taxonID	type	description	language	source
03E087EFFF83FFEB7681F8990EB0F9FF.taxon	type_taxon	Type species. Malaymetopidea maculata Taszakowski & Kim, sp. nov. (Figs 1, 2).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF83FFEB7681F8990EB0F9FF.taxon	diagnosis	Diagnosis. Recognized by the following combination of characters: top of head distinctly lower than level of highest point of pronotum; vertex width subequal to single compound eye width; compound eye large in frontal view, occupying more than half of head height; fovea antennalis positioned directly below ventral margin of compound eye; pedicel cylindrical, thickest, subequal to head width; pronotum elongate, midline length more than 1 / 2 posterior maximal width, posterior margin of pronotum nearly straight medially, angled near posterolateral corners, leading midline longer than lateral margin length; scutellum flat and slightly tumid laterally, midline shorter than pronotal midline length.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF83FFEB7681F8990EB0F9FF.taxon	description	Description. Male. Body oval, length approximately 2.80 (Fig. 1 A, B). Surface and Vestiture. Body shiny, densely punctate, covered with long, pale, setae (Figs 1 A – C, 2 A – E); head punctate and wrinkled, frons and vertex with scattered setae, posterior margin with reclining setae; antennae covered densely with long, erect setae; pronotal collar with row of setae; pronotum, mesoscutum and scutellum distinctly and densely punctate, covered with setae; thoracic pleura deeply, unevenly punctate, and covered in places with scattered setae (Fig. 2 B, D); hemelytra covered densely with long setae, unevenly punctate, clavus deeply punctate, corium shallowly punctate, embolium and cuneus impunctate; membrane covered with microtrichia; legs with short, recumbent, pale setae; abdomen covered with brown setae. Structure. Head: flattened, hypognathous, wider than high, dorsally slightly protruding pronotal collar; posterior margin of vertex convex, protruding above eye level; compound eye large in frontal view, occupying more than half of head height; vertex wide, vertex width subequal to single compound eye width; ocelli large, convex, touching compound eyes (Fig. 1 A, C – E); fovea antennalis positioned directly below ventral margin of compound eye; antenna short, total antennal length subequal to maximal pronotal width; scape short, as thick as pedicel; pedicel cylindrical, subequal to head width, longer than combined lengths of flagellomeres; flagellomeres short and thin; basiflagellomere slightly longer than scape, almost as long as distiflagellomere; labium short, reaching mesocoxa. Thorax: pronotum trapeziform in broad sense, posterior margin broadly convex, nearly straight medially, angled near posterolateral corners, leading midline longer than lateral margin length, midline length more than 1 / 2 posterior width; pronotal collar thin; calli slightly marked, separated by fossa, disc convex, lateral margin distinctly carinate; mesoscutum short; scutellum slightly tumid, medium-sized, midline length shorter than pronotal length, flat; scent gland evaporative area triangular; hemelytron lateral margin slightly rounded; embolium wide; hypocostal lamina wide; cuneus broad, inner margin almost straight; membrane with two cells; mesofemora with three well-developed trichobothria; metafemora with four trichobothria (three well-developed and one reduced); tarsi two-segmented; claws with subapical tooth. Abdomen: elongate, not reaching apex of cuneus. Genitalia: pygophore with parameres similar in size; left paramere scythe-shaped, with long apical process and well-developed sensory lobe, apical process vertically curved, sensory lobe with few setae (Figs 1 D, 2 H); right paramere slightly smaller than left paramere, with knee-shaped sensory lobe, apical process short, broad, and apically blunted (Figs 1 E, 2 H); aedeagus delicate, endosoma sacciform and membranous, weakly sclerotized inside (Fig. 1 F).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF83FFEB7681F8990EB0F9FF.taxon	etymology	Etymology. The name combines “ Malay ” (the type locality, Malay Peninsula) with the suffix “ metopidea ”) which is part of the generic name Isometopidea. Gender feminine.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF83FFEB7681F8990EB0F9FF.taxon	discussion	Remarks. The affiliation of the newly described genus to Gigantometopini is evidenced by the elongate oval body; the presence of mesial fossa (incision) in the calli region; the claval commissure length shorter than scutellum length; similar length of parameres (at least right paramere longer than 1 / 2 left paramere); and scythe-shaped right paramere. Additionally, recently published morphology-based phylogenetic framework of the Isometopinae (Kim et al. 2025) recovered Isometopini as sister to Gigantometopini and revealed a trend wherein species with a relatively large compound-eye-to-head-height ratio tend to occupy earlier-diverging positions within Gigantometopini. Although Malaymetopidea gen. nov. was not included in that analysis, it shares key features with early-diverging members of the tribe (e. g., Kohnometopus Yasunaga, 2005, Isometopidea lieweni Poppius, 1913), including a large compound eye occupying more than half of the head height. Notably, it also resembles Isometopini in its relatively high eye / head ratio and trichobothria number. These features do not constitute direct phylogenetic evidence, but they are morphologically consistent with the pattern observed in our tree and may suggest that Malaymetopidea gen. nov. represents a lineage near the base of Gigantometopini, retaining ancestral traits shared with its sister tribe. While further data, particularly molecular evidence, are needed to confirm its exact phylogenetic position, the combination of Gigantometopinidefining features and Isometopini-like traits renders Malaymetopidea gen. nov. a taxon of potential significance for understanding the early divergence and character evolution within these two tribes. The new genus, Malaymetopidea gen. nov., does not resemble any of the previously described genera of Gigantometopini. The set of structural features mentioned in the diagnosis clearly distinguishes it from the other genera in the Gigantometopini.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF81FFEC7681F96D0EE0F983.taxon	diagnosis	Diagnosis. See the generic diagnosis.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF81FFEC7681F96D0EE0F983.taxon	description	Description. Coloration. Brown with yellowish macula (Fig. 1 A, B). Head: vertex brown; compound eyes silvery; frons, clypeus and gena dark brown; scape, pedicel and flagellomeres brown (Fig. 1 A – B); labium brownish. Thorax: pronotum mostly brown, anterior part dark brown, posterior one brown, lateral margin pale, posterior margin narrowly pale; exposed part of mesoscutum dark brown; scutellum brown, with apex and lateral angles yellowish; thoracic pleura brown; ostiolar peritreme and evaporative brown; hemelytra generally brown, clavus with pale spot in posterior half, endocorium with pale spot, embolium pale, brownish in distal part, semitransparent, hypocostal lamina whitish with brown spot in 1 / 3 length and brownish distal part; cuneus pale brown, with large whitish spot medially; membrane brownish; coxae pale, femora brown with pale apex, pro- and mesotibiae yellowish, metatibiae brown, lighter in the distal part, tarsi yellowish. Abdomen: dark brown. Surface and Vestiture. Body shiny, densely punctate, covered with pale, long setae; head punctate and wrinkled, frons and vertex with scattered setae, posterior margin with reclining setae; antennae covered densely with long, erect setae; pronotal collar with row of setae; pronotum, mesoscutum and scutellum distinctly and densely punctate, covered with setae; thoracic pleura deeply, unevenly punctate, and covered in places with scattered setae; hemelytra covered densely with long setae, unevenly punctate, clavus deeply punctate, corium shallowly punctate, embolium and cuneus impunctate; membrane covered with microtrichia; legs with short, adherent, pale setae; abdomen covered with brown setae. Structure. Body oval, length 2.82. Head: flattened, hypognathous, wider than high, head high 0.78 x as width across eyes, dorsally slightly protruding pronotal collar; posterior margin of vertex convex, protruding above eye level; compound eye 0.65 x as high as head high; vertex wide, 0.35 x as head width; ocelli large, convex, touching compound eyes; fovea antennalis positioned directly below ventral margin of compound eye; antenna short, total antennal length equal to maximal pronotal width; scape short, as thick as pedicel; pedicel cylindrical, subequal to head width, longer than combined lengths of flagellomeres; flagellomeres short and thin; basiflagellomere slightly longer than scape, almost as long as distiflagellomere; proportion of scape to distiflagellomere 0.15: 0.62: 0.18: 0.17; labium short, reaching mesocoxa. Thorax: pronotum trapeziform in broad sense, posterior margin broadly convex, nearly straight medially, angled near posterolateral corners, leading midline longer than lateral margin length, midline length 0.56 x as posterior width; pronotal collar thin, as thin as diameter of distiflagellomere; exposed part of mesoscutum short; scutellum slightly tumid, medium-sized, 0.68 x as long as pronotal length, flat; scent gland evaporative area triangular; lateral margin of hemelytra slightly rounded; outer cuneal margin 0.28 x as long as embolial margin. Abdomen: elongate, not reaching apex of cuneus. Genitalia: left paramere scythe-shaped with long apical process and well-developed sensory lobe, apical process vertically curved with depressed part medially, sensory lobe with few setae (Figs 1 D, 2 H); right paramere 0.79 x as long as left one, with knee-shaped sensory lobe, apical process short, broad (Figs 1 E, 2 H); endosoma sacciform and membranous (Fig. 1 F). Measurements. Body. Length, clypeus-apex of membrane: 2.82; width 1.25. Head. Length: 0.16; width, including compound eyes: 0.69; vertex width: 0.24; eye dorsal width: 0.23; height in frontal view: 0.54. Antenna. Total length: 1.12; length of scape: 0.15, pedicel: 0.62, flagellomere I: 0.18, flagellomere II: 0.17. Labium. Total length: 1.23; length of segment I: 0.33; II: 0.30; III: 0.26; IV: 0.37. Pronotum. Mesal length: 0.63; posterior maximal width (straight): 1.12. Mesoscutum. Mesal length: 0.11. Scutellum. Anterior width: 0.51; mesal length: 0.43. Hemelytron. Length: 2.08; claval commissure length: 0.35; outer embolial margin length (straight): 1.28; outer cuneal margin length (straight): 0.36; cuneus width: 0.36. Metatarsus. Total length: 0.27; length of tarsomeres I: II: 0.10: 0.25.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF81FFEC7681F96D0EE0F983.taxon	etymology	Etymology. From Late Latin maculâta (‘ stained, spotted’).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF81FFEC7681F96D0EE0F983.taxon	materials_examined	Type material. Holotype (♂): ’ MALAYSIA – Pahang \ Cameroon Highlands \ Tanah Rata cca 1600 m \ J. Horák lgt. 11 – 27 Febr. 2000 \ ’ (ZJC).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF81FFEC7681F96D0EE0F983.taxon	distribution	Distribution. Malaysia, Malay Peninsula (Fig. 3 L).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF81FFEC7681F96D0EE0F983.taxon	biology_ecology	Biology. Unknown.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF86FFED7681F9B10CC5FE5B.taxon	diagnosis	Diagnosis. Distinguished by the following combination of characters: dorsum and pleurites of thorax with deep and dense punctures; head vertical, flattened, not punctured but wrinkled and distinctly higher than wide, as high as pronotum height; vertex convex, protruding above eye level, width equal or slightly larger than eye width; compound eye elongate oval, tapered apically in front view, height subequal to 1 / 2 head height; pedicel subequal to or shorter than combined lengths of flagellomeres; labium reaching third abdominal segment; anterior margin of pronotum convex; punctures on scutellum weaker than those on pronotum; calli slightly marked, tarsi two segmented, claw without subapical tooth or with very small, barely noticeable apical tooth; abdomen short, not reaching to apex of cuneus; left paramere scythe-shaped, apical process tapered to apex, sensory lobe broadly developed; right paramere angled, apical process thick, sensory lobe knee-shaped, longer than 1 / 2 left paramere length; endosoma membranous with long sclerotized appendage and spinules (updated from Taszakowski et al. 2020).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF86FFED7681F9B10CC5FE5B.taxon	discussion	Remarks. Taszakowski et al. (2020) presented the genital structures of Planicapitus luteus. However, a mistake was made during the preparation of the manuscript, and the presented photos (fig. 5 A – E in Taszakowski et al. 2020), in fact, belong to an undescribed species of Sophianus Distant, 1904. Unfortunately, the genitalia of the P. luteus holotype are missing; however, we have found another specimen of this species, which allows us to describe it below (Fig. 5). Moreover, Taszakowski et al. (2020) stated that the type specimen of P. luteus is deposited in RBINS. The actual place of deposition of the specimen is ZMAN.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF84FFE07681F99B0EE0FB47.taxon	description	(Fig. 4)	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF84FFE07681F99B0EE0FB47.taxon	diagnosis	Diagnosis. Distinguished from P. luteus by the following combination of characters: The bottom half of the head brown, except for a pale spot in the middle (Fig 4 G); gena yellowish brown; pedicel shorter than combined lengths of flagellomeres; corium entirely dark brown; posterior part of cuneus brown; right paramere 0.77 x as long as left one, apical process forms an angle of 130 ° with the basal process.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF84FFE07681F99B0EE0FB47.taxon	description	Description. Male. Body oval, length 3.59 (Fig. 4 A, B). Coloration. Yellowish brown (Fig. 4 A, B, G). Head: vertex reddish; the area around the ocelli red; compound eyes pale; frons whitish, with two dark brown spots ventrally extending into large, V-shaped brown macula (Fig. 4 G); gena yellowish brown; scape and pedicel yellowish, basiflagellomere brown (Fig. 4 A, B); labium yellowish, apical part of fourth segment brown. Thorax: pronotum mostly brown, anterior and lateral parts yellowish brown, central and posterior parts brown; exposed part of mesoscutum pale brown; scutellum reddish brown, with apex whitish and lateral angles narrowly pale yellowish; thoracic pleura brown; ostiolar peritreme yellowish, evaporative area pale brown; hemelytra generally brown (Fig. 4 A, B); corium entirely brown; embolium pale brown, semitransparent; hypocostal lamina brown except yellowish anterior part; cuneus pale brown, with pale spot in basal inner corner; membrane brown; coxae pale, femora and tibiae yellowish brown with brown spots, tarsi yellowish. Abdomen: brown. Surface and Vestiture. Body shiny, unevenly punctate, covered with scattered, pale, long setae (Fig. 4 A, B, I); head impunctate but wrinkled, frons and vertex with tiny scattered setae, posterior margin with long, reclining setae (Fig. 4 H, I); antennae with erect setae; pronotal collar with row of setae; pronotum, distinctly and densely punctate, covered with setae, calli slightly punctate; mesoscutum punctate and almost glabrous; scutellum punctate in medial part, in lateral part impunctate but covered with setae; thoracic pleura deeply, unevenly punctate, and covered in places with scattered setae (Fig. 4 I); hemelytra shallowly and unevenly punctate, covered with setae; legs with short, adherent, pale-brown setae; abdomen covered with pale-brown setae. Structure. Head: flattened, hypognathous, distinctly higher than width across eyes, dorsally extending to level of highest point of pronotum; vertex convex, protruding above eye level; compound eyes 0.5 x as high as head high; vertex wide, 0.45 x as head width; ocelli small, spaced apart, touching compound eyes (Fig. 4 G – I); fovea antennalis positioned at middle of the ventral margin of compound eye and apex of clypeus; scape short, as thick as pedicel; pedicel slightly arcuate, flagellomeres linear, thin; proportion of scape to distiflagellomere 0.12: 0.64: 0.80: missing; labium reaching third abdominal segment. Thorax: pronotum trapeziform, midline length almost equal to 1 / 2 posterior width; pronotal collar thin; calli slightly marked, separated by shallow fossa, disc relatively flat, lateral margin carinate, posterior margin weakly sinuate with a small backwards-pointing process in the middle, posterior angles weakly angled; mesoscutum long; scutellum tumid, large, 0.8 x as long as pronotal length, baso-medially clearly depressed; scent gland evaporative area triangular; ostiolar peritreme small, strongly swollen, directed upward and anteriorly; hemelytron lateral margin slightly rounded; hypocostal lamina wide, cuneus broad, inner margin straight; membrane with two cells; tarsi two-segmented; claws without a subapical tooth. Abdomen: elongate, not reaching apex of cuneus. Genitalia: aedeagus delicate, endosoma sacciform and membranous, weakly sclerotized inside, outer subapical and apical part more sclerotic, clothed with dense spinules (Fig. 4 C, D); left paramere scythe-shaped, with long apical process and slightly developed sensory lobe, apical process vertically curved, sensory lobe with few setae (Fig. 4 E); right paramere 0.77 x as long as left one, with knee-shaped sensory lobe, apical process long, slightly curved at apex; apical process forms an angle of 130 ° with the basal process (Fig. 4 F). Measurements. Body. Length, clypeus-apex of membrane: 2.59; width 1.20. Head. Length: 0.24; width, including compound eyes: 0.56; vertex width: 0.25; eye dorsal width: 0.16; height in frontal view: 0.81. Antenna. Length of scape: 0.12, pedicel: 0.64, flagellomere I: 0.80, flagellomere II: –. Labium. Total length: 1.18; length of segment I: 0.30; II: 0.32; III: 0.25; IV: 0.37. Pronotum. Mesal length: 0.54; posterior maximal width (straight): 1.13. Mesoscutum. Mesal length: 0.13. Scutellum. Anterior width: 0.53; mesal length: 0.43. Hemelytron. Length: 1.98; claval commissure length: 0.27; outer embolial margin length (straight): 1.23; outer cuneal margin length (straight): 0.24; cuneus width: 0.35. Metatarsus. Total length: 0.23; length of tarsomeres I: II: 0.09: 0.18.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF84FFE07681F99B0EE0FB47.taxon	etymology	Etymology. From Late Latin brűnus (‘ brown’).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF84FFE07681F99B0EE0FB47.taxon	materials_examined	Type material. Holotype (♂): ’ THAILAND, pr. Trang \ Ton Pan waterfall \ W of Ban Nawong \\ 26. XI. 2003., No. 27 \ leg. A. Orosz et \ Gy. Sziráki’ (HNHM).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF84FFE07681F99B0EE0FB47.taxon	distribution	Distribution. South Thailand, Malay Peninsula (Fig. 3 J).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF84FFE07681F99B0EE0FB47.taxon	biology_ecology	Biology. Unknown.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8AFFE07681FAF50EE0F858.taxon	diagnosis	Diagnosis. Distinguished from P. brunneus by the following combination of characters: bottom half of the head large Y-shaped brown macula; gena whitish yellow; pedicel subequal to than combined lengths of flagellomeres; corium brown with a pale spot medially; embolium and posterior part of cuneus yellowish, right paramere 0.56 x as long as left one; apical process forms an angle of 110 ° with the basal process.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8AFFE07681FAF50EE0F858.taxon	description	Description. See the original description in Taszakowski et al. (2020). Male genitalia: genital capsule impunctate, covered with long setae; aedeagus delicate; endosoma sacciform and membranous, weakly sclerotized inside, outer subapical and apical part more sclerotic, clothed with dense spinules (Fig. 5 B); left paramere scythe-shaped, apical process long, vertically curved, slightly widened in distal part, sensory lobe slightly developed, with few setae (Fig. 5 C); right paramere 0.56 x as long as left one, with knee-shaped sensory lobe, apical process long, slightly curved at apex; apical process forms an angle of 110 ° with the basal process (Fig. 5 D).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8AFFE07681FAF50EE0F858.taxon	materials_examined	Material examined. Holotype (♂): ‘ Borneo, Sabah / Danum Valley / 70 km W Lahad Datu / M. J. & J. P. Duffels // East Ridge Trail / 150 m / 2. XII. 1989 // sample Sab. 53 / understory rainforest, / at light’ (ZMAN); ♂: ‘ BRUNEI: Ulu / Temburong ridge / ii. – iii. 1982 // M. C. Day / B. M. 1983 – 75. ’ (NHM).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8AFFE07681FAF50EE0F858.taxon	distribution	Distribution. Brunei, Malaysia (North Borneo) (Fig. 3 N, P).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8AFFE07681FAF50EE0F858.taxon	biology_ecology	Biology. Unknown.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8BFFE17681FBF80EB6F929.taxon	materials_examined	Material examined: ♀ ‘ INDONESIA: \ SULAWESI UTARA. \ Dumoga-Bone N. P. \ February 1985. \\ Fog 5 \ 400 m. 11. ii. 85 \ BMNH Plot C \\ R. Ent. Soc Lond. \ PROJECT WALLACE \ B. M. 1985 - 10 \\ TRAY \ 94 ’ (NHM); ♂ ‘ 8. viii – i. x. 85 \ INDONESIA: \ SULAWESI UTARA, \ Dumoga-Bone N. P. \\ Malaise trap \\ ‘ Edwards’ Camp \ Lowland forest \ 664 m. ’ (NHM); ♂ ‘ INDONESIA: \ SULAWESI UTARA, \ Dumoga-Bone N. P. \ mid- 20 June 1985 \\ Malaise trap \\ Lowland forest \ edge ca 200 m’ (NHM); ♂ ‘ INDONESIA: \ SULAWESI UTARA, \ Dumoga-Bone N. P. \ 9 - 16 May 1985. \\ Malaise trap \\ Toraunt. \ Edge of forest \ 24 Trap days’ (NHM); ♂ ‘ INDONESIA: \ SULAWESI UTARA, \ Dumoga-Bone N. P. \ 21 - 26 July 1985. \\ Malaise trap \\ ‘ Edwards’ Camp \ Lowland forest \ 664 m. ’ (NHM); ♂ ‘ INDONESIA: \ SULAWESI UTARA, \ Dumoga-Bone N. P. \ 6 - 21 July 1985. \\ Malaise trap \\ ‘ Edwards’ Camp \ Lowland forest \ 664 m. ’ (NHM); ♂ ‘ ’’ Edwards’ ’ 680 m \ 26. IV – 7. VI. 1985 \ Toraunt. J. M. Martin \\ Malaise trap \ up tree’ (NHM).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8BFFE17681FBF80EB6F929.taxon	distribution	Distribution. This species is known from Bogani Nani Wartabone National Park (formerly known as Dumoga Bone National Park) (Fig. 3 R).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8BFFE27681F9170B93F817.taxon	type_taxon	Type species. Utarametopus linuspitti Taszakowski & Kim, sp. nov. (Figs. 6, 7).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8BFFE27681F9170B93F817.taxon	diagnosis	Diagnosis. Recognized by the following combination of characters: dorsum densely and deeply punctuate, with uniformly distributed pale, shiny, semierect long setae; head vertically elongate, flattened in front, slightly higher than or as high as pronotal height; compound eye relatively small, height less than 1 / 2 head height; lateral margin of frons under compound eye U-shaped; scape as thick as pedicel in distal part; pedicel slightly arcuate and clavate; pronotum somewhat elongate in dorsal view, moderate in thickness in lateral view, height as long as length, posterior margin weakly concave on either side, with a median projection; calli weakly swollen; scutellum partially tumid, impunctate, except baso-medial part clearly depressed with punctures; exposed part of mesoscutum punctate (Figs 7 E, 8 E); embolium and cuneus impunctate.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8BFFE27681F9170B93F817.taxon	description	Description. Female. Body oval, length approximately 2.60 – 3.40 (Figs 6 A, 8 A, B). Surface and Vestiture. Body shiny, densely and deeply punctate; except glabrous head, covered densely with pale, shiny, setae (Figs 6 A – C, 8 A – C); along costal vein row of punctures, embolium and cuneus impunctate, antennae with sparse setae; pronotal collar with row of setae; scutellum punctate only in the midline (Figs 6 E, 8 E); legs with short, recumbent, pale-brown setae. Structure. Head: flattened, hypognathous, distinctly higher than width across eyes, dorsally almost reaching or extending level of the highest point of pronotum; vertex flat, at same level or slightly protruding above eyes level; compound eyes height shorter than 1 / 2 head height; vertex wide, subequal to single compound eye width; ocelli small to medium-sized, spaced apart, touching compound eyes (Figs 6 B – D, 8 A, C); fovea antennalis positioned at middle of ventral margin of compound eye and apex of clypeus; scape short, as thick as pedicel in distal part; pedicel slightly arcuate and clavate; flagellomeres linear, thin; labium long, reaching fourth to five abdominal segment. Thorax: pronotum trapeziform, midline length about 1 / 2 posterior width, lateral margin carinate, posterior margin weakly concave on either side, with median projection, posterior angles weakly angled; pronotal collar broad; calli slightly marked, disc slightly convex; scutellum partially tumid, relatively large, midline length shorter than pronotal length, width subequal to 1 / 2 pronotal maximum width, baso-medially clearly depressed (Fig. 6 E); scent gland evaporative area triangular; ostiolar peritreme swollen, directed upward and anteriorly; hemelytron lateral margin rounded; cuneus broad, inner margin almost straight; membrane with two cells. Abdomen: elongate, reaching or exceeding apex of cuneus.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8BFFE27681F9170B93F817.taxon	etymology	Etymology. The name combines “ Utara ” (the type locality, province Sulawesi Utara) with a suffix “ metopus ” which is part of the generic name Isometopus, the type genus of the subfamily. Gender masculine.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8BFFE27681F9170B93F817.taxon	discussion	Remarks. The placement of the newly described genus in Gigantometopini is also based on the following characters: a body elongate oval; the presence of a mesial fossa (incision) in the calli region; and the claval commissure length shorter than the scutellum length, although only female individuals are known so far. Within Gigantometopini, the new genus, Utarametopus gen. nov. can be easily distinguished from the genera Isometopidea Poppius, 1913 and Kohnometopus Yasunaga, 2005 by the fovea antennalis clearly separated from compound eye (vs. placed near compound eye); from the genera Bruneimetopus Taszakowski et al., 2020, Planicapitus Taszakowski et al., 2020, and Sulawesimetopus Herczek et al., 2018 by the small compound eye relative to head height, compound eye height shorter than 1 / 2 head height (vs. subequal or longer than 1 / 2 head height); from Gigantometopus Schwartz & Schuh, 1990 and Megalofaciatus Taszakowski et al., 2021 by weakly developed pronotal calli (vs. strongly developed calli region); and Astroscopometopus Yasunaga & Hayashi, 2002 by partially tumid scutellum (vs. strongly tumid and projected scutellum). Additionally, this new genus can be separated from the genera newly described in this study based on the diagnostic characters. This new genus is currently considered to be most similar and closely related to Sulawesimetopus, among the genera belonging to Gigantometopini, due to overall similarities in morphological features and coloration patterns. Notably, sexual dimorphism has been confirmed in Sulawesimetopus (although this was not mentioned in the original description, it has been verified through our additional examination of specimens), and the female exhibits head structures that resemble those of the present genus, which is described here based solely on female specimens. Nevertheless, the two genera can be structurally distinguished by the following morphological characters: U-shaped outer margin of head under compound eye in front view (Figs 6 C, 7 C, 8 C) (vs. V-shaped outer margin; see fig 2 A in Herczek et al. (2017 )); distinctly small compound eye relative to head height (vs. compound eye height more than 1 / 2 head height in male and subequal to 1 / 2 head height in female); pronotum relatively moderate in thickness in lateral view, length subequal to height (Figs 6 B, 7 A) (vs. distinctly thick in lateral view, length shorter than height; fig. 1 C in Herczek et al. (2017 )); pronotum elongate with a sinuate posterior margin (Figs 6 A, 8 A) (vs. distinctly wide with a rounded posterior margin; fig. 1 A, B in Herczek et al. (2017 )); scutellum moderately tumid, with a narrow longitudinal median depression, punctures restricted to the depressed area only (Figs 6 A, 7 E, 8 A, E) (vs. scutellum strongly tumid, entirely punctate; fig. 1 in Herczek et al. (2017 )); and embolium and cuneus impunctate (Figs 6 A, 8 A) (vs. embolium and cuneus distinctly punctate; fig. 1 A, B in Herczek et al. (2017 )).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF89FFE57681FF580EE0FD07.taxon	description	(Fig. 6, 7)	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF89FFE57681FF580EE0FD07.taxon	diagnosis	Diagnosis. Distinguished from U. striatus by the following combination of characters: body medium-sized, 3.4 in length, ocelli medium-sized, diameter 0.25 x as vertex width, pronotum length 0.9 x as basiflagellomere length; body mostly dark brown; flagellomeres dark, embolium in basal part dark, cuneus with small, dark spot.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF89FFE57681FF580EE0FD07.taxon	description	Description. Female. Body oval, length 3.37 (Fig. 6 A). Coloration. Mostly dark brown (Fig. 6 A – C). Head: vertex reddish brown; compound eyes reddish brown; frons and clypeus dark brown (Fig. 6 C); maxillary plate greyish brown; gena chestnut brown, posterior margin whitish; scape and pedicel yellowish, flagellomeres brown (Fig. 6 A, B); labium brown except for yellowish apical part of second segment and basal 2 / 3 of fourth one (Fig. 6 B). Thorax: pronotum dark brown, lateral margins pale, semitransparent, posterior margin pale; exposed part of mesoscutum dark brown; scutellum brown, with pale apex; thoracic pleura mostly dark brown; ostiolar peritreme whitish, evaporative area dark brown; hemelytra bicolored, semitransparent (Fig. 6 A); clavus brown with greyish stripe along claval vein; endocorium brown with large greyish spot and greyish stripe along R + M vein; embolium pale; hypocostal lamina pale with brown spot in anterior part; cuneus transparent, with brown spot in middle; membrane pale brown; legs distinctly bicolored; coxa partially pale and brown; profemora unavailable; mosofemora pale in basal half, brown in distal one; metafemora pale with two brown stripes: broad antero-dorsal and narrow posterior; metatibiae yellowish except basal 1 / 3 dark and apex brown. Abdomen: dark brown. Surface and Vestiture. Body shiny, densely and deeply punctate; except glabrous head, covered densely with pale setae (Fig. 6 A – C); along costal vein row of punctures, embolium and cuneus impunctate, antennae with sparse setae; pronotal collar with row of setae; scutellum punctate only in the midline (Fig. 6 E); legs with short, adherent, pale brown setae. Structure. Head: flattened, hypognathous, distinctly higher than width across eyes, dorsally extending level of the highest point of pronotum; vertex flat, slightly protruding above eyes level; compound eyes 0.4 x as high as head high; vertex wide, 0.39 x as head width; ocelli medium-sized, diameter 0.25 x as vertex width, spaced apart, touching compound eyes (Fig. 6 B – D); fovea antennalis positioned at middle of the ventral margin of compound eye and apex of clypeus; scape 0.5 x as long as distiflagellomere, thick; pedicel slightly arcuate, thinner in basal part, slightly thickened in distal one; flagellomeres linear, thin; labium long, reaching fourth abdominal segment. Thorax: pronotum trapeziform, midline length about 1 / 2 posterior width; pronotal collar broad; calli slightly marked, disc slightly convex, lateral margin carinate, posterior margin weakly sinuate, posterior angles weakly angled; exposed part of mesoscutum short; scutellum tumid, large, 0.67 x as long as pronotal length, baso-medially clearly depressed (Fig. 6 E); scent gland evaporative area triangular; ostiolar peritreme swollen, directed upward and anteriorly; hemelytron lateral margin rounded; cuneus broad, inner margin almost straight; membrane with two cells; mesofemora with five trichobothria (four well-developed and one reduced); metafemora with six trichobothria. Abdomen: elongate, reaching apex of cuneus. Measurements. Body. Length, clypeus-apex of membrane: 3.37; width 1.46. Head. Length: 0.34; width, including compound eyes: 0.54; vertex width: 0.21; eye dorsal width: 0.20; height in frontal view: 0.93. Antenna. Total length: 1.79; length of scape: 0.12, pedicel: 0.63, flagellomere I: 0.83, flagellomere II: 0.21. Labium. Total length: 1.51; length of segment I: 0.32; II: 0.38; III: 0.37; IV: 0.47. Pronotum. Mesal length: 0.73; posterior maximal width (straight): 1.34. Mesoscutum. Mesal length: 0.11. Scutellum. Anterior width: 0.58; mesal length: 0.49. Hemelytron. Claval commissure length: 0.44; heme length: 2.45; outer embolial margin length (straight): 1.49; outer cuneal margin length (straight): 0.38; cuneus width: 0.44.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF89FFE57681FF580EE0FD07.taxon	etymology	Etymology. This species is named after Linus Pitt, a fictional lecturer in natural history from Blood of Elves (the first novel in The Witcher series by Andrzej Sapkowski), of which the first author is a big fan.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF89FFE57681FF580EE0FD07.taxon	materials_examined	Type material. Holotype (♀) ‘ INDONESIA: \ SULAWESI UTARA. \ Dumoga-Bone N. P. \ March 1985. \\ Insecticide \ Fogging: 9 \ KOPA \\ R. Ent. Soc. Lond. \ PROJECT WALLACE \ B. M. 1985 - 10 \\ TRAY \ 32 ’ (NHM).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF89FFE57681FF580EE0FD07.taxon	distribution	Distribution. Indonesia, North Sulawesi, Bogani Nani Wartabone National Park (formerly known as Dumoga Bone National Park) (Fig. 3 R).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF89FFE57681FF580EE0FD07.taxon	biology_ecology	Biology. Unknown.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8FFFE77681FD340B94FECB.taxon	description	(Fig. 8.)	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8FFFE77681FD340B94FECB.taxon	diagnosis	Diagnosis. Distinguished from U. linuspitti by the following combination of characters: body small, 2.65 in length, ocelli small, diameter 0.18 x as vertex width, pronotum length 0.75 x as basiflagellomere length; body mostly brown, flagellomeres pale brown, hemelytra with distinct whitish stripes; embolium in basal part pale brown, cuneus with a blurry, indistinct brown spot.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8FFFE77681FD340B94FECB.taxon	description	Description. Female. Body oval, length 2.65 (Fig. 8 A, B). Coloration. Shades of brown with dark punctation (Fig. 8 A – C). Head: vertex reddish; compound eyes pale reddish; frons between compound eyes pale reddish brown, below the eyes chestnut brown; clypeus chestnut brown (Fig. 8 C); gena chestnut brown, posterior margin pale; scape and pedicel yellowish, flagellomeres pale brown (Fig. 8 A, B); labium brown except for yellowish apical part of second segment and basal 2 / 3 of fourth one (Fig. 8 B). Thorax: pronotum mostly brown, lateral margins pale, semitransparent, posterior margin pale; pronotal collar brown, callosite region chestnut brown, disc brown; exposed part of mesoscutum chestnut brown; scutellum brown, with whitish apex; thoracic pleura chestnut brown; ostiolar peritreme whitish, evaporative area whitish in antero-ventral part, brown in central and pale brown in posterior part; hemelytra bicolored (Fig. 8 A); clavus brown with whitish stripe along claval vein; endocorium brown with large whitish spot and whitish stripe along R + M vein; embolium whitish; hypocostal lamina pale with brown spot in anterior part; cuneus brown, basal inner corner and lateral margin whitish; membrane brown; coxae pale; metacoxa tinged with red; profemora yellowish, tinged with red; meso- and metafemora missing; protibiae brown with two whitish bands; protarsi brownish. Abdomen: reddish pale brown. Surface and Vestiture. Body shiny, densely and deeply punctate; except glabrous head, covered densely with, pale setae (Fig. 8 A – C); along R + M and costal veins row of punctures, embolium and cuneus impunctate, antennae with sparse, erect setae; pronotal collar with row of setae; scutellum punctate only in the midline (Fig. 8 E); legs with short, adherent, pale-brown setae. Structure. Head: flattened, hypognathous, distinctly higher than width across eyes, dorsally not extending to level of highest point of pronotum; vertex flat, nearly at same level as eyes level; compound eyes 0.45 x as high as head high; vertex wide, 0.37 x as head width; ocelli small, spaced apart, touching compound eyes (Fig. 8 A, C); fovea antennalis positioned at middle of the ventral margin of compound eye and apex of clypeus; scape 0.5 x as long as distiflagellomere, thick; pedicel slightly arcuate, thinner in basal part, in distal part as thick as scape; flagellomeres linear, thin; labium long, reaching fifth abdominal segment. Thorax: pronotum trapeziform, midline length equal to 1 / 2 posterior width; lateral margin carinate, posterior margin weakly sinuate with a small backwards-pointing process in middle, posterior angles weakly angled; pronotal collar broad; calli slightly marked, disc slightly convex; exposed part of mesoscutum long; scutellum tumid, large, 0.73 x as long as pronotal length, baso-medially clearly depressed; scent gland evaporative area triangular; ostiolar peritreme small, swollen, directed upward and anteriorly; hemelytron lateral margin slightly rounded; cuneus broad, inner margin straight; membrane with two cells; tarsi three-segmented; claws without a subapical tooth. Abdomen: elongate, exceeding apex of cuneus. Measurements. Body. Length, clypeus-apex of membrane: 2.65; width 1.15. Head. Length: 0.21; width, including compound eyes: 0.51; vertex width: 0.19; eye dorsal width: 0.19; height in frontal view: 0.80. Antenna. Total length: 1.57; length of scape: 0.10, pedicel: 0.54, flagellomere I: 0.74, flagellomere II: 0.19. Labium. Total length: 1.43; length of segment I: 0.36; II: 0.38; III: 0.34; IV: 0.41. Pronotum. Mesal length: 0.55; posterior maximal width (straight): 1.10. Mesoscutum. Mesal length: 0.17. Scutellum. Anterior width: 0.50; mesal length: 0.40. Hemelytron. Length: 1.90; claval commissure length: 0.37; outer embolial margin length (straight): 1.30; outer cuneal margin length (straight): 0.31; cuneus width: 0.33.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8FFFE77681FD340B94FECB.taxon	etymology	Etymology. From Latin striâtus (‘ grooved, fluted, striated’).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8FFFE77681FD340B94FECB.taxon	materials_examined	Type material. Holotype (♀): ’ INDONESIA: \ SULAWESI UTARA, \ Dumoga-Bone N. P. \ March 1985 \\ Plot A. M, T. (raised) \ 20 - 27 / iii. 85 ’ (NHM).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8FFFE77681FD340B94FECB.taxon	distribution	Distribution. Indonesia, North Sulawesi, Bogani Nani Wartabone National Park (formerly known as Dumoga Bone National Park). (Fig. 3 R).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8FFFE77681FD340B94FECB.taxon	biology_ecology	Biology. Unknown.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8FFFE77681FD340B94FECB.taxon	discussion	Remarks. Both species described above are similar to each other. Apart from the fact that they have a clear difference in body size (3.37 vs 2.65), they differ in the size of the ocelli, the length and width of the pronotum and the coloration. The sum of these features led us to describe each individual as a separate species.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8DFFF97681FE790D71FE5B.taxon	type_taxon	Type species. Wallacemetopus anteriotenebrosus Taszakowski & Kim sp. nov.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8DFFF97681FE790D71FE5B.taxon	diagnosis	Diagnosis. Recognized by the following combination of characters: dorsum densely covered with long golden hairs; head inversely triangular in front view; vertex wide, protruding above eye level; compound eye height shorter than 1 / 2 head height; ocelli large; pedicel slightly arcuate and clavate, clearly shorter than combined lengths of flagellomeres; calli well developed, clearly punctate, covered long setae; scutellum elongate, midlength subequal to pronotum midline length, strongly tumid and projected, medially depressed; protarsi three-segmented; metatarsi two-segmented.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8DFFF97681FE790D71FE5B.taxon	description	Description. Male. Body oval, length approximately 2.20 (Fig. 9 A – C). Surface and Vestiture. Body shiny, unevenly punctate, covered with pale, long setae (Fig. 9 A – C, 10 A, B, F); head deeply punctate, somewhat wrinkled, frons and vertex with tiny scattered setae, posterior margin with few long, reclining setae (Fig. 10 C); antennae with erect setae; pronotal collar with row of setae; pronotum, calli region, and mesoscutum distinctly and densely punctate, covered with setae; scutellum unevenly punctate, covered with scattered setae; thoracic pleura deeply punctate, and covered unevenly with scattered setae (Fig. 10 B, C, E); hemelytra unevenly punctate, covered with setae; clavus and corium strongly punctate; embolium impunctate; legs with short, pale-brown setae; abdomen covered with pale setae. Structure. Head: flattened, hypognathous, inversely triangular in front view, distinctly higher than width across eyes, dorsally equal to calli level; vertex wide, flat, subequal to single compound eye width, protruding above eye level; compound eyes somewhat small, height shorter than 1 / 2 head height; ocelli large, touching compound eyes (Fig. 10 A, D, E); fovea antennalis positioned at middle of the ventral margin of compound eye and apex of clypeus (Fig. 10 B – D); scape short, as thick as pedicel in distal part; pedicel slightly arcuate and clavate, clearly shorter than combined lengths of flagellomeres; flagellomeres linear, thin; labium reaching abdomen. Thorax: pronotum trapeziform, midline length longer than 1 / 2 posterior width; pronotal collar relatively broad, diameter subequal to diameter of basal part of basiflagellomere; calli well developed, separated by distinct fossa, disc convex, lateral margin carinate, posterior margin sinuate with a small backwards-pointing process in the middle; exposed part of mesoscutum long; scutellum elongate, length slightly shorter than pronotal length, width subequal to 1 / 2 pronotal maximum width, tumid with two projections, medially depressed (Fig. 10 F); scent gland evaporative area triangular; ostiolar peritreme small, swollen, directed upward and anteriorly; hemelytron lateral margin slightly rounded; embolium wide; hypocostal lamina wide; cuneus broad, inner margin straight; membrane with two cells; protarsi three-segmented; metatarsi two-segmented (mesotarsi damaged); claws without a subapical tooth. Abdomen: elongate, reaching the apex of cuneus. Genitalia: aedeagus delicate, endosoma sacciform and membranous, weakly sclerotized inside, outer subapical and apical part more sclerotic, clothed with dense spinules (Fig. 9 E); left paramere scythe-shaped, sensory lobe slightly developed (Fig. 9 F), apical process missing; right paramere with knee-shaped sensory lobe, apical process short, slightly curved at apex (Fig. 9 G).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8DFFF97681FE790D71FE5B.taxon	etymology	Etymology. The proposed generic name is a compound of two words, the first being a patronym honoring Alfred Russel Wallace for his pioneering work on biogeography (especially the Malay Archipelago), with the suffix “ metopus ” which is part of the generic name Isometopus, the type genus of the subfamily. Gender masculine.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF8DFFF97681FE790D71FE5B.taxon	discussion	Remarks. The affiliation of Wallacemetopus gen. nov. as a member of Gigantometopini is evident by an elongate oval body; the presence of mesial fossa (incision) in calli region; the claval commissure length shorter than scutellum length; the similar length of parameres (at least right paramere longer than 1 / 2 left paramere); and the scythe-shaped right paramere. Although the apical part of the left paramere was missing, the remaining portion is intact, and the right paramere is fully preserved. The genital capsule, particularly the parameres, had been glued to the wings of the second pair and slightly crushed, which resulted in damage to the left paramere during preparation. While the structure of the parameres is highly conserved across most genera of Gigantometopini, the external morphology is not only diagnostically distinct at the generic level but also remarkably stable at the species level. Given that the Wallacemetopus gen. nov. exhibits a consistent set of external features comparable to those found in other genera of the tribe, its placement within Gigantometopini appears well supported based on the available morphological evidence. The new genus, Wallacemetopus gen. nov., exhibits a set of structural features that are uncommon even within Gigantometopini. In particular, the calli region is distinctly developed, resembling that of Gigantometopus Schwartz & Schuh, 1990 or Megalofaciatus Taszakowski et al., 2021. The strongly tumid and projected scutellum also shows similarity to Megalofaciatus, and based on these shared characteristics, Wallacemetopus gen. nov. is considered to be most closely related to this genus.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF93FFFB7681FD890EE0FEEE.taxon	diagnosis	Diagnosis See the generic diagnosis.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF93FFFB7681FD890EE0FEEE.taxon	description	Description. Coloration. Yellowish and brown. Head: vertex reddish; compound eyes silverish; frons and gena above fovea antennalis dark brown; gena below fovea antennalis, mandibular and maxillary plates and clypeus brown; scape pale brown, pedicel and flagellomeres yellowish; labium yellowish, apical part of fourth segment brownish. Thorax: anterior part of pronotum dark brown, posterior part pale brownish with ivory markings in posterior corners; exposed part of mesoscutum dark brown; scutellum laterally reddish brown, medial part and apex pale; thoracic pleura mostly brown; propleuron dark brown except yellowish posterior upper corner and dorsal margin, mesepisternum and mesepimeron dark brown, metepisternum brown; ostiolar peritreme pale yellowish, evaporative area yellowish brown; hemelytra generally yellowish, semitransparent, hypocostal lamina pale, except brown spot in anterior part; membrane pale; legs entirely pale yellowish. Abdomen: pale brown, tinged with red dorsally. Body oval, length 2.25. Surface and Vestiture. Body shiny, somewhat metallic, unevenly punctate, covered with pale, long setae; head deeply punctate, somewhat wrinkled, frons and vertex with tiny scattered setae, posterior margin with few long, reclining setae; antennae with erect setae; pronotal collar with row of setae; pronotum and mesoscutum distinctly and densely punctate, covered with setae; scutellum unevenly punctate, covered with scattered setae; thoracic pleura deeply punctate, and covered unevenly with scattered setae; hemelytra unevenly punctate, covered with setae; legs with short, pale-brown setae; abdomen covered with pale setae. Structure. Head: flattened, hypognathous, distinctly higher than width across eyes, dorsally equal to calli level; vertex flat, protruding above eye level; compound eyes 0.4 x as high as head high; vertex wide, 0.37 x as head width; ocelli large, touching compound eyes; fovea antennalis positioned at middle of the ventral margin of compound eye and apex of clypeus; scape short, as thick as pedicel in distal part; pedicel slightly arcuate; flagellomeres linear; basiflagellomere longer than distiflagellomere; proportion of scape to distiflagellomere 0.12: 0.48: 0.67: 0.23; labium reaching second abdominal segment. Thorax: pronotum trapeziform, midline length 0.55 x as posterior width; pronotal collar relatively broad; calli well developed, separated by distinct fossa, disc convex, lateral margin carinate, posterior margin sinuate with a small backwards-pointing process in the middle; exposed part of mesoscutum long; scutellum large, 0.75 x as long as pronotal length, with two projections; scent gland evaporative area triangular; ostiolar peritreme small, swollen, directed upward and anteriorly; hemelytron lateral margin slightly rounded; embolium broad, broader posteriorly; hypocostal lamina wide; cuneus broad, inner margin straight; membrane with two cells; protarsi three-segmented; metatarsi two-segmented; claws without subapical tooth. Abdomen: elongate, reaching the apex of cuneus. Genitalia: aedeagus delicate, endosoma sacciform and membranous, weakly sclerotized inside, outer subapical and apical part more sclerotic, clothed with dense spinules (Fig. 9 E); left paramere scythe-shaped, sensory lobe narrow, slightly developed (Fig. 9 F); right paramere with knee-shaped sensory lobe, apical process short, as thick as basal process, slightly curved at apex, apex sharp (Fig. 9 G). Measurements. Body. Length, clypeus-apex of membrane: 2.25; width 0.99. Head. Length: 0.24; width, including compound eyes: 0.52; vertex width: 0.19; eye dorsal width: 0.19; height in frontal view: 0.57. Antenna. Total length: 1.50; length of scape: 0.12, pedicel: 0.48, flagellomere I: 0.67, flagellomere II: 0,23. Labium. Total length: 1.09; length of segment I: 0.27; II: 0.27; III: 0.23; IV: 0.34. Pronotum. Mesal length: 0.48; posterior maximal width (straight): 0.88. Mesoscutum. Mesal length: 0.05. Scutellum. Anterior width: 0.38; mesal length: 0.36. Hemelytron. Length: 1.65; claval commissure length: 0.29; outer embolial margin length (straight): 1.05; outer cuneal margin length (straight): 0.29; cuneus width: 0.20. Metatarsus. Total length: 0.23; length of tarsomeres I: II: 0.08: 0.20.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF93FFFB7681FD890EE0FEEE.taxon	etymology	Etymology. From Latin anterior (“ that is before, foremost ”) and tenebrôsus (dark, gloomy) in reference to the dark front part of the body.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF93FFFB7681FD890EE0FEEE.taxon	materials_examined	Type material. Holotype (♂): ’ INDONESIA: \ SULAWESI UTARA, \ Dumoga-Bone N. P.: \ July 1985. \\ Fog 15 \ 400 m, 19. vii. 85. \ BMNH Plot C \\ TRAY \ 93 ’ (NHM).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF93FFFB7681FD890EE0FEEE.taxon	distribution	Distribution. Indonesia, North Sulawesi, Bogani Nani Wartabone National Park (formerly known as Dumoga Bone National Park) (Fig. 3 R).	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
03E087EFFF93FFFB7681FD890EE0FEEE.taxon	biology_ecology	Biology. Unknown.	en	Taszakowski, Artur, Kim, Junggon, Jindra, Zdeněk (2025): Further records of the tribe Gigantometopini (Hemiptera, Heteroptera, Miridae, Isometopinae) with descriptions of three new genera and five new species. Zootaxa 5725 (4): 451-474, DOI: 10.11646/zootaxa.5725.4.1, URL: https://doi.org/10.11646/zootaxa.5725.4.1
