taxonID	type	description	language	source
03E487C87B01452CFE3E4152BF05FA8A.taxon	diagnosis	DIAGNOSIS The genus is easily separable from all other genera of Cydnidae in having strongly swollen pronotal umbones concealing posterolateral margins of pronotum (Figs 3 ­ 4), apex of peritreme with more or less blunt triangular lobe posteriorly (Fig. 6), and head submargins bearing only hair­like setae (Figs 1 ­ 2), entirely without pegs or peg­like setae.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B01452CFE3E4152BF05FA8A.taxon	description	DESCRIPTION B o d y. More or less ovate, from 4.7 to 12.7 mm in length, and 2.7 to 6.7 mm in width; general appearance as in Figs 99, 143 and 279. H e a d. Broadly rounded, margins carinate; dorsal surface more or less punctate; clypeus free, usually parallel­sided, sometimes slightly tapering apicad, with or without a pair of subapical hair­like setae; each paraclypeus with two (M. lobatus) or three (all other species) primary hair­like setae (Fig. 1), submarginally without or with 1 ­ 10 secondary hair­like setae (Fig. 2); eyes large, more or less protruding, almost triangular (Fig. 1), or narrowed (Fig. 2), each bearing a single very short apical seta, about third or fourth eye length (Figs 1 ­ 2); ocelli clearly visible; bucculae large, entirely coarsely punctate or impunctate in anterior part; rostrum 4 ­ segmented; antennae 5 ­ segmented. P r o t h o r a x. Pronotum broader than long, more or less narrowed apically; anterior margin angularly insinuated, disc usually punctate anteriorly, laterally, and behind its middle, transverse discal impression behind calli from absent to distinctly developed; lateral margins with submarginal setigerous punctures (absent only in M. lobatus), pronotal disc with three primary setigerous punctures bearing long hair­like setae (absent only in M. lobatus); umbones strongly swollen and protruding caudolaterally, concealing posterolateral margins (Figs 3 ­ 4); pronotal disc usually more or less unicolorous, in some species sharply bicoloured with umbones and / or posterior margin yellow or yellowish brown. Prosternal median carina low, basally stout, coarsely punctate; subcarinal smooth areas large and clearly visible; median part of posterior margin of prosternum straight or almost straight, sometimes bearing small triangular process; propleural depression deep and coarsely punctate; anterior and posterior convexities of propleuron distinctly developed. Mesothorax. Scutellum triangular, longer than broad, sharpened apically; basal and lateral rows of punctures clearly developed, punctures in rows usually connected by incised lines; disc more or less punctate. Posterior wedge­shaped polished band on mesopleural evaporatorium well developed, impunctate or punctate; mesopleural evaporatorium large, occupying almost entire mesopleural surface, bearing more or less clearly visible longitudinal ridges (Fig. 5). Metathorax. Corium divided into clavus, meso­ and exocorium; clavus usually with one complete and one or two incomplete rows of punctures; mesocorium entirely separated from exocorium, with two rows of punctures parallelling clavo­corial suture; costa from entirely separated to entirely fused with exocorium, either with or without setigerous punctures; membrane normally developed, large; membranal suture more or less bisinuate. Metapleural evaporative areas large, bearing more or less clearly visible longitudinal ridges (Fig. 5); apex of peritreme rounded, its anterior margin clearly differentiated from evaporatorium surface or prolonged to fuse with it, its posterior margin with more or less blunt triangular lobe (Fig. 6); scent gland opening large, clearly visible, placed posteriorly on peritreme; posterior area of metapleuron with broader or narrower band of coarse punctures. L e g s. More or less unicolorous, in some species femora bicoloured, yellow or yellowish brown with median dark stripe; fore tibiae expanded apically, margins armed with spines; male hind femora usually with one subapical tooth on dorsal margin, and several more less distinct teeth or spines on ventral margin; female hind femora with or without small subapical tooth on dorsal margin, and several setae on ventral margin; hind tibiae slender, basally more or less reflected and usually with tubercles or carinae in males. Abdomen. Sterna from almost impunctate to distinctly punctate and wrinkled, usually with at least patch of tiny punctures close to spiracles; terga impunctate or punctate, always pilose. Male and female genitalia of the common geotomine type.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B01452CFE3E4152BF05FA8A.taxon	distribution	DISTRIBUTION The members of the genus are distributed only in the Old World warm climate; they reach 46 ° S and 46 ° N latitudes (Fig. 7).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B0D452EFE2F4152BA80FA50.taxon	description	(Figs 8 ­ 14)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B0D452EFE2F4152BA80FA50.taxon	description	DESCRIPTION Body 8.05 ­ 8.65 mm in length, 4.39 ­ 4.55 mm in width, from castaneous to almost black, sometimes corium and apices of pronotal umbones slightly brown or yellowish brown. Head dorsally with tiny and hardly visible punctures; clypeus subapically without a pair of setigerous punctures, each paraclypeus with a single submarginal preocular puncture bearing hair­like seta (Fig. 9); ocular index 2.19 ­ 2.50; interocellar index 3.7 ­ 4.6; length of antennal segments (in mm): 0.35 ­ 0.38: 0.43 ­ 0.46: 0.55 ­ 0.59: 0.73 ­ 0.84: 0.87 ­ 0.94. Pronotal disc weakly punctate, transverse discal impression behind calli absent; each lateral margin with 4 ­ 5 submarginal setigerous punctures (Fig. 8). Propleural depression with numerous coarse punctures. Puncturation of a scutellar disc very weak, better visible only in its median part. Mesocorium punctate with large punctures basally and the smaller apically; exocorium densely punctate; costa narrow, almost entirely separated from exocorium, bearing no setigerous punctures (Fig. 8). Hind femora with 1 ­ 2 small teeth subapically, hind tibiae with neither denticles nor emargination on inner margins (Fig. 10). Abdominal sterna smooth and polished; sutures laterally with small punctures, each segment with a patch of punctures posterior to spiracle, and a few punctures lateral and anterior to it; pregenital segment almost impunctate. Male genitalia as in Figs 11 ­ 14.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B0D452EFE2F4152BA80FA50.taxon	materials_examined	TYPE DATA Holotype male: Papua New Guinea, East. High. Prov., Alyura (BMNH). MATERIAL EXAMINED Holotype, 3 paratypes, and 3 other specimens: Papua New Guinea: East. High. Prov., Alyura, 3 paratypes (BMNH, UO); Star Range, Sibil, 3 exx. (NNML, UO).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B0D452EFE2F4152BA80FA50.taxon	distribution	DISTRIBUTION Papua New Guinea.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B0D4533FE3D45A2B87AFD09.taxon	description	(Figs 15 ­ 21)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B0D4533FE3D45A2B87AFD09.taxon	description	DESCRIPTION Body 8.43 ­ 11.27 mm in length, 4.56 ­ 5.88 mm in width, ochraceous to almost black. Head dorsally with several punctures on paraclypei; clypeus without a pair of subapical hair­like setae, each paraclypeus with a single preocular submarginal setigerous puncture (Fig. 16); ocular index 2.20 ­ 3.10; interocellar index 4.8 ­ 7.0; length of antennal segments (in mm): 0.39 ­ 0.45: 0.53 ­ 0.67: 0.60 ­ 0.76: 0.78 ­ 0.98: 0.94 ­ 1.04. Pronotal transverse discal impression behind calli usually conspicuously developed; anterior margin, lateral parts and posterior lobe more or less distinctly punctate; lateral margins with 5 ­ 6 submarginal setigerous punctures (Fig. 15). Propleural depression and basal half of posterior convexity distinctly punctate. Scutellum punctate, except anterolateral angles and apex; punctures of the same size as those on pronotum. Clavus with one complete and two partial rows of punctures; mesocorial disc almost evenly punctate, punctures almost of the same size as those on pronotum; exocorium with very dense puncturation, punctures smaller than those on mesocorium; costa narrow, almost entirely separated from exocorium, bearing 2 setigerous punctures (Fig. 15). Male hind femora with large subapical tooth on dorsal margin, and several smaller teeth on ventral margin, hind tibiae carinated and bearing small subbasal tubercles (Fig. 17); female hind femora with several strong setae on ventral margin, hind tibiae without tubercle. Abdominal sterna distinctly punctate in lateral two thirds, sometimes puncturation of last two segments less distinct. Male genitalia as in Figs 18 ­ 21. TYPE DATA Lectotype female (designated by LIS 1994 a: 213): Hong Kong (BMNH).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B0D4533FE3D45A2B87AFD09.taxon	materials_examined	MATERIAL EXAMINED Lectotype, and 87 other specimens: Bhutan: British Bhutan, 8 exx. (IRSNB, UO); Mt Durel, 4 exx. (MNHN, UO). China: Canton, 2 exx. (MNHU); Kushan, Foochow, Fuz­zhen, 5 exx. (ZlP, UO); San­nen­Kai, Yunnan­Fou, 8 exx. (NNML). Hong Kong: 2 exx. (BMNH). India: Ajmer­Rajpudana, 1 ex. (IRSNB); Karikal, Karumbagaram, 1 ex. (IRSNB); Yercaud, Shevaroy Hills, 3 exx. (IRSNB, UO); Cinchona, Anamalai Hills, 16 exx. (IRSNB, ZMA, UO); Trichinopoli, 6 exx. (MNHN, UO); Madura, 1 ex. (HNHM); Bombay, 1 ex. (HNHM); Calcutta, 1 ex. (HNHM); Matheran, 4 exx. (HNHM); Pedong, Darjeeling, 3 exx. (BMNH, UO); Agumbe Ghat, Shimoga Dist., Mysore St., 1 ex. (NSMT); Madras State, Anamalai Hills, Kadamparai, 1 ex. (NNML); Kalimpong, 1 ex. (ZIP). Laos: Ban Ban, 1 ex. (BMNH). Nepal: Birethanti, Kaski Dist., 1 ex. (NSMT); Kathmandu Valley, Godavari, 1 ex. (TLM); Central Nepal, road Pokhara ­ Goropani, 1 ex. (UO); Barabhise reg., Ting­Sang­La, 2 exx. (TLM); South Dhaulagiri, Myangdi Kholan Muri, 1 ex. (UO); Kathmandu­Balaju, 1 ex. (UO). Sri Lanka: Punduloya, 1 ex. (OXUM); Uva P. near Madulsima, 1 ex. (BMNH); Ceylan, Nuw Eliya, 1 ex. (MNHN). Thailand: Chieng Mai Province, Doi Inthanon National Park, 1 ex. (ZMUC); Doi Inthanon, 3 exx. (NSMT, UO); Shan States, Manders, 1 ex. (BMNH). Vietnam: Tonkin, reg. de Hoa­Binh, 1 ex. (MNHN); Tam Dao, 1 ex. (ZIP).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B0D4533FE3D45A2B87AFD09.taxon	distribution	DISTRIBUTION Bhutan, Cambodia, China (Yunnan, Guangdong, Fujian), Hong Kong, India, Laos, Nepal, Sri Lanka, Thailand, Vietnam.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B0D4533FE3D45A2B87AFD09.taxon	discussion	REMARK A status of the species was clarified by LIS (1994 a).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B104535FE31430BB903FB36.taxon	description	(Figs 22 ­ 28)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B104535FE31430BB903FB36.taxon	description	DESCRIPTION Body 7.61 ­ 8.83 mm in length, 4.02 ­ 4.96 mm in width, pale castaneous to almost black, posterior margin and umbones of pronotum, exocorium, basal part of mesocorium and clavus clearly yellowish brown. Head dorsally slightly wrinkled in lateral parts of paraclypei; clypeus without a pair of subapical hair­like setae, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 23); ocular index 2.08 ­ 2.36, interocellar index 5.8 ­ 7.5; length of antennal segments (in mm): 0.35 ­ 0.42: 0.34 ­ 0.46: 0.58 ­ 0.67: 0.68 ­ 0.78: 0.74 ­ 0.84. Pronotal disc without or with shallow transverse impression behind calli, dorsal surface of pronotum generally weakly punctate; lateral margins with 3 ­ 6 submarginal setigerous punctures (Fig. 22). Propleural depression with distinct large punctures. Scutellum almost impunctate, bearing only few punctures in its middle. Clavus with three rows of punctures (one complete and two partial); mesocorium evenly punctate; puncturation of exocorium denser than that of mesocorium; costa narrow, almost entirely separated from exocorium, bearing a single setigerous puncture (Fig. 22). Legs clearly paler than remaining parts of the body, yellowish brown with broad dark brown stripe on femora; tibia darker near tarsus, spines castaneous, tarsi yellowish brown; hind tibiae with neither denticles nor emargination on inner margin, hind femora with a few small teeth (Fig. 24). Abdominal sterna smooth and polished, with punctures close to sutures; each segment with a triangular patch of dense fine punctures posterior to spiracle; sterna III­VI with several punctures also anterior to each spiracle. Male genitalia as in Figs 25 ­ 28.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B104535FE31430BB903FB36.taxon	materials_examined	TYPE DATA Lectotype male (designated by LIS, 1999 c: 207) of Macroscytus annulipes HORVÁTH: New Guinea, Friedrich­Wilhelmshafen [= Madang] (HNHM). Holotype male of Macroscytus cheesmanae J. A. LIS: Papua New Guinea, Kokoda (BMNH). MATERIAL EXAMINED Lectotype of Macroscytus annulipes, Holotype of Macroscytus cheesmanae, 17 paratypes of Macroscytus cheesmanae, and 54 other specimens: Bismarck Archipelago: 2 exx. (MNHN); New Britain, Volupai, 1 ex. (BMH). Papua New Guinea: Kokoda, 16 paratypes of M. cheesmanae (BMNH, UO); Madang District, Finisterre Mts., Moro, 1 paratype of M. cheesmanae: (BMNH); Madang Province, Nagada Harbour, 51 exx. (OXUM).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B104535FE31430BB903FB36.taxon	distribution	DISTRIBUTION Bismarck Archipelago (Mussau, New Britain), Papua New Guinea.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B104535FE31430BB903FB36.taxon	discussion	REMARKS Personal examination of the type material of M. annulipes allowed to treat M. cheesmanae as a junior synonym of the former. Figures 10 and 11 in the paper on New Guinean species of Macroscytus (LIS, 1993 b: 45) actually represent M. annulipes (not M. astrolabicus).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B164537FE4C44B0BA16FA56.taxon	description	(Figs 29 ­ 35)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B164537FE4C44B0BA16FA56.taxon	description	DESCRIPTION Body 7.17 ­ 8.76 mm in length, 3.96 ­ 4.75 mm in width, from pale castaneous to dark castaneous, sometimes almost black; entire umbones and posterior margin of pronotum more or less broadly yellowish brown or whitish yellow, making pronotal disc sharply bicoloured; hemelytra paler than pronotum and scutellum, basal third of mesocorium and entire exocorium yellowish brown, yellow or whitish yellow. Head with dorsal surface at first sight looking impunctate, bearing only small and hardly visible punctures; clypeus subapically without a pair of setigerous punctures, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 31); ocular index 2.10 ­ 2.50; interocellar index 5.2 ­ 7.7; length of antennal segments (in mm): 0.29 ­ 0.33: 0.33 ­ 0.42: 0.57 ­ 0.69: 0.63 ­ 0.78: 0.78 ­ 0.88. Puncturation of pronotal disc more or less distinct; transverse discal impression behind calli absent; lateral margins of pronotum with 4 ­ 5 submarginal setigerous punctures bearing long hair­like setae (Fig. 29). Anterior convexity of propleuron punctate anteriorly; posterior convexity with a few irregularly scattered large punctures; propleural depression with a row of large, coarse punctures. Puncturation of scutellar disc more or less dense, well visible in the median part. Mesocorium almost evenly punctate; exocorium sparsely punctate, punctures brown or dark brown in basal half, almost colourless or colourless in the posterior; costa very narrow, almost entirely separated from exocorium, without or with a single setigerous puncture (Fig. 29). Femora yellowish brown, yellow or whitish yellow, sometimes basally brown; tibiae yellowish brown, yellow or whitish yellow, getting darker towards tarsi; dorsal margin of hind male femur with a row of very small teeth (Fig. 30); male hind tibia with a few very small, almost indistinct dark denticles on its inner margin (Fig. 30); female hind femur and tibia without denticles. Abdominal sterna almost impunctate, usually with fine and almost indistinct scattered punctures close to lateral margins, around trichobothria and posterior to spiracles. Male genitalia as in Figs 32 ­ 35.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B164537FE4C44B0BA16FA56.taxon	materials_examined	TYPE DATA Holotype male: Australia, North Queensland, Tully Falls (QMB). MATERIAL EXAMINED Holotype, 20 paratypes, and 10 other specimens: Australia: Queensland, Flying Fish Pt., 1 paratype (QMB); Queensland, Tully Falls, 4 paratypes (QMB, UO); Queensland, Bellenden Ker Base Stn., 5 paratypes (QMB, UO); Queensland, Bellenden Ker Range, 7 paratypes (QMB, UO); Queensland, Stone Ck., 2 paratypes (QMB, UO); Queensland, Kirrama Range, 1 paratype (QMB); Queensland, Mt. Webb Nat. Park, 50 km N. Cooktown, 10 exx. (QMB, UO).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B164537FE4C44B0BA16FA56.taxon	distribution	DISTRIBUTION Australia.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B144536FE3445AEB9D4FAD5.taxon	description	(Figs 36 ­ 37)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B144536FE3445AEB9D4FAD5.taxon	description	DESCRIPTION Body 7.78 ­ 8.01 mm in length, 4.46 ­ 4.65 mm in width, from piceous to black. Head with dorsal surface impunctate; clypeus subapically without a pair of setigerous punctures; each paraclypeus submarginally with a single preocular setigerous puncture; ocular index 2.20 ­ 2.21; length of antennal segments (in mm): 0.36: 0.44: 0.45: 0.56, 5 th missing. Anterior half of pronotal disc impunctate except for few scattered small punctures laterally and behind head; transverse discal impression behind calli absent; posterior half of disc with scattered small punctures; lateral margins of pronotum with 6 submarginal setigerous punctures bearing long hair­like setae. Anterior convexity of propleuron impunctate, except for a patch of small punctures; posterior convexity virtually impunctate; propleural depression with a few punctures. Puncturation of scutellar disc sparse, especially well visible in apical third; punctures a little larger than those of pronotum. Mesocorium almost evenly punctate; exocorium with punctures finer than those of scutellum; costa somewhat flattened, with a single setigerous puncture bearing long hair­like seta. Legs unicolorous; male tibiae with weak basal emargination followed by 5 ­ 7 very small denticles; female hind tibiae simple. Abdominal sterna almost impunctate, weakly and irregularly wrinkled laterally; each segment with a row of punctures along segmental suture, and a patch of small punctures anterior and posterior to spiracle. Male genitalia as in Figs 36 ­ 37.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B144536FE3445AEB9D4FAD5.taxon	materials_examined	TYPE DATA Holotype male: Bismarck Islands, New Ireland, Lemkanin (ZMUC). MATERIAL EXAMINED Holotype, and 1 paratype: Bismarck Archipelago: New Ireland, Lemkanin, 1 paratype (ZMUC).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B144536FE3445AEB9D4FAD5.taxon	distribution	DISTRIBUTION Bismarck Archipelago (New Ireland).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B154538FE354510BA16F9D6.taxon	description	(Figs 38 ­ 44)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B154538FE354510BA16F9D6.taxon	description	DESCRIPTION Body 7.23 ­ 8.17 mm in length, 3.83 ­ 4.32 mm in width, from dark castaneous to almost black. Head with dorsal surface bearing only tiny and hardly visible punctures; clypeus subapically without a pair of setigerous punctures, each paraclypeus submarginally with 3 ­ 4 (very rarely with 2) setigerous punctures bearing long hair­like setae (Fig. 39); ocular index 2.23 ­ 2.40; interocellar index 6.7 ­ 10.8; length of antennal segments (in mm): 0.30 ­ 0.38: 0.42 ­ 0.51: 0.41 ­ 0.50: 0.57 ­ 0.63: 0.65 ­ 0.72. Puncturation of pronotal disc weak; transverse discal impression behind calli absent or very shallow; lateral margins of pronotum with 5 ­ 7 submarginal setigerous punctures bearing long hair­like setae (Fig. 38). Anterior convexity of propleuron impunctate, sometimes with a few small punctures, posterior convexity weakly punctate with large punctures, propleural depression densely and coarsely punctate. Scutellum with coarse large punctures. Mesocorial disc almost evenly punctate; exocorium densely punctate; costa very narrow, entirely separated from exocorium, bearing 1 ­ 2 setigerous punctures (Fig. 38). Male and female hind femur with a row of small tubercles on its dorsal margin, subapical teeth very small, almost indistinct (Fig. 40); male hind tibia with small denticles (Fig. 40), female hind tibia simple. Abdominal sterna more or less polished, almost totally impunctate except for a row of punctures close to each segmental suture, and a few punctures posterior to spiracles and trichobothria; female specimens sometimes with distinct wrinkles laterally to spiracles. Male genitalia as in Figs 41 ­ 44.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B154538FE354510BA16F9D6.taxon	materials_examined	TYPE DATA Holotype male: Australia, Northern Territory, Arnhem Hwy at South Alligator R. (QMB). MATERIAL EXAMINED Holotype, and 27 paratypes: Australia: Northern Territory, Sth. Alligator Inn, 10 paratypes (QMB, UO); Northern Territory, South Alligator R., 5 paratypes (NTM, UO); Northern Territory, 12 km NNW of Mt. Cahill, 1 paratype (NTM); Northern Territory, Limestone Gorge, 2 paratypes (NTM); Northern Territory, Nourlangie Creek, 1 paratype (NTM); Northern Territory, Darwin, East Pl. Res., 1 paratype (NTM); Queensland, Moreton Is., Tertiary Dune Nr Blue Lagoon, 1 paratype (QMB); Queensland, Taroom District, 4 paratypes (QMB, UO); Queensland, Rocky Knob, 10 km SE of Toowoomba, 1 paratype (QMB); Queensland, South Percy Is., 1 paratype (QMB).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B154538FE354510BA16F9D6.taxon	distribution	DISTRIBUTION Australia.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B18453DFE4340A8B83EFE69.taxon	description	(Figs 45 ­ 51)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B18453DFE4340A8B83EFE69.taxon	description	DESCRIPTION Body 7.57 ­ 8.12 mm in length, 4.25 ­ 4.46 mm in width, head, pronotum and scutellum from blackish brown to black, hemelytra castaneous or dark castaneous, entirely umbones and extreme posterior margin of pronotum clearly yellowish brown. Head dorsally with numerous hardly visible tiny punctures, paraclypei striated and wrinkled, impunctate or bearing single clearly visible punctures; clypeus without a pair of subapical hair­like setae, each paraclypeus with two submarginal setigerous punctures (Fig. 46); ocular index 2.65 ­ 2.80, interocellar index 3.0 ­ 4.0; length of antennal segments (in mm): 0.37 ­ 0.42: 0.43 ­ 0.49: 0.52 ­ 0.55: 0.68 ­ 0.72: 0.83 ­ 0.85. Pronotal disc without transverse impression behind calli, its surface punctate with small punctures behind head and in lateral parts, and with large punctures behind calli; lateral margins with 4 ­ 5 submarginal setigerous punctures (Fig. 45). Anterior convexity of propleuron with punctures only in its apical part, posterior convexity punctate only basally, propleural depression with a row or band of large coarse punctures. Scutellar disc with a patch of few large punctures in its median part. Clavus with two clearly visible rows of punctures (one long and one short); mesocorium sparsely punctate; exocorium with dense puncturation; costa narrow, almost entirely separated from exocorium, bearing a single setigerous puncture (Fig. 45). Legs clearly paler than remaining body parts, yellowish brown with broad dark brown stripe on femora; each tibia getting darker towards tarsus, the latter yellowish brown; male hind tibiae with a row of very small tubercles on inner margin, male hind femora with a row of well visible small teeth on dorsal margin (Fig. 47); female hind tibiae simple, female hind femora with a row of small teeth on dorsal margin. Abdominal sterna smooth and polished, with punctures close to sutures; each segment laterally wrinkled and bearing a triangular patch of dense fine punctures anterior and posterior to spiracle. Male genitalia as in Figs 48 ­ 51.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B18453DFE4340A8B83EFE69.taxon	materials_examined	TYPE DATA Lectotype male (present designation): N. Guinea, Biró 1901; Mt Hansemann, Astrolabe B.; astrolabicus det. Horváth; Lectotype Macroscytus astrolabicus Horv., labelled by R. C. Froeschner ̕ 59 (HNHM). MATERIAL EXAMINED Lectotype, and 3 other specimens: Bismarck Archipelago: 1 ex. (UO); Papua New Guinea: NE Morobe, Wau, 2 exx. (BMH).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B18453DFE4340A8B83EFE69.taxon	distribution	DISTRIBUTION Bismarck Archipelago (first record), Papua New Guinea.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B1E453FFE354197B843FA97.taxon	description	(Figs 52 ­ 58)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B1E453FFE354197B843FA97.taxon	description	DESCRIPTION Body 5.41 ­ 7.03 mm in length, 3.09 ­ 4.18 mm in width, from castaneous to almost black, sometimes posterior margin of pronotum and entire corium slightly paler in shade. Head dorsally wrinkled and slightly punctate, clypeus subapically with a pair of setigerous punctures bearing long hair­like setae (Fig. 53); each paraclypeus with a row of 4 ­ 7 submarginal setigerous punctures bearing long hair­like setae (Fig. 53); ocular index 3.18 ­ 4.30; interocellar index 5.5 ­ 8.0; length of antennal segments (in mm): 0.34 ­ 0.41: 0.35 ­ 0.43: 0.36 ­ 0.45: 0.40 ­ 0.45: 0.43 ­ 0.47. Pronotal disc densely punctate, except calli and posterior margin; transverse discal impression behind calli absent or very shallow, almost indistinct; lateral margins with 8 ­ 12 submarginal setigerous punctures bearing long hair­like setae (Fig. 52); anterior convexity of propleuron sometimes with a few small punctures anteriorly, posterior convexity impunctate or with a few scattered punctures, propleural depression with a band of coarse punctures. Scutellum densely punctate with exception of its apex and anterolateral angles. Mesocorial disc densely evenly punctate; exocorium evenly punctate; costa moderately broad, separated from exocorium only in its basal two thirds, bearing 3 ­ 5 setigerous punctures with long hair­like setae (Fig. 52). Male hind femur with numerous well visible denticles on dorsal margin, male hind tibia with a row of small tubercles (Fig. 54); female hind femur without denticles on dorsal margin; female hind tibia without tubercles. Abdominal sterna almost totally impunctate, except for rows of punctures paralleling segmental sutures, and patches of small punctures around trichobothria, as well as anteriorly and posteriorly to spiracles. Male genitalia as in Figs 55 ­ 58.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B1E453FFE354197B843FA97.taxon	materials_examined	TYPE DATA Lectotype female (designated by LIS 1999 c: 207) of Cydnus australis ERICHSON: Vandiemensland [= Tasmania] (MNHU). Lectotype female (designated by LIS 1996: 221) of Aethus leptospermi BUTLER: New Zealand (BMNH). Lectotype male (designated by LIS 1996: 221) of Aethus lifuanus MONTROUZIER: Lifu (NHMW). Lectotype female (designated by SYNAVE 1969: 2, as Holotype) of Geotomus lansbergi SIGNORET: Indonesia, Java Oriental, M. Ardjoeno (IRSNB). MATERIAL EXAMINED Lectotype of Cydnus australis, Lectotype of Aethus leptospermi, Lectotype of Aethus lifuanus, Lectotype of Geotomus lansbergi, and 37 other specimens: Australia: Queensland, Bakerville, 4 exx. (DPI, UO); Queensland, Bellenden Ker Range, 1 ex. (QMB); Queensland, Lake Broadwater via Dalby, 1 ex. (QMB); Queensland, Boggom, via Taroom, 1 ex. (QMB); Queensland, Port Denison, 2 exx. (IRSNB); New South Wales, Ryde, 2 exx. (HNHM); New South Wales, Richmond River, 2 exx. (BMH); New South Wales, Sydeny, 1 ex. (BMNH); New South Wales, Upper Tumut Gorge, Snowy Mts., 1 ex. (QMB); Northern Territory, Ayers Rock, 1 ex. (TLM); Lord Howe I., 2 exx. (SAM); Tasmania, 3 exx. (BMNH, MNHN); Western Australia, Tallering Stn., 1 ex. (WAM); Western Australia, Burnbury, 2 exx. (WAM); Western Australia, Capel, 4 exx. (WAM, UO); Western Australia, Lake Gwelup, 1 ex. (NTM); Western Australia, Clover Dale Hasd., 1 ex. (NTM);. Bismarck Archipelago: New Britain, Birara, 1 ex. (BMNH). New Caledonia: Canala, 2 exx. (MNHN); Anse Vata, 1 ex. (BMH); Hienghene, 1 ex. (HNHM). New Zealand: Motuihi I., Nr. Auckland, 1 ex. (BMH); Port Waikato, 1 ex. (BMNH).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B1E453FFE354197B843FA97.taxon	distribution	DISTRIBUTION Australia (incl. Tasmania and Lord Howe Island), Indonesia (Java), New Britain (first record), New Caledonia, New Zealand.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B1C4501FE4845D0BA16F995.taxon	description	(Figs 59 ­ 65)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B1C4501FE4845D0BA16F995.taxon	description	DESCRIPTION Body 6.08 ­ 7.13 mm in length, 3.71 ­ 4.55 mm in width, from dark castaneous to blackish brown, posterior margin of pronotum slightly more yellowish brown than remaining parts of its disc. Head with dorsal surface bearing numerous small, almost evenly spaced punctures, laterally striated; clypeus subapically with a pair of setigerous punctures bearing long hair­like setae (Fig. 60); each paraclypeus with a submarginal row of 7 setigerous punctures bearing long hair­like setae (Fig. 60); ocular index 2.48 ­ 3.03; interocellar index 7.8 ­ 9.1; length of antennal segments (in mm): 0.28 ­ 0.35: 0.35 ­ 0.44: 0.36 ­ 0.42: 0.43 ­ 0.51: 0.44 ­ 0.50. Pronotal disc almost impunctate, bearing only several punctures of two types (smaller and larger), calli impunctate; discal transverse impression behind calli absent or very shallow, almost indistinct, bearing several coarse punctures and numerous wrinkles; lateral margins of pronotum with 12 ­ 15 submarginal setigerous punctures bearing long hair­like setae (Fig. 59); anterior convexity of propleuron alutaceous and impunctate, posterior convexity with a few punctures close to propleural depression, the latter with rows of well separated coarse punctures. Scutellum densely striated and punctate. Mesocorial disc weakly, evenly punctate; puncturation of exocorium more dense than that of mesocorium; costa separated from exocorium only in its basal two thirds, its margin with a row of 5 ­ 8 (very rarely 4) setigerous punctures bearing long hair­like setae (Fig. 59). Femora brown or castaneous, tibiae whitish yellow or yellowish brown, tarsi yellowish brown or pale brown; male hind femur with numerous well visible denticles, subapical teeth clearly developed (Fig. 61); male hind tibia with large subbasal lobe­like denticle followed by smaller denticle and then several smaller tubercles (Fig. 61). Abdominal sterna almost totally impunctate, except for rows of punctures paralleling segmental sutures, and a few punctures close to spiracles and trichobothria. Male genitalia as in Figs 62 ­ 65.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B1C4501FE4845D0BA16F995.taxon	materials_examined	TYPE DATA Holotype male: South Australia, Coorong, S. of Meninge (BMNH). MATERIAL EXAMINED Holotype, 3 paratypes, and 14 other specimens: Australia: New South Wales, Nygan distr., 3 exx. (UQIC, UO); South Australia, Underdale, 3 paratypes (SAM, UO); South Australia, Along railway line Goyder Siding yo Bowmans Junctn., 1 ex. (SAM); Queensland, Brisbane, 3 exx. (UQIC, UO); Queensland, Taroom District, Vine Thicket on hill, 1 ex (QMB); Queensland, Taroom District, Boggomoss, 4 exx. (QMB, UO); Queensland, Texas, 1 ex. (HNHM); Western Australia, Barnabie, 1 ex. (SAM).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B1C4501FE4845D0BA16F995.taxon	distribution	DISTRIBUTION Australia.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B234502FE2B40A8B941FB49.taxon	description	(Figs 66 ­ 72)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B234502FE2B40A8B941FB49.taxon	description	DESCRIPTION Body 5.70 ­ 7.83 mm in length, 3.15 ­ 4.44 mm in width, from pale brown to black­brown, corium paler; sometimes apical parts of umbones and posterior margin of pronotum slightly lighter than remaining surface of pronotal disc. Head dorsally impunctate, except for setigerous punctures and several almost indistinct tiny punctures on paraclypei; clypeus without subapical setigerous punctures, each paraclypeus with a row of 4 ­ 8 submarginal hair­like setae (Fig. 67); ocular index 2.10 ­ 2.80; interocellar index 9.0 ­ 15.0; length of antennal segments (in mm): 0.24 ­ 0.35: 0.32 ­ 0.50: 0.27 ­ 0.42: 0.36 ­ 0.53: 0.43 ­ 0.54. Pronotum weakly or moderately punctate, calli and posterior fifth of disc impunctate, transverse discal impression behind calli absent or shallow; lateral margins with 9 ­ 13 submarginal setigerous punctures (Fig. 66). Anterior convexity of propleuron apically sparsely punctate; depression and posterior convexity densely punctate. Puncturation of scutellum denser than that of pronotal disc, punctures slightly larger than those on the latter. Corium distinctly punctate; clavus with one complete and one or two partial rows of punctures; mesocorial disc and exocorium almost evenly punctate; costa entirely separated from exocorium, bearing 4 ­ 7 setigerous punctures (Fig. 66). Hind femora with large subapical tooth on dorsal margin (Fig. 68); male hind tibiae carinate, without denticles (Fig. 68). Abdominal sterna slightly transversely wrinkled, impunctate or with small punctures posterior to spiracles. Male genitalia as in Figs 69 ­ 72.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B234502FE2B40A8B941FB49.taxon	materials_examined	TYPE DATA Lectotype male (designated by LIS, 1991 a: 184) of Aethus badius WALKER: India (BMNH). Syntype (s) of Macroscytus expansus SIGNORET: India, Bombay (BMNH, lost). MATERIAL EXAMINED Lectotype of Aethus badius, and 131 other specimens: Burma: Palon, Pegu, 1 ex. (MCSN); Schwego­Myo, 1 ex. (MCSN). India: Bengale, 2 exx. (MNHN); Inde, Arrah, 1 ex. (MNHN); Madras Airport, 11 exx. (NNML); Karumbagaram, Karikal Terr., 3 exx. (UO); Karikal, 4 exx. (NNML); Pondichery, 1 ex. (UO); Cote de Malabar, Mahe, 3 exx. (MNHN); Maissour, Silvapoora, 1 ex. (MNHN); Malabar, Walayar Forests, 1 ex. (IRSNB); Ajmer­Rajpudana, 1 ex. (IRSNB); Barway, 1 ex. (IRSNB); Madras State, Coimbatore, 15 exx. (ZMA, UO, NSMT, NNML); Delhi, 2 exx. (ZIP); Orissa, Konarak, 2 exx. (HNHM); Mettupalayam, Tamil Nadu, 2 exx. (HNHM); Orissa, Bhubaneswar town, 4 exx. (HNHM, UO); Chipurupalle, Vizagapatan Distr., 1 ex. (BMNH); Bhimavaram, Kistna Distr., 1 ex. (UO); Serampur, 1 ex. (UO); Bombay, 1 ex. (BMNH); Pilant, Rajasthan, 1 ex. (BMNH); Chikkaballappura, 1 ex. (BMNH); Mysore, 1 ex. (BMNH); Bandra, 1 ex. (BMNH); Annandale, Madras, 1 ex. (BMNH); Kolimpong, 1 ex. (ZIP); Kerala State, Trivandrum Distr., Poonmundi Range, 1 ex. (UO); Anamalai Hills, Cinchona, 1 ex. (NNML); Santiniketan, Birbhum, Bengal, 3 exx. (BMH); Meghalaya state, West Garo Hills, Bagmara, 2 exx. (NHMW); Dehra Dun, Kumaon Himalaya, 1 ex. (ZSMC). Maldive Islands: Gan Addu Atoll, 11 exx. (BMNH, UO); Villingilli, 16 exx. (TLM, UO). Nepal: Hetaura, 1 ex. (HNHM). Sri Lanka: Ceylon, Diyatalawa, 1 ex. (BMNH); Ceylon, Hambantota, 2 exx. (BMNH, UO); Ceylon, Yala, 1 ex. (BMNH); Elephant Pass N. P., 1 ex. (BMNH); Bopikiya, 1 ex. (BMNH); Arugam Bay, 1 ex. (SMNS); Uppuvell, 1 ex. (SMNS); Kandy, 1 ex. (ZIP); National Park Wilpattu, Talawila, 8 exx. (ZIP, UO); National Park Yala, Mahasilawa, 5 exx. (ZlP, UO); 9 km ESE Maradankadawalkka, 6 exx. (ZlP, UO); Colombo, 1 ex. (ZIP).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B234502FE2B40A8B941FB49.taxon	distribution	DISTRIBUTON Burma, India, Maldive Islands, Nepal (first record), Sri Lanka. The occurrence of this species in China (WU, 1935) still remains unconfirmed and seems rather doubtful (LIS 1994 a). The records from Japan pertain to Aethus pseudindicus (see LIS 1994 a).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B214504FE3544B7BA5BFA76.taxon	description	(Figs 73 ­ 79)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B214504FE3544B7BA5BFA76.taxon	description	DESCRIPTION Body 9.36 ­ 11.81 mm in length, 4.95 ­ 5.96 mm in width, black, corium usually paler, umbones of pronotum intensely ochraceous or yellowish brown. Head dorsally distinctly punctate, except clypeus and vertex; clypeus without a subapical pair of setigerous punctures, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 74); ocular index 2.40 ­ 3.22; interocellar index 4.7 ­ 8.0; length of antennal segments: 0.40 ­ 0.55: 0.59 ­ 0.74: 0.71 ­ 0.84: 0.98 ­ 1.14: 1.09 ­ 1.26. Pronotal disc with large deep punctures; umbones and posterior margin impunctate; transverse discal impression behind calli absent; lateral margins with 5 ­ 6 submarginal setigerous punctures bearing long hair­like setae (Fig. 73). Propleural depression and posterior convexity with deep large punctures, anterior convexity with smaller punctures laterally. Scutellum punctate with large deep punctures. Clavus with one complete and two partial rows of punctures; mesocorial disc almost evenly punctate; exocorium densely punctate; all punctures smaller than those on scutellum and pronotum; costa narrow, entirely separated from exocorium, bearing a single setigerous puncture (Fig. 73). Male hind femur with large subapical tooth on dorsal margin (Fig. 75), female hind femur with small tooth; hind tibiae in both sexes without tubercles and denticles on inner margin (Fig. 75). Abdominal sterna with distinct punctures anteriorly, posteriorly and mesad of spiracles; each segmental suture accompanied by a row of dense punctures. Male genitalia as in Figs 76 ­ 79.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B214504FE3544B7BA5BFA76.taxon	materials_examined	TYPE DATA Holotype male: Malaysia, Borneo, Sabah, Mts. Crocker, 16 miles NW of Keningau (NSMT). MATERIAL EXAMINED Holotype, 7 paratypes, and 6 other specimens: Malaysia: Borneo, Sabah, Mts. Crocker, 16 miles NW of Keningau, 2 paratypes, 2 exx. (NSMT, UO); Sabah, 1 / 2 way Rd Melligan ­ Long Pa Sian, 1 paratype (NNML); Sabah, Gn Kinabalu, Sg. Liwagu nr Kundasang bridges, 1 paratype (NNML); Sabah, Long Pa Sian between LPS­L Samando (2 km from Sarawak border), 2 paratype (NNML); Borneo Sabah, Crocker Mt., Gununag Emas, 2 exx. (USMB, UO); Malaya, Pahang, Cameron Highlands, 2 km S Tanah Rata, on Tapah Road, 1 ex. (HNHM). Indonesia: East Borneo, Sanga Sanga, Moorjawa, 1 paratype (UO); O. Borneo, Samarinda, 1 ex. (MZLU).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B214504FE3544B7BA5BFA76.taxon	distribution	DISTRIBUTION Indonesia (Borneo Kalimantan), Malaysia (Borneo Sabah, Malaya ­ first record).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B274509FE3A45F0BA16FE29.taxon	description	(Figs 80 ­ 86)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B274509FE3A45F0BA16FE29.taxon	description	DESCRIPTION Body 8.68 ­ 10.15 mm in length, 4.70 ­ 5.67 mm in width, from castaneous to almost black, entire umbones and posterior margin of pronotum clearly yellowish brown or whitish yellow, basal third of mesocorium and entire exocorium always yellowish brown, yellow or whitish yellow. Head with dorsal surface at first sight looking impunctate, but bearing small, irregularly scattered punctures; clypeus subapically without setigerous punctures, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 81); ocular index 2.18 ­ 2.30; interocellar index 6.7 ­ 9.0; length of antennal segments (in mm): 0.38 ­ 0.44: 0.54 ­ 0.67: 0.65 ­ 0.78: 0.85 ­ 0.99: 0.97 ­ 1.13. Puncturation of pronotal disc more or less distinct, usually better visible behind calli; transverse discal impression absent; lateral margins of pronotum with 4 (rarely 5) submarginal setigerous punctures bearing long hair­like setae (Fig. 80). Anterior convexity of propleuron more or less punctate, posterior convexity distinctly punctate with irregularly scattered coarse punctures, propleural depression with a row of coarse punctures. Scutellum with puncturation better visible in its median part. Mesocorial disc almost evenly punctate with large punctures basally and small punctures apically; exocorium more or less densely punctate; costa narrow, entirely separated from exocorium, basally flattened and bearing two setigerous punctures (Fig. 80). Dorsal margin of male hind femur with large subbasal tooth followed by 1 ­ 2 smaller teeth, and clearly visible apical tooth (Fig. 82); male hind tibia without denticles on its inner margin, its basal third distinctly bent outwards (Fig. 82); dorsal margin of female hind femur with more or less blunt apical denticle; female hind tibia straight, without denticles on its inner margin. Abdominal sterna polished, each with a row of punctures close to segmental sutures, median two thirds of segments impunctate, lateral third with small, crowded punctures around trichobothria and posterior to spiracles. Male genitalia as in Figs 83 ­ 86.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B274509FE3A45F0BA16FE29.taxon	materials_examined	TYPE DATA Holotype male: Australia, North Queensland, Little Cedar Ck., Mt. Spec (QMB). MATERIAL EXAMINED Holotype, 17 paratypes, and 12 other specimens: Australia: Queensland, Little Cedar Ck., Mt. Spec, 4 paratypes (QMB, UO); Queensland, Mt Misery summit, 1 paratype (QMB); Queensland, Saddle Mtn, 3 paratypes (QMB, UO); Queensland, Leo Creek Road, McIlwrith Range, 30 km N. E. of Coen, 3 paratypes (QMB, UO); Queensland, East Caludie R., Iron Range, 2 paratypes (QMB); Queensland, Karnak­Devil ̕ s Thumb, 8 ­ 12 Km NW Mossman, 1 paratype (QMB); Queensland, Mt Hayward, 2 paratypes (QMB); Queensland, Mt Hayward, 1 paratype (QMB); Queensland, Kirrama State Forest, via Cardwell, 4 exx. (UQIC, UO); Queensland, Iron Range, Cape York Pen., 2 exx. (UQIC, UO); Queensland, Emerald Ck., Mareeba, 2 exx. (DPI, UO); Queensland, the Boulders via Babinda, 1 ex. (UQIC); Queensland, Massey River, McIlwraith R., 1 ex. (DPI); Queensland, Churchill Ck., Mt. Lewis Road, Via Julatten, 1 ex. (UQIC).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B274509FE3A45F0BA16FE29.taxon	distribution	DISTRIBUTION Australia.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B2A450BFE304257B9CAFDE9.taxon	description	(Figs 87 ­ 93)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B2A450BFE304257B9CAFDE9.taxon	description	DESCRIPTION Body 8.95 ­ 12.05 mm in length, 4.70 ­ 6.08 mm in width, from castaneous to almost black, corium usually more brownish than pronotum and scutellum. Head dorsally with numerous evenly spaced and almost invisible tiny punctures; clypeus without subapical pair of setigerous punctures, each paraclypeus with a single preocular submarginal setigerous puncture (Fig. 88); ocular index 1.73 ­ 2.38; interocellar index 5.7 ­ 8.4; length of antennal segments (in mm): 0.45 ­ 0.52: 0.52 ­ 0.68: 0.81 ­ 0.97: 0.99 ­ 1.29: 1.18 ­ 1.32. Pronotum with transverse discal impression behind calli deep and distinct only laterally, bearing numerous punctures; anterior lobe of pronotal disc with a patch of punctures behind head and several punctures laterally, posterior lobe with numerous scattered punctures; lateral margins with 3 ­ 5 submarginal setigerous punctures (Fig. 87). Propleural depression and posterior convexity with large punctures, anterior convexity dulled by numerous tiny punctures. Scutellum distinctly punctate, except anterolateral angles; apex with a few small punctures. Clavus with one complete, and 1 ­ 2 partial rows of punctures; mesocorium almost evenly punctate; exocorium almost impunctate; punctures on corium smaller than those on pronotum and scutellum; costa narrow, almost entirely separated from exocorium, bearing 1 ­ 2 setigerous punctures (Fig. 87). Hind femora with numerous small teeth on ventral margin, and distinct subapical tooth on the dorsal (Fig. 89); male hind tibia with several well visible tubercles on inner margin (Fig. 89). Abdominal sterna wrinkled laterally, and with tiny punctures anterior and posterior to spiracles; segmental sutures with tiny punctures. Male genitalia as in Figs 90 ­ 93.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B2A450BFE304257B9CAFDE9.taxon	materials_examined	TYPE DATA Holotype male: Malaysia, North Borneo, Samarang, Nr Sandakan (BMNH). MATERIAL EXAMINED Holotype, 24 paratypes, and 7 other specimens: Brunei: Ulu, Temburong, 2 paratypes (BMNH, UO). Malaysia: Malaya, Kuala Lumpur, 17 paratypes (BMNH, UO); Malay Peninsula, Selangor, Bukit Kutu, 1 paratype (BMNH); Sarawak, Kuching, 1 paratype (BMNH); Sabah nr Long Pa Sia, 1 paratype (NNML); Sabah, Beaufort, 2 paratypes (NNML); Borneo, Sabah, 5 exx. (MNHN, UO); Sabah, Sipitang, Mendolong, 2 exx. (MZLU).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B2A450BFE304257B9CAFDE9.taxon	distribution	DISTRIBUTION Brunei, Malaysia (Malaya, Borneo).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B28450CFE4C4217B8E1FE4A.taxon	description	(Figs 94 ­ 98)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B28450CFE4C4217B8E1FE4A.taxon	description	DESCRIPTION Body 6.00 ­ 9.13 mm in length, 3.25 ­ 4.86 mm in width, from pale brown to blackish brown, corium usually paler than pronotum and scutellum. In general appearance (habitus and body puncturation) very similar to M. badius. Head dorsally impunctate, sometimes with distinct wrinkles and several almost indistinct tiny punctures on paraclypei; clypeus without subapical setigerous punctures (in several aberrant specimens one or two setigerous punctures on clypeus were found); each paraclypeus with a row of 4 ­ 8 submarginal hair­like setae (Fig. 95); ocular index 2.20 ­ 3.35; length of antennal segments (in mm): 0.30 ­ 0.43: 0.38 ­ 0.61: 0.28 ­ 0.51: 0.48 ­ 0.71: 0.54 ­ 0.77. Pronotum weakly punctate, sometimes almost impunctate, transverse discal impression behind calli absent or very shallow; lateral margins with 8 ­ 13 submarginal setigerous punctures (Fig. 94). Anterior convexity of propleuron impunctate or almost impunctate, posterior convexity with a few punctures, propleural depression striated, sometimes bearing a row of more or less distinct punctures. Scutellum more or less punctate, puncturation usually denser than that of pronotal disc. Corium distinctly punctate; clavus with one complete and two partial rows of punctures; mesocorial disc almost evenly punctate with small punctures; exocorium almost evenly punctate, punctures larger than those on mesocorium; costa narrow, almost entirely separated from exocorium, bearing 4 ­ 7 setigerous punctures (Fig. 94). Hind femora with clearly visible subapical tooth on dorsal margin, that may vary in size; hind tibiae simple (Fig. 96). Abdominal sterna laterally slightly transversely wrinkled, impunctate, except for a few small punctures posterior to spiracles. Male genitalia as in Figs 97 ­ 98.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B28450CFE4C4217B8E1FE4A.taxon	materials_examined	TYPE DATA Lectotype male of Cydnus brunneus FABRICIUS (designated by LIS, 1994 a: 239): Morocco, Tanger (ZMUC). Lectotype male of Macroscytus scutellaris HORVÁTH (designated by LINNAVUORI, 1977: 41): Mozambique, East Africa, Zambesi (HNHM). Syntype (s) of Cydnus proximus RAMBUR: Spain, env. of Cadiz (BMNH). Syntype (s) of Aethus opacus STÃL: South Africa, Caffraria (NNML). Lectotype male of Macroscytus exiguus HORVÁTH (designated by LINNAVUORI, 1977: 41): South Africa, Natal (HNHM). Lectotype male of Macroscytus exiguus HORVÁTH (designated by LINNAVUORI, 1977: 41): South Africa, Natal (HNHM). MATERIAL EXAMINED Lectotype of Cydnus brunneus, Lectotype of Macroscytus scutellaris, Lectotype of Macroscytus exiguus, and 379 other specimens: Afghanistan: Kizyl­Kha, 2 exx. (UO). Algeria: Laghouat, 1 ex. (MNHN); Randon, 1 ex. (MNHN). Botswana: Gaborone, 1 ex. (MZLU). Canary Islands: Gran Canaria, Artedara b. Fataja, 2 exx. (TLM); Gran Canaria, Ginamar, 3 exx. (MNHN). Cyprus: Paphos, 11 exx. (CMNH, UO). Egypt: Cairo, 2 exx. (NNML). Greece: Corfu, 3 exx. (NNML). Iran: 25 km S Minab, 1 ex. (NHMW); Bandar Langeh, 12 exx. (MNHN, UO). Iraq: Bagdad, 3 exx. (HNHM, UO). Israel: Askhelon, 6 exx. (CMNH); Hullah Valley, 2 exx. (ZSMC); EnGedi, 1 ex. (ZSMC). Italy: Sardinia, 1 ex. (HNHM). Jordan: Wadi Sir, n. Amman, 1 ex. (HNHM); Arda Road, 1 ex. (HNHM). Morocco: O El Moddah, 1 ex. (UO); Safi­Agadir road, 1 ex. (UO); Oudjda, 3 exx. (MNHN, UO). Mozambique: Lourenco Marques, Matola, Vila Salazar, 1 ex. (JEC). Namibia: Bushmanland, Klein Dobe, 11 exx. (MNHU, UO); Fish­River­Canyon, 6 exx. (MNHU, UO); Etoscha N. P., Namutoni, 6 exx. (MNHU, UO); Kavango, Popa Falls, 3 exx. (MNHU); East Caprivi, Mudumu N. P., Makatwa, 3 exx. (MNHU, UO); East Caprivi, Katima Mulilo, 2 exx. (MNHU); Ongongo Fall, Warmbad, Kaokoveld, 3 exx. (MNHU, UO); Kunene, 9 km W Ruacana, 2 exx. (MNHU). Oman: Ruwi, 2 exx. (MZLU). Pakistan: Karachi, 4 exx. (NHMUK); Baluchistan, Quetta, 1 ex. (BMNH). Portugal: Algarve, Sta Barbara de N., 17 exx. (MNHN, UO). Saudi Arabia: Najd Region, 50 mi SE Riyadh, 6 exx. (JEC). Senegal: Dakar Bel­Air, 1 ex. (MNHN). South Africa: Natal, Empangeni area, 4 exx (NMP, UO); Natal, Mntunzini Garland Fm, 1 ex. (NMP); Natal, Mkuzi Game Reserve, 1 ex. (NMP); Mkuze Game Res., 1 ex. (UO); Natal, Itala Game Reserve, Louwsburg, 2 exx. (MNHU); Cape Prov., Oranje River, Vioolsdrif, 1 ex. (MNHU); Transvaal, Zoutpansberg [= Soutpansberg], 1 ex. (ZMPA). Tunisia: Boughrara, 4 exx. (HNHM); Maknassy, 8 exx. (MNHN, UO). Turkey: Prov. Urfa, Halfeti, valley of Euphrat, 2 exx. (HNHM). Turkmenistan: Kopet­Dagh Mts, 6 km S of Ipay­Kala, 1 ex. (HNHM); Kopet­Dagh Mts, 5 km S of Chull, 1 ex. (HNHM). Zambia: South Luangwa N. P., Mfuwe, Crocodile Farm, 1 ex. (MNHU); Muchinga Escarpment, 1 ex. (MNHU). Zimbabwe: St. Benedict, 30 km NE Macheke, 2 exx. (TLM); Chinhoyi Caves, 1 ex. (NMP); Victoria Falls, Zambesi Nat. Park, 1 ex. (MNHU); Matabeleland, Lake Mutirikwi bei Masvingo, Kyle N. P., 11 exx. (MNHU, UO); Matabeleland, Matopos Nat. Park, 25 exx. (MNHU, UO); S. Rhodesia, Kariba, 182 exx. (JEC, UO).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B28450CFE4C4217B8E1FE4A.taxon	distribution	DISTRIBUTION Afghanistan, Albania, Algeria, Azerbaidjan, Botswana, Cameroon, Canary Islands, Cape Verde Islands, Central African Republic,? China (North), Congo, Croatia, Cyprus, Egypt, Ethiopia, France, Gambia, Ghana, Greece, Guinea, Iran, Iraq, Israel, Italy, Ivory Coast, Jordan (first record), Kazakhstan, Kenya, Liban, Libya, Macedonia, Mali, Malta, Morocco, Mozambique, Namibia (first record), Nigeria, Oman (first record), Pakistan, Portugal, Saudi Arabia, Senegal, Socotra, Somalia, South Africa, Spain, Sudan, Switzerland, Syria, Tadzhikistan, Tanzania (incl. Zanzibar), Turkey, Turkmenistan, Tunisia, Uganda, Uzbekistan, Yemen, Yugoslavia, Zaire, Zambia (first record), Zimbabwe (first record).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B2F450FFE3D41B2BAD0FB36.taxon	description	(Figs 99)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B2F450FFE3D41B2BAD0FB36.taxon	description	DESCRIPTION Body 8.05 ­ 9.80 mm in length, 4.75 ­ 5.25 mm in width, black or blackish brown, except broad transverse yellowish brown band at posterior margin of pronotum (Fig. 99) and pale brown corium. Head dorsally impunctate; clypeus without a pair of subapical setigerous punctures; each paraclypeus submarginally with a single preocular setigerous puncture; ocular index 3.15 ­ 3.52; interocellar index 3.3 ­ 4.2; length of antennal segments (in mm): 0.48 ­ 0.49: 0.55 ­ 0.56: 0.66 ­ 0.67: 0.81 ­ 0.86: 0.98 ­ 0.99. Lateral parts of pronotum somewhat flattened (Fig. 99); transverse discal impression behind calli absent; lateral margins with 3 submarginal setigerous punctures; anterior part of a disc with numerous small punctures forming an arcuate subapical transverse band, and irregular patch on either side; posterior half with a transverse broad band of small punctures not reaching the posterior margin. Propleuron with numerous small punctures on anterior convexity, depression with crowded coarse punctures which become finer and sparser posteriorly. Scutellar disc with large coarse punctures. Clavus with one complete and two partial rows of punctures; mesocorium and exocorium densely punctate; costa somewhat flattened, separated from exocorium only basally, bearing no setigerous punctures. Hind femora subapically with small tooth. Abdominal sterna laterally slightly wrinkled, each bearing a patch of punctures posterior and anterior to spiracle.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B2F450FFE3D41B2BAD0FB36.taxon	materials_examined	TYPE DATA Syntype (s): female (s), South Celebes, Loka (lost or destroyed). Neotype (present designation): Indonesia, Sulawesi Is, 1 female, X­XI 1988, S. Nagai (NSMT). MATERIAL EXAMINED Neotype, and 2 other specimens: Indonesia: Sulawesi Utara, Dumoga­Bone National Park, Poniki trail, 1 ex. (IRSNB); North Sulawesi, Dumoga Bone National Park, 1 ex. (NNML).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B2F450FFE3D41B2BAD0FB36.taxon	distribution	DISTRIBUTION Indonesia (Sulawesi).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B2F450FFE3D41B2BAD0FB36.taxon	discussion	REMARK Male unknown.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B2C4510FE2444A4BA15FE29.taxon	description	(Figs 100 ­ 106) DESCRIPTION H e a d. Black, but lateral margins brownish in shade; dorsal surface slightly alutaceous, impunctate, bearing only a few punctures laterally; gular plate impunctate in anterior half, posterior part distinctly punctate; bucculae entirely coarsely punctate; clypeus free, as long as paraclypei, subapically without setigerous punctures, each paraclypeus submarginally with two setigerous punctures bearing hair­like setae (Fig. 101); eyes brown, ocular index 2.21; ocelli reddish brown, interocellar index 9.1, ocellar index 5.5; antennae brown; rostrum brown, reaching mid coxae. Prothorax. Pronotum dark castaneous; pronotal disc with large punctures in its posterior half, calli impunctate and distinctly elevated; transverse discal impression shallow, with medially interrupted row of several coarse punctures behind calli; lateral margins of pronotum with 6 ­ 7 submarginal setigerous punctures bearing long hair­like setae (Fig. 100). Prosternal carinae low, weakly punctate; anterior and posterior convexities of propleuron punctate with large punctures, propleural depression with a band of coarse punctures. Mesothorax. Scutellum dark castaneous, its disc weakly punctate, punctures as large as those on pronotum, irregularly scattered. Mesopleural evaporatorium with coarse longitudinal rugae, its posterior wedge­shaped polished band somewhat dulled, bearing a few large punctures. Metathorax. Hemelytra dark castaneous, punctate with almost colourless punctures; clavus with one complete and two incomplete rows of punctures; mesocorial disc almost evenly punctate, but puncturation in basal half weaker than in the apical; exocorium with puncturation more dense close to mesocorium; mesocorium and exocorium separated along their entire length; costa narrow, slightly convex and separated from exocorium only in its basal half, costal margin with two setigerous punctures bearing long hair­like setae (Fig. 100); membranal suture slightly bisinuate; membrane semihyaline, somewhat embrowned, its tip slightly surpassing the posterior extremity of abdomen. Metapleural evaporatorium with longitudinal rugae; lateral area of metapleuron impunctate, posterior area with a band of coarse punctures. L e g s. Femora and tibiae castaneous, tarsi pale brown or brown; male hind femur (Fig. 102) with 2 subapical teeth on ventral margin, its dorsal margin with numerous denticles, and a single small subapical tooth; male hind tibia with a row of small tubercles (Fig. 102). Abdomen. Sterna dark castaneous, distinctly punctate in lateral two thirds, punctures large and coarse. Ventral surface of male pygophore almost impunctate, bearing only single punctures laterally, its dorsal surface pilose, longitudinal sclerites on infolding of its dorsal rim well developed; opening of genital capsule as in Fig. 103, penis as in Fig. 105, second conjunctival appendage as in Fig. 106, paramere as in Fig. 104. Measurements (in mm). Male only: body length 8.08; body width 4.31; head length 1.32; head width 1.97; pronotum length 2.27; pronotum width 3.91; scutellum length 3.07; scutellum width 2.47; antennal segments: 0.35: 0.55: 0.57, 4 th and 5 th segments missing.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B2C4510FE2444A4BA15FE29.taxon	materials_examined	TYPE MATERIAL EXAMINED Holotype male: Comoros: Grande Comore, 4 / 5 km E. de Nioumbadjou, Bandalamadji, 640 m, 11 / 14 ­ III­ 1980, P. Viette, Muséum Paris (MNHN).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B2C4510FE2444A4BA15FE29.taxon	discussion	COMPARATIVE NOTES At first glance the species is similar to M. mayottensis. Nevertheles, it can easily be separated from the latter by its puncturation of abdominal sterna (punctures present in lateral third in M. myottensis, in lateral two thirds in M. comorensis), and puncturation of gular plate (almost impunctate, punctures present only in its posterior third in M. myottensis, punctures in the posterior half in M. comorensis). Moreover, both species differ also in their male genitalia and edentation of male hind femora.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B2C4510FE2444A4BA15FE29.taxon	distribution	DISTRIBUTION Comoros.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B334512FE334257BF07FD09.taxon	description	(Figs 107 ­ 113)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B334512FE334257BF07FD09.taxon	description	DESCRIPTION Body 9.36 ­ 11.57 mm in length, 4.71 ­ 5.88 mm in width, from castaneous to blackish brown, corium usually more brownish than pronotum and scutellum. Head dorsally slightly wrinkled on paraclypei, impunctate; clypeus without a pair of submarginal setigerous punctures, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 108); ocular index 1.77 ­ 2.32; interocellar index 6.6 ­ 8.0; length of antennal segments (in mm): 0.43 ­ 0.56: 0.55 ­ 0.73: 0.69 ­ 0.89: 0.93 ­ 1.14: 1.02 ­ 1.25. Pronotum with distinct transverse discal impression behind calli, anterior lobe with small patch of punctures behind head, and a few punctures laterally, posterior lobe with punctures forming a transverse band; lateral margins with 4 ­ 6 submarginal setigerous punctures (Fig. 107). Propleuron alutaceous, depression and basal part of posterior convexity with numerous coarse punctures. Scutellum distinctly punctate with numerous punctures in the middle of its disc. Clavus with one complete and two partial rows of punctures; mesocorial disc with small, evenly spaced punctures; exocorium weakly punctate or almost impunctate; costa narrow, almost entirely separated from exocorium, bearing 2 setigerous punctures (Fig. 107). Male hind femora with distinct subapical tooth on dorsal margin, and several smaller teeth on ventral margin (Fig. 109); male hind tibiae with distinct subbasal tubercle, followed by a few more or less visible small denticles (Fig. 109); female hind femora with small subapical tooth on dorsal margin; female hind tibiae simple. Abdominal sterna more or less wrinkled, with triangular patches of punctures posterior to spiracles. Male genitalia as in Figs 110 ­ 113.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B334512FE334257BF07FD09.taxon	materials_examined	TYPE DATA Holotype male: Indonesia, Java, Wynkoopsbay (IRSNB). MATERIAL EXAMINED Holotype, 20 paratypes, and 1 other specimen: Indonesia: Indonesia, Java, Wynkoopsbay, 1 paratype (UO); Java, Ardja­Sari, Preanger, 1 paratype (ZMA); Java, Soekaboemi, 1 paratype (NNML); East Kalimantan, Mt Pemantus, 45 km West of Sotek, 2 paratypes (NSMT). Malaysia: Malay Peninsula, 1 paratype (BMNH); Malay Peninsula, 1 paratype (UO); North Borneo, Sabah, Sandakan Distr., Rumidi, 1 paratype (UO); Sabah, Beaufort, confl. Pa Sia ­ Matang, 3 paratypes (NNML); Nature Trail, Borneo Sabah, Danum Valley, 70 km W Lahat Datu, 1 paratype (ZMA); Sabah, Long Pa Sia, Airstrip along S Pa Sia, 1 paratype (NNML); Sabah, Long Pa Sia, 1 paratype (NNML); Sabah, 10 km SE Ranau, 1 paratype (NNML); Sabah, Sipitang, Mendolong, 1 ex. (MZLU); Borneo, Sarawak, 16 km N of Bario, Long Rapun, 1 paratype (NNML). Philippines: Balabac, Dalawan Bay, 1 paratype (ZMUC); Trident Mines, Central Palawan, 3 paratypes (NSMT, UO).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B334512FE334257BF07FD09.taxon	distribution	DISTRIBUTION Malaysia (Malaya, Sabah, Sarawak), Indonesia (Java, Kalimantan), Philippines.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B314514FE4C4377BA66FBF7.taxon	description	(Figs 114 ­ 120)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B314514FE4C4377BA66FBF7.taxon	description	DESCRIPTION Body 9.05 ­ 12.25 mm in length, 5.01 ­ 6.61 mm in width, from castaneous to blackish brown. Head dorsally weakly punctate; clypeus without submarginal setigerous punctures, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 115); ocular index 1.82 ­ 2.40; interocellar index 7.4 ­ 11.0; length of antennal segments (in mm): 0.41 ­ 0.47: 0.46 ­ 0.72: 0.71 ­ 0.94: 0.95 ­ 1.11: 1.09 ­ 1.26. Pronotum with well developed transverse discal impression behind calli, usually bearing a row of punctures; anterior lobe with distinct punctures subapically and laterally; posterior lobe with irregularly scattered punctures; lateral margins with 5 ­ 6 submarginal setigerous punctures (Fig. 114). Propleuron alutaceous, depression and basal part of posterior convexity densely punctate. Scutellum with numerous irregularly scattered punctures, base and apex impunctate. Corium with punctures smaller than those on scutellum; meso­ and exocorium almost evenly punctate; costa narrow, separated from exocorium in its anterior four fifths, slightly convex, costal margin with 2 setigerous punctures (Fig. 114). Male hind femora with distinct subapical tooth on dorsal margin, and a few teeth on the ventral, tibiae with several subbasal tubercles (Fig. 116); female hind femora with small, sometimes indistinct subapical tooth on dorsal margin, and several setae on the ventral, tibiae without tubercles. Abdominal sterna smooth in the middle, sides distinctly wrinkled and bearing irregularly spaced coarse punctures. Male genitalia as in Figs 117 ­ 120.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B314514FE4C4377BA66FBF7.taxon	materials_examined	TYPE DATA Holotype male: Vietnam, Cho­Ganh (MNHN). MATERIAL EXAMINED Holotype, 1 paratype, and 28 other specimens: Cambodia: Kompong Kedey, 1 ex. (BMNH). China: South China, Haut­Tonkin et Bas Yunnan, Entce Man­Hao, Muong­Hum (pres Lao­Kay) et Ban­Nam­Coun, 1 ex. (MNHN); Yunnan, env. Dadugan, 1 ex. (ZIP). Hong Kong: Hong Kong, 1 ex. (BMNH). India: Kairhana, Assam, 1 ex. (ZIP); Barodabri, 2 exx. (ZlP, UO). Laos: Luang Prabang, Ko Kieng, 3 exx. (BMNH, UO). Malaysia: Malaya, Pahang, Cameron Highlands, 7 exx. (BMNH, UO). Thailand: Loei Province, Phu Luang Wildlife Sanctuary, 5 exx. (ZMUC, UO); 240 km NW Bangkok, 25 km NW Lan­Sak, 1 ex. (NHMW); 170 km NW Bangkok, 3 km W Ban Rain 150 m, 1 ex. (NHMW). Vietnam: Cho­Ganh, 1 paratype (UO); Tonkin, reg. de Hoa­Binh, 3 exx. (MNHN); Sud Vietnam, Bobla, 1 ex. (NSMT).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B314514FE4C4377BA66FBF7.taxon	distribution	DISTRIBUTION Cambodia, China (Yunnan), Hong Kong, India, Laos, Malaya, Vietnam, Thailand.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B374516FE764471B906FBD7.taxon	description	(Figs 121 ­ 125)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B374516FE764471B906FBD7.taxon	description	DESCRIPTION Body 7.10 ­ 9.22 mm in length, 4.26 ­ 5.20 mm in width, castaneous to dark castaneous, sometimes almost black. Head dorsally impunctate or almost impunctate; clypeus without subapical setigerous punctures, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 121); ocular index 2.70 ­ 3.20; interocellar index 5.1 ­ 7.3; length of antennal segments (in mm): 0.34 ­ 0.42: 0.38 ­ 0.52: 0.47 ­ 0.60: 0.55 ­ 0.71: 0.61 ­ 0.76. Pronotum without transverse discal impression behind calli; disc with an arcuate patch of punctures behind head, a broad band of many punctures behind the middle, and numerous punctures antero­laterally; lateral margins with 5 ­ 7 submarginal setigerous punctures. Propleural depression with punctures. Scutellum densely punctate with exception of antero­lateral angles and extreme apex. Corium conspicuously punctate; clavus with one complete and two partial rows of punctures; mesocorial disc densely punctate; exocorium with weaker puncturation; costa narrow, separated from exocorium in its anterior fourth fifths, slightly convex, bearing 2 setigerous punctures. Male hind femora with almost indistinct subbasal tooth on dorsal margin; male hind tibia with very small subbasal tubercle; female hind legs not modified. Abdominal sterna wrinkled laterally, punctures present only behind and sometimes lateral to spiracles. Male genitalia as in Figs 122 ­ 125.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B374516FE764471B906FBD7.taxon	materials_examined	TYPE DATA Lectotype male (designated by JOSIFOV & KERZHNER, 1978: 189) of Macroscytus fraterculus HORVÁTH: China, Peking (HNHM). Holotype male of Macroscytus confusus J. A. LIS: Japan, Honshu, Kanagawa pref., Hayama (ZMA). MATERIAL EXAMINED Lectotype of Macroscytus fraterculus, Holotype of Macroscytus confusus, 7 paratypes of Macroscytus confusus, and 6 other specimens: China: Fokien [Fujian], 3 paratypes (UO); Fukien, Shaowu, 1 ex. (BMH); Fukien, ChungAn, Bohea Hills, 3 exx. (BMH, UO); Peking [Beijing], 1 paratypes (UO). Japan: Honshu, Osaka, Rokkusan, 3 paratypes (UO); Kobe, 1 ex. (BMH); Hekinan, Gyõyõ­jinsha, Gyõyõ­chõ, Nishio city, Aichi Pref., 1 ex. (NSMT).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B374516FE764471B906FBD7.taxon	distribution	DISTRIBUTION China (Fujian, Beijing), Japan.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B354518FE164410BA82FBB6.taxon	description	(Figs 129 ­ 130)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B354518FE164410BA82FBB6.taxon	description	DESCRIPTION Body 7.47 ­ 7.53 mm in length, 4.06 ­ 4.10 mm in width, from dark castaneous to blackish brown, posterior margins of umbones slightly lighter in shade, hemelytra yellowish brown or pale brown. Head dorsally impunctate or with single small punctures on paraclypei; clypeus subapically without a pair of setigerous punctures, each paraclypeus submarginally with 3 ­ 4 setigerous punctures bearing long hair­like setae (Fig. 130); ocular index 2.35 ­ 2.65; interocellar index 7.4 ­ 8.7; length of antennal segments (in mm): 0.24 ­ 0.34: 0.50 ­ 0.53: 0.49 ­ 0.56: 0.67: 0.76. Puncturation of pronotal disc weak, several punctures present behind calli and in the posterior half of disc, discal transverse impression absent; lateral margins of pronotum with 6 ­ 9 submarginal setigerous punctures bearing long hair­like setae (Fig. 129). Anterior convexity of propleuron alutaceous, sparsely punctate with small punctures, posterior convexity distinctly punctate with punctures larger than those on anterior convexity, propleural depression coarsely punctate. Scutellar disc with almost evenly spaced punctures as large as those on pronotum, basal fifth of scutellar disc impunctate. Corium densely punctate; clavus with one complete and two incomplete rows of punctures; mesocorial disc densely and almost evenly punctate; exocorium densely punctate; costa flattened, separated from exocorium only in its basal half, costal margin with 2 setigerous punctures bearing hair­like setae (Fig. 129). Female hind femora and tibiae without denticles and tubercles. Abdominal sterna impunctate, except for rows of punctures close to segmental sutures, and patches of dense puncturation around trichobothria and spiracles.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B354518FE164410BA82FBB6.taxon	materials_examined	TYPE DATA Holotype (by monotypy): Seychelles Islands, Aldabra (BMNH). MATERIAL EXAMINED Holotype, and 1 other specimen: Comoros: Moheli, Lac Iconi, 1 female (MNHN).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B354518FE164410BA82FBB6.taxon	distribution	DISTRIBUTION Comoros (first record), Seychelles.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B354518FE164410BA82FBB6.taxon	discussion	REMARK Abdomen is missing in the holotype specimen, its sex was also not given in the original description.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B3B451AFE594430BA29FD69.taxon	description	(Figs 131 ­ 135)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B3B451AFE594430BA29FD69.taxon	description	DESCRIPTION Body 8.00 ­ 10.78 mm in length, 4.22 ­ 5.49 mm in width, from dark castaneous to almost black, corium and apical part of umbones usually more brownish than scutellum. Head dorsally polished, impunctate, sometimes only slightly wrinkled in lateral parts of paraclypei; clypeus subapically without a pair of setigerous punctures, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 132); ocular index 1.85 ­ 2.45; interocellar index 6.0 ­ 11.0; length of antennal segments (in mm): 0.41 ­ 0.43: 0.47 ­ 0.60: 0.63 ­ 0.85: 0.82 ­ 1.01: 0.94 ­ 1.12. Pronotum divided into two lobes by distinct transverse impression behind calli bearing numerous punctures; lateral margins with 4 ­ 5 submarginal setigerous punctures (Fig. 131). Propleuron with numerous punctures in depression and in basal part of posterior lobe. Scutellum almost evenly punctate, its basal part and apex impunctate. Corium almost evenly punctate with punctures smaller than those on scutellum; clavus with one complete and two partial rows of punctures; exocorium weakly punctate; costa narrow, almost entirely separated from exocorium, bearing 2 setigerous punctures (Fig. 131). Male hind tibiae with small subbasal tubercles, hind femora with distinct subapical teeth (Fig. 133); female hind femora with very small, sometimes indistinct subapical tooth. Abdominal sterna smooth, bearing more or less developed wrinkles, each segment laterally with a triangular patch of dense tiny punctures posterior to spiracles, sterna III and IV with very few tiny punctures also anterior to spiracle; sometimes lateral third of sterna with a few large clearly visible punctures. Male genitalia as in Figs 134 ­ 135.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B3B451AFE594430BA29FD69.taxon	materials_examined	TYPE DATA Syntype (s) of Hahnia gibbula ELLENRIEDER: Sumatra, Lahat (lost or destroyed). Neotype male (present designation) of Hahnia gibbula ELLENRIEDER: Sumatra, Bindjei­Medan (UO). Holotype male of Macroscytus sumatranus LIS: Indonesia, Java, Banjoewangi (ZMA). MATERIAL EXAMINED Neotype of Hahnia gibbula, Holotype of Macroscytus sumatranus, 62 paratypes of Macroscytus sumatranus, and 1 other specimen: Burma: Katha, 2 paratypes (MCSN, ZIZM); Tenasserim, Thagata, 1 paratype (MCSN); Rangoon, 1 paratype (MNHN). China: Nauning, Kwangsi, 2 paratypes (NNML). India: Calcutta, 1 paratype (ZIZM); Gopaldhara, Darjeeling, 1 paratype (BMNH); Tenmalai, Travancore, 1 paratype (BMNH); Barodabri, 2 paratypes (ZIP, UO); Kalimpong, 1 paratype (ZIP). Indonesia: Buitenzorg, Java, 2 paratypes (ZIZM); Java, 1 paratype (NHMW); Sumatra, Tandigeng bij, Moeara Enim, 1 paratype (ZMA); Bengkalis, 1 paratype (MNHN); Timor, 1 paratype (ZMA); Tandjong Morawa, Serdang, NO Sumatra, 3 paratypes (NNML, UO); Gedanga, 2 paratypes (NNML); Palembang, 1 paratype (NNML); Timor, Oesusu, 1 ex. (NSMT). Laos: Luang Prabang, 2 paratypes (BMNH, MNHN); Vientian, 1 paratype (ZIP); Xieng Khouang, Ta Trang, 3 paratypes (BMNH, UO). Malaysia: Pahang, Taman Negara, 3 paratypes (BMNH, UO); Malaya, Kuala Lumpur, 1 paratype (BMNH); Malay Penin., 4 paratypes (BMNH, UO); Malay Penin., West Coast, Langkawi Is., 1 paratype (BMNH); Borneo, E. Malaysia, Sepilok, Sabah, 2 paratypes (NSMT); N. Borneo, Sabah, 10 km SE Ranau, 1 paratype (NNML); Malaysia, Sabah, Long Pa Sia, 1 paratype (NNML); Borneo, Sarawak, trail Pa Lungan ­ Long Rapun, 1 paratype (NNML). Singapore: 1 paratype (BMNH). Thailand: Thailand, 1 paratype (BMNH); Siam, 1 paratype (UO); Siam, Nan, 1 paratype (BMNH). Vietnam: Tonkin Centr., Region de Chim­Hoa et de Tuyen­Quan, 1 paratype (UO); Tonkin, Env. de Hanoi (Demange), 1 paratype (MNHN); Tonkin, reg. de Hoa­Binh, 5 paratypes (MNHN, UO); Saigon, 2 paratypes (MNHN); N­Vietnam, Cuc Phuong, 2 paratypes (NHMW); Annam, Phuc­Son, 3 paratypes (ZIZM).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B3B451AFE594430BA29FD69.taxon	distribution	DISTRIBUTION Burma, China (Guangxi), India, Indonesia, Laos, Malaysia, Singapore, Thailand, Vietnam.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B39451CFE4B4297BA16FD57.taxon	description	(Figs 136 ­ 142)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B39451CFE4B4297BA16FD57.taxon	description	DESCRIPTION Body 8.30 ­ 8.88 mm in length, 4.75 ­ 5.01 mm in width, from blackish brown to black, umbones and posterior margin of pronotum dark castaneous, exocorium and basal part of mesocorium castaneous in shade. Head dorsally polished, almost smooth, bearing a few tiny and hardly visible punctures; clypeus subapically without a pair of setigerous punctures, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 137); ocular index 2.00 ­ 2.15; interocellar index 8.6 ­ 9.7; length of antennal segments (in mm): 0.33 ­ 0.36: 0.49 ­ 0.57: 0.50 ­ 0.60: 0.64 ­ 0.79: 0.64 ­ 0.74. Pronotal disc impunctate, or with a few irregularly scattered punctures; transverse discal impression behind calli absent; lateral margins of pronotum with 5 ­ 6 submarginal setigerous punctures bearing long hair­like setae (Fig. 136). Anterior convexity of propleuron punctate anteriorly and laterally, posterior convexity with a few irregularly scattered punctures, propleural depression with a broad band of coarse punctures. Scutellum with irregular puncturation, punctures well visible in its median part. Mesocorial disc densely and almost evenly punctate; exocorium densely punctate; costa narrow, almost entirely separated from exocorium, bearing a single setigerous puncture (Fig. 136). Femora castaneous or dark castaneous, tibiae basally yellowish brown or brown, getting darker towards tarsi; the latter brown or pale brown; dorsal margin of male hind femur with numerous small denticles (Fig. 138); male hind tibia with a row of small denticles on its inner margin (Fig. 138); inner margins of female hind femur and tibia without denticles. Abdominal sterna polished, punctate close to lateral margins, around trichobothria and posterior to spiracles. Male genitalia as in Figs 139 ­ 142.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B39451CFE4B4297BA16FD57.taxon	materials_examined	TYPE DATA Holotype male: Australia, North Queensland, Mt. Misery Summit (QMB). MATERIAL EXAMINED Holotype, and 3 paratypes: Australia: Queensland, Mt. Boolbun South, 1 paratype (QMB); Queensland, Bellenden Ker Range, Cableway Base Stn., 1 paratype (UO); Queensland, Mt Blackwood, 1 paratype (QMB).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B39451CFE4B4297BA16FD57.taxon	distribution	DISTRIBUTION Australia.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B3F4561FE074291BA77FE29.taxon	description	(Figs 143 ­ 149) DESCRIPTION H e a d. Black, lateral margins brownish in shade; dorsal surface alutaceous, bearing numerous tiny and hardly visible punctures, lateral parts of paraclypei striated and punctate with small punctures; gular plate coarsely punctate in its posterior half, anterior half impunctate; bucculae with a few punctures only; clypeus free, as long as paraclypei, subapically without setigerous punctures, each paraclypeus submarginally with 7 ­ 9 setigerous punctures bearing hair­like setae (Fig. 144); eyes dark reddish brown, ocular index 2.85; ocelli reddish brown, interocellar index about 4.5, ocellar index 7.0; antennae yellowish brown; rostrum pale brown, reaching middle of mesosternum. P r o t h o r a x. Pronotum black, posterior and lateral margins castaneous in shade; puncturation of pronotal disc well visible behind head, in lateral parts and behind calli, transverse discal impression shallow, bearing a row of punctures (Fig. 143); lateral margins of pronotum with 10 ­ 11 setigerous punctures bearing long hair­like setae (Fig. 143). Prosternal carinae low, coarsely punctate; anterior convexity of propleuron with a patch of punctures close to head margin, posterior convexity punctate with a few coarse punctures, propleural depression with a row of several coarse punctures. Mesothorax. Scutellum black, its apex castaneous; basal and apical parts of scutellar disc impunctate, remaining surface densely punctate with numerous punctures larger than those on pronotum (Fig. 143). Mesopleural evaporatorium with longitudinal rugae, its posterior wedge­shaped polished band impunctate. Metathorax. Hemelytra dark castaneous, distinctly punctate; clavus with one complete and two partial rows of punctures; mesocorial disc almost evenly punctate, punctures slightly smaller than those on scutellum; exocorium with puncturation more dense than those of mesocorium; mesocorium and exocorium separated almost along their entire length; costa basally broad, almost entirely separated from exocorium, costal margin with 6 setigerous punctures bearing long hair­like setae (Fig. 143); membranal suture almost straight; membrane semihyaline, milky brown, its tip clearly surpassing the posterior extremity of abdomen. Metapleural evaporatorium without longitudinal rugae; lateral area of metapleuron impunctate, posterior area with a band of coarse punctures. L e g s. Femora and tibiae dark castaneous, tarsi yellowish brown; dorsal margin of mid femur with a row of 10 ­ 11 sharp black teeth; hind femur basally with a row of tubercles bearing long setae, subapical denticles and teeth absent; hind tibia subbasally with large tooth (Fig. 147). Abdomen. Sterna blackish brown, laterally distinctly striated and coarsely punctate, segmental sutures accompanied by rows of punctures. Ventral surface of male pygophore with small punctures, dorsal surface pilose, longitudinal sclerites on infolding of its dorsal rim well developed; penis as in Fig. 148, second conjunctival appendage as in Fig. 149, paramere as in Fig. 146, opening of genital capsule as in Fig. 145. M e a s u r e m e n t s (in mm). Male only: body length 7.32; body width 4.45; head length 1.21; head width 1.92; pronotum length 2.34; pronotum width 4.12; scutellum length 2.83; scutellum width 2.77; antennal segments: 0.35: 0.47: 0.36: 0.44: 0.47.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B3F4561FE074291BA77FE29.taxon	materials_examined	TYPE MATERIAL EXAMINED Holotype male: Kenya: N­Kenia, Umg. Nairobi, VII 1981, HEISS, collection E. HEISS, Innsbruck (TLM).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B3F4561FE074291BA77FE29.taxon	discussion	COMPARATIVE NOTES This new Afrotropical species is very similar to Oriental M. nigroaeneus and Australian M. piceus, especially in its general body outline, its vestiture and puncturation. Nevertheless, it can easily be separated from the two species by characters given in the key.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B3F4561FE074291BA77FE29.taxon	distribution	DISTRIBUTION Kenya.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B424563FE35424DB9AEFE29.taxon	description	(Figs 150 ­ 156)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B424563FE35424DB9AEFE29.taxon	description	DESCRIPTION Body 7.95 ­ 8.67 mm in length, 4.16 ­ 5.10 mm in width, from dark castaneous to black brown; umbones and posterior margin of pronotum with narrow yellowish brown transverse band; exocorium, clavus and basal part of mesocorium yellow or yellowish brown, the remaining part of mesocorium castaneous or dark castaneous. Head dorsally polished, sometimes slightly wrinkled and with several small punctures laterally; clypeus without subapical pair of setigerous punctures, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 151); ocular index 2.30 ­ 2.60; interocellar index 4.9 ­ 6.3; length of antennal segments (in mm); 0.32 ­ 0.40: 0.42 ­ 0.46: 0.58 ­ 0.66: 0.70 ­ 0.81: 0.84 ­ 0.94. Pronotum without transverse discal impression behind calli, pronotal disc with several scattered punctures; lateral margins with 5 ­ 6 submarginal setigerous punctures (Fig. 150). Propleuron with numerous distinct punctures. Scutellum with several punctures in the middle of its disc. Clavus with one complete and two partial rows of punctures; mesocorium in its basal third punctate with large punctures, remaining two thirds with smaller punctures; exocorium with puncturation denser than that of mesocorium; costa narrow, almost entirely separated from exocorium, costal margin without (Fig. 150) or very rarely with a single setigerous puncture. Femur from yellowish brown to yellow, bearing dark median stripe; male hind femora with small, sometimes indistinct, subapical spines (Fig. 152); male hind tibia with a few small denticles (Fig. 152). Abdominal sterna glossy, each with a triangular patch of dense fine punctures posterior to spiracle; areas anterior to and mesad of spiracles with a few punctures; lateral parts of segments additionally irregularly wrinkled. Male genitalia as in Figs 153 ­ 156.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B424563FE35424DB9AEFE29.taxon	materials_examined	TYPE DATA Holotype male: Solomon Islands, Guadalcanal I., Honiara (BMNH). MATERIAL EXAMINED Holotype, 4 paratypes, and 4 other specimens: Bismarck Archipelago: New Britain, Volupai, 1 ex. (BMH). Solomon Islands: Guadalcanal I., Honiara, 1 paratype (BMNH); Tulagi, 2 paratypes (BMNH, UO); Guadalcanal, en route Gallego camp, 1 paratype (BMNH); Bougainville, Kukugai Vill., 3 exx. (BMH, UO).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B424563FE35424DB9AEFE29.taxon	distribution	DISTRIBUTION New Britain (first record), Solomon Islands.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B404565FE174257BA8CFDA8.taxon	description	(Figs 157 ­ 163)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B404565FE174257BA8CFDA8.taxon	description	DESCRIPTION Body 7.62 ­ 8.42 mm in length, 4.60 ­ 5.21 mm in width, from dark castaneous to black, apices of umbones and the posterior margin of pronotum narrowly yellowish brown (in dark castaneous specimens only the apices of umbones are slightly brightened), hemelytra usually more brownish in shade than pronotum and scutellum. Head dorsally impunctate; clypeus subapically without setigerous punctures, each paraclypeus with a row of 3 ­ 6 submarginal setigerous punctures bearing long hair­like setae (Fig. 158); ocular index 2.00 ­ 2.60; interocellar index 5.0 ­ 6.0; length of antennal segments (in mm): 0.29 ­ 0.32: 0.42 ­ 0.56: 0.49 ­ 0.58: 0.60 ­ 0.66: 0.62 ­ 0.72. Pronotal disc with a few small, hardly visible punctures; posterior half of pronotal disc with tiny and almost invisible transverse wrinkles; lateral margins with 7 ­ 10 submarginal setigerous punctures bearing long hair­like setae (Fig. 157). Propleural depression with a row of coarse punctures, anterior and posterior convexities of propleuron with few small punctures. Scutellar disc with small, almost invisible punctures, and a few tiny transverse wrinkles. Corium with numerous irregular wrinkles, at first sight looking almost impunctate; mesocorial disc entirely impunctate or sparsely punctate in its posterior half; exocorium broad, expanded laterally, impunctate; costa narrow, flat, almost entirely separated from exocorium, its posterior fifth fused with the latter, costal margin with 2 ­ 4 setigerous punctures bearing long hair­like setae (Fig. 157). Femora dark brown or brown, tibiae usually paler than femora; male and female hind femora simple, spines and teeth absent (Fig. 159); male and female tibiae without denticles on inner margins (Fig. 159). Abdominal sterna almost impunctate, bearing only rows of small punctures close to segmental sutures, and triangular patches of well visible punctures posterior to spiracles. Male genitalia as in Figs 160 ­ 163.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B404565FE174257BA8CFDA8.taxon	materials_examined	TYPE DATA Holotype male: Australia, Lord Howe Island, the Clear Place (QMB). MATERIAL EXAMINED Holotype, and 4 paratypes: Lord Howe Island: Stevens Reserve, 1 paratype (QMB); The Clear Place, 1 paratype (QMB); Lorhiti Lodge, 1 paratype (QMB); Base of Mt Eliza, 1 paratype (UO).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B404565FE174257BA8CFDA8.taxon	distribution	DISTRIBUTION Lord Howe Island.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B464567FE1542D6BA15FAB7.taxon	description	(Figs 164 ­ 170) DESCRIPTION H e a d. Castaneous, lateral parts of paraclypei somewhat darker in shade; dorsal surface alutaceous, impunctate, bearing only a few punctures laterally; gular plate almost entirely coarsely punctate, except for its anterior fifth; bucculae entirely coarsely punctate; clypeus free, as long as paraclypei, subapically without setigerous punctures, each paraclypeus submarginally with two setigerous punctures bearing hair­like setae (Fig. 165); eyes dark brown, ocular index 2.40; ocelli reddish brown, distance between ocelli about 7.5 times longer than the distance between ocellus and eye, ocellar index 6.0; antennae pale brown; rostrum yellowish brown, reaching mid coxae. P r o t h o r a x. Pronotum castaneous; pronotal disc with a row of small punctures behind head, and a band of large punctures behind calli, the latter impunctate and elevated; transverse discal impression deep, well developed, bearing row of coarse punctures interrupted medially; lateral margins of pronotum with 5 submarginal setigerous punctures bearing long hair­like setae (Fig. 164). Prosternal carinae low, punctate; anterior convexity of propleuron almost entirely punctate with numerous small punctures, posterior convexity of propleuron punctate with large coarse punctures, propleural depression with a band of coarse punctures. Mesothorax. Scutellum castaneous, basal fourth of its disc and scutellar apex impunctate, its remaining surface punctate with numerous punctures larger than those on pronotum. Mesopleural evaporatorium with distinct longitudinal rugae, its posterior wedge­shaped polished band with over a dozen large punctures. Metathorax. Hemelytra castaneous, puncturation colourless; clavus with one complete and two incomplete rows of punctures; mesocorial disc almost evenly punctate, punctures smaller than those on scutellum; puncturation of exocorium dense; mesocorium and exocorium separated almost along their entire length; costa broad, flattened, separated from exocorium only in its basal half, costal margin with two setigerous punctures bearing long hair­like setae (Fig. 164); membranal suture almost straight; membrane semihyaline, somewhat embrowned, its tip slightly surpassing the posterior extremity of abdomen. Metapleural evaporatorium with longitudinal rugae; lateral area of metapleuron impunctate, except for a band of punctures close to evaporatorium margin, posterior area with a band of coarse punctures. L e g s. Femora and tibiae castaneous, tarsi pale brown; ventral margin of hind femur with a few denticles, dorsal margin with 2 ­ 3 subapical small teeth (Fig. 166); hind tibia with several small tubercles (Fig. 166). Abdomen. Sterna castaneous, distinctly punctate in lateral two thirds, punctures around spiracles and trichobothria clearly smaller than remaining ones. Ventral surface of male pygophore impunctate, its dorsal surface pilose, longitudinal sclerites on infolding of its dorsal rim well developed; penis as in Fig. 169, second conjunctival appendages as in Fig. 170, paramere as in Fig. 168, opening of genital capsule as in Fig. 167. M e a s u r e m e n t s (in mm). Male only: body length 7.92; body width 4.55; head length 1.35; head width 1.99; pronotum length 2.02; pronotum width 4.02; scutellum length 2.95; scutellum width 2.61; antennal segments: 0.35: 0.56: 0.57: 0.75, 5 th segment missing.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B464567FE1542D6BA15FAB7.taxon	materials_examined	TYPE MATERIAL EXAMINED Holotype male: Comoros: Moroni, Gde Comore, Malet, Coll. E. Fleutiaux, 1919, Muséum Paris (MNHN).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B464567FE1542D6BA15FAB7.taxon	discussion	COMPARATIVE NOTES The new species is similar to M. fryeri, M. mayottensis and M. comorensis, but it can easily be distinguished from all of them by deep and well developed transverse impression on its pronotal disc (transverse impression is shallow or absent in the other three species), and by its almost entirely punctate gular plate of head (it is punctate only in posterior half or third in the other three species).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B464567FE1542D6BA15FAB7.taxon	distribution	DISTRIBUTION Comoros.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B444569FE24452FB987FC09.taxon	description	(Figs 126 ­ 128)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B444569FE24452FB987FC09.taxon	description	DESCRIPTION Body 8.10 ­ 10.5 mm in length, 4.65 ­ 6.00 mm in width, from castaneous to almost black, umbones and posterior margin of pronotum sometimes slightly paler than the disc. Head almost impunctate, bearing tiny punctures, lateral parts sometimes wrinkled and with a few punctures; clypeus without a subapical pair of hair­like setae, each paraclypeus submarginally with a single preocular setigerous puncture bearing hair­like seta (Fig. 126); ocular index 1.98 ­ 3.00; interocellar index 5.5 ­ 9.1; length of antennal segments (in mm): 0.29 ­ 0.46: 0.43 ­ 0.63: 0.51 ­ 0.69: 0.69 ­ 0.88: 0.72 ­ 0.94. Pronotum with transverse impression behind calli absent or only slightly marked on either side and interrupted medially; pronotal disc with punctures behind head, in lateral parts and behind calli; lateral margins with 5 ­ 6 submarginal setigerous punctures. Anterior convexity of propleuron moderately punctate, posterior convexity with large punctures, propleural depression with a row of coarse punctures. Scutellum evenly punctate, punctures slightly larger than those on pronotum, apex and basal part impunctate. Clavus with one complete and two or three partial rows of punctures; exocorium and middle part of mesocorium almost evenly densely punctate, punctures smaller than those on scutellum; costa almost entirely separated from exocorium, bearing 2 setigerous punctures. Male hind tibiae slightly carinate, and subbasally with distinct small tubercles, hind femora with several small teeth on ventral margin and one subapical tooth on the dorsal; female hind femora without teeth, hind tibiae without tubercles. Abdominal sterna with punctures forming patches posterior to spiracles, sutures and surface laterally of spiracles with a few punctures. Male genitalia as in Figs 127 ­ 128.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B444569FE24452FB987FC09.taxon	materials_examined	TYPE DATA Lectotype female (designated by LIS, 1994 a: 222) of Macroscytus japonensis SCOTT: Japan (BMNH). Lectotype female (designated by LIS, 1994 a: 222) of Macroscytus niponensis SIGNORET: Japan (NNML). MATERIAL EXAMINED Lectotype of Macroscytus japonensis, Lectotype of Macroscytus niponensis, and 57 other specimens: Burma: North Burma, Adung Valley, 1 ex. (BMNH). China: North China, 1 ex. (BMNH); Gan Chouen Fou, Anshunfu, 2 exx. (MNHN, UO); Peking, 3 exx. (MNHU); Kiautschou, Tsingtau, 1 ex. (MNHU); Chekiang, Hangtcheou [= Hangzhou], 1 ex. (MNHN); Chen­Si, 3 exx. (MNHN, UO); Chenfu­Wu, 1 ex. (BMNH); Shanghai, 2 exx. (NNML); Sichuan, Ulian, Utschan­Tschou, 1 ex. (ZIP); Kouy­Tcheou, Kouy Yang, 1 ex. (MNHN); Central China, 1 ex. (NNML); Kansu, Chengshien, 1 ex. (NNML); Nanning Kwangtung, 1 ex. (NNML); 35 km NW Peking, Fragrant Hills, 1 ex. (NHMW); Taiwan, 1 ex. (BMNH); Taiwan, Taihanroku, 1 ex. (BMNH); Taiwan, Teraso, 3 exx. (HNHM); Taiwan, Kaohsiung, Shanping, 1 ex. (CMNH). Japan: 1 ex. (BMNH); 2 exx. (MNHU); Honshu, Miura, Kanagawa, 3 exx. (NSMT, UO); Ryukyus, Funaura, Iriomote I., 3 exx. (NSMT, UO); Shikoku, Mt. Nakatsuno­mine Tokushima, 4 exx. (NSMT, UO). Korea: East Korea, Prov. Kanwan, Kum­gang­son, 4 exx. (HNHM, UO); Sarivon, 1 ex. (HNHM); South Korea, Song Panaku, Halla San, Cheju Do Is., 2 exx. (NSMT). Vietnam: Tonkin Centr., Env. de Yen­Bai, 1 ex. (MNHN); Tonkin, Reg. de Yen­Bay, 1 ex. (MNHN); Tonkin, Hoa­Binh, 1 ex. (MNHN); Haut­Tonkin, Lao­Kay, 1 ex. (MNHN); Pr. Annam, Than Hoa, 1 ex. (IRSNB); Tonkin, Phu Lang Huong, 1 ex. (MNHN); Hanoi, 4 exx. (NhMW, BMNH, UO).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B444569FE24452FB987FC09.taxon	distribution	DISTRIBUTION Burma (north), Korea, Japan, China (Beijing, Fujian, Gansu, Guangdong, Guizhou, Henan, Hubei, Hunan, Shandong, Shanghai, Shanxi, Sichuan, Taiwan, Zhejiang), Russia (Far East), Vietnam (north).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B4A456AFE164477BA84FDC9.taxon	description	(Figs 171 ­ 177)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B4A456AFE164477BA84FDC9.taxon	description	DESCRIPTION Body 7.30 ­ 9.15 mm in length, 3.85 ­ 5.01 mm in width, from castaneous to blackish brown, corium usually slightly paler than pronotum and scutellum, posterior margin of pronotum sometimes light brown or yellowish brown. Head dorsally impunctate, or with a few tiny punctures, slightly wrinkled in lateral parts of paraclypei; clypeus without subapical setigerous punctures, each paraclypeus with 3 ­ 4 submarginal setigerous punctures (Fig. 172); ocular index 2.00 ­ 2.40; interocellar index 7.5 ­ 11.5; length of antennal segments (in mm): 0.33 ­ 0.42: 0.38 ­ 0.49: 0.56 ­ 0.63: 0.65 ­ 0.81: 0.72 ­ 0.85. Pronotum without or sometimes with slightly developed transverse impression behind calli; pronotal disc with dense puncturation laterally and behind impunctate calli; lateral margins with 5 ­ 8 submarginal setigerous punctures (Fig. 171). Anterior convexity of propleuron with numerous small punctures, posterior convexity with large punctures, propleural depression coarsely punctate. Scutellum densely punctate, with exception of anterolateral angles and apex. Corium distinctly punctate; clavus with one complete and two partial rows of punctures, basal part sometimes with a patch of small punctures; mesocorial disc with distinctly separated punctures smaller than those on scutellum; costa moderately convex, almost entirely separated from exocorium, bearing 2 ­ 3 (rarely 4) setigerous punctures (Fig. 171). Male hind femora with a distinct subapical tooth on dorsal margin, and a few smaller teeth on the ventral (Fig. 173); male hind tibiae carinate and with distinct subbasal triangular tubercle (Fig. 173); female hind femora with subapical tooth on dorsal margin, female hind tibiae without subbasal tubercle. Abdominal sterna III and IV with small punctures anterior and posterior to spiracles, sterna V­VII with punctures posterior to spiracles only; single small punctures sometimes present also laterally to spiracles. Male genitalia as in Figs 174 ­ 177.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B4A456AFE164477BA84FDC9.taxon	materials_examined	TYPE DATA Lectotype male (designated by LIS, 1994 a: 240): Indonesia, Java (NHMW). MATERIAL EXAMINED Lectotype, and 94 other specimens: Indonesia: Sumatra, 1 ex. (NNML); Tandjong Morawa, Serdang, N. O. Sumatra, 1 ex. (NNML); Sumatra, Fort de Kock, 2 exx. (NNML); Sumatra, Labuan, 2 exx. (ZMHU); Sumatra, Medan, Doloc Baroe, 1 ex. (IRSNB); Sumatra, Bindjel­Medan, 4 exx. (ZMA, UO); Sumatra, Palembang, 3 exx. (ZMA, NNML); Sumatra, Padang, 4 exx. (ZMA, MNHN, NNML); Indrapura­Estate, Sumatra, 1 ex. (ZIZM); Sumatra, Dolok Merangir, 7 exx. (NNML, AMNH, UO); Sumatra, Lahat, 5 exx. (NNML); Sumatra, Malakoni, Insel Engano, 2 exx., (ZIZM); Java, 15 exx. (ZMHU, MNHN, HNHM, NNML); Java, Malang, 1 ex. (NNML); Java, Batavia, 1 ex. (MNHN); Java, Soekaboemi, 2 exx. (NNML); Java, Wellri Plellen, Siloewok Sawangan, 4 exx. (ZMA, UO); Java, Banjoewangi, 4 exx. (ZMA, ZIP, UO); Java, Ardja­Sari, Preanger, 1 ex. (ZMA); Java, Slawi Tegal, 1 ex. (ZMA); Java, Soerabaya, 7 ex. (ZMA, NNML); Java, Djakjarta, 1 ex. (ZMA); Java, Buitenzorg, 2 exx. (ZMA, NNML); Java, Mons Gede, 1 ex. (MNHN); Java, Mts Kawie, 1 ex. (ZIP); Java, Surabaja, 1 ex. (ZIP); Java, Blavan, Bondowoso, 1 ex. (ZIP); Java, Blawan, Indjen­Plateau, 2 exx. (ZIP, UO); Java, Bogor, 3 exx. (BMNH, NNML); Java, Temangoeng, 1 ex. (ZMA); Java, Garoet, Preanger, 2 exx. (NNML); Java, Lembang, 1 ex. (JEC); Celebes, Minahassa, Tomohon, 1 ex. (UO); S. Lombok, Ekas, 1 ex. (MNHU); O. Soembawa, Dompoe, 1 ex. (MNHU); Adonara, Solor lsl., 1 ex. (BMNH). Malaysia: Malay Peninsula, Kuala Lumpur, 3 exx. (BMNH, UO). Singapore: Singapore, 1 ex. (ZIZM). Thailand: Thailand, Khao Sok National Park, Phanom Surat Thani, 1 ex. (NSMT).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B4A456AFE164477BA84FDC9.taxon	distribution	DISTRIBUTION Indonesia (Sumatra, Java, Lombok — first record, Engano, Solor, Sumbawa — first record, Sulawesi — first record), Malaysia (Malaya), Singapore, Thailand (first record). Records from China and Japan (SIGNORET, 1881, 1883 a) related to M. japonensis (see LIS, 1994 a).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B49456DFE384237BA87FDC8.taxon	description	(Figs 178 ­ 181)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B49456DFE384237BA87FDC8.taxon	description	DESCRIPTION Body 7.98 ­ 8.02 mm in length, 4.52 ­ 4.67 mm in width, blackish brown; umbones and extreme posterior margin of pronotum yellowish brown; corium brown, clearly paler than pronotum and scutellum. Head dorsally with very small, almost invisible punctures and several rugae; clypeus subapically without setigerous punctures, each paraclypeus with two submarginal setigerous punctures (Fig. 178); ocular index 2.51 ­ 2.55; interocellar index 3.0 ­ 5.0; length of antennal segments (in mm): 0.35 ­ 0.42: 0.49 ­ 0.52: 0.50 ­ 0.53: 0.63 ­ 0.74: 0.84 ­ 0.92. Pronotum undivided into lobes; calli and the posterior quarter impunctate, remaining surface sparsely punctate with clearly visible punctures; lateral margins with 4 submarginal setigerous punctures bearing long hair­like setae (Fig. 179). Apical part of anterior convexity of propleuron dulled by numerous small punctures; depression and basal part of posterior convexity with large coarse punctures. Scutellar disc weakly punctate, bearing a median patch of few punctures (Fig. 179). Clavus with two rows of punctures (one complete, one partial); mesocorial disc evenly moderately punctate; exocorium with very dense puncturation; costa narrow, moderately convex, entirely separated from exocorium, costal margin without setigerous punctures. Legs yellowish brown or pale brown, femora with broad median dark stripe, apical half of anterior tibiae dark; dorsal margins of hind femora with several small teeth. Abdominal sterna medially smooth, lateral parts of each segment with a triangular patch of small punctures posterior to spiracle. Male genitalia as in Figs 180 ­ 181.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B49456DFE384237BA87FDC8.taxon	materials_examined	TYPE DATA Holotype male: Papua New Guinea, Madang Prov., Nagada Harbour (OXUM). MATERIAL EXAMINED Holotype, and 1 paratype: Papua New Guinea: Madang Prov., Nagada Harbour, paratype (OXUM).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B49456DFE384237BA87FDC8.taxon	distribution	DISTRIBUTION Papua New Guinea.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B4F456CFE2740A8B950F9A1.taxon	description	(Figs 182 ­ 188)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B4F456CFE2740A8B950F9A1.taxon	description	DESCRIPTION Body 11.32 ­ 11.61 mm in length, 6.43 ­ 6.63 mm in width, from dark castaneous to almost black. Head dorsally at first sight looking impunctate, bearing small hardly visible irregularly scattered punctures, lateral parts of paraclypei slightly wrinkled, sometimes with a few large punctures; clypeus subapically without a pair of setigerous punctures, each paraclypeus only with 2 primary setigerous punctures, submarginal setigerous punctures absent (Fig. 183); ocular index 2.30 ­ 2.45; interocellar index 3.5 ­ 5.7; length of antennal segments (in mm): 0.51 ­ 0.55: 0.88 ­ 1.01: 0.78 ­ 0.93: 1.15, 5 th segment missing. Pronotum with clearly visible puncturation, especially behind calli, punctures large and coarse; transverse discal impression behind calli well developed, deep, bearing large coarse, crowded punctures; lateral margins of pronotum without submarginal setigerous punctures, primary setigerous punctures on pronotal disc also absent (Fig. 182). Anterior and posterior convexities of propleuron coarsely punctate, punctures on the posterior convexity larger than those on the anterior, propleural depression with numerous coarse and deep punctures. Scutellar disc densely coarsely punctate, punctures almost of the same size as those on pronotum, antero­lateral parts and apex impunctate. Clavus with one almost indistinct and two well visible rows of punctures, its basal part with punctures between rows; mesocorial disc densely and almost evenly punctate, punctures smaller than those on scutellum; exocorium broad, densely and evenly punctate; costa broad, almost totally flatt, separated from exocorium only basally, costal margins punctate, but without setigerous punctures (Fig. 182). Dorsal margin of male and female hind femur with a row of small, usually dark denticles (Fig. 184); male hind tibia with a few small denticles on its inner margin (Fig. 184); female hind tibia without denticles on the inner margin. Abdominal sterna distinctly coarsely punctate in lateral two thirds. Male genitalia as in Figs 185 ­ 188.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B4F456CFE2740A8B950F9A1.taxon	materials_examined	TYPE DATA Lectotype male (designated by LIS, 1999 d): Bourbon (NHMW). MATERIAL EXAMINED Lectotype, and 2 other specimens: Réunion: La Réunion, Ile Bourbon, 2 exx. (MNHN, UO).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B4F456CFE2740A8B950F9A1.taxon	distribution	DISTRIBUTION Réunion (Bourbon Island).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B4D4570FE0540A8BA80FBF7.taxon	description	(Figs 189 ­ 195) DESCRIPTION H e a d. Black, dorsal surface alutaceous, impunctate, except for setigerous punctures; gular plate impunctate in its anterior half, the posterior part coarsely punctate; bucculae coarsely punctate; clypeus free, as long as paraclypei, subapically without setigerous punctures, each paraclypeus submarginally with 2 ­ 3 setigerous punctures bearing hair­like setae (Fig. 190); eyes brown, small, ocular index 3.70 ­ 4.35; ocelli brown, small, interocellar index 3.8 ­ 4.2, ocellar index 8.4 ­ 8.6; antennae brown or dark brown, apical part of 4 th and entire 5 th segment yellowish brown; rostrum brown or dark brown, surpassing mid coxae. P r o t h o r a x. Pronotum black; its disc polished, impunctate, except for setigerous punctures and numerous small, hardly visible punctures; transverse discal impression behind calli absent; lateral margins of pronotum with 4 ­ 5 submarginal setigerous punctures bearing long hair­like setae (Fig. 189). Prosternal carinae low, weakly punctate, but coarsely striated; anterior convexity of propleuron alutaceous with small punctures apically; posterior convexity and propleural depression with coarse punctures. Mesothorax. Scutellum black, its disc at first looking sight impunctate, bearing numerous hardly visible tiny punctures, and a few large punctures close to apex. Mesopleural evaporatorium with coarse longitudinal rugae, its posterior wedge­shaped polished band slightly dulled, impunctate. Metathorax. Hemelytra almost black, meso­ and exocorium blackish brown; clavus with one complete and two partial short rows of punctures; mesocorial disc sparsely punctate with small punctures; exocorium densely punctate, punctures larger than those on mesocorium; mesocorium and exocorium separated along their entire length; costa narrow, convex, almost entirely separated from exocorium, costal margin punctate, but without setigerous punctures (Fig. 189); membranal suture almost straight; membrane semihyalnie, embrowned, its tip clearly surpassing the posterior extremity of abdomen. Metapleural evaporatorium with longitudinal rugae; lateral area of metapleuron impunctate, except for a small patch of punctures close to evaporatorium margin; posterior area with a row of coarse punctures. L e g s. Femora and tibiae dark castaneous, tarsi brown; male hind femur with a row of setae on the ventral margin, and a row of very small denticles on the dorsal margin, subapical tooth absent (Fig. 191); male hind tibiae simple, without denticles and tubercles (Fig. 191). Abdomen. Sterna almost black, distinctly punctate with large and coarse punctures only around spiracles and along segmental sutures. Ventral surface of male pygophore impunctate, its dorsal surface pilose, longitudinal sclerites on infolding of its dorsal rim well developed; penis as in Fig. 194, second conjunctival appendages as in Fig. 195, paramere as in Fig. 193, opening of genital capsule as in Fig. 192. M e a s u r e m e n t s (in mm). Males only: body length 7.37 ­ 7.68; body width 4.01 ­ 4.11; head length 1.14 ­ 1.17; head width 1.71 ­ 1.82; pronotum length 2.19 ­ 2.24; pronotum width 3.65 ­ 3.68; scutellum length 2.68 ­ 2.71; scutellum width 2.22 ­ 2.32; antennal segments: 0.32 ­ 0.33: 0.40 ­ 0.46: 0.47 ­ 0.49: 0.65 ­ 0.67: 0.76 ­ 0.79.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B4D4570FE0540A8BA80FBF7.taxon	materials_examined	TYPE DATA Holotype male: Papua New Guinea, New Guinea (NE), Wau, 8. IX. 1968, NG­W. R. 19, leg. Dr. J. Loksa (HNHM). MATERIAL EXAMINED Holotype, and 1 paratype: Papua New Guinea: New Guinea (NE), Wau, 8. IX. 1968, NG­W. R. 19, leg. Dr. J. Loksa (HNHM).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B4D4570FE0540A8BA80FBF7.taxon	discussion	COMPARATIVE NOTES M. loksai is at first glance similar to another New Guinean species, M. papuanus, especially in its body puncturation and the absence of setigerous punctures on costal margins. Nevertheless, the new species can easily be separated from the latter by a number of submarginal setigerous punctures on head submargins (2 ­ 3 on each paraclypeus in M. loksai, only 1 in M. papuanus). Both species differ also in a shape of paramere and 2 nd conjunctival appendages of penis.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B4D4570FE0540A8BA80FBF7.taxon	distribution	DISTRIBUTION Papua New Guinea.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B534572FE7B4470BA36FB56.taxon	description	(Figs 196 ­ 202)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B534572FE7B4470BA36FB56.taxon	description	DESCRIPTION Body 11.51 ­ 12.68 in length, 5.79 ­ 6.73 in width, from castaneous to almost black, hemelytra paler than pronotum and scutellum, exocorium usually yellowish brown. Head dorsally impunctate; clypeus subapically without a pair of setigerous punctures, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 197); ocular index 2.50 ­ 2.90; interocellar index 4.0 ­ 6.0; length of antennal segments (in mm): 0.44 ­ 0.50: 0.72 ­ 0.83: 0.73 ­ 0.87: 0.97 ­ 1.14: 1.18 ­ 1.32. Pronotal disc impunctate, except for a row or band of large coarse punctures behind calli; transverse discal impression very shallow or absent; lateral margins of pronotum with 3 ­ 4 submarginal setigerous punctures bearing long hair­like setae (Fig. 196). Anterior convexity of propleuron alutaceous, coarsely punctate anteriorly and laterally; posterior convexity alutaceous, with irregularly scattered large punctures; propleural depression with a broad band of large, coarse and deep punctures. Puncturation of scutellar disc dense, punctures larger than those on pronotum, basal fifth and extreme apex of scutellar disc impunctate. Clavus with one complete and two partial rows of punctures; mesocorial disc densely and evenly punctate, punctures distinctly smaller than those on scutellum; exocorium densely punctate, punctures slightly smaller than those on mesocorial disc; costa narrow, separated from exocorium only basally, costal margins punctate, but without setigerous punctures (Fig. 196). Ventral margin of male hind femur with a row of small denticles bearing short setae, and with large subapical tooth, its dorsal margin with a row of small denticles (Fig. 198); male hind tibia carinate on its inner margin (Fig. 198); ventral and dorsal margin of female hind femur with a row of short setae; female hind tibia without carina on its inner margin. Abdominal sterna polished, punctate in lateral thirds, around trichobothria and spiracles. Male genitalia as in Figs 199 ­ 202.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B534572FE7B4470BA36FB56.taxon	materials_examined	TYPE DATA Lectotype male (designated by LIS, 1999 d): Madagascar (NHMW). MATERIAL EXAMINED Lectotype, and 7 other specimens: Madagascar: 2 exx. (MNHN); envir. de Perinet, 2 exx. (MNHN, UO); Sambirano, Manongarivo, 1 ex. (MNHN); Lac Andramdse, 1 ex. (MNHN); massif du Marojejy, 1 ex. (MNHN).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B534572FE7B4470BA36FB56.taxon	distribution	DISTRIBUTION Madagascar.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B514574FE294490BAEDFB96.taxon	description	(Figs 203 ­ 209)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B514574FE294490BAEDFB96.taxon	description	DESCRIPTION Body 6.63 ­ 6.88 mm in length, 3.78 ­ 3.96 mm in width, from dark brown to blackish brown, corium usually brown or dark brown exocorium, clavus and apices of umbones; sometimes pale brown; dorsal surface generally weakly punctate. Head dorsally at first sight looking impunctate, but bearing hardly visible tiny punctures; clypeus subapically without setigerous punctures, each paraclypeus with a row of 3 ­ 4 submarginal setigerous punctures bearing long hair­like setae (Fig. 204); ocular index 2.40 ­ 2.65; interocellar index 5.0 ­ 6.5; length of antennal segments (in mm): 0.25 ­ 0.35: 0.39 ­ 0.42: 0.40 ­ 0.46: 0.53 ­ 0.57: 0.59 ­ 0.65. Pronotum without transverse discal impression behind calli; pronotal disc weakly punctate; lateral margins with 6 ­ 9 submarginal setigerous punctures (Fig. 203). Propleuron polished, depression with numerous coarse punctures. Scutellum weakly punctate with large clearly visible punctures. Clavus with one complete and two partial rows of punctures; mesocorial disc with evenly spaced punctures smaller than those on scutellum; costa convex, entirely separated from exocorium, bearing a single setigerous puncture (Fig. 203). Male hind femora with small, blunt subapical denticle, male hind tibiae only slightly carinate (Fig. 205); female hind femora and tibiae simple. Abdominal sterna polished, almost totally impunctate, except for surface close to spiracles bearing irregular patches of tiny punctures accompanied by numerous wrinkles and striae. Male genitalia as in Figs 206 ­ 209.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B514574FE294490BAEDFB96.taxon	materials_examined	TYPE DATA Holotype female: New Caledonia, Tiwaka, Poindimié (MNHN). MATERIAL EXAMINED Holotype, 2 paratypes, and 2 other specimens: New Caledonia: env. Poindimié, 2 paratypes (MNHN, UO); La Crouen, 1 ex. (BMH); Grotte de Ninrin­Reu nr. Poya, 1 ex. (SAM).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B514574FE294490BAEDFB96.taxon	distribution	DISTRIBUTION New Caledonia.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B574576FE3544D0BA15FB56.taxon	description	(Figs 210 ­ 216)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B574576FE3544D0BA15FB56.taxon	description	DESCRIPTION Body 7.80 ­ 9.37 mm in length, 4.55 ­ 5.15 mm in width, from dark castaneous to almost black. Head dorsally at first sight looking impunctate, but bearing a few small punctures in lateral parts; clypeus subapically without a pair of setigerous punctures, each paraclypeus submarginally with 2 ­ 3 setigerous punctures bearing long hair­like setae (Fig. 211); ocular index 2.17 ­ 2.40; interocellar index 6.0 ­ 6.3; length of antennal segments (in mm): 0.37 ­ 0.47: 0.55 ­ 0.69: 0.60 ­ 0.67: 0.76 ­ 0.94: 0.90 ­ 1.08. Puncturation of pronotal disc weak, punctures irregularly scattered, better visible behind head and in shallow transverse impression behind calli; lateral margins of pronotum with 6 submarginal setigerous punctures bearing long hair­like setae (Fig. 210). Anterior convexity of propleuron alutaceous, punctate laterally and apically, posterior convexity distinctly coarsely punctate with large punctures, propleural depression coarsely punctate. Scutellar disc with almost evenly spaced punctures as large as those on pronotum, apex and basal fifth of scutellar disc impunctate. Punctures on corium distinctly smaller than those on scutellum; clavus with one complete and two partial rows of punctures; mesocorial disc densely and almost evenly punctate; exocorium densely punctate; costa narrow, flattened, separated from exocorium only in its basal half, costal margin with 2 setigerous punctures (Fig. 210). Male hind femur with a row of tubercles on its dorsal margin, and a row of denticles on ventral margin, subapical teeth well visible (Fig. 212); male hind tibia with a row of denticles (Fig. 212); female hind femora with a row of setae on margins, female tibiae simple. Abdominal sterna more or less polished, punctate only close to spiracles and trichobothria. Male genitalia as in Figs 213 ­ 216.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B574576FE3544D0BA15FB56.taxon	materials_examined	TYPE DATA Holotype male: Comoros, Mayotte, Mamoutzou (MNHN). MATERIAL EXAMINED Holotype, and 2 paratypes: Comoros: Grande Comore, 1 paratype (UO); Grande Comore, Nioumbadjou, Raharizonina, 1 paratype (MNHN).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B574576FE3544D0BA15FB56.taxon	distribution	DISTRIBUTION Comoros.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B55457BFE274490BA16FE29.taxon	description	(Figs 217 ­ 223)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B55457BFE274490BA16FE29.taxon	description	DESCRIPTION Body 4.75 ­ 5.70 mm in length, 2.73 ­ 3.19 mm in width, from dark castaneous to blackish brown, corium from castaneous to dark castaneous with punctures darker than the corial disc. Head dorsally slightly wrinkled, at first glance looking impunctate, but bearing numerous hardly visible tiny punctures; clypeus subapically without a pair of setigerous punctures, each paraclypeus submarginally with 6 ­ 8 (usually 7) setigerous punctures bearing long hair­like setae (Fig. 218); ocular index 2.42 ­ 2.75; interocellar index 10.0 ­ 16.0; length of antennal segments (in mm): 0.20 ­ 0.28: 0.22 ­ 0.33: 0.26 ­ 0.35: 0.33 ­ 0.39: 0.35 ­ 0.43. Pronotal disc distinctly punctate in its anterior and lateral parts and behind calli; transverse discal impression absent; lateral margins of pronotum with 7 ­ 9 submarginal setigerous punctures bearing long hair­like setae (Fig. 217). Anterior convexity of propleuron impunctate, posterior convexity impunctate except for a few punctures, propleural depression coarsely punctate. Scutellar disc densely punctate with punctures larger than those on pronotum, anterolateral angles and apex impunctate. Mesocorial disc entirely evenly punctate; exocorium densely punctate; costa narrow, almost entirely separated from exocorium, bearing 4 ­ 6 setigerous punctures (Fig. 217). Margins of male and female hind femur without tubercles, subapical teeth absent (Fig. 219); male and female hind tibia simple, not specifically modified (Fig. 219). Abdominal sterna almost totally impunctate, except for a row of punctures along each segmental suture, and a few coarse wrinkles and punctures posterior to spiracles and trichobothria. Male genitalia as in Figs 220 ­ 223.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B55457BFE274490BA16FE29.taxon	materials_examined	TYPE DATA Holotype male: Australia, South Queensland, Taroom District, Vine Thicket on hill (QMB). MATERIAL EXAMINED Holotype, 18 paratypes, and 34 other specimens: Australia: South Queensland, Taroom District, Vine Thicket on hill, 1 paratype (QMB); Queensland, Cunnamulla, Caravan Pk., 11 paratypes (SAM, UO); Queensland, Annamulla, Caravan Park, 6 paratypes (SAM, UO); Queensland, Brisbane, 1 ex. (BMNH); Northern Territory, Bowerbird, Magela Creek, 3 exx. (NTM, UO); Northern Territory, Gimbat Stn., 1 ex. (NTM); Northern Territory, Umbrawarra Gorge, 2 exx. (NTM); Northern Territory, Alice Springs, 3 exx. (NTM, UO); Northern Territory, Palm Valley, Finke Gorge National Park, 1 ex. (NTM); Northern Territory, Ormiston Gorge, Jn. Ormiston and Pioneer Creeks, 1 ex. (NTM); Northern Territory, Giles Ck. or Wattie Ck., 7 km N Wave Hill Police Stn., 1 ex. (NTM); Northern Territory, Lake Argyle, 1 ex. (NTM); Northern Territory, Top of Ayers Rock, 2 exx. (WAM); South Australia, Coopers Creek, Ferry Crossing, 2 exx. (SAM); South Australia, Hallett Cove, 20 km SW of Adelaide, 1 ex. (UO); South Australia, 5 1 / 2 km WNW Myrtle Springs, 1 ex. (SAM); South Australia, W coast, SE Westalt Point, Smooth Point, 1 ex. (SAM); South Australia, Wirreandah Crk., Crossing, 30 km S Hawker, 1 ex. (SAM); South Australia, Yumbarra Nat. Pk., 1 ex. (SAM); New South Wales, Wilcannia, 1 ex. (SAM); New South Wales, Narrabeen, 3 exx. (HMNH); New South Wales, Sydney, 1 ex. (BMNH); Western Australia, Boorabbin Rock, 1 ex. (WAM); Western Australia, Mt. Ant, 1 ex. (WAM); Western Australia, Marandoo Camp, 2 exx. (WAM, UO); Western Australia, Wurarga, Marloo Stat., 1 ex. (MNHU); Western Australia, Kununurra via Wyndham, 1 ex. (HNHM).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B55457BFE274490BA16FE29.taxon	distribution	DISTRIBUTION Australia.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B58457DFE394257BA16FDC8.taxon	description	(Figs 224 ­ 230)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B58457DFE394257BA16FDC8.taxon	description	DESCRIPTION Body 6.63 ­ 7.57 mm in length, 3.66 ­ 4.21 mm in width, from castaneous to almost black, corium paler than pronotum and scutellum. Head dorsally polished, at first glance looking impunctate, bearing very small irregularly scattered punctures; clypeus subapically without a pair of setigerous punctures, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 225); ocular index 2.23 ­ 2.51; interocellar index 6.0 ­ 7.7; length of antennal segments (in mm): 0.29 ­ 0.33: 0.33 ­ 0.43: 0.39 ­ 0.47: 0.50 ­ 0.65: 0.57 ­ 0.75. Pronotal disc almost impunctate, bearing a few small irregularly scattered punctures; transverse discal impression behind calli absent; lateral margins of pronotum with 5 ­ 6 submarginal setigerous punctures bearing long hair­like setae (Fig. 224). Anterior convexity of propleuron more or less punctate anteriorly and laterally, posterior convexity with a few irregularly scattered punctures, propleural depression with a broad band of coarse punctures. Puncturation of scutellar disc irregular, well visible in its median part. Mesocorial disc almost evenly punctate with punctures as large as those on scutellum; exocorium densely punctate; costa narrow, convex and raised, almost entirely separated from exocorium, bearing a single setigerous puncture (Fig. 224). Femora castaneous or dark castaneous, tibiae basally brown or yellowish brown getting darker towards tarsi; male hind femur with a few very small, almost indistinct denticles bearing short setae on dorsal margin (Fig. 226); male hind tibia with a few very small, sometimes almost indistinct denticles on its inner margin (Fig. 226); female hind femur and tibia without denticles on inner margins. Abdominal sterna polished, slightly wrinkled, coarsely punctate only close to lateral margins of segments, around trichobothria and posterior to spiracles. Male genitalia as in Figs 227 ­ 230.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B58457DFE394257BA16FDC8.taxon	materials_examined	TYPE DATA Holotype male: Australia, North Queensland, Mt Abbott, RF Gully (QMB). MATERIAL EXAMINED Holotype, and 7 paratypes: Australia: Queensland, Mossman Bluff Track, 5 ­ 10 km W Mossman, 1 paratype (QMB); Queensland, Mt French National Park, 3 paratypes (QMB, UO); Queensland, Yarraman, 2 paratypes (QMB, UO); Queensland, „ Glen Witheren ”, Main Scrub, 1 paratype (QMB).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B58457DFE394257BA16FDC8.taxon	distribution	DISTRIBUTION Australia.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B5E457FFE444236BAB0FB77.taxon	description	(Figs 231 ­ 233)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B5E457FFE444236BAB0FB77.taxon	description	DESCRIPTION Body 8.40 ­ 10.69 mm in length, 4.70 ­ 6.45 mm in width, from blackish brown to black, corium usually more brown than pronotum and scutellum; sometimes umbones of pronotum light brown apically. Head dorsally wrinkled, distinctly punctate on paraclypei, vertex smooth; clypeus without a pair of subapical hair­like setae, each paraclypeus with 4 ­ 8 submarginal setigerous punctures bearing long hair­like setae (Fig. 231); ocular index 2.45 ­ 2.90; interocellar index 8.0 ­ 13.0; length of antennal segments (in mm): 0.35 ­ 0.49: 0.49 ­ 0.77: 0.47 ­ 0.70: 0.65 ­ 0.87: 0.70 ­ 0.91. Pronotal transverse postmedian impression absent or only faintly developed; calli more or less elevated; puncturation of disc dense, well visible medially and posteriorly; lateral margins with 12 ­ 16 submarginal setigerous punctures bearing long hair­like setae (Fig. 231). Scutellum densely punctate, with exception of anterolateral angles and apex. Corium punctate; clavus with one complete and two partial rows of punctures; mesocorial disc and exocorium densely punctate; costa narrow, almost entirely separated from exocorium, bearing 4 ­ 9 setigerous punctures (Fig. 231). Propleural depression, apical part of anterior convexity, and almost entire posterior convexity distinctly punctate. Male hind femora with conspicuous subapical tooth, male hind tibiae with large subbasal tubercle. Abdominal sterna glossy in the middle, lateral two thirds of sterna III­VI distinctly punctate and sometimes also transversely striated, sternum VII almost impunctate. Male genitalia as in Figs 232 ­ 233.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B5E457FFE444236BAB0FB77.taxon	materials_examined	TYPE DATA Lectotype female (designated by LIS 1991 a: 185): Ceylon (BMNH). MATERIAL EXAMINED Lectotype, and 94 other specimens: India: Coimbatore, Madras St., 25 exx. (IRSNB, NNML, ZMA, NSMT, UO); Point Calimere, Madras St., 1 ex. (NSMT); Anamalai Hills, Cinchona, 1 ex. (NNML); Pondicherry State, Karikal, 3 exx. (NNML); Kerala, Trichur Distr., Peechi, 2 exx. (UO); Trichinopoli, 1 ex. (MNHN); Trevandrum, 1 ex. (MNHN); Coromandel, Genji, 1 ex. (MNHN); Ramandorogm, 1 ex. (HNHM); Madras, Presid, Ramandroog, 2 ex. (ZIZM, NHMW); Tenmalai, Travancore, 3 exx. (BMNH); Dhikala, Naini Tal, 1 ex. (BMNH); Cote de Malabar, 2 exx. (MNHN); Karnataka, Abiathi, 1 ex. (ZSMC). Nepal: Amiekhgani, Therai, 1 ex. (UO). Sri Lanka: Ceylon, 3 xx. (NhMW, HNHM, BMNH); Anuradhapura, 2 exx. (ZMA, ZSMC); Wirawila, 2 exx. (SMNS); Paradeniya, 1 ex. (NHMW); Hambantota, 2 exx. (BMNH); Galge Ruhunu Nat. Park, 1 ex. (UO); Vavuniya NP., 1 ex. (BMNH); Batticaloa, 1 ex. (BMNH); Polonuruwa, 2 exx. (TLM); Nat. Park Wilpattu, Talawila, 13 km W Maradanmaduwa, 17 exx. (ZIP); Nat. Park Yala, Mahasilawa, 8 km ENE Palatupana, 8 exx. (ZIP); Southern Prov., Yala Nat. Res., 1 ex. (HNHM); Noert Western Prov., Galkulama, SE Anuradhapura, 3 exx. (HNHM, UO); Katagarama, Buttala Road, 3 exx. (HNHM, UO); Dambulla env., 1 ex. (ZSMC).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B5E457FFE444236BAB0FB77.taxon	distribution	DISTRIBUTION India, Nepal, Sri Lanka.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B5C457EFE5144F1BA3EFBB6.taxon	description	(Figs 234 ­ 236)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B5C457EFE5144F1BA3EFBB6.taxon	description	DESCRIPTION Body 7.65 ­ 8.40 mm in length, 3.77 ­ 4.01 in mm in width, chocolate brown, corium a little paler than pronotum and scutellum. Head dorsally polished and impunctate; clypeus without a subapical pair of hair­like setae, each paraclypeus submarginally with a single preocular setigerous puncture bearing long hair­like seta (Fig. 234); ocular index 1.89 ­ 1.98; ocelli large, interocellar index about 8.4 ­ 8.5: length of antennal segments (in mm): 0.36 ­ 0.40: 0.44 ­ 0.46: 0.62 ­ 0.67: 0.78 ­ 0.79: 0.87 ­ 1.02. Puncturation of pronotal disc well visible, especially laterally and posteriorly; transverse impression behind calli deep, bearing a row of coarse punctures interrupted medially, impression continues laterally backwards to umbones; lateral margins with 4 setigerous punctures bearing long hair­like setae (Fig. 234). Anterior convexity of propleuron with small punctures, depression strong with crowded coarse punctures which become finer and sparser posteriorly. Scutellar disc with numerous coarse irregularly spaced punctures, its basal part and apex impunctate. Clavus with one complete row of punctures and few punctures basally; mesocorial disc with scattered almost colourless small punctures; exocorium with very few punctures finer than those on mesocorium; costa almost entirely separated from exocorium, bearing 2 setigerous punctures (Fig. 234). Male hind femora with short tooth subapically, hind tibiae ventrally with a weak basal emargination and small tubercles. Abdominal sterna shining, almost smooth, lateral parts weakly wrinkled; each segment with triangular patch of dense fine punctures posterior to spiracle. Male genitalia as in Figs 235 ­ 236.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B5C457EFE5144F1BA3EFBB6.taxon	materials_examined	TYPE DATA Holotype male: Philippines, Balabac, Dalawan Bay (ZMUC). MATERIAL EXAMINED Holotype, and 1 paratype: Philippines: Balabac, Dalawan Bay, 1 paratype (ZMUC).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B5C457EFE5144F1BA3EFBB6.taxon	distribution	DISTRIBUTION Philippines.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B5D4540FE324433BA80FBB6.taxon	description	(Figs 237 ­ 243)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B5D4540FE324433BA80FBB6.taxon	description	DESCRIPTION Body 6.98 ­ 8.86 mm in length, 3.75 ­ 4.65 mm in width, almost black, corium and posterior margin of pronotal disc blackish brown. Head dorsally looking impunctate, but bearing hardly visible tiny punctures; clypeus subapically without a pair of setigerous punctures, each paraclypeus submarginally with a single preocular puncture bearing long hair­like seta (Fig. 238); ocular index 2.75 ­ 3.80; interocellar index 3.7 ­ 5.0; length of antennal segments (in mm): 0.35 ­ 0.42: 0.40 ­ 0.54: 0.49 ­ 0.72: 0.63 ­ 0.91: 0.76 ­ 0.98. Pronotal disc almost impunctate, with punctures better visible in its posterior half; transverse discal impression behind calli absent or only faintly developed; lateral margins with 4 submarginal setigerous punctures (Fig. 237). Anterior convexity of propleuron alutaceous, impunctate; posterior convexity and propleural depression with numerous coarse punctures. Scutellar disc almost impunctate, except for median patch of punctures (Fig. 239). Mesocorium punctate with several large punctures basally, apically almost impunctate (Fig. 239); exocorium densely punctate with large punctures; costa narrow, almost entirely separated from exocorium, without setigerous punctures (Fig. 239). Hind femora with small subapical teeth in both sexes, hind tibiae with neither denticles nor emargination on inner margins. Abdominal sterna smooth and polished; sutures laterally with small punctures, each sternite with a patch of punctures posterior to spiracle, and a few punctures anterior to it. Male genitalia as in Figs 240 ­ 243.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B5D4540FE324433BA80FBB6.taxon	materials_examined	TYPE DATA Lectotype male (designated by LIS 1996: 223): New Guinea, Lemien Berlinenhafen (HNHM). MATERIAL EXAMINED Lectotype, and 7 other specimens: Papua New Guinea: Kokoda, 1 ex. (BMNH); Madang Dist., Finisterre Mts, Damanti, 2 exx., Budemu 3 exx., Moro 1 ex. (BMNH, UO).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B5D4540FE324433BA80FBB6.taxon	distribution	DISTRIBUTION Papua New Guinea.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B634542FE384430B96DFBB6.taxon	description	(Figs 244 ­ 250)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B634542FE384430B96DFBB6.taxon	description	DESCRIPTION Body 10.67 ­ 12.17 mm in length, 6.02 ­ 6.32 mm in width, almost black, corium brownish black. Head dorsally impunctate; clypeus as long as paraclypei, without subapical setigerous punctures, each paraclypeus submarginally with a single preocular setigerous punctures (Fig. 245); ocular index 1.66 ­ 2.08; ocelli large, interocellar index 5.5 ­ 8.7; length of antennal segments (in mm): 0.43 ­ 0.55: 0.56 ­ 0.70: 0.85 ­ 1.13: 1.09 ­ 1.25: 1.15 ­ 1.35. Pronotal disc with well visible puncturation; transverse discal impression behind calli present, accompanied by a row of coarse punctures; lateral margins with 4 ­ 6 submarginal setigerous punctures (Fig. 244). Propleuron punctate, anterior convexity dulled by numerous tiny punctures, depression and posterior convexity with larger punctures. Scutellum sparsely punctate, its basal part and apex impunctate. Corium distinctly punctate with punctures smaller than those on pronotum and scutellum; clavus with one complete and one partial row of punctures; mesocorial disc almost evenly punctate; costa moderately convex, almost entirely separated from exocorium, bearing 1 ­ 2 setigerous punctures (Fig. 244). Male hind femora with large subapical tooth on dorsal margin, and numerous small teeth on the ventral (Fig. 246); hind tibiae with small subbasal tubercles (Fig. 246); female hind femora with small conspicuous subapical tooth on dorsal margin, hind tibiae without tubercles. Abdominal sterna alutaceous, slightly wrinkled; sterna V­VII with patches of tiny punctures posterior and anterior to spiracle, sternum III and sometimes also sternum IV with very few punctures anteriorly of the spiracle. Male genitalia as in Figs 247 ­ 250.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B634542FE384430B96DFBB6.taxon	materials_examined	TYPE DATA Lectotype female (designated by LIS 1994 a: 225): Borneo (NHMW). MATERIAL EXAMINED Lectotype, and 11 other specimens: Indonesia: North Sumatra, Bivonac One, Mt. Bandahara, 9 exx. (NNML, UO). Malaysia: Borneo, Sarawak, Semongok, 1 ex. (UO). Vietnam: Tonkin, Montes Mauson, 1 ex. (NhMW).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B634542FE384430B96DFBB6.taxon	distribution	DISTRIBUTION Borneo, Sumatra, Vietnam.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B614547FE314430B802FAE9.taxon	description	(Figs 251 ­ 257)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B614547FE314430B802FAE9.taxon	description	DESCRIPTION Body 6.08 ­ 8.75 mm in length, 3.68 ­ 5.32 mm in width, from castaneous to black, posterior margin of pronotum sometimes slightly lighter in shade. Head dorsally striated and wrinkled, at first glance looking impunctate, but bearing numerous hardly visible punctures; clypeus subapically without a pair of setigerous punctures, each paraclypeus submarginally with 4 ­ 10 (usually 6 ­ 9) setigerous punctures bearing long hair­like setae (Fig. 252); ocular index 2.38 ­ 2.90; interocellar index 4.0 ­ 7.0; length of antennal segments (in mm): 0.26 ­ 0.45: 0.37 ­ 0.57: 0.34 ­ 0.62: 0.37 ­ 0.46: 0.40 ­ 0.50. Puncturation of pronotal disc well developed, punctures coarse and large, better visible behind calli; transverse discal impression behind calli absent or very shallow; lateral margins of pronotum with 10 ­ 15 submarginal setigerous punctures bearing long hair­like setae (Fig. 251). Anterior convexity of propleuron alutaceous, its anterior half distinctly punctate; posterior convexity with a few large punctures close to propleural depression, the latter densely and coarsely punctate. Scutellar disc with large crowded punctures, except basal fourth and extreme apex. Mesocorial disc densely and evenly punctate; exocorium densely punctate; costa broad, flat, separated from exocorium in its basal two thirds, and bearing 7 ­ 13 setigerous punctures (Fig. 251). Male hind femora with a few more or less visible denticles, subapical teeth absent (Fig. 253); male hind tibia with large subbasal tooth (Fig. 253); female hind tibia and femur simple. Abdominal sterna alutaceous, more or less polished, each segmental suture accompanied by a row of punctures, median parts of sterna impunctate, lateral thirds with numerous large punctures. Male genitalia as in Figs 254 ­ 257.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B614547FE314430B802FAE9.taxon	materials_examined	TYPE DATA Lectotype female of Cydnus piceus WESTWOOD (designated by LIS 1995 b: 145): N [ova] H [ollandia] (OXUM). Lectotype male of Cydnus dilatatus SIGNORET (designated by LIS 1995 b: 144): Australia, Somerset (NHMW). MATERIAL EXAMINED Lectotype of Cydnus piceus, Lectotype of Cydnus dilatatus, and 140 other specimens: Australia: Western Australia, Yanchep, 32 m. N. of Perth, 1 ex. (BMNH); Western Australia, Yardie Ck., Carnarvon, 3 exx. (NMV, UO); Western Australia, Carnarvon, 6 exx. (NMV, UO); Western Australia, Bungabiddy Rockhole, Walter James Ra., 3 exx. (WAM, UO); Western Australia, Irrunytju Rockhole, Hinckley Range, 3 exx. (WAM, UO); Western Australia, Mt Yokine, 2 exx. (NMV); Western Australia, 4 km SW Giles Creek, X­ring, Giles­Docker R Rd., 1 ex. (WAM); Western Australia, Nullagine, 1 ex. (WAM); Western Australia, Roebuck Bay, 1 ex. (BMNH); Western Australia, Enneaba, 1 ex. (WAM); Northern Territory, 2.8 mls S. of Renner Springs, 12 exx. (SAM, UO); Northern Territory, Kings Canyon, George Gill Range, 5 exx. (NTM, UO); Northern Territory, Junction of Arnhem Hwy & Oenpelli Road, 2 exx. (NTM); Northern Territory, South Attack Ck, 6 exx. (NTM, UO); Northern Territory, Attack Ck., 2 exx. (NTM); Northern Territory, Murgenella, 3 exx. (NTM, UO); Northern Territory, 1 km W of Baralili Ck, crossing on Arnhem Hwy nr Jabiru, 2 exx. (NTM); Northern Territory, White Range, Arltunga, 3 exx. (NTM, UO); Northern Territory, 1 km N of Barrow Creek, 1 ex. (NTM); Northern Territory, Groote, Eylandt, 1 ex. (NTM); Northern Territory, Groote, Eylandt, 1 ex. (BMNH); Northern Territory, Kakadu NP., c. 1 km S of Arnhem Hwy on Pine Creek Rd., 1 ex. (NTM); Northern Territory, Corndorl billabong nr Jabiru, 1 ex. (NTM); Northern Territory, 9 km N by E of Mudginberri, 4 exx. (NTM, UO); Northern Territory, Swim Ck., Crossing Point Stuart Stn., 1 ex. (NTM); Northern Territory, Koongara, 15 km E of Mt. Cahill, 2 exx. (NTM); Northern Territory, Cooper Ck., 15 km S by W of Nimbuwah Rock, 1 ex. (NTM); Northern Territory, 1 km N of Cahills Crossing, East Alligator R., 1 ex. (NTM); Northern Territory, Melville Island, Pularumpi, 4 exx. (NTM, UO); Northern Territory, Nourlangie Ck., 8 km N of Mt. Cahill, 1 ex. (NTM); Northern Territory, 4.6 km S of Renner Springs, 2 exx. (SAM); Northern Territory, Elliot, 3 exx. (NTM, UO); Northern Territory, Roderick Creek, Gregory N. P., 1 ex. (NTM); Northern Territory, Keep River NP., Gorge carpark, 1 ex. (NTM); Northern Territory, Ooraminna Rockhole, 50 km SSE Alice Springs, 1 ex. (NTM); Northern Territory, Green Ant Ck., 40 ml. S of Adelaide River, 1 ex. (NTM); Northern Territory, Elsey Ck., 10 ml. E of Mataranka on Roper Bar Rd., 1 ex. (NTM); South Australia, 22 km WN. V of Moralana HS, 2 exx. (SAM); South Australia, 10 km NW Maynards Well. Nthn Flinders Rgs., 2 exx. (SAM); South Australia, Flinders Rg., Brachina Gorge, 2 exx. (SAM); South Australia, Mudla Bore, 35 km NNE Billa Kalina HS, 3 exx. (SAM); South Australia, Coopers Creek, Ferry Crossing, 2 exx. (SAM); South Australia, Levi Crk., 8 km NW Big Perry Spring, 20 exx. (SAM, UO); South Australia, L. Torrens nr. Beda Hill. 2 exx. (SAM); South Australia, Mid E of L. Frome coast, Sandhills, Desert Oak., 3 exx. (SAM, UO); South Australia, 10 km ENE Stuart Crk. HS., 1 ex. (SAM); South Australia, Mt. Remarkable, 2 exx. (SAM); South Australia, Margaret R., 10 km SE Coward Springs, 1 ex. (SAM); South Australia, Ediacara, 1 ex. (SAM); Queensland, Telegraph Line Crossing, Jardine R., Cape York, 1 ex. (UQIC); Queensland, Bamaga, Cape York, 1 ex. (QMB); Queensland, Moreton Is., 1 ex. (QMB).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B614547FE314430B802FAE9.taxon	distribution	DISTRIBUTION Australia (incl. Tasmania), Indonesia (Sumba Island).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B644549FE284517B901FA56.taxon	description	(Figs 258 ­ 264)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B644549FE284517B901FA56.taxon	description	DESCRIPTION Body 6.81 ­ 9.12 mm in length, 3.87 ­ 4.85 mm in width, from castaneous to blackish brown, corium and umbones of pronotum usually slightly paler than scutellum. Head dorsally at first glance looking impunctate, but bearing hardly visible tiny punctures; clypeus without subapical setigerous punctures, each paraclypeus submarginally with a single preocular setigerous punctures (Fig. 259); ocular index 2.00 ­ 2.40; interocellar index 8.0 ­ 12.0; length of antennal segments (in mm): 0.31 ­ 0.42: 0.40 ­ 0.49: 0.49 ­ 0.65: 0.63 ­ 0.80: 0.68 ­ 0.92. Pronotal transverse impression behind calli more or less shallow, with a row of punctures interrupted medially; lateral margins with 5 ­ 8 submarginal setigerous punctures bearing long hair­like setae (Fig. 258); puncturation of anterior and posterior lobe of pronotal disc clearly visible. Propleural depression and basal part of posterior convexity with numerous punctures, anterior convexity alutaceous. Scutellum with numerous irregularly scattered punctures, apex and anterolateral angles impunctate. Corium with distinct punctures smaller than those on pronotum and scutellum; clavus with one complete and one or two partial rows of punctures; mesocorial disc almost evenly punctate; costa narrow, moderately convex, almost entirely separated from exocorium, bearing 2 ­ 4 setigerous punctures (Fig. 258). Male hind femora with distinct subapical tooth on dorsal margin and several smaller teeth on the ventral, hind tibiae with small subbasal tubercles (Fig. 260); female hind femora with small subapical tooth on dorsal margin, ventral margin with setae only, hind tibiae without tubercles. Abdominal sterna smooth or weakly wrinkled laterally, with tiny punctures along sutures; sterna III and IV with a few tiny punctures anterior and posterior to spiracles, sterna V­VII with tiny punctures only posterior to spiracles. Male genitalia as in Figs 261 ­ 264.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B644549FE284517B901FA56.taxon	materials_examined	TYPE DATA Holotype male: Vietnam, prov. Kien Giang, isl. Tho Tu (UO). MATERIAL EXAMINED Holotype, 1 paratype and 14 other specimens: China: Prov. Fo­kien, 1 ex. (ZIZM). Laos: Mg. Xoi Prov., 1 ex. (NNML). Vietnam: Lao­Kay, Tonkin, paratype (MNHN); Tonkin, Ha­Giang, 1 ex. (MNHN); Prov. Ha­Tinh, foretsiere Hüöng­sön, 4 exx. (HNHM, UO); Moc Chau, 2 exx. (HNHM, UO); Ha Son­Binh, 10 km SW Hoa­Binh, 1 ex. (UO); Poulocondore, 1 ex. (NNML); Nha­Trang, South Annam, 3 exx. (ZlP, UO).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B644549FE284517B901FA56.taxon	distribution	DISTRIBUTION China (Fujian), Laos, Vietnam.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B6A454BFE334590BAB3F9B6.taxon	description	(Figs 265 ­ 271)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B6A454BFE334590BAB3F9B6.taxon	description	DESCRIPTION Body 8.39 ­ 9.85 mm in length, 4.60 ­ 5.45 mm in width, from castaneous to black, corium usually paler than pronotum and scutellum, at least costa and a part of exocorium brown or yellowish brown, sometimes also apices of pronotal umbones pale brown or yellowish brown. Head dorsally alutaceous and impunctate except for setigerous punctures and sometimes a few very small punctures laterally; clypeus subapically without a pair of setigerous punctures, each paraclypeus submarginally with 2 ­ 5 setigerous punctures bearing long hair­like setae (Fig. 266); ocular index 2.10 ­ 2.70; interocellar index 7.0 ­ 12.0; length of antennal segments (in mm): 0.34 ­ 0.40: 0.45 ­ 0.60: 0.55 ­ 0.69: 0.76 ­ 0.88: 0.77 ­ 0.88. Puncturation of pronotal disc well visible, punctures small, better visible behind head and calli, discal transverse impression behind calli absent or very shallow; lateral margins of pronotum with 6 ­ 10 submarginal setigerous punctures bearing long hair­like setae (Fig. 265). Anterior convexity of propleuron impunctate or almost impunctate, posterior convexity distinctly coarsely punctate close to propleural depression, the latter with a band of coarse punctures. Scutellar disc densely punctate with almost evenly spaced punctures larger than those on pronotum, anterolateral angles of a disc impunctate. Corium densely punctate; clavus with one complete and two incomplete rows of punctures; mesocorial disc densely and almost evenly punctate; exocorium more or less densely punctate; costa flattened, separated from exocorium only in its basal third, costal margin with 2 ­ 3 setigerous punctures (Fig. 265). Femora from castaneous to almost black, tibiae yellow or yellowish brown, tibial spines dark brown or black; male hind femur with a row of teeth on its dorsal margin, among them three subapical, ventral margin with a row of well visible teeth, among them one large subbasal and three subapical (Fig. 267); male hind tibia with a row of very small denticles (Fig. 267); female hind femora with a row of setae on margins, female tibiae simple. Abdominal sterna more or less polished, almost totally impunctate, bearing only a patch of few small punctures posterior to spiracles. Male genitalia as in Figs 268 ­ 271. TYPE DATA Lectotype male (designated by LIS 1999 d): Madagascar, Finarantsoa (HNHM).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B6A454BFE334590BAB3F9B6.taxon	materials_examined	MATERIAL EXAMINED Lectotype, and 20 other specimens: Madagascar: Madagascar, 3 exx. (MNHN, UO); Madagascar Sambirano, Nord­est de Maromandia, Base du Manongarivo, 2 exx. (MNHN, UO); Madagascar Sud, plateau Mahafaly, 11 / 12 km Ouest d ̕ Ankalirano, 1 ex. (MNHN); Madagascar Ouest, Ankarafantsika, Ampijoroa, 1 ex. (MNHN); N. d ̕ Ivohibe, Station Forest d ̕ Angodongodona, 1 ex. (MNHN); Région de Sud, Pays Androy, Jehara, 1 ex. (MNHN); Région de Sud, Andrahomana, 1 ex. (UO); Province de Fort­Dauphin, Antanimora, 1 ex. (MNHN); Lambomakandro, Tuléar, 1 ex. (MNHN); Ikongo, 1 ex. (MNHN); Périnet, 1 ex. (ZIP); Mahajanga prov., Mahajamba riv., Ampatika env. 2 exx. (USMB, UO); Mahajanga env., Mahajanga distr., 2 ex. (USMB); Mahajanga distr., Ambodimanga env., 2 exx. (USMB, UO).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B6A454BFE334590BAB3F9B6.taxon	distribution	DISTRIBUTION Madagascar, Rodriguez.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B69454AFE4540A8BA16F9A1.taxon	description	(Figs 272 ­ 278)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B69454AFE4540A8BA16F9A1.taxon	description	DESCRIPTION Body 6.93 ­ 7.38 mm in length, 4.16 ­ 4.55 mm in width, dark castaneous, posterior margin of pronotum narrowly yellowish brown, corium more yellow­brown in shade than scutellum, punctate with dark punctures. Head dorsally slightly wrinkled and bearing numerous small, almost evenly spaced punctures; clypeus subapically with a pair of setigerous punctures bearing long hair­like setae, each paraclypeus submarginally with a row of 8 ­ 9 setigerous punctures bearing long hair­like setae (Fig. 273); ocular index 2.69 ­ 2.80; interocellar index 7.1 ­ 9.3; length of antennal segments (in mm): 0.35: 0.42 ­ 0.44: 0.38 ­. 40: 0.53 ­ 0.54: 0.52 ­ 0.54. Pronotal disc with scattered punctures or different size, calli impunctate; transverse discal impression behind calli shallow and almost indistinct, bearing medially interrupted row of several coarse punctures; lateral margins of pronotum with 16 ­ 21 submarginal setigerous punctures bearing long hair­like setae (Fig. 272). Anterior convexity of propleuron impunctate, except for a few small punctures anteriorly, posterior convexity with a few large punctures close to propleural depression, the latter with a row of well separated coarse punctures. Scutellar disc with almost evenly spaced punctures larger than those on pronotum; basal fourth of scutellar disc impunctate. Mesocorial disc evenly punctate with exception of its middle fifth; exocorium with very dense puncturation; costa broad and flat, separated from exocorium only in its basal two thirds, costal margin with 8 ­ 11 setigerous punctures bearing long hair­like setae (Fig. 272). Femora castaneous, tibiae brown or yellowish brown; male hind femur with 2 subapical teeth on ventral margin, and a row of denticles on dorsal margin (Fig. 274); female hind femur with a row of small denticles on dorsal margin; male hind tibia with large subbasal lobe­like denticle and numerous smaller tubercles (Fig. 274); female hind tibia simple. Abdominal sterna almost totally impunctate, except for rows of punctures parallel to segmental sutures, and longitudinal patches of punctures posterior to each spiracle. Male genitalia as in Figs 275 ­ 278.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B69454AFE4540A8BA16F9A1.taxon	materials_examined	TYPE DATA Holotype male: Australia, Western Australia, Yardie Creek (WAM). MATERIAL EXAMINED Holotype, and 1 paratype: Australia: Western Australia, Southern Cross, 1 paratype (WAM).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B69454AFE4540A8BA16F9A1.taxon	distribution	DISTRIBUTION Australia.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B6F4551FE2440ABB918FAC9.taxon	description	(Figs 279 ­ 282)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B6F4551FE2440ABB918FAC9.taxon	description	DESCRIPTION Body 9.25 ­ 11.90 mm in length, 5.04 ­ 6.59 mm in width, from yellowish brown to almost black, sometimes corium slightly paler than pronotum and scutellum. Head dorsally at first sight looking impunctate, but bearing hardly visible tiny punctures, lateral parts of paraclypei wrinkled; clypeus subapically without setigerous punctures, each paraclypeus submarginally with 2 ­ 4 setigerous puncture bearing long hair­like setae (Fig. 279); ocular index 2.10 ­ 2.64; interocellar index 4.0 ­ 8.5; length of antennal segments (in mm): 0.34 ­ 0.63: 0.59 ­ 0.76: 0.73 ­ 0.85: 0.90 ­ 1.13: 0.94 ­ 1.15. Puncturation of pronotal disc more or less dense, punctures small, clearly visible; calli, umbones and posterior fourth of pronotal disc impunctate; discal transverse impression behind calli shallow, developed only laterally, usually acompanied by a row of coarse punctures; lateral margins of pronotum with 6 ­ 8 submarginal setigerous punctures bearing long hair­like setae (Fig. 279). Anterior convexity of propleuron apically punctate with small punctures, posterior convexity distinctly anteriorly punctate with large punctures, propleural depression with a band of coarse punctures and wrinkles. Scutellar disc punctate with almost evenly spaced punctures. Corium densely punctate; clavus with one complete and two partial rows of punctures; mesocorial disc densely and almost evenly punctate; exocorium densely punctate; costa flattened, basally expanded, separated from exocorium only in its basal half, costal margin with 2 setigerous punctures bearing long hair­like setae (Fig. 279). Male mid femur with a row of small teeth; male hind femur with a row of well visible teeth and denticles on margins, subbasal and subapical teeth large (variability of edentation was presented by LINNAVUORI, 1993), male hind tibia with a row of numerous denticles; female hind femur with small (sometimes indistinct) subapical tooth, female tibia simple. Abdominal sterna more or less polished, punctate only around spiracles and trichobothria. Male genitalia as in Figs 280 ­ 282.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B6F4551FE2440ABB918FAC9.taxon	materials_examined	TYPE DATA Syntype (s) male (s) of Macroscytus reflexus SIGNORET: South Africa (lost or destroyed). Neotype male (present designation) of Macroscytus reflexus SIGNORET: South Africa, Natal, Empangeni area, 28 ° 45 ̕ S 31 ° 55 ” E, 20 ­ II­ 86, 110 m, P. Reavell, light (NMP). Lectotype female (designated by LINNAVUORI 1993: 25) of Macroscytus acutus SIGNORET: Kenya, Mombasa (BMNH). Lectotype male (designated by LINNAVUORI 1993: 22, as Holotype) of Macroscytus excavatus SIGNORET: East Africa, Lac Hyarsa [= Nyassa Lake] (BMNH). Lectotype male (designated by LINNAVUORI 1977: 41) of Macroscytus inermipes HORVÁTH: Sao Thomé (HNHM). MATERIAL EXAMINED Neotype of Macroscytus reflexus, Lectotype of Macroscytus acutus, Lectotype of Macroscytus excavatus, Lectotype of Macroscytus inermipes, and 229 other specimens: Angola: 1 ex. (MNHN). Burkina Faso: Ouagadougou, 1 ex. (ZIP). Cameroon: Nkolbisson, 3 exx. (MNHN, UO); Yaoundé, 1 ex. (MNHN). Central African Republic: La Maboke, 16 exx. (MNHN, UO); Boukoko, 3 exx. (MNHN). Congo: Bouenza, 1 ex. (MNHN); Brazzaville, 85 exx. (HNHM, MNHN, UO); Sibiti, 1 ex. (MNHN); Odzala, 1 ex. (MNHN); Dimonika, Mayumbe, 7 exx. (MNHN). Eritrea: Ghinda, 2 exx. (NNML, ZIP). Ethiopia: Bahor Dar, 10 exx. (SMNS, UO); Gidole, Prov. Gamu Gofa, 5 exx. (SMNS, UO); Wollo­Ataye, 1 ex. (MNHN); Kébré­Mengist, 1 ex. (MNHN); Lake Langano, Shoa, 3 exx. (ZSMC). Gabon: Ovenda near Libreville, 1 ex. (ZIZM). Guinea: Mt. Nimba, Plateau de Zouguepo, 1 ex. (MNHN). Ivory Coast: Lamto, 1 ex. (MNHN). Kenya: Malindi, 3 exx. (MNHU); Mombasa, 2 exx. (MNHU, SMNS); Mt. Elgon National Park, near Chapnyalil Cave, 1 ex. (HNHM); Ukunda, Diani Beach, 1 ex. (HNHM); Tsavo West National Park, 1 ex. (HNHM). Liberia: 1 ex. (ZIZM). Mozambique: 1 ex. (ZSMC); Beira, 1 ex. (MNHN); Lourenco Marques, Matola, Vila Salazar, 1 ex. (JEC). Namibia: Kavango, Popa­Falls, 6 exx. (MNHU, MNHU, UO); East Caprivi, Katima Mulilo, 3 exx. (MNHU, UO). Ruanda: Kissenyi, 1 ex. (MNHN). Senegal: Kedougou, 2 exx. (MNHN). Sierra Leone: Makeni, 12 exx. (MZLU, UO). South Africa: Natal, Empangeni area, 14 exx. (NMP, UO); Natal, Kube Yini Game Reserve, 3 exx. (NMP, UO); Natal, Mkuzi Game Reserve, 7 exx. (MNHU, NMP, UO). St. Thomé Island: Bombaim, 1 ex. (USMB). Tanzania: Arusha distr., Naberera env., 3 exx. (USMB, UO); Morogoro, 4 exx. (HNHM, UO). Togo: Lama Kara, 1 ex. (ZIZM). Uganda: Nyabyeya, 3 exx. (MNHU, UO). Zambia: Chisamba nr Lusaka, 1 ex. (NMP); Kafue National Park, Chunga Camp, 1 ex. (MNHU); 5 km S Choma, 1 ex. (MNHU). Zimbabwe: Matabeleland, Matopos National Park, 3 exx. (MNHU, UO); Zimbabwe Ruins, 1 ex. (NMP); Kariba, 5 exx. (JEC, UO).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B6F4551FE2440ABB918FAC9.taxon	distribution	DISTRIBUTION Angola, Burkina Faso (first record), Cameroon, Central African Republic,? Chad, Congo, Eritrea, Ethiopia, Gabon (first record), Gambia, Ghana, Guinea (first record), Ivory Coast, Kenya, Liberia (first record), Mozambique (first record), Namibia (first record), Nigeria, Ruanda (first record), Senegal (first record), Sierra Leone, South Africa, St. Thomé Island, Sudan, Tanzania, Togo (first record), Uganda, Zaire, Zambia, Zimbabwe.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B6F4551FE2440ABB918FAC9.taxon	discussion	REMARK The type material of Macroscytus reflexus from South Africa was not found in the collections containing SIGNORET ̕ s specimens (it was apparently lost or destroyed). Since the other three species (Macroscytus acutus, M. excavatus, M. inermipes) are found to be conspecific with the former, it was necessary to designate a neotype for M. reflexus.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B724552FE364537B8CCFC48.taxon	description	(Figs 287 ­ 294)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B724552FE364537B8CCFC48.taxon	description	DESCRIPTION Body 9.46 ­ 12.00 mm in length, 5.44 ­ 6.69 mm in width, from castaneous to dark castaneous, corium usually paler than pronotum and scutellum. Head dorsally punctate and wrinkled, especially in lateral parts of paraclypei; clypeus subapically without a pair of setigerous punctures, each paraclypeus submarginally with a single preocular setigerous puncture bearing long hair­like seta (Fig. 288); ocular index 2.30 ­ 2.62; interocellar index 4.0 ­ 6.0; length of antennal segments (in mm): 0.40 ­ 0.49: 0.54 ­ 0.71: 0.76 ­ 0.97: 1.00 ­ 1.19: 1.07 ­ 1.21. Puncturation of pronotal disc well visible, dense, punctures small, calli and posterior fourth of pronotal disc impunctate; transverse discal impression behind calli absent or very shallow, developed only laterally, calli somewhat elevated; lateral margins of pronotum with 4 ­ 6 submarginal setigerous punctures bearing long hair­like setae (Fig. 287). Anterior convexity of propleuron laterally and apically punctate with numerous small punctures, posterior convexity distinctly coarsely punctate with punctures larger than those of anterior convexity, propleural depression with a band of coarse punctures and wrinkles. Scutellar disc densely punctate with almost evenly spaced punctures smaller than those on pronotum and corium, basal fifth and apex impunctate. Corium densely punctate; clavus with one complete and two partial rows of punctures; mesocorial disc densely and almost evenly punctate; exocorium densely punctate; costa somewhat flattened, separated from exocorium only in its basal two thirds, costal margin with 2 setigerous punctures (Fig. 287). Male mid femur with a row of small teeth; male hind femur with a row of well visible teeth and denticles on margins, subapical teeth well developed (Figs 289 ­ 290); male hind tibia without denticles; female hind femora with small subapical tooth, female tibia simple. Abdominal sterna polished, more or less densely punctate in lateral thirds. Male genitalia as in Figs 291 ­ 294.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B724552FE364537B8CCFC48.taxon	materials_examined	TYPE DATA Syntype (s) of Macroscytus ruficornis SIGNORET: Guinea (lost). Neotype male (present designation) of Macroscytus ruficornis SIGNORET: Guinea, Coyah; III­V. 1967, leg. Ferenczy (HNHM). Syntype of Macroscytus punctiventris SIGNORET: Kenya, Mombasa (BMNH, lost). Syntypes of Macroscytus rubiginosus STICHEL: Cameroon, Joko (not located, lost or destroyed). MATERIAL EXAMINED Neotype of Macroscytus ruficornis, and 16 other specimens: Benin: Dahomey, Ouidam, 1 ex. (MNHN); Plat. De Zaguanado & Ketou, 1 ex. (MNHN); Palteau de Kaguanado, 1 ex. (MNHN). Cameroon: Nanga Eboko, 1 ex. (HNHM). Central African Republic: Lamaboké, 3 exx. (MNHN, UO). Congo: Sanga, 1 ex. (MNHN). Guinea: Mt. Nimba, Savana de Serengbara, 2 exx. (MNHN); Mt. Nimba, Bord du Zougué, 1 ex. (MNHN); Mt. Nimba ,, 3 exx. (MNHN, UO). Ivory Coast: Lamto, Toumodi, 1 ex. (MNHN). Togo: Bismarckburg (= Yegue), 1 ex. (MNHN).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B724552FE364537B8CCFC48.taxon	distribution	DISTRIBUTION Benin (first record), Cameroon, Central African Republic (first record), Congo (first record), Guinea, Ivory Coast, Kenya, Nigeria, Sierra Leone, Tanzania (Zanzibar), Togo.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B724552FE364537B8CCFC48.taxon	discussion	REMARKS I have not been able to trace the type material of Macroscytus ruficornis from Guinea in the collections containing SIGNORET ̕ s specimens. Nevertheless, only two species of Macroscytus are known from Guinea — M. brunneus and another one that should be treated as M. ruficornis (the latter described originally just from this country). Since the type material of M. ruficornis was apparently lost or destroyed, and it was hitherto treated as a species of uncertain position it was necessary to designate a neotype for it. Syntypes of two other species, namely Macroscytus punctiventris from Kenya and M. rubiginosus from Cameroon were also lost or destroyed (LINNAVUORI, 1993; LIS, unpublished data), but original descriptions and distribution data allow to treat both as junior synonyms of M. ruficornis. Specimens from Kilimandjaro recorded by JEANNEL (1913: 15) as M. ruficornis represent Geocnethus nitidus SIGNORET (see LINNAVUORI 1993).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B714554FE2643B1BA36FBB6.taxon	description	(Figs 295 ­ 301)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B714554FE2643B1BA36FBB6.taxon	description	DESCRIPTION Body 10.34 ­ 11.02 mm in length, 5.64 ­ 5.99 mm in width, from dark castaneous to black, entire pronotal umbones yellow­brown, thus making pronotal disc sharply bicoloured. Head dorsally distinctly punctate with large, coarse punctures; clypeus subapically without a pair of setigerous punctures, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 296); ocular index 2.60 ­ 2.75; interocellar index 4.5 ­ 5.6; length of antennal segments (in mm): 0.49 ­ 0.53: 0.72 ­ 0.76: 0.80 ­ 0.87, 4 th and 5 th segment missing. Puncturation of pronotal disc clearly visible, punctures larger than those on head; calli and posterior margin impunctate; transverse discal impression behind calli absent; lateral margins of pronotum with 4 ­ 5 submarginal setigerous punctures bearing long hair­like setae (Fig. 295). Anterior convexity of propleuron alutaceous, anteriorly and laterally coarsely punctate with small punctures; posterior convexity alutaceous, with numerous large punctures; propleural depression with large and coarse punctures. Scutellum almost evenly punctate, punctures almost of the same size as those on pronotum; antero­lateral angles impunctate. Clavus with one complete and two partial rows of punctures; mesocorial disc almost evenly punctate, all punctures distinctly smaller than those on scutellum; exocorium distinctly punctate, punctures of the same size as those on mesocorial disc; costa narrow, entirely separated from exocorium, costal margin with a single setigerous puncture bearing long hair­like seta (Fig. 295). Ventral margin of male hind femur with a subapical denticle, its dorsal margin with large subapical tooth (Fig. 297); male hind tibia without denticles on its inner margin (Fig. 297); ventral margin of female hind femur with small subapical denticle, its dorsal margin with subapical tooth; female hind tibia without denticles on its inner margin. Abdominal sterna polished, bearing large punctures in the lateral third, around trichobothria and spiracles. Male genitalia as in Figs 298 ­ 301.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B714554FE2643B1BA36FBB6.taxon	materials_examined	TYPE DATA Holotype male: Madagascar, Tamatave prov., Moramanga env. (USMB). MATERIAL EXAMINED Holotype, and 3 paratypes: Madagascar: Tamatave prov., Moramanga env., 3 paratypes (USMB, UO).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B714554FE2643B1BA36FBB6.taxon	distribution	DISTRIBUTION Madagascar.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B774557FE374430BA5FFB96.taxon	description	(Figs 283 ­ 286)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B774557FE374430BA5FFB96.taxon	description	DESCRIPTION Body 8.45 ­ 9.24 mm in length, 4.70 ­ 5.05 mm in width, from dark castaneous to almost black, corium slightly paler than pronotum and scutellum. Head dorsally at first glance looking impunctate, but bearing hardly visible tiny punctures; clypeus without subapical setigerous punctures, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 283); ocular index 2.30 ­ 2.57; interocellar index 6.9 ­ 9.0; length of antennal segments (in mm): 0.37 ­ 0.46: 0.42 ­ 0.57: 0.51 ­ 0.70: 0.66 ­ 0.85: 0.80 ­ 1.01. Pronotal transverse impression behind calli absent, disc undivided into lobes, posterior half with more or less broad band of punctures; lateral margins with 3 ­ 6 submarginal setigerous punctures (Fig. 283). Propleural depression with numerous coarse punctures. Scutellum almost impunctate or with a few clearly separated punctures. Clavus with one complete and two partial rows of punctures; exocorium and mesocorium almost evenly punctate; costa moderately convex, basally flattened, almost entirely separated from exocorium, bearing two setigerous punctures (Fig. 283). Hind male femur with one large and two small teeth basally and one tooth subapically (Fig. 286). Abdominal sterna polished and smooth, sometimes slightly wrinkled close to segmental sutures; lateral third of segments finely punctate, and bearing triangular patches of dense tiny punctures posterior to spiracle and several punctures anterior to it. Male genitalia as in Figs 283 ­ 285.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B774557FE374430BA5FFB96.taxon	materials_examined	TYPE DATA Lectotype male (designated by LIS 1994 a: 228): Philippines (BMNH). MATERIAL EXAMINED Lectotype, and 2 other specimens: Indonesia: Flores, Larantoeka, 1 ex. (ZMA). Thailand: Siam, 1 ex. (MNHN).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B774557FE374430BA5FFB96.taxon	distribution	DISTRIBUTION Indonesia (Flores), Philippines, Thailand. Records from other countries were based on misidentifications (see LIS 1994 a).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B744559FE3E44D0BA36FCB7.taxon	description	(Figs 302 ­ 306)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B744559FE3E44D0BA36FCB7.taxon	description	DESCRIPTION Body 12.58 mm in length, 6.42 mm in width, almost entirely black, only corium dark castaneous to some extent. Head dorsally polished, slightly wrinkled in lateral parts of paraclypei, at first glance looking impunctate, but bearing numerous hardly visible tiny punctures; clypeus without subapical hair­like setae, each paraclypeus submarginally with a single preocular setigerous puncture bearing long hair­like seta (Fig. 303); ocular index 1.90; interocellar index 6.7; length of antennal segments (in mm): 0.60: 0.76: 1.07: 1.39, 5 th segment missing. Puncturation of pronotal disc well visible, especially behind head and callal areas; transverse discal impression behind calli well developed, deep and punctate; lateral margins of pronotum with 4 ­ 5 submarginal setigerous punctures bearing long hair­like setae (Fig. 302). Anterior convexity of propleuron dulled and densely punctate with small punctures, posterior convexity with large scattered punctures, propleural depression densely and coarsely punctate. Scutellar disc with irregularly scattered punctures, its basal fourth impunctate. Clavus with one complete and two partial rows of punctures; mesocorial disc almost evenly punctate; exocorium narrow, weakly punctate; costa narrow, slightly convex, somewhat flattened basally, almost entirely separated from exocorium, its margin with two setigerous punctures bearing long hair­like setae (Fig. 302). Hind femur with well visible tubercles on its ventral margin, and large subapical tooth on dorsal margin (Fig. 304); hind tibia with a row of subbasal denticles (Fig. 304). Abdominal sterna punctate laterally, around spiracles and trichobothria. Male genitalia as in Figs 305 ­ 306.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B744559FE3E44D0BA36FCB7.taxon	materials_examined	TYPE DATA Holotype male: Madagascar, Tamatave prov. Moramanga env. (USMB). MATERIAL EXAMINED Holotype.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B744559FE3E44D0BA36FCB7.taxon	distribution	DISTRIBUTION Madagascar.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7A455BFE4D4330BACBFCD7.taxon	description	(Figs 307 ­ 313)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7A455BFE4D4330BACBFCD7.taxon	description	DESCRIPTION Body 9.14 ­ 10.98 mm in length, 4.85 ­ 5.75 mm in width, from pale brown to black­brown. Head dorsally impunctate, except for three primary setigerous punctures, among them a single preocular, clypeus without a subapical pair of hair­like setae (Fig. 308); ocular index 1.98 ­ 2.69; interocellar index 6.3 ­ 8.0; length of antennal segments (in mm): 0.42 ­ 0.52: 0.56 ­ 0.68: 0.73 ­ 0.80: 1.02 ­ 1.11: 1.09 ­ 1.12. Pronotum with discal transverse impression behind calli abruptly marked, bearing a row of punctures; lateral margins of pronotum with 4 submarginal setigerous punctures (Fig. 307); anterior lobe subapically and laterally densely punctate; posterior lobe distinctly punctate with exception of umbones and posterior margin. Propleuron distinctly punctate; depression and posterior convexity with large coarse punctures, anterior convexity with smaller punctures. Scutellum densely punctate, anterolateral angles and apex impunctate. Corium with punctures a little smaller than those on pronotum; clavus with one complete and two partial rows of punctures; mesocorium almost evenly punctate; exocorium very densely punctate; costa narrow, entirely separated from exocorium, bearing 2 setigerous punctures (Fig. 307). Male hind femora with distinct spines on margins, male hind tibiae with distinct carina (Fig. 309); female hind femora with clearly visible subapical spine, female hind tibiae simple. Abdominal sterna glossy in the middle, laterally densely wrinkled and bearing very small, almost indistinct punctures between wrinkles. Male genitalia as in Figs 310 ­ 313.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7A455BFE4D4330BACBFCD7.taxon	materials_examined	TYPE DATA Holotype male: Burma, Tenasserim, Kawkareet (ISNB). MATERIAL EXAMINED Holotype, and 8 other specimens: India: Indes Or., 1 ex. (MNHN); Assam, Kazirachi res., 2 exx. (ZIP, UO); Kalimpong, 1 ex. (ZIP). Burma: Katha, 1 ex. (MCSN); Schwego­Myo, 1 ex. (UO); Tenasserim, Kawkareet, 1 ex. (MCSN); Palon, Pegu, 1 ex. (ZIP).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7A455BFE4D4330BACBFCD7.taxon	distribution	DISTRIBUTION Burma, India.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B78455DFE574311B94AFA56.taxon	description	(Figs 314 ­ 320)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B78455DFE574311B94AFA56.taxon	description	DESCRIPTION Body 7.35 ­ 10.88 mm in length, 3.92 ­ 5.74 mm in width, from dark brown to blackish brown, posterior margin and umbones of pronotum intensely yellowish brown or milky yellow, corium paler than pronotum and scutellum, sometimes clavus and exocorium yellowish brown or ochraceous. Head dorsally with hardly visible tiny punctures, usually wrinkled, sometimes with single large punctures in lateral parts of paraclypei; clypeus without a subapical pair of hair­like setae, each paraclypeus submarginally with a single preocular setigerous puncture (Fig. 315); ocular index 2.15 ­ 2.90; interocellar index 4.2 ­ 6.2; length of antennal segments (in mm): 0.38 ­ 0.49: 0.42 ­ 0.66: 0.50 ­ 0.87: 0.60 ­ 1.03. 0.80 ­ 1.12. Pronotal disc clearly punctate, transverse discal impression behind calli absent; lateral margins with 5 ­ 6 submarginal setigerous punctures (Fig. 314). Propleural depression and basal half of posterior convexity with numerous large punctures, anterior convexity dulled by numerous small punctures and wrinkles. Scutellum almost evenly punctate, basal part and apex impunctate. Clavus with one complete and one or two partial row of dark punctures; mesocorial disc and exocorium evenly punctate; costa narrow, entirely separated from exocorium, bearing 2 (rarely 3) setigerous punctures (Fig. 314). Male hind femora with distinct subapical tooth on dorsal margin, and more or less distinct subbasal tooth on ventral margin, hind tibiae without tubercles (Fig. 316); female hind femora with more or less visible subapical tooth on dorsal margin, hind tibiae without tubercles. Abdominal sterna glossy in the middle, lateral parts wrinkled and punctate around spiracles. Male genitalia as in Figs 317 ­ 320.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B78455DFE574311B94AFA56.taxon	materials_examined	TYPE DATA Lectotype female (designated by LIS, 1991 a: 187) of Cydnus transversus BURMEISTER: Philippines, Manila (MNHU). Lectotype male (designated by LIS, 1994 a: 232) of Acatalectus transversus WALKER: Indonesia, Aru (BMNH). MATERIAL EXAMINED Lectotype of Cydnus transversus, Lectotype and paralectotype of Acatalectus transversus, and 44 other specimens: Indonesia: Celebes, 1 paralectotype (BMNH); Java, Banjoewangi, 1 ex. (ZIP); Java, Soekaboemi, 1 ex. (NNML); Sulawesi Utara, Dumoga­Bone Nat. Park, 6 exx. (IRSNB, UO); N. Sulawesi, Dumoga Bone NP, Malibagu Rd., 1 ex. (NNML); Sulawesi Tenggara, S. Sanggona, Foot of Gn Watuwila, Mokowu river, 1 ex. (NNML); C. Sulawesi, Palu Region, Tawaeli ­ Tomboli rd (km 34), 1 ex. (NNML); Sulawesi Tengah, Nr. Morowali, Ranu River Area, 19 exx. (BMNH, UO). Philippines: Surigao, Mindanao, 1 ex. (ZMA); Leyte, 2 exx. (ZMA, UO); Davao, Mindanao, 1 ex. (UO); Luzon, Manila, 1 ex. (BMNH); Negro lsl., 3 exx. (AMNH, UO); Philippines, Trident Mines, nr Narwan, C. Palawan lsl., 3 exx. (NSMT, UO); Baracatan, Mindanao, 1 ex. (NSMT); Mindoro, 20 km W Calapan, Hidden Parad., 2 ex. (NHMW).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B78455DFE574311B94AFA56.taxon	distribution	DISTRIBUTION Bismarck Archipelago (New Ireland), Indonesia (Aru, Java, Moluccas, Sulawesi, West Irian), Papua New Guinea, Philippines. Reported also from China, Sri Lanka and Japan, but all those records were based on misidentifications (see LIS 1994 a). Records from Australia related to M. bisetosus (see LIS 1999 a).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7E455CFE324587BA80FAB5.taxon	description	(Figs 321 ­ 327)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7E455CFE324587BA80FAB5.taxon	description	DESCRIPTION Body 10.34 mm in length, 5.94 mm in width, blackish brown, umbones of pronotum yellowish brown, posterior pronotal margin castaneous, corium brown. Head dorsally slightly wrinkled and bearing numerous hardly visible tiny punctures; clypeus subapically without setigerous punctures, each paraclypeus submarginally with 3 setigerous punctures bearing long hair­like setae (Fig. 322); ocular index 2.66; interocellar index 3.5; length of antennal segments (in mm): 0.56: 0.76: 0.67: 0.90: 1.07. Puncturation of pronotal disc weak, punctures irregularly scattered; transverse discal impression behind calli absent; lateral margins of pronotum with 5 ­ 7 submarginal setigerous punctures bearing long hair­like setae (Fig. 321). Anterior convexity of propleuron densely punctate with small punctures, posterior convexity with large scattered punctures, propleural depression densely and coarsely punctate. Scutellar disc medially with coarse punctures, its basal and apical thirds impunctate. Clavus with one complete and two incomplete rows of punctures; mesocorium weakly punctate; exocorium densely punctate, punctures larger than those on mesocorial disc; costa broad, basally flattened, separated from exocorium in its basal three fourths, bearing a single setigerous puncture (Fig. 321). Femora bicoloured, yellow or yellowish brown with broad dark median stripe; hind femur with a row of tubercles and denticles (Fig. 323); hind tibia with small subbasal denticles (Fig. 323). Abdominal sterna punctate only close to lateral margins, and around spiracles and trichobothria. Male genitalia as in Figs 324 ­ 327.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7E455CFE324587BA80FAB5.taxon	materials_examined	TYPE DATA Holotype male: New Guinea NE, Wau (BMH). MATERIAL EXAMINED Holotype.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7E455CFE324587BA80FAB5.taxon	distribution	DISTRIBUTION Papua New Guinea.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7F455EFE254527B84DFAB9.taxon	description	(Figs 328 ­ 334)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7F455EFE254527B84DFAB9.taxon	description	DESCRIPTION Body 6.85 ­ 7.81 mm in length, 3.68 ­ 4.05 mm in width, from brown to almost black; hemelytra, umbones and posterior margin of pronotum lighter in shade (from yellowish brown to brown). Head dorsally at first glance looking impunctate, but bearing numerous almost indistinct punctures, lateral parts of paraclypei slightly wrinkled; clypeus subapically without a pair of setigerous punctures, each paraclypeus bearing submarginally a single preocular setigerous puncture (Fig. 329); ocular index 2.00 ­ 2.40; interocellar index 5.5 ­ 8.3; length of antennal segments (in mm): 0.30 ­ 0.40: 0.38 ­ 0.47: 0.49 ­ 0.53: 0.57 ­ 0.67: 0.71 ­ 0.78. Pronotal transverse impression behind calli faintly developed (sometimes almost indistinct), bearing medially interrupted row of punctures; anterior lobe with distinct punctures forming subapical transverse band, and with irregular patch of punctures on either side; posterior lobe with several scattered punctures; lateral margins with 5 ­ 6 submarginal setigerous punctures (Fig. 328). Propleural depression with coarse punctures. Scutellar disc with numerous coarse punctures; apex and base impunctate. Clavus with one complete and two partial rows of punctures; mesocorial disc densely evenly punctate; exocorium sparsely punctate; costa narrow, almost entirely separated from exocorium, bearing 2 ­ 3 setigerous punctures (Fig. 328). Male hind femora with several small teeth on ventral margin, and one distinct subapical tooth on dorsal margin, hind tibiae with small tubercles (Fig. 330); female hind femora with small subapical tooth on dorsal margin, hind tibiae without tubercles. Abdominal sterna weakly, irregularly wrinkled; each segment with a triangular patch of dense fine punctures posterior to spiracles. Male genitalia as in Figs 331 ­ 334.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7F455EFE254527B84DFAB9.taxon	materials_examined	TYPE DATA Holotype male: Indonesia, Sulawesi Utara, Dumoga­Bone National Park (ISNB). MATERIAL EXAMINED Holotype, 3 paratypes and 9 other specimens: Indonesia: Sulawesi Utara, Dumoga­Bone Nat. Park, 3 paratypes (IRSNB, UO); Sulawesi, Minahassa, Tomohon, 1 ex. (BMNH): Sulawesi Tengah, NW of Luwuk, Hanga­hanga, 1 ex. (ZMA); N. Sulawesi, Dumoga Bone NP, Malibagu Rd., 1 ex. (NNML); Iles Moluques, Amboine, 1 ex. (IRSNB); Luksag, Peleng Is., 3 exx. (NSMT, UO); E. Kalimantan, Balikpapan, 2 exx. (NSMT).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7F455EFE254527B84DFAB9.taxon	distribution	DISTRIBUTION Indonesia (Borneo, Sulawesi, Ambon, Peleng).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7D45A0FE4B453BB805FAB6.taxon	description	(Figs 335 ­ 341)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7D45A0FE4B453BB805FAB6.taxon	description	DESCRIPTION Body 8.86 ­ 10.19 mm in length, 4.55 ­ 5.65 mm in width, from dark castaneous to black, corium slightly paler than scutellum and pronotum. Head dorsally with a few wrinkles and several small almost indistinct irregularly scattered punctures; clypeus subapically without hair­like setae, each paraclypeus with 2 submarginal setigerous punctures (Fig. 336); ocular index 2.10 ­ 2.23; interocellar index 6.6 ­ 10.4; length of antennal segments (in mm): 0.4 ­ 0.45: 0.45 ­ 0.59: 0.70 ­ 0.78: 0.84 ­ 1.05: 0.92 ­ 1.08. Pronotum with transverse impression behind calli distinctly developed, sometimes interrupted medially; puncturation of pronotal disc clearly visible, especially laterally and behind calli; lateral margins with 4 ­ 5 submarginal setigerous punctures (Fig. 335). Propleural depression densely coarsely punctate, posterior convexity of propleuron sparsely punctate, apical part of its anterior convexity with smaller punctures. Scutellum evenly punctate with exception of apex and anterolateral angles; punctures larger than those on pronotum. Clavus with one complete and two partial rows of punctures; mesocorial disc almost evenly punctate, punctures of the same size as those on scutellum; exocorium densely punctate; costa narrow, moderately convex, almost entirely separated from exocorium, bearing 2 setigerous punctures (Fig. 335). Hind male femora with several distinct teeth and tubercles, hind tibiae with distinct subbasal tooth (Fig. 337). Abdominal sterna III­VI glossy in the middle, distinctly punctate in lateral two thirds. Male genitalia as in Figs 338 ­ 341.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7D45A0FE4B453BB805FAB6.taxon	materials_examined	TYPE DATA Holotype male: Vietnam, Saigon (MNHN). MATERIAL EXAMINED Holotype, 8 paratype, and 2 other specimens: Burma: Palon, Pegu, 1 paratype (UO). India: Kalimpong, 1 ex. (ZIP). Laos: Vientiane, 1 paratype, 1 ex. (( BMNH, ZSMC). Thailand: Siam, 1 paratype (MNHN). Vietnam: Cochinchine, 1 paratype (UO); Tonkin, reg. de Hoa­Binh, 1 paratype (MNHN); Vung Tau, 1 paratype (MNHN); Tonkin, 2 paratypes (BMNH, UO).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B7D45A0FE4B453BB805FAB6.taxon	distribution	DISTRIBUTION Burma, India (first record), Laos, Thailand, Vietnam.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B8345A2FE144530BABCF9D6.taxon	description	(Figs 342 ­ 348)	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B8345A2FE144530BABCF9D6.taxon	description	DESCRIPTION Body 5.44 ­ 7.33 mm in length, 3.24 ­ 4.05 mm in width, from dark castaneous to black, corium and posterior margin of pronotum somewhat paler in shade. Head dorsally impunctate, except for setigerous punctures and sometimes a few small punctures laterally; clypeus subapically without setigerous punctures, each paraclypeus submarginally with 3 ­ 5 setigerous punctures bearing long hair­like setae (Fig. 343); ocular index 2.40 ­ 2.85; interocellar index 8.0 ­ 13.0; length of antennal segments (in mm): 0.24 ­ 0.31: 0.32 ­ 0.50: 0.33 ­ 0.47: 0.43 ­ 0.57: 0.50 ­ 0.60. Puncturation of pronotal disc well developed, punctures small, especially clearly visible behind head and calli, the latter impunctate, discal transverse impression behind calli absent; lateral margins of pronotum with 6 ­ 10 submarginal setigerous punctures bearing long hair­like setae (Fig. 342). Anterior convexity of propleuron impunctate or almost impunctate, posterior convexity distinctly coarsely punctate close to propleural depression, the latter coarsely punctate. Scutellum with almost evenly spaced punctures as large as those on pronotum, basal fifth of scutellar disc impunctate. Corium densely punctate, punctures darker and larger than those on scutellum; clavus with one complete and two partial rows of punctures; mesocorial disc densely and almost evenly punctate; exocorium densely punctate; costa narrow, entirely separated from exocorium, bearing 2 ­ 4 setigerous punctures (Fig. 342). Male hind femur with subbasal tubercles and large subapical toth on its dorsal margin (Fig. 344); male hind tibia with a row of small denticles (Fig. 344); female hind femora with a row of setae on margins, female tibiae simple. Abdominal sterna more or less polished, impunctate or with a few punctures posterior to spiracles. Male genitalia as in Figs 345 ­ 348.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B8345A2FE144530BABCF9D6.taxon	materials_examined	TYPE DATA Holotype male: Comoros, Grande Comore, N ̕ Tsoudjini (MNHN). MATERIAL EXAMINED Holotype, and 18 paratypes: Comoros: Grande Comore, N ̕ Tsoudjini, 3 paratypes (MNHN, UO); Grande Comore, 1 paratype (MNHN); Moroni, Gde Comore, Malet, 1 paratype (MNHN); Djouma­Dougna, Mohéli, 2 paratypes (MNHN, UO); Mohéli, Bandamalé, 1 paratype (MNHN); Mayotte, Convalescense, 1 paratype (MNHN). Madagascar: 3 paratypes (MNHN, UO); Madagascar Est, district de Maroantsetra, station forestiére de Farankaraina, toute de Navana, km 16.5, vallée d ̕ Antoroka, 3 paratypes (MNHN, UO); Baie d ̕ Antongil, 1 paratype (MNHN); Région du Sud­Est, Fort­Dauphin, 1 paratype (MNHN); Mora Mora, Tulear, 1 paratype (MZLU).	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
03E487C87B8345A2FE144530BABCF9D6.taxon	distribution	DISTRIBUTION Comoros, Madagascar.	en	Lis, J. A. (2000): A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae). Genus (Wrocław) 11 (3), No. 359 - 509: 459-509
