G. Rozen leg.; ex Monardella lanceolata; EMEC • 1 ♀; same data as for preceding; 28 Jun. 1954; J. D. Latin leg.; EMEC • 1 ♀; San Bernardino Co., 2 mi. NE of Highland; [34.138 ° N, 117.191 ° W]; 16 Jul. 1957; J. T. Doyen leg.; EMEC • 1 ♂; San Bernardino Co., Mojave River at Apple Valley; [34.511 ° N, 117.260 ° W]; 29 Jun. 1940; P. H. Timberlake leg.; UCRC • 1 ♀; same data as for preceding; 29 Jun. 1940; P. H. Timberlake leg.; ex Melilotus albus; UCRC • 1 ♀; San Bernardino Co., Redlands; [34.055 ° N, 117.182 ° W]; F. R. Cole leg.; USNM • 1 ♀; San Diego Co., 6 mi. E of Banner; [33.066 ° N, 116.441 ° W]; 26 Jun. 1963; C. H. Frady leg.; OSAC • 1 ♂; San Diego Co., Cardiff; [33.021 ° N, 117.279 ° W]; 19 Jun. 1940; P. H. Timberlake leg.; UCRC • 1 ♀; same data as for preceding; 19 Jun. 1940; P. H. Timberlake leg.; UCRC • 1 ♂; same data as for preceding; 19 Jun. 1965; J. Powell leg.; EMEC • 1 ♀; San Diego Co., Cuyamaca Park; [32.928 ° N, 116.568 ° W]; 4 May 1943; E. W. Clark leg.; EMEC • 1 ♂; San Diego Co., Del Mar; [32.955 ° N, 117.263 ° W]; 6 May 1979; A. S. Menke leg.; USNM • 1 ♀; San Diego Co., Geo. H. Field; SDMC • 1 ♀; San Diego Co., Jacumba; [32.617 ° N, 116.188 ° W]; 3 Oct. 1925; C. L. Fox leg.; CAS • 1 ♀; San Diego Co., La Jolla; [32.840 ° N, 117.277 ° W]; 4 Jul. 1938; Cockerell leg.; USNM • 1 ♀; San Diego Co., Lake Cuyamaca; [32.987 ° N, 116.583 ° W]; 7 May 1945; A. L. Melander leg.; UCRC • 1 ♀; San Diego Co., Mesa Grande; [33.180 ° N, 116.769 ° W]; 10 Jul. 1908; F. E. Blaisdell leg.; CAS • 1 ♀; San Diego Co., Pine Valley; [32.822 ° N, 116.530 ° W]; 2 Aug. 1926; F. W. Kelsey leg.; SDMC • 1 ♀; San Diego Co., San Diego; [32.72 ° N, 117.16 ° W]; 1930; F. W. Kelsey leg.; SDMC • 1 ♂; same data as for preceding; 3 Jul. 1945; EMEC • 1 ♀; San Diego Co., Solana Beach; [32.991 ° N, 117.269 ° W]; 19 Jun. 1963; J. Powell leg.; EMEC • 1 ♂; same data as for preceding; 19 Jun. 1963; J. Powell leg.; ex Heliotropium curassavicum; EMEC • 2 ♀♀; same data as for preceding; 19 Jun. 1963; J. Powell leg.; ex Heliotropium curassavicum; EMEC • 1 ♂; same data as for preceding; 19 Jun. 1963; P. D. Hurd leg.; ex Heliotropium curassavicum; EMEC • 16 ♀♀; same data as for preceding; 19 Jun. 1963; P. D. Hurd leg.; ex Heliotropium curassavicum; EMEC • 2 ♂♂; same data as for preceding; 19 Jun. 1963; P. D. Hurd leg.; EMEC • 1 ♀; San Diego Co., Sorrente Valley; [32.903 ° N, 117.185 ° W]; 1 Sep. 1980; Faulkner leg.; SDMC • 1 ♀; San Diego Co., Torrey Pines; [32.892 ° N, 117.240 ° W]; 30 May 1955; W. E. Simonds leg.; CSCA • 2 ♀♀; San Diego Co., Warner Hot Spring; [33.282 ° N, 116.634 ° W]; 18 Aug. 1938; C. M. Dammers leg.; UCRC • 1 ♀; San Diego Co., Warner Springs; [33.282 ° N, 116.634 ° W]; 27 Jul. 1921; SDMC • 1 ♀; same data as for preceding; 1 Aug. 1921; SDMC • 1 ♀; San Diego Co., Warner Springs, Agua Caliente Creek; [33.279 ° N, 116.720 ° W]; 23 – 25 Aug. 1980; M. Wasbauer, P. Adams leg.; CSCA • 1 ♀; San Diego Co.; [33.03 ° N, 116.77 ° W]; 12 Jul. 1929; SDMC • 1 ♀; same data as for preceding; 25 Jun. – 14 Jul. 1931; F. W. Kelsey leg.; SDMC • 1 ♀; same data as for preceding; 7 Oct. 1928; SDMC • 1 ♀; San Diego Co., San Felipe Creek; [33.170 ° N, 115.822 ° W]; 5 Jun. 1936; P. H. Timberlake leg.; ex Heliotropium curassavicum; UCRC • 1 ♀; San Francisco Co., Presidio; [37.789 ° N, 122.460 ° W]; 4 May 1927; C. L. Fox leg.; CAS • 1 ♀; San Francisco Co., Sigmund Stern Grove; [37.736 ° N, 122.479 ° W]; 17 Aug. 1958; P. H. Arnaud Jr leg.; ex Hypochoeris radicata; CAS • 1 ♀; San Francisco Co., San Francisco; [37.778 ° N, 122.416 ° W]; 15 Aug. 1928; C. L. Fox leg.; CAS • 1 ♂; same data as for preceding; 8 Nov. 1925; C. L. Fox leg.; CAS • 1 ♀; San Joaquin Co., 10 mi. S of Stockton; [37.810 ° N, 121.293 ° W]; 18 Oct. 1962; A. E. Michelbacher and M. M. Michelbacher leg.; ex banana squash; USNM • 1 ♂; same data as for preceding; 18 Oct. 1962; A. E. Michelbacher and M. M. Michelbacher leg.; ex banana squash; USNM • 1 ♀; San Joaquin Co., 4 mi. N of Tracy; [37.795 ° N, 121.425 ° W]; 30 Jul. 1960; W. A. Steffan leg.; EMEC • 1 ♂; San Joaquin Co., Tracy; [37.738 ° N, 121.434 ° W]; 21 Jun. 1949; R. F. Smith leg.; ex Melilotus; EMEC • 1 ♀; same data as for preceding; 5 May 1949; ex Brassica; EMEC • 1 ♀; same data as for preceding; 21 Jun. 1949; R. F. Smith leg.; ex Melilotus; EMEC • 1 ♀; same data as for preceding; 13 Jun. 1949; J. W. MacSwain leg.; EMEC • 1 ♀; same data as for preceding; 20 May 1949; R. F. Smith leg.; ex Brassica; EMEC • 1 ♂; same data as for preceding; 7 Jun. 1949; J. W. MacSwain leg.; EMEC • 1 ♂; same data as for preceding; 13 Jun. 1949; J. W. MacSwain leg.; ex Asclepias; EMEC • 1 ♀; same data as for preceding; 20 May 1949; R. F. Smith leg.; ex Convolvulus; EMEC • 1 ♀; San Luis Obispo Co., Oso Flaco Lake; [35.029 ° N, 120.624 ° W]; 14 Jul. 1965; J. Powell leg.; EMEC • 1 ♀; San Luis Obispo Co., 10 mi. SE of Creston; [35.416 ° N, 120.398 ° W]; 25 Apr. 1968; J. Powell leg.; EMEC • 1 ♀; San Luis Obispo Co., 10 mi. W of Simmler; [35.351 ° N, 120.163 ° W]; 5 May 1962; P. D. Hurd leg.; ex Malacothrix californica; EMEC • 1 ♀; San Luis Obispo Co., Cayucos Creek; [35.449 ° N, 120.908 ° W]; 8 Aug. 1962; J. C. Hall leg.; UCRC • 1 ♀; San Luis Obispo Co., Dune Lakes, 3 mi. S of Oceano; [35.055 ° N, 120.612 ° W]; 11 Jul. 1973; R. Coville leg.; EMEC • 1 ♀; San Luis Obispo Co., Grover City; [35.12 ° N, 120.61 ° W]; 28 Apr. 1959; J. W. MacSwain leg.; ex Camissoniopsis cheiranthifolia (verbatim: Oenothera cheiranthifolia var. cheiranthifolia); EMEC • 1 ♀; San Luis Obispo Co., Morro Bay; [35.367 ° N, 120.847 ° W]; 2 May 1962; C. A. Toschi leg.; EMEC • 2 ♀♀; San Luis Obispo Co., Oso Flaco Lake, 5 mi. S of Oceano; [35.029 ° N, 120.620 ° W]; 27 Apr. 1968; J. A. Chemsak leg.; EMEC • 1 ♂; San Luis Obispo Co., Oso Flaco Lake, Costal dunes; [35.029 ° N, 120.626 ° W]; 29 Jun. 1967; M. E. Irwin, T. Cronin and S. Larisch leg.; UCRC • 1 ♀; San Luis Obispo Co., Pismo; [35.145 ° N, 120.639 ° W]; 8 May 1936; I. McCracken leg.; ex morning glory; CAS • 1 ♀; San Mateo Co., Bean Hollow; [37.23 ° N, 122.42 ° W]; 4 Aug. 1960; P. S. Bartholomew leg.; CAS • 1 ♀; San Mateo Co., Menlo Park; [37.453 ° N, 122.183 ° W]; 29 Jul. 1937; T. X. Alillium leg.; UCRC • 1 ♀; San Mateo Co., Millbrae; [37.6 ° N, 122.401 ° W]; 1 Sep. 1912; E. C. Van Dyke leg.; CAS • 1 ♀; San Mateo Co., Redwood City; [37.483 ° N, 122.236 ° W]; 18 Aug. 1956; R. R. Snelling leg.; EMEC • 1 ♂; same data as for preceding; 15 Jun. 1946; P. H. Arnaud leg.; EMEC • 1 ♀; San Mateo Co., Salada Beach; [37.63 ° N, 122.49 ° W]; 21 Apr. 1918; E. P. Van Duzee leg.; CAS • 1 ♀; San Mateo Co., San Bruno; [37.625 ° N, 122.425 ° W]; 12 Jul. 1965; R. P. Allen leg.; CSCA • 1 ♀; Santa Barbara Co., 3 mi. N of Refugio Beach; [34.504 ° N, 120.074 ° W]; 21 Jun. 1965; J. Powell leg.; EMEC • 1 ♂; Santa Barbara Co., 2 mi. E of Solvang; [34.595 ° N, 120.102 ° W]; 28 Jun. 1965; R. W. Gear leg.; EMEC • 1 ♀; same data as for preceding; 28 Jun. 1965; J. Powell leg.; EMEC • 1 ♂; Santa Barbara Co., Cañada del Puerto, Santa Cruz Island; [34.019 ° N, 119.681 ° W]; 15 Feb. 1964; T. Fisher leg.; UCRC • 1 ♀; Santa Barbara Co., Carpinteria; [34.399 ° N, 119.519 ° W]; 14 Aug. 1957; P. D. Hurd leg.; ex Hemizonia; EMEC • 1 ♀; Santa Barbara Co., Coast; [34.423 ° N, 119.645 ° W]; W. M. Giffard leg.; CAS • 1 ♀; Santa Barbara Co., Goleta; [34.436 ° N, 119.828 ° W]; 23 Jun. 1959; C. A. Campbell leg.; EMEC • 1 ♀; same data as for preceding; 22 Jun. 1959; C. A. Campbell leg.; EMEC • 1 ♂; Santa Barbara Co., Goleta, U. C; [34.434 ° N, 119.814 ° W]; 30 Jun. 1965; J. Powell leg.; EMEC • 1 ♀; Santa Barbara Co., Montecito; [34.433 ° N, 119.633 ° W]; 3 Jun. 1927; P. H. Timberlake leg.; UCRC • 1 ♀; same data as for preceding; 3 Jun. 1927; P. H. Timberlake leg.; ex Eriophyllum confertiflorum; UCRC • 2 ♀♀; Santa Barbara Co., Potter hotel (near hedge); [34.414 ° N, 119.692 ° W]; 3 Jun. 1915; Harold Morrison leg.; USNM • 1 ♀; Santa Barbara Co., Prisoner’s Harbor, Santa Cruz Island; [34.019 ° N, 119.685 ° W]; 15 Sep. 1964; E. I. Schlinger leg.; UCRC • 1 ♀; Santa Barbara Co., Santa Cruz Island; [34.004 ° N, 119.726 ° W]; 17 Aug. 1939; L. M. Martin leg.; EMEC • 1 ♀; same data as for preceding; 14 Aug. 1939; UCRC • 1 ♀; Santa Clara Co., Fleming Avenue; [37.368 ° N, 121.806 ° W]; 9 Jul. 1953; O. H. Swezey leg.; OSAC • 1 ♀; Santa Clara Co., Palo Alto; [37.429 ° N, 122.138 ° W]; 4 Sep. 1921; E. P. Van Duzee leg.; CAS • 1 ♀; same data as for preceding; 1 Aug. 1891; EMEC • 1 ♂; Santa Clara Co., Salt Marshes, Palo Alto; [37.461 ° N, 122.102 ° W]; 21 Sep. 1913; UCRC • 1 ♀; Santa Clara Co., San Francisquito Creek, Stanford University; [37.439 ° N, 122.18 ° W]; 19 Jul. 1910; EMEC • 2 ♀♀; Santa Clara Co., San Jose; [37.336 ° N, 121.890 ° W]; USNM • 1 ♀; same data as for preceding; 1910; EMEC • 1 ♀; same data as for preceding; S. Graenicher leg.; MCZ • 1 ♀; Santa Clara Co., Stanford University; [37.43 ° N, 122.17 ° W]; 25 Apr. 1961; P. H. Arnaud Jr leg.; CAS • 1 ♀; same data as for preceding; 1 – 30 May 1930; EMEC • 2 ♀♀; same data as for preceding; 29 Jun. 1910; EMEC • 1 ♀; same data as for preceding; 14 Aug. 1917; ex Centaurea imperialis; EMEC • 1 ♀; same data as for preceding; 24 Jun. 1910; EMEC • 2 ♂♂; Santa Clara Co.; [37.29 ° N, 121.74 ° W]; Baker leg.; USNM • 2 ♀♀; same data as for preceding; EMEC • 1 ♂; same data as for preceding; EMEC • 1 ♀; Santa Cruz Co., Mission Springs; [37.066 ° N, 122.032 ° W]; 25 Jul. 1956; M. D. Stage leg.; EMEC • 1 ♀; Santa Cruz Co., Watsonville; [36.920 ° N, 121.764 ° W]; Aug. 29; J. B. U. Cockwell leg.; ex Aster; UCRC • 1 ♀; Shasta Co., 5 mi. E of Burney; [40.8811 ° N, 121.565 ° W]; 9 Jun. 1941; C. D. Michener leg.; EMEC • 1 ♀; Shasta Co., Hat Creek; [40.79 ° N, 121.474 ° W]; 1 Jun. 1941; C. D. Michener leg.; EMEC • 1 ♀; Shasta Co., Redding; [40.583 ° N, 122.367 ° W]; 4 Jul. 1975; CSCA • 1 ♂; Shasta Co., Redding; [40.583 ° N, 122.367 ° W]; 1 Jul. 1975; T. R. Haig leg.; CSCA • 1 ♀; Shasta Co., Redding to Igo; [40.54 ° N, 122.4683 ° W]; elev. 304.8 m; 23 Jul. 1941; H. A. Scullen leg.; OSAC • 1 ♂; same data as for preceding; 23 Jul. 1941; H. A. Scullen leg.; OSAC • 1 ♀; Shasta Co., Shingletown; [40.496 ° N, 121.864 ° W]; 2 Jun. 1941; EMEC • 1 ♀; Shasta Co., Viola; [40.5179 ° N, 121.678 ° W]; 27 Jun. 1947; T. F. Leigh leg.; EMEC • 1 ♀; Shasta Co., Whiskeytown; [40.635 ° N, 122.561 ° W]; 21 Jun. 1975; T. R. Haig leg.; CSCA • 1 ♀; Siskiyou Co., 1 mi. SE of Bartle; [41.247 ° N, 121.8076 ° W]; 8 – 10 Jun. 1974; R. Coville leg.; EMEC • 1 ♀; same data as for preceding; 8 – 10 Jun. 1974; J. Chemask leg.; EMEC • 1 ♀; Siskiyou Co., Montague; [41.7283 ° N, 122.5274 ° W]; 1 Jun. 1966; R. P. Allen leg.; CSCA • 1 ♀; Solano Co., 4 mi. NE of Benicia; [38.091 ° N, 122.103 ° W]; 12 Sep. 1966; J. Powell leg.; EMEC • 1 ♀; Solano Co., Benicia; [38.063 ° N, 122.156 ° W]; 14 Aug. 1910; J. C. Bridwell leg.; USNM • 1 ♀; Sonoma Co., 2 mi. N of Two Rock; [38.293 ° N, 122.794 ° W]; 28 Apr. 1978; R. W. Thorp leg.; ex Limnanthes douglasii; UCDC • 1 ♀; Sonoma Co., Cazadero; [38.533 ° N, 123.085 ° W]; 29 Mar. 1935; R. M. Bohart and G. E. Bohart leg.; CAS • 1 ♂; Sonoma Co., Pitkin Ranch, 6 mi. N of Sebastopol; [38.4868 ° N, 122.824 ° W]; 25 Jul. 1965; J. W. MacSwain leg.; ex Clarkia imbricata; EMEC • 1 ♀; Sonoma Co., Sonoma Coast State Park, Shell Beach; [38.4174 ° N, 123.1056 ° W]; 5 May 1962; J. Doyen leg.; EMEC • 1 ♀; Stanislaus Co., 10 mi. SW of Turlock; [37.391 ° N, 120.975 ° W]; 12 May 1952; R. R. Snelling leg.; EMEC • 1 ♀; Stanislaus Co., Adobe Creek W; [37.3958 ° N, 121.3751 ° W]; 20 Apr. 1945; EMEC • 3 ♀♀; Stanislaus Co., Del Puerto Canyon; [37.488 ° N, 121.207 ° W]; 7 May 1970; R. R. Snelling leg.; ex Centaurea; EMEC • 1 ♀; Stanislaus Co., La Grange; [37.664 ° N, 120.464 ° W]; 30 May 1962; R. P. Allen leg.; EMEC • 1 ♀; Stanislaus Co., San Joaquin River, Newman; [37.342 ° N, 120.979 ° W]; 1 – 30 Jun. 1916; J. M. Aldrich leg.; USNM • 1 ♀; Stanislaus Co., Turlock; [37.495 ° N, 120.847 ° W]; 8 Jul. 1956; M. D. Stage leg.; EMEC • 1 ♀; same data as for preceding; 22 Jun. 1952; ex Heliotropium curassavicum; EMEC • 1 ♀; same data as for preceding; 15 Apr. 1951; Snelling leg.; EMEC • 1 ♀; same data as for preceding; 31 Jul. 1952; R. R. Snelling leg.; EMEC • 2 ♀♀; same data as for preceding; 3 May 1952; R. R. Snelling leg.; EMEC • 1 ♀; same data as for preceding; 23 Mar. 1953; R. R. Snelling leg.; ex Eschscholzia californica; EMEC • 1 ♀; same data as for preceding; 29 Apr. 1953; R. R. Snelling leg.; ex Rubus; EMEC • 1 ♀; same data as for preceding; 6 Apr. 1955; R. R. Snelling leg.; ex Amsinckia; EMEC • 37 ♀♀; same data as for preceding; 31 Jul. 1952; R. R. Snelling leg.; ex Centromadia pungens (verbatim: Hemizonia pungens); EMEC • 1 ♀; same data as for preceding; 31 Jul. 1952; J. I. Stage leg.; ex Centromadia pungens (verbatim: Hemizonia pungens); EMEC • 1 ♀; same data as for preceding; 14 May 1952; R. R. Snelling leg.; ex Centromadia pungens (verbatim: Hemizonia pungens); EMEC • 1 ♂; same data as for preceding; 5 Jul. 1952; R. R. Snelling leg.; ex Heliotropium; EMEC • 1 ♂; same data as for preceding; 3 Jun. 1951; R. R. Snelling leg.; EMEC • 1 ♂; same data as for preceding; 22 Jun. 1952; R. R. Snelling leg.; ex morning glory; EMEC • 1 ♂; same data as for preceding; 19 Jun. 1954; R. R. Snelling leg.; ex Solidago velutina subsp. californica (verbatim: Solidago californica); EMEC • 1 ♂; same data as for preceding; 23 Nov. 1951; R. R. Snelling leg.; EMEC • 2 ♂♂; same data as for preceding; 3 Jun. 1955; P. D. Hurd leg.; EMEC • 1 ♀; Tehama Co., Lassen National Forest, Lost Creek; [40.56 ° N, 121.5 ° W]; elev. 1524 m; 11 Aug. 1966; H. G. Real leg.; CAS • 1 ♂; same data as for preceding; 11 Aug. 1966; H. G. Real leg.; CAS • 1 ♀; Tehama Co., Rio Vsta. REM; 39.878 ° N, 122.0603 ° W; 15 May 2012; N. Williams leg.; WRME • 1 ♀; Trinity Co., 7 mi. S of Trinity Center; [40.888 ° N, 122.7089 ° W]; 26 Jul. 1954; J. Powell leg.; EMEC • 1 ♀; Trinity Co., 3 mi. E of Burnt Ranch; [40.808 ° N, 123.417 ° W]; 11 Jul. 1954; R. O. Schuster leg.; EMEC • 1 ♀; Trinity Co., Carrville; [41.065 ° N, 122.704 ° W]; 15 May 1934; E. C. Van Dyke leg.; CAS • 1 ♀; same data as for preceding; 19 Jun. 1951; E. C. Van Dyke leg.; CAS • 1 ♀; same data as for preceding; 8 Jun. 1913; E. C. Van Dyke leg.; CAS • 2 ♀♀; same data as for preceding; 17 Jun. 1913; E. C. Van Dyke leg.; CAS • 1 ♀; Trinity Co., Junction City; [40.74 ° N, 123.06 ° W]; 21 Apr. 1977; T. R. Haig leg.; CSCA • 1 ♀; Tulare Co., 3 mi. W of Three Rivers; [36.437 ° N, 118.958 ° W]; 1 Apr. 1939; P. H. Timberlake leg.; ex Anisocoma acaulis; UCRC • 1 ♀; Tulare Co., Lemon Cove; [36.381 ° N, 119.026 ° W]; 29 Mar. 1947; P. D. Hurd leg.; EMEC • 2 ♀♀; Tulare Co., Wood Lake; [36.414 ° N, 119.099 ° W]; 10 Jul. 1947; N. W. Frazier leg.; EMEC • 1 ♀; same data as for preceding; 29 May 1947; N. W. Frazier leg.; EMEC • 1 ♂; same data as for preceding; 6 Jun. 1935; A. L. Melander leg.; MCZ • 1 ♀; Ventura Co., Santa Paula; [34.356 ° N, 119.068 ° W]; 23 Jul. 1923; D. Burk leg.; EMEC • 1 ♀; Ventura Co., Saticoy; [34.283 ° N, 119.149 ° W]; 24 May 1926; EMEC • 1 ♀; same data as for preceding; 19 May 1926; EMEC • 1 ♀; same data as for preceding; 1 Aug. 1924; USNM • 1 ♀; Yolo Co., Davis; [38.554 ° N, 121.738 ° W]; 24 Nov. 1935; R. M. Bohart leg.; MCZ • 1 ♀; Yolo Co., Yolo; 38.605833 ° N, 121.968611 ° W; 22 Jun. 2011; S. Kaiser leg.; ex Sonchus oleraceus; WRME • 1 ♀; Igo to Oak; [39.9 ° N, 122.1 ° W]; 23 Jul. 1941; H. A. Scullen leg.; ex Eryngium articulatum; OSAC • 1 ♀; R. P. Allen leg.; CSCA. – Idaho • 1 ♀; Canyon Co., Deer Flat National Wildlife Refuge, DFO; 43.562 ° N, 116.66801 ° W; 12 Apr. 2012; A. Mohler leg.; OSAC • 4 ♀♀; Canyon Co., Deer Flat National Wildlife Refuge, DFS; 43.55281 ° N, 116.64137 ° W; 5 – 6 Jul. 2012; S. Seidel leg.; OSAC (Fig. 66) • 1 ♀; Camas Co., Fairfield; [43.346 ° N, 114.791 ° W]; elev. 1543.8 m; 3 Aug. 1928; USNM • 3 ♀♀; Gooding Co., Wood River, 1 mi. NE of Gooding; [42.9483 ° N, 114.699 ° W]; 6 – 7 Jul. 1980; M. S. Wasbauer and K. M. Wasbauer leg.; CSCA. – Nevada • 1 ♀; Washoe Co., 1 mi. N of Washoe; [38.32 ° N, 122.736 ° W]; 3 Sep. 1964; A. G. Raske leg.; EMEC • 1 ♀; Washoe Co., 10 mi. S of Reno on hyw 395; [39.625 ° N, 119.913 ° W]; 19 Jun. 1959; G. I. Stage leg.; ex Camissonia claviformis subsp. cruciformis (verbatim: Oenothera clavaeformis var. cruciformis); EMEC • 1 ♀; Washoe Co., Patrick; [39.547 ° N, 119.578 ° W]; 16 Jun. 1964; R. E. Scott leg.; UCRC • 1 ♀; Washoe Co., Sparks; [39.554 ° N, 119.736 ° W]; 28 Jun. 1927; E. P. Van Duzee leg.; CAS. – Oregon • 1 ♀; Baker Co., Haines; [44.912 ° N, 117.940 ° W]; 22 May 1935; H. P. Lanohester leg.; EMEC • 2 ♀♀; Benton Co., Corvallis; [44.567 ° N, 123.283 ° W]; 11 Jun. 1925; E. P. Van Duzee leg.; CAS • 1 ♀; same data as for preceding; 15 Jun. 1936; G. Ferguson leg.; EMEC • 1 ♀; same data as for preceding; 18 Apr. 1945; OSAC • 1 ♀; same data as for preceding; 14 Apr. 1929; H. A. Scullen leg.; ex Taraxacum officinale (verbatim: Taraxacum vulgare); OSAC • 1 ♀; same data as for preceding; 9 Aug. 1900; OSAC • 1 ♀; same data as for preceding; 24 May 1935; G. Ferguson leg.; OSAC • 1 ♀; same data as for preceding; 5 Aug. 1935; G. Ferguson leg.; OSAC • 1 ♀; same data as for preceding; 2 Jun. 1935; G. Ferguson leg.; USNM • 1 ♀; same data as for preceding; 10 May 1936; N. P. Larson leg.; USNM • 1 ♀; same data as for preceding; 6 Jun. 1998; USNM • 1 ♀; same data as for preceding; 7 Aug. 1998; USNM • 1 ♀; same data as for preceding; 8 Jun. 1998; USNM • 1 ♀; Benton Co., Corvallis, Ore. Exp. Sta; [44.566 ° N, 123.277 ° W]; 25 Apr. 1926; H. A. Scullen leg.; OSAC • 1 ♀; Benton Co., Monroe; [44.315 ° N, 123.297 ° W]; 19 Jun. 1930; G. Bennett leg.; OSAC • 2 ♀♀; Columbia Co., Scappoose; [45.754 ° N, 122.875 ° W]; 5 – 6 May 1937; OSAC • 1 ♀; same data as for preceding; 1 Jun. 1937; OSAC • 1 ♀; same data as for preceding; 28 Jun. 1937; R. E. Rieder leg.; OSAC • 2 ♀♀; Curry Co., Brookings; [42.058 ° N, 124.286 ° W]; 8 Jul. 1925; H. A. Scullen leg.; OSAC • 1 ♀; Curry Co., Gold Beach; [42.377 ° N, 124.414 ° W]; 15 Jul. 1935; OSAC • 1 ♀; Curry Co., Sixes River; [42.85 ° N, 124.54 ° W]; 13 Jul. 1925; H. A. Scullen leg.; OSAC • 1 ♀; Deschutes Co., Bend; [44.05 ° N, 121.3 ° W]; 25 Jun. 1954; W. P. Stephen leg.; OSAC • 1 ♀; Deschutes Co., Tumalo State Park; [44.129 ° N, 121.327 ° W]; 27 Jun. 1961; D. R. Smith leg.; OSAC • 1 ♀; Harney Co., 8 mi. S of Roaring Springs; [42.540 ° N, 118.986 ° W]; 21 Jun. 1947; F. Ellertson leg.; OSAC • 2 ♀♀; Harney Co., Blitzen River; [43.29 ° N, 118.82 ° W]; 6 Jul. 1906; OSAC • 2 ♀♀; Harney Co., Malheur National Wildlife Refuge, Tower; 43.2365 ° N, 118.8421 ° W; 18 – 30 Jun. 2011; T. Coe leg.; OSAC • 1 ♀; Harney Co., Steens Mountain; [42.636 ° N, 118.577 ° W]; 23 Jun. 1922; E. C. Van Dyke leg.; CAS • 4 ♀♀; Hood River Co., Hood River; [45.709 ° N, 121.512 ° W]; 12 Jun. 1920; E. C. Van Dyke leg.; CAS • 1 ♀; Jackson Co., Gold Hill; [42.434 ° N, 123.054 ° W]; 13 Jul. 1930; H. A. Scullen leg.; OSAC • 1 ♀; Jackson Co., Madford; [42.332 ° N, 122.862 ° W]; 15 Jul. 1930; H. A. Scullen leg.; OSAC • 1 ♂; same data as for preceding; 15 Jul. 1930; H. A. Scullen leg.; OSAC • 4 ♀♀; Josephine Co., Cave Junction; [42.167 ° N, 123.647 ° W]; 18 Aug. 1941; P. H. Timberlake leg.; ex Hypochaeris radicata; UCRC • 1 ♀; Josephine Co., Lake Selmac; [42.259 ° N, 123.577 ° W]; 6 Jul. 1964; P. Rude leg.; EMEC • 2 ♀♀; Klamath Co., Eagle Ridge; [42.418 ° N, 121.953 ° W]; 27 May 1924; C. L. Fox leg.; CAS • 1 ♀; Klamath Co., Eagle Ridge Klamath Lake; [42.389 ° N, 121.975 ° W]; 27 May 1924; C. L. Fox leg.; CAS • 1 ♀; Klamath Co., Eagle Ridge, Harrimans Lodge; [42.468 ° N, 122.1003 ° W]; C. L. Fox leg.; CAS • 3 ♀♀; same data as for preceding; 6 Aug. 1924; C. L. Fox leg.; CAS • 1 ♀; Klamath Co., Klamath Falls; [42.225 ° N, 121.782 ° W]; 1 – 31 Aug. 1923; EMEC • 1 ♀; Klamath Co., Lamis Mill, Eagle Ridge Klamath Lake; [42.407 ° N, 121.979 ° W]; 24 May 1924; C. L. Fox leg.; CAS • 1 ♀; Linn Co., Shedd; [44.462 ° N, 123.109 ° W]; 11 Jun. 1929; J. E. Davis leg.; OSAC • 1 ♀; Linn Co., Sodaville; [44.484 ° N, 122.868 ° W]; 1903; Herbert leg.; OSAC • 1 ♂; Linn Co.; [44.53 ° N, 122.57 ° W]; 17 Aug. 1929; J. E. Davis leg.; OSAC • 1 ♀; Marion Co., Salem; [44.939 ° N, 123.033 ° W]; 16 Aug. 1919; EMEC • 1 ♀; Multnomah Co., Portland; [45.49 ° N, 122.61 ° W]; 1 – 30 Jun. 1925; H. A. Scullen leg.; OSAC • 1 ♀; Multnomah County Co., Oxbow Meadow; 45.485 ° N, 122.302 ° W; 15 Jun. 2021; J. D. Engler leg.; ex Leucanthemum; OSAC • 1 ♀; Multnomah County Co., Oxbow Meadow; 45.4844 ° N, 122.3015 ° W; 7 Aug. 2020; J. D. Engler leg.; OSAC • 1 ♀; Polk Co., Independence; [44.853 ° N, 123.191 ° W]; 16 Jul. 1934; N. P. Larson leg.; OSAC • 2 ♀♀; Sherman Co., Maryhill Ferry, 3 mi. E of Briggs; [45.693 ° N, 120.806 ° W]; 3 Jun. 1963; P. H. Raven leg.; ex Oenothera pallida; EMEC • 19 ♀♀; same data as for preceding; 19 May 1963; P. H. Raven leg.; ex Oenothera pallida; EMEC • 2 ♀♀; Umatilla Co., Milton; [45.936 ° N, 118.391 ° W]; 22 Jun. 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♀; Wasco Co., 5 mi. S of The Dalles; [45.528 ° N, 121.185 ° W]; 5 May 1938; K. Gray and J. Schuh leg.; OSAC • 1 ♂; Wasco Co., Tygh Valley; [45.25 ° N, 121.17 ° W]; 2 Jul. 1941; R. E. Rieder leg.; ex sweet clover; OSAC. – Utah • 1 ♀; Box Elder Co., Mouth Bear River; [41.472 ° N, 112.294 ° W]; 24 Jul. 1915; A. Wetmore leg.; USNM • 1 ♀; Utah Co., Provo; [40.244 ° N, 111.66 ° W]; 1 – 31 May 1937; H. Chandler leg.; CAS • 1 ♀; Salt Lake Co., Salt Lake City; [40.76 ° N, 111.89 ° W]; S. Garman leg.; MCZ. – Washington • 1 ♀; Adams Co., Ritzville; [47.126 ° N, 118.377 ° W]; 28 Aug. 1923; M. C. Lane leg.; USNM • 1 ♂; Asotin Co., Asotin; [46.339 ° N, 117.046 ° W]; 21 Jun. 1932; J. M. Aldrich leg.; ex wild parsnip; USNM • 1 ♀; Clark Co., Ridgefield National Wildlife Refuge, Wig Lk; 45.817 ° N, 122.773 ° W; 24 May 2011; A. Chmielewski leg.; OSAC • 1 ♀; Grant Co., Grand Coulee, Osborn’s Ranch; [47.904 ° N, 119.064 ° W]; 8 Jul. 1902; USNM • 1 ♀; Island Co., Saratoga Beach; [48.087 ° N, 122.517 ° W]; 10 Jul. 1938; E. Dalley leg.; OSAC • 1 ♀; King Co., Seattle; [47.61 ° N, 122.33 ° W]; 19 Jun. 1920; E. P. Van Duzee leg.; CAS • 1 ♀; same data as for preceding; 18 May 1941; Lewis leg.; OSAC • 1 ♀; same data as for preceding; 25 Jul. 1928; ex Grindelia tarweed; OSAC • 3 ♀♀; same data as for preceding; 18 Aug. 1927; ex tarweed Grindelia; OSAC • 1 ♀; same data as for preceding; 17 May 1912; OSAC • 1 ♀; same data as for preceding; 1 – 31 May 1925; OSAC • 1 ♀; same data as for preceding; 29 Jul. 1928; ex Grindelia tarweed; OSAC • 1 ♂; same data as for preceding; 7 Aug. 1927; ex Grindelia tarweed; OSAC • 1 ♂; same data as for preceding; 18 Aug. 1927; ex Grindelia tarweed; OSAC • 1 ♂; same data as for preceding; 4 Aug. 1927; ex Chamerion angustifolium subsp. angustifolium (verbatim: fireweed, Epilobium spicatum); OSAC • 1 ♀; King Co., Seattle; 47.5131 ° N, 122.3065 ° W; 14 Jul. 2014; E. A. Sugden and W. Peterman leg.; WRME • 1 ♀; King Co., Seattle; 47.4301 ° N, 122.3051 ° W; 8 Aug. 2015; E. A. Sugden and W. Peterman leg.; WRME • 1 ♀; same data as for preceding; 17 Apr. 2016; E. A. Sugden and W. Peterman leg.; WRME • 1 ♀; same data as for preceding; 7 Aug. 2016; E. A. Sugden and W. Peterman leg.; WRME • 1 ♀; King Co., Seattle; 47.4295 ° N, 122.3048 ° W; 10 May 2017; E. A. Sugden and W. Peterman leg.; WRME • 1 ♀; Lincoln Co., Siso-pollinator baseline monitoring, Sage, Downs Rd near Tan Rd E of Lamona; 47.376919 ° N, 118.389973 ° W; elev. 547.88 m; 42571; T. D. Hatten leg.; WRME • 1 ♀; Lincoln Co., Siso-pollinator baseline monitoring, Sage, Downs Rd N., S of Coal Coule Lamona; 47.377576 ° N, 118.391616 ° W; elev. 547.88 m; 42508; T. D. Hatten leg.; WRME • 1 ♀; Pacific Co., Nahcotta; [46.50 ° N, 124.03 ° W]; 1 Aug. 1952; EMEC • 2 ♀♀; same data as for preceding; 25 Jun. 1952; EMEC • 1 ♀; same data as for preceding; 20 Jul. 1951; EMEC • 1 ♀; same data as for preceding; 15 Jun. 1952; EMEC • 1 ♀; same data as for preceding; 10 Aug. 1951; EMEC • 1 ♀; same data as for preceding; 11 Jul. 1951; EMEC • 1 ♀; same data as for preceding; 16 Jul. 1951; EMEC • 1 ♀; same data as for preceding; 30 Jul. 1951; EMEC • 1 ♀; same data as for preceding; 14 Aug. 1951; EMEC • 1 ♂; same data as for preceding; 25 Jul. 1952; EMEC • 2 ♂♂; same data as for preceding; 26 Aug. 1952; EMEC • 1 ♂; same data as for preceding; 23 Aug. 1952; EMEC • 3 ♀♀; same data as for preceding; 30 May 1953; OSAC • 8 ♀♀; same data as for preceding; 9 Jun. 1953; ex Hypochaeris radicata; OSAC • 2 ♀♀; same data as for preceding; 15 May 1954; OSAC • 1 ♀; same data as for preceding; 21 Jun. 1953; OSAC • 5 ♀♀; same data as for preceding; 16 Jun. 1953; ex Symphyotrichum subspicatum (verbatim: Aster douglassi); OSAC • 1 ♀; same data as for preceding; 14 Jun. 1953; OSAC • 2 ♀♀; same data as for preceding; 15 Jun. 1953; OSAC • 1 ♀; same data as for preceding; 5 Aug. 1954; OSAC • 1 ♀; same data as for preceding; 30 Jun. 1954; OSAC • 1 ♀; same data as for preceding; 2 Jul. 1954; ex Hypochaeris radicata; OSAC • 1 ♀; same data as for preceding; 16 Jun. 1954; OSAC • 1 ♀; same data as for preceding; 12 May 1954; OSAC • 1 ♀; same data as for preceding; 22 Jul. 1954; OSAC • 3 ♀♀; same data as for preceding; 1 Jun. 1954; ex Hypochaeris radicata; OSAC • 1 ♀; same data as for preceding; 22 Jun. 1954; OSAC • 1 ♀; same data as for preceding; 10 May 1954; OSAC • 2 ♀♀; same data as for preceding; 2 Jun. 1954; ex Hypochaeris radicata; OSAC • 1 ♀; same data as for preceding; 21 Jun. 1954; OSAC • 1 ♀; same data as for preceding; 25 Jun. 1955; OSAC • 1 ♀; same data as for preceding; 10 May 1953; OSAC • 1 ♀; same data as for preceding; 12 May 1954; SDMC • 1 ♀; same data as for preceding; 8 Aug. 1954; RBCM • 1 ♀; same data as for preceding; 11 Jul. 1954; RBCM • 1 ♀; same data as for preceding; 4 May 1953; RBCM • 1 ♀; same data as for preceding; 1 Jun. 1954; ex Hypochaeris radicata; RBCM • 1 ♀; same data as for preceding; Jul. 15; RBCM • 1 ♀; same data as for preceding; 25 Jul. 1954; RBCM • 1 ♀; same data as for preceding; 16 Jun. 1953; ex Symphyotrichum subspicatum (verbatim: Aster douglassi); RBCM • 1 ♀; same data as for preceding; 21 Jun. 1953; RBCM • 1 ♀; same data as for preceding; 13 Jul. 1953; RBCM • 1 ♀; same data as for preceding; 18 May 1954; OSAC • 1 ♂; same data as for preceding; 10 Aug. 1953; OSAC • 1 ♀; Pacific Co., Willapa National Wildlife Refuge, OSB 3; 46.3689 ° N, 124.0273 ° W; 22 Aug. 2017; W. Ritchie leg.; OSAC • 1 ♀; Pacific Co., Willapa National Wildlife Refuge, TRL; 46.36886 ° N, 124.02731 ° W; 27 Apr. 2015; W. Ritchie leg.; OSAC • 1 ♀; same data as for preceding; 16 May 2016; W. Ritchie leg.; OSAC • 1 ♀; Pierce Co., Fort Lewis; [47.109 ° N, 122.555 ° W]; 2 Aug. 1957; R. R. Snelling leg.; EMEC • 1 ♀; Spokane Co., Turnbull National Wildlife Refuge, SSA; 47.4208 ° N, 117.6203 ° W; 22 – 23 Jun. 2016; D. Cousins leg.; OSAC • 1 ♀; same data as for preceding; 27 – 28 Apr. 2016; D. Cousins leg.; OSAC • 1 ♀; Thurston Co., Olympia; [47.038 ° N, 122.901 ° W]; 10 Jun. 1895; USNM • 1 ♀; Walla Walla Co., McNary National Wildlife Refuge, WAL; 46.069 ° N, 118.8985 ° W; 1 Jul. 2011; M. Gregg leg.; OSAC • 2 ♀♀; Walla Walla Co., Walla Walla; [46.067 ° N, 118.339 ° W]; 29 May 1937; R. M. Bohart and G. E. Bohart leg.; CAS • 1 ♀; Yakima Co., N. Yakima; [46.620 ° N, 120.544 ° W]; 24 May 1903; Eldred Jenne leg.; USNM.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF70FF5CFE12FCC1FC7AFAD9.taxon	description	Description GENITALIA (Figs 68, 120 L). Gonostylus elongate, length approximately twice breadth, with setae short. Retrorse lobe broad, and curved, about 3.5 times as long as broad, pointed at apex, covered in fine, sparse setae.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF70FF5CFE12FCC1FC7AFAD9.taxon	distribution	Range Canada (British Columbia), Mexico (Baja California), USA (California, Idaho, Nevada, Oregon, Utah, Washington) (Fig. 69).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF70FF5CFE12FCC1FC7AFAD9.taxon	biology_ecology	Biological notes The robber fly Megaphorus guildianus (Williston, 1885) (Asilidae) was recorded as a predator of L. kincaidii (Moure & Hurd 1987). Floral hosts AIZOACEAE: Carpobrotus sp. • APIACEAE: Eryngium articulatum Hook., Foeniculum vulgare Mill., Pastinaca sativa L. • APOCYNACEAE: Asclepias sp., Asclepias speciosa Torr. • ASTERACEAE: Ageratum sp., Anisocoma acaulis Torr. & A. Gray, Argyranthemum frutescens (L.) Sch. Bip., Aster sp., Baccharis sp., Calendula sp., Centaurea sp., Centromadia pungens Greene, Chrysothamnus sp., Chrysothamnus viscidiflorus subsp. typicus H. M. Hall & Clem., Coreopsis lanceolata L., Coreopsis tinctoria Nutt., Cosmos sp., Deinandra sp., Encelia farinosa A. Gray ex Torr., Eriophyllum confertiflorum (DC.) A. Gray, Grindelia sp., Helenium bigelovii A. Gray, Helianthus sp., Hemizonia sp., Heterotheca grandiflora Nutt., Hypochaeris radicata L., Jensia yosemitana (Parry ex A. Gray) B. G. Baldwin, Leucanthemum × superbum (Bergmans ex J. W. Ingram) D. H. Kent, Leucanthemum sp., Malacothrix californica DC., Picris echioides L., Solidago velutina subsp. californica (Nutt.) Semple, Sonchus oleraceus L., Stephanomeria exigua Nutt., Symphyotrichum subspicatum (Nees) G. L. Nesom, Taraxacum officinale F. H. Wigg. • BORAGINACEAE: Amsinckia sp., Heliotropium sp., Heliotropium curassavicum L. • BRASSICACEAE: Brassica sp., Lobularia maritima (L.) Desv., Nasturtium officinale W. T. Aiton, Sisymbrium irio • CONVOLVULACEAE: Calystegia occidentalis (A. Gray) Brummitt, Convolvulus sp., Ipomoea purpurea (L.) Roth • CUCURBITACEAE: Cucurbita maxima Duchesne • FABACEAE: Acmispon glaber (Vogel) Brouillet, Medicago sativa L., Melilotus sp., Melilotus albus Medik. • FRANKENIACEAE: Frankenia salina (Molina) I. M. Johnst. • GERANIACEAE: Erodium moschatum (L.) L’Hér. • HYDROPHYLLACEAE: Phacelia californica Cham. • IRIDACEAE: Sisyrinchium sp. • LAMIACEAE: Mentha spicata L., Mentha sp., Monardella lanceolata A. Gray • LILIACEAE: Narcissus sp. • LIMNANTHACEAE: Limnanthes douglasii R. Br. • ONAGRACEAE: Camissonia campestris (Greene) P. H. Raven subsp. campestris, Camissoniopsis cheiranthifolia (Hornem. ex Spreng.) W. L. Wagner & Hoch, Camissonia claviformis (Torr. & Frém.) P. H. Raven subsp. cruciformis (Kellogg) P. H. Raven, Camissonia contorta (Douglas ex Lehm.) Kearney, Taraxia tanacetifolia (Torr. & A. Gray) Piper, Chamerion angustifolium (L.) Holub subsp. angustifolium, Oenothera pallida Lindl., Oenothera sp. • PAPAVERACEAE: Eschscholzia californica Cham. • POLEMONIACEAE: Gilia achilleifolia subsp. multicaulis (Benth.) A. D. Grant & V. E. Grant • POLYGONACEAE: Polygonum lapathifolium L. • RHAMNACEAE: Frangula californica (Eschsch.) A. Gray subsp. californica • ROSACEAE: Photinia arbutifolia Lindl., Rubus sp. • RUBIACEAE: Kelloggia galioides Torr. • SALICACEAE: Salix gooddingii C. R. Ball, Salix nigra Marshall, Salix sp. • SOLANACEAE: Datura inoxia Mill. • VERBENACEAE: Phyla lanceolata (Michx.) Greene, Phyla nodiflora (L.) Greene • ZYGOPHYLLACEAE R. Br.: Tribulus sp., Tribulus terrestris L.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF70FF5CFE12FCC1FC7AFAD9.taxon	discussion	DNA barcodes Three full DNA barcodes are available, all from British Columbia, Washington, and Oregon. They form a discrete cluster with little intraspecific variation. Sequences show similarity to the other members of the pectorale species group and are most similar to other members of the Lasioglossum pectorale species group, including L. nelumbonis and several Neotropical species.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF61FF63FE27FA8AFE3EFDEB.taxon	description	urn: lsid: zoobank. org: act: EE 37 F 974 - CA 4 D- 4 D 5 C-A 5 AD- 1 ED 4 E 3 CB 0729 Figs 70 – 73, 95 G, 103 A, 104 A, 111 B, 112 A, 120 M	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF61FF63FE27FA8AFE3EFDEB.taxon	diagnosis	Diagnosis Female L. pathiranae sp. nov. can be recognised by the combination of head long, length / width ratio 1.18 ± 0.01 SD (shorter in all the other similar species); mesoscutum polished (rougher in L. robertsoni), moderately dense (IS = 1 – 2 PD) puncture density submedially (sparser in L. angustius; denser in L. angustius and L. robertsoni), double punctate distinctive on anteromedially (no double punctation in L. angustius and L. robertsoni); propodeal triangle with rugae reaching posterior margin (separating from species in which the rugae do not reach the posterior margin), propodeal triangle posterior margin sharply edged with a distinct raised apical carina (this excludes all species with a smoothly rounded posterior margin); T 1 strongly coriarious on anterior face with a fan bearing a narrow dorsal opening (T 1 anterior face mostly polished with broad dorsal opening in the fan in L. angusticulum sp. nov.); and T 2 – T 3 with distinct basolateral tomentum (removes all species lacking basal patches or bands of tomentum on T 2 – T 3, such as L. nitidiusculum group). Males L. pathiranae sp. nov. can be recognised by the combination of head long, length / width ratio 1.12 (± 0.05 SD) (shorter in all the other similar species); female-like antennae mesoscutum mostly polished with dense (IS ≤ 1 PD) punctures (moderately sparser (IS = 1 – 2 PD) in L. angusticulum sp. nov.); metapostnotal rugae reaching posterior margin (separating from species in which the rugae do not reach the posterior margin); and propodeal triangle with prominently raised apical carina (indistinct in L. angusticulum).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF61FF63FE27FA8AFE3EFDEB.taxon	etymology	Etymology The species epithet pathiranae is a heartfelt tribute to TH’s loving mother, Padmini Pathirana. This new bee species is named in her honour, a meaningful gesture to express TH’s deep appreciation and love.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF61FF63FE27FA8AFE3EFDEB.taxon	materials_examined	Material examined Holotype UNITED STATES – California • ♀; Riverside Co., Tabquitz Canyon; [33.501 ° N, 116.308 ° W]; 8 Jun. 1957; Menke, Stange and Bromley leg.; LACM (Fig. 70). Specimen labels “ June 8, 1957 / Tabquitz Canyon / Riverside Co., Cal. / Menke & Stange & Bromley // Lasioglossum GIBBS – 14412 // HOLOTYPE / Lasioglossum (Hemihalictus) pathiranae Hettiarachchi and Gibbs ”. Paratypes MEXICO – Baja California Sur • 2 ♂♂; Sierra Laguna, Big Canyon; [23.567 ° N, 110.000 ° W]; 13 Oct. 1941; E. Ross and G. Bohart leg.; CAS (Fig. 72). UNITED STATES – Arizona • 1 ♂; Graham Co., Thatcher; [32.833 ° N, 109.763 ° W]; 11 Aug. 1958; Jon McBride leg.; ASUHIC (Fig. 71) • 1 ♀; same data as for preceding; 11 Aug. 1958; Jon McBride leg.; ASUHIC • 1 ♂; Pima and Santa Cruz Co., Santa Rita Mountains; [31.826 ° N, 110.775 ° W]; elev. 1219.2 m; 1 – 31 Oct. 1927; C. T. Vorhies leg.; ASUHIC • 1 ♀; same data as for preceding; 14 Jun. 1942; elev. over 1524 m; H. A. Scullen leg.; OSAC • 2 ♀♀; Pima Co., Tucson; [32.222 ° N, 110.926 ° W]; 13 Mar. 1955; F. Werner leg.; ASUHIC • 1 ♀; same data as for preceding; 15 – 18 Jun. 1962; G. D. Butler leg.; ASUHIC • 1 ♀; same data as for preceding; 19 – 26 Aug. 1962; G. D. Butler leg.; ASUHIC • 1 ♀; same data as for preceding; 19 May 1955; G. D. Butler leg.; ex Physaria gordonii (verbatim: Lesquerella gordonii); ASUHIC • 1 ♀; same data as for preceding; 30 May 1954; G. D. Butler leg.; ASUHIC • 1 ♀; same data as for preceding; 25 Sep. 1923; ASUHIC • 1 ♀; Pima Co., Tucson, 1207 E Speedway; [32.236 ° N, 110.953 ° W]; 11 Feb. 1962; J. C. Bequaert leg.; ex Taraxacum officinale; ASUHIC • 1 ♀; Pima Co., Tucson; [32.222 ° N, 110.926 ° W]; 3 Oct. 1923; ASUHIC • 1 ♀; Pima Co., Sabino Canyon; [32.322 ° N, 110.810 ° W]; 30 May 1953; G. D. Butler leg.; ASUHIC • 1 ♀; Yavapai Co., Atascosa Mountains, Sycamore Canyon; [31.432 ° N, 111.188 ° W]; 14 Jun. 1968; Flint and Menke leg.; ASUHIC. – California • 1 ♀; Riverside Co., Whitewater Canyon; [33.968 ° N, 116.652 ° W]; 26 Jul. 1963; E. I. Schlinger leg.; LACM • 1 ♀; Riverside Co., Palm Springs; [33.8246 ° N, 116.5403 ° W]; 3 Apr. 1925; E. C. Van Dyke leg.; CAS • 1 ♀; Riverside Co., Bautista Canyon near Hemet; [33.698 ° N, 116.853 ° W]; 24 Apr. 1972; Ali Tabet leg.; UCRC. – Texas • ♀; El Paso Co., Santa Fe Bridge; [31.748 ° N, 106.487 ° W]; 19 Feb. 1945; ex strawberry; USNM. Other material excluded from paratypes UNITED STATES – California • 1 ♀; Orange Co., Starr Ranch Audubon Sanctuary, Santa Ana Mountains; [33.632 ° N, 117.555 ° W]; elev. 274.3 m; 17 – 20 Jun. 1980; D. C. Michener leg.; ex Keckiella cordifolia; LACM.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF61FF63FE27FA8AFE3EFDEB.taxon	description	Description Female BODY MEASUREMENTS. Length 7.92 mm (± 0.65 SD). Head length 2.01 mm (± 0.05 SD). Face length 1.65 mm (± 0.03 SD). Head width 1.69 mm (± 0.04 SD). Wing length 5.04 mm (± 0.14 SD). n = 5. COLOURATION. Head and mesosoma dark brown. Clypeus apex dark brown to brown. Labrum brown. Mandible brown basally, becoming light brown medially and reddish-brown apically. Flagellum brown dorsally, light brown ventrally. Pronotal lobe brown. Tegula light brown, sometime with brown base. Wing membrane faintly brownish, veins with subcosta brown, otherwise light brown. Legs brown. Metasoma dark brown or brown with rims of terga, sterna and downcurved lateral areas of terga broadly light brown. PUBESCENCE. Body hair colour off-white. Tomentum dense on pronotal angle and lobe, space between pronotal lobe and tegula, metanotum anteromedially, propodeum lateral surface anterodorsally, and T 2 – T 4 basolaterally; moderately dense on anterolateral angle and marginally on mesoscutum; very sparse on supraclypeal area dorsolaterally, paraocular area, gena near eye margin, propodeum lateral (except anterodorsal area) and posterior surface mostly on dorsally. Mesosoma covered by appressed, simple, very short hairs (<0.25 OD) mixed with sparse, erect, weakly plumose, long hairs (<3 OD) on mesoscutum and erect, plumose, long hairs (<2.5 OD) on other areas. Metasomal sterna with moderately sparse, erect, weakly plumose, long (<3 OD) hairs. Metafemoral scopa with dense plumose hairs. Wing hairs brown, very short and sparse. T 1 with a fan, narrow dorsal opening (1 OD), composed of overlapping sub-appressed simple hairs (<1.5 OD). SURFACE SCULPTURING. Clypeus imbricate to tessellate, becoming weakly imbricate apical ⅓; punctures irregularly sparse (IS <4 PD) apically and medially, becoming smaller and denser (IS <1 PD) basolaterally. Supraclypeal area tessellate; punctures fine, dense (IS <1 PD). Frons reticulate-punctate with an imbricate microsculpture present throughout the surface; punctures crowded (IS = 0 PD). Ocellocular area polished; punctures dense (IS ≤ 1 PD) and diversopunctate posteriorly, becoming crowded (IS = 0 PD) anteriorly. Gena rugulose to weakly lineolate, becoming lineolate posteriorly; punctures dense (IS <1 PD) or obscure in some angles. Postgena weakly imbricate to weakly lineolate. Tegula punctures sparse (IS = 1 – 3 PD) in the anterior half and basally, becoming impunctate posteriorly. Mesoscutum polished, becoming imbricate anteromedially and on median line; punctures dense (IS <1 PD), becoming moderately dense (IS = 1 – 2 PD) submedially, doubly punctate, but only conspicuous on anteromedially. Mesoscutellum polished, becoming rugulose posteriorly and laterally; punctures dense (IS ≤ 1 PD) marginally and on median line, sparse submedially (IS = 1 – 4 PD), diversopunctate. Metanotum rugulose, punctures obscure. Propodeal triangle with rugae strong, reaching posterior margin, strongly anastomosing; interspace sculpture shiny to weakly imbricate. Preëpisternum reticulate-rugose. Hypoepimeron weakly imbricate, but somewhat polished ventrally, becoming rugulose dorsally; punctures dense (IS <1 PD). Mesepisternum reticulate-rugose, becoming rugose ventrally; impunctate. Propodeum lateral face strongly imbricate, becoming rugulose posteriorly; posterior face rugose, punctures obscure. T 1 anterior face dull, strongly coriarious, punctures present laterally; dorsal surface mostly weakly coriarious to coriarious, punctures sparse (IS = 1 – 3 PD), becoming more sparsely punctate (IS = 2 – 6 PD) in small subapicolateral boss and punctures fine, minute, and sparse (IS = 1 – 4 PD) on rim, becoming impunctate apical third and medially on rim. T 2 disc coriarious to polished, punctures moderately dense (IS = 1 – 2 PD) basally, becoming sparse (IS = 1 – 3 PD) apically; rim coriarious, punctures minute and sparse (IS = 1 – 3 PD), becoming impunctate narrowly after premarginal line. STRUCTURE. Head long, length / width ratio 1.18 (± 0.01 SD). Face length / head width ratio 0.97 (± 0.02 SD). Clypeus projecting ~ 89 % below suborbital tangent; clypeus length / width ratio 1.10 (± 0.08 SD). Mandible long, nearly reaching opposing mandibular base. Eyes convergent below [UOD / LOD ratio 1.16 (± 0.04 SD)]. Ocelli normal. Gena / eye width ratio 0.83 (± 0.34 SD). Pronotal angle obtuse. Intertegular span 1.40 mm (± 0.11 SD). Mesoscutum length / width ratio 0.90 (± 0.06 SD); mesoscutum / scutellum length ratio 2.98 (± 0.13 SD); scutellum / metanotum length ratio 1.55 (± 0.14 SD); metanotum / propodeal triangle length ratio 0.88 (± 0.16 SD). Protibial spur with apical serrations longer or subequal in length to width of malus; malus not reaching much beyond velum. Inner metatibial spur pectinate, teeth 3 – 4, basal teeth longer than width of rachis. Propodeal triangle posterior margin angled onto posterior propodeal surface; apical carina well distinct and high. Propodeal lateral carina reaching dorsolateral slope; oblique carina obscure from background sculpturing. n = 6. VARIATION. One female we observed from Orange County has an exceptionally shiny mesoscutum. Male BODY MEASUREMENTS. Length 5.55 mm (± 0.56 SD). Head length 1.65 mm (± 0.05 SD). Face length 1.42 mm (± 0.08 SD). Head width 1.48 mm (± 0.09 SD). Wing length 4.33 mm (± 0.23 SD). n = 3. COLOURATION. Head and mesosoma dark brown to black. Clypeus brown. Labrum dark brown. Mandible dark brown basally, becoming light brown medially and slightly reddish brown apically. Flagellum dark brown dorsally, brown ventrally. Pronotal lobe brown. Tegula mostly dark brown, light brown medially. Wing membrane faintly brownish, veins mostly brownish. Legs brown. Metasoma dark brown to brown with rims of terga, sterna and downcurved lateral areas of terga narrowly light brown. PUBESCENCE. Body hair colour off-white. Tomentum dense on pronotal angle, and lobe, space between pronotal lobe and tegula, metanotum anteromedially, and T 2 – T 3 basolaterally; sparse on paraocular area and propodeum posterior face. Mesosoma covered by appressed to erect, simple, very short hairs (<0.25 OD) mixed with very sparse, erect, weakly plumose, long hairs (<1 OD) on mesoscutum and erect, plumose, long hairs (<2.5 OD) on other areas. Metasomal sterna with dense, erect and sub-appressed, plumose, long (<2 OD) hairs. Wing hairs brown, very short and sparse. SURFACE SCULPTURE. Clypeus polished, becoming imbricate basally and sometime polished throughout; punctures irregularly sparse (IS <5 PD) apically and medially, becoming denser (IS <2 PD) basolaterally. Supraclypeal area imbricate, becoming weakly so medially; punctures dense (IS <1 PD), becoming little bit sparser anteromedially (IS ≤ 1 PD). Frons reticulate-punctate; punctures crowded (IS = 0 PD). Ocellocular area polished; punctures moderately dense (IS = 1 – 2 PD) and diversopunctate posteriorly, becoming crowded (IS = 0 PD) anteriorly. Gena weakly lineolate, becoming lineolate posteriorly; punctures dense (IS <1 PD), becoming obscure posteriorly. Postgena weakly lineolate. Tegula broadly impunctate, minute punctures present anteriorly and basally, sparse (IS = 1 – 3 PD). Mesoscutum polished, becoming imbricate anteromedially and halfway on median line; punctures dense (IS ≤ 1 PD), doubly punctate (or diversopunctate). Mesoscutellum polished, becoming rugulose posteriorly; punctures dense (IS ≤ 1 PD) marginally and on median line, sparse submedially (IS = 1 – 3 PD), diversopunctate. Metanotum rugulose, punctures obscure. Propodeal triangle with rugae strong, reaching posterior margin and join with sharply carinate posterior border, moderately anastomosing medially and weakly so laterally; interspace sculpture shiny. Preëpisternum reticulate-rugose. Hypoepimeron rugulose dorsally, becoming weakly so ventrally; punctures crowded (IS = 0 PD). Mesepisternum reticulate-rugose, becoming rugose ventrally. Propodeum lateral face rugulose, becoming rugose posteriorly; posterior face strongly rugose. T 1 anterior face polished, punctures present laterally; dorsal surface polished, punctures sparse (IS = 1 – 3 PD), becoming sparser towards premarginal line (IS = 1 – 4 PD) and becoming nearly impunctate in large subapicolateral boss; punctures minute, and sparse (IS = 1 – 4 PD) on rim, becoming impunctate apical half and impunctate medially on rim. T 2 disc polished, punctures sparse (IS = 1 – 3 PD) basally, becoming sparse (IS = 1 – 4 PD) apically and nearly impunctate or very sparse on subapicolateral boss (IS = 2 – 6 PD); rim polished, punctures minute and sparse (IS = 1 – 3 PD), becoming narrowly impunctate apically. STRUCTURE. Head long, length / width ratio 1.12 (± 0.05 SD). Face length / width ratio 0.96 (± 0.02 SD). Clypeus projecting ~ 74 % below suborbital tangent. Mandible long, just reaching opposing mandibular base. Flagellomeres short, F 1 length / pedicel length ratio 1.05 (± 0.10 SD); F 2 length / F 1 length ratio 1.18 (± 0.16 SD); F 2 length / F 2 width ratio 1.02 (± 0.08 SD). Gena / eye width ratio 0.87 (± 0.04 SD). Forewing with 3 submarginal cells. Pronotal angle obtuse, quite sharp angle. Intertegular span 0.99 mm (± 0.15 SD). Mesoscutum length / width ratio 0.95 (± 0.01 SD). Propodeal triangle posterior border sharply carinate, well distinct and high. Propodeum lateral carinae reaching dorsolateral slope; oblique carina obscure from background sculpturing. n = 3. GENITALIA (Figs 72, 120 M). Gonostylus elongate, approximately 3 times as long as broad, bearing short setae. Retrorse lobe broad, about 5 times as long as broad, covered in short sparse setae.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF61FF63FE27FA8AFE3EFDEB.taxon	distribution	Range Southern California through to southern Arizona, west of Texas (Fig. 73).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF61FF63FE27FA8AFE3EFDEB.taxon	biology_ecology	Floral hosts ASTERACEAE: Taraxacum officinale F. H. Wigg • BRASSICACEAE: Physaria gordonii (A. Gray) O’Kane & Al-Shehbaz • ROSACEAE: Fragaria sp. • SCROPHULARIACEAE: Keckiella cordifolia (Benth.) Straw.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF5EFF6CFE7FFDC4FCD0FC07.taxon	description	Figs 74 – 77, 93 D, 101 D, 104 G, 109 C, 111 C, 120 N	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF5EFF6CFE7FFDC4FCD0FC07.taxon	etymology	Etymology Cockerell (1895) states that this species has been “ confounded with H. pectoralis, which it much resembles ”. He contrasts the two directly in his description. The suffix ‘ - oides ’, means ‘ resembling’.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF5EFF6CFE7FFDC4FCD0FC07.taxon	diagnosis	Diagnosis Both sexes of L. pectoraloides can be easily recognised by the combination of mesepisternum shiny with a combination of rugulose and distinct punctation (rugose in L. pectorale and L. kincaidii); tegula ovoid (enlarged in L. kincaidii); metasoma with basolateral tomentum (absent in L. sequoia e and L. villosulum); propodeal triangle with rugae reaching posterior margin, this margin with a distinct apical carina forming a sharp edge; and apical impressed areas polished and almost impunctate (in contrast to L. pectorale).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF5EFF6CFE7FFDC4FCD0FC07.taxon	materials_examined	Type material Syntype of Halictus pectoraloides ♀, not examined; current location unknown. Holotype of Halictus pectoraloides var. beatulus (by original designation; examined by photos) UNITED STATES – New Mexico • ♀; Doña Ana Co., Mesilla; [32.27 ° N, 106.80 ° W]; 25 Jun.; T. D. Cockerell leg.; ex Chloracantha spinosa (verbatim: Aster spinosus); USNM Type No. 22953. Specimen labels “ Mesilla / June 25. / (Ckll) // on Aster spinosus // Type No. / 22953 / U. S. N. M. // Halictus / pectoraloides / v. beatulus / Ckll. TYPE // USNMENT / 01735665 ”. Holotype of Halictus (Evylaeus) grindeliae (by original designation; not examined) UNITED STATES – Colorado • 1 ♀; Hudson; 1 Oct. 1933; UCMC. Type notes Cockerell (1895) stated that L. pectoraloides was very abundant and recorded it from Mesilla Valley and Las Cruces, New Mexico, but did not explicitly designate a holotype. The location of the type series is unknown. Cockerell (1918) later documented the variety beatulus from Mesilla and Las Cruces, New Mexico, and referred specifically to a single type specimen. Subsequently, Cockerell (1934) reported the location of a type specimen of Halictus grindeliae and gave no indication that additional type specimens existed. On this basis, we interpret the name as having been established from a single specimen and therefore treat it as based on a holotype by monotypy. Material examined MEXICO – Chihuahua • 1 ♀; Villa Ahumada, 3 mi. S of Chihuahua; [30.5645 ° N, 106.5058 ° W]; 10 Jul. 1964; J. Powell leg.; EMEC. – Sonora • 1 ♀; 10 mi. S of Caborca; [30.5687 ° N, 112.1619 ° W]; 11 Jun. 1962; D. H. Janzen leg.; EMEC • 2 ♀♀; 30 km E of Agua Prieta; 31.3258 ° N, 109.2881 ° W; 6 Apr. 2006; L. Julier leg.; ex Physaria gordonii (verbatim: Lesquerella gordonii); PCYU • 1 ♀; 30 km E of Agua Prieta; 31.333333 ° N, 109.259722 ° W; 20 Aug. 2003; R. L. Minckley leg.; ex Baccharis; ASUHIC • 1 ♀; Nogales; [31.3402 ° N, 110.9361 ° W]; 4 Sep. 1965; A. E. Michelbacher leg.; EMEC • 2 ♀♀; Valley of Mexico; [19.67 ° N, 98.87 ° W]; 1 – 31 Aug. 1929; Y. Mexia leg.; CAS. UNITED STATES – Arizona • 2 ♀♀; Cochise Co., 1 mi. NNW of Jct. State Line Rd and Hwy 80, along irrigation ditch; [31.44 ° N, 109.83 ° W]; 9 Aug. 1973; S. I. Frommer and S. L. Frommer leg.; UCRC • 1 ♀; Cochise Co., 1.7 mi. NW of Portal, San Simon Road; [31.9307 ° N, 109.1619 ° W]; 26 Jul. 1973; S. I. Frommer and S. L. Frommer leg.; UCRC • 1 ♀; Cochise Co., 17 mi. E of Douglas; [31.3418 ° N, 109.2575 ° W]; 8 Aug. 1958; P. D. Hurd leg.; ex Lepidium thurberi; EMEC • 1 ♀; Cochise Co., 2 mi. NE of Portal; [31.9308 ° N, 109.1163 ° W]; 14 Jul. 1964; J. H. Puckle, M. A. Mortenson and M. A. Cazier leg.; ex Baccharis salicifolia (verbatim: Baccharis glutinosa); ASUHIC • 1 ♀; Cochise Co., 27 mi. SE of Willcox; [31.9598 ° N, 109.5075 ° W]; 31 Aug. 1980; McGinley and Woodley leg.; ex Gaillardia; MCZ • 3 ♀♀; Cochise Co., 4 mi. N of Elfrida; [31.743 ° N, 109.688 ° W]; 20 May 1961; R. H. Painter and E. M. Painter leg.; ASUHIC • 1 ♀; Cochise Co., 5 mi. SE of Apache; [31.638 ° N, 109.072 ° W]; 11 Aug. 1958; P. D. Hurd leg.; ex Eriogonum; EMEC • 1 ♀; Cochise Co., 5 mi. SE of San Simon; [32.215 ° N, 109.167 ° W]; 22 Aug. 1967; E. Schlinger leg.; UCRC • 1 ♀; Cochise Co., Benson; [31.956 ° N, 110.307 ° W]; 14 – 15 Sep. 1974; E. G. Linsley and J. M. Linsley leg.; ex Helianthus annuus; EMEC • 1 ♀; same data as for preceding; 7 Aug. 1961; P. H. Johnson leg.; ASUHIC • 1 ♀; Cochise Co., Cochise; [32.114 ° N, 109.922 ° W]; elev. 1280.2 m; 26 May 1961; H. A. Scullen leg.; EMEC • 1 ♀; Cochise Co., Douglas; [31.34 ° N, 109.55 ° W]; 9 Aug. 1944; W. W. Jones leg.; EMEC • 7 ♀♀; same data as for preceding; 16 Jun. 1942; H. A. Scullen leg.; EMEC • 1 ♀; Cochise Co., Douglas, Huachucha Mountains; [31.4 ° N, 110.3 ° W]; 8 Jun. 1933; W. W. Jones leg.; EMEC • 1 ♀; Cochise Co., Portal; [31.914 ° N, 109.141 ° W]; 18 Jul. 1963; A. Raske leg.; ex Datura metaloides; EMEC • 1 ♀; Cochise Co., Portal, Chiricahua Mountains; [31.929 ° N, 109.378 ° W]; elev. 1432.6 m; 14 Aug. 1965; G. R. Ballmer leg.; UCRC • 1 ♀; Cochise Co., Rucker Canyon, Chiricahua Mountains; [31.753 ° N, 109.43 ° W]; 25 Aug. 1970; EMEC • 1 ♀; Cochise Co.; [31.88 ° N, 109.75 ° W]; 23 Jun. 1916; V. W. Owen leg.; CAS • 3 ♀♀; Cochise Co., St David; [31.8936 ° N, 110.2284 ° W]; 27 Jul. 1955; G. Butler and F. Werner leg.; ex Lepidium virginicum subsp. menziesii (verbatim: Lepidium medium); ASUHIC • 1 ♀; same data as for preceding; 27 Jul. 1955; G. Butler and F. Werner leg.; ASUHIC • 1 ♀; same data as for preceding; 27 Jul. 1955; G. Buttler, F. Werner leg.; ex Lepidium; ASUHIC • 1 ♀; same data as for preceding; 2 May 1956; G. Butler and F. Werner leg.; ASUHIC • 1 ♀; same data as for preceding; 2 May 1956; F. Werner and G. Buttler leg.; ASUHIC • 3 ♀♀; same data as for preceding; 2 May 1956; F. Werner and G. Buttler leg.; ex alfalfa; ASUHIC • 1 ♀; Coconino Co., 10 mi. N of Sedona; [35.0115 ° N, 111.7598 ° W]; 27 Jul. 1956; G. D. Butler and Gerhardt leg.; ex Melilotus alba; ASUHIC • 1 ♀; same data as for preceding; 27 Jul. 1956; G. D. Butler and Gerhardt leg.; ASUHIC • 2 ♀♀; Coconino Co., 24 mi. N of Cameron; [36.22 ° N, 111.411 ° W]; 28 May 1954; F. Werner leg.; ASUHIC • 1 ♀; Coconino Co., 26 mi. N of Cameron; [36.246 ° N, 111.412 ° W]; 28 May 1954; F. Werner leg.; ASUHIC • 3 ♀♀; Coconino Co., Oak Creek Canon; [34.91 ° N, 111.73 ° W]; elev. 1828.8 m; F. H. Snow leg.; SEMC • 1 ♂; same data as for preceding; Aug; F. H. Snow leg.; USNM • 2 ♀♀; same data as for preceding; Jul.; F. H. Snow leg.; USNM • 1 ♀; Doña Ana Co., Las Cruces; [32.3199 ° N, 106.7637 ° W]; 10 May; Townsend leg.; ex Dimorphocarpa wislizeni; CMNH • 7 ♀♀; Gila Co., Gila River, 2 mi. SW of Christmas; [33.027 ° N, 110.738 ° W]; 5 Jun. 1962; J. C. Bequaert leg.; ASUHIC (Fig. 74 A – B) • 1 ♂; same data as for preceding; 5 Jun. 1962; Bequaert and Werner leg.; ASUHIC • 1 ♀; Gila Co., San Carlos; [33.351 ° N, 110.452 ° W]; 15 Jul. 1955; N. J. Nerney leg.; ex alfalfa; ASUHIC • 2 ♀♀; Graham Co., Central; [32.868 ° N, 109.793 ° W]; 24 Jun. 1954; G. D. Butler leg.; ASUHIC • 2 ♀♀; Graham Co., Geronimo; [33.077 ° N, 110.035 ° W]; 15 Sep. 1957; P. H. Timberlake leg.; ex Conyza canadensis var. canadensis (verbatim: Erigeron canadensis); UCRC • 3 ♀♀; same data as for preceding; 15 Sep. 1957; P. H. Timberlake leg.; UCRC • 1 ♀; Graham Co., Gila Valley; [33.183 ° N, 110.217 ° W]; 29 Jul. 1933; CAS • 1 ♀; Graham Co., Safford; [32.833 ° N, 109.715 ° W]; 19 Jul. 1956; C. Williams leg.; ex cotton (Gossypium); ASUHIC • 1 ♀; same data as for preceding; 3 May 1956; F. Werner, G. Buttler leg.; ex alfalfa; ASUHIC • 1 ♀; Maricopa Co., 3 mi. W of Palo Verde; [33.348 ° N, 112.73 ° W]; 21 Feb. 1961; J. C. Bequaert leg.; ex Dimorphocarpa wislizeni (verbatim: Dithyrea wislizeni); ASUHIC • 2 ♀♀; Maricopa Co., Tempe; [33.4128 ° N, 111.9431 ° W]; 31 Jul. 2017; Wheeler leg.; MCZ • 1 ♀; same data as for preceding; 1 Jul.; J. Bequaert leg.; MCZ • 1 ♀; Maricopa Co., Buckeye; [33.37 ° N, 112.59 ° W]; 28 May 1933; P. H. Timberlake leg.; ex Salix nigra; UCRC • 1 ♀; Maricopa Co., Granite Reef Dam; [33.516 ° N, 111.691 ° W]; 6 Apr. 1961; C. R. Brehmer leg.; ASUHIC • 1 ♀; Maricopa Co., Laveen; [33.378 ° N, 112.168 ° W]; 31 May 1955; O. L. Barnes leg.; ex alfalfa; v • 1 ♀; Maricopa Co., Mesa; [33.422 ° N, 111.823 ° W]; elev. 365.8 m; 21 Mar. 1926; A. A. Nichol leg.; ASUHIC • 1 ♀; Maricopa Co., Palo Verde; [33.348 ° N, 112.678 ° W]; 11 Jun. 1919; A. Wetmore leg.; USNM • 1 ♀; Maricopa Co., St Johns; [34.502 ° N, 109.372 ° W]; 26 Jul. 1956; G. D. Butler and Gerhardt leg.; ASUHIC • 1 ♀; Maricopa Co., Tempe; [33.4128 ° N, 111.9431 ° W]; 1 – 31 Jul. 1931; J. Bequaert leg.; MCZ • 1 ♀; same data as for preceding; 20 Nov. 1958; J. McBride leg.; ASUHIC • 5 ♀♀; Maricopa Co., Tolleson; [33.45 ° N, 112.259 ° W]; 29 May 1933; P. H. Timberlake leg.; ex Sphaeralcea; UCRC • 2 ♀♀; same data as for preceding; 28 May 1933; P. H. Timberlake leg.; ex Sphaeralcea; UCRC • 10 ♀♀; Navajo Co., Snowflake; [34.51 ° N, 110.08 ° W]; 25 Jul. 1956; G. D. Butler and Gerhardt leg.; ASUHIC • 3 ♀♀; same data as for preceding; 25 Jul. 1956; Gerhardt and Butler leg.; ASUHIC • 1 ♀; Navajo Co., Taylor; [34.47 ° N, 110.1 ° W]; 25 Jul. 1956; G. D. Butler and Gerhardt leg.; ex alfalfa; ASUHIC • 1 ♂; same data as for preceding; 25 Jun. 1956; G. D. Butler and Gerhardt leg.; ASUHIC (Fig. 75) • 1 ♀; Navajo Co., Laguna Creek Kayenta; [36.86 ° N, 109.82 ° W]; 13 Jul. 1967; J. H. Davidson, J. M. Davidson and M. A. Cazier leg.; ASUHIC • 2 ♀♀; Pima Co., 10 mi. E of Tucson; [32.2498 ° N, 110.8037 ° W]; 30 Jul. 1955; G. D. Butler leg.; ex Lepidium; ASUHIC • 1 ♂; Pima Co., Continental; [31.85 ° N, 110.97 ° W]; 14 Jun. 1955; G. D. Butler leg.; ASUHIC • 1 ♀; Pima Co., Continental; [31.851 ° N, 110.975 ° W]; 14 Jun. 1955; G. D. Butler leg.; ex Verbesina encelioides; ASUHIC • 1 ♀; Pima Co., 7.5 mi. W of Arivaca; [31.576 ° N, 111.459 ° W]; 31 Jul. 1952; H. B. Leech and J. W. Green leg.; ex Verbesina encelioides; CAS • 2 ♀♀; Pima Co., Aguirre Lake; [31.589 ° N, 111.508 ° W]; 5 May 1963; M. L. Noller leg.; ex Neltuma juliflora (verbatim: Prosopis juliflora); ASUHIC • 1 ♀; Pima Co., Sahuarita; [31.9291 ° N, 110.9822 ° W]; 17 Apr. 1956; G. D. Butler leg.; ex alfalfa; ASUHIC (Fig. 74 D) • 1 ♀; Pima Co., Tucson; [32.222 ° N, 110.926 ° W]; 2 – 7 Jun. 1962; G. D. Butler leg.; ASUHIC • 1 ♀; same data as for preceding; 2 Jun. 1962; G. D. Butler leg.; ex Calendula; ASUHIC • 2 ♀♀; same data as for preceding; 30 May 1953; G. D. Butler leg.; ex crownbeard (Verbesina); ASUHIC • 1 ♂; same data as for preceding; 6 Jul. 1962; G. D. Butler leg.; ASUHIC • 2 ♀♀; same data as for preceding; 11 Jun. 1964; G. D. Butler leg.; ASUHIC • 1 ♂; same data as for preceding; 6 Jul. 1962; G. D. Butler leg.; ex suck / cotton; ASUHIC • 1 ♀; same data as for preceding; 11 May 1953; G. D. Butler leg.; ex desert marigold; ASUHIC • 1 ♀; same data as for preceding; 30 May 1953; G. D. Butler leg.; ex bachelor’s button; ASUHIC • 4 ♀♀; same data as for preceding; 30 May 1953; G. D. Butler leg.; ex sun flower; ASUHIC • 1 ♀; same data as for preceding; 21 May 1962; G. D. Butler leg.; ex safflower; ASUHIC • 1 ♀; same data as for preceding; 24 May 1953; G. D. Butler leg.; ex Calendula; ASUHIC • 1 ♀; same data as for preceding; 30 May 1954; G. D. Butler leg.; ex Haplopappus; ASUHIC • 1 ♀; same data as for preceding; 29 May 1953; G. D. Butler leg.; ex Calendula; ASUHIC • 1 ♀; same data as for preceding; 30 May 1964; G. D. Butler leg.; ex safflower; ASUHIC • 1 ♀; same data as for preceding; 31 May 1962; G. D. Butler leg.; ex safflower; ASUHIC • 1 ♀; same data as for preceding; 20 Jun. 1964; G. D. Butler leg.; ex safflower; ASUHIC • 3 ♀♀; same data as for preceding; 1 – 30 Jun. 1965; G. D. Butler leg.; ex safflower; ASUHIC • 1 ♀; same data as for preceding; 23 May 1962; G. D. Butler leg.; ex safflower; ASUHIC • 1 ♀; same data as for preceding; 8 Apr. 1955; G. D. Butler leg.; ex Baileya multiradiata; ASUHIC • 1 ♀; same data as for preceding; 23 Aug. 1955; G. D. Butler leg.; ASUHIC • 1 ♀; same data as for preceding; 8 Jun. 1962; G. D. Butler leg.; ex safflower; ASUHIC • 1 ♀; same data as for preceding; 17 Jul. 1955; G. D. Butler leg.; ex Wislizenia; ASUHIC • 2 ♀♀; same data as for preceding; 1 Aug. 1975; E. G. Linsley and J. M. Linsley leg.; ex Tamarix chinensis; EMEC • 1 ♀; same data as for preceding; 10 Apr. 1955; T. Adams leg.; ASUHIC • 8 ♀♀; same data as for preceding; 12 Oct. 1963; ASUHIC • 1 ♀; Pima Co., San Xavier Mission; [32.107 ° N, 111.008 ° W]; 15 Oct. 1936; E. P. Van Duzee leg.; EMEC • 3 ♀♀; Santa Cruz Co., 19 mi. NW of Nogales; [31.5334 ° N, 111.1622 ° W]; 27 May 1961; R. H. Painter and E. M. Painter leg.; ASUHIC • 3 ♀♀; Santa Cruz Co., 3 mi. S of Patagonia; [31.4977 ° N, 110.7552 ° W]; 29 Jul. 1962; M. Noller leg.; ex Baccharis salicifolia (verbatim: Baccharis glutinosa); ASUHIC • 1 ♀; Santa Cruz Co., 5 mi. N of Patagonia; [31.6126 ° N, 110.76 ° W]; 5 May 1961; R. H. Painter and E. M. Painter leg.; ASUHIC • 5 ♀♀; Santa Cruz Co., Amado; [31.705 ° N, 111.066 ° W]; elev. 914.4 m; 14 Jun. 1963; Scullen and Bolinger leg.; EMEC • 1 ♀; Santa Cruz Co., Patagonia; [31.54 ° N, 110.76 ° W]; 28 May 1995; B. Brown and E. Wilk leg.; PMAE • 5 ♀♀; same data as for preceding; 26 Jul. 1962; Werner, Bequaert and Noller leg.; ASUHIC • 8 ♀♀; same data as for preceding; 29 Jul. 1962; Noller, Bequaert, Johnson leg.; ex Lepidium thurberi; ASUHIC • 1 ♀; same data as for preceding; 26 Mar. 1955; G. Butler and F. Werner leg.; ex Barkleyanthus salicifolius (verbatim: Senecio salignus); ASUHIC • 1 ♂; same data as for preceding; 26 Jul. 1962; Werner, Bequaert and Noller leg.; ex suck / cotton; ASUHIC • 1 ♂; same data as for preceding; 3 Aug. 1958; M. Adachi leg.; ASUHIC • 3 ♀♀; same data as for preceding; 25 Aug. 1955; F. G. Werner and G. D. Butler leg.; ex Cleomella jonesii (verbatim: Cleome jonesii); ASUHIC • 1 ♀; same data as for preceding; 10 Aug. 1958; F. G. Werner and G. D. Butler leg.; ASUHIC • 1 ♀; same data as for preceding; 3 Aug. 1958; M. Adachi leg.; ASUHIC • 1 ♀; same data as for preceding; 21 May 1955; G. D. Butler leg.; ex alfalfa; ASUHIC • 1 ♀; same data as for preceding; 21 May 1955; G. D. Butler leg.; ex mustard; ASUHIC • 1 ♀; same data as for preceding; 21 May 1955; G. D. Butler leg.; ex Sphaeralcea; ASUHIC • 1 ♀; same data as for preceding; 8 Aug. 1940; P. H. Timberlake leg.; ex Cleome lutea; UCRC • 1 ♀; same data as for preceding; 9 Aug. 1940; P. H. Timberlake leg.; UCRC • 1 ♀; same data as for preceding; 15 Aug. 1953; P. H. Timberlake leg.; UCRC • 2 ♂♂; Santa Cruz Co., Patagonia Sta; [31.49 ° N, 110.85 ° W]; 25 Jul. 1975; E. G. Linsley and J. M. Linsley leg.; ex Sapindus saponaria; EMEC (Fig. 76) • 1 ♀; Santa Cruz Co., Patagonia, Sonoita Creek Reserve; [31.52 ° N, 110.77 ° W]; 15 Jul. 1995; B. Brown and E. Wilk leg.; PMAE • 1 ♂; Santa Cruz Co., Tubac; [31.626 ° N, 111.052 ° W]; 21 Aug. 1935; R. H. Beamer leg.; SEMC • 1 ♀; Yuma Co., Roll; [32.75 ° N, 113.99 ° W]; 2 Jun. 1955; G. D. Butler leg.; ASUHIC • 1 ♀; same data as for preceding; 14 Apr. 1955; G. D. Butler and Gerhardt leg.; ex Heliotropium; ASUHIC • 1 ♀; Bill Williams Fork; [34.272 ° N, 114.046 ° W]; F. H. Snow leg.; SEMC • 2 ♀♀; S. Arizona; [33.4 ° N, 112.51 ° W]; 1 – 31 Aug. 1902; Carl F. Baker leg.; ex Sisymbrium; USNM. – California • 1 ♀; San Bernardino Co., Needles; [34.85 ° N, 114.61 ° W]; 3 Apr. 1951; EMEC. – Colorado • 2 ♀♀; Bent Co., John Martin Reservoir; [38.073 ° N, 103.028 ° W]; 6 Jul. 1978; G. E. Bohart leg.; ex Melilotus; UCMC • 1 ♀; Boulder Co., Boulder; [40.015 ° N, 105.271 ° W]; 1 – 30 Jun. 1920; H. Andrews leg.; UCMC • 2 ♀♀; Boulder Co.; [40.14 ° N, 105.47 ° W]; 18 Jun. 1925; C. H. Hicks leg.; UCMC • 1 ♀; Clear Creek Co., Bear Creek; [33.4 ° N, 112.51 ° W]; 20 Jul.; E. J. Oslar leg.; USNM • 2 ♀♀; same data as for preceding; 15 Jul.; E. J. Oslar leg.; USNM • 1 ♀; same data as for preceding; 13 May; E. J. Oslar leg.; USNM • 11 ♀♀; same data as for preceding; USNM • 1 ♀; Clear Creek Co., Clear Creek; [39.83 ° N, 104.95 ° W]; Oslar leg.; MCZ • 2 ♀♀; same data as for preceding; N. Banks leg.; MCZ • 23 ♀♀; Clear Creek Co., Clear Creek; [39.828 ° N, 104.949 ° W]; USNM • 1 ♀; Conejos Co., Los Pinos; [37.09 ° N, 106.49 ° W]; 31 May 1999; Carl F. Baker leg.; ex Prunus; USNM • 1 ♀; same data as for preceding; 31 May 1999; Carl F. Baker leg.; USNM • 3 ♀♀; Denver Co., Denver; [39.739 ° N, 104.985 ° W]; 28 May 1998; E. J. Oslar leg.; USNM • 4 ♀♀; same data as for preceding; USNM • 1 ♀; same data as for preceding; 3 Aug; E. J. Oslar leg.; USNM • 47 ♀♀; Douglas Co., Platte Canyon; [39.485 ° N, 105.106 ° W]; USNM • 1 ♂; Elbert Co., Box Elder Creek E of Aurora; [40.332 ° N, 104.511 ° W]; 14 Aug. 1947; H. G. Rodeck leg.; UCMC • 5 ♀♀; Logan Co., Sterling; [40.626 ° N, 103.211 ° W]; 21 Jul. 1923; G. Sandhouse leg.; USNM • 2 ♀♀; same data as for preceding; 3 Jul. 1923; G. Sandhouse leg.; ex marigold; USNM • 6 ♀♀; same data as for preceding; 28 Jun. 1923; G. Sandhouse leg.; USNM • 1 ♀; same data as for preceding; 20 Jul. 1923; G. Sandhouse leg.; USNM • 1 ♂; same data as for preceding; 20 Jul. 1923; G. Sandhouse leg.; USNM • 1 ♀; same data as for preceding; 17 Jun. 1923; G. Sandhouse leg.; USNM • 3 ♀♀; same data as for preceding; 13 Jun. 1923; G. Sandhouse leg.; USNM • 1 ♀; same data as for preceding; 6 Jul. 1923; G. Sandhouse leg.; USNM • 1 ♀; same data as for preceding; 18 Jun. 1923; G. Sandhouse leg.; USNM • 1 ♀; same data as for preceding; 23 Jun. 1923; G. Sandhouse leg.; USNM • 1 ♀; same data as for preceding; 11 Jul. 1921; G. Sandhouse leg.; USNM • 2 ♀♀; Mesa Co., Colorado National Monument; [39.0425 ° N, 108.6861 ° W]; elev. 1371.6 m; 16 May 1963; C. J. McCoy leg.; ex Sphaeralcea parvifolia; UCMC • 1 ♀; same data as for preceding; 24 May 1963; C. J. McCoy leg.; ex Sphaeralcea; UCMC (Fig. 74 C) • 1 ♀; same data as for preceding; 28 May 1963; C. J. McCoy leg.; UCMC • 1 ♀; Moffat Co., Castle Park, Dinosaur National Monument; [40.533 ° N, 108.983 ° W]; elev. 1569.7 m; 8 Jul. 1949; H. G. Rodeck leg.; ex Solidago; UCMC • 2 ♀♀; Morgan Co., 2 mi. E of Roggen; [40.1685 ° N, 104.3339 ° W]; 26 May 1963; U. N. Lanham et al. leg.; UCMC • 3 ♀♀; Morgan Co., 3 mi. N of Orchard; [40.3734 ° N, 104.118 ° W]; 10 May 1963; B. Vogel leg.; UCMC • 1 ♀; Pueblo Co., Pueblo; [38.27 ° N, 104.62 ° W]; 21 May 1918; A. Wetmore leg.; USNM • 1 ♀; Weld Co., 10 mi. NE of Roggen; [40.2708 ° N, 104.2377 ° W]; 25 May 1963; U. N. Lanham leg.; ex Salix; UCMC • 3 ♂♂; Weld Co., Owl Creek, 12 mi. NE of Nunn; [40.576 ° N, 104.636 ° W]; 4 Aug. 1983; H. E. Evans leg.; UCMC • 1 ♀; Weld Co., Roggen; [40.1683 ° N, 104.3717 ° W]; 19 May 1930; W. P. Cockerell leg.; UCRC • 1 ♂; same data as for preceding; 8 Jul. 1984; H. E. Evans leg.; UCMC • 1 ♀; Weld Co., Pawnee National Grassland, Cow Valley Rec. Area; 40.645 ° N, 104.343 ° W; elev. 1480 m; 39977; K. Perez, S. McCubbin, J. Cossey and A. Borisenko leg.; CBG • 1 ♀; Yamu Co., Beecher Island; [39.87 ° N, 102.187 ° W]; 21 Jun. 1979; U. N. Lanham leg.; ex Melilotus; UCMC • 4 ♀♀; White Rocks (?); 29 Jun. 1939; P. H. Timberlake leg.; ex Monarda pectinata; UCRC • 1 ♀; same data as for preceding; 28 Jun. 1939; P. H. Timberlake leg.; UCRC • 1 ♂; Carl F. Baker leg.; USNM • 94 ♀♀; same data as for preceding; Carl F. Baker leg.; USNM. – Nevada • 2 ♀♀; Clark Co., Las Vegas; [36.175 ° N, 115.136 ° W]; 20 Jun. 1939; P. H. Timberlake leg.; ex Cirsium; UCRC • 6 ♀♀; Lincoln Co., 3 mi. NW of Caliente; [37.6448 ° N, 114.5504 ° W]; 22 Jun. 1982; W. J. Pulawski leg.; CAS. – New Mexico • 1 ♀; Bernalillo Co., Oslar, Albuquerque; [35.111 ° N, 106.593 ° W]; 17 Jul. 1902; USNM • 3 ♀♀; Chaves Co., 5 mi. E of Roswell; [33.3917 ° N, 104.4367 ° W]; 11 Sep. 1961; P. D. Hurd leg.; ex Lepidium integrifolium (verbatim: Lepidium montanum var. integrifolium); EMEC • 3 ♀♀; same data as for preceding; 11 Sep. 1961; P. D. Hurd leg.; ex Isocoma pluriflora (verbatim: Haplopappus pluriflorus); EMEC • 2 ♂♂; same data as for preceding; 11 Sep. 1961; P. D. Hurd leg.; ex Lepidium integrifolium (verbatim: Lepidium montanum var. integrifolium); EMEC • 2 ♀♀; Chaves Co., 5 mi. W of Roswell; [33.3965 ° N, 104.608 ° W]; 11 Sep. 1961; P. H. Timberlake leg.; UCRC • 1 ♀; same data as for preceding; 11 Sep. 1961; P. H. Timberlake leg.; ex Lepidium montanum; UCRC • 1 ♀; Cochise Co., Southwest Research Station; 31.8840 ° N, 109.2070 ° W]; 19 Aug. 2024; J. Gibbs leg.; WRME • 1 ♀; same data as for preceding; 15 Aug. 2024; J. Gibbs leg.; WRME • 1 ♀; Doña Ana Co., Las Cruces; [32.3199 ° N, 106.7637 ° W]; Townsend leg.; MCZ • 1 ♀; same data as for preceding; 4872; Cockerell leg.; ex Salix; MCZ • 1 ♀; same data as for preceding; Cockerell leg.; ex Melilotus; MCZ • 1 ♀; same data as for preceding; Cockerell leg.; ex Salix; MCZ • 1 ♀; same data as for preceding; Townsend leg.; EMEC • 1 ♀; same data as for preceding; 3 May; T. D. A. Cockerell leg.; ex Salix; EMEC • 2 ♀♀; same data as for preceding; 18 Jun. 1942; E. C. Van Dyke leg.; CAS • 1 ♀; same data as for preceding; 5 Oct. 1995; Cockerell leg.; USNM • 1 ♀; same data as for preceding; 29 Sep. 1995; Cockerell leg.; USNM • 1 ♀; same data as for preceding; May 11; Townsend leg.; USNM • 1 ♀; same data as for preceding; USNM • 1 ♀; same data as for preceding; 10 May; Townsend leg.; USNM • 3 ♀♀; Doña Ana Co., Las Cruces; [32.3199 ° N, 106.7637 ° W]; elev. 1182.6 m; 18 Jun. 1942; H. A. Scullen leg.; EMEC • 1 ♀; Doña Ana Co., Mesilla; [32.27 ° N, 106.80 ° W]; 1 – 30 Apr. 1917; ex plum; USNM • 27 ♀♀; same data as for preceding; 1 – 30 Apr. 1909; C. N. Ainslie leg.; USNM • 10 ♀♀; same data as for preceding; 4 May 1909; C. N. Ainslie leg.; USNM • 6 ♀♀; same data as for preceding; C. N. Ainslie leg.; USNM • 1 ♂; Doña Ana Co., Mesilla Park; [32.252 ° N, 106.748 ° W]; 8 Jul. 1898; W. P. Cockerell leg.; UCRC • 1 ♀; same data as for preceding; 18 May 1900; Cockerell leg.; USNM • 1 ♀; Doña Ana Co., Mesilla; [32.27 ° N, 106.80 ° W]; 1 – 31 May 1910; ex Hymenoxys; UCRC • 1 ♀; Eddy Co., 5 mi. N of Carlsbad; [32.49 ° N, 104.229 ° W]; 21 Sep. 1956; J. W. MacSwain leg.; ex Chrysothamnus; EMEC • 1 ♂; Grant Co., 16 mi. NW of Silver City; [32.9324 ° N, 108.4755 ° W]; 13 Jun. 1961; H. A. Scullen leg.; ex Asclepias; OSAC • 1 ♀; Guadalupe Co., 2 mi. W of Santa Ross; [34.9368 ° N, 104.7172 ° W]; 10 Jul. 1929; V. E. Romney leg.; ex Croton texensis; USNM • 1 ♀; Hidalgo Co., 15 mi. S of Animas; [31.729 ° N, 108.8066 ° W]; 18 Aug. 1980; McGinley and Woodley leg.; ex Cleome; MCZ • 2 ♀♀; Hidalgo Co., 16 mi. S of Animas; 31.7200 ° N, 108.822 ° W; 16 Aug. 2024; J. Gibbs leg.; WRME • 1 ♂; same data as for preceding; 16 Aug. 2024; J. Gibbs leg.; WRME • 1 ♀; Hidalgo Co., 29 mi. S of Animas; [31.5275 ° N, 108.8114 ° W]; 27 Aug. 1980; McGinley and Woodley leg.; ex Sphaeralcea; MCZ • 1 ♀; same data as for preceding; 27 Aug. 1980; McGinley and Woodley leg.; ex Sphaeralcea; MCZ • 28 ♀♀; Hidalgo Co., 15 mi. S of Animas; [31.729 ° N, 108.8066 ° W]; 15 Aug. 1978; N. S. Wasbauer leg.; ex Baccharis salicifolia (verbatim: Baccharis glutinosa); CSCA • 1 ♀; Los Alamos Co., Frijoles Canyon; Bandelier National Monument; [36.44 ° N, 105.503 ° W]; 14 Jun. 1960; Burks, Kinzer leg.; USNM • 1 ♀; Maricopa Co., 5 mi. SW of Avondale; [33.4303 ° N, 112.4365 ° W]; 1 Apr. 1967; G. I. Stage leg.; USNM • 1 ♀; San Miguel Co., Apache Springs; [35.372 ° N, 105.193 ° W]; 13 Jul. 1959; E. G. Linsley leg.; EMEC • 1 ♀; Sandoval Co., Peña Blanca; [35.573 ° N, 106.337 ° W]; 7 Jun. 1965; U. N. Lanham leg.; UCMC • 2 ♀♀; Santa Fe Co., 10 km S of La Cienega, Hyw 85; 35.5122 ° N, 106.22415 ° W; 17 Aug. 2007; C. Sheffield and J. Gibbs leg.; PMAE • 6 ♀♀; Socorro Co., 38 mi. W of Magdalena; [34.0923 ° N, 107.904 ° W]; elev. 2164.1 m; 5 Aug. 1946; H. A. Scullen leg.; EMEC • 2 ♀♀; Valencia Co., Los Lunas, 20 mi. W of Carrizo Arroyo; [34.809 ° N, 106.735 ° W]; 1 – 23 Aug. 1977; S. and J. Peck leg.; PMAE • 2 ♀♀; 2.2 mi. E of (?); 16 Sep. 1965; P. H. Timberlake leg.; UCRC. – Oklahoma • 1 ♀; Cimarron Co., Lake Carl Etling, Black Mesa State Park, 25 mi. by road NW. of Boise City; [36.84 ° N, 102.882 ° W]; 14 Aug. 1967; H. B. Leech leg.; CAS. – Texas • 1 ♀; Brewster Co., 1 mi. W of Lajitas; [29.2613 ° N, 103.7931 ° W]; 23 – 30 Apr. 1963; H. E. Evans leg.; MCZ • 1 ♂; Brewster Co., Rio Grande; [26.381 ° N, 98.818 ° W]; 13 – 17 Jun. 1908; Mitchell and Cushman leg.; ex Neltuma glandulos a (verbatim: Prosopis glandulosa); USNM • 2 ♀♀; Brewster Co., Santa Elena Canyon, Big Bend National Park; [29.165 ° N, 103.612 ° W]; 24 Apr. 1953; B. J. Adelson leg.; EMEC • 1 ♀; same data as for preceding; 21 Apr. 1953; M. Wasbauer leg.; EMEC • 1 ♀; Brewster Co., Santa Elena Canyon, Big Bend National Park; [29.165 ° N, 103.614 ° W]; 24 Apr. 1953; M. Wasbauer leg.; EMEC • 2 ♀♀; same data as for preceding; 22 Apr. 1953; B. J. Adelson leg.; EMEC • 2 ♀♀; El Paso Co., El Paso; [31.759 ° N, 106.489 ° W]; 26 Apr. 1927; J. O. Martin leg.; CAS • 1 ♀; same data as for preceding; 25 Apr. 1927; J. O. Martin leg.; CAS • 1 ♂; Starr Co., Santa Elena; [26.7601 ° N, 98.4864 ° W]; 22 Apr. 1953; M. Wasbauer leg.; EMEC • 2 ♀♀; Val Verde Co., Langtry; [29.809 ° N, 101.559 ° W]; 14 Jun. 1980; J. L. Neff leg.; ASUHIC. – Utah • 1 ♀; Salt Lake Co., Salt Lake; [40.761 ° N, 111.891 ° W]; 21 Aug. 1913; P. H. Timberlake leg.; UCRC • 2 ♀♀; Salt Lake Co., Salt Lake City; [40.76 ° N, 111.89 ° W]; 9 Jun. 1926; E. C. VanDyke leg.; CAS • 1 ♀; San Juan Co., 4 mi. S of Blanding; [37.565 ° N, 109.475 ° W]; 24 Aug. 1967; J. C. Hall leg.; ex Grindelia; UCRC • 1 ♀; Sevier Co., Elsinore; [38.684 ° N, 112.148 ° W]; 18 Jun. 1933; J. T. Howell leg.; CAS • 1 ♀; Uintah Co., Watson; [39.8802 ° N, 109.1574 ° W]; elev. 1615.4 m; 1 – 31 Aug. 1910; F. M. Carpenter leg.; MCZ • 1 ♂; same data as for preceding; 1 – 31 Jul. 1922; F. M. Carpenter leg.; MCZ. – Wyoming • 2 ♀♀; Natrona Co., South Fork Powder River at Hwy 20, 1.8 mi. W of Town of Powder River; [43.0316 ° N, 107.0227 ° W]; 21 Aug. 1965; H. B. Leech leg.; CAS.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF5EFF6CFE7FFDC4FCD0FC07.taxon	description	Description GENITALIA (Figs 76, 120 N). Gonostylus elongate, approximately 2.5 times as long as broad, and bears short setae. Retrorse lobe broad, about 2 times as long as broad, covered in fine setae.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF5EFF6CFE7FFDC4FCD0FC07.taxon	distribution	Range Southwestern US and northern Mexico. Mexico (Chihuahua, Sonora), USA (Arizona, California, Colorado, Nevada, New Mexico, Oklahoma, Texas, Utah, Wyoming) (Fig. 77).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF5EFF6CFE7FFDC4FCD0FC07.taxon	biology_ecology	Floral hosts APOCYNACEAE: Asclepias sp. • ASTERACEAE: Aphanostephus ramosissimus DC., Aster sp., Baccharis salicifolia (Ruiz & Pav.) Pers., Baccharis sp., Baileya multiradiata Harv. & A. Gray, Barkleyanthus salicifolius (Kunth) H. Rob. & Brettell, Calendula sp., Carthamus tinctorius L., Centaurea cyanus L., Chloracantha spinosa (Benth.) G. L. Nesom, Chrysothamnus sp., Cirsium sp., Conyza canadensis var. canadensis Cronquist, Encelia sp., Flaveria sp., Grindelia decumbens Greene, Grindelia sp., Haplopappus sp., Helianthus annuus L., Helianthus sp., Hymenoxys richardsonii (Hook.) Cockerell, Hymenoxys sp., Isocoma pluriflora Greene, Solidago canadensis L., Solidago sp., Tagetes sp., Verbesina encelioides (Cav.) Benth. & Hook. f. ex A. Gray, Verbesina sp. • BORAGINACEAE: Heliotropium sp. • BRASSICACEAE: Descurainia pinnata (Walter) Britton, Descurainia sp., Dimorphocarpa wislizeni (Engelm.) Rollins, Lepidium integrifolium Nutt., Lepidium montanum Nutt., Lepidium thurberi Wooton, Lepidium sp., Lepidium virginicum subsp. menziesii (DC.) Thell., Physaria gordonii (A. Gray) O’Kane & Al-Shehbaz, Brassica nigra (L.) W. D. J. Koch, Sisymbrium sp. • CAPPARACEAE: Cleome lutea Hook., Cleome sp., Cleomella jonesii (J. F. Macbr.) J. C. Hall & Roalson, Wislizenia sp. • EUPHORBIACEAE: Croton texensis (Klotzsch) Müll. Arg. • FABACEAE: Medicago sativa L., Melilotus albus Medik., Melilotus indicus (L.) All., Melilotus sp., Neltuma glandulos a (Torr.) Britton & Rose, Neltuma juliflora (Sw.) Raf. • LAMIACEAE: Monarda pectinata Nutt. • MALVACEAE: Gossypium sp., Sphaeralcea sp., Sphaeralcea parvifolia A. Nelson • POLYGONACEAE: Eriogonum sp. • ROSACEAE: Prunus domestica L., Prunus sp. • SALICACEAE: Salix nigra Marshall, Salix sp. • SAPINDACEAE: Sapindus saponaria L. • SOLANACEAE: Datura inoxia Mill., Solanum elaeagnifolium Cav. • TAMARICACEAE: Tamarix chinensis Lour.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF5EFF6CFE7FFDC4FCD0FC07.taxon	discussion	DNA barcodes Two full DNA barcodes and one partial barcode were analyzed all from Sonora. They are indistinguishable from L. pectorale (Smith, 1853) (Gibbs et al. 2013). The two species can be unambiguously separated using morphology and have distinct distributions.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF51FF70FE6AFBE1FDB9FB53.taxon	description	Figs 78, 79, 104 B, 106 A	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF51FF70FE6AFBE1FDB9FB53.taxon	diagnosis	Diagnosis Female L. robertsoni can be recognised by the combination of head long, length / width ratio 1.07 (± 0.06 SD) (remarkably long in L. pathiranae sp. nov.); mesoscutum well imbricate and dull (shinier in all the other similar species), with contiguous (IS <1 PD) punctures (not contiguous in L. angustius); hypoepimeron rugulose (partly polished in L. angustius), punctures obscure (distinct in L. angustius and L. pathiranae); propodeal triangle with rugae reaching posterior margin (separating from species in which the rugae do not reach posterior margin), propodeal triangle posterior margin sharply edged with a distinct apical carina (this excludes all species with a smoothly rounded posterior margin); T 1 strongly coriarious on anterior face with a fan bearing a narrow dorsal opening (T 1 anterior face mostly polished with broad dorsal opening in the fan in L. angusticulum sp. nov.); and T 2 – T 3 with distinct basolateral tomentum (removes all species lacking basal patches or bands of tomentum on T 2 – T 3, such as L. nitidiusculum group). Males are unknown.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF51FF70FE6AFBE1FDB9FB53.taxon	etymology	Etymology The species was named for Charles Robertson (Carlinville, Illinois) for his “ excellent descriptions and notes on Illinois bees ”.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF51FF70FE6AFBE1FDB9FB53.taxon	materials_examined	Type material Syntypes of Halictus Robertsoni (“ type ” and paratype ” status not compliant with ICZN; examined) UNITED STATES – Texas • 7 ♀♀. United States, Texas, Victoria Co., Victoria; [28.80 ° N, 97.01 ° W]; 24 Feb. 1904; J. C. Crawford leg.; USNM Type No. 12039 (Fig. 78). Specimen labels “ Victoria / Tex // Feb. 24. 4 // J. C. Crawford collector // Type No. 12039 U. S. N. M. // Halictus Robertsoni Cwfd. ♀ Type // USNMENT 01735668 ” on one female; “ Victoria / Tex 4. 04 // J. C. Crawford collector // Type No. 12039 U. S. N. M. // Paratype No. 12039 U. S. N. M. // Halictus Robertsoni Cwfd. ♀ Paratype ” on remaining females. Type notes The original type series was a syntype series. Crawford wrote “ Type, Victoria, Texas, Febr. 24, 1904. Crawford collector ”. There is no reference to additional material; however, someone labeled additional specimens in the USNM, with the same collection data, as paratypes. It is not possible to be certain that it was Crawford who did so, and eve n if so, it cannot be determined whether this occurred before or after the species description was published, and regardless, there was no designation of a particular individual specimen in print at any point, as required by the Code. This does not comply with either Articles 74.5 or 74.6 of the Code, so even though one specimen is labeled as ‘ type’, it is not a validly-designated lectotype under the Code, and the entire set of 7 females is all still syntypes. Material examined UNITED STATES – Texas • 1 ♀; Bexar Co., San Antonio; [29.425 ° N, 98.495 ° W]; 11 Apr. 2006; F. C. Pratt leg.; ex Pyrrhopappus grandiflorus [verbatim: Sitilias grandiflora]; USNM • 1 ♀; same data as for preceding; 14 May 2006; USNM • 1 ♀; same data as for preceding; ex Sonchus; USNM • 1 ♀; Texas Hill Co., New Braunfels; [29.70 ° N, 98.13 ° W]; 7 Jun. 1942; E. S. Ross leg.; CAS.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF51FF70FE6AFBE1FDB9FB53.taxon	description	Redescription Female BODY MEASUREMENTS. Length 7.98 mm (± 0.30 SD). Head length 2.00 mm (± 0.04 SD). Face length 1.61 mm (± 0.06 SD). Head width 1.85 mm (± 0.09 SD). Wing length 5.23 mm (± 0.30 SD). n = 6. COLOURATION. Head and mesosoma dark brown. Clypeus apex dark brown. Labrum brown. Mandible dark brown basally, becoming light brown medially and reddish brown apically. Flagellum brown dorsally, light brown ventrally. Pronotal lobe brown. Tegula brown. Wing membrane faintly brownish, veins with subcosta brown, otherwise light brown. Legs brown. Metasoma dark brown with rims of terga, sterna and downcurved lateral areas of terga broadly light brown. PUBESCENCE. Body hair colour off-white. Tomentum dense on pronotal angle and lobe, space between pronotal lobe and tegula, pronotal collar, metanotum anteromedially, propodeum lateral surface anterodorsally, T 2 – T 3 basolaterally, and T 4 basally; moderately dense on supraclypeal area dorsolaterally, paraocular area, gena near eye margin, anterolateral angle and marginally of mesoscutum, and propodeum posterior surface dorsally; sparse on propodeum lateral (except anterodorsal area). Mesosoma covered by appressed, simple, very short hairs (<0.25 OD) mixed with sparse, erect, weakly plumose, long hairs (<1 OD) on mesoscutum and erect, plumose, long hairs (<2 OD) on other areas. Metasomal sterna with moderately sparse, erect, weakly plumose, long (<2.5 OD) hairs. Metafemoral scopa with dense plumose hairs. Wing hairs brown, very short and sparse. T 1 with a fan, narrow dorsal opening (0.5 OD), composed of overlapping sub-appressed simple hairs (<1.5 OD). SURFACE SCULPTURING. Clypeus imbricate to tessellate, becoming polished apical ⅓; punctures irregularly sparse (IS <3 PD) apically and medially, becoming smaller and denser (IS <1 PD) basolaterally. Supraclypeal area tessellate medially, imbricate posterolaterally; punctures fine, irregularly sparse (IS <2 PD) medially, becoming denser (IS <1 PD) posterolaterally. Frons reticulate-punctate punctures crowded (IS = 0 PD). Ocellocular area weakly rugulose; punctures dense (IS <1 PD). Gena polished, becoming lineolate posteriorly; punctures moderately sparse (IS = 1 – 2 PD) or obscure from some angles, becoming obscure posteriorly. Postgena lineolate. Tegula punctures sparse (IS = 1 – 3 PD) in the anterior half and basally, becoming impunctate posteriorly. Mesoscutum dull, well imbricate throughout, sometime becoming weakly so laterally; punctures dense (IS <1 PD). Mesoscutellum polished submedially, becoming weakly rugulose to imbricate marginally; punctures dense (IS <1 PD) marginally and on median line, sparse submedially (IS = 1 – 4 PD), diversopunctate. Metanotum rugulose, punctures obscure. Propodeal triangle with moderately strong rugae moderately strong, reaching posterior margin and join with the oblique carina, anastomosing, lateral rugae extending to dorsolateral slope of propodeum lateral face; interspace sculpture shiny to weakly imbricate. Preëpisternum rugose. Hypoepimeron rugulose; punctures obscure. Mesepisternum rugose dorsally, becoming rugulose ventrally. Propodeum lateral face strongly imbricate, becoming rugulose posteriorly; posterior face imbricate-rugulose. T 1 anterior face dull, strongly coriarious; dorsal surface weakly polished, becoming weakly coriarious on small subapicolateral boss and coriarious on rim, punctures sparse (IS = 1 – 3 PD), becoming more sparsely punctate (IS = 2 – 6 PD) in small subapicolateral boss and punctures fine, minute, and sparse (IS = 1 – 4 PD) on rim, becoming impunctate apical half and medially on rim. T 2 disc coriarious to polished, punctures moderately dense (IS = 1 – 2 PD) basally, becoming sparse (IS = 1 – 3 PD) apically; rim coriarious, punctures minute and sparse (IS = 3 – 6 PD) or nearly impunctate after premarginal line, becoming closer (IS = 1 – 3 PD) apically. STRUCTURE. Head long, length / width ratio 1.07 (± 0.06 SD). Face length / width ratio 0.87 (± 0.02 SD). Clypeus projecting ~ 85 % below suborbital tangent; clypeus length / width ratio 1.01 (± 0.05 SD). Mandible short, not extending much beyond opposing clypeal angle. Eyes convergent below, UOD / LOD ratio 1.15 (± 0.03 SD). Ocelli normal. Gena / eye width ratio 0.82 (± 0.04 SD). Pronotal angle obtuse. Intertegular span 1.40 mm (± 0.11 SD). Mesocutum length / width ratio 0.92 (± 0.05 SD); mesoscutum / scutellum length ratio 2.70 (± 0.25 SD); scutellum / metanotum length ratio 1.50 (± 0.08 SD); metanotum / propodeal triangle length ratio 0.96 (± 0.03 SD). Protibial spur with apical serrations shorter or subequal in length to width of malus; malus extending beyond the velum, nearly ½ the length of velum. Inner metatibial spur pectinate, teeth 3 – 5, basal teeth length mostly subequal or longer than width of rachis and sometime shorter than width of rachis. Propodeal triangle posterior margin carinate, apical carina distinct from background sculpturing. Propodeal lateral carina reaching dorsolateral slope. n = 6. Male Unknown	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF51FF70FE6AFBE1FDB9FB53.taxon	distribution	Range Rare. Texas, including Victoria County, Bexar County, and Texas Hill County, especially both coastal and central regions of Texas (Fig. 79).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF51FF70FE6AFBE1FDB9FB53.taxon	biology_ecology	Floral host ASTERACEAE: Pyrrhopappus grandiflorus (Nutt.) Nutt., Sonchus sp.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF51FF70FE6AFBE1FDB9FB53.taxon	discussion	Remarks Rare. Lasioglossum robertsoni may include cryptic species, but the limited number of specimens makes it difficult to conclusively separate them with confidence. These specimens are not included in the L. robertsoni observed material section but are listed separately below. A conspicuous characteristic distinguishing them from L. robertsoni is the presence of abruptly depressed rims on the terga, along with dense (IS <1 PD) larger punctures on the discs. When run through the key, these potential new species might also be identified as L. robertsoni, as they share the same additional characters outlined in the key.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF51FF70FE6AFBE1FDB9FB53.taxon	materials_examined	UNITED STATES — Texas • 1 ♀; Travis Co., Austin; [30.27 ° N, 97.74 ° W]; 8 Jun. 1979; J. L. Neff leg.; ex Cirsium texanum; SEMC • 2 ♀♀; Kerr Co.; Kerrville; [30.05 ° N, 99.14 ° W]; 13 Aug. 1963; N. L. H. Krauss leg.; USNM (Fig. 79)	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4DFF76FE13FB3CFEE0F940.taxon	description	Figs 80 – 83, 100 B, 106 F, 117 B, 120 O	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4DFF76FE13FB3CFEE0F940.taxon	diagnosis	Diagnosis Female L. sequoiae can be recognized by the combination of head round, length / width ratio 0.96 (± 0.02 SD); inner metatibial spur denticulate-serrate, teeth shorter than width of rachis; supraclypeal area with dense (IS <1 PD) punctures throughout; and propodeal triangle with rugae reaching posterior margin. Male L. sequoiae can be recognized by the combination of head long, length / width ratio 1.03 (± 0.04 SD); clypeus yellow in apical half; supraclypeal area with dense (IS <1 PD) punctures throughout; mesoscutum polished and with dense (IS <1 PD) punctures; and propodeal triangle with rugae reaching posterior margin. Both sexes of L. sequoiae cannot be easily mixed with other species in western North America.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4DFF76FE13FB3CFEE0F940.taxon	etymology	Etymology Michener (1936) did not state the derivation of the epithet. The name sequoiae is interpreted here as a possible reference to Sequoia (e. g., the coast redwood, Sequoia sempervirens Endl.), given that the type series originates from northern California localities within the broader redwood region.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4DFF76FE13FB3CFEE0F940.taxon	materials_examined	Type material Holotype of Halictus (Evylaeus) sequoiae (by original designation; examined by JG ca 2013 and re-examined from photographs) UNITED STATES – California • 1 ♀; Marin Co., Ross; [37.96 ° N, 122.56 ° W]; 28 Apr. 1918; E. P. Van Duzee leg.; CAS Type No. 4199. Specimen labels “ Ross Cal / Marin Co. / Apr. 28 ’ 18 // EPVan Duzee / Collector // Halictus / sequoiae / Mich. Type // California Academy / of Sciences / Type No. 4199 ”. Paratypes of Halictus (Evylaeus) sequoiae (by original designation; examined) UNITED STATES – California • 2 ♀♀; Sonoma Co., Mark West Springs; [37.339 ° N, 121.442 ° W]; 23 Apr. 1935; E. P. Van Duzee leg.; ex Nemophila; CAS • 1 ♀; Marin Co., Fairfax; [37.339 ° N, 121.442 ° W]; 12 Apr. 1925; C. L. Fox leg.; CAS • 1 ♀; Marin Co., Ross; [37.339 ° N, 121.442 ° W]; 18 Apr. 1928; E. P. Van Duzee leg.; CAS • 1 ♀; Mendocino Co., Yorkville; [37.339 ° N, 121.442 ° W]; 8 May 1935; E. P. Van Duzee leg.; ex Gilia; CAS. Specimen labels “ Mark West / Spgs Cal. / IV- 23 - 35 // EPVanDuzee / Collector // Nemophila tricolor // Halictus / sequoiae / Paratype Mich // Lasioglossum / GIBBS- 12853 and 12854 ”. “ Fairfax, / IV- 12 - 25 / MarinCo. Cal. // C. L. Fox // C. L. Fox Collection // Halictus / sequoiae / Paratype Mich // Lasioglossum / GIBBS- 12855 ”. “ Ross Cal / Marin Co. / Apr. 28 ’ 18 // EPVanDuzee / Collector // Halictus / sequoiae / Paratype Mich // Lasioglossum / GIBBS- 12932 ”. “ Yorkville Cal / Mndcno. Co. / V- 8 - 35 // EPVanDuzee / Collector // White / Gilia // Halictus / sequoiae / Paratype Mich // Lasioglossum / GIBBS- 12856 ”. Material examined UNITED STATES – California • 1 ♀; Alameda Co., Leona Heights; [37.789 ° N, 122.178 ° W]; 6 Apr. 1913; J. C. Bridwell leg.; EMEC • 1 ♀; Calaveras Co., Mokelumne Hill; [37.339 ° N, 121.442 ° W]; F. E. Blaisdell leg.; CAS • 1 ♀; same data as for preceding; Apr; F. E. Balaisdell leg.; CAS • 1 ♀; Clara Co., Santa Cruz Mountains; [37.117 ° N, 121.850 ° W]; 25 Apr. 1913; J. C. Bridwell leg.; EMEC • 1 ♀; Contra Costa Co., Russelman Park; [37.912 ° N, 121.906 ° W]; 22 Jun. 1954; P. D. Hurd leg.; EMEC • 1 ♀; El Dorado Co., 2 mi. SE of Latrobe; [38.538 ° N, 120.958 ° W]; 18 Apr. 1976; S. C. Kuba leg.; CSCA • 2 ♀♀; El Dorado Co., Blodgett Forest, 13 mi. E of Georgetown; [38.892 ° N, 120.663 ° W]; 10 Jul. 1967; W. J. Turner leg.; EMEC • 2 ♀♀; El Dorado Co., Cool; [38.887 ° N, 121.015 ° W]; 4 Apr. 1964; R. W. Thorp leg.; ex Lasthenia (verbatim: Baeria); EMEC • 4 ♀♀; same data as for preceding; 29 Mar. 1964; J. Powell leg.; EMEC • 1 ♀; Humboldt Co., 4 mi. N of Garberville; [40.158 ° N, 123.795 ° W]; 3 Jun. 1963; L. D. Thurman leg.; ex Oenothera; EMEC • 1 ♀; Lake Co., Elk Hill; [37.339 ° N, 121.442 ° W]; 1 – 31 May 1938; CAS • 1 ♀; Lake Co., Lakeport; [37.339 ° N, 121.442 ° W]; 1 – 30 Jun. 1917; J. Bentley leg.; EMEC • 1 ♀; Lake Co., Whispering Pines; [38.814 ° N, 122.713 ° W]; 14 Apr. 1964; C. N. Slobodchikoff leg.; EMEC • 9 ♀♀; Los Angeles Co., Claremont; [34.097 ° N, 117.719 ° W]; Baker leg.; EMEC • 2 ♀♀; Los Angeles Co., Mts near Claremont; [34.161 ° N, 117.716 ° W]; Baker leg.; EMEC • 1 ♀; Madera Co., Oakhurst; [37.328 ° N, 119.649 ° W]; 1 Jun. 1942; EMEC • 1 ♀; same data as for preceding; 19 May 1942; C. Kennett leg.; ex Ranunculus; EMEC • 1 ♀; Madera Co., San Joaquin Experimental Range; [37.083 ° N, 119.716 ° W]; 18 Apr. 1953; H. E. Childs leg.; EMEC • 1 ♀; Madera Co., 2 mi. NE Bass Lake; [37.355 ° N, 119.558 ° W]; 14 Jun. 1976; W. W. Middlekauff leg.; EMEC • 1 ♀; Madera Co., Coarsegold; [37.262 ° N, 119.701 ° W]; 12 May 1942; A. J. Walz leg.; UCRC • 1 ♀; Marin Co., Alpine Lake; [37.339 ° N, 121.442 ° W]; 26 May 1955; R. O. Schuster leg.; EMEC • 1 ♀; Marin Co., Mount Tamalpias; [37.929 ° N, 122.578 ° W]; 25 May 1952; LACM • 1 ♀; Marin Co., San Anselmo; [37.975 ° N, 122.562 ° W]; 7 Apr. 1939; EMEC • 2 ♀♀; Marin Co., San Geronimo; [38.013 ° N, 122.664 ° W]; 23 Apr. 1913; J. C. Bridwell leg.; EMEC • 1 ♀; Mendocino Co., 3 air mi. NW of Beaver Glade Station. Maple Creek; [39.962 ° N, 123.034 ° W]; elev. 975.4 m; 16 Jun. 1972; J. Powell leg.; ex Triteleia hyacinthina (verbatim: Brodiaea hyacinthina); EMEC • 1 ♀; Mendocino Co., 3 mi. N of Branscob; [39.695 ° N, 123.626 ° W]; 25 May 1976; S. Szerlip leg.; ex Ranunculus; EMEC • 1 ♀; Mendocino Co., 8 mi. N of Ukiah; [37.339 ° N, 121.442 ° W]; 21 Apr. 1935; E. P. Van Duzee leg.; CAS • 1 ♀; Mendocino Co., Hopland Field Station; [39.001 ° N, 123.080 ° W]; 1 May 1958; W. W. Middlekauff leg.; EMEC • 1 ♀; Mendocino Co., Ryan Creek; [39.476 ° N, 123.381 ° W]; 13 Apr. 1941; N. F. Hardman leg.; EMEC • 1 ♂; same data as for preceding; 19 Jun. 1949; EMEC • 1 ♀; Monterey Co., 5 mi. NE of Arroyo Seco Guard Station; [36.293 ° N, 121.435 ° W]; elev. 243.8 m; 4 May 1975; J. Powell leg.; ex Cryptantha; EMEC • 1 ♀; Monterey Co., 8 mi. W of Jolon; [35.970 ° N, 121.341 ° W]; 2 Apr. 1959; C. W. O’Brien leg.; EMEC • 1 ♀; Monterey Co., Arroyo Seco, 14 mi. SW of Greenfield; [36.184 ° N, 121.408 ° W]; elev. 396.2 m; 12 Jun. 1963; M. L. Noller leg.; ex Eriophyllum confertiflorum; ASUHIC • 1 ♀; Monterey Co., Bradley; [37.339 ° N, 121.442 ° W]; 23 Apr. 1917; E. P. Van Duzee leg.; CAS • 1 ♀; Monterey Co., Hastings Reservation, near Jamesburg; [36.372 ° N, 121.586 ° W]; 3 May 1958; D. D. Linsdale leg.; EMEC (Fig. 81) • 1 ♂; same data as for preceding; 8 Jul. 1958; D. D. Linsdale leg.; EMEC • 1 ♀; Monterey Co., Horse Bridge, 1.5 air mi. SW of Arroyo Seco Guard Station; [36.222 ° N, 121.495 ° W]; 6 May 1975; R. Wharton leg.; EMEC • 1 ♀; same data as for preceding; 6 May 1975; J. Powell leg.; EMEC • 1 ♀; Monterey Co., Indians Guard Staion, 17 air mi. NW of Jolon; [36.144 ° N, 121.392 ° W]; elev. 640.1 m; 9 May 1975; R. Wharton leg.; EMEC • 1 ♀; Monterey Co., Jamesburg; [36.370 ° N, 121.590 ° W]; 22 Mar. 1940; E. G. Linsley leg.; EMEC • 1 ♀; Monterey Co., Paloma Creek, 3 air mi. NE Arroyo Seco Guard Station; [36.273 ° N, 121.4499 ° W]; elev. 274.3 m; 5 May 1975; J. Powell leg.; EMEC • 3 ♀♀; Monterey Co., Robinson Canyon Road, Carmel Valley; [37.339 ° N, 121.442 ° W]; 27 Apr. 1955; J. Powell leg.; EMEC • 1 ♀; Monterey Co., Tassajara Hot Springs; [37.339 ° N, 121.442 ° W]; 27 May 1920; L. S. Slevin leg.; CAS • 1 ♀; Monterey Co., Tularcitos Ranch, Carmel Valley; [36.442 ° N, 121.646 ° W]; 27 Apr. 1955; J. Powell leg.; EMEC • 1 ♂; Napa Co., Mount Saint Helena; [37.339 ° N, 121.442 ° W]; 9 Jun. 1918; E. P. Van Duzee leg.; CAS (Fig. 82) • 1 ♀; Nevada Co., 2 mi. S of Grass Valley; [39.19 ° N, 121.061 ° W]; 27 Jun. 1967; J. Powell leg.; EMEC • 1 ♀; Riverside Co., 12 mi. NW Oak Grove on Hwy 79; [33.469 ° N, 116.946 ° W]; 5 Apr. 1969; S. Frommer, K. Reed, M. Reed leg.; UCRC • 1 ♀; Riverside Co., Riverside; [33.9556 ° N, 117.3722 ° W]; elev. 290 m; 10 May 1926; P. H. Timberlake leg.; ex Eschscholzia californica; UCRC • 1 ♀; same data as for preceding; 13 Apr. 1927; P. H. Timberlake leg.; ex Lasthenia gracilis (verbatim: Baeria gracilis); UCRC • 1 ♀; same data as for preceding; 15 Jul. 1934; P. H. Timberlake leg.; ex Gilia achilleifolia subsp. multicaulis (verbatim: Gilia multicaulis); UCRC • 1 ♀; San Diego Co., Cuyumaca; [32.946 ° N, 116.576 ° W]; 19 Apr. 1950; E. G. Linsley leg.; ex Lasthenia (verbatim: Baeria); EMEC • 1 ♀; same data as for preceding; 19 Apr. 1950; J. W. MacSwain leg.; ex Gilia; EMEC • 1 ♀; San Diego Co., Lake Cuyamaca; [32.987 ° N, 116.583 ° W]; 5 Jul. 1945; A. Melander leg.; UCRC • 1 ♀; San Diego Co., Lake Henshaw; [33.242 ° N, 116.763 ° W]; 10 Apr. 1950; P. E. Hurd leg.; EMEC • 1 ♀; San Diego Co., Mount Palomar; [33.363 ° N, 116.836 ° W]; 28 Jun. 1963; P. D. Hurd leg.; ex Cryptantha intermedia; EMEC • 1 ♀; San Francisco Co., Portola; [37.726 ° N, 122.409 ° W]; 25 Apr. 1915; EMEC • 1 ♀; San Luis Obispo Co., 10 mi. SE of Creston; [35.416 ° N, 120.398 ° W]; 1 May 1962; J. Powell leg.; EMEC • 1 ♀; San Luis Obispo Co., 7 mi. SW of Atascadero; [35.488 ° N, 120.814 ° W]; 4 Apr. 1968; C. A. Toschi leg.; EMEC • 1 ♀; same data as for preceding; 4 Apr. 1968; J. Powell leg.; ex Plagiobothrys nothofulvus; EMEC • 1 ♀; San Luis Obispo Co., 8 mi. W of Atascadero; [35.489 ° N, 120.834 ° W]; 3 Jul. 1956; P. D. Hurd leg.; ex Anthemis cotula; EMEC • 2 ♀♀; San Luis Obispo Co., Atascadero; [37.339 ° N, 121.442 ° W]; 26 Apr. 1919; E. P. Van Duzee leg.; CAS • 3 ♀♀; San Luis Obispo Co., Pozo; [37.339 ° N, 121.442 ° W]; 1 May 1962; P. D. Hurd leg.; ex Lupinus nanus; EMEC • 1 ♀; same data as for preceding; 30 Apr. 1962; R. L. Langston leg.; EMEC • 1 ♀; San Mateo Co., Kings Mtn; [37.421 ° N, 122.323 ° W]; 12 Apr. 1948; P. E. Hurd leg.; ex Frangula californica (verbatim: Rhamnus californica); EMEC • 2 ♀♀; San Mateo Co., Portola; [37.339 ° N, 121.442 ° W]; 30 May 1959; P. H. Arnaud Jr leg.; ex Verbena hastata; CAS • 1 ♀; Santa Barbara Co., Cachuma campground; Los Padres National Forest; [34.574 ° N, 119.956 ° W]; 24 – 25 May 1976; ex Frangula californica (verbatim: Rhamnus californica); LACM • 2 ♀♀; Santa Barbara Co., Los Prietos; [34.541 ° N, 119.802 ° W]; 14 Mar. 1967; J. Powell leg.; ex Plagiobothrys; EMEC • 1 ♀; same data as for preceding; 14 Mar. 1967; J. Powell leg.; EMEC • 10 ♀♀; Santa Clara Co., 11 mi. E of Mount Hamilton; [37.339 ° N, 121.442 ° W]; 19 Apr. 1956; D. J. Burdick leg.; EMEC • 9 ♀♀; Santa Clara Co., 5 mi. E of Mount Hamilton; [37.341 ° N, 121.551 ° W]; 16 Apr. 1955; D. J. Burdick leg.; EMEC • 18 ♀♀; Santa Clara Co., 8 mi. E of Mount Hamilton; [37.342 ° N, 121.496 ° W]; 16 Apr. 1955; D. J. Burdick leg.; EMEC • 1 ♀; Santa Clara Co., Mount Hamilton; [37.342 ° N, 121.643 ° W]; 15 Apr. 1947; G. E. Bohart leg.; ex Mimulus; EMEC • 19 ♀♀; same data as for preceding; 16 Apr. 1955; D. J. Burdick leg.; EMEC • 3 ♀♀; Santa Clara Co., San Antonio Valley; [37.344 ° N, 121.453 ° W]; 8 Apr. 1947; R. F. Smith leg.; EMEC • 3 ♀♀; same data as for preceding; 20 Apr. 1947; R. F. Smith leg.; EMEC • 2 ♀♀; Santa Clara Co., Stanford University; [37.430 ° N, 122.170 ° W]; 19 May 19 (??); UCRC • 2 ♀♀; Sonoma Co., Cazadero; [38.533 ° N, 123.085 ° W]; 29 Mar. 1935; R. M. Bohart and G. C. Bohart leg.; CAS • 1 ♀; Stanislaus Co., Adobe Creek W; [37.396 ° N, 121.375 ° W]; 20 Apr. 1948; R. F. Smith leg.; EMEC • 1 ♀; Tulare Co., 3 mi. W of Three Rivers; [36.437 ° N, 118.958 ° W]; 1 Apr. 1939; P. H. Timberlake leg.; ex Phacelia douglasii; UCRC (Fig. 80 C) • 1 ♀; Tulare Co., 4 mi. N of Kaweah; [36.528 ° N, 118.918 ° W]; 13 May 1963; S. W. Earnshaw leg.; EMEC • 1 ♀; Tulare Co., Three Rivers; [36.454 ° N, 118.886 ° W]; 1 Apr. 1939; P. H. Timberlake leg.; ex (?); UCRC (Fig. 80 A – B, D) • 1 ♀; Tuolumne Co., 2 mi. N of Sonora; [38.016 ° N, 120.382 ° W]; 29 Apr. 1952; ex Nemophila menziesii; LACM • 1 ♀; Tuolumne Co., Ackerson Meadow; [37.339 ° N, 121.442 ° W]; elev. 1524 m; 24 May 1969; T. W. Davies leg.; CAS • 1 ♀; Tuolumne Co., Basin Creek Campground; [37.339 ° N, 121.442 ° W]; 1 Jun. 1963; P. H. Arnaud Jr leg.; CAS • 1 ♀; Tuolumne Co., Chinese Camp; [37.870 ° N, 120.434 ° W]; 1 Apr. 1961; ex Ranunculus; LACM • 1 ♀; same data as for preceding; 2 Apr. 1960; ex Ranunculus; LACM • 1 ♀; Tuolumne Co., Sonora Pass; [38.328 ° N, 119.636 ° W]; 26 Apr. 1951; P. E. Hurd leg.; EMEC • 1 ♀; Tuolumne Co., Tuolumne; [37.962 ° N, 120.237 ° W]; elev. 1066.8 m; 9 Jun. 1938; N. F. Hardman leg.; EMEC • 1 ♂; Tuolumne Co., Twain-Harte; [37.339 ° N, 121.442 ° W]; elev. 1219.2 m; 1 – 31 Jul. 1937; F. E. Blaisdell leg.; CAS. – Oregon • 1 ♀; Josephine Co., Merlin; [42.518 ° N, 123.420 ° W]; 1 May 1970; F. A. Rushmore leg.; OSAC.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4DFF76FE13FB3CFEE0F940.taxon	description	Description GENITALIA (Figs 82, 120 O). Lasioglossum sequoiae has diagnostically distinct genital morphology when compared to other species. Gonostylus gradually widens distally, terminating in a broad apex, with long setae. The gonocoxite apex with elongate setae oriented toward broad gonostylus. Retrorse lobe curved, narrow at base, medially expanded, then gradually tapering to a pointed apex; about 4 times as long as broad and covered in fine, sparse setae.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4DFF76FE13FB3CFEE0F940.taxon	distribution	Range Almost exclusively recorded from California. USA (California, Oregon) (Fig 83).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4DFF76FE13FB3CFEE0F940.taxon	biology_ecology	Floral hosts ASTERACEAE: Anthemis cotula L., Eriophyllum confertiflorum (DC.) A. Gray, Eriophyllum lanatum (Pursh) J. Forbes, Lasthenia gracilis (DC.) Greene, Lasthenia sp., Layia platyglossa (Fisch. & C. A. Mey.) A. Gray • BORAGINACEAE: Cryptantha intermedia (A. Gray) Greene, Cryptantha sp., Plagiobothrys nothofulvus (A. Gray) A. Gray, Plagiobothrys sp. • FABACEAE: Lupinus nanus Douglas ex Benth. • HYDROPHYLLACEAE: Nemophila sp., Nemophila menziesii Hook. & Arn., Phacelia douglasii (Benth.) Torr. • IRIDACEAE: Sisyrinchium bellum S. Watson • LAMIACEAE: Salvia carduacea Benth. • LILIACEAE: Triteleia hyacinthina (Lindl.) Greene • ONAGRACEAE: Oenothera sp. • PAPAVERACEAE: Eschscholzia sp. • POLEMONIACEAE: Gilia achilleifolia subsp. multicaulis (Benth.) A. D. Grant & V. E. Grant, Gilia sp., Leptosiphon parviflorus Benth. • RANUNCULACEAE: Ranunculus sp., Ranunculus californicus Benth. • RHAMNACEAE: Frangula californica (Eschsch.) A. Gray • SCROPHULARIACEAE: Mimulus sp. • VERBENACEAE: Verbena hastata L.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4BFF7DFE71F923FB84FAD2.taxon	description	Figs 84 – 87, 100 D, 103 B, 107 B, 115 F, 117 E, 119 D, 120 P	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4BFF7DFE71F923FB84FAD2.taxon	etymology	Etymology The specific epithet is treated as a Latin adjective combing ‘ obscurus’ for ‘ obscure’ and prefix ‘ sub - ’ meaning ‘ slightly’ or ‘ somewhat’.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4BFF7DFE71F923FB84FAD2.taxon	diagnosis	Diagnosis Females of L. subobscurum can be recognised by the combination of small size, length 5.86 mm (± 0.20 SD); head wide, length / width ratio 0.88 (± 0.01 SD); mesoscutum imbricate-tessellate throughout with moderately sparse (IS = 1 – 2 PD) fine punctures; mesoscutum conspicuously covered by a layer of appressed short (<0.5 OD) pubescence; legs brown, except tarsi, bases and apices of tibiae light brown; metasoma brown with pale marginal areas; and terga with basal tomentose bands. Males of L. subobscurum can be recognised by the combination of small size, body length 5.13 mm (± 0.38 SD); wide to round head, length / width ratio 0.93 (± 0.08 SD); clypeus with yellow apical ¾; mandible short, nearly reaching opposing clypeal angle; mesoscutum largely imbricate with moderately sparse (IS = 1 – 2 PD), fine punctures; mesoscutum conspicuously covered by a layer of appressed short (<0.25 OD) pubescence; and legs brown, except tarsi, bases and apices of tibiae yellow. Females of L. subobscurum are superficially similar to L. swenki, but the latter species are usually larger, body length 7.18 mm (± 0.78 SD), possibly with orange metasoma and have doubly punctate mesoscutum, sharply angled pronotal ridge, and inner metatibial spur with basal teeth longer than width of rachis. Mesoscutum of female L. subobscurum have no double punctation on mesoscutum, and inner metatibial spur with basal teeth shorter than width of rachis. Males and females of L. subobscurum closely resemble the L. arizonense group. However, L. subobscurum is characterized by a brown metasoma with pale marginal areas, mesoscutum and overall body with a dull appearance due to microsculpturing; and a mesepisternum without distinct punctures.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4BFF7DFE71F923FB84FAD2.taxon	materials_examined	Type material Syntypes of Halictus subobscurus (only the first specimen examined) UNITED STATES – New Mexico • 1 ♀; Doña Ana Co., Mesilla; [32.270 ° N, 106.801 ° W]; 12 Apr. 1895; W. P. Cockerell leg.; ex Yellow Sisymbrium; USNM • 1 ♀; same data as for preceding; Las Cruces “ College Farm, Mesilla Valley ”; [32.28 ° N, 106.75 ° W]; 12 Apr. 1895; W. P. Cockerell leg.; ex Descurainia pinnata (verbatim: Sisymbrium canescens); USNM Type No. 3706. Specimen labels “ On yellow / Sisymbrium // Mesilla / apr. 12. / (Ckll.) // Halictus subobscurus // Cotype No 3706 / U. S. N. M. // Halictus subobscurus Ckll. ” Syntypes of Halictus Cockerelli Not examined; current location unknown. ♀. Type notes Cockerell does not designate a type explicitly. A USNM specimen labeled by Cockerell as a Halictus subobscurus co-type was examined. Following the Code (Article 72.4.1.1), even though the label does not match Cockerell’s text verbatim, since Cockerell labeled it as a cotype, it is considered part of the type series, and we treat this as a syntype; it may be that there are no other extant specimens from the type series, but this is difficult to determine. The location of the other cotype (s) of L. subobscurum is unknown and was not reported by Moure & Hurd (1987). The original description of Halictus cockerelli was based on specimens collected in New Mexico (Santa Fe and Mesilla) from 22 April to July, on old flower clusters of Virginia creeper and flowers of yellow Sisymbrium (Cockerell 1895). A syntype belonging to UCMC was not found among their material, and its current whereabouts are unknown. Material examined MEXICO – Sonora • 1 ♀; 30 km E of Agua Prieta; 31.300278 ° N, 109.2625 ° W; 3 May 2002; A. Romero leg.; ex Brassica?; PCYU • 1 ♀; Rancho San Bernardino, 28 km E of Agua Prieta; [31.327 ° N, 109.255 ° W]; 6 Jul. 2000; R. Minckley leg.; ASUHIC • 1 ♂; 30 km E of Agua Prieta; 31.316944 ° N, 109.259167 ° W; 6 Jun. 2004; N. de la Torre leg.; ex Tamarix ramosissima; ASUHIC (Fig. 86). UNITED STATES – Arizona • 1 ♀; Cochise Co., 3 mi. SW of Portal, along Cave Creek; 31.88687 ° N, 109.17358 ° W; elev. 1550 m; 15 Jul. 1987; SEMC • 1 ♀; same data as for preceding; 25 Jul. 1987; SEMC • 1 ♀; Cochise Co., Southwestern Research Station, Chiricahua Mountains; [31.884 ° N, 109.206 ° W]; 29 Jul. 1955; P. D. Hurd leg.; ex Melilotus alba; EMEC (Fig. 84) • 1 ♀; Cochise Co., Portal; [31.914 ° N, 109.141 ° W]; 4 Jun. 1968; A. G. Raske leg.; ex Sorbus; EMEC • 1 ♀; Cochise Co., Southwestern Research Station; Chiricahua Mountains; [31.884 ° N, 109.206 ° W]; 1 Jun. 1965; V. Roth leg.; UCRC • 1 ♀; Coconino Co., 3 mi. N of Sedona; [34.903 ° N, 111.758 ° W]; 23543; J. Powell leg.; ex Nolina flowers; EMEC • 2 ♂♂; Santa Cruz Co., Nogales; [31.340 ° N, 110.936 ° W]; 24 Aug. 1955; G. D. Butler and F. G. Werner leg.; ASUHIC (Fig. 85) • 1 ♀; Yavapai Co., Camp Verde; [34.564 ° N, 111.854 ° W]; 30 Aug. 1964; C. D. Michener leg.; ex Cleome lutea; SEMC • 1 ♂; Yavapai Co., Dead Horse Ranch State Park; 34.7502 ° N, 112.0217 ° W; elev. 1001 m; 5 Jun. 2011; BioBus 2011 leg.; CBG. – New Mexico • 2 ♀♀; Hidalgo Co., 16 mi. S of Animas; 31.7200 ° N, 108.8220 ° W; 16 Aug. 2024; J. Gibbs leg.; WRME. – Texas • 4 ♀♀; Presidio Co., Presidio; [29.564 ° N, 104.372 ° W]; 2 Apr. – 9 May 1941; W. L. Lowry leg.; EMEC. – Utah • 1 ♀; Washington Co., St George; [37.104 ° N, 113.584 ° W]; 14 Apr. 1962; G. F. Knowlton leg.; ex Tamarix; SEMC.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4BFF7DFE71F923FB84FAD2.taxon	description	Redescription Female BODY MEASUREMENTS. Length 5.86 mm (± 0.22 SD). Head length 1.39 mm (± 0.01 SD). Face length 1.21 mm (± 0.03 SD). Head width 1.55 mm (± 0.04 SD). Wing length 4.00 mm (± 0.10 SD). n = 6. COLOURATION. Head and mesosoma black to dark brown. Clypeus apex dark brown. Labrum dark brown. Mandible brown basally, becoming light brown medially and slightly reddish brown apically. Flagellum brown dorsally, paler ventrally with apical segments light brown. Pronotal lobe light brown. Tegula translucent light brown. Wing membrane hyaline, faintly dusky, veins with subcosta brown, otherwise light brown. Legs brown, except tarsi, bases and apices of tibiae light brown. Metasoma dark brown or brown with rims of terga, sterna and downcurved lateral areas of terga broadly pale. PUBESCENCE. Body hair colour off-white. Tomentum dense on paraocular area, gena, pronotal angle, lobe and collar, space between pronotal lobe and tegula, anterolateral angle and marginally on mesoscutum, metanotum anteromedially, propodeum lateral surface anterodorsally, T 2 basolaterally, and T 3 – T 4 as basal bands (sometime obscured by preceding segment); moderately dense on frons, preëpisternum, and other areas of propodeum lateral surface. Mesosoma covered by appressed, simple to weakly plumose, very short hairs (<0.5 OD) mixed with sparse, erect, weakly plumose, short hairs (<1 OD) on mesoscutum and erect, plumose, long hairs (<2.5 OD) on other areas. Metasomal sterna with moderately sparse, erect, weakly plumose, long (<2.5 OD) hairs. Metafemoral scopa with dense plumose hairs. Wing hairs brown, very short and sparse. SURFACE SCULPTURING. Clypeus polished apically, becoming imbricate basally; punctures large, irregularly spaced (IS <3 PD) on apically, becoming small and sparse (IS = 1 – 2 PD) basally. Supraclypeal area imbricate throughout; punctures fine, moderately dense (IS = 1 – 2 PD) and sometime punctures very fine and not visible. Frons strongly imbricate; punctures dense (IS <1 PD). Ocellocular area imbricate; punctures moderately dense (IS = 1 – 2 PD). Gena polished, becoming weakly rugulose posteriorly; punctures moderately sparse (IS = 1 – 2 PD). Postgena polished, becoming imbricate posteriorly. Tegula broadly impunctate. Mesoscutum imbricate-tessellate throughout; punctures fine, punctures dense (IS ≤ 1 PD). Mesoscutellum somewhat polished or weakly imbricate submedially, becoming imbricate marginally and on median line, punctures dense (IS ≤ 1 PD) on marginally and on median line, sparse submedially (IS = 2 – 4 PD), diversopunctate. Metanotum imbricate to rugulose, punctures obscure. Propodeal triangle with rugae moderately strong, reaching ¾ towards posterior margin at middle, anastomosing; sculpture imbricate to tessellate. Preëpisternum rugulose. Hypoepimeron rugulose to imbricate; punctures fine and obscure. Mesepisternum rugulose dorsally, becoming imbricate ventrally; punctures obscure. Propodeum lateral face imbricate, punctures obscure; posterior face imbricate, punctures obscure. T 1 anterior face polished, becoming weakly coriarious dorsally, punctures present laterally, sparse (IS = 2 – 5 PD); dorsal surface weakly coriarious to polished, with punctures moderately dense (IS = 1 – 2 PD), becoming sparser towards premarginal line (IS = 1 – 4 PD) and nearly impunctate in small subapicolateral boss, punctures minute and moderately dense (IS = 1 – 2 PD) on rim, becoming impunctate apical half and medially on rim. T 2 disc polished, becoming coriarious apically, punctures moderately dense (IS = 1 – 2 PD) basally, becoming sparse (IS = 1 – 3 PD) apically; rim coriarious, punctures fine, minute and sparse (IS = 1 – 3 PD), becoming impunctate apically. STRUCTURE. Head wide, length / width ratio 0.88 (± 0.01 SD). Face length / width ratio 0.78 (± 0.02 SD). Clypeus projecting ~ 55 % below suborbital tangent; clypeus length / width ratio (± 0.07 SD). Mandible short, reaching opposing clypeal angle. Eyes convergent below, UOD / LOD ratio 1.13 (± 0.03 SD). Ocelli normal. Gena / eye width ratio 0.61 (± 0.07 SD). Pronotal angle obtuse. Intertegular span 1.09 mm (± 0.04 SD). Mesoscutum length / width ratio 0.83 (± 0.05 SD); mesoscutum / scutellum length ratio 2.54 (± 0.16 SD); scutellum / metanotum length ratio 1.58 (± 0.17 SD); metanotum / propodeal triangle length ratio 0.76 (± 0.14 SD). Protibial spur with apical serrations shorter than width of malus. Inner metatibial spur pectinate, teeth 3 – 4, basal teeth usually shorter than width of rachis. Propodeal triangle posterior margin angled onto posterior propodeal surface. Propodeal lateral carina not reaching dorsal margin; oblique carina absent. n = 6. Description Male BODY MEASUREMENTS. Length 5.13 mm (± 0.38 SD). Head length 1.28 mm (± 0.05 SD). Face length 1.01 mm (± 0.12 SD). Head width 1.39 mm (± 0.08 SD). Wing length 3.80 mm (± 0.26 SD). n = 3. COLOURATION. Head and mesosoma black to dark brown. Clypeus yellow on apical ¾. Labrum yellow. Mandible brown basally, becoming more yellow and reddish brown on apical ⅓. Flagellum brown dorsally, light brown ventrally. Pronotal lobe yellow. Tegula translucent light brown. Wing membrane hyaline, veins with subcosta brown, otherwise light brown. Legs brown, except tarsi, bases and apices of tibiae yellow. Metasoma brown with rims of terga, sterna and downcurved lateral areas of terga broadly light brown. PUBESCENCE. Body hair colour white. Tomentum dense on face except clypeus, frons dorsally and vertex, gena, pronotal angle, lobe and collar, space between pronotal lobe and tegula, and metanotum anteromedially; moderate or sparse on frons dorsally, gena, marginally on mesoscutum preëpisternum, and propodeum lateral surface. Mesosoma covered by appressed, simple, very short hairs (<0.25 OD) mixed with very sparse, erect, weakly plumose, short hairs (<1 OD) on mesoscutum and erect, plumose, long hairs (<2 OD) on other areas. Metasomal sterna with moderately sparse, erect, plumose, long (<2.5 OD) hairs. Wing hairs brown, very short and sparse. SURFACE SCULPTURE. Clypeus polished, becoming weakly rugulose basally, punctures obscure apically and dense basally (IS <1 PD). Supraclypeal area weakly rugulose to imbricate, punctures obscure. Frons strongly imbricate, punctures dense (IS <1 PD). Ocellocular area imbricate; punctures moderately dense (IS = 1 – 2 PD). Gena polished, becoming weakly rugulose posteriorly; punctures sparse (IS = 1 – 3 PD). Postgena polished, becoming lineolate posteriorly. Tegula broadly impunctate. Mesoscutum imbricate, becoming polished submedially; punctures fine, punctures dense (IS ≤ 1 PD). Mesoscutellum polished or weakly imbricate submedially, becoming imbricate marginally and on median line, punctures dense (IS ≤ 1 PD) marginally and on median line, sparse submedially (IS = 2 – 4 PD), diversopunctate. Metanotum rugulose, punctures obscure. Propodeal triangle with rugae weak, weakly anastomosing; sculpture imbricate to tessellate. Preëpisternum rugulose. Hypoepimeron weakly rugulose; punctures minute and sparse (IS = 1 – 3 PD) becoming obscure dorsally. Mesepisternum weakly rugulose; punctures sparse (IS = 1 – 3 PD) and obscure in some angles. Propodeum lateral face imbricate, punctures obscure; posterior face imbricate, punctures obscure. T 1 anterior face polished, punctures present laterally, sparse (IS = 1 – 3 PD); dorsal surface polished, with punctures dense (IS ≤ 1 PD), becoming moderately dense (IS = 1 – 2 PD) towards premarginal line and sparse (IS = 1 – 4 PD) in small subapicolateral boss; rim coriarious, punctures fine, minute and moderately sparse (IS = 1 – 2 PD), becoming impunctate on apical half and medially on rim. T 2 disc polished, becoming coriarious apically, punctures dense (IS ≤ 1 PD) basally, becoming moderately dense (IS = 1 – 2 PD) apically; rim coriarious, punctures fine, minute and sparse (IS = 1 – 3 PD), becoming narrowly impunctate apically. STRUCTURE. Head wide to round, length / width ratio 0.93 (± 0.08 SD). Face length / width ratio 0.73 (± 0.13 SD). Clypeus projecting ~ 45 % below suborbital tangent. Mandible short, nearly reaching opposing clypeal angle. Flagellomeres short, F 1 / pedicel length ratio 0.92 (± 0.09 SD); F 2 / F 1 length ratio 1.60 (± 0.03 SD); F 2 length / width ratio 1.03 (± 0.06 SD). Gena / eye width ratio 0.46 (± 0.07 SD). Pronotal angle obtuse. Intertegular span 0.91 mm (± 0.15 SD). Mesoscutum length / width ratio 0.90 (± 0.09 SD). Propodeal lateral carina nearly reaching dorsolateral slope; oblique carina absent. n = 3. GENITALIA (Figs 86, 120 P). Gonostylus elongate and directed towards opposite one, 2.5 times as long as broad, with long setae. A prominent medial keel marks the point where the penis valves oppose each other. Retrorse lobe curved, narrow at base, medially expanded, then gradually tapering to a pointed apex; about 4 times as long as broad and covered in fine, sparse setae.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4BFF7DFE71F923FB84FAD2.taxon	distribution	Range Southern Utah to Chihuahuan Desert of southern Arizona, New Mexico, and Texas (Fig. 87).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4BFF7DFE71F923FB84FAD2.taxon	biology_ecology	Floral hosts BRASSICACEAE: Brassica sp., Descurainia pinnata (Walter) Britton, Rorippa sinuata (Nutt.) Hitchc., Sisymbrium sp. • CAPPARACEAE: Cleome lutea Hook. • FABACEAE: Melilotus albus Medik. • LILIACEAE: Nolina sp. • ROSACEAE: Sorbus sp. • TAMARICACEAE: Tamarix sp., Tamarix ramosissima Ledeb.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF4BFF7DFE71F923FB84FAD2.taxon	discussion	DNA barcodes Five DNA barcodes are available from Arizona, which form a discrete cluster.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF40FF04FEB3FABDFE76FE5F.taxon	description	Figs 88 – 91, 100 A, 106 E, 112 F, 115 E, 119 F, 120 Q	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF40FF04FEB3FABDFE76FE5F.taxon	diagnosis	Diagnosis Lasioglossum villosulum females can be recognized by the following combination of characters: body entirely dark, head round, length / width ratio 0.95 (± 0.01 SD), mesoscutal punctation sparse (IS = 1 – 2.5 PD), inner metatibial spur with numerous short branches (subequal to length of main rachis), and metasoma without basolateral tomentum. They are most similar to L. sequoiae, which can be distinguished by the denser mesoscutal punctation (IS <1.5 PD). Male L. villosulum are similar to females but have a distinct yellow maculation on the clypeus. Further distinguishing characters are clypeus flat to medially depressed, mandibles and labrum dark; mesepisternum is distinctly punctate, mesoscutal punctation sparse, tegula ovoid, and retrorse lobe large and broad. They are perhaps most similar to dark males of L. aspilurus, which have the clypeus more evenly rounded, mandible and labrum yellow.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF40FF04FEB3FABDFE76FE5F.taxon	etymology	Etymology The specific epithet is treated as a Latin adjective, a diminutive of the Latin ‘ villosus ’ meaning ‘ hairy, shaggy’.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF40FF04FEB3FABDFE76FE5F.taxon	materials_examined	Material examined CANADA – British Columbia • 1 ♂; BBR Barrowtown; 49.0417 ° N, 122.0417 ° W; 19 Aug. 2021; A. Braz leg.; RSKM • 2 ♀♀; Coquitlam, Widgeon Marsh R. P; 49.3625 ° N, 122.6435 ° W; 21 Sep. 2020; SEM Team leg.; UBCZ • 1 ♀; Delta; 49.098611 ° N, 123.043611 ° W; 17 May 2016; M. Clausen leg.; UBCZ • 1 ♀; same data as for preceding; 2 Jun. 2016; Hengel K. leg.; UBCZ • 1 ♀; Delta; 49.058611 ° N, 123.076111 ° W; 2 Jun. 2016; M. Clausen leg.; UBCZ • 2 ♂♂; Hornby Island; 49.524 ° N, 124.64905 ° W; 3 – 5 Sep. 2015; B. Zand leg.; RSKM • 1 ♂; Hornby Island, Helliwell Provincial Park; 49.5178 ° N, 124.59608 ° W; 21 Aug. – 15 Sep. 2015; B. Zand leg.; RSKM • 1 ♀; Oakhaven Park, Central Saanich; 48.564167 ° N, 123.447778 ° W; 4 Jun. 2017; T. Kelly leg.; ex Hypochaeris radicata; SFU • 1 ♂; Vancouver, PSRP, Acadia Meadow; 49.27837 ° N, 123.24701 ° W; 17 Aug. 2020; SEM Team leg.; UBCZ • 1 ♀; Vernon, CFB Vernon; 50.24629 ° N, 119.28875 ° W; 15 Sep. 2017; T. Ehiers leg.; RSKM. UNITED STATES – Oregon • 1 ♀; Lane Co., Coburg Ridge, near Eugene; 44.101614 ° N, 122.998153 ° W; 19 Jun. 2017; E. Merlo leg.; ex Hypochaeris radicata; SFU • 2 ♀♀; Benton Co., Bluebird Strip, William L. Finley National Wildlife Refuge; 44.418872 ° N, 123.330236 ° W; 27 May 2017; E. Merlo leg.; ex Hypochaeris radicata; SFU • 1 ♀; Clackamas Co., Milwaukie, 123; 45.448933 ° N, 122.620337 ° W; 21 Jun. 2017; G. Langellotto leg.; OSAC • 1 ♀; Clackamas Co., Milwaukie, 54; 45.448933 ° N, 122.620337 ° W; 26 Jun. 2017; G. Langellotto leg.; OSAC • 2 ♀♀; Clackamas Co., West Linn, 9; 45.381328 ° N, 122.633538 ° W; 21 Jun. 2017; G. Langellotto leg.; OSAC • 1 ♂; Clackamas Co., Milwaukie, 54; 45.448933 ° N, 122.620337 ° W; 26 Jun. 2017; G. Langellotto leg.; OSAC • 2 ♂♂; Clackamas Co., West Linn, 9; 45.381328 ° N, 122.633538 ° W; 24 Jun. 2017; G. Langellotto leg.; OSAC • 1 ♀; Clackamas Co., Boring Trailhead; 45.432 ° N, 122.375 ° W; 12 Jun. 2021; J. D. Engler leg.; ex Taraxacum; OSAC • 1 ♀; Clackamas Co., Eagle Creek, Heiple Rd; 45.31666 ° N, 122.36611 ° W; 10 Jun. 2011; J. D. Engler leg.; OSAC • 1 ♀; Malheur Co., Willow Creek, near Eugene; 44.028839 ° N, 123.173722 ° W; 22 May 2017; E. Merlo leg.; ex Linum lewisii; SFU • 2 ♀♀; Marion Co., Kingston Prairie, near Stayton; 44.777214 ° N, 122.744086 ° W; 28 May 2017; E. Elle leg.; ex Hypochaeris radicata; SFU • 2 ♀♀; same data as for preceding; 28 May 2017; T. Kelly leg.; ex Potentilla gracilis; SFU • 2 ♀♀; same data as for preceding; 29 May 2017; E. Merlo leg.; ex Linum lewisii; SFU • 3 ♀♀; same data as for preceding; 29 May 2017; E. Merlo leg.; ex Hypochaeris radicata; SFU • 1 ♀; same data as for preceding; 29 May 2017; T. Kelly leg.; ex Linum lewisii; SFU • 3 ♀♀; Multnomah Co., Portland; 45.484752 ° N, 122.608543 ° W; 28 Jun. 2017; G. Langellotto leg.; OSAC • 1 ♀; same data as for preceding; 23 Jun. 2017; G. Langellotto leg.; OSAC • 1 ♀; same data as for preceding; 29 Jun. 2017; G. Langellotto leg.; OSAC • 1 ♀; same data as for preceding; 26 Jun. 2017; G. Langellotto leg.; OSAC • 1 ♀; same data as for preceding; 21 Jun. 2017; G. Langellotto leg.; OSAC • 1 ♀; same data as for preceding; 27 Jun. 2017; G. Langellotto leg.; OSAC • 3 ♂♂; same data as for preceding; 25 Jun. 2017; G. Langellotto leg.; OSAC • 1 ♀; Multnomah Co., Oxbow Meadow; 45.485 ° N, 122.302 ° W; 12 Jun. 2021; J. D. Engler leg.; ex Leucanthemum; OSAC • 1 ♀; Multnomah Co., Oxbow Meadow; 45.4844 ° N, 122.302 ° W; 28 May 2021; J. D. Engler leg.; ex Spergula arvensis; OSAC • 1 ♀; same data as for preceding; 28 May 2021; J. D. Engler leg.; ex Geranium molle; OSAC • 1 ♀; same data as for preceding; 4 Aug. 2020; J. D. Engler leg.; ex Asteraceae; OSAC • 1 ♀; same data as for preceding; 4 Aug. 2020; J. D. Engler leg.; OSAC • 2 ♀♀; same data as for preceding; 19 May 2021; J. D. Engler leg.; ex Taraxacum; OSAC • 1 ♂; Multnomah Co., Bensen Sp.; 45.577 ° N, 122.131 ° W; 30 Jun. 2021; J. D. Engler leg.; ex Tarax / Hypochaeris; OSAC • 2 ♀♀; Multnomah Co., Oxbow Meadow; 45.485 ° N, 122.302 ° W; 2 Jul. 2021; J. D. Engler leg.; OSAC • 2 ♂♂; same data as for preceding; 28 Jul. 2021; J. D. Engler leg.; ex Tarax / Hypochaeris; OSAC • 3 ♀♀; same data as for preceding; 12 Oct. 2021; J. D. Engler leg.; ex Hypochaeris; OSAC • 1 ♀; Multnomah Co., Oxbow Meadow; 45.484 ° N, 122.302 ° W; 2 Jun. 2021; J. D. Engler leg.; OSAC • 1 ♀; Multnomah Co., Oxbow Meadow; 45.485 ° N, 122.302 ° W; 7 Jun. 2021; J. D. Engler leg.; ex Hypochaeris; OSAC • 1 ♀; same data as for preceding; 15 Jun. 2022; J. D. Engler leg.; OSAC • 1 ♂; same data as for preceding; 24 Sep. 2021; J. D. Engler leg.; ex Taraxacum; OSAC (Fig. 90) • 1 ♂; same data as for preceding; 24 Sep. 2021; J. D. Engler leg.; ex Hypochaeris; OSAC • 1 ♂; same data as for preceding; 16 Sep. 2021; J. D. Engler leg.; OSAC • 4 ♂♂; same data as for preceding; 23 Jul. 2021; J. D. Engler leg.; ex Tarax / Hypochaer; OSAC • 1 ♀; Tillamook Co., Nestucca Bay; 46.1678 ° N, 123.9523 ° W; 4 – 5 Jun. 2017; W. Medlen leg.; OSAC • 1 ♀; Tillamook Co., Nestucca Bay; 46.1653 ° N, 123.9544 ° W; 4 – 5 Jun. 2017; W. Medlen leg.; OSAC • 1 ♀; Tillamook Co., Nestucca Bay; 46.1643 ° N, 123.9531 ° W; 4 – 5 Jun. 2017; W. Medlen leg.; OSAC • 1 ♀; same data as for preceding; 2 – 3 Jul. 2017; W. Medlen leg.; OSAC • 2 ♀♀; Tillamook Co., Nestucca Bay; 46.1653 ° N, 123.9544 ° W; 2 – 3 Jul. 2017; W. Medlen leg.; OSAC • 1 ♀; Tillamook Co., Nestucca Bay; 46.1643 ° N, 123.9531 ° W; 21 – 22 May 2017; W. Medlen leg.; OSAC • 4 ♀♀; Tillamook Co., Nestucca Bay; 46.1653 ° N, 123.9544 ° W; 4 – 5 Jul. 2017; W. Medlen leg.; OSAC • 1 ♀; Tillamook Co., Nestucca Bay; 46.1643 ° N, 123.9531 ° W; 4 – 5 Jul. 2017; W. Medlen leg.; OSAC • 1 ♀; same data as for preceding; 18 – 19 Jun. 2017; W. Medlen leg.; OSAC • 6 ♀♀; Tillamook Co., Nestucca Bay; 46.1643 ° N, 123.9523 ° W; 21 – 22 Jun. 2017; W. Medlen leg.; OSAC • 2 ♀♀; Tillamook Co., Nestucca Bay; 46.1643 ° N, 123.9531 ° W; 30 Apr. – 1 May 2017; W. Medlen leg.; OSAC • 1 ♀; Tillamook Co., Nestucca Bay; 46.1653 ° N, 123.9544 ° W; 30 Apr. – 1 May 2017; W. Medlen leg.; OSAC • 3 ♀♀; Tillamook Co., Nestucca Bay; 46.1643 ° N, 123.9531 ° W; 4 – 5 May 2017; W. Medlen leg.; OSAC. – Washington • 1 ♀; Clark Co., Steigerwald Lake National Wildlife Refuge; 45.57215 ° N, 122.30483 ° W; 6 Jun. 2011; G. Fuquay leg.; OSAC • 1 ♀; same data as for preceding; 24 Jun. 2011; G. Fuquay leg.; OSAC • 1 ♀; Clark Co., Steigerwald Lake National Wildlife Refuge; 45.5698 ° N, 122.2981 ° W; 6 Jun. 2011; G. Fuquay leg.; OSAC • 2 ♀♀; Clark Co., Ridgefield National Wildlife Refuge, Wig Lk; 45.817 ° N, 122.773 ° W; 24 May 2011; A. Chmielewski leg.; OSAC • 2 ♀♀; Clark Co., Lacamas Lake Regional Park, near Camas; 45.601733 ° N, 122.398961 ° W; 1 Jun. 2017; T. Kelly leg.; ex Hypochaeris radicata; SFU • 1 ♀; same data as for preceding; 31 May 2017; E. Merlo leg.; ex Hypochaeris radicata; SFU • 1 ♀; same data as for preceding; 1 Jun. 2017; E. Merlo leg.; ex Hypochaeris radicata; SFU • 1 ♀; same data as for preceding; 31 May 2017; E. Merlo leg.; ex Hypochaeris radicata; SFU • 2 ♀♀; King Co., Seattle; 47.4238 ° N, 122.3071 ° W; 8 Aug. 2015; R. Redmond and E. A. Sugden leg.; WRME • 1 ♀; same data as for preceding; 29 Sep. 2014; R. Redmond and E. A. Sugden leg.; WRME • 1 ♀; same data as for preceding; 29 Apr. 2019; R. Redmond and E. A. Sugden leg.; WRME • 1 ♀; King Co., Seattle; 47.9307 ° N, 122.2853 ° W; 12 Jul. 2015; E. A. Sugden and W. Peterman leg.; WRME • 1 ♀; King Co., Seattle; 47.4301 ° N, 122.3051 ° W; 11 Jul. 2015; R. Redmond and E. A. Sugden leg.; WRME • 1 ♀; King Co., Seattle; 47.4233 ° N, 122.3085 ° W; 29 May 2019; E. A. Sugden and W. Peterman leg.; WRME • 1 ♀; King Co., Seattle; 47.4292 ° N, 122.3035 ° W; 29 May 2019; E. A. Sugden and W. Peterman leg.; WRME • 1 ♀; King Co., Seattle; 47.423 ° N, 122.3076 ° W; 29 May 2019; E. A. Sugden and W. Peterman leg.; WRME • 1 ♀; King Co., Seattle; 47.4236 ° N, 122.3102 ° W; 29 May 2019; E. A. Sugden and W. Peterman leg.; WRME • 1 ♀; King Co., Seattle; 47.4285 ° N, 122.3063 ° W; 29 May 2019; E. A. Sugden and W. Peterman leg.; WRME • 2 ♂♂; King Co., Seattle; 47.4263 ° N, 122.3061 ° W; 6 Aug. 2018; E. A. Sugden and W. Peterman leg.; WRME • 1 ♂; King Co., Seattle; 47.4285 ° N, 122.3063 ° W; 5 Jul. 2019; E. A. Sugden and W. Peterman leg.; WRME • 5 ♂♂; same data as for preceding; 31 Jul. 2019; E. A. Sugden and W. Peterman leg.; WRME • 3 ♂♂; King Co., Seattle; 47.4276 ° N, 122.3087 ° W; 21 Sep. 2019; E. A. Sugden and W. Peterman leg.; WRME • 3 ♂♂; King Co., Seattle; 47.506 ° N, 122.2674 ° W; 11 Jul. 2015; E. A. Sugden and W. Peterman leg.; WRME • 1 ♂; same data as for preceding; 11 Jul. 2015; R. Redmond and W. Peterman leg.; WRME • 2 ♂♂; King Co., Seattle; 47.5039 ° N, 122.2757 ° W; 11 Jul. 2015; R. Redmond and W. Peterman leg.; WRME • 1 ♂; King Co., Seattle; 47.5029 ° N, 122.2851 ° W; 14 Jul. 2014; R. Redmond and W. Peterman leg.; WRME • 1 ♂; King Co., Seattle; 47.5053 ° N, 122.2655 ° W; 16 Jul. 2016; R. Redmond and W. Peterman leg.; WRME • 3 ♂♂; King Co., Seattle; 47.4301 ° N, 122.3051 ° W; 11 Jul. 2015; R. Redmond and W. Peterman leg.; WRME (Fig. 89) • 1 ♂; King Co., Seattle; 47.4292 ° N, 122.3035 ° W; 31 Jul. 2019; E. A. Sugden and W. Peterman leg.; WRME • 2 ♀♀; Pacific Co., Wallapa National Wildlife Refuge; 46.3689 ° N, 124.0273 ° W; 10 Jun. 2017; W. Ritchie leg.; OSAC • 1 ♀; Pacific Co., Wallapa National Wildlife Refuge; 46.36886 ° N, 124.02731 ° W; 30 May 2016; W. Ritchie leg.; OSAC • 3 ♀♀; Pacific Co., Wallapa National Wildlife Refuge, OSB 3; 46.3689 ° N, 124.0273 ° W; 21 Apr. 2017; W. Ritchie leg.; OSAC • 7 ♀♀; Pacific Co., Wallapa National Wildlife Refuge, TRL; 46.36886 ° N, 124.02731 ° W; 8 Jun. 2015; W. Ritchie leg.; OSAC • 2 ♂♂; Pierce Co.; 47.013 ° N, 122.448 ° W; elev. 115 m; 24 Jul. 2009; C. Fimbel leg.; PCYU • 7 ♀♀; Pierce Co.; 47.056 ° N, 122.457 ° W; elev. 120 m; 23 Jun. 2009; C. Fimbel leg.; PCYU • 1 ♀; Pierce Co.; 47.03828 ° N, 122.504 ° W; elev. 117 m; 23 Jun. 2009; C. Fimbel leg.; PCYU • 1 ♀; Pierce Co.; 47.012 ° N, 122.429 ° W; elev. 125 m; 24 Jul. 2009; C. Fimbel leg.; PCYU • 3 ♀♀, 1 ♂; Pierce Co.; 47.105 ° N, 122.505 ° W; elev. 88 m; 24 Jul. 2009; C. Fimbel leg.; PCYU • 1 ♀; Pierce Co.; 47.105 ° N, 122.505 ° W; elev. 88 m; 39974; C. Fimbel leg.; PCYU • 1 ♀; Snohomish Co., Everett; 47.9297 ° N, 122.285 ° W; 29 May 2019; E. A. Sugden and W. Peterman leg.; WRME • 1 ♀; Snohomish Co., Everett; 47.93 ° N, 122.2852 ° W; 29 May 2019; E. A. Sugden and W. Peterman leg.; WRME • 1 ♂; Snohomish Co., Everett; 47.933 ° N, 122.2741 ° W; 30 Jul. 2018; E. A. Sugden and W. Peterman leg.; WRME • 1 ♂; Snohomish Co., Everett; 47.4285 ° N, 122.3036 ° W; 7 Jul. 2018; E. A. Sugden and W. Peterman leg.; WRME • 4 ♂♂; Snohomish Co., Everett; 47.93 ° N, 122.2852 ° W; 5 Jul. 2019; E. A. Sugden and W. Peterman leg.; WRME • 3 ♂♂; Snohomish Co., Everett; 47.9297 ° N, 122.285 ° W; 31 Jul. 2019; E. A. Sugden and W. Peterman leg.; WRME • 1 ♂; Snohomish Co., Everett; 48.9328 ° N, 122.2748 ° W; 31 Jul. 2019; E. A. Sugden and W. Peterman leg.; WRME • 1 ♂; Thurston Co.; 46.914 ° N, 122.718 ° W; elev. 120 m; 30 May 2009; C. Fimbel leg.; PCYU • 3 ♂♂; Thurston Co.; 46.953 ° N, 123.025 ° W; elev. 57 m; 23 May 2009; C. Fimbel leg.; PCYU • 2 ♂♂; Thurston Co.; 46.914 ° N, 122.709 ° W; elev. 170 m; 23 Jun. 2009; C. Fimbel leg.; PCYU • 2 ♂♂; Thurston Co.; 46.903 ° N, 122.739 ° W; elev. 125 m; 23 Jun. 2009; C. Fimbel leg.; PCYU • 1 ♂; Thurston Co.; 46.924 ° N, 122.735 ° W; elev. 113 m; 27 May 2009; C. Fimbel leg.; PCYU • 2 ♀♀; Wahkiakum Co., Julia Butler Hansen Refuge; 46.241 ° N, 123.4487 ° W; 31 May – 1 Jun. 2016; K. Brenner leg.; OSAC.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF40FF04FEB3FABDFE76FE5F.taxon	description	Description GENITALIA (Figs 90, 120 Q). Gonostylus broad and develops into a plate from the apical view, 1.5 times as long as broad, with setae short, clearly branched. Retrorse lobe about 2 times as long as broad, with irregular margin, and rounded at apex.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF40FF04FEB3FABDFE76FE5F.taxon	distribution	Range Pacific northwest of USA (Oregon, Washington) and southern British Columbia (Fig. 91).	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF40FF04FEB3FABDFE76FE5F.taxon	biology_ecology	Floral hosts ASTERACEAE: Hypochaeris sp., Hypochaeris radicata L., Leucanthemum sp., Taraxacum sp. or Hypochaeris sp., Taraxacum sp. • CARYOPHYLLACEAE: Spergula arvensis L. • GERANIACEAE: Geranium molle L. • LINACEAE: Linum lewisii Pursh • ROSACEAE: Potentilla gracilis Douglas ex Hook.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF40FF04FEB3FABDFE76FE5F.taxon	discussion	Remarks The earliest North American record of L. villosulum dates back to 2009 in Pierce County, Washington, based on the available data. Recent samples indicate that this species is increasing in abundance, suggesting a potential population expansion. DNA barcodes Pauly et al. (2019) analyzed DNA barcodes of L. villosulum from across its range. North American sequences from British Columbia, Oregon, and Washington match Palaearctic data. Dubious record section This species is retained in this section due to the unavailability of the holotype. The original description hints at a possible affiliation with the subgenus Evylaeus, as indicated by the metatibial spur described with 3 – 4 oblique, nearly obsolete serrations and wide head – characteristics commonly observed in the subgenus Evylaeus.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF39FF05FE1FFE08FC19F8E0.taxon	etymology	Etymology The specific epithet is a reference to the type locality, Pecos, New Mexico, with the adjectival suffix - ensis denoting place.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF39FF05FE1FFE08FC19F8E0.taxon	biology_ecology	Floral hosts ROSACEAE: Holodiscus australis A. Heller.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
03EC87C3FF39FF05FE1FFE08FC19F8E0.taxon	discussion	Remarks The location of the type is unknown and no specimens identified as this species have ever been seen. The depository should be the USNM (Moure & Hurd 1987), but it has not been located there to date. Crawford (1906 a) compares it to three species of L. (Hemihalictus), although based on the description and characters provided in a subsequent key (Crawford 1907 a), this may belong to the subgenus L. (Evylaeus). Based on the type locality it could plausibly be a junior synonym of L. amicus (Cockerell, 1897). Keys to species of L. (Hemihalictus) in North America Geographic data can help discriminate species (see Fig. 92, Table 3), but readers should refer to the following key and the species diagnoses to confirm identifications.	en	Hettiarachchi, Thilina, Gibbs, Jason (2026): Review of the bee subgenus Lasioglossum (Hemihalictus) in western North America (Hymenoptera: Halictidae). European Journal of Taxonomy 1085: 1-232, DOI: 10.5852/ejt.2026.1085.3316, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3316/14568
