identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E02C60CF53DE00E3B12FB548F0E828.text	03E02C60CF53DE00E3B12FB548F0E828.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus Erichson 1839	<div><p>Key to species of the genus  Diochus of Indonesia, Malaysia and Singapore</p><p>(Figs 1 −11)</p><p>1 Antennomeres longer: antennomere 3 more elongate, distinctly longer than antennomere 2; antennomere 4 more elongate, more than twice as long as wide.............................................................................. 2</p><p>- Antennomeres shorter: antennomere 3 less elongate, nearly equal to antennomere 2; antennomere 4 less elongate, less than twice as long as wide.................................................................................. 7</p><p>2 Tibiae partly in black color; known only from Borneo (Kalimantan, Indonesia).................................................................................................................  D. borneensis Cameron, 1933</p><p>- Tibiae not in black color................................................................................ 3</p><p>3 Body larger, longer than 6 mm; posterior margin of male sternite VIII emarginate medially, and with shallower emarginations on both sides......................................................................................... 4</p><p>- Body smaller, shorter than 5.5 mm; posterior margin of male sternite VIII only emarginate medially, or with deeper emarginations on both sides......................................................................................... 5</p><p>4 Posterior margin of male sternite VIII with medial emargination deep and rounded, posterior angles strongly sharpened.....................................................................  D. paradoxus Huang, Janák &amp; Zhou,  sp. nov.</p><p>- Posterior margin of male sternite VIII with medial emargination shallow, not rounded, posterior angle slightly sharpened................................................................  D. paraparadoxus Huang, Janák &amp; Zhou,  sp. nov.</p><p>5 Posterior margin of male sternite VIII with medial emargination very deep, posterior angles strongly sharpened and projecting; known only from Borneo (Kalimantan, Indonesia)......................  D. parellipticus Huang, Janák &amp; Zhou,  sp. nov.</p><p>- Posterior margin of male sternite VIII with medial emargination shallow, not so rounded, with posterior angles not sharply projecting........................................................................................... 6</p><p>6 Head and pronotum more elongate; male sternite VIII distinctly narrowed at apex; aedeagus with parameres shorter and thicker........................................................  D. sulawesiensis Huang, Janák &amp; Zhou,  sp. nov.</p><p>- Head and pronotum less elongate; male sternite VIII not distinctly narrowed at apex; aedeagus with parameres longer and thinner.................................................................  D. punctipennis (Motschulsky, 1858)</p><p>7 Elytra not in a uniform color, but in bicolor; basal transverse carina of male and female sternite VIII interrupted or absent................................................................................  D. pulchellus Cameron, 1918</p><p>- Elytra in a uniform color; basal transverse carina of male or female sternite VIII entire.............................. 8</p><p>8 Posterior margin of male sternite VIII nearly straight at middle, with slight emargination on both sides; basal margin of male sternite IX narrower than apical margin...............................  D. lombokensis Huang, Janák &amp; Zhou,  sp. nov.</p><p>- Posterior margin of male sternite VIII emarginate at middle; basal margin of male sternite IX wider than apical margin..... 9</p><p>9 Posterior margin of male sternite VIII emarginate at middle and on both sides; aedeagus with parameres very short........................................................................  D. bogorensis Huang, Janák &amp; Zhou,  sp. nov.</p><p>- Posterior margin of male sternite VIII only emarginate at middle; aedeagus with parameres thin and long.............. 10</p><p>10 Pronotum more elongate; posterior margin of male sternite VIII with medial emargination deeper; basal transverse carina of male sternite VIII deeply produced at middle...........................  D. uvellipticus Huang, Janák &amp; Zhou,  sp. nov.</p><p>- Pronotum not so elongate; posterior margin of male sternite VIII with medial emargination shallower; basal transverse carina of male sternite VIII shallowly produced at middle.......................................................... 11</p><p>11 Basal transverse carina of male sternite VIII sharp at middle; aedeagus with a T-shaped sclerite distally; body darker...............................................................................  D. antennatus (Motschulsky, 1858)</p><p>- Basal transverse carina of male sternite VIII rounded at middle; aedeagus without T-shaped sclerite distally; body lighter....................................................................  D. acehensis Huang, Janák &amp; Zhou,  sp. nov.</p></div>	https://treatment.plazi.org/id/03E02C60CF53DE00E3B12FB548F0E828	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang (2025): Revision of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) from Indonesia, Malaysia and Singapore. Zootaxa 5632 (3): 441-479, DOI: 10.11646/zootaxa.5632.3.2, URL: https://doi.org/10.11646/zootaxa.5632.3.2
03E02C60CF50DE04E3B12DD84856EB7A.text	03E02C60CF50DE04E3B12DD84856EB7A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus acehensis Huang, Janak & Zhou 2025	<div><p>1.  Diochus acehensis Huang, Janák &amp; Zhou,  sp. nov.</p><p>(Figs 1L; 2-1; 2-2)</p><p>Type locality. Indonesia: Sumatra: Aceh.</p><p>Type material.   Holotype: male, INDONESIA: Sumatra: Aceh #25a,  Mt. Leuser NP, 300–500 m,  Ketambe, 23–30. XI. 1989, Löbl, Agosti, Burckhardt (MHNG)  .   Paratypes: INDONESIA: Sumatra: 5 males, Aceh #26a,  Mt. Leuser NP, Ketambe, 800 m, 28. XI. 1989, Löbl, Agosti, Burckhardt (MHNG, JJRC) ;   2 males,  W Sum., Padangpanjan, 600 m, 17. XI. 1989, Agosti, Löbl, Burckhardt (MHNG) ;   1 female, W Sum. #18,  Anai V. Nat. Res., 10 km W Padangpanjan, 250 m, 17. XI. 1989, Löbl, Agosti, Burckhardt (MHNG) ;   1 female, W Sum. #19,  Anai V. Nat. Res., 10 km W Padangpanjan, 200 m, 17. XI. 1989, Löbl, Agosti, Burckhardt (MHNG) ;   2 females, W Sum. #21,  Palopo Nat. Res. N, Bukittinggi, 900 m, 18–20. XI. 1989, Löbl, Agosti, Burckhardt (MHNG, JJRC)  .</p><p>Measurements. BL = 3.93 mm, FL = 1.73 mm, HL = 0.50 mm, HW = 0.40 mm, EyL = 0.10 mm, TL = 0.29 mm, ANL = 1.00 mm, ANT1 = 0.15 mm, ANT2 = 0.09 mm, ANT3 = 0.09 mm, ANT4 = 0.07 mm, ANT 11 = 0.11 mm, PL = 0.65 mm, PW = 0.52 mm, EL = 0.61 mm, EW = 0.71 mm, SL = 0.44 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, small-sized. Body light brown, abdominal intersegmental membrane lighter. Legs yellowish brown, tarsi distinctly lighter. Antennae brown to light brown, apex of each antennomere lighter. Maxillary and labial palpi yellowish brown.</p><p>Head (Fig 2-1A). Trapezoid in shape, 1.27 times as long as wide. Tempora distinctly widened backwards, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of head with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye small-sized, not distinctly protruding laterad, longitudinal diameter distinctly shorter than half the length of tempora (eye: tempora = 0.35). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.11 mm) distinctly longer than distance from antenna to eye (0.09 mm). Ventral surface with sparsely scattered punctures, interspaces between them as large as about 4 puncture diameters. Gular sutures deep, approaching to each other before base, but not confluent.</p><p>Antennae (Fig 2-1B). Scape rod-shaped, slightly thickened apically, slightly shorter than two subsequent antennomeres combined; antennomere 2 not distinctly elongate; antennomere 3 about the length of 2; length of antennomere 4 about 1.3 times of width, antennomere 4 to 10 nearly equal in length, but gradually widened; antennomere 11 distinctly longer than 10.</p><p>Mouthparts (Fig 2-1A). Labrum nearly hexagonal, lateral margins rounded, widest at basal 1/2; anterior margin about half of width at the widest point. Mandibles (Fig 2-1D) falciform, left one with three small teeth on inner margin, whereas right one with two.</p><p>Neck (Fig 2-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.13 mm, slightly shorter than 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.</p><p>Prothorax (Fig 2-1A). Pronotum elongate (PL to PW ratio 1.26), distinctly longer and wider than head. Anterior region near anterior angles slightly deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures composed of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards.</p><p>Pterothorax (Fig 2-1A). Mesoscutellum triangular and small, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; paired oblique furrows short but deep; carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.</p><p>Elytra (Fig 2-1A). Elytra distinctly transverse (EL to EW ratio 0.86), shorter but wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 5 punctures on disc, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 2-1C). First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi about the length of 2−4 together, that of meso-, meta- slightly shorter than the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal microsculpture, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male (Figs 2-1E–J; 2-2A–E). Head without elevation between eyes. Posterior margin of sternite VIII slightly emarginate medially, basal transverse carina wave-shaped, emarginate medially (Figs 2-1E; 2-2B). Posterior margin of tergite VIII arcuate (Figs 2-1F; 2-2C). Tergite IX (Figs 2-1H; 2-2D) symmetrical, connected mediobasally, narrowed apically. Sternite IX (Figs 2-1G; 2-2E) symmetrical, widest at basal 1/3 to 1/2; basal margin deeply emarginate, apical margin slightly emarginate; width of apical margin slightly wider than basal margin. Tergite X (Figs 2-1H; 2-2D) symmetrical, triangular. Aedeagus (Figs 2-1 I−J; 2-2A) asymmetrical, small-sized, ca. 0.38 mm long, normally sclerotized. Parameres symmetrical, reaching above top of median lobe. Internal structures sclerotized, with a heart-shaped sclerite and two elongated sclerites in middle. Sperm pump coiled, thinned toward apex.</p><p>Female (Figs 2-2 F−I). Head without elevation between eyes. Sternite VIII (Fig 2-2F) with posterior margin rounded. Tergite IX bearing numerous setae, elongate and sharply pointed apically. Sternite IX (Fig 2-2H) symmetrical, with deep concaved margin in middle. Tergite X broad, basal margin curved. Sternite X (Fig 2-2G) slender and rhomboidal, with numerous setae in lower half. Tube of spermatheca thin and long, with a loop apically (Fig 2-2I).</p><p>Distribution. Indonesia (Sumatra).</p><p>Diagnosis. This new species is small in size, with light coloration and short antennae, and males have a head without an elevation between eyes. Males can be easily separated from congeneric species but are similar to  D. antennatus, which also has a similar aedeagus. However, this species is distinctly smaller in size, and the aedeagus lacks a T-shaped sclerite distally.</p><p>Etymology. The species epithet is derived from the name of type locality, Aceh (Indonesia, Sumatra).</p></div>	https://treatment.plazi.org/id/03E02C60CF50DE04E3B12DD84856EB7A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang (2025): Revision of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) from Indonesia, Malaysia and Singapore. Zootaxa 5632 (3): 441-479, DOI: 10.11646/zootaxa.5632.3.2, URL: https://doi.org/10.11646/zootaxa.5632.3.2
03E02C60CF54DE0BE3B12E3E4850EB56.text	03E02C60CF54DE0BE3B12E3E4850EB56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus bogorensis Huang, Janak & Zhou 2025	<div><p>2.  Diochus bogorensis Huang, Janák &amp; Zhou,  sp. nov.</p><p>(Figs 1H; 3-1; 3-2)</p><p>Type locality. Indonesia: Java: Bogor.</p><p>Type material.  Holotype: male, INDONESIA: Java: Bogor, 28. XI. 1991.78, J. T. Huber collected (MHNG).</p><p>Measurements. BL = 4.52 mm, FL = 2.01 mm, HL = 0.54 mm, HW = 0.44 mm, EyL = 0.13 mm, TL = 0.32 mm, ANT1 = 0.17 mm, ANT2 = 0.11 mm, ANT3 = 0.11 mm, ANT4 = 0.09 mm, PL = 0.76 mm, PW = 0.59 mm, EL = 0.71 mm, EW = 0.75 mm, SL = 0.54 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, medium-sized. Body light brown, abdominal intersegmental membrane lighter. Legs yellowish brown, tarsi distinctly lighter. Antennae yellowish brown, apex of each antennomere lighter. Maxillary and labial palpi yellowish brown.</p><p>Head (Fig 3-1A). Oblong in shape, 1.24 times as long as wide. Tempora straight and parallel, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of head with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye small-sized, distinctly protruding laterad, longitudinal diameter slightly shorter than half the length of tempora (eye: tempora = 0.42). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.10 mm) distinctly longer than distance from antenna to eye (0.07 mm). Ventral surface with sparsely scattered punctures, interspaces between them as large as about 4 puncture diameters. Gular sutures deep, confluent near base.</p><p>Antennae. Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 slightly elongate; antennomere 3 nearly equal to 2; length of antennomere 4 about 1.3 times of width.</p><p>Mouthparts (Fig 3-1A). Labrum oval-shaped, lateral margins rounded, widest at basal 1/2; anterior margin about 1/3 of width at the widest point. Mandibles falciform, left one with three small teeth on inner margin, whereas right one with two.</p><p>Neck (Fig 3-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.16 mm, slightly longer than 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.</p><p>Prothorax (Fig 3-1A). Pronotum elongate (PL to PW ratio 1.29), distinctly longer and wider than head.Anterior region near anterior angles deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures composed of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards.</p><p>Pterothorax (Fig 3-1A). Mesoscutellum triangular, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; paired oblique furrows short but deep; carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.</p><p>Elytra (Fig 3-1A). Elytra distinctly transverse (EL to EW ratio 0.89), shorter but wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 6 punctures on disc, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 3-1B). First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi slightly longer than 2−4 together, that of meso-, meta- slightly shorter than the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal microsculpture, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male (Figs 3-1D–I; 3-2A–E). Head without elevation between eyes. Posterior margin of sternite VIII slightly emarginate medially, also deeply emarginate on each side, with posterior angles sharpened (Figs 3-1D; 3-2B). Posterior margin of tergite VIII arcuate (Figs 3-1E; 3-2C). Tergite IX (Figs 3-1G; 3-2D) symmetrical, connected mediobasally, narrowed apically. Sternite IX (Figs 3-1F; 3-2E) symmetrical, widest at basal 1/3 to 1/2; basal margin deeply emarginate, apical margin slightly emarginate; width of apical margin about 2/3 of basal margin. Tergite X (Figs 3-1G; 3-2D) symmetrical, triangular. Aedeagus (Figs 3-1H−I; 3-2A) symmetrical, small-sized, ca. 0.56 mm long, normally sclerotized. Parameres symmetrical, reaching near middle of median lobe. Internal structures sclerotized: two paired irregular sclerites located on both sides; an Y-shaped sclerite and an irregular sclerite in middle. Sperm pump coiled, thinned toward apex.</p><p>Female. Unknown.</p><p>Distribution. Indonesia (Java).</p><p>Diagnosis. This new species is similar to  D. acehensis sp. nov. and  D. pulchellus which are small in size and with light coloration. The posterior margin of male sternite VIII in this new species is emarginate at middle and also strongly emarginate on both sides, while in  D. acehensis sp. nov. and  D. pulchellus it is only emarginate medially.</p><p>Etymology. The species epithet is derived from the name of type locality, Bogor (Indonesia, Java Is.).</p></div>	https://treatment.plazi.org/id/03E02C60CF54DE0BE3B12E3E4850EB56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang (2025): Revision of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) from Indonesia, Malaysia and Singapore. Zootaxa 5632 (3): 441-479, DOI: 10.11646/zootaxa.5632.3.2, URL: https://doi.org/10.11646/zootaxa.5632.3.2
03E02C60CF58DE0FE3B129E5499FE8DE.text	03E02C60CF58DE0FE3B129E5499FE8DE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus lombokensis Huang, Janak & Zhou 2025	<div><p>3.  Diochus lombokensis Huang, Janák &amp; Zhou,  sp. nov.</p><p>(Figs 1I; 4-1; 4-2)</p><p>Type locality. Indonesia:  Lombok .</p><p>Type material.   Holotype: male, INDONESIA: LOMBOK:  Mt. Rinjiani, Senaro, 400 m, Waterfalls, 5. XI. 1991, I. Löbl (MHNG)  .  Paratypes: INDONESIA: LOMBOK: 1 male, same data as holotype (MHNG);   Bali: 1 female,  Lake Buyan, 1200 m, 8 - 9. XI. 1991, forest, I. Löbl, litter (MHNG) ;   Timor NTT: 1 male, 3 females, 11 km N Soe, 30. 3. 1991, F91856, D. Agosti, everg. for. (MHNG, JJRC)  .</p><p>Measurements. BL = 4.00 mm, FL = 1.96 mm, HL = 0.56 mm, HW = 0.42 mm, EyL = 0.15 mm, TL = 0.30 mm, ANL = 1.08 mm, ANT1 = 0.15 mm, ANT2 = 0.10 mm, ANT3 = 0.11 mm, ANT4 = 0.08 mm, ANT 11 = 0.12 mm, PL = 0.72 mm, PW = 0.58 mm, EL = 0.73 mm, EW = 0.78 mm, SL = 0.57 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, small to medium-sized. Body brown to dark brown, abdominal intersegmental membrane lighter. Legs yellowish brown, tarsi distinctly lighter. Antennae dark brown to light brown, apex of each antennomere lighter. Maxillary and labial palpi yellowish brown.</p><p>Head (Fig 4-1A). Rounded rectangle in shape, 1.33 times as long as wide. Tempora straight and parallel, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of head with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye medium-sized, distinctly protruding laterad, longitudinal diameter about half the length of tempora (eye: tempora = 0.51). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.13 mm) distinctly longer than distance from antenna to eye (0.07 mm). Ventral surface with scattered punctures, interspaces between them as large as about 4 puncture diameters. Gular sutures deep, confluent at basal 1/6, then extended to base.</p><p>Antennae (Fig 4-1B). Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 elongate; antennomere 3 slightly longer than 2; length of antennomere 4 about 1.5 times of width, antennomere 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.</p><p>Mouthparts (Fig 4-1A). Labrum oval-shaped, lateral margins rounded, widest at basal 1/2; anterior margin narrow, about 1/5 of width at the widest point. Mandibles (Fig 4-1D) falciform, left one with three small teeth on inner margin, whereas right one with two.</p><p>Neck (Fig 4-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.13 mm, shorter than 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.</p><p>Prothorax (Fig 4-1A). Pronotum elongate (PL to PW ratio 1.23), distinctly longer and wider than head. Anterior region near anterior angles slightly deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures composed of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards.</p><p>Pterothorax (Fig 4-1A). Mesoscutellum triangular and small, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; paired oblique furrows short but deep; carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.</p><p>Elytra (Fig 4-1A). Elytra slightly transverse (EL to EW ratio 0.95), longer and wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 5 punctures on disc, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 4-1C). First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi slightly shorter than 2−4 together, that of meso-, meta- slightly shorter than the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal microsculpture, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male (Figs 4-1E–J; 4-2A–E). Head with small round elevation between eyes. Posterior margin of sternite VIII not emarginate medially, but nearly straight, with several short setae, each side truncate (Figs 4-1E; 4-2B). Posterior margin of tergite VIII arcuate (Figs 4-1F; 4-2C). Tergite IX (Figs 4-1H; 4-2D) symmetrical, connected mediobasally, narrowed apically. Sternite IX (Figs 4-1G; 4-2E) symmetrical, widest at 1/2 to 1/3; basal margin straight, apical margin normally emarginate; width of apical margin about 1.5 times of basal margin. Tergite X (Figs 4-1H; 4-2D) symmetrical, triangular. Aedeagus (Figs 4-1I−J; 4-2A) symmetrical, medium-sized, ca. 0.60 mm long, normally sclerotized. Parameres symmetrical, extremely thick and short, reaching to about half of median lobe. Internal structures sclerotized, paired L-shaped sclerites on upper middle, in black color; two paired strips shaped sclerites located on both sides, in black color; 3 sclerites in middle: paired rectangular sclerites in brown color, one Y-shaped sclerite in brown to black color. Sperm pump coiled, thinned toward apex.</p><p>Female (Figs 4-2F−I). Head without elevation between eyes. Sternite VIII (Fig 4-2F) with posterior margin rounded. Tergite IX bearing numerous setae, elongate and sharply pointed apically. Sternite IX (Fig 4-2H) symmetrical, with deep concave margin in middle. Tergite X broad, basal margin curved. Sternite X (Fig 4-2G) slender and rhomboidal, with numerous setae in lower half. Tube of spermatheca sclerotized, long and coiled (Fig 4-2I).</p><p>Distribution. Indonesia.</p><p>Diagnosis. This species is related to the Australian species  D. octavii Fauvel, 1877 and  D. pubiventris Lea, 1929 based on the similar aedeagal structure, also with similar shape of male sternite VIII: medial part of posterior margin straight and with short setae. Compared to  D. octavii, this new species has distinctly lighter coloration, posterior margin of male sternite VIII with medial straight part distinctly wider, and male aedeagus with parameres thicker and shorter. Compared to  D. pubiventris, male sternite VIII of this new species is with posterior angles truncate, not rounded, and basal margin of sternite IX is wider.</p><p>Etymology. The species epithet is derived from the name of type locality, Lombok (Indonesia).</p></div>	https://treatment.plazi.org/id/03E02C60CF58DE0FE3B129E5499FE8DE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang (2025): Revision of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) from Indonesia, Malaysia and Singapore. Zootaxa 5632 (3): 441-479, DOI: 10.11646/zootaxa.5632.3.2, URL: https://doi.org/10.11646/zootaxa.5632.3.2
03E02C60CF5FDE12E3B12D81494CEBA2.text	03E02C60CF5FDE12E3B12D81494CEBA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus paradoxus Huang, Janak & Zhou 2025	<div><p>4.  Diochus paradoxus Huang, Janák &amp; Zhou,  sp. nov.</p><p>(Figs 1B; 5-1; 5-2)</p><p>Type locality. Indonesia: Sumatra: Aceh.</p><p>Type material.   Holotype: male, INDONESIA: Sumatra: Aceh #25b,  Mt. Leuser NP, 300–500 m,  Ketambe, 23–30. XI. 1989, Löbl, Agosti, Burckhardt (MHNG)  .   Paratypes: INDONESIA: Sumatra: 1 male, 1 female, Aceh #25a,  Mt. Leuser NP, 300–500 m,  Ketambe, 23–30. XI. 1989, Löbl, Agosti, Burckhardt (MHNG)  .</p><p>Measurements. BL = 6.75 mm, FL = 2.59 mm, HL = 0.74 mm, HW = 0.54 mm, EyL = 0.19 mm, TL = 0.44 mm, ANL = 1.62 mm, ANT1 = 0.27 mm, ANT2 = 0.13 mm, ANT3 = 0.18 mm, ANT4 = 0.14 mm, ANT 11 = 0.17 mm, PL = 1.00 mm, PW = 0.78 mm, EL = 0.92 mm, EW = 1.04 mm, SL = 0.66 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, large-sized. Body dark brown, abdominal intersegmental membrane lighter. Legs reddish brown, tarsi distinctly lighter. Antennae brown to reddish brown, apex of each antennomere lighter. Maxillary and labial palpi yellowish brown.</p><p>Head (Fig 5-1A). Oblong in shape, 1.38 times as long as wide. Tempora straight and parallel, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of head with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye small-sized, distinctly protruding laterad, longitudinal diameter slightly shorter than half the length of tempora (eye: tempora = 0.43). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.16 mm) distinctly longer than distance from antenna to eye (0.07 mm). Ventral surface with sparsely scattered punctures, interspaces between them as large as about 5 puncture diameters. Gular sutures deep, confluent near basal 1/4 to 1/3, then extended to base.</p><p>Antennae (Fig 5-1B). Scape rod-shaped, slightly thickened apically, slightly shorter than two subsequent antennomeres combined; antennomere 2 elongate; antennomere 3 distinctly longer than 2; length of antennomere 4 about twice of width, antennomere 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.</p><p>Mouthparts (Fig 5-1A). Labrum nearly hexagonal, lateral margins rounded, widest at basal 1/2; anterior margin about half of width at the widest point. Mandibles (Fig 5-1D) falciform, left one with three small teeth on inner margin, whereas right one with two.</p><p>Neck (Fig 5-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.18 mm, about 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.</p><p>Prothorax (Fig 5-1A). Pronotum elongate (PL to PW ratio 1.27), distinctly longer and wider than head.Anterior region near anterior angles deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures composed of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards.</p><p>Pterothorax (Fig 5-1A). Mesoscutellum triangular and small, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; paired oblique furrows short but deep; carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.</p><p>Elytra (Fig 5-1A). Elytra distinctly transverse (EL to EW ratio 0.89), shorter but wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 5 punctures on disc, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 5-1C). First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi slightly shorter than 2−4 together, that of meso-, meta- slightly shorter than the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal microsculpture, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male (Figs 5-1E–J; 5-2A–E). Head with an elongate elevation between eyes. Posterior margin of sternite VIII emarginate medially, emargination deep and rounded, also emarginate on each side, with posterior angles sharpened (Figs 5-1E; 5-2B). Posterior margin of tergite VIII arcuate (Figs 5-1F; 5-2C). Tergite IX (Figs 5-1H; 5- 2D) symmetrical, connected mediobasally, narrowed apically. Sternite IX (Figs 5-1G; 5-2E) symmetrical, widest at basal 1/3; basal margin nearly straight, apical margin slightly emarginate; width of apical margin about 2/3 of basal margin. Tergite X (Figs 5-1H; 5-2D) symmetrical, triangular. Aedeagus (Figs 5-1I−J; 5-2A) symmetrical, medium-sized, ca. 0.67 mm long, normally sclerotized. Parameres symmetrical, reaching top of median lobe. Internal structures sclerotized, an irregular shaped large sclerite and paired rectangular shaped sclerites in middle; an elongated sclerite and a round sclerite at bottom. Sperm pump coiled, thinned toward apex.</p><p>Female (Figs 5-2F−I). Head without elevation between eyes. Sternite VIII (Fig 5-2F) with posterior margin rounded. Tergite IX bearing numerous setae, elongate and sharply pointed apically. Sternite IX (Fig 5-2H) symmetrical, with deep concaved margin in middle. Tergite X broad, basal margin curved. Sternite X (Fig 5-2G) slender and rhomboidal, with numerous setae in lower half. Tube of spermatheca sclerotized and wavy in middle part, with three bends (Fig 5-2I).</p><p>Distribution. Indonesia (Sumatra).</p><p>Diagnosis. According to nearly similar aedeagal structures, we suggest this new species and  D. paraparadoxus sp. nov. to be closely allied, but the former can be distinguished from the latter by smaller body size, distinctly deeper emargination on the posterior margin of male sternite VIII and longer parameres of the aedeagus.</p><p>Etymology. The species epithet is from the latinized Greek word paradoxus.</p></div>	https://treatment.plazi.org/id/03E02C60CF5FDE12E3B12D81494CEBA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang (2025): Revision of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) from Indonesia, Malaysia and Singapore. Zootaxa 5632 (3): 441-479, DOI: 10.11646/zootaxa.5632.3.2, URL: https://doi.org/10.11646/zootaxa.5632.3.2
03E02C60CF43DE11E3B129E54C35EB9F.text	03E02C60CF43DE11E3B129E54C35EB9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus paraparadoxus Huang, Janak & Zhou 2025	<div><p>5.  Diochus paraparadoxus Huang, Janák &amp; Zhou,  sp. nov.</p><p>(Figs 1A; 6-1; 6-2)</p><p>Type locality. Indonesia: Sumatra: Jambi.</p><p>Type material.   Holotype: male, INDONESIA: Jambi:  W Mt Tujuh Lake, 1400 m, 14. XI. 1989, Agosti, Löbl, Burckhardt (MHNG).</p><p>Measurements. BL = 7.97 mm, FL = 3.07 mm, HL = 0.87 mm, HW = 0.62 mm, EyL = 0.19 mm, TL = 0.54 mm, ANL = 1.76 mm, ANT1 = 0.26 mm, ANT2 = 0.15 mm, ANT3 = 0.19 mm, ANT4 = 0.14 mm, ANT 11 = 0.17 mm, PL = 1.18 mm, PW = 0.91 mm, EL = 1.04 mm, EW = 1.28 mm, SL = 0.74 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, large-sized. Body dark brown, abdominal intersegmental membrane lighter. Legs brown, tarsi distinctly lighter. Antennae brown, apex of each antennomere lighter. Maxillary and labial palpi yellowish brown.</p><p>Head (Fig 6-1A). Oblong in shape, distinctly elongate, 1.41 times as long as wide. Tempora straight and parallel, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of head with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye small-sized, not distinctly protruding laterad, longitudinal diameter slightly shorter than half the length of tempora (eye: tempora = 0.35). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.20 mm) distinctly longer than distance from antenna to eye (0.13 mm). Ventral surface with sparsely scattered punctures, interspaces between them as large as about 5 puncture diameters. Gular sutures deep, confluent near basal 1/3, then extended to base.</p><p>Antennae (Fig 6-1B). Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 elongate; antennomere 3 distinctly longer than 2; length of antennomere 4 about twice of width, antennomere 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.</p><p>Mouthparts (Fig 6-1A). Labrum nearly hexagonal, lateral margins rounded, widest at basal 1/2; anterior margin about half of width at the widest point. Mandibles (Fig 6-1D) falciform, left one with three small teeth on inner margin, whereas right one with two.</p><p>Neck (Fig 6-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.18 mm, about 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.</p><p>Prothorax (Fig 6-1A). Pronotum elongate (PL to PW ratio 1.30), distinctly longer and wider than head.Anterior region near anterior angles deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures composed of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards.</p><p>Pterothorax (Fig 6-1A). Mesoscutellum triangular, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; paired oblique furrows short but deep; carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.</p><p>Elytra (Fig 6-1A). Elytra distinctly transverse (EL to EW ratio 0.81), shorter but wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 5 punctures on disc, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 6-1C). First four segments of protarsi heart-shaped, slightly dilated, and those of meso- and metatarsi slender. Last segment of protarsi longer than 2−4 together, that of meso-, meta- slightly shorter than the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal microsculpture, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male (Figs 6-1E–J; 6-2A–E). Head with an elongated elevation between eyes. Posterior margin of sternite VIII emarginate medially, also shallowly emarginate on each side, with posterior angles sharpened (Figs 6-1E; 6-2B). Posterior margin of tergite VIII arcuate (Figs 6-1F; 6-2C). Tergite IX (Figs 6-1H; 6-2D) symmetrical, connected mediobasally, narrowed apically. Sternite IX (Figs 6-1G; 6-2E) symmetrical, widest at basal 1/3; basal margin nearly straight, basal angles sharpened, apical margin slightly emarginated; width of apical margin about 2/3 of basal margin. Tergite X (Figs 6-1H; 6-2D) symmetrical, triangular. Aedeagus (Figs 6-1I−J; 6-2A) symmetrical, large-sized, ca. 0.90 mm long, normally sclerotized. Parameres symmetrical, reaching at apical 1/5 of median lobe. Internal structures sclerotized: paired flag-shaped sclerites and paired rectangular shaped sclerites in middle; a small oval sclerite and a round sclerite at bottom. Sperm pump coiled, thinned toward apex.</p><p>Female. Unknown.</p><p>Distribution. Indonesia (Sumatra).</p><p>Diagnosis. This new species is extremely large in size, with distinctly elongate head and pronotum. The new species is similar to  D. paradoxus sp. nov. and  D. punctipennis in aedeagal structures (all with a round sclerite at base), but has shorter parameres. The medial emargination of male sternite VIII in this new species is distinctly shallower than  D. paradoxus sp. nov., and the posterior angles are sharp, which differs from  D. punctipennis . This species is externally similar to  D. borneensis, known only from females but has distinctly paler tibiae.</p><p>Etymology. The species epithet is from the Greek terms, para - and  paradoxus, referring to the great similarity with that species.</p></div>	https://treatment.plazi.org/id/03E02C60CF43DE11E3B129E54C35EB9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang (2025): Revision of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) from Indonesia, Malaysia and Singapore. Zootaxa 5632 (3): 441-479, DOI: 10.11646/zootaxa.5632.3.2, URL: https://doi.org/10.11646/zootaxa.5632.3.2
03E02C60CF46DE15E3B129E54C60EFDA.text	03E02C60CF46DE15E3B129E54C60EFDA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus parellipticus Huang, Janak & Zhou 2025	<div><p>6.  Diochus parellipticus Huang, Janák &amp; Zhou,  sp. nov.</p><p>(Figs 1D; 7-1; 7-2)</p><p>Type locality. Malaysia: Sabah.</p><p>Type material.   Holotype: male, MALAYSIA: Sabah:  Crocker Range, 1550–1650 m, 16. V. 1987, Burckhardt-Löbl (MHNG)  .  Paratypes: MALAYSIA: Sabah: 1 female, same data as holotype (MHNG) .</p><p>Measurements. BL = 5.27 mm, FL = 2.46 mm, HL = 0.66 mm, HW = 0.54 mm, EyL = 0.19 mm, TL = 0.36 mm, ANL = 1.57 mm, ANT1 = 0.21 mm, ANT2 = 0.13 mm, ANT3 = 0.18 mm, ANT4 = 0.13 mm, ANT 11 = 0.16 mm, PL = 0.85 mm, PW = 0.70 mm, EL = 0.92 mm, EW = 0.95 mm, SL = 0.68 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, medium-sized. Body reddish to dark brown, abdominal intersegmental membrane lighter. Legs yellowish brown, tarsi distinctly lighter. Antennae brown to reddish brown, apex of each antennomere lighter. Maxillary and labial palpi yellowish brown.</p><p>Head (Fig 10-1A). Oblong in shape, 1.22 times as long as wide. Tempora slightly widened backwards, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of head with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye large-sized, distinctly protruding laterad, longitudinal diameter slightly shorter than half the length of tempora (eye: tempora = 0.52). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.15 mm) distinctly longer than distance from antenna to eye (0.06 mm). Ventral surface with sparsely scattered punctures, interspaces between them as large as about 4 puncture diameters. Gular sutures deep, approaching to each other before basal 1/4, but not confluent.</p><p>Antennae (Fig 7-1B). Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 distinctly elongate; antennomere 3 distinctly longer than 2; length of antennomere 4 about 2.5 times of width, antennomere 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.</p><p>Mouthparts (Fig 7-1A). Labrum transverse, lateral margins rounded, widest at basal 1/2; anterior margin about 1/3 of width at the widest point. Mandibles (Fig 7-1D) falciform, left one with three small teeth on inner margin, whereas right one with two.</p><p>Neck (Fig 7-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.18 mm, about 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.</p><p>Prothorax (Fig 7-1A). Pronotum elongate (PL to PW ratio 1.22), distinctly longer and wider than head.Anterior region near anterior angles deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures composed of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards.</p><p>Pterothorax (Fig 7-1A). Mesoscutellum triangular, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; paired oblique furrows short but deep; carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.</p><p>Elytra (Fig 7-1A). Elytra slightly transverse (EL to EW ratio 0.96), distinctly longer and wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 5 punctures on disc, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 7-1C). First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi shorter than 2−4 together, that of meso-, meta-slightly shorter than the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal microsculpture, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male (Figs 7-1E–J; 7-2A–E). Head with a small round elevation between eyes. Posterior margin of sternite VIII emarginate medially, emargination distinctly deep and rounded, with posterior angles strongly sharpened (Figs 7-1E; 7-2B). Posterior margin of tergite VIII arcuate (Figs 7-1F; 7-2C). Tergite IX (Figs 7-1H; 7-2D) symmetrical, connected mediobasally, narrowed apically. Sternite IX (Figs 7-1G; 7-2E) symmetrical, widest at basal 1/3 to 1/2; basal margin normally emarginate, apical margin slightly emarginate; width of apical margin about half of basal margin. Tergite X (Figs 7-1H; 7-2D) symmetrical, triangular. Aedeagus (Figs 7-1I−J; 7-2A) symmetrical, medium-sized, ca. 0.73 mm long, normally sclerotized. Parameres symmetrical, reaching about apical 1/5 of median lobe. Internal structures sclerotized, paired elongated sclerites and a small symmetrical sclerite in middle; a heart-shaped sclerite and a Y-shaped sclerite at bottom. Sperm pump coiled, thinned toward apex.</p><p>Female (Figs 7-2F−I). Head without elevation between eyes. Sternite VIII (Fig 7-2F) with posterior margin rounded. Tergite IX bearing numerous setae, elongate and sharply pointed apically. Sternite IX (Fig 7-2H) symmetrical, with deep concaved margin in middle. Tergite X broad, basal margin curved. Sternite X (Fig 7-2G) slender and rhomboidal, with numerous setae in lower half. Tube of spermatheca thin and sclerotized, with two bends (Fig 7-2I).</p><p>Distribution. Malaysia (Sabah).</p><p>Diagnosis. The new species is similar to  D. uvellipticus sp. nov. and  D. sulawesiensis sp. nov. in body size and coloration. Compared to  D. uvellipticus sp. nov., this species has distinctly longer antennae, and the male sternite VIII is with medial emargination deeper and more rounded. Compared to  D. sulawesiensis sp. nov., this species has less elongate head, posterior margin of the male sternite VIII is deeply emarginate, and the aedeagus has distinctly longer parameres.</p><p>Etymology. The species epithet is derived from Greek terms, par - and ellipt-, to indicate the relatively elongate pronotum.</p></div>	https://treatment.plazi.org/id/03E02C60CF46DE15E3B129E54C60EFDA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang (2025): Revision of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) from Indonesia, Malaysia and Singapore. Zootaxa 5632 (3): 441-479, DOI: 10.11646/zootaxa.5632.3.2, URL: https://doi.org/10.11646/zootaxa.5632.3.2
03E02C60CF45DE18E3B12A854996EF6A.text	03E02C60CF45DE18E3B12A854996EF6A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus sulawesiensis Huang, Janak & Zhou 2025	<div><p>7.  Diochus sulawesiensis Huang, Janák &amp; Zhou,  sp. nov.</p><p>(Figs 1E; 8-1; 8-2)</p><p>Type locality. Indonesia: Sulawesi.</p><p>Type material.   Holotype: male, INDONESIA: Sulawesi:  Sel., W of Mamasa, 1000 m, 11. IV. 1991, D. Agosti, F91741 (MHNG).</p><p>Measurements. BL = 5.22 mm, FL = 2.38 mm, HL = 0.69 mm, HW = 0.50 mm, EyL = 0.15 mm, TL = 0.40 mm, ANL = 1.40 mm, ANT1 = 0.19 mm, ANT2 = 0.13 mm, ANT3 = 0.14 mm, ANT4 = 0.11 mm, ANT 11 = 0.13 mm, PL = 0.89 mm, PW = 0.68 mm, EL = 0.79 mm, EW = 0.95 mm, SL = 0.60 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, medium-sized. Body black, abdominal intersegmental membrane lighter. Legs dark brown, tarsi distinctly lighter. Antennae brown to black brown, apex of each antennomere lighter. Maxillary and labial palpi dark brown.</p><p>Head (Fig 8-1A). Oblong in shape, 1.38 times as long as wide. Tempora straight and parallel, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of head with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye small-sized, not distinctly protruding laterad, longitudinal diameter distinctly shorter than half the length of tempora (eye: tempora = 0.39). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.15 mm) distinctly longer than distance from antenna to eye (0.09 mm). Ventral surface with sparsely scattered punctures, interspaces between them as large as about 4 puncture diameters. Gular sutures deep, confluent near the base.</p><p>Antennae (Fig 8-1B). Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 elongate; antennomere 3 slightly longer than 2; length of antennomere 4 about twice of width, antennomere 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.</p><p>Mouthparts (Fig 8-1A). Labrum oval-shaped, lateral margins rounded, widest at basal 1/2; anterior margin about half of width at the widest point. Mandibles (Fig 8-1D) falciform, left one with three small teeth on inner margin, whereas right one with two.</p><p>Neck (Fig 8-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.15 mm, shorter than 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.</p><p>Prothorax (Fig 8-1A). Pronotum elongate (PL to PW ratio 1.31), distinctly longer and wider than head.Anterior region near anterior angles deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures composed of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards.</p><p>Pterothorax (Fig 8-1A). Mesoscutellum triangular, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; paired oblique furrows short but deep; carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.</p><p>Elytra (Fig 8-1A). Elytra distinctly transverse (EL to EW ratio 0.89), shorter but wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 5 punctures on disc, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 8-1C). First four segments of protarsi heart-shaped, slightly dilated, and those of meso- and metatarsi slender. Last segment of protarsi slightly shorter than 2−4 together, that of meso-, meta- slightly shorter than the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal microsculpture, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male (Figs 8-1E–J; 8-2A–E). Head with an oval elevation between eyes. Sternite VIII with posterior margin emarginate medially, lateral margins strongly narrowed backwards (Figs 8-1E; 8-2B). Posterior margin of tergite VIII arcuate (Figs 8-1F; 8-2C). Tergite IX (Figs 8-1H; 8-2D) symmetrical, connected mediobasally, narrowed apically. Sternite IX (Figs 8-1G; 8-2E) symmetrical, widest at basal 1/3; basal margin moderately emarginate, apical margin deeply emarginate; width of apical margin about 2/3 of basal margin. Tergite X (Figs 8-1H; 8-2D) symmetrical, triangular. Aedeagus (Figs 8-1I−J; 8-2A) symmetrical, medium-sized, ca. 0.59 mm long, normally sclerotized. Parameres symmetrical, thick and short, reaching near half of median lobe. Internal structures sclerotized: paired flag-shaped sclerites and paired rectangular shaped sclerites in middle; an elongated sclerite at bottom. Sperm pump coiled, thinned toward apex.</p><p>Female. Unknown.</p><p>Distribution. Indonesia (Sulawesi).</p><p>Diagnosis. This new species is entirely black in color and male sternite VIII is strongly narrowed to apex, nearly in the form of an inverted triangle, easily distinguishing it from other species.</p><p>Etymology. The species epithet is derived from the name of type locality, Sulawesi (Indonesia).</p></div>	https://treatment.plazi.org/id/03E02C60CF45DE18E3B12A854996EF6A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang (2025): Revision of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) from Indonesia, Malaysia and Singapore. Zootaxa 5632 (3): 441-479, DOI: 10.11646/zootaxa.5632.3.2, URL: https://doi.org/10.11646/zootaxa.5632.3.2
03E02C60CF48DE1FE3B12A154C60EFB9.text	03E02C60CF48DE1FE3B12A154C60EFB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus uvellipticus Huang, Janak & Zhou 2025	<div><p>8.  Diochus uvellipticus Huang, Janák &amp; Zhou,  sp. nov.</p><p>(Figs 1F; 9-1; 9-2)</p><p>Type locality. Indonesia: Java.</p><p>Type material.   Holotype: male, INDONESIA: Java:  Col du Puncak et Cibodas, lac Telaga Warna, 1. VIII. 84, J. Robert (MHNG)  .  Paratypes: INDONESIA: Java: 1 male, same data as holotype (MHNG) .</p><p>Measurements. BL = 5.18 mm, FL = 2.24 mm, HL = 0.62 mm, HW = 0.47 mm, EyL = 0.17 mm, TL = 0.32 mm, ANL = 1.23 mm, ANT1 = 0.19 mm, ANT2 = 0.11 mm, ANT3 = 0.12 mm, ANT4 = 0.08 mm, ANT 11 = 0.15 mm, PL = 0.77 mm, PW = 0.63 mm, EL = 0.81 mm, EW = 0.91 mm, SL = 0.65 mm.</p><p>Description. Body long, narrowed anteriorly and posteriorly, large-sized. Body dark brown, abdominal intersegmental membrane lighter. Legs brown, tarsi distinctly lighter. Antennae brown to dark brown, apex of each antennomere lighter. Maxillary and labial palpi yellowish brown.</p><p>Head (Fig 9-1A). Rounded rectangle in shape, 1.31 times as long as wide. Tempora slightly widened backwards, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of head with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye medium-sized, distinctly protruding laterad, longitudinal diameter slightly longer than half the length of tempora (eye: tempora = 0.52). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.15 mm) distinctly longer than distance from antenna to eye (0.07 mm). Ventral surface with sparsely scattered punctures, interspaces between them as large as about 4 puncture diameters. Gular sutures deep, approaching each other before basal 1/4, but not confluent, then parallel to base.</p><p>Antennae (Fig 9-1B). Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 elongate; antennomere 3 slightly longer than 2; length of antennomere 4 about 1.3 times of width, antennomere 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.</p><p>Mouthparts (Fig 9-1A). Labrum transverse, lateral margins rounded, widest at basal 1/2; anterior margin about half of width at the widest point. Mandibles (Fig 9-1D) falciform, left one with three small teeth on inner margin, whereas right one with two.</p><p>Neck (Fig 9-1A). Cylindrical, shiny, only bearing shallow microsculpture, width 0.15 mm, slightly shorter than 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.</p><p>Prothorax (Fig 9-1A). Pronotum elongate (PL to PW ratio 1.22), distinctly longer and wider than head.Anterior region near anterior angles deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures composed of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards.</p><p>Pterothorax (Fig 9-1A). Mesoscutellum triangular, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; paired oblique furrows short but deep; carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.</p><p>Elytra (Fig 9-1A). Elytra distinctly transverse (EL to EW ratio 0.89), longer and wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 5 punctures on disc, also with 4−5 rows of punctures on deflexed portion.</p><p>Legs (Fig 9-1C). First four segments of protarsi heart-shaped, dilated, and those of meso- and metatarsi slender. Last segment of protarsi about the length of 2−4 together, that of meso-, meta- slightly shorter than the length of each 2−3.</p><p>Abdomen. Broadest at segment V. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal microsculpture, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p><p>Male (Figs 9-1E–J; 9-2A–E). Head with a round elevation between eyes. Posterior margin of sternite VIII emarginate medially, basal transverse carina emarginate deeply (Figs 9-1E; 9-2B). Posterior margin of tergite VIII arcuate (Figs 9-1F; 9-2C). Tergite IX (Figs 9-1H; 9-2D) symmetrical, connected mediobasally, narrowed apically. Sternite IX (Figs 9-1G; 9-2E) symmetrical, widest near basal 1/2; basal margin deeply emarginate, apical margin slightly emarginate; width of apical margin slightly shorter than basal margin. Tergite X (Figs 9-1H; 9- 2D) symmetrical, triangular. Aedeagus (Figs 9-1I−J; 9-2A) symmetrical, large-sized, ca. 0.80 mm long, normally sclerotized. Parameres symmetrical, reaching above top of median lobe. Internal structures sclerotized: a triangle sclerite on top; paired irregular sclerites in middle. Sperm pump coiled, thinned toward apex.</p><p>Female. Unknown.</p><p>Distribution. Indonesia (Java).</p><p>Diagnosis. This species is related to the Oriental species  D. bisegmentatus Zhou &amp; Zhou, 2016 based on the similar aedeagal structure. This new species is distinctly larger in size, with more elongated pronotum and the first four segments of protarsi are not so strongly dilated. Regarding the short antennae, this new species is also similar to  D. antennatus,  D. acehensis sp. nov. and  D. pulchellus . This new species can be separated from those by the longer pronotum and the deep emargination at the middle of posterior margin of male sternite VIII.</p><p>Etymology. The species epithet is derived from Greek terms, uv - and ellipt-, to indicate the elongate pronotum.</p></div>	https://treatment.plazi.org/id/03E02C60CF48DE1FE3B12A154C60EFB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang (2025): Revision of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) from Indonesia, Malaysia and Singapore. Zootaxa 5632 (3): 441-479, DOI: 10.11646/zootaxa.5632.3.2, URL: https://doi.org/10.11646/zootaxa.5632.3.2
03E02C60CF4FDE1FE3B12D7F4EC6EB86.text	03E02C60CF4FDE1FE3B12D7F4EC6EB86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus antennatus (Motschulsky 1858)	<div><p>9.  Diochus antennatus (Motschulsky, 1858)</p><p>(Fig 1J)</p><p>Rhegmatocerus antennatus Motschulsky, 1858: 659 (Type locality: Indes orientales).</p><p>Diochus antennatus; Gemminger and Harold, 1868: 608 (catalog); Bernhauer and Schubert, 1914:319 (catalog); Bernhauer, 1922: 231 (Formosa); Bernhauer, 1928: 15 (Sumatra); Cameron, 1932: 46 (characters; Penang; Malay Peninsula); Scheerpeltz, 1933: 1323 (catalog; Malay-Halbinsel, Formosa, Neu Guinea); Cameron, 1934: 79 (type examined); Cameron, 1937: 4 (Java); Shibata, 1973: 130 (Taiwan); Coiffait, 1982: 27 (characters; Nepal); Herman, 2001: 2444 (catalog); Smetana, 2004: 624 (Taiwan; Nepal); Zhou and Zhou, 2016: 20 (characters); Huang, Janák and Zhou, 2024c: 537 (type examined, Thailand; Myanmar; Malaysia).</p><p>Syn.:  Diochus indicus Kraatz, 1859: 113 (Type locality: India orientalis); Bernhauer and Schubert, 1914: 319 (synonym of  D. antennatus); Cameron, 1931: 360 (New Guinea); Cameron, 1932: 46 (synonym of  D. antennatus); Smetana, 2004: 624 (synonym of  D. antennatus); Löbl &amp; Löbl, 2015: 1007 (Palaearctic catalog; synonym of  D. antennatus).</p><p>Material examined.   MYANMAR: Shan Prov.: 1 male, 2 females,  Hsipaw, ca. 600 m, 22. 02. 96, litter, leg. S. Kurbatov (MHNG) ;   1 male: “ Birmanie, Helfer ”, “  Darjeeling,  Sikkim ”,  “indicus Kr. type ”,  “punctipennis Mots.,  indicus Kr. ”, “Ex-Typis” (IRSNB-coll. Fauvel). THAILAND: Chanthaburi :   8 males, 3 females,  Khao Sabap Nat. Park, 150–300 m, 23–24. XI. 1985, Burckhardt-Löbl (MHNG) ;   3 males, NE Bangkok, Khao Yai Nat. Park,  Khao Khieo, 1150 m, 28. XI. 1985, Burckhardt-Löbl (MHNG) ;   1 male, 2 females, Khlong Nakha Wildlife Sanctuary,  Kapoe Dist., 30 m, 29. 1. 1991, P. Schwendinger (MHNG) ;   1 male, 4 specimens, sex indet., Chiang Mai,  Doi Suthep National Park, Nov. 2019, Mok Fa waterfall, leaf litter, 08.11.2019, leg. Kleeberg (AKBC, JJRC) ;   2 males, 1 female, Khao Lak N.P.,  Thone Chong Fa Fall, 100–300 m, 6.-15. I. 1998, leg. A. Schultz &amp; K. Vock (SMNS, JJRC)  .  MALAYSIA: 1 male, 1 female, Penang, Dr. Cameron (NHML) .</p><p>Diagnosis. This species is externally similar to  D. lombokensis sp. nov.,  D. bogorensis sp. nov. and  D. punctipennis . This species has shorter antennae, and first segments of protarsi strongly dilated, distinguishing it from  D. punctipennis . Compared to  D. lombokensis sp. nov. and  D. bogorensis sp. nov., this species only has a shallow emargination on posterior margin of male sternite VIII, and the aedeagus of this species has parameres distinctly longer.</p><p>Distribution. Myanmar, Thailand, Malaysia. Published records from other regions of Indonesia (Sumatra, Java, Kalimantan [Borneo]), China (Taiwan) and Nepal have to be checked.</p></div>	https://treatment.plazi.org/id/03E02C60CF4FDE1FE3B12D7F4EC6EB86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang (2025): Revision of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) from Indonesia, Malaysia and Singapore. Zootaxa 5632 (3): 441-479, DOI: 10.11646/zootaxa.5632.3.2, URL: https://doi.org/10.11646/zootaxa.5632.3.2
03E02C60CF4CDE22E3B129E54F32EEB2.text	03E02C60CF4CDE22E3B129E54F32EEB2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus borneensis Cameron 1933	<div><p>10.  Diochus borneensis Cameron, 1933</p><p>(Fig 1C; Fig 10)</p><p>Diochus borneensis Cameron, 1933: 344 (Type locality:  British North Borneo: Kenokok); Hammond, 1984: 205 (checklist).</p><p>Type material examined.   MALAYSIA: SABAH: Holotype: female, “Type” (printed black, rounded label with red borders), “  B.N. BORNEO. Mt. Kinabalu, Kenokok,” (printed, light reddish label), “ 3.300 ft. 23 Apr. 1929 ” (printed and handwrited, light reddish label), “  D. borneensis Cam. TYPE” (black and red, Cameronʼs handwriting), “ M. Cameron. Bequest. B.M.1955-147” (NHMUK).</p><p>Additional material examined.   MALAYSIA: SABAH: 1 female,  Poring Hot Springs, 500 m, 8. V. 1987, Burckhardt-Löbl (MHNG)  .</p><p>Note. We examined the unique female type from Cameronʼs collection (NHMUK), which we considered as a holotype as only “one example” is mentioned in the description (Cameron 1933: 344).</p><p>Measurements. BL = 5.60 mm, FL = 2.98 mm, HL = 0.87 mm, HW = 0.65 mm, EyL = 0.22 mm, TL = 0.53 mm, ANL = 1.75 mm, ANT1 = 0.29 mm, ANT2 = 0.14 mm, ANT3 = 0.19 mm, ANT4 = 0.14 mm, ANT 11 = 0.17 mm, PL = 1.11 mm, PW = 0.88 mm, EL = 1.07 mm, EW = 1.18 mm, SL = 0.79 mm.</p><p>Redescription. Body long, narrowed anteriorly and posteriorly, large-sized. Body black or dark brown, posterior parts of abdominal segments lighter, reddish brown, intersegmental membrane reddish yellow. Legs yellowish brown with apical three quarters of tibiae at least partly blackish. Antennae brown with first 1–3 antennomeres partly and apex of each antennomere lighter, light reddish brown. Maxillary and labial palpi yellowish brown.</p><p>Head (Fig 10A, E). Very oblong in shape, 1.35 times as long as wide. Tempora regularly rounded, noticeably apically narrowed in basal third of head, posterior angles indistinct. Dorsal surface shiny, bearing very shallow and less distinct transverse microstriae, also with coarse and sparse punctures and scattered micropunctures. Head with about 7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of temple and near posterior margin. Each side of head with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye medium-sized, only slightly protruding laterad, longitudinal diameter shorter than half the length of tempora (eye: tempora = 0.42). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (0.20 mm) distinctly longer than distance from antenna to eye (0.08 mm).</p><p>Antennae (Fig 10B, F). Scape rod-shaped, slightly thickened apically, slightly shorter than two subsequent antennomeres combined; antennomere 2 slightly elongate, antennomere 3 distinctly longer than 2; length of antennomere 4 about twice of width, antennomere 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.</p><p>Mouthparts (Fig 10A, E). Labrum nearly hexagonal, lateral margin straight, widest at basal 1/3 to 1/2; width of anterior margin is about 3/5 the width at the widest point.</p><p>Neck (Fig 10A, E). Cylindrical, shiny, microsculpture indistinct, width 0.16 mm, shorter than 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.</p><p>Prothorax (Fig 10A, E). Pronotum distinctly elongate (PL to PW ratio 1.26), longer and wider than head.Anterior region near anterior angles deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, with scattered micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures composed of 3 large punctures (atypically with one additional puncture in left row in holotype) and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins).</p><p>Pterothorax (Fig 10A, E). Mesoscutellum triangular, surface shiny, bearing slightly transverse very fine mesh, but without any punctures.</p><p>Elytra (Fig 10A, E). Elytra slightly transverse (EL to EW ratio 0.91), about as long as but wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5–7 punctures along suture, two rows of 4–6 punctures in median, also with 3−4 rows of punctures on deflexed portion.</p><p>Legs (Fig 10G). First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi shorter than 2−4 together, that of metatibia only slightly longer than length of first segment.</p><p>Abdomen. Broadest at segment IV. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing fine transverse or polygonal mesh, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe.</p><p>Male. Unknown.</p><p>Female (Fig 10C, H−K). Head without elevation between eyes. Tube of spermatheca with three bends. Apical circle of the tube large in size but apical lobe small.</p><p>Diagnosis. The species can be easily distinguished by the large body, narrowed pronotum and noticeably apically narrowed and partly black tibiae.</p><p>Note. The female from Poring Hot Springs is externally very similar to the holotype, but differs by having the basal part of the tube of spermatheca more symmetrical and with less pronounced bends.</p><p>Distribution. Kalimantan (Borneo).</p></div>	https://treatment.plazi.org/id/03E02C60CF4CDE22E3B129E54F32EEB2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang (2025): Revision of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) from Indonesia, Malaysia and Singapore. Zootaxa 5632 (3): 441-479, DOI: 10.11646/zootaxa.5632.3.2, URL: https://doi.org/10.11646/zootaxa.5632.3.2
03E02C60CF72DE23E3B12A6D4992EC86.text	03E02C60CF72DE23E3B12A6D4992EC86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus pulchellus , Cameron 1918	<div><p>11.  Diochus pulchellus Cameron, 1918</p><p>(Figs 1K; 11-1; 11-2)</p><p>Diochus pulchellus Cameron, 1918: 84 (Type locality: Singapore: Sembawang); Cameron, 1921: 404 (Singapore); Scheerpeltz, 1933: 1324 (catalog); Herman, 2001: 2448 (catalog); Rougemont, 2001: 62 (Hongkong); Smetana, 2004 (Palaearctic catalog; Hongkong); Löbl &amp; Löbl, 2015: 1007 (Palaearctic catalog; Hongkong); Zhou &amp; Zhou, 2016: 26 (redescription, photo and line-drawing plates).</p><p>Material examined.   SINGAPORE: Holotype: male, “Holo-, Type” (printed and handwritten black, rounded label with red borders), “TYPE” (red, handwritten), “HOLOTYPE,  Diochus,  pulchellus, Cameron, 1918, det. R.G. Booth 2013”/ “Sembawang, Singapore., Dr. Cameron.” (black, printed), “Debris” (black, printed), “ M.Cameron., Bequest., B.M.1955–147.”, “HOLOTYPE,  Diochus pulchellus Cameron, 1918, det. R.G. Booth 2013” (NHMUK).</p><p>Note. The holotype is partly damaged and was dissected and partly disarticulated before the current study. The aedeagus is partly deformed and embedded in the middle (Fig 11-2G). Due to poor condition of the specimen we decided not to remount and clean again the specimen and the aedeagus.</p><p>Additional material examined.   INDONESIA: Panaitan Island, 2 males, 1 females:  Prinsen Eil., 20.i.1922, damm (SDEI, NHMUK) ;   Bali: 1 female, Sanur, at light, II.87,  Rougemont (JJRC) ;   Lombok: 1 female (elytra missing), Segaot, 13: VI: 1984,  Rougemont (GROG) ;   CHINA:  Hong Kong: 1 male, 23.XI.1996, Mai Po, Winkler extraction, G. de Rougemont (GROG) ;   1 female, #25 of Hong Kong,  Mai Po Nature Reserve, 21.vi.-5.vii.2014, leg.: C. Barthelemy, Malaise Trap (GROG)  .   LAOS: 1 male, Viang Chan prov.,  Ban Pa Kho resort, 50 km NE Vientiane, 90m, 9.-14.VI.2007, M. ŠTRBA leg.” (SMNS)  .</p><p>Diagnosis.  D. pulchellus is a small species with bicolored elytra and a complex spermatheca divided in two tubes: a shorter lateral tube and a main tube with many bends and basally connected with a sclerotized plate (Figs 11-2M–Q). These characters and also the shape of the aedeagus and missing or incomplete basal transverse carina of the male sternite VIII can be used to separate it easily from all other species.</p><p>Redescription. A detailed redescription of the species based on the holotype was published by Zhou &amp; Zhou (2016). As some details are not visible in the schematic pictures of the male sternite VIII (incomplete, medially interrupted basal transverse carina of male sternite VIII) and the aedeagus (paired middle sclerite) published by the authors, we made photographs of these characters of the holotype (Fig 11-1L–M, Fig 11-2G–L).</p><p>Variability.  D. pulchellus shows great variability in external characters such as shape of head and pronotum and also coloration of the forebody (e.g. color of the head varies from black to dark reddish brown and the head varies from slightly to markedly elongate: Figs 11-1A, D, N, O). The basal transverse carina of male sternite VIII is interrupted in the middle in the holotype (Fig. 11-2A–B), but completely missing in all other specimens (Figs 11- 1H; Figs 11-2C–F). Most of the specimens (including the holotype) possess a specific bilobate sclerite in aedeagus (Figs 11-2G, J–K; detail in Fig 11-2L). The bilobate sclerite is not always visible (Fig 11-1M; 11-2H), but it may be covered by other darker sclerites.</p><p>Distribution. Singapore; Indonesia (Painatan Island, Bali, Lombok); China (Hong Kong), Laos.</p></div>	https://treatment.plazi.org/id/03E02C60CF72DE23E3B12A6D4992EC86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang (2025): Revision of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) from Indonesia, Malaysia and Singapore. Zootaxa 5632 (3): 441-479, DOI: 10.11646/zootaxa.5632.3.2, URL: https://doi.org/10.11646/zootaxa.5632.3.2
03E02C60CF76DE26E3B129E54E29EE8A.text	03E02C60CF76DE26E3B129E54E29EE8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diochus punctipennis (Motschulsky 1858)	<div><p>12.  Diochus punctipennis (Motschulsky, 1858)</p><p>(Fig 1G)</p><p>Rhegmatocerus punctipennis Motschulsky, 1858: 659 (Type locality: Indes orientales).</p><p>Diochus punctipennis Fauvel, 1898: 117 (Seychelles); Kolbe, 1910: 20 (Seychelles); Bernhauer and Schubert, 1914: 319 (catalog); Bernhauer, 1922: 176 (Seychelles); Cameron, 1932: 46 (characters; India); Scheerpeltz, 1933: 1324 (catalog); Cameron, 1934: 79 (type examined); Cameron, 1937: 4 (Java); Scheerpeltz, 1957: 233 (Sumba); Coiffait, 1981: 329 (Andaman Islands); Lecoq, 1990: 189 (Mascarene Islands); Huang, Janák and Zhou, 2024c: 532 (type examined; Thailand; Malaysia).</p><p>Material examined.   MALAYSIA: 2 males, 1 female,  The Gap, Selangor, F. M. S., Dr. Cameron (NHMUK, JJRC)  .</p><p>Diagnosis. This species has aedeagal structures similar to  D. paradoxus sp. nov. and  D. paraparadoxus sp. nov., all with a round sclerite at the base of the aedeagus. This species can be distinguished from the two new species by the smaller size, and also the shape of male sternite VIII, which is rounded at posterior angles. Additionally, the aedeagus of this species has longer parameres than the other two species.</p><p>Distribution. Malaysia (Selangor), Thailand. Published records from other countries (Seychelles, Mauritius, Réunion, Rodriguez, Andaman Is., India) have to be verified.</p></div>	https://treatment.plazi.org/id/03E02C60CF76DE26E3B129E54E29EE8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Huang, Tian;Janák, Jiří;Zhou, Hong-Zhang	Huang, Tian, Janák, Jiří, Zhou, Hong-Zhang (2025): Revision of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) from Indonesia, Malaysia and Singapore. Zootaxa 5632 (3): 441-479, DOI: 10.11646/zootaxa.5632.3.2, URL: https://doi.org/10.11646/zootaxa.5632.3.2
